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<document id="BABD8DCDD61490407E7F69AF58C5760D" ID-CLB-Dataset="32359" ID-DOI="10.11646/zootaxa.4308.1.1" ID-GBIF-Dataset="01ef7a56-a2ba-4493-9182-a0ca2f9a32d2" ID-ISSN="1175-5326" ID-Zenodo-Dep="846086" ID-ZooBank="254C1FE2-8233-43C0-9B83-7A10BC64A092" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1503302896227" checkinUser="plazi" docAuthor="Schneider, Clément" docDate="2017" docId="03A8760E7B1D4F7872BBFF45D07172E4" docLanguage="en" docName="zootaxa.4308.1.1.pdf" docOrigin="Zootaxa 4308 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Megalothorax nigropunctatus , Schneider &amp; D'Haese 2013" docType="treatment" docVersion="8" lastPageNumber="55" masterDocId="FF910E767B294F4E722CFFD2D33D7769" masterDocTitle="Morphological review of the order Neelipleona (Collembola) through the redescription of the type species of Acanthoneelidus, Neelides and Neelus" masterLastPageNumber="94" masterPageNumber="1" pageNumber="53" updateTime="1698466929409" updateUser="ExternalLinkService">
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<mods:title id="2848ACF5266CBC4905165F460CD8DE70">Morphological review of the order Neelipleona (Collembola) through the redescription of the type species of Acanthoneelidus, Neelides and Neelus</mods:title>
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<mods:namePart id="5569C82C7D556791C9691AB7DA5C905B">Schneider, Clément</mods:namePart>
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<mods:date id="36E329A4E14686AB38B6DF0BB840C1E6">2017</mods:date>
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<mods:number id="0F78185EFA426C0F4E592E8683346B71">4308</mods:number>
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<treatment id="03A8760E7B1D4F7872BBFF45D07172E4" ID-DOI="http://doi.org/10.5281/zenodo.6040267" ID-GBIF-Taxon="132786154" ID-Zenodo-Dep="6040267" LSID="urn:lsid:plazi:treatment:03A8760E7B1D4F7872BBFF45D07172E4" httpUri="http://treatment.plazi.org/id/03A8760E7B1D4F7872BBFF45D07172E4" lastPageId="54" lastPageNumber="55" pageId="52" pageNumber="53">
<subSubSection id="C31B94937B1D4F7A72BBFF45D2CA77BA" pageId="52" pageNumber="53" type="nomenclature">
<paragraph id="8BBEC7187B1D4F7A72BBFF45D05D77DB" blockId="52.[151,864,151,211]" box="[151,864,151,178]" pageId="52" pageNumber="53">
<heading id="D0F670747B1D4F7A72BBFF45D05D77DB" bold="true" box="[151,864,151,178]" fontSize="11" level="1" pageId="52" pageNumber="53" reason="1">
<taxonomicName id="4C01BC9B7B1D4F7A72BBFF45D05D77DB" authority="Schneider &amp; D'Haese, 2013" authorityName=", Schneider &amp; D'Haese" authorityYear="2013" box="[151,864,151,178]" class="Entognatha" family="Neelidae" genus="Megalothorax" kingdom="Animalia" order="Collembola" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="nigropunctatus">
<emphasis id="B9751B0A7B1D4F7A72BBFF45D05D77DB" bold="true" box="[151,864,151,178]" pageId="52" pageNumber="53">
<emphasis id="B9751B0A7B1D4F7A72BBFF45D13D77D8" bold="true" box="[151,512,151,177]" italics="true" pageId="52" pageNumber="53">Megalothorax nigropunctatus</emphasis>
Schneider &amp; D’Haese, 2013
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BBEC7187B1D4F7A72BBFF68D2CA77BA" blockId="52.[151,864,151,211]" box="[151,503,186,211]" pageId="52" pageNumber="53">
<figureCitation id="133ADB9D7B1D4F7A72BBFF68D3E677BA" box="[151,219,186,211]" captionStart="FIGURE 3" captionStartId="15.[151,250,1756,1778]" captionTargetBox="[175,1414,1242,1657]" captionTargetId="graphics@15.[786,1314,1251,1648]" captionTargetPageId="15" captionText="FIGURE 3. Diagram of the dorsal chaetotaxy of the head. (A) Neelides folsomi; (B) Acanthoneelidus pratensis; (C) Neelus murinus; (D) Megalothorax nigropunctatus; (E) Megalothorax perspicillum; (F) Megalothorax granulosus, integumentary channels not represented. In gray, chaetae absent in juvenile." httpUri="https://zenodo.org/record/846092/files/figure.svg" pageId="52" pageNumber="53">Figs 3</figureCitation>
D, 4D, 6, 15A, B, 20, 21.
</paragraph>
</subSubSection>
<subSubSection id="C31B94937B1D4F7A72BBFED0D76F76C3" pageId="52" pageNumber="53" type="materials_examined">
<paragraph id="8BBEC7187B1D4F7A72BBFED0D76F76C3" blockId="52.[151,1437,258,2011]" pageId="52" pageNumber="53">
<emphasis id="B9751B0A7B1D4F7A72BBFED0D2B97672" bold="true" box="[151,388,258,283]" pageId="52" pageNumber="53">Material examined.</emphasis>
Holotype. ♂ on slide (MNHN-EA
<date id="FFBFE1D87B1D4F7A710EFED0D04A7672" box="[802,887,258,283]" pageId="52" pageNumber="53" value="53-04-01">040153</date>
), Chile, Puerto Montt, Parque Nacional Alerce Andino, 41°34'58&quot;S 72°34'1&quot;W,
<date id="FFBFE1D87B1D4F7A7023FEF4D1B57656" box="[527,648,294,319]" pageId="52" pageNumber="53" value="2011-11-25">25.xi.2011</date>
, captured with aspirator under the bark of a dead tree, C. Schneider leg., [MNHNCH]. Paratypes.
<materialsCitation id="3B69CD457B1D4F7A702EFE99D21476EE" ID-GBIF-Occurrence="1586248032" collectionCode="MNHNCH" pageId="52" pageNumber="53" specimenCode="MNHN-EA040154-161" specimenCount="15" specimenCount-female="10" specimenCount-male="5" typeStatus="holotype">
<specimenCount id="9D070C917B1D4F7A702EFE99D179760B" box="[514,580,331,355]" pageId="52" pageNumber="53" type="female">10 ♀</specimenCount>
,
<specimenCount id="9D070C917B1D4F7A7077FE98D1B2760B" box="[603,655,330,355]" pageId="52" pageNumber="53" type="male">5 ♂</specimenCount>
on slides (
<specimenCode id="DBA76F637B1D4F7A710FFE98D705760A" box="[803,1080,330,355]" collectionCode="MNHN-EA" pageId="52" pageNumber="53">
MNHN-EA
<date id="FFBFE1D87B1D4F7A7181FE98D73E760A" box="[941,1027,330,355]" pageId="52" pageNumber="53" value="54-04-01">040154</date>
–161
</specimenCode>
) same data as the
<typeStatus id="54BA79BA7B1D4F7A771FFE98D6A5760A" box="[1331,1432,330,355]" pageId="52" pageNumber="53" type="holotype">holotype</typeStatus>
, [
<collectionCode id="ED105FDD7B1D4F7A72B1FEBDD21E76EE" box="[157,291,367,391]" country="Chile" httpUri="http://grbio.org/cool/qsji-8x93" name="Museo Nacional de Historia Natural de Chile" pageId="52" pageNumber="53">MNHNCH</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="3B69CD457B1D4F7A7319FEBDD6A576EF" ID-GBIF-Occurrence="1586248044" box="[309,1432,366,391]" collectionCode="MNHN" pageId="52" pageNumber="53" specimenCode="MNHN-EA040162-169" specimenCount="20" specimenCount-female="12" specimenCount-juvenile="4" specimenCount-male="4" typeStatus="holotype">
<specimenCount id="9D070C917B1D4F7A7319FEBDD25376EF" box="[309,366,367,391]" pageId="52" pageNumber="53" type="female">12 ♀</specimenCount>
,
<specimenCount id="9D070C917B1D4F7A7350FEBDD29A76EF" box="[380,423,367,391]" pageId="52" pageNumber="53" type="male">4 ♂</specimenCount>
and
<specimenCount id="9D070C917B1D4F7A73F2FEBDD16576EF" box="[478,600,366,391]" pageId="52" pageNumber="53" type="juvenile">4 juveniles</specimenCount>
on slides (
<specimenCode id="DBA76F637B1D4F7A70FEFEBCD0D476EF" box="[722,1001,366,390]" collectionCode="MNHN-EA" pageId="52" pageNumber="53">
MNHN-EA
<date id="FFBFE1D87B1D4F7A7170FEBCD08E76EF" box="[860,947,366,390]" pageId="52" pageNumber="53" value="62-04-01">040162</date>
–169
</specimenCode>
), same data as the
<typeStatus id="54BA79BA7B1D4F7A7690FEBCD61C76EF" box="[1212,1313,366,390]" pageId="52" pageNumber="53" type="holotype">holotype</typeStatus>
, [
<collectionCode id="ED105FDD7B1D4F7A771FFEBDD6AC76EF" box="[1331,1425,367,390]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="52" pageNumber="53">MNHN</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="3B69CD457B1D4F7A72BBFE41D77376C3" ID-GBIF-Occurrence="1586248050" box="[151,1102,402,427]" collectionCode="MNHN" pageId="52" pageNumber="53" specimenCode="MNHN-EA041007" specimenCount="1" typeStatus="holotype">
8 spm on mount for SEM (
<specimenCode id="DBA76F637B1D4F7A73EFFE41D19F76C2" box="[451,674,403,427]" collectionCode="MNHN-EA" pageId="52" pageNumber="53">
MNHN-EA
<date id="FFBFE1D87B1D4F7A7060FE41D19F76C2" box="[588,674,403,427]" pageId="52" pageNumber="53" value="07-04-10">041007</date>
</specimenCode>
), same data as the
<typeStatus id="54BA79BA7B1D4F7A715EFE40D0EA76C2" box="[882,983,402,427]" pageId="52" pageNumber="53" type="holotype">holotype</typeStatus>
, [
<collectionCode id="ED105FDD7B1D4F7A71C4FE41D77A76C3" box="[1000,1095,403,426]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="52" pageNumber="53">MNHN</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31B94937B1D4F7872EBFE64D07172E4" lastPageId="54" lastPageNumber="55" pageId="52" pageNumber="53" type="description">
<paragraph id="8BBEC7187B1D4F7A72EBFE64D07376A6" blockId="52.[151,1437,258,2011]" box="[199,846,438,463]" pageId="52" pageNumber="53">
<emphasis id="B9751B0A7B1D4F7A72EBFE64D26476A6" bold="true" box="[199,345,438,463]" pageId="52" pageNumber="53">Description.</emphasis>
Addendum to Schneider &amp; D’Haese (2013).
</paragraph>
<paragraph id="8BBEC7187B1D4F7A72EBFE08D0AF7552" blockId="52.[151,1437,258,2011]" pageId="52" pageNumber="53">
<emphasis id="B9751B0A7B1D4F7A72EBFE08D22E769A" bold="true" box="[199,275,474,499]" pageId="52" pageNumber="53">Adult.</emphasis>
Total length of antenna ~ 100 µm; max width (Ant. III) ~ 22.5 µm; ratio width: length = 0.225; ratio length of Ant. I: Ant. II: Ant. III: Ant. IV ~ 0.36: 0.53: 1: 1. Correction from Schneider &amp; D’Haese (2013): with six ordinary chaetae on Ant. IV (
<figureCitation id="133ADB9D7B1D4F7A702BFDF0D1657552" box="[519,600,546,571]" captionStart="FIGURE 14" captionStartId="47.[151,250,1686,1708]" captionTargetBox="[171,1412,193,1657]" captionTargetId="graphics@47.[771,1399,210,945]" captionTargetPageId="47" captionText="FIGURE 14. Antenna, genus Megalothorax. M. granulosus (A) posterior side, (B) anterior side modified after Schneider &amp; D'Haese (2013), (C) ventral side, (D), dorsal side; M. massoudi after Deharveng 1978, (E) posterior side, (F) anterior side; M. minimus (G) posterior side, (H) anterior side." httpUri="https://zenodo.org/record/846108/files/figure.svg" pageId="52" pageNumber="53">Fig. 14</figureCitation>
A, B). Testis as in
<figureCitation id="133ADB9D7B1D4F7A710BFDF0D0457552" box="[807,888,546,571]" captionStart="FIGURE 20" captionStartId="53.[151,250,1904,1926]" captionTargetBox="[177,1411,194,1880]" captionTargetId="graphics@53.[177,1411,194,1880]" captionTargetPageId="53" captionText="FIGURE 20. Megalothorax nigropunctatus. Male, (A) testis; juvenile (B) chaetotaxy of the body, dorsal side; (C) chaetotaxy of the antenna." httpUri="https://zenodo.org/record/846120/files/figure.svg" pageId="52" pageNumber="53">Fig. 20</figureCitation>
A.
</paragraph>
<paragraph id="8BBEC7187B1D4F7A72EBFD94D09C75A2" blockId="52.[151,1437,258,2011]" pageId="52" pageNumber="53">
<emphasis id="B9751B0A7B1D4F7A72EBFD94D20C7536" bold="true" box="[199,305,582,607]" pageId="52" pageNumber="53">Juvenile.</emphasis>
The difference of chaetae between juvenile and adult chaetotaxy is similar to what have been described for
<taxonomicName id="4C01BC9B7B1D4F7A7318FDBED28675EA" authorityName="Willem, 1900 sensu Schneider &amp; D'Haese" authorityYear="2013" box="[308,443,620,643]" class="Entognatha" family="Neelidae" genus="Megalothorax" kingdom="Animalia" order="Collembola" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="B9751B0A7B1D4F7A7318FDBED28675EA" box="[308,443,620,643]" italics="true" pageId="52" pageNumber="53">M. minimus</emphasis>
</taxonomicName>
with one exception: the chaetae
<emphasis id="B9751B0A7B1D4F7A7119FDB9D06F75EA" box="[821,850,619,643]" italics="true" pageId="52" pageNumber="53">β3</emphasis>
are specific to the adult of
<taxonomicName id="4C01BC9B7B1D4F7A76BDFDBED62A75EA" authorityName="Willem, 1900 sensu Schneider &amp; D'Haese" authorityYear="2013" box="[1169,1303,620,643]" class="Entognatha" family="Neelidae" genus="Megalothorax" kingdom="Animalia" order="Collembola" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="B9751B0A7B1D4F7A76BDFDBED62A75EA" box="[1169,1303,620,643]" italics="true" pageId="52" pageNumber="53">M. minimus</emphasis>
</taxonomicName>
and appear during the post-embryonic development. The supplementary chaetae on
<taxonomicName id="4C01BC9B7B1D4F7A7192FD5DD7B175CF" authorityName=", Schneider &amp; D'Haese" authorityYear="2013" box="[958,1164,655,678]" class="Entognatha" family="Neelidae" genus="Megalothorax" kingdom="Animalia" order="Collembola" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="nigropunctatus">
<emphasis id="B9751B0A7B1D4F7A7192FD5DD7B175CF" box="[958,1164,655,678]" italics="true" pageId="52" pageNumber="53">M. nigropunctatus</emphasis>
</taxonomicName>
in regard to
<taxonomicName id="4C01BC9B7B1D4F7A7734FD5DD6A175CF" authorityName="Willem, 1900 sensu Schneider &amp; D'Haese" authorityYear="2013" box="[1304,1436,655,678]" class="Entognatha" family="Neelidae" genus="Megalothorax" kingdom="Animalia" order="Collembola" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="B9751B0A7B1D4F7A7734FD5DD6A175CF" box="[1304,1436,655,678]" italics="true" pageId="52" pageNumber="53">M. minimus</emphasis>
</taxonomicName>
(on fields of labium and Abd. IV sternum) are present in the juvenile.
</paragraph>
<paragraph id="8BBEC7187B1D4F7A72EBFD04D1867586" blockId="52.[151,1437,258,2011]" box="[199,699,726,751]" pageId="52" pageNumber="53">
<emphasis id="B9751B0A7B1D4F7A72EBFD04D3C37586" bold="true" box="[199,254,726,751]" pageId="52" pageNumber="53">Size.</emphasis>
Length from labrum to anus: ~300 µm.
</paragraph>
<paragraph id="8BBEC7187B1D4F7A72EBFD28D177747A" blockId="52.[151,1437,258,2011]" box="[199,586,762,787]" pageId="52" pageNumber="53">
<emphasis id="B9751B0A7B1D4F7A72EBFD28D253747A" bold="true" box="[199,366,762,787]" pageId="52" pageNumber="53">Pigmentation.</emphasis>
White (in ethanol).
</paragraph>
<paragraph id="8BBEC7187B1D4F7A72EBFCCCD2FF74CA" blockId="52.[151,1437,258,2011]" pageId="52" pageNumber="53">
<emphasis id="B9751B0A7B1D4F7A72EBFCCCD10C745E" bold="true" box="[199,561,798,823]" pageId="52" pageNumber="53">Body shape and segmentation.</emphasis>
Orthognathous. Ratio height: length of head ~ 0.75. Total length of antenna ~ 50 µm; max width (Ant. III) ~ 8 µm; ratio width: length ~ 0.16; ratio length of Ant. I: Ant. II: Ant. III: Ant. IV ~ 0.4: 0.6: 1: 0.88. On each leg, length of coxa&gt; tibiotarsus ~ femur ~ trochanter. Length of manubrium: dp: dd: mucro ~ 0.81: 0.68: 1: 1.
</paragraph>
<paragraph id="8BBEC7187B1D4F7A72EBFC7CD2D87482" blockId="52.[151,1437,258,2011]" pageId="52" pageNumber="53">
<emphasis id="B9751B0A7B1D4F7A72EBFC7CD25F74AE" bold="true" box="[199,354,942,967]" pageId="52" pageNumber="53">Granulation.</emphasis>
Secondary granulation present. Dorsal protuberance
<emphasis id="B9751B0A7B1D4F7A71ECFC7DD0F274AF" box="[960,975,943,966]" italics="true" pageId="52" pageNumber="53">T</emphasis>
present at the posterior extremity of the dorso-median line (
<figureCitation id="133ADB9D7B1D4F7A735DFC00D2FB7482" box="[369,454,978,1003]" captionStart="FIGURE 20" captionStartId="53.[151,250,1904,1926]" captionTargetBox="[177,1411,194,1880]" captionTargetId="graphics@53.[177,1411,194,1880]" captionTargetPageId="53" captionText="FIGURE 20. Megalothorax nigropunctatus. Male, (A) testis; juvenile (B) chaetotaxy of the body, dorsal side; (C) chaetotaxy of the antenna." httpUri="https://zenodo.org/record/846120/files/figure.svg" pageId="52" pageNumber="53">Fig. 20</figureCitation>
B).
</paragraph>
<paragraph id="8BBEC7187B1D4F7A72EBFC24D1C5735A" blockId="52.[151,1437,258,2011]" pageId="52" pageNumber="53">
<emphasis id="B9751B0A7B1D4F7A72EBFC24D2CD7366" bold="true" box="[199,496,1014,1039]" pageId="52" pageNumber="53">Integumentary channels.</emphasis>
Not precisely observed, present dorsally on head, and ventrally on Th. II–Th. III but apparently less developed than in the adult (
<figureCitation id="133ADB9D7B1D4F7A70A8FBC8D1E5735A" box="[644,728,1050,1075]" captionStart="FIGURE 20" captionStartId="53.[151,250,1904,1926]" captionTargetBox="[177,1411,194,1880]" captionTargetId="graphics@53.[177,1411,194,1880]" captionTargetPageId="53" captionText="FIGURE 20. Megalothorax nigropunctatus. Male, (A) testis; juvenile (B) chaetotaxy of the body, dorsal side; (C) chaetotaxy of the antenna." httpUri="https://zenodo.org/record/846120/files/figure.svg" pageId="52" pageNumber="53">Fig. 20</figureCitation>
B).
</paragraph>
<paragraph id="8BBEC7187B1D4F7A72EBFBECD1117312" blockId="52.[151,1437,258,2011]" pageId="52" pageNumber="53">
<emphasis id="B9751B0A7B1D4F7A72EBFBECD20D733E" bold="true" box="[199,304,1086,1111]" pageId="52" pageNumber="53">Chaetae.</emphasis>
The chaetotaxic pattern and morphology are very similar to the adult though the absence of some specific chaetae and wrc (
<figureCitation id="133ADB9D7B1D4F7A7395FBB0D1307312" box="[441,525,1122,1147]" captionStart="FIGURE 20" captionStartId="53.[151,250,1904,1926]" captionTargetBox="[177,1411,194,1880]" captionTargetId="graphics@53.[177,1411,194,1880]" captionTargetPageId="53" captionText="FIGURE 20. Megalothorax nigropunctatus. Male, (A) testis; juvenile (B) chaetotaxy of the body, dorsal side; (C) chaetotaxy of the antenna." httpUri="https://zenodo.org/record/846120/files/figure.svg" pageId="52" pageNumber="53">Fig. 20</figureCitation>
B).
</paragraph>
<paragraph id="8BBEC7187B1D4F7A72EBFB54D7E573AA" blockId="52.[151,1437,258,2011]" pageId="52" pageNumber="53">
<emphasis id="B9751B0A7B1D4F7A72EBFB54D1AA73F6" bold="true" box="[199,663,1158,1183]" pageId="52" pageNumber="53">Wax secretory rods and sensory fields.</emphasis>
With 6 + 6 sensory fields like the adult. Smaller inner chaeta of
<emphasis id="B9751B0A7B1D4F7A7757FB55D6A173F6" box="[1403,1436,1159,1183]" italics="true" pageId="52" pageNumber="53">sf5</emphasis>
absent (the bigger present), inner chaetotaxy of the other sensory fields as in the adult (
<figureCitation id="133ADB9D7B1D4F7A7649FB78D78473AA" box="[1125,1209,1194,1219]" captionStart="FIGURE 20" captionStartId="53.[151,250,1904,1926]" captionTargetBox="[177,1411,194,1880]" captionTargetId="graphics@53.[177,1411,194,1880]" captionTargetPageId="53" captionText="FIGURE 20. Megalothorax nigropunctatus. Male, (A) testis; juvenile (B) chaetotaxy of the body, dorsal side; (C) chaetotaxy of the antenna." httpUri="https://zenodo.org/record/846120/files/figure.svg" pageId="52" pageNumber="53">Fig. 20</figureCitation>
B).
</paragraph>
<paragraph id="8BBEC7187B1D4F7A72EBFB1CD7F7738F" blockId="52.[151,1437,258,2011]" box="[199,1226,1230,1255]" pageId="52" pageNumber="53">
<emphasis id="B9751B0A7B1D4F7A72EBFB1CD1D3738E" bold="true" box="[199,750,1230,1255]" pageId="52" pageNumber="53">Labium, maxilla outer lobe, oral fold, labrum.</emphasis>
All chaetae observed on the adult present.
</paragraph>
<paragraph id="8BBEC7187B1D4F7A72EBFB20D127721E" blockId="52.[151,1437,258,2011]" pageId="52" pageNumber="53">
<emphasis id="B9751B0A7B1D4F7A72EBFB20D2AA7262" bold="true" box="[199,407,1266,1291]" pageId="52" pageNumber="53">Head chaetotaxy.</emphasis>
Postero-dorsal area with 9 + 9 chaetae (the 2 + 2 most anterior chaetae missing); anterodorsal area with nine pairs of chaetae and two unpaired chaetae (2 + 2 chaetae missing in regard to adult near the dorsal edge of
<emphasis id="B9751B0A7B1D4F7A736DFAE9D25C723A" box="[321,353,1339,1363]" italics="true" pageId="52" pageNumber="53">sf1</emphasis>
,
<emphasis id="B9751B0A7B1D4F7A735DFAE9D2B0723A" box="[369,397,1339,1363]" italics="true" pageId="52" pageNumber="53">a0</emphasis>
present); antero-lateral areas with 1 + 1 chaetae (
<emphasis id="B9751B0A7B1D4F7A71E8FAEED0EE723A" box="[964,979,1340,1363]" italics="true" pageId="52" pageNumber="53">L</emphasis>
;
<figureCitation id="133ADB9D7B1D4F7A71CFFAE8D714723A" box="[995,1065,1338,1363]" captionStart="FIGURE 3" captionStartId="15.[151,250,1756,1778]" captionTargetBox="[175,1414,1242,1657]" captionTargetId="graphics@15.[786,1314,1251,1648]" captionTargetPageId="15" captionText="FIGURE 3. Diagram of the dorsal chaetotaxy of the head. (A) Neelides folsomi; (B) Acanthoneelidus pratensis; (C) Neelus murinus; (D) Megalothorax nigropunctatus; (E) Megalothorax perspicillum; (F) Megalothorax granulosus, integumentary channels not represented. In gray, chaetae absent in juvenile." httpUri="https://zenodo.org/record/846092/files/figure.svg" pageId="52" pageNumber="53">Figs 3</figureCitation>
D, 20B). Ventral area with 3 + 3 post-labial mesochaetae (
<figureCitation id="133ADB9D7B1D4F7A739EFA8CD2CA721E" box="[434,503,1374,1399]" captionStart="FIGURE 4" captionStartId="16.[151,250,1717,1739]" captionTargetBox="[794,1353,684,1681]" captionTargetPageId="16" captionText="FIGURE 4. Diagram of the ventral chaetotaxy of the head. (A) Neelides folsomi; (B) Acanthoneelidus pratensis; (C) Neelus murinus; (D) Megalothorax nigropunctatus; (E) Megalothorax minimus; (F) Megalothorax granulosus." httpUri="https://zenodo.org/record/846094/files/figure.svg" pageId="52" pageNumber="53">Fig. 4</figureCitation>
D).
</paragraph>
<paragraph id="8BBEC7187B1D4F7A72EBFA50D0267142" blockId="52.[151,1437,258,2011]" pageId="52" pageNumber="53">
<emphasis id="B9751B0A7B1D4F7A72EBFA50D28672F2" bold="true" box="[199,443,1410,1435]" pageId="52" pageNumber="53">Antenna chaetotaxy.</emphasis>
Ant. I with a chaeta, Ant II with four chaetae, Ant III with 10 ordinary chaetae, two long S-chaetae and two short, swollen S-chaetae (
<emphasis id="B9751B0A7B1D4F7A70BEFA75D1E572D6" box="[658,728,1447,1471]" italics="true" pageId="52" pageNumber="53">S1–S4</emphasis>
, Ant. III sensory organ) and a long S-chaeta
<emphasis id="B9751B0A7B1D4F7A76F4FA75D7DB72D7" box="[1240,1254,1447,1470]" italics="true" pageId="52" pageNumber="53">S</emphasis>
(one missing in regard to adult; Figs 6, 20C). Ant. IV with six ordinary chaetae as the adult; seven S-chaetae including five thin and long (
<emphasis id="B9751B0A7B1D4F7A72F5FA3DD3DA716F" box="[217,231,1519,1542]" italics="true" pageId="52" pageNumber="53">S</emphasis>
, three missing in regard to adult), a short and thin (
<emphasis id="B9751B0A7B1D4F7A7107FA3DD078716F" box="[811,837,1519,1542]" italics="true" pageId="52" pageNumber="53">Sx</emphasis>
) and a long and thick (
<emphasis id="B9751B0A7B1D4F7A7667FA3DD758716F" box="[1099,1125,1519,1542]" italics="true" pageId="52" pageNumber="53">Sy</emphasis>
); two apical and sub-apical sensory rods (
<emphasis id="B9751B0A7B1D4F7A7318F9C6D27F7142" box="[308,322,1556,1579]" italics="true" pageId="52" pageNumber="53">a</emphasis>
,
<emphasis id="B9751B0A7B1D4F7A737CF9C6D2547142" box="[336,361,1556,1579]" italics="true" pageId="52" pageNumber="53">sa</emphasis>
) and a short organite
<emphasis id="B9751B0A7B1D4F7A7071F9C1D1417143" box="[605,636,1555,1578]" italics="true" pageId="52" pageNumber="53">Or</emphasis>
(
<figureCitation id="133ADB9D7B1D4F7A70A0F9C0D1E37142" box="[652,734,1554,1579]" captionStart="FIGURE 20" captionStartId="53.[151,250,1904,1926]" captionTargetBox="[177,1411,194,1880]" captionTargetId="graphics@53.[177,1411,194,1880]" captionTargetPageId="53" captionText="FIGURE 20. Megalothorax nigropunctatus. Male, (A) testis; juvenile (B) chaetotaxy of the body, dorsal side; (C) chaetotaxy of the antenna." httpUri="https://zenodo.org/record/846120/files/figure.svg" pageId="52" pageNumber="53">Fig. 20</figureCitation>
C, 6).
</paragraph>
<paragraph id="8BBEC7187B1D4F7A72EBF9E4D0F37006" blockId="52.[151,1437,258,2011]" pageId="52" pageNumber="53">
<emphasis id="B9751B0A7B1D4F7A72EBF9E4D17A7126" bold="true" box="[199,583,1590,1615]" pageId="52" pageNumber="53">Th. II–Abd. V terga chaetotaxy.</emphasis>
A total of 26 + 26 ordinary chaetae (15 + 15 missing in regard to adult), τchaetae not observed but probably present, 3 + 3 s-chaetae (
<emphasis id="B9751B0A7B1D4F7A716FF989D061711A" box="[835,860,1627,1651]" italics="true" pageId="52" pageNumber="53">s1</emphasis>
,
<emphasis id="B9751B0A7B1D4F7A7140F989D0B9711A" box="[876,900,1627,1651]" italics="true" pageId="52" pageNumber="53">s2</emphasis>
,
<emphasis id="B9751B0A7B1D4F7A71B8F989D091711A" box="[916,940,1627,1651]" italics="true" pageId="52" pageNumber="53">s3</emphasis>
), 4 + 4 sensory fields and 5 + 5 wrc (3 + 3 missing in regard to adult;
<figureCitation id="133ADB9D7B1D4F7A73EEF9ACD12971FE" box="[450,532,1662,1687]" captionStart="FIGURE 20" captionStartId="53.[151,250,1904,1926]" captionTargetBox="[177,1411,194,1880]" captionTargetId="graphics@53.[177,1411,194,1880]" captionTargetPageId="53" captionText="FIGURE 20. Megalothorax nigropunctatus. Male, (A) testis; juvenile (B) chaetotaxy of the body, dorsal side; (C) chaetotaxy of the antenna." httpUri="https://zenodo.org/record/846120/files/figure.svg" pageId="52" pageNumber="53">Fig. 20</figureCitation>
B). Dorso-lateral area of Th. II with 5 + 5 chaetae (
<emphasis id="B9751B0A7B1D4F7A7664F9ADD75871FE" box="[1096,1125,1663,1687]" italics="true" pageId="52" pageNumber="53">a3</emphasis>
,
<emphasis id="B9751B0A7B1D4F7A765EF9ADD7B371FE" box="[1138,1166,1663,1687]" italics="true" pageId="52" pageNumber="53">p2</emphasis>
,
<emphasis id="B9751B0A7B1D4F7A76B0F9ADD78571FE" box="[1180,1208,1663,1687]" italics="true" pageId="52" pageNumber="53">p3</emphasis>
missing in regard to adult). Precoxal areas of Th. II with 1 + 1 s-chaetae (
<emphasis id="B9751B0A7B1D4F7A70C2F971D03571D2" box="[750,776,1699,1723]" italics="true" pageId="52" pageNumber="53">s1</emphasis>
) and 2 + 2 chaetae (
<emphasis id="B9751B0A7B1D4F7A71DDF971D73171D3" box="[1009,1036,1699,1722]" italics="true" pageId="52" pageNumber="53">a7</emphasis>
,
<emphasis id="B9751B0A7B1D4F7A7637F971D70A71D2" box="[1051,1079,1699,1723]" italics="true" pageId="52" pageNumber="53">a8</emphasis>
missing in regard to adult;
<figureCitation id="133ADB9D7B1D4F7A775CF970D38A71B6" captionStart="FIGURE 20" captionStartId="53.[151,250,1904,1926]" captionTargetBox="[177,1411,194,1880]" captionTargetId="graphics@53.[177,1411,194,1880]" captionTargetPageId="53" captionText="FIGURE 20. Megalothorax nigropunctatus. Male, (A) testis; juvenile (B) chaetotaxy of the body, dorsal side; (C) chaetotaxy of the antenna." httpUri="https://zenodo.org/record/846120/files/figure.svg" pageId="52" pageNumber="53">Fig. 20</figureCitation>
B). Dorso-lateral area of Th. III–anterior abdomen with 7 + 7 chaetae (
<emphasis id="B9751B0A7B1D4F7A71FAF915D0CF71B7" box="[982,1010,1735,1758]" italics="true" pageId="52" pageNumber="53">a5</emphasis>
,
<emphasis id="B9751B0A7B1D4F7A762DF915D72171B6" box="[1025,1052,1735,1759]" italics="true" pageId="52" pageNumber="53">p2</emphasis>
,
<emphasis id="B9751B0A7B1D4F7A7600F915D77571B6" box="[1068,1096,1735,1759]" italics="true" pageId="52" pageNumber="53">p3</emphasis>
,
<emphasis id="B9751B0A7B1D4F7A767BF915D74E71B6" box="[1111,1139,1735,1759]" italics="true" pageId="52" pageNumber="53">p4</emphasis>
,
<emphasis id="B9751B0A7B1D4F7A76ADF915D7A371B6" box="[1153,1182,1735,1759]" italics="true" pageId="52" pageNumber="53">α1</emphasis>
,
<emphasis id="B9751B0A7B1D4F7A7681F915D7F771B6" box="[1197,1226,1735,1759]" italics="true" pageId="52" pageNumber="53">α2</emphasis>
,
<emphasis id="B9751B0A7B1D4F7A76F5F915D7C971B6" box="[1241,1268,1735,1759]" italics="true" pageId="52" pageNumber="53">β1</emphasis>
,
<emphasis id="B9751B0A7B1D4F7A772FF915D61D71B6" box="[1283,1312,1735,1759]" italics="true" pageId="52" pageNumber="53">β2</emphasis>
missing in regard to adult) and 3 + 3 wrc (
<emphasis id="B9751B0A7B1D4F7A73D4F939D112706A" box="[504,559,1771,1795]" italics="true" pageId="52" pageNumber="53">wrc2</emphasis>
,
<emphasis id="B9751B0A7B1D4F7A7011F939D149706B" box="[573,628,1771,1794]" italics="true" pageId="52" pageNumber="53">wrc7</emphasis>
missing in regard to adult;
<figureCitation id="133ADB9D7B1D4F7A7186F938D0C0706A" box="[938,1021,1770,1795]" captionStart="FIGURE 20" captionStartId="53.[151,250,1904,1926]" captionTargetBox="[177,1411,194,1880]" captionTargetId="graphics@53.[177,1411,194,1880]" captionTargetPageId="53" captionText="FIGURE 20. Megalothorax nigropunctatus. Male, (A) testis; juvenile (B) chaetotaxy of the body, dorsal side; (C) chaetotaxy of the antenna." httpUri="https://zenodo.org/record/846120/files/figure.svg" pageId="52" pageNumber="53">Fig. 20</figureCitation>
B). Precoxal areas of Th. III with 2 + 2 chaetae (
<emphasis id="B9751B0A7B1D4F7A733FF8DDD212704E" box="[275,303,1807,1831]" italics="true" pageId="52" pageNumber="53">a8</emphasis>
missing in regard to adult) and 1 + 1 wrc (
<emphasis id="B9751B0A7B1D4F7A710EF8DDD064704F" box="[802,857,1807,1830]" italics="true" pageId="52" pageNumber="53">wrc5</emphasis>
missing;
<figureCitation id="133ADB9D7B1D4F7A71E4F8DCD720704E" box="[968,1053,1806,1831]" captionStart="FIGURE 20" captionStartId="53.[151,250,1904,1926]" captionTargetBox="[177,1411,194,1880]" captionTargetId="graphics@53.[177,1411,194,1880]" captionTargetPageId="53" captionText="FIGURE 20. Megalothorax nigropunctatus. Male, (A) testis; juvenile (B) chaetotaxy of the body, dorsal side; (C) chaetotaxy of the antenna." httpUri="https://zenodo.org/record/846120/files/figure.svg" pageId="52" pageNumber="53">Fig. 20</figureCitation>
B). Posterior area of the abdomen (until Abd. V included) with 10 + 10 chaetae (
<emphasis id="B9751B0A7B1D4F7A70B7F8E1D18A7022" box="[667,695,1843,1867]" italics="true" pageId="52" pageNumber="53">η2</emphasis>
missing in regard to adult), 1 + 1 wrc and 1 + 1 s-chaetae (
<emphasis id="B9751B0A7B1D4F7A7764F8E1D65C7022" box="[1352,1377,1843,1867]" italics="true" pageId="52" pageNumber="53">s2</emphasis>
;
<figureCitation id="133ADB9D7B1D4F7A7743F8E0D38A7006" captionStart="FIGURE 20" captionStartId="53.[151,250,1904,1926]" captionTargetBox="[177,1411,194,1880]" captionTargetId="graphics@53.[177,1411,194,1880]" captionTargetPageId="53" captionText="FIGURE 20. Megalothorax nigropunctatus. Male, (A) testis; juvenile (B) chaetotaxy of the body, dorsal side; (C) chaetotaxy of the antenna." httpUri="https://zenodo.org/record/846120/files/figure.svg" pageId="52" pageNumber="53">Fig. 20</figureCitation>
B).
<emphasis id="B9751B0A7B1D4F7A72F1F885D3CB7006" box="[221,246,1879,1903]" italics="true" pageId="52" pageNumber="53">s1</emphasis>
short, tubular with rounded apex,
<emphasis id="B9751B0A7B1D4F7A705BF885D1AD7006" box="[631,656,1879,1903]" italics="true" pageId="52" pageNumber="53">s2</emphasis>
and
<emphasis id="B9751B0A7B1D4F7A70EBF885D1DD7006" box="[711,736,1879,1903]" italics="true" pageId="52" pageNumber="53">s3</emphasis>
spherical (
<figureCitation id="133ADB9D7B1D4F7A7177F884D0927006" box="[859,943,1878,1903]" captionStart="FIGURE 20" captionStartId="53.[151,250,1904,1926]" captionTargetBox="[177,1411,194,1880]" captionTargetId="graphics@53.[177,1411,194,1880]" captionTargetPageId="53" captionText="FIGURE 20. Megalothorax nigropunctatus. Male, (A) testis; juvenile (B) chaetotaxy of the body, dorsal side; (C) chaetotaxy of the antenna." httpUri="https://zenodo.org/record/846120/files/figure.svg" pageId="52" pageNumber="53">Fig. 20</figureCitation>
B).
</paragraph>
<paragraph id="8BBEC7187B1D4F7A72EBF8A8D28B70B2" blockId="52.[151,1437,258,2011]" pageId="52" pageNumber="53">
<emphasis id="B9751B0A7B1D4F7A72EBF8A8D2B370FA" bold="true" box="[199,398,1914,1939]" pageId="52" pageNumber="53">Legs chaetotaxy.</emphasis>
With two chaetae on trochanter I (one missing in regard to adult), no chaeta on coxa II and III (one missing on each in regard to adult) and seven chaetae on femur III (one missing in regard to adult;
<figureCitation id="133ADB9D7B1D4F7A7740F84CD38F70B2" captionStart="FIGURE 21" captionStartId="54.[151,250,1083,1105]" captionTargetBox="[177,1409,193,1056]" captionTargetId="graphics@54.[719,931,197,1056]" captionTargetPageId="54" captionText="FIGURE 21. Megalothorax nigropunctatus, juvenile. Chaetotaxy of the legs (A) leg I anterior side; (B) leg II posterior side; (C) leg III posterior side." httpUri="https://zenodo.org/record/846122/files/figure.svg" pageId="52" pageNumber="53">Figs 21</figureCitation>
A–C,
<tableCitation id="C683F2A37B1D4F7A72DAF810D26870B2" box="[246,341,1986,2011]" captionStart="TABLE 9" captionStartId="32.[151,239,1300,1323]" captionText="TABLE 9. Summary of the chaetotaxic characters on the leg I among various representatives of Neelipleona." pageId="52" pageNumber="53">Tables 9</tableCitation>
,
<tableCitation id="C683F2A37B1D4F7A734DF811D24270B2" box="[353,383,1987,2011]" captionStart="TABLE 10" captionStartId="33.[151,239,152,175]" captionText="TABLE 10. Summary of the chaetotaxic characters on the leg II among various representatives of Neelipleona." pageId="52" pageNumber="53">10</tableCitation>
,
<tableCitation id="C683F2A37B1D4F7A73A7F811D29570B2" box="[395,424,1987,2011]" captionStart="TABLE 11" captionStartId="33.[151,239,692,715]" captionText="TABLE 11. Summary of the chaetotaxic characters on the leg III among various representatives of Neelipleona." pageId="52" pageNumber="53">11</tableCitation>
).
</paragraph>
<caption id="DF7E97907B1C4F7B72BBF8A2D21070CC" httpUri="https://zenodo.org/record/846120/files/figure.svg" pageId="53" pageNumber="54" targetBox="[177,1411,194,1880]" targetPageId="53">
<paragraph id="8BBEC7187B1C4F7B72BBF8A2D21070CC" blockId="53.[151,1436,1904,1957]" pageId="53" pageNumber="54">
<emphasis id="B9751B0A7B1C4F7B72BBF8A2D21D70EF" bold="true" box="[151,288,1904,1926]" pageId="53" pageNumber="54">FIGURE 20.</emphasis>
<taxonomicName id="4C01BC9B7B1C4F7B730BF8A2D16870EF" authorityName=", Schneider &amp; D'Haese" authorityYear="2013" box="[295,597,1904,1926]" class="Entognatha" family="Neelidae" genus="Megalothorax" kingdom="Animalia" order="Collembola" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="nigropunctatus">
<emphasis id="B9751B0A7B1C4F7B730BF8A2D16870EF" box="[295,597,1904,1926]" italics="true" pageId="53" pageNumber="54">Megalothorax nigropunctatus</emphasis>
</taxonomicName>
. Male,
<emphasis id="B9751B0A7B1C4F7B7089F8A2D1F570EF" bold="true" box="[677,712,1904,1926]" pageId="53" pageNumber="54">(A)</emphasis>
testis; juvenile
<emphasis id="B9751B0A7B1C4F7B7146F8A2D0B170EF" bold="true" box="[874,908,1904,1926]" pageId="53" pageNumber="54">(B)</emphasis>
chaetotaxy of the body, dorsal side;
<emphasis id="B9751B0A7B1C4F7B7728F8A2D61A70EF" bold="true" box="[1284,1319,1904,1926]" pageId="53" pageNumber="54">(C)</emphasis>
chaetotaxy of the antenna.
</paragraph>
</caption>
<caption id="DF7E97907B1F4F7872BBFBE9D2A87319" httpUri="https://zenodo.org/record/846122/files/figure.svg" pageId="54" pageNumber="55" targetBox="[177,1409,193,1056]" targetPageId="54">
<paragraph id="8BBEC7187B1F4F7872BBFBE9D2A87319" blockId="54.[151,1436,1083,1136]" pageId="54" pageNumber="55">
<emphasis id="B9751B0A7B1F4F7872BBFBE9D21C7338" bold="true" box="[151,289,1083,1105]" pageId="54" pageNumber="55">FIGURE 21.</emphasis>
<taxonomicName id="4C01BC9B7B1F4F787304FBE9D16A7338" authorityName=", Schneider &amp; D'Haese" authorityYear="2013" box="[296,599,1083,1105]" class="Entognatha" family="Neelidae" genus="Megalothorax" kingdom="Animalia" order="Collembola" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="nigropunctatus">
<emphasis id="B9751B0A7B1F4F787304FBE9D16A7338" box="[296,599,1083,1105]" italics="true" pageId="54" pageNumber="55">Megalothorax nigropunctatus</emphasis>
</taxonomicName>
, juvenile. Chaetotaxy of the legs
<emphasis id="B9751B0A7B1F4F78719FFBE9D0EA7338" bold="true" box="[947,983,1083,1105]" pageId="54" pageNumber="55">(A)</emphasis>
leg I anterior side;
<emphasis id="B9751B0A7B1F4F78768EFBE9D7F97338" bold="true" box="[1186,1220,1083,1105]" pageId="54" pageNumber="55">(B)</emphasis>
leg II posterior side;
<emphasis id="B9751B0A7B1F4F7872BBFB88D3877319" bold="true" box="[151,186,1114,1136]" pageId="54" pageNumber="55">(C)</emphasis>
leg III posterior side.
</paragraph>
</caption>
<paragraph id="8BBEC7187B1F4F7872EBFB4ED00D73DC" blockId="54.[151,1436,1180,1421]" box="[199,816,1180,1205]" pageId="54" pageNumber="55">
<emphasis id="B9751B0A7B1F4F7872EBFB4ED29073DC" bold="true" box="[199,429,1180,1205]" pageId="54" pageNumber="55">Claws morphology.</emphasis>
Similar to the adult (
<figureCitation id="133ADB9D7B1F4F7870B7FB4ED1D673DC" box="[667,747,1180,1205]" captionStart="FIGURE 21" captionStartId="54.[151,250,1083,1105]" captionTargetBox="[177,1409,193,1056]" captionTargetId="graphics@54.[719,931,197,1056]" captionTargetPageId="54" captionText="FIGURE 21. Megalothorax nigropunctatus, juvenile. Chaetotaxy of the legs (A) leg I anterior side; (B) leg II posterior side; (C) leg III posterior side." httpUri="https://zenodo.org/record/846122/files/figure.svg" pageId="54" pageNumber="55">Fig. 21</figureCitation>
A–C).
</paragraph>
<paragraph id="8BBEC7187B1F4F7872EBFB12D6B47394" blockId="54.[151,1436,1180,1421]" pageId="54" pageNumber="55">
<emphasis id="B9751B0A7B1F4F7872EBFB12D20D73B0" bold="true" box="[199,304,1216,1241]" pageId="54" pageNumber="55">Abd. VI.</emphasis>
Tergum anterior row with 2 + 2 chaetae and an unpaired chaeta, posterior row with 2 + 2 chaetae, anal valve with one chaeta, as the adult (
<figureCitation id="133ADB9D7B1F4F787071FB36D18C7394" box="[605,689,1252,1277]" captionStart="FIGURE 20" captionStartId="53.[151,250,1904,1926]" captionTargetBox="[177,1411,194,1880]" captionTargetId="graphics@53.[177,1411,194,1880]" captionTargetPageId="53" captionText="FIGURE 20. Megalothorax nigropunctatus. Male, (A) testis; juvenile (B) chaetotaxy of the body, dorsal side; (C) chaetotaxy of the antenna." httpUri="https://zenodo.org/record/846120/files/figure.svg" pageId="54" pageNumber="55">Fig. 20</figureCitation>
B). Sternum with 5 + 5 chaetae including one on each anal valve.
</paragraph>
<paragraph id="8BBEC7187B1F4F7872EBFADAD7A87248" blockId="54.[151,1436,1180,1421]" box="[199,1173,1288,1313]" pageId="54" pageNumber="55">
<emphasis id="B9751B0A7B1F4F7872EBFADAD2A47248" bold="true" box="[199,409,1288,1313]" pageId="54" pageNumber="55">Abd. IV sternum.</emphasis>
With 3 + 3 chaetae and 2 + 2 neosminthuroid chaetae (as the adult).
</paragraph>
<paragraph id="8BBEC7187B1F4F7872EBFAFED07172E4" blockId="54.[151,1436,1180,1421]" pageId="54" pageNumber="55">
<emphasis id="B9751B0A7B1F4F7872EBFAFED2D7722C" bold="true" box="[199,490,1324,1349]" pageId="54" pageNumber="55">Abdominal appendages.</emphasis>
As the adult: manubrium with 2 + 2 posterior chaetae; dens proximal part with a posterior chaeta, distal part with a posterior chaeta, four posterior spines and three anterior spines; tenaculum with 4 + 4 apical teeth, ventral tube with 2 + 2 apical mesochaetae.
</paragraph>
</subSubSection>
</treatment>
</document>
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