578 lines
67 KiB
XML
578 lines
67 KiB
XML
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<mods:title id="754F86D9B588EC15982FF779627B4A6C">Transfer of two South American ant species from Tapinoma Foerster 1850 to Forelius Emery 1888 (Hymenoptera: Formicidae: Dolichoderinae)</mods:title>
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<mods:date id="8116F5CD518B77A8606EF23A80620578">2021</mods:date>
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<taxonomicName id="4C0CA588024E281B0896FC0AFE1FEB9C" authority="(Forel 1902)" authorityName="Guerrero" authorityYear="2021" baseAuthorityName="Santos-Sousa" baseAuthorityYear="1902" box="[151,495,978,1005]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="heyeri" status="comb. nov.">
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<emphasis id="B9780219024E281B0896FC0AFEA0EB9D" bold="true" box="[151,336,978,1005]" italics="true" pageId="5" pageNumber="433">Forelius heyeri</emphasis>
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(
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<bibRefCitation id="EF9DA3FA024E281B095EFC0AFE17EB9C" author="Forel, A." box="[351,487,978,1004]" pageId="5" pageNumber="433" pagination="293 - 298" refId="ref7411" refString="Forel, A. (1902) [Untitled. Strumigenys Lujae Forel n. sp., Tapinoma Heyeri Forel n. sp.]. In: Wasmann, E. (Ed.), Neues uber die zusammengesetzten Nester und gemischten Kolonien der Ameisen [part]. Allgemeine Zeitschrift fur Entomologie, 7, 293 - 298." type="journal article" year="1902">Forel 1902</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="A24BBF62024E281B09F6FC0BFD8DEB9D" box="[503,637,979,1005]" pageId="5" pageNumber="433" rank="species">comb. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3168D80024E281B0896FC2FFEE9EC61" box="[151,281,1014,1041]" pageId="5" pageNumber="433" type="description">
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<paragraph id="8BB3DE0B024E281B0896FC2FFEE9EC61" blockId="5.[151,637,978,1041]" box="[151,281,1014,1041]" pageId="5" pageNumber="433">
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(
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<figureCitation id="1337C28E024E281B089EFC2FFEE1EC61" box="[159,273,1014,1041]" captionStart="FIGURE 5–10" captionStartId="7.[151,250,1715,1739]" captionTargetBox="[155,1433,381,1691]" captionTargetId="figure-148@7.[155,1433,381,1691]" captionTargetPageId="7" captionText="FIGURE 5–10. Forelius heyeri workers; type specimens of F. heyeri and its junior synonyms (F. heyeri risii and F. pusillus). 5–6) Full face view of the head, and profile of the Forelius heyeri lectotype worker (CASENT0909771). 7–8) Full face view of the head, and profile of the Forelius heyeri risii holotype worker (CASENT0909772). 9–10) Full face view of the head, and profile view of the Forelius pusillus syntype worker (CASENT0911547). Images 5–8 by Zach Lieberman, and images 9–10 by Will Ericson. All images from www.antweb.org." figureDoi="http://doi.org/10.5281/zenodo.4478199" httpUri="https://zenodo.org/record/4478199/files/figure.png" pageId="5" pageNumber="433">Figs 5–10</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3168D80024E281B0896FBE4FEF2EDB3" pageId="5" pageNumber="433" type="reference_group">
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<taxonomicName id="4C0CA588024E281B0896FBE4FE1CEC23" ID-CoL="54T6M" authority="Forel 1902: 296" authorityName="Forel" authorityPageNumber="296" authorityYear="1902" box="[151,492,1083,1107]" class="Insecta" family="Formicidae" genus="Tapinoma" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="heyeri">
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<emphasis id="B9780219024E281B0896FBE4FEB1EC23" box="[151,321,1083,1107]" italics="true" pageId="5" pageNumber="433">Tapinoma heyeri</emphasis>
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<bibRefCitation id="EF9DA3FA024E281B0949FBE3FE1CEC23" author="Forel, A." box="[328,492,1083,1107]" pageId="5" pageNumber="433" pagination="293 - 298" refId="ref7411" refString="Forel, A. (1902) [Untitled. Strumigenys Lujae Forel n. sp., Tapinoma Heyeri Forel n. sp.]. In: Wasmann, E. (Ed.), Neues uber die zusammengesetzten Nester und gemischten Kolonien der Ameisen [part]. Allgemeine Zeitschrift fur Entomologie, 7, 293 - 298." type="journal article" year="1902">Forel 1902: 296</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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||
<typeStatus id="54B760A9024E281B09F7FBE3FDB9EC23" box="[502,585,1083,1107]" pageId="5" pageNumber="433" type="syntype">Syntype</typeStatus>
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<materialsCitation id="3B64D456024E281B0A4EFBE3FE4EEC02" ID-GBIF-Occurrence="3033771305" collectionCode="MHNG" country="Brazil" location="Leopoldo" pageId="5" pageNumber="433" specimenCount="1" stateProvince="Rio Grande do Sul" typeStatus="syntype">
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||
workers [two workers], S„o
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<location id="8ED388D0024E281B0B6EFBE3FC23EC23" LSID="urn:lsid:plazi:treatment:03A56F1D024E28190896FC0AFCE4E915:8ED388D0024E281B0B6EFBE3FC23EC23" box="[879,979,1083,1107]" country="Brazil" name="Leopoldo" pageId="5" pageNumber="433" stateProvince="Rio Grande do Sul">Leopoldo</location>
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,
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<collectingRegion id="49C810E9024E281B0BDCFBE3FB6CEC23" box="[989,1180,1083,1107]" country="Brazil" name="Rio Grande do Sul" pageId="5" pageNumber="433">Rio Grande do Sul</collectingRegion>
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,
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<collectingCountry id="F31B9E9B024E281B0CA6FBE3FB15EC23" box="[1191,1253,1083,1107]" name="Brazil" pageId="5" pageNumber="433">Brazil</collectingCountry>
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(Heyer) [
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<collectionCode id="ED1D46CE024E281B0D47FBE3FA69EC23" box="[1350,1433,1083,1107]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="5" pageNumber="433" type="Museum">MHNG</collectionCode>
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, examined]. One
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||
<typeStatus id="54B760A9024E281B096EFB82FE4EEC02" box="[367,446,1114,1138]" pageId="5" pageNumber="433" type="syntype">syntype</typeStatus>
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</materialsCitation>
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worker (CASENT0909771) here designated
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<emphasis id="B9780219024E281B0B8BFB82FC1BEC02" bold="true" box="[906,1003,1114,1138]" pageId="5" pageNumber="433">
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||
<typeStatus id="54B760A9024E281B0B8BFB82FC1BEC02" box="[906,1003,1114,1138]" pageId="5" pageNumber="433" type="lectotype">lectotype</typeStatus>
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</emphasis>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BB3DE0B024E281B0896FBA1FE6BECDF" blockId="5.[151,1437,1083,1475]" pageId="5" pageNumber="433">
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<treatmentCitationGroup id="AB1CF925024E281B0896FBA1FE6BECDF" pageId="5" pageNumber="433">
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<treatmentCitation id="0AADF81A024E281B0896FBA1FDC6ECE0" author="Forel, A." box="[151,566,1144,1169]" page="58" pageId="5" pageNumber="433" year="1912">
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||
<taxonomicName id="4C0CA588024E281B0896FBA1FDC6ECE0" ID-CoL="5LF2K" authority="Forel 1912: 58" authorityName="Forel" authorityPageNumber="58" authorityYear="1912" box="[151,566,1144,1169]" class="Insecta" family="Formicidae" genus="Tapinoma" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="variety" species="heyeri" variety="risii">
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||
<emphasis id="B9780219024E281B0896FBA1FEB0ECE0" box="[151,320,1144,1168]" italics="true" pageId="5" pageNumber="433">Tapinoma heyeri</emphasis>
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||
<materialsCitation id="3B64D456024E281B0947FBA1FDC6ECE0" ID-GBIF-Occurrence="3033771301" box="[326,566,1144,1169]" collectionCode="MHNG" collectorName="D. Ris" country="Uruguay" location="Montevideo" pageId="5" pageNumber="433" specimenCount="1" stateProvince="Montevideo" typeStatus="holotype">
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||
var.
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||
<emphasis id="B9780219024E281B0970FBA1FE6AECE0" box="[369,410,1145,1168]" italics="true" pageId="5" pageNumber="433">risii</emphasis>
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||
<bibRefCitation id="EF9DA3FA024E281B09A1FBA0FDC6ECE0" author="Forel, A." box="[416,566,1144,1169]" pageId="5" pageNumber="433" pagination="33 - 58" refId="ref7495" refString="Forel, A. (1912) Formicides neotropiques. Part V. 4 me sous-famille Dolichoderinae Forel. Memoires de la Societe Entomologique de Belgique, 20, 33 - 58. https: // doi. org / 10.5962 / bhl. part. 19251" type="journal article" year="1912">Forel 1912: 58</bibRefCitation>
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</materialsCitation>
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</taxonomicName>
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.
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<typeStatus id="54B760A9024E281B0A41FBA0FD6EECE0" box="[576,670,1144,1168]" pageId="5" pageNumber="433" type="holotype">Holotype</typeStatus>
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worker (by monotypy),
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||
<collectingRegion id="49C810E9024E281B0B95FBA0FBE1ECE0" box="[916,1041,1144,1168]" country="Uruguay" name="Montevideo" pageId="5" pageNumber="433">Montevideo</collectingRegion>
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,
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<collectingCountry id="F31B9E9B024E281B0C1BFBA1FB83ECE1" box="[1050,1139,1145,1169]" name="Uruguay" pageId="5" pageNumber="433">Uruguay</collectingCountry>
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(
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<collectorName id="26F9BBDD024E281B0C80FBA1FB32ECE0" box="[1153,1218,1144,1169]" pageId="5" pageNumber="433">D. Ris</collectorName>
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) [MHNG, examined] (CASENT0909772).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BB3DE0B024E281B0896FB6EFD81EC9C" blockId="5.[151,1437,1083,1475]" pageId="5" pageNumber="433">
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<taxonomicName id="4C0CA588024E281B0896FB6EFD56ECBE" authority="Santschi 1922: 374" authorityName="Santschi" authorityPageNumber="374" authorityYear="1922" box="[151,678,1206,1230]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="variety" species="breviscapus" variety="pusilla">
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||
<emphasis id="B9780219024E281B0896FB6EFE99ECBE" box="[151,361,1206,1230]" italics="true" pageId="5" pageNumber="433">Forelius breviscapus</emphasis>
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||
<materialsCitation id="3B64D456024E281B096EFB6EFD56ECBE" ID-GBIF-Occurrence="3033771306" box="[367,678,1206,1230]" collectionCode="NHMB" country="Argentina" location="Cordoba" pageId="5" pageNumber="433" specimenCount="2" typeStatus="syntype">
|
||
var.
|
||
<emphasis id="B9780219024E281B0998FB6EFE2EECBE" box="[409,478,1206,1230]" italics="true" pageId="5" pageNumber="433">pusilla</emphasis>
|
||
Santschi 1922: 374
|
||
</materialsCitation>
|
||
</taxonomicName>
|
||
.
|
||
<typeStatus id="54B760A9024E281B0AAEFB6EFCF2ECBE" box="[687,770,1206,1230]" pageId="5" pageNumber="433" type="syntype">Syntype</typeStatus>
|
||
worker(s),
|
||
<location id="8ED388D0024E281B0B74FB6EFC3FECBE" LSID="urn:lsid:plazi:treatment:03A56F1D024E28190896FC0AFCE4E915:8ED388D0024E281B0B74FB6EFC3FECBE" box="[885,975,1206,1230]" country="Argentina" name="Cordoba" pageId="5" pageNumber="433">Cordoba</location>
|
||
,
|
||
<collectingCountry id="F31B9E9B024E281B0BD7FB6EFBCBECBE" box="[982,1083,1206,1230]" name="Argentina" pageId="5" pageNumber="433">Argentina</collectingCountry>
|
||
(Biraben) [NHMB, AntWeb image of
|
||
<typeStatus id="54B760A9024E281B08E2FB0DFEC2EC9D" box="[227,306,1237,1261]" pageId="5" pageNumber="433" type="syntype">syntype</typeStatus>
|
||
examined, CASENT0911547].
|
||
</paragraph>
|
||
<paragraph id="8BB3DE0B024E281B0896FB2BFD2FED5A" blockId="5.[151,1437,1083,1475]" pageId="5" pageNumber="433">
|
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<treatmentCitationGroup id="AB1CF925024E281B0896FB2BFD2FED5A" pageId="5" pageNumber="433">
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||
<taxonomicName id="4C0CA588024E281B0896FB2BFE1AED7B" authority="Santschi. Status" authorityName="Santschi. Status" authorityYear="1922" box="[151,490,1267,1291]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="pusillus">
|
||
<emphasis id="B9780219024E281B0896FB2BFEB2ED7B" box="[151,322,1267,1291]" italics="true" pageId="5" pageNumber="433">Forelius pusillus</emphasis>
|
||
Santschi. Status
|
||
</taxonomicName>
|
||
as species, senior synonym of
|
||
<taxonomicName id="4C0CA588024E281B0B29FB2BFB5CED7B" ID-CoL="6JGWW" authority="(Kusnezov 1953)" baseAuthorityName="Kusnezov" baseAuthorityYear="1953" box="[808,1196,1267,1291]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="tucumanus">
|
||
<emphasis id="B9780219024E281B0B29FB2BFC02ED7B" box="[808,1010,1267,1291]" italics="true" pageId="5" pageNumber="433">Forelius tucumanus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9DA3FA024E281B0BFEFB2BFB54ED7B" author="Kusnezov, N." box="[1023,1188,1267,1291]" pageId="5" pageNumber="433" pagination="327 - 339" refId="ref7629" refString="Kusnezov, N. (1953) Lista de las hormigas de Tucuman con descripcion de dos nuevos generos (Hymenoptera, Formicidae). Acta Zoologica Lilloana, 13, 327 - 339." type="journal article" year="1953">Kusnezov 1953</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
:
|
||
<treatmentCitation id="0AADF81A024E281B0CB7FB2BFA83ED7B" author="Cuezzo, F." box="[1206,1395,1267,1291]" page="263" pageId="5" pageNumber="433" year="2000">
|
||
<bibRefCitation id="EF9DA3FA024E281B0CB7FB2BFA83ED7B" author="Cuezzo, F." box="[1206,1395,1267,1291]" pageId="5" pageNumber="433" pagination="197 - 277" refId="ref7217" refString="Cuezzo, F. (2000) Revision del genero Forelius (Hymenoptera: Formicidae: Dolichoderinae). Sociobiology, 35, 197 - 277." type="journal article" year="2000">Cuezzo 2000: 263</bibRefCitation>
|
||
</treatmentCitation>
|
||
. Junior synonym of
|
||
<taxonomicName id="4C0CA588024E281B0974FACBFE20ED5A" ID-CoL="54T6M" authorityName="Forel" authorityYear="1902" box="[373,464,1298,1322]" class="Insecta" family="Formicidae" genus="Tapinoma" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="heyeri">
|
||
<emphasis id="B9780219024E281B0974FACBFE74ED5A" box="[373,388,1299,1322]" italics="true" pageId="5" pageNumber="433">F</emphasis>
|
||
.
|
||
<emphasis id="B9780219024E281B0990FACAFE20ED5A" box="[401,464,1298,1322]" italics="true" pageId="5" pageNumber="433">heyeri</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B9780219024E281B09DEFACAFDCCED5A" bold="true" box="[479,572,1298,1322]" pageId="5" pageNumber="433">syn. nov.</emphasis>
|
||
): present study.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BB3DE0B024E281B0896FAE8FA6CED38" blockId="5.[151,1437,1083,1475]" box="[151,1436,1328,1353]" pageId="5" pageNumber="433">
|
||
<taxonomicName id="4C0CA588024E281B0896FAE8FE40ED38" authority="(Forel)" baseAuthorityName="Forel" baseAuthorityYear="1912" box="[151,432,1328,1352]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="subSpecies" species="heyeri" subSpecies="risii">
|
||
<emphasis id="B9780219024E281B0896FAE8FE91ED38" box="[151,353,1328,1352]" italics="true" pageId="5" pageNumber="433">Forelius heyeri risii</emphasis>
|
||
(Forel)
|
||
</taxonomicName>
|
||
. Combination in
|
||
<taxonomicName id="4C0CA588024E281B0A5EFAE8FD44ED38" ID-CoL="4KRL" authorityName="Emery" authorityYear="1888" box="[607,692,1328,1352]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9780219024E281B0A5EFAE8FD44ED38" box="[607,692,1328,1352]" italics="true" pageId="5" pageNumber="433">Forelius</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B9780219024E281B0AC2FAE9FCC6ED39" bold="true" box="[707,822,1329,1353]" pageId="5" pageNumber="433">comb. nov.</emphasis>
|
||
) and junior synonym of
|
||
<taxonomicName id="4C0CA588024E281B0C33FAE9FB7DED38" authorityName="Guerrero" authorityYear="2021" baseAuthorityName="Santos-Sousa" baseAuthorityYear="1902" box="[1074,1165,1328,1352]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="heyeri">
|
||
<emphasis id="B9780219024E281B0C33FAE9FBB1ED38" box="[1074,1089,1329,1352]" italics="true" pageId="5" pageNumber="433">F</emphasis>
|
||
.
|
||
<emphasis id="B9780219024E281B0C4FFAE8FB7DED38" box="[1102,1165,1328,1352]" italics="true" pageId="5" pageNumber="433">heyeri</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B9780219024E281B0C9DFAE9FB09ED39" bold="true" box="[1180,1273,1329,1353]" pageId="5" pageNumber="433">syn. nov.</emphasis>
|
||
): present study.
|
||
</paragraph>
|
||
<paragraph id="8BB3DE0B024E281B0896FA97FB4BEDF6" blockId="5.[151,1437,1083,1475]" pageId="5" pageNumber="433">
|
||
<treatmentCitationGroup id="AB1CF925024E281B0896FA97FB4BEDF6" pageId="5" pageNumber="433">
|
||
<treatmentCitation id="0AADF81A024E281B0896FA97FDA8ED17" author="Kusnezov, N." box="[151,600,1359,1383]" page="330" pageId="5" pageNumber="433" year="1953">
|
||
<taxonomicName id="4C0CA588024E281B0896FA97FDA8ED17" authority="Kusnezov 1953: 330" authorityName="Kusnezov" authorityPageNumber="330" authorityYear="1953" box="[151,600,1359,1383]" class="Insecta" family="Formicidae" genus="Neoforelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="tucumanus">
|
||
<emphasis id="B9780219024E281B0896FA97FE70ED17" box="[151,384,1359,1383]" italics="true" pageId="5" pageNumber="433">Neoforelius tucumanus</emphasis>
|
||
<bibRefCitation id="EF9DA3FA024E281B0984FA88FDA8ED17" author="Kusnezov, N." box="[389,600,1359,1383]" pageId="5" pageNumber="433" pagination="327 - 339" refId="ref7629" refString="Kusnezov, N. (1953) Lista de las hormigas de Tucuman con descripcion de dos nuevos generos (Hymenoptera, Formicidae). Acta Zoologica Lilloana, 13, 327 - 339." type="journal article" year="1953">Kusnezov 1953: 330</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
, figs. 1–12.
|
||
<typeStatus id="54B760A9024E281B0AD3FA97FCC0ED17" box="[722,816,1359,1383]" pageId="5" pageNumber="433" type="holotype">Holotype</typeStatus>
|
||
male, nontype workers, queens:
|
||
<collectingCountry id="F31B9E9B024E281B0C78FA97FB11ED17" box="[1145,1249,1359,1383]" name="Argentina" pageId="5" pageNumber="433">Argentina</collectingCountry>
|
||
,
|
||
<collectingRegion id="49C810E9024E281B0CEBFA97FABBED17" box="[1258,1355,1359,1383]" country="Argentina" name="Tucuman" pageId="5" pageNumber="433">Tucumán</collectingRegion>
|
||
,
|
||
<date id="FFB2F8CB024E281B0D55FA97FF0AEDF6" pageId="5" pageNumber="433" value="1949-12-14">14 Dec 1949</date>
|
||
(Kusnezov) [not examined]. Junior synonym of
|
||
<taxonomicName id="4C0CA588024E281B0AE9FAB6FC02EDF6" authority="Santschi" authorityName="Santschi. Status" authorityYear="1922" box="[744,1010,1390,1414]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="pusillus">
|
||
<emphasis id="B9780219024E281B0AE9FAB6FC62EDF6" box="[744,914,1390,1414]" italics="true" pageId="5" pageNumber="433">Forelius pusillus</emphasis>
|
||
Santschi
|
||
</taxonomicName>
|
||
:
|
||
<treatmentCitation id="0AADF81A024E281B0BFAFAB6FB47EDF6" author="Cuezzo, F." box="[1019,1207,1390,1414]" page="263" pageId="5" pageNumber="433" year="2000">
|
||
<bibRefCitation id="EF9DA3FA024E281B0BFAFAB6FB47EDF6" author="Cuezzo, F." box="[1019,1207,1390,1414]" pageId="5" pageNumber="433" pagination="197 - 277" refId="ref7217" refString="Cuezzo, F. (2000) Revision del genero Forelius (Hymenoptera: Formicidae: Dolichoderinae). Sociobiology, 35, 197 - 277." type="journal article" year="2000">Cuezzo 2000: 263</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BB3DE0B024E281B0896FA54FEF2EDB3" blockId="5.[151,1437,1083,1475]" pageId="5" pageNumber="433">
|
||
<treatmentCitationGroup id="AB1CF925024E281B0896FA54FEF2EDB3" pageId="5" pageNumber="433">
|
||
<taxonomicName id="4C0CA588024E281B0896FA54FE2FEDD5" ID-CoL="6JGWW" authority="(Kusnezov)" baseAuthorityName="Kusnezov" baseAuthorityYear="1953" box="[151,479,1420,1445]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="tucumanus">
|
||
<emphasis id="B9780219024E281B0896FA54FE90EDD4" box="[151,352,1420,1444]" italics="true" pageId="5" pageNumber="433">Forelius tucumanus</emphasis>
|
||
(Kusnezov)
|
||
</taxonomicName>
|
||
. Combination in
|
||
<treatmentCitation id="0AADF81A024E281B0A8DFA54FC58EDD4" author="Shattuck, S. O." box="[652,936,1420,1445]" page="95" pageId="5" pageNumber="433" year="1992">
|
||
<taxonomicName id="4C0CA588024E281B0A8DFA54FC58EDD4" ID-CoL="4KRL" authority=": Shattuck 1992: 95" authorityName=": Shattuck" authorityPageNumber="95" authorityYear="1992" box="[652,936,1420,1445]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9780219024E281B0A8DFA54FD11EDD4" box="[652,737,1420,1444]" italics="true" pageId="5" pageNumber="433">Forelius</emphasis>
|
||
:
|
||
<bibRefCitation id="EF9DA3FA024E281B0AEFFA54FC58EDD4" author="Shattuck, S. O." box="[750,936,1420,1445]" pageId="5" pageNumber="433" pagination="1 - 181" refId="ref7795" refString="Shattuck, S. O. (1992) Generic revision of the ant subfamily Dolichoderinae. Sociobiology, 21, 1 - 181." type="journal article" year="1992">Shattuck 1992: 95</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
. Junior synonym of
|
||
<taxonomicName id="4C0CA588024E281B0C74FA55FB20EDD4" authorityName="Guerrero" authorityYear="2021" baseAuthorityName="Santos-Sousa" baseAuthorityYear="1902" box="[1141,1232,1420,1444]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="heyeri">
|
||
<emphasis id="B9780219024E281B0C74FA55FB74EDD4" box="[1141,1156,1421,1444]" italics="true" pageId="5" pageNumber="433">F</emphasis>
|
||
.
|
||
<emphasis id="B9780219024E281B0C90FA54FB20EDD4" box="[1169,1232,1420,1444]" italics="true" pageId="5" pageNumber="433">heyeri</emphasis>
|
||
</taxonomicName>
|
||
: present study (
|
||
<emphasis id="B9780219024E281B0D70FA55FF03EDB3" bold="true" pageId="5" pageNumber="433">syn. nov.</emphasis>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3168D80024E281B0896FA29FA72EE07" pageId="5" pageNumber="433" type="materials_examined">
|
||
<paragraph id="8BB3DE0B024E281B0896FA29FB81EE5F" blockId="5.[151,1437,1521,2016]" pageId="5" pageNumber="433">
|
||
<emphasis id="B9780219024E281B0896FA29FDE0EE7B" bold="true" box="[151,528,1521,1547]" pageId="5" pageNumber="433">Additional Material examined</emphasis>
|
||
.
|
||
<materialsCitation id="3B64D456024E281B0A27FA2AFB9DEE5F" ID-GBIF-Occurrence="3033771302" collectingDate="2016-06-07" collectionCode="CBUMAG" collectorName="RJ Guerrero" country="Uruguay" elevation="4" latitude="-34.284054" location="Lavalleja" longLatPrecision="1" longitude="-55.257694" pageId="5" pageNumber="433" specimenCount="1" stateProvince="Lavalleja">
|
||
<collectingCountry id="F31B9E9B024E281B0A27FA2AFD7CEE7C" box="[550,652,1522,1548]" name="Uruguay" pageId="5" pageNumber="433">Uruguay</collectingCountry>
|
||
:
|
||
<collectingRegion id="49C810E9024E281B0A9EFA29FCFBEE7B" box="[671,779,1521,1547]" country="Uruguay" name="Lavalleja" pageId="5" pageNumber="433">Lavalleja</collectingRegion>
|
||
;
|
||
<geoCoordinate id="EE38B8CC024E281B0B1FFA29FC45EE7B" box="[798,949,1521,1547]" degrees="34" direction="south" minutes="17" orientation="latitude" pageId="5" pageNumber="433" precision="1" seconds="02.6" value="-34.284054">34º17’02.6”S</geoCoordinate>
|
||
<geoCoordinate id="EE38B8CC024E281B0BC5FA29FB98EE7C" box="[964,1128,1521,1548]" degrees="55" direction="west" minutes="15" orientation="longitude" pageId="5" pageNumber="433" precision="1" seconds="27.7" value="-55.257694">55º15’27.7”W</geoCoordinate>
|
||
;
|
||
<quantity id="4CF473EE024E281B0C7AFA29FB41EE7C" box="[1147,1201,1521,1548]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="433" unit="m" value="4.0">
|
||
<elevation id="00213938024E281B0C7AFA29FB41EE7C" box="[1147,1201,1521,1548]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="433" unit="m" value="4.0">4 m</elevation>
|
||
</quantity>
|
||
; manual collection;
|
||
<date id="FFB2F8CB024E281B0896F9CDFED3EE5F" box="[151,291,1557,1583]" pageId="5" pageNumber="433" value="2016-06-07">
|
||
<collectingDate id="EFF60123024E281B0896F9CDFED3EE5F" box="[151,291,1557,1583]" pageId="5" pageNumber="433" value="2016-06-07">07.Jun.2016</collectingDate>
|
||
</date>
|
||
; foraging on coarse sand dune;
|
||
<collectorName id="26F9BBDD024E281B0A87F9CEFCFFEE5F" box="[646,783,1557,1584]" pageId="5" pageNumber="433">RJ Guerrero</collectorName>
|
||
RJGF-0185 (3w) (
|
||
<collectionCode id="ED1D46CE024E281B0BE5F9CDFB96EE5F" box="[996,1126,1557,1583]" pageId="5" pageNumber="433">CBUMAG</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB3DE0B024E281B08C6F9E1FA72EE07" blockId="5.[151,1437,1521,2016]" pageId="5" pageNumber="433">
|
||
<emphasis id="B9780219024E281B08C6F9E1FBD8EE23" bold="true" box="[199,1064,1593,1619]" pageId="5" pageNumber="433">Additional specimen images examined, provided by www.antweb.org</emphasis>
|
||
.
|
||
<materialsCitation id="3B64D456024E281B0C3CF9E2FA8FEE07" ID-GBIF-Occurrence="3033771307" collectingDate="2002-11-20" collectionCode="ALWC" collectorName="A. L. Wild" country="Paraguay" elevation="212" latitude="-24.205587" location="Tendal" longLatPrecision="1" longitude="-55.56588" pageId="5" pageNumber="433" specimenCode="CASENT0173739" specimenCount="1" stateProvince="Canindeyu">
|
||
<collectingCountry id="F31B9E9B024E281B0C3CF9E2FB5AEE24" box="[1085,1194,1594,1620]" name="Paraguay" pageId="5" pageNumber="433">Paraguay</collectingCountry>
|
||
:
|
||
<collectingRegion id="49C810E9024E281B0CBAF9E1FAC7EE23" box="[1211,1335,1593,1619]" country="Paraguay" name="Canindeyu" pageId="5" pageNumber="433">Canindeyú</collectingRegion>
|
||
,
|
||
<location id="8ED388D0024E281B0D48F9E1FA68EE23" LSID="urn:lsid:plazi:treatment:03A56F1D024E28190896FC0AFCE4E915:8ED388D0024E281B0D48F9E1FA68EE23" box="[1353,1432,1593,1619]" country="Paraguay" latitude="-24.205587" longLatPrecision="1" longitude="-55.56588" name="Tendal" pageId="5" pageNumber="433" stateProvince="Canindeyu">Tendal</location>
|
||
,
|
||
<geoCoordinate id="EE38B8CC024E281B0896F985FEBBEE07" box="[151,331,1629,1655]" degrees="24" direction="south" minutes="12" orientation="latitude" pageId="5" pageNumber="433" precision="1" seconds="20.112" value="-24.205587">24°12’20.112”S</geoCoordinate>
|
||
<geoCoordinate id="EE38B8CC024E281B0952F985FDE5EE08" box="[339,533,1629,1656]" degrees="55" direction="west" minutes="33" orientation="longitude" pageId="5" pageNumber="433" precision="1" seconds="57.167" value="-55.56588">55°33’57.167”W</geoCoordinate>
|
||
;
|
||
<quantity id="4CF473EE024E281B0A20F985FD94EE07" box="[545,612,1629,1655]" metricMagnitude="2" metricUnit="m" metricValue="2.12" pageId="5" pageNumber="433" unit="m" value="212.0">
|
||
<elevation id="00213938024E281B0A20F985FD94EE07" box="[545,612,1629,1655]" metricMagnitude="2" metricUnit="m" metricValue="2.12" pageId="5" pageNumber="433" unit="m" value="212.0">212m</elevation>
|
||
</quantity>
|
||
;
|
||
<date id="FFB2F8CB024E281B0A6EF985FCF3EE07" box="[623,771,1629,1655]" pageId="5" pageNumber="433" value="2002-11-20">
|
||
<collectingDate id="EFF60123024E281B0A6EF985FCF3EE07" box="[623,771,1629,1655]" pageId="5" pageNumber="433" value="2002-11-20">20.Nov.2002</collectingDate>
|
||
</date>
|
||
.
|
||
<collectorName id="26F9BBDD024E281B0B0DF986FC8DEE07" box="[780,893,1629,1656]" pageId="5" pageNumber="433">A.L. Wild</collectorName>
|
||
AWL1710 (1w) (
|
||
<collectionCode id="ED1D46CE024E281B0C43F985FB69EE07" box="[1090,1177,1629,1655]" pageId="5" pageNumber="433">ALWC</collectionCode>
|
||
).
|
||
<specimenCode id="DBAA7670024E281B0CADF985FA8FEE07" box="[1196,1407,1629,1655]" collectionCode="ALWC" pageId="5" pageNumber="433">CASENT0173739</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3168D80024E281B08C6F959FB58EEEB" box="[199,1192,1665,1692]" pageId="5" pageNumber="433" type="description">
|
||
<paragraph id="8BB3DE0B024E281B08C6F959FB58EEEB" blockId="5.[151,1437,1521,2016]" box="[199,1192,1665,1692]" pageId="5" pageNumber="433">
|
||
<emphasis id="B9780219024E281B08C6F959FE2EEEEB" bold="true" box="[199,478,1665,1691]" pageId="5" pageNumber="433">Worker measurements.</emphasis>
|
||
<typeStatus id="54B760A9024E281B09E4F95AFDA9EEEC" box="[485,601,1666,1692]" pageId="5" pageNumber="433" type="lectotype">Lectotype</typeStatus>
|
||
: HL 0.57, HW 0.51, SL 0.47, WL 0.69, CI 90, SI 92.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3168D80024E281B08C6F97DFA9AEECF" box="[199,1386,1701,1728]" pageId="5" pageNumber="433" type="materials_examined">
|
||
<paragraph id="8BB3DE0B024E281B08C6F97DFA9AEECF" blockId="5.[151,1437,1521,2016]" box="[199,1386,1701,1728]" pageId="5" pageNumber="433">
|
||
<emphasis id="B9780219024E281B08C6F97DFE8AEECF" bold="true" box="[199,378,1701,1727]" pageId="5" pageNumber="433">Other material</emphasis>
|
||
(n= 7): HL 0.45–0.53, HW 0.39–0.51, SL 0.32–0.43, WL 0.48–0.68, CI 83–96, SI 80–91.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3168D80024E281B08C6F911FC80EF3F" pageId="5" pageNumber="433" type="diagnosis">
|
||
<paragraph id="8BB3DE0B024E281B08C6F911FC80EF3F" blockId="5.[151,1437,1521,2016]" pageId="5" pageNumber="433">
|
||
<emphasis id="B9780219024E281B08C6F911FECAEE93" bold="true" box="[199,314,1737,1763]" pageId="5" pageNumber="433">Diagnosis</emphasis>
|
||
. Posterior cephalic margin completely straight. Mandible with five teeth and one denticle. Scape much shorter (±
|
||
<quantity id="4CF473EE024E281B0954F935FE4BEF78" box="[341,443,1773,1800]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="5" pageNumber="433" unit="mm" value="0.43">0.43 mm</quantity>
|
||
) than that of
|
||
<taxonomicName id="4C0CA588024E281B0A54F936FD02EF77" authorityName="Guerrero" authorityYear="2021" baseAuthorityName="Guerrero" baseAuthorityYear="2021" box="[597,754,1774,1800]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="antarcticus">
|
||
<emphasis id="B9780219024E281B0A54F936FD96EF77" box="[597,614,1774,1799]" italics="true" pageId="5" pageNumber="433">F</emphasis>
|
||
.
|
||
<emphasis id="B9780219024E281B0A74F936FD02EF77" box="[629,754,1774,1799]" italics="true" pageId="5" pageNumber="433">antarcticus</emphasis>
|
||
</taxonomicName>
|
||
(SL Ż
|
||
<quantity id="4CF473EE024E281B0B42F935FC5DEF78" box="[835,941,1773,1800]" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" pageId="5" pageNumber="433" unit="mm" value="0.69">0.69 mm</quantity>
|
||
), below or barely reaching the posterior cephalic margin by a length less than the width of the pedicel. Pronotum with two thick dark erect setae that contrast with the light integument, each seta arising from a distinct fovea.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3168D80024E281808C6F881FCA6EB9D" lastPageId="6" lastPageNumber="434" pageId="5" pageNumber="433" type="description">
|
||
<paragraph id="8BB3DE0B024E281808C6F881FB3EEA4D" blockId="5.[151,1437,1521,2016]" lastBlockId="6.[151,1437,151,2014]" lastPageId="6" lastPageNumber="434" pageId="5" pageNumber="433">
|
||
<emphasis id="B9780219024E281B08C6F881FEA9EF03" bold="true" box="[199,345,1881,1907]" pageId="5" pageNumber="433">Description.</emphasis>
|
||
Head slightly longer than broad to subquadrate, with subparallel sides, slightly straight above the middle of the eyes to almost the most posterior region of the head where the margin curves inward; posterolateral corners blunt but differentiated; dorsal and ventral faces flattened, surface finely punctate and covered with abundant, very short, decumbent setae; four long erect setae arising from between the frontal carinae; eyes flattened, relatively large (~
|
||
<quantity id="4CF473EE024D28180968FF4FFE21E8C1" box="[361,465,151,177]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="434" unit="mm" value="0.16">0.16 mm</quantity>
|
||
), and located within the head capsule some distance from the lateral margin in frontal view; maxillary palps long and digitiform, reaching the middle of the occipital foramen, 2nd to 6th segment almost the same length. Mandibles relatively wide, with well differentiated semiparallel external and basal margins; masticatory margin reddish and armed with a very long, pointed and falcate apical tooth, followed by a subapical tooth ¼ of the length of the apical, a third tooth of equal size to the subapical, a fourth tooth followed by robust denticle, a diastema crenulate, and usually a small basal tooth; mandibular surface smooth and opaque, with widely spaced setae and coarse foveae. Clypeus with the anterior margin entire and convex; with setae of relatively equal length directed downwards, a very long median clypeal seta (
|
||
<quantity id="4CF473EE024D28180AFEFE4BFC95E9DD" box="[767,869,403,429]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="434" unit="mm" value="0.14">0.14 mm</quantity>
|
||
in the
|
||
<typeStatus id="54B760A9024D28180BB5FE4BFBD9E9DD" box="[948,1065,403,429]" pageId="6" pageNumber="434" type="lectotype">Lectotype</typeStatus>
|
||
), curved downwards just beyond the closed mandibles and flanked by two shorter frontally directed setae. Antennae with scapes slightly widening distally; pedicel elongated, almost twice the length of the second flagellomere, 2nd to 7th flagellomeres semirectangular and equal in length, the remaining four increasing in length and width, with the last flagellomere very enlarged and almost twice the length of the preceding one; abundant short, decumbent and separate setae.
|
||
</paragraph>
|
||
<paragraph id="8BB3DE0B024D281808C6FD9FFB86EB49" blockId="6.[151,1437,151,2014]" pageId="6" pageNumber="434">In lateral view, promesonotum continuous, slightly elevated in the region where these sclerites join, mesonotum flat, in lateral view metanotal groove marked but not deeply impressed, mesonotum and propodeum almost continuous with a slight sinuosity where meet them (in dorsal view fully fused); dorsal face of the propodeum almost the same length as the propodeal declivity, without an angle that differentiates the two faces; spiracle large and circular, aligned with upper margin of propodeal declivity; dorsum of head with long, thin appressed hairs, integument with very short, spaced hairs; mesopleuron and part of the metapleuron smooth and shiny, the latter with a few long yellow hairs separated from each other, extending over the surface of the metapleural bulla.</paragraph>
|
||
<paragraph id="8BB3DE0B024D281808C6FC9BFD3BEBB9" blockId="6.[151,1437,151,2014]" pageId="6" pageNumber="434">Petiolar scale very small, projecting forward, generally concealed by the front of the first gastral segment; gaster surface finely punctate, pilosity like that of the head; first and second gastral terga with no erect setae, third and fourth gastral tergites with four and six thick erect setae, respectively; gastral sternites 1–4 with 3 to 4 erect setae on each plate, the last sternum with two central setae.</paragraph>
|
||
<paragraph id="8BB3DE0B024D281808C6FC0BFCA6EB9D" blockId="6.[151,1437,151,2014]" box="[199,854,979,1005]" pageId="6" pageNumber="434">Dull yellow to light yellowish brown throughout the body.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3168D80024D281908C6FC2EFCE4E915" lastPageId="7" lastPageNumber="435" pageId="6" pageNumber="434" type="discussion">
|
||
<paragraph id="8BB3DE0B024D281808C6FC2EFE1CECD1" blockId="6.[151,1437,151,2014]" pageId="6" pageNumber="434">
|
||
<emphasis id="B9780219024D281808C6FC2EFEB8EC60" bold="true" box="[199,328,1014,1040]" pageId="6" pageNumber="434">Comments</emphasis>
|
||
. The name
|
||
<taxonomicName id="4C0CA588024D281809D1FC2FFD7DEC61" authorityName="Forel" authorityYear="1902" box="[464,653,1015,1041]" class="Insecta" family="Formicidae" genus="Tapinoma" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="species" species="heyeri">
|
||
<emphasis id="B9780219024D281809D1FC2FFD7DEC61" box="[464,653,1015,1041]" italics="true" pageId="6" pageNumber="434">Tapinoma heyeri</emphasis>
|
||
</taxonomicName>
|
||
is transferred to the genus
|
||
<taxonomicName id="4C0CA588024D28180BC4FC2FFBD3EC61" authorityName=": Shattuck" authorityYear="1992" box="[965,1059,1015,1041]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9780219024D28180BC4FC2FFBD3EC61" box="[965,1059,1015,1041]" italics="true" pageId="6" pageNumber="434">Forelius</emphasis>
|
||
</taxonomicName>
|
||
as the
|
||
<typeStatus id="54B760A9024D28180C74FC2FFB55EC61" box="[1141,1189,1015,1041]" pageId="6" pageNumber="434">type</typeStatus>
|
||
workers have a completely convex and entire anterior margin of the clypeus, and they have three long setae on the anterior clypeal vertex which are equal in length to the semi-closed mandibles. Furthermore, the masticatory margin of the mandible bears five conspicuous teeth and two denticles, and the basal margin is smooth and differentiated from the masticatory margin by an acute angle.
|
||
</paragraph>
|
||
<paragraph id="8BB3DE0B024D281808C6FB73FD19EDB1" blockId="6.[151,1437,151,2014]" pageId="6" pageNumber="434">
|
||
Although the
|
||
<typeStatus id="54B760A9024D28180962FB73FE63ECB5" box="[355,403,1195,1221]" pageId="6" pageNumber="434">type</typeStatus>
|
||
specimens of
|
||
<taxonomicName id="4C0CA588024D28180A39FB73FD14ECB5" authorityName="Guerrero" authorityYear="2021" baseAuthorityName="Santos-Sousa" baseAuthorityYear="1902" box="[568,740,1195,1221]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="species" species="heyeri">
|
||
<emphasis id="B9780219024D28180A39FB73FD14ECB5" box="[568,740,1195,1221]" italics="true" pageId="6" pageNumber="434">Forelius heyeri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0CA588024D28180B1CFB73FC48ECB4" baseAuthorityName="Forel" baseAuthorityYear="1912" box="[797,952,1195,1221]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="subSpecies" species="heyeri" subSpecies="risii">
|
||
<emphasis id="B9780219024D28180B1CFB73FCDEECB4" box="[797,814,1195,1220]" italics="true" pageId="6" pageNumber="434">F</emphasis>
|
||
.
|
||
<emphasis id="B9780219024D28180B3CFB73FC48ECB4" box="[829,952,1195,1221]" italics="true" pageId="6" pageNumber="434">heyeri risii</emphasis>
|
||
</taxonomicName>
|
||
differ in size, they both share two pairs of long thick setae on the dorsum of the head, the first pair on the oblique-lateral side of the posterior portion of the clypeus, and the second pair near to the upper end of the frontal carinae. In both cases the setae are directed anteriorly. Both taxa have a continuous mesosomal profile, with the posterior end of the propodeum rounding evenly into the oblique posterior face, and short and appressed setae on the surface of the body. Two erect pronotal setae are visible in
|
||
<taxonomicName id="4C0CA588024D28180909FA87FE9FED09" authorityName="Guerrero" authorityYear="2021" baseAuthorityName="Santos-Sousa" baseAuthorityYear="1902" box="[264,367,1375,1401]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="species" species="heyeri">
|
||
<emphasis id="B9780219024D28180909FA87FEE9ED08" box="[264,281,1375,1400]" italics="true" pageId="6" pageNumber="434">F</emphasis>
|
||
.
|
||
<emphasis id="B9780219024D28180928FA87FE9FED09" box="[297,367,1375,1401]" italics="true" pageId="6" pageNumber="434">heyeri</emphasis>
|
||
</taxonomicName>
|
||
, but in the
|
||
<typeStatus id="54B760A9024D281809F2FA87FDD3ED09" box="[499,547,1375,1401]" pageId="6" pageNumber="434">type</typeStatus>
|
||
specimen of
|
||
<taxonomicName id="4C0CA588024D28180ABCFA87FCA9ED08" baseAuthorityName="Forel" baseAuthorityYear="1912" box="[701,857,1375,1401]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="subSpecies" species="heyeri" subSpecies="risii">
|
||
<emphasis id="B9780219024D28180ABCFA87FD3EED08" box="[701,718,1375,1400]" italics="true" pageId="6" pageNumber="434">F</emphasis>
|
||
.
|
||
<emphasis id="B9780219024D28180ADFFA87FCA9ED08" box="[734,857,1375,1401]" italics="true" pageId="6" pageNumber="434">heyeri risii</emphasis>
|
||
</taxonomicName>
|
||
these are difficult to evaluate (CASENT0909772). Three workers collected in
|
||
<collectingRegion id="49C810E9024D281809CBFA5BFDC2EDED" box="[458,562,1411,1437]" country="Uruguay" name="Lavalleja" pageId="6" pageNumber="434">Lavalleja</collectingRegion>
|
||
(
|
||
<collectingCountry id="F31B9E9B024D28180A40FA5BFD56EDED" box="[577,678,1411,1437]" name="Uruguay" pageId="6" pageNumber="434">Uruguay</collectingCountry>
|
||
) are relatively similar to the
|
||
<typeStatus id="54B760A9024D28180BEBFA5BFBEAEDED" box="[1002,1050,1411,1437]" pageId="6" pageNumber="434">type</typeStatus>
|
||
<taxonomicName id="4C0CA588024D28180C23FA5BFB4BEDEC" baseAuthorityName="Forel" baseAuthorityYear="1912" box="[1058,1211,1411,1437]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="subSpecies" species="heyeri" subSpecies="risii">
|
||
<emphasis id="B9780219024D28180C23FA5BFBC3EDEC" box="[1058,1075,1411,1436]" italics="true" pageId="6" pageNumber="434">F</emphasis>
|
||
.
|
||
<emphasis id="B9780219024D28180C40FA5BFB4BEDEC" box="[1089,1211,1411,1437]" italics="true" pageId="6" pageNumber="434">heyeri risii</emphasis>
|
||
</taxonomicName>
|
||
but all bear the pair of erect pronotal setae found on the
|
||
<typeStatus id="54B760A9024D28180A2AFA7FFDABEDB1" box="[555,603,1447,1473]" pageId="6" pageNumber="434">type</typeStatus>
|
||
of
|
||
<taxonomicName id="4C0CA588024D28180A81FA7FFD12EDB1" authorityName="Guerrero" authorityYear="2021" baseAuthorityName="Santos-Sousa" baseAuthorityYear="1902" box="[640,738,1447,1473]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="species" species="heyeri">
|
||
<emphasis id="B9780219024D28180A81FA7FFD12EDB1" box="[640,738,1447,1473]" italics="true" pageId="6" pageNumber="434">F. heyeri</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB3DE0B024D281808C6FA13FC27EF59" blockId="6.[151,1437,151,2014]" pageId="6" pageNumber="434">
|
||
There are morphometric differences between the specimens associated with both names (
|
||
<figureCitation id="1337C28E024D28180CB6FA13FAF3ED95" box="[1207,1283,1483,1509]" captionStart="FIGURE 11" captionStartId="8.[151,250,928,952]" captionTargetBox="[192,1403,193,889]" captionTargetId="figure-16@8.[169,1418,181,904]" captionTargetPageId="8" captionText="FIGURE 11. Bivariate plot relating the head length (HL) to both the head width (HW, left) and the scape length (SL, right) of Forelius heyeri. Small circles: three workers from a series recently collected in Uruguay. Medium circle: second syntype of F. heyeri. Large circle: F. heyeri lectotype. Square: F. heyeri risii lectotype. Small triangle: worker from Paraguay identified as F. pusillus on AntWeb (CASENT0173739). Large triangle: syntype of F. pusillus." figureDoi="http://doi.org/10.5281/zenodo.4478201" httpUri="https://zenodo.org/record/4478201/files/figure.png" pageId="6" pageNumber="434">Fig 11</figureCitation>
|
||
). The
|
||
<typeStatus id="54B760A9024D28180D4CFA13FA8DED95" box="[1357,1405,1483,1509]" pageId="6" pageNumber="434">type</typeStatus>
|
||
of
|
||
<taxonomicName id="4C0CA588024D28180896FA37FEDCEE78" baseAuthorityName="Forel" baseAuthorityYear="1912" box="[151,300,1519,1545]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="subSpecies" species="heyeri" subSpecies="risii">
|
||
<emphasis id="B9780219024D28180896FA37FEDCEE78" box="[151,300,1519,1545]" italics="true" pageId="6" pageNumber="434">F. heyeri risii</emphasis>
|
||
</taxonomicName>
|
||
is much smaller than the
|
||
<typeStatus id="54B760A9024D28180A48FA37FD89EE79" box="[585,633,1519,1545]" pageId="6" pageNumber="434">type</typeStatus>
|
||
of
|
||
<taxonomicName id="4C0CA588024D28180A9CFA37FCF2EE79" authorityName="Guerrero" authorityYear="2021" baseAuthorityName="Santos-Sousa" baseAuthorityYear="1902" box="[669,770,1519,1545]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="species" species="heyeri">
|
||
<emphasis id="B9780219024D28180A9CFA37FD5EEE78" box="[669,686,1519,1544]" italics="true" pageId="6" pageNumber="434">F</emphasis>
|
||
.
|
||
<emphasis id="B9780219024D28180ABDFA37FCF2EE79" box="[700,770,1519,1545]" italics="true" pageId="6" pageNumber="434">heyeri</emphasis>
|
||
</taxonomicName>
|
||
(WL = 0.49 vs. 0.69, respectively). The head of
|
||
<taxonomicName id="4C0CA588024D28180D1FFA37FA73EE79" authorityName="Guerrero" authorityYear="2021" baseAuthorityName="Santos-Sousa" baseAuthorityYear="1902" box="[1310,1411,1519,1545]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="species" species="heyeri">
|
||
<emphasis id="B9780219024D28180D1FFA37FADFEE78" box="[1310,1327,1519,1544]" italics="true" pageId="6" pageNumber="434">F</emphasis>
|
||
.
|
||
<emphasis id="B9780219024D28180D3CFA37FA73EE79" box="[1341,1411,1519,1545]" italics="true" pageId="6" pageNumber="434">heyeri</emphasis>
|
||
</taxonomicName>
|
||
is broader (CIŻ 90), with subparallel sides converging slightly anteriorly, and a straight vertex with an almost impeceptible medial depression, while the head of the worker of
|
||
<taxonomicName id="4C0CA588024D28180B32F9EFFC3CEE20" baseAuthorityName="Forel" baseAuthorityYear="1912" box="[819,972,1591,1617]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="subSpecies" species="heyeri" subSpecies="risii">
|
||
<emphasis id="B9780219024D28180B32F9EFFCB4EE20" box="[819,836,1591,1616]" italics="true" pageId="6" pageNumber="434">F</emphasis>
|
||
.
|
||
<emphasis id="B9780219024D28180B53F9EFFC3CEE20" box="[850,972,1591,1617]" italics="true" pageId="6" pageNumber="434">heyeri risii</emphasis>
|
||
</taxonomicName>
|
||
is narrower (CI= 83), with convex lateral margins, continuing in a straight vertex. Both specimens also differ in the length of the scape relative to head width. In
|
||
<taxonomicName id="4C0CA588024D281808B7F9A7FEECEEE9" authorityName="Guerrero" authorityYear="2021" baseAuthorityName="Santos-Sousa" baseAuthorityYear="1902" box="[182,284,1663,1689]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="species" species="heyeri">
|
||
<emphasis id="B9780219024D281808B7F9A7FF37EEE8" box="[182,199,1663,1688]" italics="true" pageId="6" pageNumber="434">F</emphasis>
|
||
.
|
||
<emphasis id="B9780219024D281808D7F9A7FEECEEE9" box="[214,284,1663,1689]" italics="true" pageId="6" pageNumber="434">heyeri</emphasis>
|
||
</taxonomicName>
|
||
the scape is relatively long (SI = 81–83) surpassing the vertex by a very short distance, just
|
||
<quantity id="4CF473EE024D28180D30F9A7FA68EEE9" box="[1329,1432,1663,1689]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="434" unit="mm" value="0.04">0.04 mm</quantity>
|
||
, while in
|
||
<taxonomicName id="4C0CA588024D281808FBF97BFE64EECC" baseAuthorityName="Forel" baseAuthorityYear="1912" box="[250,404,1699,1725]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="subSpecies" species="heyeri" subSpecies="risii">
|
||
<emphasis id="B9780219024D281808FBF97BFEFBEECC" box="[250,267,1699,1724]" italics="true" pageId="6" pageNumber="434">F</emphasis>
|
||
.
|
||
<emphasis id="B9780219024D28180918F97BFE64EECC" box="[281,404,1699,1725]" italics="true" pageId="6" pageNumber="434">heyeri risii</emphasis>
|
||
</taxonomicName>
|
||
the apex of the scape is well below the cephalic vertex (SI = 67). The coloration is variable among the specimens studied, from dull yellow in
|
||
<taxonomicName id="4C0CA588024D28180AC5F91FFCD7EE91" authorityName="Guerrero" authorityYear="2021" baseAuthorityName="Santos-Sousa" baseAuthorityYear="1902" box="[708,807,1735,1761]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="species" species="heyeri">
|
||
<emphasis id="B9780219024D28180AC5F91FFD25EE90" box="[708,725,1735,1760]" italics="true" pageId="6" pageNumber="434">F</emphasis>
|
||
.
|
||
<emphasis id="B9780219024D28180AE0F91FFCD7EE91" box="[737,807,1735,1761]" italics="true" pageId="6" pageNumber="434">heyeri</emphasis>
|
||
</taxonomicName>
|
||
to light brown in
|
||
<taxonomicName id="4C0CA588024D28180BEBF91FFB71EE90" baseAuthorityName="Forel" baseAuthorityYear="1912" box="[1002,1153,1735,1761]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="subSpecies" species="heyeri" subSpecies="risii">
|
||
<emphasis id="B9780219024D28180BEBF91FFC0BEE90" box="[1002,1019,1735,1760]" italics="true" pageId="6" pageNumber="434">F</emphasis>
|
||
.
|
||
<emphasis id="B9780219024D28180C09F91FFB71EE90" box="[1032,1153,1735,1761]" italics="true" pageId="6" pageNumber="434">heyeri risii</emphasis>
|
||
</taxonomicName>
|
||
. Although the
|
||
<typeStatus id="54B760A9024D28180D22F91FFAA3EE91" box="[1315,1363,1735,1761]" pageId="6" pageNumber="434">type</typeStatus>
|
||
specimens show differences, the specimens recently collected in
|
||
<collectingCountry id="F31B9E9B024D28180B2BF933FC7CEF75" box="[810,908,1771,1797]" name="Uruguay" pageId="6" pageNumber="434">Uruguay</collectingCountry>
|
||
show a relative continuity in the morphological and metric traits between the
|
||
<typeStatus id="54B760A9024D281809E3F8D7FDEDEF59" box="[482,541,1807,1833]" pageId="6" pageNumber="434">types</typeStatus>
|
||
of
|
||
<taxonomicName id="4C0CA588024D28180A43F8D7FD54EF59" authorityName="Guerrero" authorityYear="2021" baseAuthorityName="Santos-Sousa" baseAuthorityYear="1902" box="[578,676,1807,1833]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="species" species="heyeri">
|
||
<emphasis id="B9780219024D28180A43F8D7FD54EF59" box="[578,676,1807,1833]" italics="true" pageId="6" pageNumber="434">F. heyeri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0CA588024D28180ADBF8D7FC9FEF58" baseAuthorityName="Forel" baseAuthorityYear="1912" box="[730,879,1807,1833]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="subSpecies" species="heyeri" subSpecies="risii">
|
||
<emphasis id="B9780219024D28180ADBF8D7FC9FEF58" box="[730,879,1807,1833]" italics="true" pageId="6" pageNumber="434">F. heyeri risii</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1337C28E024D28180B7FF8D7FC39EF59" box="[894,969,1807,1833]" captionStart="FIGURE 11" captionStartId="8.[151,250,928,952]" captionTargetBox="[192,1403,193,889]" captionTargetId="figure-16@8.[169,1418,181,904]" captionTargetPageId="8" captionText="FIGURE 11. Bivariate plot relating the head length (HL) to both the head width (HW, left) and the scape length (SL, right) of Forelius heyeri. Small circles: three workers from a series recently collected in Uruguay. Medium circle: second syntype of F. heyeri. Large circle: F. heyeri lectotype. Square: F. heyeri risii lectotype. Small triangle: worker from Paraguay identified as F. pusillus on AntWeb (CASENT0173739). Large triangle: syntype of F. pusillus." figureDoi="http://doi.org/10.5281/zenodo.4478201" httpUri="https://zenodo.org/record/4478201/files/figure.png" pageId="6" pageNumber="434">Fig 11</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB3DE0B024D281908C6F8EBFCE4E915" blockId="6.[151,1437,151,2014]" lastBlockId="7.[151,1437,151,357]" lastPageId="7" lastPageNumber="435" pageId="6" pageNumber="434">
|
||
<bibRefCitation id="EF9DA3FA024D281808C6F8EBFE9DEF3D" author="Cuezzo, F." box="[199,365,1843,1869]" pageId="6" pageNumber="434" pagination="197 - 277" refId="ref7217" refString="Cuezzo, F. (2000) Revision del genero Forelius (Hymenoptera: Formicidae: Dolichoderinae). Sociobiology, 35, 197 - 277." type="journal article" year="2000">Cuezzo (2000)</bibRefCitation>
|
||
raised
|
||
<taxonomicName id="4C0CA588024D281809BFF8EBFD6BEF3D" authority="Santschi" authorityName="Santschi. Status" authorityYear="1922" box="[446,667,1843,1869]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="species" species="pusillus">
|
||
<emphasis id="B9780219024D281809BFF8EBFE3FEF3C" box="[446,463,1843,1868]" italics="true" pageId="6" pageNumber="434">F</emphasis>
|
||
.
|
||
<emphasis id="B9780219024D281809DFF8EBFDC5EF3D" box="[478,565,1843,1869]" italics="true" pageId="6" pageNumber="434">pusillus</emphasis>
|
||
Santschi
|
||
</taxonomicName>
|
||
to the species level, offering a diagnosis and measurements that differentiated this
|
||
<taxonomicName id="4C0CA588024D28180944F88FFE53EF01" authorityName=": Shattuck" authorityYear="1992" box="[325,419,1879,1905]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9780219024D28180944F88FFE53EF01" box="[325,419,1879,1905]" italics="true" pageId="6" pageNumber="434">Forelius</emphasis>
|
||
</taxonomicName>
|
||
species from all other species in South America. The taxonomic delimitation of
|
||
<taxonomicName id="4C0CA588024D28180D27F88FFA6CEF01" authorityName="Santschi. Status" authorityYear="1922" box="[1318,1436,1879,1905]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="species" species="pusillus">
|
||
<emphasis id="B9780219024D28180D27F88FFAC7EF00" box="[1318,1335,1879,1904]" italics="true" pageId="6" pageNumber="434">F</emphasis>
|
||
.
|
||
<emphasis id="B9780219024D28180D44F88FFA6CEF01" box="[1349,1436,1879,1905]" italics="true" pageId="6" pageNumber="434">pusillus</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EF9DA3FA024D281808B8F8A3FEAEEFE5" author="Cuezzo, F." box="[185,350,1915,1941]" pageId="6" pageNumber="434" pagination="197 - 277" refId="ref7217" refString="Cuezzo, F. (2000) Revision del genero Forelius (Hymenoptera: Formicidae: Dolichoderinae). Sociobiology, 35, 197 - 277." type="journal article" year="2000">Cuezzo (2000)</bibRefCitation>
|
||
together with the measurements recorded here from a
|
||
<typeStatus id="54B760A9024D28180BB8F8A3FC19EFE5" box="[953,1001,1915,1941]" pageId="6" pageNumber="434">type</typeStatus>
|
||
specimen of
|
||
<taxonomicName id="4C0CA588024D28180C7DF8A3FB02EFE5" authorityName="Santschi. Status" authorityYear="1922" box="[1148,1266,1915,1941]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="species" species="pusillus">
|
||
<emphasis id="B9780219024D28180C7DF8A3FB7DEFE4" box="[1148,1165,1915,1940]" italics="true" pageId="6" pageNumber="434">F</emphasis>
|
||
.
|
||
<emphasis id="B9780219024D28180C9AF8A3FB02EFE5" box="[1179,1266,1915,1941]" italics="true" pageId="6" pageNumber="434">pusillus</emphasis>
|
||
</taxonomicName>
|
||
and a non-type worker from
|
||
<collectingCountry id="F31B9E9B024D28180929F847FE60EFC9" box="[296,400,1951,1977]" name="Paraguay" pageId="6" pageNumber="434">Paraguay</collectingCountry>
|
||
(measurements generated from images with codes CASENT0911547 and CASENT0173739, respectively) were compared with the diagnosis and description provided here for
|
||
<taxonomicName id="4C0CA588024D28180C3BF81BFB50EFAD" authorityName="Guerrero" authorityYear="2021" baseAuthorityName="Santos-Sousa" baseAuthorityYear="1902" box="[1082,1184,1987,2013]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="species" species="heyeri">
|
||
<emphasis id="B9780219024D28180C3BF81BFBBBEFAC" box="[1082,1099,1987,2012]" italics="true" pageId="6" pageNumber="434">F</emphasis>
|
||
.
|
||
<emphasis id="B9780219024D28180C5BF81BFB50EFAD" box="[1114,1184,1987,2013]" italics="true" pageId="6" pageNumber="434">heyeri</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C0CA588024D28180CAEF81BFA9CEFAD" authorityName="Santschi. Status" authorityYear="1922" box="[1199,1388,1987,2013]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="434" phylum="Arthropoda" rank="species" species="pusillus">
|
||
<emphasis id="B9780219024D28180CAEF81BFA9CEFAD" box="[1199,1388,1987,2013]" italics="true" pageId="6" pageNumber="434">Forelius pusillus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0CA588024C28190896FF4FFF0DE8C1" authorityName="Guerrero" authorityYear="2021" baseAuthorityName="Santos-Sousa" baseAuthorityYear="1902" box="[151,253,151,177]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="435" phylum="Arthropoda" rank="species" species="heyeri">
|
||
<emphasis id="B9780219024C28190896FF4FFF58E8C0" box="[151,168,151,176]" italics="true" pageId="7" pageNumber="435">F</emphasis>
|
||
.
|
||
<emphasis id="B9780219024C281908B6FF4FFF0DE8C1" box="[183,253,151,177]" italics="true" pageId="7" pageNumber="435">heyeri</emphasis>
|
||
</taxonomicName>
|
||
fully agree in their morphology as follows: the workers of
|
||
<taxonomicName id="4C0CA588024C28190B9BFF4FFBE0E8C1" authorityName="Santschi. Status" authorityYear="1922" box="[922,1040,151,177]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="435" phylum="Arthropoda" rank="species" species="pusillus">
|
||
<emphasis id="B9780219024C28190B9BFF4FFC5BE8C0" box="[922,939,151,176]" italics="true" pageId="7" pageNumber="435">F</emphasis>
|
||
.
|
||
<emphasis id="B9780219024C28190BB8FF4FFBE0E8C1" box="[953,1040,151,177]" italics="true" pageId="7" pageNumber="435">pusillus</emphasis>
|
||
</taxonomicName>
|
||
also have two thick, erect pronotal setae, as well as a mesosomal profile similar to that found in
|
||
<taxonomicName id="4C0CA588024C28190B58FF63FC30E8A5" authorityName="Guerrero" authorityYear="2021" baseAuthorityName="Santos-Sousa" baseAuthorityYear="1902" box="[857,960,187,213]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="435" phylum="Arthropoda" rank="species" species="heyeri">
|
||
<emphasis id="B9780219024C28190B58FF63FC9AE8A4" box="[857,874,187,212]" italics="true" pageId="7" pageNumber="435">F</emphasis>
|
||
.
|
||
<emphasis id="B9780219024C28190B7BFF63FC30E8A5" box="[890,960,187,213]" italics="true" pageId="7" pageNumber="435">heyeri</emphasis>
|
||
</taxonomicName>
|
||
; the measurements are relatively variable throughout the distribution of these two putative taxa, with a similar morphometric space between those named
|
||
<taxonomicName id="4C0CA588024C28190D85FF07FEE0E96C" baseAuthorityName="Forel" baseAuthorityYear="1912" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="435" phylum="Arthropoda" rank="subSpecies" species="heyeri" subSpecies="risii">
|
||
<emphasis id="B9780219024C28190D85FF07FA65E888" box="[1412,1429,223,248]" italics="true" pageId="7" pageNumber="435">F</emphasis>
|
||
.
|
||
<emphasis id="B9780219024C28190896FEDBFEE0E96C" box="[151,272,259,285]" italics="true" pageId="7" pageNumber="435">heyeri risii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0CA588024C28190947FEDBFE4BE96D" authorityName="Santschi. Status" authorityYear="1922" box="[326,443,259,285]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="435" phylum="Arthropoda" rank="species" species="pusillus">
|
||
<emphasis id="B9780219024C28190947FEDBFEA7E96C" box="[326,343,259,284]" italics="true" pageId="7" pageNumber="435">F</emphasis>
|
||
.
|
||
<emphasis id="B9780219024C28190965FEDBFE4BE96D" box="[356,443,259,285]" italics="true" pageId="7" pageNumber="435">pusillus</emphasis>
|
||
</taxonomicName>
|
||
, but the nest-mate workers collected in
|
||
<collectingCountry id="F31B9E9B024C28190B77FEDBFC28E96D" box="[886,984,259,285]" name="Uruguay" pageId="7" pageNumber="435">Uruguay</collectingCountry>
|
||
agree as much with those of
|
||
<taxonomicName id="4C0CA588024C28190D1BFEDBFA8FE96D" authorityName="Guerrero" authorityYear="2021" baseAuthorityName="Santos-Sousa" baseAuthorityYear="1902" box="[1306,1407,259,285]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="435" phylum="Arthropoda" rank="species" species="heyeri">
|
||
<emphasis id="B9780219024C28190D1BFEDBFADBE96C" box="[1306,1323,259,284]" italics="true" pageId="7" pageNumber="435">F</emphasis>
|
||
.
|
||
<emphasis id="B9780219024C28190D38FEDBFA8FE96D" box="[1337,1407,259,285]" italics="true" pageId="7" pageNumber="435">heyeri</emphasis>
|
||
</taxonomicName>
|
||
as those of
|
||
<taxonomicName id="4C0CA588024C281908F9FEFFFE9EE931" authorityName="Santschi. Status" authorityYear="1922" box="[248,366,295,321]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="435" phylum="Arthropoda" rank="species" species="pusillus">
|
||
<emphasis id="B9780219024C281908F9FEFFFEF9E930" box="[248,265,295,320]" italics="true" pageId="7" pageNumber="435">F</emphasis>
|
||
.
|
||
<emphasis id="B9780219024C28190916FEFFFE9EE931" box="[279,366,295,321]" italics="true" pageId="7" pageNumber="435">pusillus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1337C28E024C2819097CFEFFFE38E931" box="[381,456,295,321]" captionStart="FIGURE 11" captionStartId="8.[151,250,928,952]" captionTargetBox="[192,1403,193,889]" captionTargetId="figure-16@8.[169,1418,181,904]" captionTargetPageId="8" captionText="FIGURE 11. Bivariate plot relating the head length (HL) to both the head width (HW, left) and the scape length (SL, right) of Forelius heyeri. Small circles: three workers from a series recently collected in Uruguay. Medium circle: second syntype of F. heyeri. Large circle: F. heyeri lectotype. Square: F. heyeri risii lectotype. Small triangle: worker from Paraguay identified as F. pusillus on AntWeb (CASENT0173739). Large triangle: syntype of F. pusillus." figureDoi="http://doi.org/10.5281/zenodo.4478201" httpUri="https://zenodo.org/record/4478201/files/figure.png" pageId="7" pageNumber="435">Fig 11</figureCitation>
|
||
). In this sense, I do not find morphological evidence to differentiate between these taxa and I propose
|
||
<taxonomicName id="4C0CA588024C28190936FE93FE5DE915" authorityName="Santschi. Status" authorityYear="1922" box="[311,429,331,357]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="435" phylum="Arthropoda" rank="species" species="pusillus">
|
||
<emphasis id="B9780219024C28190936FE93FEB8E914" box="[311,328,331,356]" italics="true" pageId="7" pageNumber="435">F</emphasis>
|
||
.
|
||
<emphasis id="B9780219024C28190957FE93FE5DE915" box="[342,429,331,357]" italics="true" pageId="7" pageNumber="435">pusillus</emphasis>
|
||
</taxonomicName>
|
||
as junior synonym of
|
||
<taxonomicName id="4C0CA588024C28190AA9FE93FCFDE915" authorityName="Guerrero" authorityYear="2021" baseAuthorityName="Santos-Sousa" baseAuthorityYear="1902" box="[680,781,331,357]" class="Insecta" family="Formicidae" genus="Forelius" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="435" phylum="Arthropoda" rank="species" species="heyeri">
|
||
<emphasis id="B9780219024C28190AA9FE93FD49E914" box="[680,697,331,356]" italics="true" pageId="7" pageNumber="435">F</emphasis>
|
||
.
|
||
<emphasis id="B9780219024C28190AC6FE93FCFDE915" box="[711,781,331,357]" italics="true" pageId="7" pageNumber="435">heyeri</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |