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<document id="237BFEE5BE202F2E312975716936F78B" ID-DOI="10.11646/zootaxa.3923.1.1" ID-GBIF-Dataset="2822ae22-53bb-45c1-b3a3-49f7b6c0fb53" ID-ISSN="1175-5326" ID-Zenodo-Dep="241936" ID-ZooBank="2305A96C-0A03-4524-93AA-90359893A4DD" IM.bibliography_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460998774137" checkinUser="plazi" docAuthor="Kaczmarek, Łukasz, Michalczyk, Łukasz &amp; Mcinnes, Sandra J." docDate="2015" docId="038FA02EFFDE7A7E4A85BC1E506DF440" docLanguage="en" docName="zt03923p107.pdf" docOrigin="Zootaxa 3923 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Macrobiotus hibiscus" docType="treatment" docVersion="13" lastPageNumber="67" masterDocId="FFB6D856FF9C7A3C4A12B8175470F34E" masterDocTitle="Annotated zoogeography of non-marine Tardigrada. Part II: South America" masterLastPageNumber="107" masterPageNumber="1" pageNumber="67" updateTime="1698564022090" updateUser="plazi">
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<mods:title id="A9981E31E0CC106ACDC27885DF978CC5">Annotated zoogeography of non-marine Tardigrada. Part II: South America</mods:title>
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<heading id="D0D1A654FFDE7A7E4A85BC1E56B5F76C" bold="true" box="[151,709,1033,1058]" fontSize="10" level="2" pageId="66" pageNumber="67" reason="2">
<emphasis id="B952CD2AFFDE7A7E4A85BC1E56B5F76C" bold="true" box="[151,709,1033,1058]" pageId="66" pageNumber="67">
164.
<taxonomicName id="4C266ABBFFDE7A7E4ADDBC1E55B7F76C" box="[207,455,1033,1058]" class="Heterotardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="66" pageNumber="67" phylum="Tardigrada" rank="species" species="hibiscus">
<emphasis id="B952CD2AFFDE7A7E4ADDBC1E55B7F76C" bold="true" box="[207,455,1033,1058]" italics="true" pageId="66" pageNumber="67">Macrobiotus hibiscus</emphasis>
</taxonomicName>
de
<bibRefCitation id="EFB76CC9FFDE7A7E4BE3BC1E56E9F76C" author="Barros" box="[497,665,1033,1058]" pageId="66" pageNumber="67" refString="De Barros, R. (1942 b) Tardigrados de Estado de Sao Paulo, Brasil. II. Genero Macrobiotus. Revista Brasileira de Biologia, 2, 373 - 386." type="journal article" year="1942" yearSuffix="b">Barros, 1942b</bibRefCitation>
[T]
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<taxonomicName id="4C266ABBFFDE7A7E4AD5BC3955B5F70D" authority="Barros, 1942" authorityName="Barros" authorityYear="1942" box="[199,453,1069,1091]" class="Heterotardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="66" pageNumber="67" phylum="Tardigrada" rank="species" species="hibiscus">
<emphasis id="B952CD2AFFDE7A7E4AD5BC39554CF70D" box="[199,316,1069,1091]" italics="true" pageId="66" pageNumber="67">M. hibiscus</emphasis>
Barros, 1942
</taxonomicName>
(
<bibRefCitation id="EFB76CC9FFDE7A7E4BC0BC3A56D1F70D" author="Claps" box="[466,673,1069,1091]" pageId="66" pageNumber="67" refString="Claps, M. C. &amp; Rossi, G. C. (1981) Contribucion al conocimiento de los tardigrados de Argentina. II. Revista de la Sociedad Entomologica Argentina, 40 (1 - 4), 107 - 114." type="journal article" year="1981">Claps &amp; Rossi 1981</bibRefCitation>
, 1984)
</paragraph>
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<taxonomicName id="4C266ABBFFDE7A7E4AD5BC5B55D0F72C" box="[199,416,1100,1122]" class="Heterotardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="66" pageNumber="67" phylum="Tardigrada" rank="species" species="hibiscus" status="sp. nov.">
<emphasis id="B952CD2AFFDE7A7E4AD5BC5B55D0F72C" box="[199,416,1100,1122]" italics="true" pageId="66" pageNumber="67">Macrobiotus hibiscus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2617051FFDE7A7E4BB5BC5A55AEF72C" box="[423,478,1101,1122]" pageId="66" pageNumber="67" rank="species">n. sp.</taxonomicNameLabel>
(de
<bibRefCitation id="EFB76CC9FFDE7A7E4818BC5A56E4F72C" author="Barros" box="[522,660,1100,1122]" pageId="66" pageNumber="67" refString="De Barros, R. (1942 b) Tardigrados de Estado de Sao Paulo, Brasil. II. Genero Macrobiotus. Revista Brasileira de Biologia, 2, 373 - 386." type="journal article" year="1942" yearSuffix="b">Barros 1942b</bibRefCitation>
)
</paragraph>
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<subSubSection id="C33C42B3FFDE7A7E4AD5BC7D572DF55C" pageId="66" pageNumber="67" type="reference_group">
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<taxonomicName id="4C266ABBFFDE7A7E4AD5BC7D577CF7CE" authority="Barros, 1942 (Rossi &amp; Claps 1980)" authorityName="Barros, 1942 (Rossi &amp; Claps" authorityYear="1980" box="[199,780,1130,1152]" class="Heterotardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="66" pageNumber="67" phylum="Tardigrada" rank="species" species="hibiscus">
<emphasis id="B952CD2AFFDE7A7E4AD5BC7D55D0F7CE" box="[199,416,1130,1152]" italics="true" pageId="66" pageNumber="67">Macrobiotus hibiscus</emphasis>
Barros, 1942 (
<bibRefCitation id="EFB76CC9FFDE7A7E4825BC7D5775F7CE" author="Rossi" box="[567,773,1130,1152]" pageId="66" pageNumber="67" refString="Rossi, G. C. and Claps, M. C. (1980) Contribucion al conocimiento de los tardigrados de Argentina. I. Revista de la Sociedad Entomologica Argentina, 39, 243 - 250" type="journal article" year="1980">Rossi &amp; Claps 1980</bibRefCitation>
)
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<emphasis id="B952CD2AFFDE7A7E4AD5BC9D5534F7D1" bold="true" box="[199,324,1161,1183]" italics="true" pageId="66" pageNumber="67">Terra typica</emphasis>
:
<collectingCountry id="F33151A8FFDE7A7E4B40BC9E55E4F7D1" box="[338,404,1161,1183]" name="Brazil" pageId="66" pageNumber="67">Brazil</collectingCountry>
(South
<collectingCountry id="F33151A8FFDE7A7E4BFBBC9E5637F7D1" box="[489,583,1161,1183]" name="United States of America" pageId="66" pageNumber="67">America</collectingCountry>
)
</emphasis>
</paragraph>
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<collectingCountry id="F33151A8FFDE7A7E4A85BCBF5579F7F0" box="[151,265,1192,1214]" name="Argentina" pageId="66" pageNumber="67">Argentina</collectingCountry>
:
</emphasis>
</paragraph>
<paragraph id="8B991138FFDE7A7E4A85BCDF55DCF7B5" blockId="66.[151,1437,1033,1806]" pageId="66" pageNumber="67">
<emphasis id="B952CD2AFFDE7A7E4A85BCDF5582F792" box="[151,498,1222,1245]" italics="true" pageId="66" pageNumber="67">
•
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,
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;
<quantity id="4CDEBCDDFFDE7A7E4B9EBCD155BDF793" box="[396,461,1222,1245]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="66" pageNumber="67" unit="m" value="400.0">400 m</quantity>
asl
</emphasis>
: Salta Province, road from Las Lajitas to J.V. González, lichens on tree in the mountains.
<bibRefCitation id="EFB76CC9FFDE7A7E4ADBBCF255DCF7B5" author="Claps" box="[201,428,1253,1275]" pageId="66" pageNumber="67" refString="Claps, M. C. &amp; Rossi, G. C. (1984) Contribucion al conocimiento de los tardigrados de Argentina. IV. Acta Zoologica Lilloana, 38, 45 - 50." type="journal article" year="1984">
<emphasis id="B952CD2AFFDE7A7E4ADBBCF255DCF7B5" bold="true" box="[201,428,1253,1275]" pageId="66" pageNumber="67">Claps &amp; Rossi (1984)</emphasis>
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<emphasis id="B952CD2AFFDE7A7E4A85BD1255BFF654" box="[151,463,1284,1306]" italics="true" pageId="66" pageNumber="67">
•
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,
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; 0 m asl
</emphasis>
: Buenos Aires Province, San Clemente del Tuy, lichens on tree (
<taxonomicName id="4C266ABBFFDE7A7E4E4EBD1350BDF654" box="[1116,1229,1284,1306]" class="Heterotardigrada" family="Myrtaceae" genus="Eucalyptus" kingdom="Animalia" order="Myrtales" pageId="66" pageNumber="67" phylum="Tardigrada" rank="genus">
<emphasis id="B952CD2AFFDE7A7E4E4EBD1350BDF654" box="[1116,1229,1284,1306]" italics="true" pageId="66" pageNumber="67">Eucalyptus</emphasis>
</taxonomicName>
coppice) in full sun.
<bibRefCitation id="EFB76CC9FFDE7A7E4ADBBD3555DCF676" author="Rossi" box="[201,428,1314,1336]" pageId="66" pageNumber="67" refString="Rossi, G. C. and Claps, M. C. (1980) Contribucion al conocimiento de los tardigrados de Argentina. I. Revista de la Sociedad Entomologica Argentina, 39, 243 - 250" type="journal article" year="1980">
<emphasis id="B952CD2AFFDE7A7E4ADBBD3555DCF676" bold="true" box="[201,428,1314,1336]" pageId="66" pageNumber="67">Rossi &amp; Claps (1980)</emphasis>
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</paragraph>
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<emphasis id="B952CD2AFFDE7A7E4A85BD555581F619" box="[151,497,1345,1367]" italics="true" pageId="66" pageNumber="67">
•
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,
<geoCoordinate id="EE1277FFFFDE7A7E4B37BD56550DF619" box="[293,381,1345,1367]" direction="west" orientation="longitude" pageId="66" pageNumber="67" precision="925" value="-71.63333">71°38′W</geoCoordinate>
;
<quantity id="4CDEBCDDFFDE7A7E4B99BD5655BBF619" box="[395,459,1345,1367]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="66" pageNumber="67" unit="m" value="900.0">900 m</quantity>
asl
</emphasis>
: Neuquén Province, Peninsula de Quetrihue, semi-wet and dry, mosses on tree in coppice, mosses on rock and soil in the forest and lichens on trees (4 samples).
<bibRefCitation id="EFB76CC9FFDE7A7E4983BD775004F638" author="Claps" box="[913,1140,1376,1398]" pageId="66" pageNumber="67" refString="Claps, M. C. &amp; Rossi, G. C. (1981) Contribucion al conocimiento de los tardigrados de Argentina. II. Revista de la Sociedad Entomologica Argentina, 40 (1 - 4), 107 - 114." type="journal article" year="1981">
<emphasis id="B952CD2AFFDE7A7E4983BD775004F638" bold="true" box="[913,1140,1376,1398]" pageId="66" pageNumber="67">Claps &amp; Rossi (1981)</emphasis>
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</paragraph>
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<collectingCountry id="F33151A8FFDE7A7E4A85BD6954ADF6DA" box="[151,221,1406,1428]" name="Brazil" pageId="66" pageNumber="67">Brazil</collectingCountry>
:
</emphasis>
</paragraph>
<paragraph id="8B991138FFDE7A7E4A85BD8950E0F6FD" blockId="66.[151,1437,1033,1806]" box="[151,1168,1437,1459]" pageId="66" pageNumber="67">
<emphasis id="B952CD2AFFDE7A7E4A85BD895599F6FD" box="[151,489,1437,1459]" italics="true" pageId="66" pageNumber="67">
•
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,
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;
<quantity id="4CDEBCDDFFDE7A7E4B95BD8A55B5F6FD" box="[391,453,1437,1459]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="66" pageNumber="67" unit="m" value="700.0">700 m</quantity>
asl
</emphasis>
:
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<typeStatus id="549DAF9AFFDE7A7E4BE5BD8A565CF6FD" box="[503,556,1437,1459]" pageId="66" pageNumber="67">Type</typeStatus>
Locality
</emphasis>
: São Paulo State, Casa Branca.
<emphasis id="B952CD2AFFDE7A7E49DDBD8A50E0F6FD" bold="true" box="[975,1168,1437,1459]" pageId="66" pageNumber="67">
de
<bibRefCitation id="EFB76CC9FFDE7A7E49FCBD8A50E0F6FD" author="Barros" box="[1006,1168,1437,1459]" pageId="66" pageNumber="67" refString="De Barros, R. (1942 b) Tardigrados de Estado de Sao Paulo, Brasil. II. Genero Macrobiotus. Revista Brasileira de Biologia, 2, 373 - 386." type="journal article" year="1942" yearSuffix="b">Barros (1942b)</bibRefCitation>
</emphasis>
</paragraph>
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<collectingCountry id="F33151A8FFDE7A7E4A85BDAB5488F69C" box="[151,248,1468,1490]" name="Ecuador" pageId="66" pageNumber="67">Ecuador</collectingCountry>
:
</emphasis>
</paragraph>
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•
<geoCoordinate id="EE1277FFFFDE7A7E4AD5BDCD5566F6BE" box="[199,278,1498,1520]" direction="south" orientation="latitude" pageId="66" pageNumber="67" precision="925" value="-2.1833334">02°11′S</geoCoordinate>
,
<geoCoordinate id="EE1277FFFFDE7A7E4B33BDCD5509F6BE" box="[289,377,1498,1520]" direction="west" orientation="longitude" pageId="66" pageNumber="67" precision="925" value="-79.88333">79°53′W</geoCoordinate>
; 0 m asl:
</emphasis>
Guayaquil, mixture of moss and lichen (
<taxonomicName id="4C266ABBFFDE7A7E4967BDCD57B5F6BE" box="[885,965,1498,1520]" class="Heterotardigrada" family="Physciaceae" genus="Physcia" kingdom="Animalia" order="Teloschistales" pageId="66" pageNumber="67" phylum="Tardigrada" rank="genus">
<emphasis id="B952CD2AFFDE7A7E4967BDCD57B5F6BE" box="[885,965,1498,1520]" italics="true" pageId="66" pageNumber="67">Physcia</emphasis>
</taxonomicName>
sp.) on trees.
<bibRefCitation id="EFB76CC9FFDE7A7E4E46BDCD51F8F6BE" author="Schuster" box="[1108,1416,1498,1520]" pageId="66" pageNumber="67" refString="Schuster, R. O. &amp; Grigarick, A. A. (1966) Tardigrada from the Galapagos and Cocos Islands. Proceedings of the California Academy of Sciences, 34, 315 - 328." type="journal article" year="1966">
<emphasis id="B952CD2AFFDE7A7E4E46BDCD51F8F6BE" bold="true" box="[1108,1416,1498,1520]" pageId="66" pageNumber="67">Schuster &amp; Grigarick (1966)</emphasis>
</bibRefCitation>
</paragraph>
<paragraph id="8B991138FFDE7A7E4A85BDEE572DF55C" blockId="66.[151,1437,1033,1806]" box="[151,861,1529,1554]" pageId="66" pageNumber="67">
<emphasis id="B952CD2AFFDE7A7E4A85BDEE5518F55C" bold="true" box="[151,360,1529,1554]" pageId="66" pageNumber="67">Record numbers:</emphasis>
<collectingCountry id="F33151A8FFDE7A7E4B7DBDEE5593F55C" box="[367,483,1529,1554]" name="Argentina" pageId="66" pageNumber="67">Argentina</collectingCountry>
: 3,
<collectingCountry id="F33151A8FFDE7A7E481BBDEE5622F55C" box="[521,594,1529,1554]" name="Brazil" pageId="66" pageNumber="67">Brazil</collectingCountry>
: 1,
<collectingCountry id="F33151A8FFDE7A7E486ABDEE56AAF55C" box="[632,730,1529,1554]" name="Ecuador" pageId="66" pageNumber="67">Ecuador</collectingCountry>
: 1;
<emphasis id="B952CD2AFFDE7A7E4913BDEE572DF55C" bold="true" box="[769,861,1529,1554]" pageId="66" pageNumber="67">total: 5.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33C42B3FFDE7A7E4A85BE0A506DF440" pageId="66" pageNumber="67" type="discussion">
<paragraph id="8B991138FFDE7A7E4A85BE0A506DF440" blockId="66.[151,1437,1033,1806]" pageId="66" pageNumber="67">
<emphasis id="B952CD2AFFDE7A7E4A85BE0A557DF578" bold="true" box="[151,269,1565,1590]" pageId="66" pageNumber="67">Remarks:</emphasis>
There has been some discussion over whether
<taxonomicName id="4C266ABBFFDE7A7E4905BE0A5077F578" box="[791,1031,1565,1590]" class="Heterotardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="66" pageNumber="67" phylum="Tardigrada" rank="species" species="hibiscus">
<emphasis id="B952CD2AFFDE7A7E4905BE0A5077F578" box="[791,1031,1565,1590]" italics="true" pageId="66" pageNumber="67">Macrobiotus hibiscus</emphasis>
</taxonomicName>
is a junior synonym of
<taxonomicName id="4C266ABBFFDE7A7E4F1CBE0A5541F514" class="Heterotardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="66" pageNumber="67" phylum="Tardigrada" rank="species" species="hufelandi">
<emphasis id="B952CD2AFFDE7A7E4F1CBE0A5541F514" italics="true" pageId="66" pageNumber="67">Macrobiotus hufelandi</emphasis>
</taxonomicName>
as the morphology of this species falls within the
<taxonomicName id="4C266ABBFFDE7A7E4985BE5650EBF514" box="[919,1179,1601,1626]" class="Heterotardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="66" pageNumber="67" phylum="Tardigrada" rank="species" species="hufelandi">
<emphasis id="B952CD2AFFDE7A7E4985BE5650EBF514" box="[919,1179,1601,1626]" italics="true" pageId="66" pageNumber="67">Macrobiotus hufelandi</emphasis>
</taxonomicName>
group diagnosis (
<emphasis id="B952CD2AFFDE7A7E4F66BE5551ECF517" box="[1396,1436,1602,1625]" italics="true" pageId="66" pageNumber="67">e.g.</emphasis>
<bibRefCitation id="EFB76CC9FFDE7A7E4AD5BE725506F530" author="Kathman" box="[199,374,1637,1662]" pageId="66" pageNumber="67" refString="Kathman, R. D. (1990) Eutardigrada from Vancouver Island, British Columbia, Canada, including a description of Platicrista cheleusis n. sp. Canadian Journal of Zoology, 68 (9), 1880 - 1895. http: // dx. doi. org / 10.1139 / z 90 - 268" type="journal article" year="1990">Kathman 1990</bibRefCitation>
). This is a group that requires careful taxonomic analysis, including egg morphology, for correct identification (see
<bibRefCitation id="EFB76CC9FFDE7A7E4BE1BE9E575DF5EC" author="Bertolani" box="[499,813,1673,1698]" pageId="66" pageNumber="67" refString="Bertolani, R. &amp; Rebecchi, L. (1993) A revision of the Macrobiotus hufelandi group (Tardigrada, Macrobiotidae), with some observations on the taxonomic characters of eutardigrades. Zoologica Scripta, 22, 127 - 152. http: // dx. doi. org / 10.1111 / j. 1463 - 6409.1993. tb 00347. x" type="journal article" year="1993">Bertolani &amp; Rebecchi 1993</bibRefCitation>
for a diagnostic key and other papers for more recent species descriptions:
<emphasis id="B952CD2AFFDE7A7E4BA4BEB955AEF58B" box="[438,478,1710,1733]" italics="true" pageId="66" pageNumber="67">e.g.</emphasis>
<bibRefCitation id="EFB76CC9FFDE7A7E4BF4BEBA56C9F588" author="Pilato" box="[486,697,1709,1734]" pageId="66" pageNumber="67" refString="Pilato, G., Kaczmarek, L., Michalczyk, L. &amp; Lisi, O. (2003 b) Macrobiotus polonicus, a new species of Tardigrada from Poland (Eutardigrada, Macrobiotidae, ' hufelandi group'). Zootaxa, 258, 1 - 8." type="journal article" year="2003" yearSuffix="b">
Pilato
<emphasis id="B952CD2AFFDE7A7E4822BEB95619F588" box="[560,617,1709,1734]" italics="true" pageId="66" pageNumber="67">et al.</emphasis>
2003b
</bibRefCitation>
, Kaczmarek &amp; Michalczyk 2004,
<bibRefCitation id="EFB76CC9FFDE7A7E4E2FBEB95155F588" author="Fontoura" box="[1085,1317,1709,1734]" pageId="66" pageNumber="67" refString="Fontoura, P., Pilato, G. &amp; Lisi, O. (2008) New records of eutardigrades (Tardigrada) from Faial and Pico Islands, the Azores, with the description of two new species. Zootaxa, 1778, 37 - 47." type="journal article" year="2008">
Fontoura
<emphasis id="B952CD2AFFDE7A7E4EBBBEB95092F588" box="[1193,1250,1709,1734]" italics="true" pageId="66" pageNumber="67">et al.</emphasis>
2008
</bibRefCitation>
,
<bibRefCitation id="EFB76CC9FFDE7A7E4F23BEBA5558F5A4" author="Bartels" pageId="66" pageNumber="67" refString="Bartels, P., Pilato, G., Lisi, O. &amp; Nelson, D. R. (2009) Macrobiotus (Eutardigrada, Macrobiotidae) from the Great Smoky Mountains National Park, Tennessee / North Carolina, USA (North America): two new species and six new records. Zootaxa, 2022, 45 - 57." type="journal article" year="2009">
Bartels
<emphasis id="B952CD2AFFDE7A7E4F9ABEB95494F5A4" italics="true" pageId="66" pageNumber="67">et al.</emphasis>
2009
</bibRefCitation>
,
<bibRefCitation id="EFB76CC9FFDE7A7E4B27BEC656C4F5A4" author="Kaczmarek" box="[309,692,1745,1770]" pageId="66" pageNumber="67" refString="Kaczmarek, L. &amp; Michalczyk, L. (2009 b) Two new species of Macrobiotidae, Macrobiotus szeptyckii (harmsworthi group) and Macrobiotus kazmierskii (hufelandi group) from Argentina. Acta Zoologica Cracoviensia, 52 B, 87 - 99. http: // dx. doi. org / 10.3409 / azc. 52 b _ 1 - 2.87 - 99" type="journal article" year="2009" yearSuffix="b">Kaczmarek &amp; Michalczyk 2009b</bibRefCitation>
,
<bibRefCitation id="EFB76CC9FFDE7A7E48D0BEC657D8F5A4" author="Guidetti" box="[706,936,1745,1770]" pageId="66" pageNumber="67" refString="Guidetti R., Peluffo, J. R., Rocha, A. M., Cesari, M. &amp; Moly de Peluffo, M. C. (2013) The morphological and molecular analyses of a new South American urban tardigrade offer new insights on the biological meaning of the Macrobiotus hufelandi group of species (Tardigrada: Macrobiotidae). Journal of Natural History, 47 (37 - 38), 2409 - 2426. http: // dx. doi. org / 10.1080 / 00222933.2013.800610" type="journal article" year="2013">
Guidetti
<emphasis id="B952CD2AFFDE7A7E4935BEC55712F5A4" box="[807,866,1745,1770]" italics="true" pageId="66" pageNumber="67">et al.</emphasis>
2013
</bibRefCitation>
). Further work will be required on the
<typeStatus id="549DAF9AFFDE7A7E4F7EBEC551ECF5A4" box="[1388,1436,1746,1770]" pageId="66" pageNumber="67">type</typeStatus>
material or
<typeStatus id="549DAF9AFFDE7A7E4B5ABEE15508F440" box="[328,376,1782,1806]" pageId="66" pageNumber="67">type</typeStatus>
locale to clarify the status and biogeography of this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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