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<document id="F90B0D4682D0CC91AD826BE8B3D715B4" ID-DOI="10.11646/zootaxa.3923.1.1" ID-GBIF-Dataset="2822ae22-53bb-45c1-b3a3-49f7b6c0fb53" ID-ISSN="1175-5326" ID-Zenodo-Dep="241936" ID-ZooBank="2305A96C-0A03-4524-93AA-90359893A4DD" IM.bibliography_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460998774137" checkinUser="plazi" docAuthor="Kaczmarek, Łukasz, Michalczyk, Łukasz &amp; Mcinnes, Sandra J." docDate="2015" docId="038FA02EFF887A284A85BB3556F8F653" docLanguage="en" docName="zt03923p107.pdf" docOrigin="Zootaxa 3923 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Echiniscus spiniger Richters 1904" docType="treatment" docVersion="13" lastPageNumber="21" masterDocId="FFB6D856FF9C7A3C4A12B8175470F34E" masterDocTitle="Annotated zoogeography of non-marine Tardigrada. Part II: South America" masterLastPageNumber="107" masterPageNumber="1" pageNumber="21" updateTime="1698564022090" updateUser="plazi">
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<mods:title id="97FB6AE36B509F4621AAA6103933B176">Annotated zoogeography of non-marine Tardigrada. Part II: South America</mods:title>
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<subSubSection id="C33C42B3FF887A284A85BB35577AF075" box="[151,778,802,827]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="8B991138FF887A284A85BB35577AF075" blockId="20.[151,1436,802,1309]" box="[151,778,802,827]" pageId="20" pageNumber="21">
<heading id="D0D1A654FF887A284A85BB35577AF075" bold="true" box="[151,778,802,827]" fontSize="10" level="2" pageId="20" pageNumber="21" reason="2">
<emphasis id="B952CD2AFF887A284A85BB35577AF075" bold="true" box="[151,778,802,827]" pageId="20" pageNumber="21">
45.
<taxonomicName id="4C266ABBFF887A284AD3BB355613F075" ID-CoL="38G3J" authority="Richters, 1904" authorityName="Richters" authorityYear="1904" box="[193,611,802,827]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="20" pageNumber="21" phylum="Tardigrada" rank="species" species="spiniger">
<emphasis id="B952CD2AFF887A284AD3BB3555D4F075" bold="true" box="[193,420,802,827]" italics="true" pageId="20" pageNumber="21">Echiniscus spiniger</emphasis>
<bibRefCitation id="EFB76CC9FF887A284BBEBB355613F075" author="Richters" box="[428,611,802,827]" pageId="20" pageNumber="21" refString="Richters, F. (1904 c) Beitrag zur Verbreitungen der Tardigraden im sudlichen Skandinavien und an der mecklenburgischen Kuste. Zoologischer Anzeiger, 28, 347 - 352." type="journal article" year="1904" yearSuffix="c">Richters, 1904c</bibRefCitation>
</taxonomicName>
<emphasis id="B952CD2AFF887A284878BB3456AFF075" bold="true" box="[618,735,802,827]" italics="true" pageId="20" pageNumber="21">sensu lato</emphasis>
[T]
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33C42B3FF887A284AD5BB5156A4F7FE" pageId="20" pageNumber="21" type="reference_group">
<paragraph id="8B991138FF887A284AD5BB5156F1F012" blockId="20.[151,1436,802,1309]" box="[199,641,838,861]" pageId="20" pageNumber="21">
<taxonomicName id="4C266ABBFF887A284AD5BB5156F1F012" ID-CoL="38G3J" authority="Richters. (Heinis 1914)" authorityName="Richters. (Heinis" authorityYear="1914" box="[199,641,838,861]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="20" pageNumber="21" phylum="Tardigrada" rank="species" species="spiniger">
<emphasis id="B952CD2AFF887A284AD5BB5155FEF013" box="[199,398,838,861]" italics="true" pageId="20" pageNumber="21">Echiniscus spiniger</emphasis>
Richters. (
<bibRefCitation id="EFB76CC9FF887A284BEEBB51560AF012" author="Heinis" box="[508,634,838,860]" pageId="20" pageNumber="21" refString="Heinis, F. (1914) Die Moosfauna Columbiens. Memoires de la Societe des Sciences Naturelles de Neuchatel, 5, 713 - 724." type="journal article" year="1914">Heinis 1914</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8B991138FF887A284AD5BB725722F035" blockId="20.[151,1436,802,1309]" box="[199,850,869,891]" pageId="20" pageNumber="21">
<taxonomicName id="4C266ABBFF887A284AD5BB7256E6F035" ID-CoL="38G3J" authority="Richt., 1904" authorityName="Richt." authorityYear="1904" box="[199,662,869,891]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="20" pageNumber="21" phylum="Tardigrada" rank="species" species="spiniger" subGenus="Echiniscus">
<emphasis id="B952CD2AFF887A284AD5BB725545F035" box="[199,309,869,891]" italics="true" pageId="20" pageNumber="21">Echiniscus</emphasis>
(
<emphasis id="B952CD2AFF887A284B51BB7255C1F035" box="[323,433,869,891]" italics="true" pageId="20" pageNumber="21">Echiniscus</emphasis>
)
<emphasis id="B952CD2AFF887A284BD2BB715663F035" box="[448,531,870,891]" italics="true" pageId="20" pageNumber="21">spiniger</emphasis>
Richt., 1904
</taxonomicName>
(de
<bibRefCitation id="EFB76CC9FF887A2848D0BB71573BF035" author="Barros" box="[706,843,869,891]" pageId="20" pageNumber="21" refString="De Barros, R. (1942 a) Tardigrados de Estado de Sao Paulo, Brasil. I. Introducao. Generos Echiniscus e Pseudechiniscus. Revista Brasileira de Biologia, 2, 257 - 269." type="journal article" year="1942" yearSuffix="a">Barros 1942a</bibRefCitation>
)
</paragraph>
<paragraph id="8B991138FF887A284AD5BB925721F0D4" blockId="20.[151,1436,802,1309]" box="[199,849,900,922]" pageId="20" pageNumber="21">
<taxonomicName id="4C266ABBFF887A284AD5BB925587F0D4" ID-CoL="38G3J" authority="Richters, 1904" authorityName="Richters" authorityYear="1904" box="[199,503,900,922]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="20" pageNumber="21" phylum="Tardigrada" rank="species" species="spiniger" subGenus="Echiniscus">
<emphasis id="B952CD2AFF887A284AD5BB9254ADF0D4" box="[199,221,901,922]" italics="true" pageId="20" pageNumber="21">E.</emphasis>
(
<emphasis id="B952CD2AFF887A284AF7BB92548BF0D4" box="[229,251,901,922]" italics="true" pageId="20" pageNumber="21">E.</emphasis>
)
<emphasis id="B952CD2AFF887A284B18BB92552DF0D4" box="[266,349,901,922]" italics="true" pageId="20" pageNumber="21">spiniger</emphasis>
Richters, 1904
</taxonomicName>
(du
<bibRefCitation id="EFB76CC9FF887A284837BB93573BF0D4" author="Bois-Reymond" box="[549,843,900,922]" pageId="20" pageNumber="21" refString="Bois-Reymond Marcus, E. du. (1944) Sobre tardigrados brasileiros. Comunicaciones Zoologicas del Museo de Historia Natural de Montevideo, 1 (13), 1 - 19." type="journal article" year="1944">Bois-Reymond Marcus 1944</bibRefCitation>
)
</paragraph>
<paragraph id="8B991138FF887A284AD5BBB4567CF0F6" blockId="20.[151,1436,802,1309]" box="[199,524,930,952]" pageId="20" pageNumber="21">
<emphasis id="B952CD2AFF887A284AD5BBB4567CF0F6" bold="true" box="[199,524,930,952]" pageId="20" pageNumber="21">
<emphasis id="B952CD2AFF887A284AD5BBB45534F0F6" bold="true" box="[199,324,930,952]" italics="true" pageId="20" pageNumber="21">Terra typica</emphasis>
:
<collectingCountry id="F33151A8FF887A284B40BBB555D5F0F6" box="[338,421,930,952]" name="Sweden" pageId="20" pageNumber="21">Sweden</collectingCountry>
(Europe)
</emphasis>
</paragraph>
<paragraph id="8B991138FF887A284A85BBD65492F099" blockId="20.[151,1436,802,1309]" box="[151,226,961,983]" pageId="20" pageNumber="21">
<emphasis id="B952CD2AFF887A284A85BBD65492F099" bold="true" box="[151,226,961,983]" pageId="20" pageNumber="21">
<collectingCountry id="F33151A8FF887A284A85BBD654ADF099" box="[151,221,961,983]" name="Brazil" pageId="20" pageNumber="21">Brazil</collectingCountry>
:
</emphasis>
</paragraph>
<paragraph id="8B991138FF887A284A85BBF65008F75A" blockId="20.[151,1436,802,1309]" pageId="20" pageNumber="21">
<emphasis id="B952CD2AFF887A284A85BBF6558FF0B8" box="[151,511,992,1014]" italics="true" pageId="20" pageNumber="21">
<geoCoordinate id="EE1277FFFF887A284ADABBF75569F0B8" box="[200,281,992,1014]" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="925" value="-22.733334">22°44S</geoCoordinate>
,
<geoCoordinate id="EE1277FFFF887A284B36BBF7550CF0B8" box="[292,380,992,1014]" direction="west" orientation="longitude" pageId="20" pageNumber="21" precision="925" value="-45.583332">45°35W</geoCoordinate>
;
<quantity id="4CDEBCDDFF887A284B9BBBF755ABF0B8" box="[393,475,992,1014]" metricMagnitude="0" metricUnit="m" metricValue="1.65" pageId="20" pageNumber="21" unit="m" value="1.65">1,650 m</quantity>
asl
</emphasis>
: São Paulo State, Campos do Jordão, mosses or aquatic plants (du
<bibRefCitation id="EFB76CC9FF887A284EA2BBF7548DF75A" author="Bois-Reymond" pageId="20" pageNumber="21" refString="Bois-Reymond Marcus, E. du. (1944) Sobre tardigrados brasileiros. Comunicaciones Zoologicas del Museo de Historia Natural de Montevideo, 1 (13), 1 - 19." type="journal article" year="1944">Bois-Reymond Marcus 1944</bibRefCitation>
) and mosses (de
<bibRefCitation id="EFB76CC9FF887A284BB9BBE85647F75A" author="Barros" box="[427,567,1022,1044]" pageId="20" pageNumber="21" refString="De Barros, R. (1942 a) Tardigrados de Estado de Sao Paulo, Brasil. I. Introducao. Generos Echiniscus e Pseudechiniscus. Revista Brasileira de Biologia, 2, 257 - 269." type="journal article" year="1942" yearSuffix="a">Barros 1942a</bibRefCitation>
).
<emphasis id="B952CD2AFF887A28485ABBE95008F75A" bold="true" box="[584,1144,1022,1044]" pageId="20" pageNumber="21">
de
<bibRefCitation id="EFB76CC9FF887A284875BBE95779F75A" author="Barros" box="[615,777,1022,1044]" pageId="20" pageNumber="21" refString="De Barros, R. (1942 a) Tardigrados de Estado de Sao Paulo, Brasil. I. Introducao. Generos Echiniscus e Pseudechiniscus. Revista Brasileira de Biologia, 2, 257 - 269." type="journal article" year="1942" yearSuffix="a">Barros (1942a)</bibRefCitation>
, du
<bibRefCitation id="EFB76CC9FF887A284925BBE95008F75A" box="[823,1144,1022,1044]" pageId="20" pageNumber="21" refString="Bois-Reymond Marcus, E. du. (1944) Sobre tardigrados brasileiros. Comunicaciones Zoologicas del Museo de Historia Natural de Montevideo, 1 (13), 1 - 19." type="journal article">Bois-Reymond Marcus (1944)</bibRefCitation>
</emphasis>
</paragraph>
<paragraph id="8B991138FF887A284A85BC09579EF77D" blockId="20.[151,1436,802,1309]" box="[151,1006,1053,1075]" pageId="20" pageNumber="21">
<emphasis id="B952CD2AFF887A284A85BC0955BFF77D" box="[151,463,1053,1075]" italics="true" pageId="20" pageNumber="21">
<geoCoordinate id="EE1277FFFF887A284AD5BC0A5568F77D" box="[199,280,1053,1075]" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="925" value="-23.95">23°57S</geoCoordinate>
,
<geoCoordinate id="EE1277FFFF887A284B30BC0A550AF77D" box="[290,378,1053,1075]" direction="west" orientation="longitude" pageId="20" pageNumber="21" precision="925" value="-46.316666">46°19W</geoCoordinate>
; 0 m asl
</emphasis>
: São Paulo State, Santos, mosses.
<emphasis id="B952CD2AFF887A28493CBC0A579EF77D" bold="true" box="[814,1006,1053,1075]" pageId="20" pageNumber="21">
de
<bibRefCitation id="EFB76CC9FF887A28495CBC0A579EF77D" author="Barros" box="[846,1006,1053,1075]" pageId="20" pageNumber="21" refString="De Barros, R. (1942 a) Tardigrados de Estado de Sao Paulo, Brasil. I. Introducao. Generos Echiniscus e Pseudechiniscus. Revista Brasileira de Biologia, 2, 257 - 269." type="journal article" year="1942" yearSuffix="a">Barros (1942a)</bibRefCitation>
</emphasis>
</paragraph>
<paragraph id="8B991138FF887A284A85BC2B5579F71C" blockId="20.[151,1436,802,1309]" box="[151,265,1084,1106]" pageId="20" pageNumber="21">
<emphasis id="B952CD2AFF887A284A85BC2B5579F71C" bold="true" box="[151,265,1084,1106]" pageId="20" pageNumber="21">
<collectingCountry id="F33151A8FF887A284A85BC2B5574F71C" box="[151,260,1084,1106]" name="Colombia" pageId="20" pageNumber="21">Colombia</collectingCountry>
:
</emphasis>
</paragraph>
<paragraph id="8B991138FF887A284A85BC4C50F7F7C1" blockId="20.[151,1436,802,1309]" pageId="20" pageNumber="21">
<emphasis id="B952CD2AFF887A284A85BC4C5659F73E" box="[151,553,1114,1136]" italics="true" pageId="20" pageNumber="21">
<geoCoordinate id="EE1277FFFF887A284ADABC4D556DF73E" box="[200,285,1114,1136]" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="925" value="4.483333">04°29N</geoCoordinate>
,
<geoCoordinate id="EE1277FFFF887A284B23BC4D55F9F73E" box="[305,393,1114,1136]" direction="west" orientation="longitude" pageId="20" pageNumber="21" precision="925" value="-73.95">73°57W</geoCoordinate>
;
<quantity id="4CDEBCDDFF887A284BB2BC4D558CF73E" box="[416,508,1114,1136]" metricMagnitude="0" metricUnit="m" metricValue="2.3" pageId="20" pageNumber="21" unit="m" value="2.3">2,300 m</quantity>
asl
</emphasis>
: Cundinamarca Department, near Ubaque, below Páramo Cruz Verde, mosses (
<emphasis id="B952CD2AFF887A284AC3BC6E5651F7C1" box="[209,545,1145,1167]" italics="true" pageId="20" pageNumber="21">Erythrodontrium consanguineum</emphasis>
,
<taxonomicName id="4C266ABBFF887A28483CBC6E5605F7C1" box="[558,629,1145,1167]" class="Heterotardigrada" family="Pottiaceae" genus="Tortula" kingdom="Animalia" order="Pottiales" pageId="20" pageNumber="21" phylum="Tardigrada" rank="genus">
<emphasis id="B952CD2AFF887A28483CBC6E5605F7C1" box="[558,629,1145,1167]" italics="true" pageId="20" pageNumber="21">Tortula</emphasis>
</taxonomicName>
sp.,
<taxonomicName id="4C266ABBFF887A2848B5BC6E5756F7C1" box="[679,806,1145,1167]" class="Heterotardigrada" family="Dicranaceae" genus="Campylopus" kingdom="Animalia" order="Dicranales" pageId="20" pageNumber="21" phylum="Tardigrada" rank="genus">
<emphasis id="B952CD2AFF887A2848B5BC6E5756F7C1" box="[679,806,1145,1167]" italics="true" pageId="20" pageNumber="21">Campylopus</emphasis>
</taxonomicName>
sp.,
<taxonomicName id="4C266ABBFF887A284947BC6E57B0F7C1" box="[853,960,1145,1167]" class="Heterotardigrada" family="Sphagnaceae" genus="Sphagnum" kingdom="Animalia" order="Sphagnales" pageId="20" pageNumber="21" phylum="Tardigrada" rank="genus">
<emphasis id="B952CD2AFF887A284947BC6E57B0F7C1" box="[853,960,1145,1167]" italics="true" pageId="20" pageNumber="21">Sphagnum</emphasis>
</taxonomicName>
sp.).
<bibRefCitation id="EFB76CC9FF887A2849EBBC6E50F7F7C1" author="Heinis" box="[1017,1159,1145,1167]" pageId="20" pageNumber="21" refString="Heinis, F. (1914) Die Moosfauna Columbiens. Memoires de la Societe des Sciences Naturelles de Neuchatel, 5, 713 - 724." type="journal article" year="1914">
<emphasis id="B952CD2AFF887A2849EBBC6E50F7F7C1" bold="true" box="[1017,1159,1145,1167]" pageId="20" pageNumber="21">Heinis (1914)</emphasis>
</bibRefCitation>
</paragraph>
<paragraph id="8B991138FF887A284A85BC8056A4F7FE" blockId="20.[151,1436,802,1309]" box="[151,724,1175,1200]" pageId="20" pageNumber="21">
<emphasis id="B952CD2AFF887A284A85BC805518F7FE" bold="true" box="[151,360,1175,1200]" pageId="20" pageNumber="21">Record numbers:</emphasis>
<collectingCountry id="F33151A8FF887A284B7DBC8055C8F7FE" box="[367,440,1175,1200]" name="Brazil" pageId="20" pageNumber="21">Brazil</collectingCountry>
: 2,
<collectingCountry id="F33151A8FF887A284BCCBC805621F7FE" box="[478,593,1175,1200]" name="Colombia" pageId="20" pageNumber="21">Colombia</collectingCountry>
: 1;
<emphasis id="B952CD2AFF887A28486ABC8056A4F7FE" bold="true" box="[632,724,1175,1200]" pageId="20" pageNumber="21">total: 3.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33C42B3FF887A284A85BCAB56F8F653" pageId="20" pageNumber="21" type="discussion">
<paragraph id="8B991138FF887A284A85BCAB56F8F653" blockId="20.[151,1436,802,1309]" pageId="20" pageNumber="21">
<emphasis id="B952CD2AFF887A284A85BCAB557DF79B" bold="true" box="[151,269,1212,1237]" pageId="20" pageNumber="21">Remarks:</emphasis>
This species known from numerous European localities (
<taxonomicName id="4C266ABBFF887A2849C2BCAA502BF79A" box="[976,1115,1213,1236]" class="Heterotardigrada" family="Lycaenidae" genus="Terra" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="90" phylum="Tardigrada" rank="species" species="typica">
<emphasis id="B952CD2AFF887A2849C2BCAA502BF79A" box="[976,1115,1213,1236]" italics="true" pageId="20" pageNumber="21">terra typica</emphasis>
</taxonomicName>
) and single sites in Asia,
<collectingCountry id="F33151A8FF887A284AD5BCC85547F7B6" box="[199,311,1247,1272]" name="Indonesia" pageId="20" pageNumber="21">Indonesia</collectingCountry>
,
<collectingCountry id="F33151A8FF887A284B54BCC855C0F7B6" box="[326,432,1247,1272]" name="Australia" pageId="20" pageNumber="21">Australia</collectingCountry>
,
<collectingCountry id="F33151A8FF887A284BD2BCF7562EF7B6" box="[448,606,1247,1272]" name="New Zealand" pageId="20" pageNumber="21">New Zealand</collectingCountry>
, North and South
<collectingCountry id="F33151A8FF887A28492FBCC857EDF7B6" box="[829,925,1247,1272]" name="United States of America" pageId="20" pageNumber="21">America</collectingCountry>
(
<bibRefCitation id="EFB76CC9FF887A2849A3BCF75019F7B6" author="McInnes" box="[945,1129,1248,1272]" pageId="20" pageNumber="21" refString="McInnes, S. J. (1994 a) Zoogeographic distribution of terrestrial / freshwater tardigrades from current literature. Journal of Natural History, 28, 257 - 352. http: // dx. doi. org / 10.1080 / 00222939400770131" type="journal article" year="1994" yearSuffix="a">McInnes 1994a</bibRefCitation>
). Such a wide geographic distribution suggests a species complex.
</paragraph>
</subSubSection>
</treatment>
</document>