274 lines
35 KiB
XML
274 lines
35 KiB
XML
<document id="AC60CAAD825543A0ADDE7D5D7BAB2BA4" ID-CLB-Dataset="7670" ID-DOI="10.11646/zootaxa.4942.3.8" ID-GBIF-Dataset="0bf4c020-2726-4d2a-aff0-7a5164aae728" ID-ISSN="1175-5326" ID-Zenodo-Dep="4604476" ID-ZooBank="DACB7FDD-75FD-4E10-BF98-E1509B5F2B6A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615798512074" checkinUser="plazi" docAuthor="Rakovič, Miloslav & Anichtchenko, Alexander" docDate="2021" docId="03A887A7FFA9FF83FF1BF8FDFD4CFA07" docLanguage="en" docName="zootaxa.4942.3.8.pdf" docOrigin="Zootaxa 4942 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Odochilus (Parodochilus) shavrini Rakovic & Anichtchenko 2021, new species" docType="treatment" docVersion="8" lastPageNumber="449" masterDocId="FF91FFDFFFA9FF81FF8CFFD6FF9BFF9A" masterDocTitle="Two new species of the genus Odochilus Harold, 1877 (Coleoptera: Scarabaeidae Aphodiinae: Odochilini)" masterLastPageNumber="450" masterPageNumber="447" pageNumber="447" updateTime="1698933438600" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3200B378D00BB6C60C285427E3781C2B">Two new species of the genus Odochilus Harold, 1877 (Coleoptera: Scarabaeidae Aphodiinae: Odochilini)</mods:title>
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<mods:namePart id="FAB14650003BC6529D9E4432842F635C">Rakovič, Miloslav</mods:namePart>
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<mods:namePart id="C5E7C6494BA85B1E222F98B21DE60C24">Anichtchenko, Alexander</mods:namePart>
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<mods:title id="26B341D23733BBF80056321505F0A8D4">Zootaxa</mods:title>
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<mods:part id="F958C4FD31D556273A345885321DBF5B">
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<mods:date id="6561BBC6709FCCEEE8BD69D3C01E21BD">2021</mods:date>
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<mods:number id="C2B473BAD04236BB5A56E6E4D3EE56B2">2021-03-15</mods:number>
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<mods:number id="0525B01BD8D94D2D4264EC9A863CD338">4942</mods:number>
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<treatment id="03A887A7FFA9FF83FF1BF8FDFD4CFA07" ID-DOI="http://doi.org/10.5281/zenodo.4619724" ID-GBIF-Taxon="180591608" ID-Zenodo-Dep="4619724" LSID="urn:lsid:plazi:treatment:03A887A7FFA9FF83FF1BF8FDFD4CFA07" httpUri="http://treatment.plazi.org/id/03A887A7FFA9FF83FF1BF8FDFD4CFA07" lastPageId="2" lastPageNumber="449" pageId="0" pageNumber="447">
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<subSubSection id="C31B653AFFA9FF81FF1BF8FDFB81F8DF" box="[151,1050,1835,1862]" pageId="0" pageNumber="447" type="nomenclature">
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<paragraph id="8BBE36B1FFA9FF81FF1BF8FDFB81F8DF" blockId="0.[151,1050,1835,1897]" box="[151,1050,1835,1862]" pageId="0" pageNumber="447">
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<heading id="D0F681DDFFA9FF81FF1BF8FDFB81F8DF" bold="true" box="[151,1050,1835,1862]" fontSize="11" level="1" pageId="0" pageNumber="447" reason="1">
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<emphasis id="B975EAA3FFA9FF81FF1BF8FDFB81F8DF" bold="true" box="[151,1050,1835,1862]" pageId="0" pageNumber="447">
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<taxonomicName id="4C014D32FFA9FF81FF1BF8FDFCE5F8DF" authority="Rakovic & Anichtchenko" authorityName="Rakovic & Anichtchenko" authorityYear="2021" box="[151,894,1835,1862]" class="Insecta" family="Aphodiidae" genus="Odochilus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="447" phylum="Arthropoda" rank="species" species="shavrini" status="sp. nov." subGenus="Parodochilus">
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<emphasis id="B975EAA3FFA9FF81FF1BF8FDFE88F8DC" bold="true" box="[151,275,1835,1862]" italics="true" pageId="0" pageNumber="447">Odochilus</emphasis>
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(
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<emphasis id="B975EAA3FFA9FF81FEA8F8FDFE5EF8DC" bold="true" box="[292,453,1835,1862]" italics="true" pageId="0" pageNumber="447">Parodochilus</emphasis>
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)
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<emphasis id="B975EAA3FFA9FF81FE5BF8FDFDA1F8DC" bold="true" box="[471,570,1835,1862]" italics="true" pageId="0" pageNumber="447">shavrini</emphasis>
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Rakovič & Anichtchenko
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24657D8FFA9FF81FC07F8FDFB81F8DF" box="[907,1050,1835,1861]" pageId="0" pageNumber="447" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31B653AFFA9FF81FF1BF899FE89F8F3" box="[151,274,1871,1897]" pageId="0" pageNumber="447" type="description">
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<paragraph id="8BBE36B1FFA9FF81FF1BF899FE89F8F3" blockId="0.[151,1050,1835,1897]" box="[151,274,1871,1897]" pageId="0" pageNumber="447">
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(
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<figureCitation id="133A2A34FFA9FF81FF13F899FE92F8F3" box="[159,265,1871,1897]" captionStart="FIGURES 1–6" captionStartId="1.[151,264,1825,1849]" captionTargetBox="[303,1299,179,1799]" captionTargetId="figure-19@1.[288,1299,179,1799]" captionTargetPageId="1" captionText="FIGURES 1–6. Habitus of Odochilus (Parodochilus) species. 1, O. (P.) shavrini Rakovič & Anichtchenko, new species, male (holotype), habitus in dorsal view; 2, O. (P.) shavrini new species, male (holotype), habitus in dorsolateral view; 3, O. (P.) shavrini new species, male (holotype), habitus in ventral view; 4, O. (P.) buruensis Boucomont, 1926, drawing of pronotum overprinted from Rakovič, 1997 (permitted by Acta Societatis Zoologicae Bohemicae); 5, O. (P.) borneensis Rakovič & Anich- tchenko, new species, female (holotype), habitus in dorsal view; 6, O. (P.) jani Bezděk & Křivan, 2001, paratype no. 3, habitus in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4604478" httpUri="https://zenodo.org/record/4604478/files/figure.png" pageId="0" pageNumber="447">Figs. 1–3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31B653AFFA9FF81FF1BF841FF64F863" pageId="0" pageNumber="447" type="materials_examined">
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<paragraph id="8BBE36B1FFA9FF81FF1BF841FF64F863" blockId="0.[151,1437,1943,2041]" pageId="0" pageNumber="447">
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<emphasis id="B975EAA3FFA9FF81FF1BF841FED1F82B" bold="true" box="[151,330,1943,1969]" pageId="0" pageNumber="447">Type material.</emphasis>
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<materialsCitation id="3B693CECFFA9FF81FEDAF841FC28F84F" ID-GBIF-Occurrence="3056489302" collectingDate="2019-05-04" collectionCode="V, DUBC" country="Philippines" county="Candalaga Mts." elevation="700" location="Mindanao" municipality="Marangig Falls" pageId="0" pageNumber="447" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54BA8813FFA9FF81FEDAF841FE5AF82B" box="[342,449,1943,1969]" pageId="0" pageNumber="447" type="holotype">Holotype</typeStatus>
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, male: “
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<collectingCountry id="F3167621FFA9FF81FDA8F841FD33F82B" box="[548,680,1943,1969]" name="Philippines" pageId="0" pageNumber="447">Philippines</collectingCountry>
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,
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<location id="8EDE606AFFA9FF81FD34F841FCB7F82B" LSID="urn:lsid:plazi:treatment:03A887A7FFA9FF83FF1BF8FDFD4CFA07:8EDE606AFFA9FF81FD34F841FCB7F82B" box="[696,812,1943,1969]" country="Philippines" county="Candalaga Mts." municipality="Marangig Falls" name="Mindanao" pageId="0" pageNumber="447">Mindanao</location>
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,
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<location id="8EDE606AFFA9FF81FCB1F84EFC27F828" LSID="urn:lsid:plazi:treatment:03A887A7FFA9FF83FF1BF8FDFD4CFA07:8EDE606AFFA9FF81FCB1F84EFC27F828" box="[829,956,1944,1970]" country="Philippines" county="Candalaga Mts." municipality="Marangig Falls" name="Maragusan" pageId="0" pageNumber="447">Maragusan</location>
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,
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<quantity id="4CF99B54FFA9FF81FC40F841FB94F82B" box="[972,1039,1943,1969]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="0" pageNumber="447" unit="m" value="700.0">
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<elevation id="002CD182FFA9FF81FC40F841FB94F82B" box="[972,1039,1943,1969]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="0" pageNumber="447" unit="m" value="700.0">700m</elevation>
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</quantity>
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,
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<collectingMunicipality id="6BDAACCBFFA9FF81FB92F841FB4BF82B" box="[1054,1232,1943,1969]" pageId="0" pageNumber="447">Marangig Falls</collectingMunicipality>
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,
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<collectingCounty id="62DF4E3DFFA9FF81FB6DF841FA03F828" box="[1249,1432,1943,1970]" pageId="0" pageNumber="447">Candalaga Mts.</collectingCounty>
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, 7˚20’32.2”N 126˚10’24.0”E, 3-
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<date id="FFBF1071FFA9FF81FE7BF86DFDF5F84C" box="[503,622,1979,2006]" pageId="0" pageNumber="447" value="2019-05-04">
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<collectingDate id="EFFBE999FFA9FF81FE7BF86DFDF5F84C" box="[503,622,1979,2006]" pageId="0" pageNumber="447" value="2019-05-04">4.05.2019</collectingDate>
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</date>
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, Shavrin A.
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<collectionCode id="ED10AE74FFA9FF81FD7CF86DFC9DF84F" box="[752,774,1979,2005]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="0" pageNumber="447" type="Museum">V</collectionCode>
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. leg.” (
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<collectionCode id="ED10AE74FFA9FF81FCDBF86DFC30F84F" box="[855,939,1979,2005]" pageId="0" pageNumber="447">DUBC</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="3B693CECFFA9FF81FC32F86AFF60F863" ID-GBIF-Occurrence="3056489301" collectionCode="DUBC" country="Philippines" pageId="0" pageNumber="447" specimenCount="1" typeStatus="paratype">
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<typeStatus id="54BA8813FFA9FF81FC32F86AFBB8F84C" box="[958,1059,1980,2006]" pageId="0" pageNumber="447" type="paratype">Paratype</typeStatus>
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, female, with the same label data (
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<collectionCode id="ED10AE74FFA9FF81FF13F809FF68F863" box="[159,243,2015,2041]" pageId="0" pageNumber="447">DUBC</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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||
</subSubSection>
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<caption id="DF7E6639FFA8FF80FF1BF8F7FEB7F877" ID-DOI="http://doi.org/10.5281/zenodo.4604478" ID-Zenodo-Dep="4604478" httpUri="https://zenodo.org/record/4604478/files/figure.png" pageId="1" pageNumber="448" startId="1.[151,264,1825,1849]" targetBox="[303,1299,179,1799]" targetPageId="1">
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<paragraph id="8BBE36B1FFA8FF80FF1BF8F7FEB7F877" blockId="1.[151,1437,1825,2029]" pageId="1" pageNumber="448">
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<emphasis id="B975EAA3FFA8FF80FF1BF8F7FEA0F8A3" bold="true" box="[151,315,1825,1849]" pageId="1" pageNumber="448">FIGURES 1–6.</emphasis>
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Habitus of
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<taxonomicName id="4C014D32FFA8FF80FE3FF8F7FD21F8A3" authority="" box="[435,698,1825,1849]" class="Insecta" family="Aphodiidae" genus="Odochilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="subGenus" subGenus="Parodochilus">
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<emphasis id="B975EAA3FFA8FF80FE3FF8F7FD80F8A3" box="[435,539,1825,1849]" italics="true" pageId="1" pageNumber="448">Odochilus</emphasis>
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(
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<emphasis id="B975EAA3FFA8FF80FDA6F8F7FD29F8A3" box="[554,690,1825,1849]" italics="true" pageId="1" pageNumber="448">Parodochilus</emphasis>
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)
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</taxonomicName>
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species. 1,
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<taxonomicName id="4C014D32FFA8FF80FCBDF8F7FB41F8A3" authority="Rakovic & Anichtchenko" authorityName="Rakovic & Anichtchenko" authorityYear="2021" box="[817,1242,1825,1849]" class="Insecta" family="Aphodiidae" genus="Odochilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="species" species="shavrini" status="sp. nov." subGenus="Parodochilus">
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<emphasis id="B975EAA3FFA8FF80FCBDF8F7FCD1F8A2" box="[817,842,1825,1848]" italics="true" pageId="1" pageNumber="448">O.</emphasis>
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(
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<emphasis id="B975EAA3FFA8FF80FCD5F8F4FCF6F8A3" box="[857,877,1826,1849]" italics="true" pageId="1" pageNumber="448">P.</emphasis>
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)
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<emphasis id="B975EAA3FFA8FF80FCF7F8F7FC55F8A3" box="[891,974,1825,1849]" italics="true" pageId="1" pageNumber="448">shavrini</emphasis>
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Rakovič & Anichtchenko
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24657D8FFA8FF80FB68F8F7FAFAF8A3" box="[1252,1377,1825,1849]" pageId="1" pageNumber="448" rank="species">new species</taxonomicNameLabel>
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, male (holotype), habitus in dorsal view; 2,
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<taxonomicName id="4C014D32FFA8FF80FDAFF893FD5FF8C7" authority="Rakovič & Anichtchenko, 2021" authorityName="Rakovič & Anichtchenko" authorityYear="2021" box="[547,708,1861,1885]" class="Insecta" family="Aphodiidae" genus="Odochilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="species" species="shavrini" status="sp. nov." subGenus="Parodochilus">
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||
<emphasis id="B975EAA3FFA8FF80FDAFF893FDA7F8C6" box="[547,572,1861,1884]" italics="true" pageId="1" pageNumber="448">O.</emphasis>
|
||
(
|
||
<emphasis id="B975EAA3FFA8FF80FDC1F890FDFBF8C7" box="[589,608,1862,1885]" italics="true" pageId="1" pageNumber="448">P.</emphasis>
|
||
)
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||
<emphasis id="B975EAA3FFA8FF80FDFDF893FD5FF8C7" box="[625,708,1861,1885]" italics="true" pageId="1" pageNumber="448">shavrini</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A24657D8FFA8FF80FD40F893FCD7F8C7" box="[716,844,1861,1885]" pageId="1" pageNumber="448" rank="species">new species</taxonomicNameLabel>
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, male (holotype), habitus in dorsolateral view; 3,
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<taxonomicName id="4C014D32FFA8FF80FADBF893FF71F81B" authority="Rakovič & Anichtchenko, 2021" authorityName="Rakovič & Anichtchenko" authorityYear="2021" class="Insecta" family="Aphodiidae" genus="Odochilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="species" species="shavrini" status="sp. nov." subGenus="Parodochilus">
|
||
<emphasis id="B975EAA3FFA8FF80FADBF893FAEBF8C6" box="[1367,1392,1861,1884]" italics="true" pageId="1" pageNumber="448">O.</emphasis>
|
||
(
|
||
<emphasis id="B975EAA3FFA8FF80FA0DF890FA0EF8C7" box="[1409,1429,1862,1885]" italics="true" pageId="1" pageNumber="448">P.</emphasis>
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||
)
|
||
<emphasis id="B975EAA3FFA8FF80FF1BF8BFFF71F81B" box="[151,234,1897,1921]" italics="true" pageId="1" pageNumber="448">shavrini</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A24657D8FFA8FF80FF7EF8BFFEEAF81B" box="[242,369,1897,1921]" pageId="1" pageNumber="448" rank="species">new species</taxonomicNameLabel>
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||
, male (holotype), habitus in ventral view; 4,
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<taxonomicName id="4C014D32FFA8FF80FCC9F8BFFB2EF81B" authority="Boucomont, 1926" authorityName="Boucomont" authorityYear="1926" box="[837,1205,1897,1921]" class="Insecta" family="Aphodiidae" genus="Odochilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="species" species="buruensis" subGenus="Parodochilus">
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||
<emphasis id="B975EAA3FFA8FF80FCC9F8BFFCC5F81A" box="[837,862,1897,1920]" italics="true" pageId="1" pageNumber="448">O.</emphasis>
|
||
(
|
||
<emphasis id="B975EAA3FFA8FF80FCE2F8BCFC19F81B" box="[878,898,1898,1921]" italics="true" pageId="1" pageNumber="448">P.</emphasis>
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||
)
|
||
<emphasis id="B975EAA3FFA8FF80FC1DF8BFFC6FF81B" box="[913,1012,1897,1921]" italics="true" pageId="1" pageNumber="448">buruensis</emphasis>
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<bibRefCitation id="EF904B40FFA8FF80FC70F8BFFB2EF81B" author="Boucomont, A." box="[1020,1205,1897,1921]" pageId="1" pageNumber="448" pagination="110" refId="ref2276" refString="Boucomont, A. (1926) Fauna Buruana. Coleoptera, Fam. Lamellicornia Coprophaga. Treubia, 7, 110." type="journal article" year="1926">Boucomont, 1926</bibRefCitation>
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</taxonomicName>
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||
, drawing of pronotum overprinted from
|
||
<bibRefCitation id="EF904B40FFA8FF80FEC5F85BFE46F83F" author="Rakovic, M." box="[329,477,1933,1957]" pageId="1" pageNumber="448" pagination="157 - 167" refId="ref2497" refString="Rakovic, M. (1997) A review of the genus Odochilus (Coleoptera: Aphodiidae: Odochilinae). Acta Societatis Zoologicae Bohemicae, 61, 157 - 167." type="journal article" year="1997">Rakovič, 1997</bibRefCitation>
|
||
(permitted by Acta Societatis Zoologicae Bohemicae); 5,
|
||
<taxonomicName id="4C014D32FFA8FF80FBABF85BFAACF83F" authority="Rakovic" authorityName="Rakovic & Anichtchenko" authorityYear="2021" box="[1063,1335,1933,1957]" class="Insecta" family="Aphodiidae" genus="Odochilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="species" species="borneensis" subGenus="Parodochilus">
|
||
<emphasis id="B975EAA3FFA8FF80FBABF85BFBDBF83E" box="[1063,1088,1933,1956]" italics="true" pageId="1" pageNumber="448">O.</emphasis>
|
||
(
|
||
<emphasis id="B975EAA3FFA8FF80FBC2F858FBF9F83F" box="[1102,1122,1934,1957]" italics="true" pageId="1" pageNumber="448">P.</emphasis>
|
||
)
|
||
<emphasis id="B975EAA3FFA8FF80FBE3F85BFB46F83F" box="[1135,1245,1933,1957]" italics="true" pageId="1" pageNumber="448">borneensis</emphasis>
|
||
Rakovič
|
||
</taxonomicName>
|
||
& Anichtchenko, new species, female (holotype), habitus in dorsal view; 6,
|
||
<taxonomicName id="4C014D32FFA8FF80FCCDF867FB37F853" authority="Bezdek & Krivan, 2001" authorityName="Bezdek & Krivan" authorityYear="2001" box="[833,1196,1969,1993]" class="Insecta" family="Aphodiidae" genus="Odochilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="species" species="jani" subGenus="Parodochilus">
|
||
<emphasis id="B975EAA3FFA8FF80FCCDF867FCC1F852" box="[833,858,1969,1992]" italics="true" pageId="1" pageNumber="448">O.</emphasis>
|
||
(
|
||
<emphasis id="B975EAA3FFA8FF80FCE5F864FCE7F853" box="[873,892,1970,1993]" italics="true" pageId="1" pageNumber="448">P.</emphasis>
|
||
)
|
||
<emphasis id="B975EAA3FFA8FF80FC06F864FC2AF853" box="[906,945,1970,1993]" italics="true" pageId="1" pageNumber="448">jani</emphasis>
|
||
<bibRefCitation id="EF904B40FFA8FF80FC34F867FB37F853" author="Bezdek, A. & Krivan, V." box="[952,1196,1969,1993]" pageId="1" pageNumber="448" pagination="65 - 67" refId="ref2232" refString="Bezdek, A. & Krivan, V. (2001) Odochilus (Parodochilus) jani sp. nov. from Sabah (Borneo) (Coleoptera: Scarabaeidae: Apho- diinae). Entomological Problems, 32, 65 - 67." type="journal article" year="2001">Bezděk & Křivan, 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
, paratype no. 3, habitus in dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31B653AFFABFF83FF4BFF40FF4FFBAF" pageId="2" pageNumber="449" type="description">
|
||
<paragraph id="8BBE36B1FFABFF83FF4BFF40FA13FF4F" blockId="2.[151,1437,150,1437]" pageId="2" pageNumber="449">
|
||
<emphasis id="B975EAA3FFABFF83FF4BFF40FD8FFF2B" bold="true" box="[199,532,150,177]" pageId="2" pageNumber="449">
|
||
Description of the
|
||
<typeStatus id="54BA8813FFABFF83FE2AFF41FD94FF2B" box="[422,527,151,177]" pageId="2" pageNumber="449" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Small (length of
|
||
<quantity id="4CF99B54FFABFF83FD50FF41FCACFF2B" box="[732,823,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="449" unit="mm" value="2.8">2.8 mm</quantity>
|
||
), strongly convex (
|
||
<figureCitation id="133A2A34FFABFF83FB9CFF41FBE6FF2B" box="[1040,1149,151,177]" captionStart="FIGURES 1–6" captionStartId="1.[151,264,1825,1849]" captionTargetBox="[303,1299,179,1799]" captionTargetId="figure-19@1.[288,1299,179,1799]" captionTargetPageId="1" captionText="FIGURES 1–6. Habitus of Odochilus (Parodochilus) species. 1, O. (P.) shavrini Rakovič & Anichtchenko, new species, male (holotype), habitus in dorsal view; 2, O. (P.) shavrini new species, male (holotype), habitus in dorsolateral view; 3, O. (P.) shavrini new species, male (holotype), habitus in ventral view; 4, O. (P.) buruensis Boucomont, 1926, drawing of pronotum overprinted from Rakovič, 1997 (permitted by Acta Societatis Zoologicae Bohemicae); 5, O. (P.) borneensis Rakovič & Anich- tchenko, new species, female (holotype), habitus in dorsal view; 6, O. (P.) jani Bezděk & Křivan, 2001, paratype no. 3, habitus in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4604478" httpUri="https://zenodo.org/record/4604478/files/figure.png" pageId="2" pageNumber="449">Figs. 1, 2</figureCitation>
|
||
), remarkably broader behind, length-to-width ratio of 1.98, dark brown, tibiae and tarsi light brown, antennae and mouth parts yellowish.
|
||
</paragraph>
|
||
<paragraph id="8BBE36B1FFABFF83FF4BFF09FC9BFE37" blockId="2.[151,1437,150,1437]" pageId="2" pageNumber="449">Head glabrous from above, except for minute, dust-like setae along ridges above eyes. Genae moderately protruding. Clypeus margin with upper and lower edges: upper edge convex, sharp and distinct, lower edge obsolete and not observable from above. Clypeal disc convex, matte, with rather uneven, corrugately punctate surface; longitudinal ridges on clypeus barely visible. Vertex with two pairs of distinct oblique ridges. Median pair of ridges replaced by a continuous semicircular ridge. Exterior pair forming a characteristic T-shaped structure with ridge extending forward from eyes. Eyes visible from above.</paragraph>
|
||
<paragraph id="8BBE36B1FFABFF83FF4BFE61FC18FD1F" blockId="2.[151,1437,150,1437]" pageId="2" pageNumber="449">Pronotum transverse, 1.5 times as wide as long; opaque, convex, with four transverse ridges. Anterior and lateral margins, and all ridges with very small, dust-like setae. Lateral margins with two sub-angular emarginations and thus, with three moderately protruding, widely angular lobes (ponotum narrowed behind posterior lobe). Basal margin with apically slightly dilated setae. First and second pronotal ridges continuous, sharp, third ridge weakly and narrowly interrupted in the middle; fourth ridge broken into six discrete teeth; lateral longitudinal ridges small, connected with ends of third ridge. Intervals between ridges matte.</paragraph>
|
||
<paragraph id="8BBE36B1FFABFF83FF4BFD59FC3DFC8F" blockId="2.[151,1437,150,1437]" pageId="2" pageNumber="449">Elytra widely suboval, 1.1 times as long as wide; with small humeral tooth at base of costa 8. Odd costae twice as high as even ones; both odd and even costae covered by small, dust-like setae. Intercostae with large punctures; the punctures about as long as wide, distance between punctures slightly longer than their diameter. Intervals between punctures shiny. Scutellum small, elongate, with rounded apex.</paragraph>
|
||
<paragraph id="8BBE36B1FFABFF83FF4BFCC9FD13FC1B" blockId="2.[151,1437,150,1437]" pageId="2" pageNumber="449">Profemur wide, with large punctures throughout. Protibia slim at base; slightly widened toward the apex, with three equidistant teeth in apical half; apical tooth long, medial slightly shorter, basal tooth twice shorter than medial. Apical spur as long as two basal tarsomeres.</paragraph>
|
||
<paragraph id="8BBE36B1FFABFF83FF4BFC5DFB07FC53" blockId="2.[151,1437,150,1437]" pageId="2" pageNumber="449">
|
||
Ventral side (
|
||
<figureCitation id="133A2A34FFABFF83FED3FC5DFE38FC3F" box="[351,419,907,933]" captionStart="FIGURES 1–6" captionStartId="1.[151,264,1825,1849]" captionTargetBox="[303,1299,179,1799]" captionTargetId="figure-19@1.[288,1299,179,1799]" captionTargetPageId="1" captionText="FIGURES 1–6. Habitus of Odochilus (Parodochilus) species. 1, O. (P.) shavrini Rakovič & Anichtchenko, new species, male (holotype), habitus in dorsal view; 2, O. (P.) shavrini new species, male (holotype), habitus in dorsolateral view; 3, O. (P.) shavrini new species, male (holotype), habitus in ventral view; 4, O. (P.) buruensis Boucomont, 1926, drawing of pronotum overprinted from Rakovič, 1997 (permitted by Acta Societatis Zoologicae Bohemicae); 5, O. (P.) borneensis Rakovič & Anich- tchenko, new species, female (holotype), habitus in dorsal view; 6, O. (P.) jani Bezděk & Křivan, 2001, paratype no. 3, habitus in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4604478" httpUri="https://zenodo.org/record/4604478/files/figure.png" pageId="2" pageNumber="449">Fig. 3</figureCitation>
|
||
) dark brown, matte. Abdomen with ventrites 1–5 minutely punctate, ventrite 5 with four short setae; ventrite 6 densely punctate, matte, with six setae. Pygidium with six erect setae.
|
||
</paragraph>
|
||
<paragraph id="8BBE36B1FFABFF83FF4BFC05FF4FFBAF" blockId="2.[151,1437,150,1437]" pageId="2" pageNumber="449">
|
||
Female: there are no considerable differences between external characters of the male
|
||
<typeStatus id="54BA8813FFABFF83FB29FC05FA9CFC77" box="[1189,1287,979,1005]" pageId="2" pageNumber="449" type="holotype">holotype</typeStatus>
|
||
and those of the female
|
||
<typeStatus id="54BA8813FFABFF83FE9FFC21FEECFB8B" box="[275,375,1015,1041]" pageId="2" pageNumber="449" type="paratype">paratype</typeStatus>
|
||
. The body length of the female
|
||
<typeStatus id="54BA8813FFABFF83FD54FC21FCA3FB8B" box="[728,824,1015,1041]" pageId="2" pageNumber="449" type="paratype">paratype</typeStatus>
|
||
(3.0 mm) is larger compared to the male
|
||
<typeStatus id="54BA8813FFABFF83FA8BFC21FAF2FB8B" box="[1287,1385,1015,1041]" pageId="2" pageNumber="449" type="holotype">holotype</typeStatus>
|
||
(
|
||
<quantity id="4CF99B54FFABFF83FAF4FC21FF5CFBAF" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="449" unit="mm" value="2.8">2.8 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31B653AFFABFF83FF4BFBE8FC45FAAB" pageId="2" pageNumber="449" type="diagnosis">
|
||
<paragraph id="8BBE36B1FFABFF83FF4BFBE8FC45FAAB" blockId="2.[151,1437,150,1437]" pageId="2" pageNumber="449">
|
||
<emphasis id="B975EAA3FFABFF83FF4BFBE8FE56FBC2" bold="true" box="[199,461,1086,1112]" pageId="2" pageNumber="449">Differential diagnosis.</emphasis>
|
||
The new species can be compared only with
|
||
<taxonomicName id="4C014D32FFABFF83FC44FBE9FEACFBE7" authority="Boucomont, 1926" authorityName="Boucomont" authorityYear="1926" class="Insecta" family="Aphodiidae" genus="Odochilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="449" phylum="Arthropoda" rank="species" species="buruensis" subGenus="Parodochilus">
|
||
<emphasis id="B975EAA3FFABFF83FC44FBE9FBA0FBC3" box="[968,1083,1087,1113]" italics="true" pageId="2" pageNumber="449">Odochilus</emphasis>
|
||
(
|
||
<emphasis id="B975EAA3FFABFF83FBC7FBE9FB7AFBC3" box="[1099,1249,1087,1113]" italics="true" pageId="2" pageNumber="449">Parodochilus</emphasis>
|
||
)
|
||
<emphasis id="B975EAA3FFABFF83FB7DFBE9FAC5FBC3" box="[1265,1374,1087,1113]" italics="true" pageId="2" pageNumber="449">buruensis</emphasis>
|
||
<bibRefCitation id="EF904B40FFABFF83FAE8FBE9FEACFBE7" author="Boucomont, A." pageId="2" pageNumber="449" pagination="110" refId="ref2276" refString="Boucomont, A. (1926) Fauna Buruana. Coleoptera, Fam. Lamellicornia Coprophaga. Treubia, 7, 110." type="journal article" year="1926">Boucomont, 1926</bibRefCitation>
|
||
</taxonomicName>
|
||
, known from the
|
||
<collectingCountry id="F3167621FFABFF83FD8BFBB5FDECFBE7" box="[519,631,1123,1149]" name="Indonesia" pageId="2" pageNumber="449">Moluccas</collectingCountry>
|
||
, because of an unusually small elytra length-to-width ratio (the elytra are relatively very broad). The two species can be most easily differentiated from each other based on the pronotal sculpture. In
|
||
<taxonomicName id="4C014D32FFABFF83FEA5FB7DFE7DFB5F" authorityName="Boucomont" authorityYear="1926" box="[297,486,1195,1221]" class="Insecta" family="Aphodiidae" genus="Odochilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="449" phylum="Arthropoda" rank="species" species="buruensis" subGenus="Parodochilus">
|
||
<emphasis id="B975EAA3FFABFF83FEA5FB7DFEDFFB5F" box="[297,324,1195,1221]" italics="true" pageId="2" pageNumber="449">O.</emphasis>
|
||
(
|
||
<emphasis id="B975EAA3FFABFF83FED8FB7DFEF1FB5E" box="[340,362,1195,1220]" italics="true" pageId="2" pageNumber="449">P.</emphasis>
|
||
)
|
||
<emphasis id="B975EAA3FFABFF83FEF5FB7DFE7DFB5F" box="[377,486,1195,1221]" italics="true" pageId="2" pageNumber="449">buruensis</emphasis>
|
||
</taxonomicName>
|
||
ridges 1–3 are in the form of continuous, high, narrow and sharp lamellae (
|
||
<figureCitation id="133A2A34FFABFF83FAA1FB7DFAE9FB5F" box="[1325,1394,1195,1221]" captionStart="FIGURES 1–6" captionStartId="1.[151,264,1825,1849]" captionTargetBox="[303,1299,179,1799]" captionTargetId="figure-19@1.[288,1299,179,1799]" captionTargetPageId="1" captionText="FIGURES 1–6. Habitus of Odochilus (Parodochilus) species. 1, O. (P.) shavrini Rakovič & Anichtchenko, new species, male (holotype), habitus in dorsal view; 2, O. (P.) shavrini new species, male (holotype), habitus in dorsolateral view; 3, O. (P.) shavrini new species, male (holotype), habitus in ventral view; 4, O. (P.) buruensis Boucomont, 1926, drawing of pronotum overprinted from Rakovič, 1997 (permitted by Acta Societatis Zoologicae Bohemicae); 5, O. (P.) borneensis Rakovič & Anich- tchenko, new species, female (holotype), habitus in dorsal view; 6, O. (P.) jani Bezděk & Křivan, 2001, paratype no. 3, habitus in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4604478" httpUri="https://zenodo.org/record/4604478/files/figure.png" pageId="2" pageNumber="449">Fig. 4</figureCitation>
|
||
); in the new species, ridges 1–3 are convex but not in the form of lamellae and ridge 3 is rather interrupted medially (
|
||
<figureCitation id="133A2A34FFABFF83FAE3FB19FF3CFA97" captionStart="FIGURES 1–6" captionStartId="1.[151,264,1825,1849]" captionTargetBox="[303,1299,179,1799]" captionTargetId="figure-19@1.[288,1299,179,1799]" captionTargetPageId="1" captionText="FIGURES 1–6. Habitus of Odochilus (Parodochilus) species. 1, O. (P.) shavrini Rakovič & Anichtchenko, new species, male (holotype), habitus in dorsal view; 2, O. (P.) shavrini new species, male (holotype), habitus in dorsolateral view; 3, O. (P.) shavrini new species, male (holotype), habitus in ventral view; 4, O. (P.) buruensis Boucomont, 1926, drawing of pronotum overprinted from Rakovič, 1997 (permitted by Acta Societatis Zoologicae Bohemicae); 5, O. (P.) borneensis Rakovič & Anich- tchenko, new species, female (holotype), habitus in dorsal view; 6, O. (P.) jani Bezděk & Křivan, 2001, paratype no. 3, habitus in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4604478" httpUri="https://zenodo.org/record/4604478/files/figure.png" pageId="2" pageNumber="449">Fig. 5</figureCitation>
|
||
). Longitudinal V-shaped formations, present in the subgenus
|
||
<taxonomicName id="4C014D32FFABFF83FCD6FB25FC6BFA97" box="[858,1008,1267,1293]" class="Insecta" family="Aphodiidae" genus="Odochilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="449" phylum="Arthropoda" rank="subGenus" subGenus="Parodochilus">
|
||
<emphasis id="B975EAA3FFABFF83FCD6FB25FC6BFA97" box="[858,1008,1267,1293]" italics="true" pageId="2" pageNumber="449">Parodochilus</emphasis>
|
||
</taxonomicName>
|
||
instead of ridge 4, are very distinct in the former species (
|
||
<figureCitation id="133A2A34FFABFF83FEF8FAC1FE23FAAB" box="[372,440,1303,1329]" captionStart="FIGURES 1–6" captionStartId="1.[151,264,1825,1849]" captionTargetBox="[303,1299,179,1799]" captionTargetId="figure-19@1.[288,1299,179,1799]" captionTargetPageId="1" captionText="FIGURES 1–6. Habitus of Odochilus (Parodochilus) species. 1, O. (P.) shavrini Rakovič & Anichtchenko, new species, male (holotype), habitus in dorsal view; 2, O. (P.) shavrini new species, male (holotype), habitus in dorsolateral view; 3, O. (P.) shavrini new species, male (holotype), habitus in ventral view; 4, O. (P.) buruensis Boucomont, 1926, drawing of pronotum overprinted from Rakovič, 1997 (permitted by Acta Societatis Zoologicae Bohemicae); 5, O. (P.) borneensis Rakovič & Anich- tchenko, new species, female (holotype), habitus in dorsal view; 6, O. (P.) jani Bezděk & Křivan, 2001, paratype no. 3, habitus in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4604478" httpUri="https://zenodo.org/record/4604478/files/figure.png" pageId="2" pageNumber="449">Fig. 4</figureCitation>
|
||
), but rather indistinct in the new species (
|
||
<figureCitation id="133A2A34FFABFF83FC00FAC1FC4BFAAB" box="[908,976,1302,1329]" captionStart="FIGURES 1–6" captionStartId="1.[151,264,1825,1849]" captionTargetBox="[303,1299,179,1799]" captionTargetId="figure-19@1.[288,1299,179,1799]" captionTargetPageId="1" captionText="FIGURES 1–6. Habitus of Odochilus (Parodochilus) species. 1, O. (P.) shavrini Rakovič & Anichtchenko, new species, male (holotype), habitus in dorsal view; 2, O. (P.) shavrini new species, male (holotype), habitus in dorsolateral view; 3, O. (P.) shavrini new species, male (holotype), habitus in ventral view; 4, O. (P.) buruensis Boucomont, 1926, drawing of pronotum overprinted from Rakovič, 1997 (permitted by Acta Societatis Zoologicae Bohemicae); 5, O. (P.) borneensis Rakovič & Anich- tchenko, new species, female (holotype), habitus in dorsal view; 6, O. (P.) jani Bezděk & Křivan, 2001, paratype no. 3, habitus in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4604478" httpUri="https://zenodo.org/record/4604478/files/figure.png" pageId="2" pageNumber="449">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31B653AFFABFF83FF4BFAECFD4CFA07" pageId="2" pageNumber="449" type="distribution">
|
||
<paragraph id="8BBE36B1FFABFF83FF4BFAECFB73FACF" blockId="2.[151,1437,150,1437]" box="[199,1256,1338,1365]" pageId="2" pageNumber="449">
|
||
<emphasis id="B975EAA3FFABFF83FF4BFAECFEFAFACE" bold="true" box="[199,353,1338,1364]" pageId="2" pageNumber="449">Distribution.</emphasis>
|
||
Species known only from the
|
||
<typeStatus id="54BA8813FFABFF83FD3BFAEDFD7CFACF" box="[695,743,1339,1365]" pageId="2" pageNumber="449">type</typeStatus>
|
||
locality in Mindanao Island, The
|
||
<collectingCountry id="F3167621FFABFF83FBEFFAEDFB7EFACF" box="[1123,1253,1339,1365]" name="Philippines" pageId="2" pageNumber="449">Philippines</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBE36B1FFABFF83FF4BFA89FD4CFA07" blockId="2.[151,1437,150,1437]" pageId="2" pageNumber="449">
|
||
<emphasis id="B975EAA3FFABFF83FF4BFA89FE0DFAE2" bold="true" box="[199,406,1374,1401]" pageId="2" pageNumber="449">Name derivation.</emphasis>
|
||
New species named after our friend and colleague, specialist in
|
||
<taxonomicName id="4C014D32FFABFF83FBEBFA89FA03FAE0" authority="(Coleoptera)" baseAuthorityName="Coleoptera" box="[1127,1432,1375,1402]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="449" phylum="Arthropoda" rank="family">Staphylinidae (Coleoptera)</taxonomicName>
|
||
, Dr. Alexey Shavrin, who collected the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |