treatments-xml/data/03/A8/87/03A887A1A977197CAFE9FA0243F163D9.xml
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<document id="12B003FEBB2DD21D04CC2C1276B4DA51" ID-CLB-Dataset="8896" ID-DOI="10.11646/zootaxa.4868.1.4" ID-GBIF-Dataset="cfd5c06e-06f0-4f1f-ae4b-40b7b46f44a3" ID-ISSN="1175-5326" ID-Zenodo-Dep="4417281" ID-ZooBank="DC4B8EC5-CBDA-49D2-B370-381C96AF3A10" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609805123592" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2020" docId="03A887A1A977197CAFE9FA0243F163D9" docLanguage="en" docName="zootaxa.4868.1.4.pdf" docOrigin="Zootaxa 4868 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydrochus uniformis Perkins 2020, new species" docType="treatment" docVersion="8" lastPageNumber="85" masterDocId="FF91FFD9A9611964AF7EFFC8470B6435" masterDocTitle="Taxonomy of Peruvian water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)" masterLastPageNumber="89" masterPageNumber="61" pageNumber="83" updateTime="1698852338306" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="37377C46521BAF4A4443DFE666661058">Taxonomy of Peruvian water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)</mods:title>
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<mods:date id="29B4D4DE37331929FE1B390A75DBC00C">2020</mods:date>
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<mods:number id="2095BC13C094157AD94B9F4059E74FE4">2020-10-23</mods:number>
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<treatment id="03A887A1A977197CAFE9FA0243F163D9" ID-DOI="http://doi.org/10.5281/zenodo.4417480" ID-GBIF-Taxon="176581827" ID-Zenodo-Dep="4417480" LSID="urn:lsid:plazi:treatment:03A887A1A977197CAFE9FA0243F163D9" httpUri="http://treatment.plazi.org/id/03A887A1A977197CAFE9FA0243F163D9" lastPageId="24" lastPageNumber="85" pageId="22" pageNumber="83">
<subSubSection id="C31B653CA9771972AFE9FA02453161D1" box="[151,570,1482,1509]" pageId="22" pageNumber="83" type="nomenclature">
<paragraph id="8BBE36B7A9771972AFE9FA02453161D1" blockId="22.[151,570,1482,1545]" box="[151,570,1482,1509]" pageId="22" pageNumber="83">
<heading id="D0F681DBA9771972AFE9FA02453161D1" bold="true" box="[151,570,1482,1509]" fontSize="11" level="1" pageId="22" pageNumber="83" reason="1">
<emphasis id="B975EAA5A9771972AFE9FA02453161D1" bold="true" box="[151,570,1482,1509]" pageId="22" pageNumber="83">
<taxonomicName id="4C014D34A9771972AFE9FA02469061D1" ID-CoL="8Y4RR" authority="Perkins, 2020" authorityName="Perkins" authorityYear="2020" box="[151,411,1482,1509]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="83" phylum="Arthropoda" rank="species" species="uniformis" status="sp. nov.">
<emphasis id="B975EAA5A9771972AFE9FA02469061D1" bold="true" box="[151,411,1482,1509]" italics="true" pageId="22" pageNumber="83">Hydrochus uniformis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A24657DEA9771972AED4FA03453161D1" box="[426,570,1482,1509]" pageId="22" pageNumber="83" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C31B653CA9771972AFE9FA274606623C" box="[151,269,1519,1545]" pageId="22" pageNumber="83" type="description">
<paragraph id="8BBE36B7A9771972AFE9FA274606623C" blockId="22.[151,570,1482,1545]" box="[151,269,1519,1545]" pageId="22" pageNumber="83">
<figureCitation id="133A2A32A9771972AFE9FA2747E8623C" box="[151,227,1519,1545]" captionStart="FIGURE 1" captionStartId="2.[151,250,1971,1995]" captionTargetBox="[157,1430,309,1948]" captionTargetId="figure-37@2.[157,1430,309,1948]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus of Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.4417283" httpUri="https://zenodo.org/record/4417283/files/figure.png" pageId="22" pageNumber="83">Figs. 1</figureCitation>
,
<figureCitation id="133A2A32A9771972AF8FFA274606623C" box="[241,269,1519,1545]" captionStart="FIGURE 16" captionStartId="24.[151,250,1643,1667]" captionTargetBox="[177,1415,205,1605]" captionTargetId="figure-26@24.[151,1435,181,1620]" captionTargetPageId="24" captionText="FIGURE 16. Hydrochus uniformis n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.4417321" httpUri="https://zenodo.org/record/4417321/files/figure.png" pageId="22" pageNumber="83">16</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31B653CA9771972AFE9F9FF464F62AC" pageId="22" pageNumber="83" type="materials_examined">
<paragraph id="8BBE36B7A9771972AFE9F9FF464F62AC" blockId="22.[151,1437,1590,2013]" pageId="22" pageNumber="83">
<emphasis id="B975EAA5A9771972AFE9F9FF46426265" bold="true" box="[151,329,1590,1617]" pageId="22" pageNumber="83">Type Material.</emphasis>
<materialsCitation id="3B693CEAA9771972AE2CF9FF464F62AC" ID-GBIF-Occurrence="3015416353" collectingDate="1989-06-21" collectionCode="SEMC" collectorName="J. S. Ashe &amp; R. A. Leschen" country="Peru" county="Tambopata Prov." elevation="200" latitude="-12.55" location="Reserva" longLatPrecision="1293" longitude="-69.05" municipality="Dpto" pageId="22" pageNumber="83" specimenCount="1" stateProvince="Madre de Dios" typeStatus="holotype">
<emphasis id="B975EAA5A9771972AE2CF9FF45176265" bold="true" box="[338,540,1590,1617]" pageId="22" pageNumber="83">
<typeStatus id="54BA8815A9771972AE2CF9FF46B46264" box="[338,447,1591,1617]" pageId="22" pageNumber="83" type="holotype">Holotype</typeStatus>
(male):
</emphasis>
<collectingCountry id="F3167627A9771972AD51F9FF45766264" box="[559,637,1591,1617]" name="Peru" pageId="22" pageNumber="83">PERU</collectingCountry>
:
<collectingCounty id="62DF4E3BA9771972ADF6F9FF44416264" box="[648,842,1591,1617]" pageId="22" pageNumber="83">Tambopata Prov.</collectingCounty>
,
<collectingRegion id="49C5F855A9771972AC28F9FF44F46264" box="[854,1023,1591,1617]" country="Peru" name="Madre de Dios" pageId="22" pageNumber="83">Madre de Dios</collectingRegion>
<collectingMunicipality id="6BDAACCDA9771972AB79F9FF43496264" box="[1031,1090,1591,1617]" pageId="22" pageNumber="83">Dpto</collectingMunicipality>
,
<locationDeviation id="9A0C2F95A9771972AB31F9FE42936264" box="[1103,1432,1590,1617]" location="Puerto Maldonado" pageId="22" pageNumber="83">
<quantity id="4CF99B52A9771972AB31F9FE43846264" box="[1103,1167,1590,1617]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="22" pageNumber="83" unit="km" value="15.0">15km</quantity>
NE Puerto Maldonado
</locationDeviation>
,
<location id="8EDE606CA9771972AFE9F99347FA6240" LSID="urn:lsid:plazi:treatment:03A887A1A977197CAFE9FA0243F163D9:8EDE606CA9771972AFE9F99347FA6240" box="[151,241,1627,1653]" country="Peru" county="Tambopata Prov." latitude="-12.55" longLatPrecision="1293" longitude="-69.05" municipality="Dpto" name="Reserva" pageId="22" pageNumber="83" stateProvince="Madre de Dios">Reserva</location>
<collectingRegion id="49C5F855A9771972AF87F993464A6240" box="[249,321,1627,1653]" country="Peru" name="Cusco" pageId="22" pageNumber="83">Cuzco</collectingRegion>
<location id="8EDE606CA9771972AE39F99346C06240" LSID="urn:lsid:plazi:treatment:03A887A1A977197CAFE9FA0243F163D9:8EDE606CA9771972AE39F99346C06240" box="[327,459,1627,1653]" country="Peru" county="Tambopata Prov." latitude="-12.55" longLatPrecision="1293" longitude="-69.05" municipality="Dpto" name="Amazonico" pageId="22" pageNumber="83" stateProvince="Madre de Dios">Amazonico</location>
,
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,
<geoCoordinate id="EE355070A9771972AD3FF99345916240" box="[577,666,1627,1653]" degrees="69" direction="west" minutes="3" orientation="longitude" pageId="22" pageNumber="83" precision="925" value="-69.05">69°3W</geoCoordinate>
,
<quantity id="4CF99B52A9771972ADD8F99345E26240" box="[678,745,1627,1653]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="22" pageNumber="83" unit="m" value="200.0">
<elevation id="002CD184A9771972ADD8F99345E26240" box="[678,745,1627,1653]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="22" pageNumber="83" unit="m" value="200.0">200m</elevation>
</quantity>
, camp,
<date id="FFBF1077A9771972AC3EF99344D26240" box="[832,985,1627,1653]" pageId="22" pageNumber="83" value="1989-06-21">
<collectingDate id="EFFBE99FA9771972AC3EF99344D26240" box="[832,985,1627,1653]" pageId="22" pageNumber="83" value="1989-06-21">21 June 1989</collectingDate>
</date>
,
<collectorName id="26F45361A9771972AC9BF993435C6240" box="[997,1111,1627,1653]" pageId="22" pageNumber="83">J. S. Ashe</collectorName>
,
<collectorName id="26F45361A9771972AB1DF993420D6240" box="[1123,1286,1627,1653]" pageId="22" pageNumber="83">R. A. Leschen</collectorName>
, #213, ex., at light” (
<collectionCode id="ED10AE72A9771972AF98F9B7463362AC" box="[230,312,1663,1689]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="22" pageNumber="83" type="Museum">SEMC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31B653CA9771972AFB9F96A450A631C" pageId="22" pageNumber="83" type="diagnosis">
<paragraph id="8BBE36B7A9771972AFB9F96A450A631C" blockId="22.[151,1437,1590,2013]" pageId="22" pageNumber="83">
<emphasis id="B975EAA5A9771972AFB9F96A46D86289" bold="true" box="[199,467,1698,1724]" pageId="22" pageNumber="83">Differential Diagnosis.</emphasis>
Among Peruvian species, recognized by the combination of small size (ca.
<quantity id="4CF99B52A9771972AA56F96B429B6288" box="[1320,1424,1699,1725]" metricMagnitude="-3" metricUnit="m" metricValue="2.27" pageId="22" pageNumber="83" unit="mm" value="2.27">2.27 mm</quantity>
), the dense and deep dorsal punctation, the interstices with sparse to moderately dense microsetae, the pronotal punctures of very uniform size,, and the male genitalia (
<figureCitation id="133A2A32A9771972AD9EF923443F6330" box="[736,820,1771,1797]" captionStart="FIGURE 16" captionStartId="24.[151,250,1643,1667]" captionTargetBox="[177,1415,205,1605]" captionTargetId="figure-26@24.[151,1435,181,1620]" captionTargetPageId="24" captionText="FIGURE 16. Hydrochus uniformis n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.4417321" httpUri="https://zenodo.org/record/4417321/files/figure.png" pageId="22" pageNumber="83">Fig. 16</figureCitation>
; described below). Completely reliable determinations will include dissection of males.
</paragraph>
</subSubSection>
<subSubSection id="C31B653CA977197CAFB9F8FA46F063FD" lastPageId="24" lastPageNumber="85" pageId="22" pageNumber="83" type="description">
<paragraph id="8BBE36B7A9771973AFB9F8FA44C864E0" blockId="22.[151,1437,1590,2013]" lastBlockId="23.[151,1437,150,213]" lastPageId="23" lastPageNumber="84" pageId="22" pageNumber="83">
<emphasis id="B975EAA5A9771972AFB9F8FA465A6379" bold="true" box="[199,337,1842,1868]" pageId="22" pageNumber="83">Description</emphasis>
. Habitus as illustrated (
<figureCitation id="133A2A32A9771972AD27F8FB45A7637B" box="[601,684,1843,1870]" captionStart="FIGURE 16" captionStartId="24.[151,250,1643,1667]" captionTargetBox="[177,1415,205,1605]" captionTargetId="figure-26@24.[151,1435,181,1620]" captionTargetPageId="24" captionText="FIGURE 16. Hydrochus uniformis n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.4417321" httpUri="https://zenodo.org/record/4417321/files/figure.png" pageId="22" pageNumber="83">Fig. 16</figureCitation>
). Size:
<typeStatus id="54BA8815A9771972AD80F8FB446B6378" box="[766,864,1843,1869]" pageId="22" pageNumber="83" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 2.27/0.86; head width 0.58; pronotum l/w 0.52/0.55; PA 0.52; PB 0.44; elytra 1.36/0.86. Body size moderate (ca.
<quantity id="4CF99B52A9771972AFE9F8B347F563A0" box="[151,254,1915,1941]" metricMagnitude="-3" metricUnit="m" metricValue="2.27" pageId="22" pageNumber="83" unit="mm" value="2.27">2.27 mm</quantity>
). Dorsum dark brown to black, with slight iridescence, elytra without distinct black spots. Legs testaceous, with tibiofemoral joints and basal ½ of femora darker. Dorsal punctuation dense and deep. Punctation on elytra ca. 45x that of pronotum. Elytra interstriae narrow ridges forming zig-zag pattern to ca. 1x strial puncture diameter, with sparse microsetae. Interstria 9
<superScript id="7C749BFFA9761973AD65FF5E45236491" attach="left" box="[539,552,150,164]" fontSize="6" pageId="23" pageNumber="84">th</superScript>
more convex than others, overhanging 10
<superScript id="7C749BFFA9761973AC85FF5E43036491" attach="left" box="[1019,1032,150,164]" fontSize="6" pageId="23" pageNumber="84">th</superScript>
interstria. Usual area of callus on 5
<superScript id="7C749BFFA9761973AAEEFF5E42966491" attach="none" box="[1424,1437,150,164]" fontSize="6" pageId="23" pageNumber="84">th</superScript>
weakly raised, moderately elongate, ca. 5 punctures in row on each side.
</paragraph>
<caption id="DF7E663FA9761973AFE9F91344D362C6" ID-DOI="http://doi.org/10.5281/zenodo.4417319" ID-Zenodo-Dep="4417319" box="[151,984,1755,1780]" httpUri="https://zenodo.org/record/4417319/files/figure.png" pageId="23" pageNumber="84" startId="23.[151,250,1755,1779]" targetBox="[162,1435,298,1719]" targetPageId="23">
<paragraph id="8BBE36B7A9761973AFE9F91344D362C6" blockId="23.[151,984,1755,1780]" box="[151,984,1755,1780]" pageId="23" pageNumber="84">
<emphasis id="B975EAA5A9761973AFE9F913462B62C6" bold="true" box="[151,288,1755,1779]" pageId="23" pageNumber="84">FIGURE 15.</emphasis>
<taxonomicName id="4C014D34A9761973AE58F913450962C6" authority="Perkins, 2020" authorityName="Perkins" authorityYear="2020" box="[294,514,1755,1779]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="84" phylum="Arthropoda" rank="species" species="tripartitus" status="sp. nov.">
<emphasis id="B975EAA5A9761973AE58F913450962C6" box="[294,514,1755,1779]" italics="true" pageId="23" pageNumber="84">Hydrochus tripartitus</emphasis>
</taxonomicName>
<emphasis id="B975EAA5A9761973AD76F914454A62C1" bold="true" box="[520,577,1756,1780]" pageId="23" pageNumber="84">
<taxonomicNameLabel id="A24657DEA9761973AD76F914454A62C1" box="[520,577,1756,1780]" pageId="23" pageNumber="84" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, habitus and male genitalia of holotype.
</paragraph>
</caption>
<paragraph id="8BBE36B7A9761973AFB9F8EA435863B1" blockId="23.[151,1437,1826,2032]" pageId="23" pageNumber="84">Pronotum densely coarsely punctate, of quite uniform size; interstices rounded, with rather dense microsetae, length less than width, widest slightly behind anterior margin, narrowed at base, sides arcuate or very weakly sinuate, smooth; anterior margin slightly bisinuate; anterior depressions moderately deep.</paragraph>
<paragraph id="8BBE36B7A9761973AFB9F846461E63C5" blockId="23.[151,1437,1826,2032]" pageId="23" pageNumber="84">Elytra with sutural interval raised over posterior declivity; spaces between strial punctures narrow walls to ca. 0.5x their diameter; apices slightly produced, separately rounded in dorsal view, in lateral view outer margin without angulation.</paragraph>
<caption id="DF7E663FA979197CAFE9F9A344D862B1" ID-DOI="http://doi.org/10.5281/zenodo.4417321" ID-Zenodo-Dep="4417321" box="[151,979,1643,1668]" httpUri="https://zenodo.org/record/4417321/files/figure.png" pageId="24" pageNumber="85" startId="24.[151,250,1643,1667]" targetBox="[177,1415,205,1605]" targetPageId="24">
<paragraph id="8BBE36B7A979197CAFE9F9A344D862B1" blockId="24.[151,979,1643,1668]" box="[151,979,1643,1668]" pageId="24" pageNumber="85">
<emphasis id="B975EAA5A979197CAFE9F9A3462B62B1" bold="true" box="[151,288,1643,1668]" pageId="24" pageNumber="85">FIGURE 16.</emphasis>
<taxonomicName id="4C014D34A979197CAE58F9A446F762B1" authority="Perkins, 2020" authorityName="Perkins" authorityYear="2020" box="[294,508,1644,1668]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="85" phylum="Arthropoda" rank="species" species="uniformis" status="sp. nov.">
<emphasis id="B975EAA5A979197CAE58F9A446F762B1" box="[294,508,1644,1668]" italics="true" pageId="24" pageNumber="85">Hydrochus uniformis</emphasis>
</taxonomicName>
<emphasis id="B975EAA5A979197CAD7CF9A4453062B1" bold="true" box="[514,571,1644,1668]" pageId="24" pageNumber="85">
<taxonomicNameLabel id="A24657DEA979197CAD7CF9A4453062B1" box="[514,571,1644,1668]" pageId="24" pageNumber="85" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, habitus and male genitalia of holotype.
</paragraph>
</caption>
<paragraph id="8BBE36B7A979197CAFB9F97A427F62F9" blockId="24.[151,1436,1714,2028]" box="[199,1396,1714,1740]" pageId="24" pageNumber="85">Ventral characters: Mentum densely punctate, with large median fovea. Submentum with two large foveae.</paragraph>
<paragraph id="8BBE36B7A979197CAFB9F91E44FE6369" blockId="24.[151,1436,1714,2028]" pageId="24" pageNumber="85">
Male genitalia (
<figureCitation id="133A2A32A979197CAE08F91E46CD62C4" box="[374,454,1750,1777]" captionStart="FIGURE 16" captionStartId="24.[151,250,1643,1667]" captionTargetBox="[177,1415,205,1605]" captionTargetId="figure-26@24.[151,1435,181,1620]" captionTargetPageId="24" captionText="FIGURE 16. Hydrochus uniformis n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.4417321" httpUri="https://zenodo.org/record/4417321/files/figure.png" pageId="24" pageNumber="85">Fig. 16</figureCitation>
) general characters: short and rather broad, basal piece longer than parameres; paramere tips extending to level of aedeagus tip, or very slightly beyond, weakly sclerotized, widened slightly, arcuate slightly toward midline but not contacting one another; aedeagus wider than parameres, broad even at base; basal orifice distinctively shaped, broad in lateral view, with apicoventral arcuate process.
</paragraph>
<paragraph id="8BBE36B7A979197CAFB9F8AE429F63B5" blockId="24.[151,1436,1714,2028]" box="[199,1428,1894,1922]" pageId="24" pageNumber="85">Dorsal surface: pdmm contiguous with alm in distal ½, overlapping alm slightly in proximal ½; adtl and adbl absent.</paragraph>
<paragraph id="8BBE36B7A979197CAFB9F84246F063FD" blockId="24.[151,1436,1714,2028]" pageId="24" pageNumber="85">Ventral surface: pvmm slightly overlapping alm; parameres fused to one another basally; pseudoparameres large, ca. ½ length of paramere.</paragraph>
</subSubSection>
<subSubSection id="C31B653CA979197CAFB9F81A43F163D9" box="[199,1274,2002,2028]" pageId="24" pageNumber="85" type="etymology">
<paragraph id="8BBE36B7A979197CAFB9F81A43F163D9" blockId="24.[151,1436,1714,2028]" box="[199,1274,2002,2028]" pageId="24" pageNumber="85">
<emphasis id="B975EAA5A979197CAFB9F81A464563D9" bold="true" box="[199,334,2002,2028]" pageId="24" pageNumber="85">Etymology.</emphasis>
Named in reference to the uniform size of the punctures on the head and pronotum.
</paragraph>
</subSubSection>
</treatment>
</document>