180 lines
20 KiB
XML
180 lines
20 KiB
XML
<document id="AD61969878915F648186871A949C7D1C" ID-DOI="10.11646/zootaxa.5002.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5122655" ID-ZooBank="5DC2A66A-3F04-42D0-8A8C-F0686054E556" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626888359662" checkinUser="plazi" docAuthor="Motamedinia, Behnam, Skevington, Jeffrey H. & Kelso, Scott" docDate="2021" docId="03A887D05F0AFFC3FF3EEC33FD024E54" docLanguage="en" docName="zootaxa.5002.1.1.pdf" docOrigin="Zootaxa 5002 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Tomosvaryella teligera Kuznetzov 1994" docType="treatment" docVersion="3" lastPageNumber="75" masterDocId="FF91FFA85F42FF89FFA9EB42FFAF4F30" masterDocTitle="Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species" masterLastPageNumber="103" masterPageNumber="1" pageNumber="73" updateTime="1699012478263" updateUser="plazi">
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<mods:title id="5501CBFE33542E3F8C658E350DFD9827">Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species</mods:title>
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<mods:name id="52520BB9FF94046A8F87F4BD70C1DE81" type="personal">
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<mods:roleTerm id="81D0C55E1552E461AF87E5B27E389978">Author</mods:roleTerm>
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<mods:namePart id="FFEDBF361F1D9F1D1F5CC41BB768FBDE">Motamedinia, Behnam</mods:namePart>
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<mods:roleTerm id="69405C2A782082A1B65B39B6156A30F8">Author</mods:roleTerm>
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<mods:namePart id="F02137785A00FE1D1806860C15E21942">Skevington, Jeffrey H.</mods:namePart>
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<mods:nameIdentifier id="6F5003F4DFDB997B3793D17F9DBE4D22" type="email">jhskevington@gmail.com</mods:nameIdentifier>
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<mods:name id="9722C47D18626109F3EEE386515561DD" type="personal">
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<mods:roleTerm id="6955E1600226E26FBA27EC9C631FFE68">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="07DC2E839E3F642F1FAAE04F3818BF5D">Kelso, Scott</mods:namePart>
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<mods:nameIdentifier id="AC78F35E506A0D2C06C10BAF17E80926" type="email">scott.kelso@canada.ca</mods:nameIdentifier>
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<mods:typeOfResource id="57FAE7AEADF9120EF3D0307B2966DCE8">text</mods:typeOfResource>
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<mods:title id="DD44ED9D373C5F8ADA3C9F1B87D87150">Zootaxa</mods:title>
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<mods:part id="08136F86C5E34F98DE714E4CA408F18F">
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<mods:date id="BDF0C93874962704F816A6DB19D40D67">2021</mods:date>
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<mods:number id="1EBDD2369B5EA24CE4C46B9319159582">2021-07-16</mods:number>
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<mods:number id="64915982EB4A5F221B416529D47CE393">5002</mods:number>
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<mods:identifier id="89573F1C41E653CC7C21FCBA492B18DC" type="DOI">10.11646/zootaxa.5002.1.1</mods:identifier>
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<mods:identifier id="BA41F30B827E82C0F2F9AC7AE6EBE97C" type="ISSN">1175-5326</mods:identifier>
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<subSubSection id="C31B654D5F0AFFC1FF3EEC33FDD148BC" box="[151,638,1905,1932]" pageId="72" pageNumber="73" type="nomenclature">
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<paragraph id="8BBE36C65F0AFFC1FF3EEC33FDD148BC" blockId="72.[151,638,1905,1968]" box="[151,638,1905,1932]" pageId="72" pageNumber="73">
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<heading id="D0F681AA5F0AFFC1FF3EEC33FDD148BC" bold="true" box="[151,638,1905,1932]" fontSize="11" level="2" pageId="72" pageNumber="73" reason="4">
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<taxonomicName id="4C014D455F0AFFC1FF3EEC33FDD148BC" ID-CoL="7CH72" authority="Kuznetzov, 1994" authorityName="Kuznetzov" authorityYear="1994" box="[151,638,1905,1932]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="teligera">
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<emphasis id="B975EAD45F0AFFC1FF3EEC33FDD148BC" bold="true" box="[151,638,1905,1932]" pageId="72" pageNumber="73">
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<emphasis id="B975EAD45F0AFFC1FF3EEC33FE0748BC" bold="true" box="[151,424,1905,1932]" italics="true" pageId="72" pageNumber="73">Tomosvaryella teligera</emphasis>
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<bibRefCitation id="EF904B375F0AFFC1FE19EC30FDD148BC" author="Kuznetzov, S. Y." box="[432,638,1906,1932]" pageId="72" pageNumber="73" pagination="107 - 207" refId="ref57481" refString="Kuznetzov, S. Y. (1994) New Palaearctic species of Tomosvaryella Aczel (Diptera, Pipunculidae) with a redescription of T. rondanii (Collin). Dipterological Research, 5, 107 - 207." type="journal article" year="1994">Kuznetzov, 1994</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31B654D5F0AFFC1FF3EECD4FE2B4880" box="[151,388,1942,1968]" pageId="72" pageNumber="73" type="description">
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<paragraph id="8BBE36C65F0AFFC1FF3EECD4FE2B4880" blockId="72.[151,638,1905,1968]" box="[151,388,1942,1968]" pageId="72" pageNumber="73">
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<figureCitation id="133A2A435F0AFFC1FF3EECD4FE8F4880" box="[151,288,1942,1968]" captionStart="FIGURE 41" captionStartId="73.[151,250,1890,1914]" captionTargetBox="[166,1436,275,1866]" captionTargetId="figure-61@73.[151,1436,273,1866]" captionTargetPageId="73" captionText="FIGURE 41. Male genitalia of Tomosvaryella teligera (JSS51936, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122755" httpUri="https://zenodo.org/record/5122755/files/figure.png" pageId="72" pageNumber="73">Figs 41A‒E</figureCitation>
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,
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<figureCitation id="133A2A435F0AFFC1FE85ECD4FEF24880" box="[300,349,1942,1968]" captionStart="FIGURE 48" captionStartId="82.[151,250,1173,1197]" captionTargetBox="[151,1436,181,1147]" captionTargetId="figure-31@82.[151,1436,181,1149]" captionTargetPageId="82" captionText="FIGURE 48. Compound eye of Tomosvaryella species in lateral view A) T. debruyni (JSS51736, TAU), B) T. unicorna sp. nov., holotype (JSS51821, CNC), C) T. teligera (JSS51938, CNC)." figureDoi="http://doi.org/10.5281/zenodo.5122767" httpUri="https://zenodo.org/record/5122767/files/figure.png" pageId="72" pageNumber="73">48C</figureCitation>
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,
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<figureCitation id="133A2A435F0AFFC1FEC1ECD4FE2B4880" box="[360,388,1942,1968]" captionStart="FIGURE 64" captionStartId="92.[151,250,1598,1622]" captionTargetBox="[151,1436,932,1574]" captionTargetId="figure-58@92.[151,1436,932,1574]" captionTargetPageId="92" captionText="FIGURE 64. Distribution of Tomosvaryella subvirescens, T. teligera, T. transvaalensis, T. vicina in the Middle East." figureDoi="http://doi.org/10.5281/zenodo.5122799" httpUri="https://zenodo.org/record/5122799/files/figure.png" pageId="72" pageNumber="73">64</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31B654D5F0AFFC3FF3EEC9CFD024E54" lastPageId="74" lastPageNumber="75" pageId="72" pageNumber="73" type="diagnosis">
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<paragraph id="8BBE36C65F0AFFC0FF3EEC9CFE044FC9" blockId="72.[151,1436,2014,2040]" lastBlockId="73.[151,1437,151,249]" lastPageId="73" lastPageNumber="74" pageId="72" pageNumber="73">
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<emphasis id="B975EAD45F0AFFC1FF3EEC9CFEBB48C8" bold="true" box="[151,276,2014,2040]" pageId="72" pageNumber="73">Diagnosis:</emphasis>
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This species can be recognized by the distinct frontal prominence (
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<figureCitation id="133A2A435F0AFFC1FBAAEC9CFBC548C8" box="[1027,1130,2014,2040]" captionStart="FIGURE 48" captionStartId="82.[151,250,1173,1197]" captionTargetBox="[151,1436,181,1147]" captionTargetId="figure-31@82.[151,1436,181,1149]" captionTargetPageId="82" captionText="FIGURE 48. Compound eye of Tomosvaryella species in lateral view A) T. debruyni (JSS51736, TAU), B) T. unicorna sp. nov., holotype (JSS51821, CNC), C) T. teligera (JSS51938, CNC)." figureDoi="http://doi.org/10.5281/zenodo.5122767" httpUri="https://zenodo.org/record/5122767/files/figure.png" pageId="72" pageNumber="73">Fig. 48C</figureCitation>
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); shape of surstyli in dorsal view, both broadened in basal half, moderately narrowed to apical half, curved to each other at apex; left surstylus slightly longer than right one (
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<figureCitation id="133A2A435F0BFFC0FE47EBF9FDF84FE5" box="[494,599,187,213]" captionStart="FIGURE 41" captionStartId="73.[151,250,1890,1914]" captionTargetBox="[166,1436,275,1866]" captionTargetId="figure-61@73.[151,1436,273,1866]" captionTargetPageId="73" captionText="FIGURE 41. Male genitalia of Tomosvaryella teligera (JSS51936, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122755" httpUri="https://zenodo.org/record/5122755/files/figure.png" pageId="73" pageNumber="74">Fig. 41A</figureCitation>
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), gonopods slightly equal in height, right with rounded projection towards phallic guide (
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<figureCitation id="133A2A435F0BFFC0FE91EB9DFE304FC9" box="[312,415,223,249]" captionStart="FIGURE 41" captionStartId="73.[151,250,1890,1914]" captionTargetBox="[166,1436,275,1866]" captionTargetId="figure-61@73.[151,1436,273,1866]" captionTargetPageId="73" captionText="FIGURE 41. Male genitalia of Tomosvaryella teligera (JSS51936, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5122755" httpUri="https://zenodo.org/record/5122755/files/figure.png" pageId="73" pageNumber="74">Fig. 41B</figureCitation>
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).
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</paragraph>
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<caption id="DF7E664E5F0BFFC0FF3EEC20FE3B48AE" ID-DOI="http://doi.org/10.5281/zenodo.5122755" ID-Zenodo-Dep="5122755" httpUri="https://zenodo.org/record/5122755/files/figure.png" pageId="73" pageNumber="74" startId="73.[151,250,1890,1914]" targetBox="[166,1436,275,1866]" targetPageId="73">
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<paragraph id="8BBE36C65F0BFFC0FF3EEC20FE3B48AE" blockId="73.[151,1436,1890,1950]" pageId="73" pageNumber="74">
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<emphasis id="B975EAD45F0BFFC0FF3EEC20FE8F484A" bold="true" box="[151,288,1890,1914]" pageId="73" pageNumber="74">FIGURE 41.</emphasis>
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Male genitalia of
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<taxonomicName id="4C014D455F0BFFC0FE75EC20FD69484A" authorityName="Kuznetzov" authorityYear="1994" box="[476,710,1890,1914]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="teligera">
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<emphasis id="B975EAD45F0BFFC0FE75EC20FD69484A" box="[476,710,1890,1914]" italics="true" pageId="73" pageNumber="74">Tomosvaryella teligera</emphasis>
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</taxonomicName>
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(JSS51936, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view.
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</paragraph>
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</caption>
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<paragraph id="8BBE36C65F08FFC3FF6EEBD4FA324F81" blockId="74.[151,1437,150,357]" box="[199,1437,150,177]" pageId="74" pageNumber="75">
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<emphasis id="B975EAD45F08FFC3FF6EEBD4FE654F80" bold="true" box="[199,458,150,176]" pageId="74" pageNumber="75">Specimens examined:</emphasis>
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<materialsCitation id="3B693C9B5F08FFC3FE7BEBD4FADF4F81" box="[466,1392,150,177]" collectingDate="2016-07-05" country="Iran" elevation="1011" latitude="35.0" location="Dodan" longLatPrecision="1196" longitude="46.2" pageId="74" pageNumber="75" specimenCount="1" stateProvince="Kermanshah">
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<collectingCountry id="F31676565F08FFC3FE7BEBD4FDB44F80" box="[466,539,150,176]" name="Iran" pageId="74" pageNumber="75">IRAN</collectingCountry>
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:
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<collectingRegion id="49C5F8245F08FFC3FD8FEBD4FD174F80" box="[550,696,150,176]" country="Iran" name="Kermanshah" pageId="74" pageNumber="75">Kermanshah</collectingRegion>
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:
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<location id="8EDE601D5F08FFC3FD6DEBD4FCBD4F80" LSID="urn:lsid:plazi:treatment:03A887D05F0AFFC3FF3EEC33FD024E54:8EDE601D5F08FFC3FD6DEBD4FCBD4F80" box="[708,786,150,176]" country="Iran" latitude="35.0" longLatPrecision="1196" longitude="46.2" name="Dodan" pageId="74" pageNumber="75" stateProvince="Kermanshah">Dodan</location>
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,
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<geoCoordinate id="EE3550015F08FFC3FCB4EBD4FC2F4F81" box="[797,896,150,177]" degrees="35" direction="north" minutes="00" orientation="latitude" pageId="74" pageNumber="75" precision="925" value="35.0">35°00’N</geoCoordinate>
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,
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<geoCoordinate id="EE3550015F08FFC3FC24EBD4FC424F80" box="[909,1005,150,176]" degrees="46" direction="east" minutes="12" orientation="longitude" pageId="74" pageNumber="75" precision="925" value="46.2">46°12’E</geoCoordinate>
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,
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<elevation id="002CD1F55F08FFC3FC53EBD4FB224F80" box="[1018,1165,150,177]" metricMagnitude="3" metricUnit="m" metricValue="1.011" pageId="74" pageNumber="75" unit="m" value="1011.0">
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<quantity id="4CF99B235F08FFC3FC53EBD4FBE04F81" box="[1018,1103,150,177]" metricMagnitude="3" metricUnit="m" metricValue="1.011" pageId="74" pageNumber="75" unit="m" value="1011.0">1011 m</quantity>
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a.s.l.
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</elevation>
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, Garden,
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<date id="FFBF10065F08FFC3FB55EBD4FADF4F81" box="[1276,1392,150,177]" pageId="74" pageNumber="75" value="2016-07-05">
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<collectingDate id="EFFBE9EE5F08FFC3FB55EBD4FADF4F81" box="[1276,1392,150,177]" pageId="74" pageNumber="75" value="2016-07-05">5.vii.2016</collectingDate>
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</date>
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||
</materialsCitation>
|
||
, M.
|
||
</paragraph>
|
||
<paragraph id="8BBE36C65F08FFC3FF3EEBF8FA324FE5" blockId="74.[151,1437,150,357]" box="[151,1437,186,213]" pageId="74" pageNumber="75">
|
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<materialsCitation id="3B693C9B5F08FFC3FF3EEBF8FADE4FE5" accessionNumber="JSS51938" box="[151,1393,186,213]" collectingDate="2015-05-14" collectionCode="CNC" elevation="456" latitude="34.466667" longLatPrecision="1199" longitude="45.816666" pageId="74" pageNumber="75" specimenCount="1" specimenCount-male="1">
|
||
Zardouei,
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||
<collectingMethod id="52404ED15F08FFC3FEA3EBF8FE494FE5" box="[266,486,186,213]" pageId="74" pageNumber="75">funnel Malaise trap</collectingMethod>
|
||
, JSS51938 (
|
||
<specimenCount id="9D07FD4F5F08FFC3FDDBEBF8FD394FE4" box="[626,662,186,212]" pageId="74" pageNumber="75" type="male">1♂</specimenCount>
|
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,
|
||
<collectionCode id="ED10AE035F08FFC3FD0BEBF8FD4F4FE4" box="[674,736,186,212]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="74" pageNumber="75">CNC</collectionCode>
|
||
); Sarpolezahab,
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||
<geoCoordinate id="EE3550015F08FFC3FC31EBF8FC554FE4" box="[920,1018,186,212]" degrees="34" direction="north" minutes="28" orientation="latitude" pageId="74" pageNumber="75" precision="925" value="34.466667">34°28’N</geoCoordinate>
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,
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<geoCoordinate id="EE3550015F08FFC3FBAFEBF8FBC94FE5" box="[1030,1126,186,213]" degrees="45" direction="east" minutes="49" orientation="longitude" pageId="74" pageNumber="75" precision="925" value="45.816666">45°49’E</geoCoordinate>
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,
|
||
<elevation id="002CD1F55F08FFC3FBD8EBF8FB5B4FE4" box="[1137,1268,186,213]" metricMagnitude="2" metricUnit="m" metricValue="4.56" pageId="74" pageNumber="75" unit="m" value="456.0">
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<quantity id="4CF99B235F08FFC3FBD8EBF8FB174FE5" box="[1137,1208,186,213]" metricMagnitude="2" metricUnit="m" metricValue="4.56" pageId="74" pageNumber="75" unit="m" value="456.0">456 m</quantity>
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||
a.s.l.
|
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</elevation>
|
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,
|
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<date id="FFBF10065F08FFC3FAA9EBF8FADE4FE5" box="[1280,1393,186,213]" pageId="74" pageNumber="75" value="2015-05-14">
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<collectingDate id="EFFBE9EE5F08FFC3FAA9EBF8FADE4FE5" box="[1280,1393,186,213]" pageId="74" pageNumber="75" value="2015-05-14">14.v.2015</collectingDate>
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</date>
|
||
</materialsCitation>
|
||
, M.
|
||
</paragraph>
|
||
<paragraph id="8BBE36C65F08FFC3FF3EEB9CFA334E71" blockId="74.[151,1437,150,357]" pageId="74" pageNumber="75">
|
||
Zardouei, Malaise trap, JSS51936‒7 (
|
||
<specimenCount id="9D07FD4F5F08FFC3FD96EB9CFDCA4FC8" box="[575,613,222,248]" pageId="74" pageNumber="75" type="male">2♂</specimenCount>
|
||
, CNC).
|
||
<emphasis id="B975EAD45F08FFC3FF6EEA40FECC4E2C" bold="true" box="[199,355,258,284]" pageId="74" pageNumber="75">Distribution:</emphasis>
|
||
<collectingCountry id="F31676565F08FFC3FEC3EA41FE354E2D" box="[362,410,259,285]" name="Iran" pageId="74" pageNumber="75">Iran</collectingCountry>
|
||
,
|
||
<collectingCountry id="F31676565F08FFC3FE0CEA40FD874E2C" box="[421,552,258,284]" name="Kazakhstan" pageId="74" pageNumber="75">Kazakhstan</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF904B375F08FFC3FD9FEA40FD5C4E2C" author="Kuznetzov, S. Y." box="[566,755,258,284]" pageId="74" pageNumber="75" pagination="107 - 207" refId="ref57481" refString="Kuznetzov, S. Y. (1994) New Palaearctic species of Tomosvaryella Aczel (Diptera, Pipunculidae) with a redescription of T. rondanii (Collin). Dipterological Research, 5, 107 - 207." type="journal article" year="1994">Kuznetzov 1994</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF904B375F08FFC3FD56EA40FC104E2C" author="Skevington, J. H." box="[767,959,258,284]" pageId="74" pageNumber="75" refId="ref58350" refString="Skevington, J. H. (2020) " Pipunculidae ". Canadian National Collection of Insects, Arachnids and Nematodes database. Available from: https: // cnc. agr. gc. ca / taxonomy / Taxonomy. php? id = 18557 (accessed 18 October 2020) [public version of Pipunculidae catalogue maintained by J. H. Skevington]" type="url" year="2020">Skevington 2020</bibRefCitation>
|
||
) (
|
||
<figureCitation id="133A2A435F08FFC3FC7FEA40FB854E2D" box="[982,1066,258,285]" captionStart="FIGURE 64" captionStartId="92.[151,250,1598,1622]" captionTargetBox="[151,1436,932,1574]" captionTargetId="figure-58@92.[151,1436,932,1574]" captionTargetPageId="92" captionText="FIGURE 64. Distribution of Tomosvaryella subvirescens, T. teligera, T. transvaalensis, T. vicina in the Middle East." figureDoi="http://doi.org/10.5281/zenodo.5122799" httpUri="https://zenodo.org/record/5122799/files/figure.png" pageId="74" pageNumber="75">Fig. 64</figureCitation>
|
||
).
|
||
<emphasis id="B975EAD45F08FFC3FF6EEA64FEA74E70" bold="true" box="[199,264,294,320]" pageId="74" pageNumber="75">Note:</emphasis>
|
||
Based on DNA barcoding, this species is genetically similar to
|
||
<taxonomicName id="4C014D455F08FFC3FC7EEA65FBCA4E70" box="[983,1125,294,320]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="ellipiensis" status="sp. nov.">
|
||
<emphasis id="B975EAD45F08FFC3FC7EEA65FBCA4E70" box="[983,1125,294,320]" italics="true" pageId="74" pageNumber="75">T. ellipiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24657AF5F08FFC3FBC5EA64FB694E71" box="[1132,1222,294,321]" pageId="74" pageNumber="75" rank="species">
|
||
<emphasis id="B975EAD45F08FFC3FBC5EA64FB6E4E70" bold="true" box="[1132,1217,294,320]" pageId="74" pageNumber="75">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, differing by 5.7%
|
||
</paragraph>
|
||
<paragraph id="8BBE36C65F08FFC3FF3EEA08FD024E54" blockId="74.[151,1437,150,357]" box="[151,685,330,357]" pageId="74" pageNumber="75">
|
||
(pairwise divergence) (Supplementary
|
||
<tableCitation id="C683037D5F08FFC3FDE3EA08FD334E54" box="[586,668,330,356]" captionStart="TABLE 1" captionStartId="5.[151,239,810,836]" captionText="TABLE 1. Cytochrome c oxidase subunit I (COI) and internal transcribed spacer (ITS) primers." pageId="74" pageNumber="75">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |