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<document id="AFA6F060567A8B5A70464B1CF89BE02A" ID-DOI="10.5852/ejt.2024.957.2651" ID-ISSN="2118-9773" ID-Zenodo-Dep="13773385" ID-ZooBank="9957F2DF-3F31-4B58-84BC-2F8E0F45C6E3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726582870537" checkinUser="plazi" docAuthor="Selnekovič, Dávid, Jäch, Manfred A. &amp; Kodada, Ján" docDate="2024" docId="038B5001DA76E7CDFDC9FB80FE5A2513" docLanguage="en" docName="EJT.2024.957.1-229.pdf" docOrigin="European Journal of Taxonomy 957" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2651/12247" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Elmomorphus simplipes Selnekovič, Jäch &amp; Kodada, 2024, sp. nov." docType="treatment" docUuid="85D2B8A7-D3C1-4D2C-8765-A3D3181E5489" docUuidSource="ZooBank" docVersion="2" lastPageNumber="131" masterDocId="FFB22879DA08E74FFF9EFFBAFFE32440" masterDocTitle="Taxonomic revision of the genus Elmomorphus Sharp, 1888. II. Redescription of the genus and review of the species from India, Nepal, Bhutan, Myanmar, China, Thailand, Laos, Cambodia, and Vietnam (Coleoptera: Dryopidae)" masterLastPageNumber="229" masterPageNumber="1" pageNumber="127" updateTime="1726584008522" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="1AFD23A0C86F2645A5A58C767F964323">Taxonomic revision of the genus Elmomorphus Sharp, 1888. II. Redescription of the genus and review of the species from India, Nepal, Bhutan, Myanmar, China, Thailand, Laos, Cambodia, and Vietnam (Coleoptera: Dryopidae)</mods:title>
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<mods:namePart id="6C98172CE81A066B476648659D1E4396">Selnekovič, Dávid</mods:namePart>
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<mods:affiliation id="E4BC7EF6E4481E81C4F88E28EAE3F9B8">Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Ilkovičova 6, SK- 842 15 Bratislava, Slovakia.</mods:affiliation>
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<mods:namePart id="9FBDC4193E1D8B72FA5249ED5DD0C3D0">Jäch, Manfred A.</mods:namePart>
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<mods:affiliation id="0E493A7EC1BE719E429A9D5AD1E478B2">Naturhistorisches Museum Wien, Burgring 7, A- 1010 Wien, Austria.</mods:affiliation>
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<mods:namePart id="6BF54EAAC9A2C16B4F80BFB33D80650A">Kodada, Ján</mods:namePart>
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<mods:affiliation id="F1A978ACC771F0D8C9225D8D3E2DB6C1">Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Ilkovičova 6, SK- 842 15 Bratislava, Slovakia.</mods:affiliation>
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<mods:date id="207E7A29418699E1F72979EC63C16399">2024</mods:date>
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<heading id="D0D5567BDA76E731FDC9FB80FC3E2015" box="[599,989,1082,1109]" centered="true" fontSize="11" level="2" pageId="126" pageNumber="127" reason="2">
<taxonomicName id="4C229A94DA76E731FDC9FB80FC982015" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[599,891,1082,1109]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="simplipes" status="sp. nov.">
<emphasis id="B9563D05DA76E731FDC9FB80FC982015" bold="true" box="[599,891,1082,1109]" italics="true" pageId="126" pageNumber="127">Elmomorphus simplipes</emphasis>
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<paragraph id="8B9DE117DA76E731FD0EFBC5FC4020DA" blockId="126.[364,1223,1082,1178]" box="[656,931,1151,1178]" pageId="126" pageNumber="127">
<figureCitation id="1319FD92DA76E731FD0EFBC5FCF520D9" box="[656,790,1151,1177]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="127.[189,232,1966,1992]" captionStartId-1="128.[189,232,1952,1978]" captionTargetBox-0="[189,1399,265,1928]" captionTargetBox-1="[319,1240,278,1892]" captionTargetId-0="figure-15@127.[189,1399,265,1928]" captionTargetId-1="figure-21@128.[306,1262,265,1916]" captionTargetPageId-0="127" captionTargetPageId-1="128" captionText-0="Fig. 61. Elmomorphus simplipes sp. nov., paratype, male from type locality (CKB), TL: 2.81 mm." captionText-1="Fig. 62. Elmomorphus simplipes sp. nov., holotype (NMW), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13773557" figureDoi-1="http://doi.org/10.5281/zenodo.13773559" httpUri-0="https://zenodo.org/record/13773557/files/figure.png" httpUri-1="https://zenodo.org/record/13773559/files/figure.png" pageId="126" pageNumber="127">Figs 61–62</figureCitation>
, 
<figureCitation id="1319FD92DA76E731FCBCFBC5FCB420D9" box="[802,855,1151,1177]" captionStart="Fig" captionStartId="137.[189,232,1893,1919]" captionTargetBox="[200,1398,275,1860]" captionTargetId="figure-52@137.[189,1398,265,1868]" captionTargetPageId="137" captionText="Fig. 67. Bursa copulatrix and vagina. A. Elmomorphus simplipes sp. nov., paratype from type locality (CKB).B. E. donatus Kodada, Selnekovič &amp; Jäch sp. nov., holotype (NMW), lateral aspect. C. E. donatus, holotype (NMW), ventral aspect. D. E. paradonatus sp. nov., holotype (IAECAS). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773575" httpUri="https://zenodo.org/record/13773575/files/figure.png" pageId="126" pageNumber="127">67A</figureCitation>
, 
<figureCitation id="1319FD92DA76E731FCFDFB3AFC4020DA" box="[867,931,1152,1178]" captionStart="Fig" captionStartId="220.[189,232,1871,1897]" captionTargetBox="[254,1334,266,1829]" captionTargetId="figure-44@220.[189,1399,265,1830]" captionTargetPageId="220" captionText="Fig. 112. Distribution maps. A. Elmomorphus bispinosus sp. nov. B. E. nepalensis Satô, 1981. C. E. punctulatus sp. nov. D. E. simplipes sp. nov. E. E. donatus Kodada, Selnekovič &amp; Jäch sp. nov. F. E. paradonatus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13773718" httpUri="https://zenodo.org/record/13773718/files/figure.png" pageId="126" pageNumber="127">112D</figureCitation>
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<emphasis id="B9563D05DA76E731FF23FB7AFE2A209A" bold="true" box="[189,457,1216,1242]" pageId="126" pageNumber="127">Differential diagnosis</emphasis>
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<taxonomicName id="4C229A94DA76E731FF23FB54FE342148" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[189,471,1262,1288]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="simplipes" status="sp. nov.">
<emphasis id="B9563D05DA76E731FF23FB54FE342148" box="[189,471,1262,1288]" italics="true" pageId="126" pageNumber="127">Elmomorphus simplipes</emphasis>
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(
<figureCitation id="1319FD92DA76E731FDD8FB54FD792148" box="[582,666,1262,1288]" captionStart="Fig" captionStartId="127.[189,232,1966,1992]" captionTargetBox="[189,1399,265,1928]" captionTargetId="figure-15@127.[189,1399,265,1928]" captionTargetPageId="127" captionText="Fig. 61. Elmomorphus simplipes sp. nov., paratype, male from type locality (CKB), TL: 2.81 mm." figureDoi="http://doi.org/10.5281/zenodo.13773557" httpUri="https://zenodo.org/record/13773557/files/figure.png" pageId="126" pageNumber="127">Fig. 61</figureCitation>
) belongs to a group of species characterised by having a dorsal plastron distributed on the anterior portions of the head and the posterolateral portions of the elytra. Elytral punctures are scattered over the entire surface and striae are absent. Males bear conspicuous, long, erect setae or setal clusters on the labrum, prosternal process, and metaventrite. The metaventrite is strongly depressed in males. With 
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<emphasis id="B9563D05DA76E731FDFAFAC2FCF421D1" box="[612,791,1399,1425]" italics="true" pageId="126" pageNumber="127">E. paradonatus</emphasis>
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<taxonomicNameLabel id="A265807EDA76E731FC81FACDFC9A21D1" box="[799,889,1399,1425]" pageId="126" pageNumber="127" rank="species">sp. nov.</taxonomicNameLabel>
and 
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<emphasis id="B9563D05DA76E731FC30FAC2FBC921D1" box="[942,1066,1399,1425]" italics="true" pageId="126" pageNumber="127">E. donatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDA76E731FBAEFACDFB6F21D1" box="[1072,1164,1399,1425]" pageId="126" pageNumber="127" rank="species">sp. nov.</taxonomicNameLabel>
, it shares the simple metatibiae without teeth. However, 
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<emphasis id="B9563D05DA76E731FDFFFA20FD1321F3" box="[609,752,1433,1459]" italics="true" pageId="126" pageNumber="127">E. simplipes</emphasis>
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can be distinguished from the latter two species by the smaller and less convex body (TL in 
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<emphasis id="B9563D05DA76E731FDE0FA06FCED2195" box="[638,782,1467,1493]" italics="true" pageId="126" pageNumber="127">E. simplipes</emphasis>
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: 
<specimenCount id="9D242A9EDA76E731FC81FA06FCA82196" box="[799,843,1468,1494]" count="1" pageId="126" pageNumber="127" type="male">♂♂</specimenCount>
<quantity id="4CDA4CF2DA76E731FCCAFA01FBE32196" box="[852,1024,1467,1494]" metricMagnitude="-3" metricUnit="m" metricValue="2.78" metricValueMax="2.97" metricValueMin="2.59" pageId="126" pageNumber="127" unit="mm" value="2.78" valueMax="2.97" valueMin="2.59">2.59–2.97 mm</quantity>
(2.81 ±0.09, n =25), 
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<quantity id="4CDA4CF2DA76E731FAAAFA06FEC621B8" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.19" metricValueMin="2.91" pageId="126" pageNumber="127" unit="mm" value="3.05" valueMax="3.19" valueMin="2.91">2.91– 3.19 mm</quantity>
(3.01± 0.08, n= 20), in 
<taxonomicName id="4C229A94DA76E731FDA4FA65FD5421B8" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[570,695,1502,1528]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="donatus">
<emphasis id="B9563D05DA76E731FDA4FA65FD5421B8" box="[570,695,1502,1528]" italics="true" pageId="126" pageNumber="127">E. donatus</emphasis>
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: 
<specimenCount id="9D242A9EDA76E731FD58FA64FD1121B8" box="[710,754,1502,1528]" count="1" pageId="126" pageNumber="127" type="male">♂♂</specimenCount>
<quantity id="4CDA4CF2DA76E731FD66FA64FC4121B8" box="[760,930,1502,1529]" metricMagnitude="-3" metricUnit="m" metricValue="3.295" metricValueMax="3.5" metricValueMin="3.09" pageId="126" pageNumber="127" unit="mm" value="3.295" valueMax="3.5" valueMin="3.09">3.09–3.50 mm</quantity>
(3.30± 0.10, n=20), 
<specimenCount id="9D242A9EDA76E731FB04FA64FB2521B8" box="[1178,1222,1502,1528]" count="1" pageId="126" pageNumber="127" type="female">♀♀</specimenCount>
<quantity id="4CDA4CF2DA76E731FB53FA64FA9421B8" box="[1229,1399,1502,1529]" metricMagnitude="-3" metricUnit="m" metricValue="3.505" metricValueMax="3.66" metricValueMin="3.35" pageId="126" pageNumber="127" unit="mm" value="3.505" valueMax="3.66" valueMin="3.35">3.35–3.66 mm</quantity>
(3.51 ± 0.09, n= 19), in 
<taxonomicName id="4C229A94DA76E731FE42F9BBFD76225A" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[476,661,1536,1562]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="paradonatus">
<emphasis id="B9563D05DA76E731FE42F9BBFD76225A" box="[476,661,1536,1562]" italics="true" pageId="126" pageNumber="127">E. paradonatus</emphasis>
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<specimenCount id="9D242A9EDA76E731FD3FF9BBFD2E225B" box="[673,717,1537,1563]" count="1" pageId="126" pageNumber="127" type="male">♂♂</specimenCount>
<quantity id="4CDA4CF2DA76E731FD47F9BAFC6B225B" box="[729,904,1536,1563]" metricMagnitude="-3" metricUnit="m" metricValue="3.275" metricValueMax="3.58" metricValueMin="2.97" pageId="126" pageNumber="127" unit="mm" value="3.275" valueMax="3.58" valueMin="2.97">2.97–3.58 mm</quantity>
(3.24± 0.14, n=20), 
<specimenCount id="9D242A9EDA76E731FB0EF9BBFB5F225B" box="[1168,1212,1537,1563]" count="1" pageId="126" pageNumber="127" type="female">♀♀</specimenCount>
<quantity id="4CDA4CF2DA76E731FB56F9BAFA94225B" box="[1224,1399,1536,1563]" metricMagnitude="-3" metricUnit="m" metricValue="3.465" metricValueMax="3.79" metricValueMin="3.14" pageId="126" pageNumber="127" unit="mm" value="3.465" valueMax="3.79" valueMin="3.14">3.14–3.79 mm</quantity>
(3.34 ± 0.14, n=20) (
<figureCitation id="1319FD92DA76E731FE2AF998FDE9227C" box="[436,522,1570,1596]" captionStart="Fig" captionStartId="127.[189,232,1966,1992]" captionTargetBox="[189,1399,265,1928]" captionTargetId="figure-15@127.[189,1399,265,1928]" captionTargetPageId="127" captionText="Fig. 61. Elmomorphus simplipes sp. nov., paratype, male from type locality (CKB), TL: 2.81 mm." figureDoi="http://doi.org/10.5281/zenodo.13773557" httpUri="https://zenodo.org/record/13773557/files/figure.png" pageId="126" pageNumber="127">Fig. 61</figureCitation>
) and by the arrangement of the spines of the bursa copulatrix (
<figureCitation id="1319FD92DA76E731FB65F998FA89227C" box="[1275,1386,1570,1596]" captionStart="Fig" captionStartId="137.[189,232,1893,1919]" captionTargetBox="[200,1398,275,1860]" captionTargetId="figure-52@137.[189,1398,265,1868]" captionTargetPageId="137" captionText="Fig. 67. Bursa copulatrix and vagina. A. Elmomorphus simplipes sp. nov., paratype from type locality (CKB).B. E. donatus Kodada, Selnekovič &amp; Jäch sp. nov., holotype (NMW), lateral aspect. C. E. donatus, holotype (NMW), ventral aspect. D. E. paradonatus sp. nov., holotype (IAECAS). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773575" httpUri="https://zenodo.org/record/13773575/files/figure.png" pageId="126" pageNumber="127">Fig. 67A</figureCitation>
). 
<taxonomicName id="4C229A94DA76E731FF23F9FFFE2D221E" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[189,462,1605,1631]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="curvipes" status="sp. nov.">
<emphasis id="B9563D05DA76E731FF23F9FFFE2D221E" box="[189,462,1605,1631]" italics="true" pageId="126" pageNumber="127">Elmomorphus curvipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDA76E731FE45F9FFFDDB221F" box="[475,568,1605,1631]" pageId="126" pageNumber="127" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="4C229A94DA76E731FDD6F9FFFD2E221F" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[584,717,1605,1631]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="dentipes" status="sp. nov.">
<emphasis id="B9563D05DA76E731FDD6F9FFFD2E221F" box="[584,717,1605,1631]" italics="true" pageId="126" pageNumber="127">E. dentipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDA76E731FD49F9FFFCD7221F" box="[727,820,1605,1631]" pageId="126" pageNumber="127" rank="species">sp. nov.</taxonomicNameLabel>
, and 
<taxonomicName id="4C229A94DA76E731FCE6F9FFFC0B221F" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[888,1000,1605,1631]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
<emphasis id="B9563D05DA76E731FCE6F9FFFC0B221F" box="[888,1000,1605,1631]" italics="true" pageId="126" pageNumber="127">E. similis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDA76E731FC71F9FFFBAF221F" box="[1007,1100,1605,1631]" pageId="126" pageNumber="127" rank="species">sp. nov.</taxonomicNameLabel>
differ in the presence of metatibial teeth in males (
<figureCitation id="1319FD92DA76E731FE67F9DDFDB622C1" box="[505,597,1639,1665]" captionStart="Fig" captionStartId="140.[189,232,1969,1995]" captionTargetBox="[189,1399,265,1928]" captionTargetId="figure-16@140.[189,1399,265,1928]" captionTargetPageId="140" captionText="Fig. 68. Elmomorphus similis sp. nov., male from China, locality CWBS 354 (CKB), TL: 3.13 mm." figureDoi="http://doi.org/10.5281/zenodo.13773577" httpUri="https://zenodo.org/record/13773577/files/figure.png" pageId="126" pageNumber="127">Figs 68</figureCitation>
, 
<figureCitation id="1319FD92DA76E731FDFAF9DDFD6722C1" box="[612,644,1639,1665]" captionStart="Fig" captionStartId="144.[189,232,1968,1994]" captionTargetBox="[189,1399,265,1928]" captionTargetId="figure-15@144.[189,1399,265,1928]" captionTargetPageId="144" captionText="Fig. 70. Elmomorphus dentipes sp. nov., paratype, male from type locality (NMW), TL: 2.90 mm." figureDoi="http://doi.org/10.5281/zenodo.13773583" httpUri="https://zenodo.org/record/13773583/files/figure.png" pageId="126" pageNumber="127">70</figureCitation>
, 
<figureCitation id="1319FD92DA76E731FD0CF9DDFD5222C1" box="[658,689,1639,1665]" captionStart="Fig" captionStartId="149.[189,232,1973,1999]" captionTargetBox="[188,1398,265,1928]" captionTargetId="figure-15@149.[188,1398,265,1928]" captionTargetPageId="149" captionText="Fig. 72. Elmomorphus curvipes sp. nov., paratype, male from type locality (NMW), TL: 3.20 mm." figureDoi="http://doi.org/10.5281/zenodo.13773589" httpUri="https://zenodo.org/record/13773589/files/figure.png" pageId="126" pageNumber="127">72</figureCitation>
) and the arrangement of the spines of the bursa copulatrix (
<figureCitation id="1319FD92DA76E731FF58F933FEFC22E3" box="[198,287,1673,1699]" captionStart="Fig" captionStartId="151.[189,232,1871,1897]" captionTargetBox="[199,1382,273,1811]" captionTargetId="figure-50@151.[189,1395,265,1828]" captionTargetPageId="151" captionText="Fig. 74. Bursa copulatrix and vagina. A. Elmomorphus similis sp. nov., specimen from China, locality CWBS 378 (NMW). B. E. dentipes sp. nov., specimen from China, locality CWBS 354 (NMW). C. E. curvipes sp. nov., specimen from China, locality CWBS 260 (NMW). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773597" httpUri="https://zenodo.org/record/13773597/files/figure.png" pageId="126" pageNumber="127">Fig. 74</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C338B29CDA76E731FF23F970FE762374" pageId="126" pageNumber="127" type="etymology">
<paragraph id="8B9DE117DA76E731FF23F970FEA722A4" blockId="126.[189,324,1738,1764]" box="[189,324,1738,1764]" pageId="126" pageNumber="127">
<heading id="D0D5567BDA76E731FF23F970FEA722A4" bold="true" box="[189,324,1738,1764]" fontSize="11" level="3" pageId="126" pageNumber="127" reason="3">
<emphasis id="B9563D05DA76E731FF23F970FEA722A4" bold="true" box="[189,324,1738,1764]" pageId="126" pageNumber="127">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DA76E731FF23F94DFE762374" blockId="126.[189,1398,1783,1844]" pageId="126" pageNumber="127">The epithet, a Latin adjective in the nominative singular, refers to the tibiae lacking teeth, swellings, or apical expansions.</paragraph>
</subSubSection>
<subSubSection id="C338B29CDA76E7CEFF23F8E0FD7825F2" lastPageId="129" lastPageNumber="130" pageId="126" pageNumber="127" type="materials_examined">
<paragraph id="8B9DE117DA76E731FF23F8E0FE8F2334" blockId="126.[189,364,1882,1908]" box="[189,364,1882,1908]" pageId="126" pageNumber="127">
<heading id="D0D5567BDA76E731FF23F8E0FE8F2334" bold="true" box="[189,364,1882,1908]" fontSize="11" level="3" pageId="126" pageNumber="127" reason="3">
<emphasis id="B9563D05DA76E731FF23F8E0FE8F2334" bold="true" box="[189,364,1882,1908]" pageId="126" pageNumber="127">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DA76E731FF74F832FEBF23E2" blockId="126.[189,1399,1928,2025]" box="[234,348,1928,1954]" pageId="126" pageNumber="127">
<emphasis id="B9563D05DA76E731FF74F832FEBF23E2" bold="true" box="[234,348,1928,1954]" pageId="126" pageNumber="127">
<typeStatus id="54995FB5DA76E731FF74F832FEBF23E2" box="[234,348,1928,1954]" pageId="126" pageNumber="127" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B9DE117DA76E731FF23F810FE4F23A9" blockId="126.[189,1399,1928,2025]" pageId="126" pageNumber="127">
<materialsCitation id="3B4AEB4ADA76E731FF23F810FE4F23A9" collectingDate="1996-06-08" collectionCode="NMW" collectorName="Dembicky" country="Vietnam" location="Vinh Phuc Province" pageId="126" pageNumber="127" specimenCount="1" specimenCount-male="1" stateProvince="Vinh Phuc" typeStatus="holotype">
<collectingCountry id="F335A187DA76E731FF23F810FEA72384" box="[189,324,1962,1988]" name="Vietnam" pageId="126" pageNumber="127">VIETNAM</collectingCountry>
– 
<emphasis id="B9563D05DA76E731FEF9F810FD862384" bold="true" box="[359,613,1961,1988]" pageId="126" pageNumber="127">
<collectingRegion id="49E62FF5DA76E731FEF9F810FD862384" box="[359,613,1961,1988]" country="Vietnam" name="Vinh Phuc" pageId="126" pageNumber="127">Vinh Phúc Province</collectingRegion>
</emphasis>
• 
<specimenCount id="9D242A9EDA76E731FD1AF811FD7E2385" box="[644,669,1963,1989]" count="1" pageId="126" pageNumber="127" type="male">♂</specimenCount>
; “N-VIETNAM: Tam Dao (2) 1.- 
<date id="FF9CC7D7DA76E731FBB5F810FB7F2384" box="[1067,1180,1962,1988]" pageId="126" pageNumber="127" value="1996-06-08">
<collectingDate id="EFD83E3FDA76E731FBB5F810FB7F2384" box="[1067,1180,1962,1988]" pageId="126" pageNumber="127" value="1996-06-08">8.6.1996</collectingDate>
</date>
leg. 
<collectorName id="26D784C1DA76E731FB45F810FAB62384" box="[1243,1365,1962,1988]" pageId="126" pageNumber="127">Dembicky</collectorName>
&amp; Pacholatko”; 
<collectionCode id="ED3379D2DA76E731FEC2F875FE4B23A9" box="[348,424,1999,2025]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="126" pageNumber="127">NMW</collectionCode>
.
</materialsCitation>
</paragraph>
<caption id="DF5DB19FDA77E730FF23F814FAD92389" ID-DOI="http://doi.org/10.5281/zenodo.13773557" ID-Zenodo-Dep="13773557" box="[189,1338,1966,1993]" httpUri="https://zenodo.org/record/13773557/files/figure.png" pageId="127" pageNumber="128" startId="127.[189,232,1966,1992]" targetBox="[189,1399,265,1928]" targetPageId="127">
<paragraph id="8B9DE117DA77E730FF23F814FAD92389" blockId="127.[189,1338,1966,1993]" box="[189,1338,1966,1993]" pageId="127" pageNumber="128">
<emphasis id="B9563D05DA77E730FF23F814FEFA2388" bold="true" box="[189,281,1966,1992]" pageId="127" pageNumber="128">Fig. 61.</emphasis>
<taxonomicName id="4C229A94DA77E730FEBEF814FDD82388" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[288,571,1966,1992]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="127" pageNumber="128" phylum="Arthropoda" rank="species" species="simplipes" status="sp. nov.">
<emphasis id="B9563D05DA77E730FEBEF814FDD82388" box="[288,571,1966,1992]" italics="true" pageId="127" pageNumber="128">Elmomorphus simplipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDA77E730FDDCF815FD432389" box="[578,672,1967,1993]" pageId="127" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, male from type locality (CKB), TL: 2.81 mm.
</paragraph>
</caption>
<caption id="DF5DB19FDA88E7CFFF23F81AFE11239E" ID-DOI="http://doi.org/10.5281/zenodo.13773559" ID-Zenodo-Dep="13773559" httpUri="https://zenodo.org/record/13773559/files/figure.png" pageId="128" pageNumber="129" startId="128.[189,232,1952,1978]" targetBox="[319,1240,278,1892]" targetPageId="128">
<paragraph id="8B9DE117DA88E7CFFF23F81AFE11239E" blockId="128.[189,1399,1952,2014]" pageId="128" pageNumber="129">
<emphasis id="B9563D05DA88E7CFFF23F81AFEFE23FA" bold="true" box="[189,285,1952,1978]" pageId="128" pageNumber="129">Fig. 62.</emphasis>
<taxonomicName id="4C229A94DA88E7CFFEB9F81AFDA623FA" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[295,581,1952,1978]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="128" pageNumber="129" phylum="Arthropoda" rank="species" species="simplipes" status="sp. nov.">
<emphasis id="B9563D05DA88E7CFFEB9F81AFDA623FA" box="[295,581,1952,1978]" italics="true" pageId="128" pageNumber="129">Elmomorphus simplipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDA88E7CFFDD1F81BFD4C23FB" box="[591,687,1953,1979]" pageId="128" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (NMW), aedeagus. 
<emphasis id="B9563D05DA88E7CFFB83F81AFBD123FA" bold="true" box="[1053,1074,1952,1978]" pageId="128" pageNumber="129">A</emphasis>
. Lateral aspect. 
<emphasis id="B9563D05DA88E7CFFB63F81BFAF223FB" bold="true" box="[1277,1297,1953,1979]" pageId="128" pageNumber="129">B</emphasis>
. Ventral aspect. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8B9DE117DA89E7CEFF74FEB3FE842563" blockId="129.[189,1399,265,325]" box="[234,359,265,291]" pageId="129" pageNumber="130">
<emphasis id="B9563D05DA89E7CEFF74FEB3FE842563" bold="true" box="[234,359,265,291]" pageId="129" pageNumber="130">
<typeStatus id="54995FB5DA89E7CEFF74FEB3FE842563" box="[234,359,265,291]" pageId="129" pageNumber="130" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B9DE117DA89E7CEFF23FE90FA942505" blockId="129.[189,1399,265,325]" box="[189,1399,297,325]" pageId="129" pageNumber="130">
<materialsCitation id="3B4AEB4ADA89E7CEFF23FE90FA902505" box="[189,1395,297,325]" collectingDate="1996-06-08" collectionCode="CKB, NMW" collectorName="Dembicky" country="Vietnam" location="Vinh Phuc Province" pageId="129" pageNumber="130" specimenCount="191" specimenCount-female="111" specimenCount-male="80" stateProvince="Vinh Phuc" typeStatus="paratype">
<collectingCountry id="F335A187DA89E7CEFF23FE90FEA72504" box="[189,324,298,324]" name="Vietnam" pageId="129" pageNumber="130">VIETNAM</collectingCountry>
– 
<emphasis id="B9563D05DA89E7CEFEC0FE90FD872504" bold="true" box="[350,612,297,324]" pageId="129" pageNumber="130">
<collectingRegion id="49E62FF5DA89E7CEFEC0FE90FDB72504" box="[350,596,297,324]" country="Vietnam" name="Vinh Phuc" pageId="129" pageNumber="130">Vinh Phúc Province</collectingRegion>
•
</emphasis>
<specimenCount id="9D242A9EDA89E7CEFDF4FE90FD582505" box="[618,699,298,325]" count="80" pageId="129" pageNumber="130" type="male">80 ♂♂</specimenCount>
, 
<specimenCount id="9D242A9EDA89E7CEFD59FE90FCC02505" box="[711,803,298,325]" count="111" pageId="129" pageNumber="130" type="female">111 ♀♀</specimenCount>
; same collection data as for holotype; 
<collectionCode id="ED3379D2DA89E7CEFB43FE90FAFF2504" box="[1245,1308,298,324]" pageId="129" pageNumber="130">CKB</collectionCode>
, 
<collectionCode id="ED3379D2DA89E7CEFAB9FE91FA902505" box="[1319,1395,299,325]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="129" pageNumber="130">NMW</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9DE117DA89E7CEFF23FED0FEBE25C4" blockId="129.[189,349,362,388]" box="[189,349,362,388]" pageId="129" pageNumber="130">
<heading id="D0D5567BDA89E7CEFF23FED0FEBE25C4" bold="true" box="[189,349,362,388]" fontSize="11" level="3" pageId="129" pageNumber="130" reason="3">
<emphasis id="B9563D05DA89E7CEFF23FED0FEBE25C4" bold="true" box="[189,349,362,388]" pageId="129" pageNumber="130">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DA89E7CEFF23FE2DFD7825F2" blockId="129.[189,667,407,434]" box="[189,667,407,434]" pageId="129" pageNumber="130">
<collectingCountry id="F335A187DA89E7CEFF23FE2DFEC025F1" box="[189,291,407,433]" name="Vietnam" pageId="129" pageNumber="130">Vietnam</collectingCountry>
, 
<collectingRegion id="49E62FF5DA89E7CEFEB0FE2DFDFF25F1" box="[302,540,407,433]" country="Vietnam" name="Vinh Phuc" pageId="129" pageNumber="130">Vinh Phúc Province</collectingRegion>
, Tam Dao.
</paragraph>
</subSubSection>
<subSubSection id="C338B29CDA89E7CEFF23FE6DFC7123A8" pageId="129" pageNumber="130" type="description">
<paragraph id="8B9DE117DA89E7CEFF23FE6DFEAD25B1" blockId="129.[189,334,471,497]" box="[189,334,471,497]" pageId="129" pageNumber="130">
<heading id="D0D5567BDA89E7CEFF23FE6DFEAD25B1" bold="true" box="[189,334,471,497]" fontSize="11" level="3" pageId="129" pageNumber="130" reason="3">
<emphasis id="B9563D05DA89E7CEFF23FE6DFEAD25B1" bold="true" box="[189,334,471,497]" pageId="129" pageNumber="130">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DA89E7CEFF23FDBFFC3226E6" blockId="129.[189,1399,516,678]" pageId="129" pageNumber="130">
Measurements (mm): TL: 
<specimenCount id="9D242A9EDA89E7CEFE66FDBFFDC7265F" box="[504,548,517,543]" count="1" pageId="129" pageNumber="130" type="male">♂♂</specimenCount>
2.59–2.97 (2.81± 0.09, n= 25), 
<specimenCount id="9D242A9EDA89E7CEFC3CFDBFFC2D265F" box="[930,974,517,543]" count="1" pageId="129" pageNumber="130" type="female">♀♀</specimenCount>
2.91–3.19 (3.01± 0.08, n= 20), PL: 0.58–0.68 (0.64 ± 0.03, n= 25), 
<specimenCount id="9D242A9EDA89E7CEFDA4FD9DFD852601" box="[570,614,551,577]" count="1" pageId="129" pageNumber="130" type="female">♀♀</specimenCount>
0.63–0.70 (0.66 ±0.02, n=20); PW: 
<specimenCount id="9D242A9EDA89E7CEFBB0FD9DFBB92601" box="[1070,1114,551,577]" count="1" pageId="129" pageNumber="130" type="male">♂♂</specimenCount>
1.15–1.33 (1.26± 0.04, n= 25), 
<specimenCount id="9D242A9EDA89E7CEFE81FDF3FEA82623" box="[287,331,585,611]" count="1" pageId="129" pageNumber="130" type="female">♀♀</specimenCount>
1.25–1.43 (1.35± 0.04, n =20); EL: 
<specimenCount id="9D242A9EDA89E7CEFC8EFDF3FCDF2623" box="[784,828,585,611]" count="1" pageId="129" pageNumber="130" type="male">♂♂</specimenCount>
2.01–2.31 (2.17 ±0.06, n=25), 
<specimenCount id="9D242A9EDA89E7CEFB56FDF3FB172623" box="[1224,1268,585,611]" count="1" pageId="129" pageNumber="130" type="female">♀♀</specimenCount>
2.24–2.50 (2.35 ± 0.06, n=20), EW: 
<specimenCount id="9D242A9EDA89E7CEFE71FDD0FDF826C4" box="[495,539,618,644]" count="1" pageId="129" pageNumber="130" type="male">♂♂</specimenCount>
1.36–1.56 (1.49 ±0.04, n =25), 
<specimenCount id="9D242A9EDA89E7CEFC08FDD0FC2126C4" box="[918,962,618,644]" count="1" pageId="129" pageNumber="130" type="female">♀♀</specimenCount>
1.53–1.69 (1.60± 0.05, n =20); PhL: 0.88–1.00 (0.95± 0.02, n =25); PrL: 0.58–0.64 (0.61± 0.02, n= 25).
</paragraph>
<paragraph id="8B9DE117DA89E7CEFF23FD76FA89270B" blockId="129.[189,1398,716,844]" pageId="129" pageNumber="130">
Body narrowly oval, convex dorsally, widest before midlength of elytra (
<figureCitation id="1319FD92DA89E7CEFBA3FD76FB7A26A6" box="[1085,1177,716,742]" captionStart="Fig" captionStartId="127.[189,232,1966,1992]" captionTargetBox="[189,1399,265,1928]" captionTargetId="figure-15@127.[189,1399,265,1928]" captionTargetPageId="127" captionText="Fig. 61. Elmomorphus simplipes sp. nov., paratype, male from type locality (CKB), TL: 2.81 mm." figureDoi="http://doi.org/10.5281/zenodo.13773557" httpUri="https://zenodo.org/record/13773557/files/figure.png" pageId="129" pageNumber="130">Fig. 61</figureCitation>
). Integument dark brown to black; mouthparts, antennae, trochanters, and tarsi reddish brown. Pubescence consists of very short sparse yellowish setae. Plastron present on anterior portions of head, apicolateral portions of elytra, and on ventral surface, except prosternal process, middle of metaventrite, and middle of ventrites 1–2.
</paragraph>
<paragraph id="8B9DE117DA89E7CEFF23FCCBFEF12016" blockId="129.[189,1399,881,1110]" pageId="129" pageNumber="130">
Head with small round setiferous punctures, slightly smaller than eye facets, separated by about one puncture diameter; microreticulation absent; plastron covering anterior portions. Labrum transverse, anterior margin slightly rounded, surface microreticulate, males with conspicuous setae about twice as long as exposed portion of labrum. Anterior margin of clypeus straight. Antennae 9-segmented. Eyes oval, interfacetal setae short; ID: 
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<quantity id="4CDA4CF2DA89E7CEFD25FC42FC842053" box="[699,871,1016,1043]" metricMagnitude="-4" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="129" pageNumber="130" unit="mm" value="0.425" valueMax="0.45" valueMin="0.4">0.40–0.45 mm</quantity>
(
<quantity id="4CDA4CF2DA89E7CEFCE8FC42FC022053" box="[886,993,1016,1043]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="129" pageNumber="130" unit="mm" value="0.43">0.43 mm</quantity>
±0.01, n=25), 
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<quantity id="4CDA4CF2DA89E7CEFB52FC42FA942053" box="[1228,1399,1016,1043]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="4.9" metricValueMin="4.1" pageId="129" pageNumber="130" unit="mm" value="0.44999999999999996" valueMax="0.49" valueMin="0.41">0.41–0.49 mm</quantity>
(
<quantity id="4CDA4CF2DA89E7CEFF5BFBA0FED52074" box="[197,310,1050,1076]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="129" pageNumber="130" unit="mm" value="0.46">0.46 mm</quantity>
± 0.02, n= 20); APD/ID: 
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1.58–1.78 (1.70 ± 0.04, n=25), 
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1.62–1.82 (1.71 ± 0.05, n= 20).
</paragraph>
<paragraph id="8B9DE117DA89E7CEFF23FBC6FB85226B" blockId="129.[189,1399,1147,1580]" pageId="129" pageNumber="130">
Pronotum wider than long, widest at base; PW/PL: 
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1.85–2.17 (1.99± 0.08, n=20), 
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1.92–2.10 (2.04 ± 0.04, n=20); surface with round setiferous punctures; microreticulation absent; plastron absent; rim of anterior margin as wide as three eye facets; anterior angles moderately deflexed, prominent; lateral pronotal sides convergent, slightly rounded before anterior angles. Lateral margins of prosternal process divergent and straight; posterior edge broadly V-shaped; lateral portions slightly raised, with clusters of long setae in males; median keel slightly arcuate. Scutellum longer than wide, with sparse round punctures, microreticulation absent. Disc of metaventrite in males slightly depressed, with two groups of long setae; lateral margins of metaventral process raised, separated by shallow ridge. Elytra oval, widest before middle, EL/EW: 
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1.41–1.51 (1.45 ±0.02, n= 25), 
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1.43–1.52 (1.47± 0.02, n= 20); dorsal surface with round setiferous punctures; microreticulation very fine; plastron confined to apicolateral portions and to epipleura. Tibiae slightly curved; protibia ca 1.3× as long as protarsus; metatibia simple, without tooth; PrTL/PL: 
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1.09–1.56 (1.19± 0.09, n =25), 
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1.04–1.15 (1.09± 0.03, n= 20). Terminal protarsomere as long as three preceding tarsomeres combined.
</paragraph>
<paragraph id="8B9DE117DA89E7CEFF23F9EBFBA52317" blockId="129.[189,1399,1617,1880]" pageId="129" pageNumber="130">
Ventrites covered with plastron, except for middle of ventrites 1–2. Apical margin of ventrite 5 rounded in males, with small median triangular excision in females; in both sexes with short longitudinal keel before apex. Aedeagus (
<figureCitation id="1319FD92DA89E7CEFD83F92FFD9922EF" box="[541,634,1685,1711]" captionStart="Fig" captionStartId="128.[189,232,1952,1978]" captionTargetBox="[319,1240,278,1892]" captionTargetId="figure-21@128.[306,1262,265,1916]" captionTargetPageId="128" captionText="Fig. 62. Elmomorphus simplipes sp. nov., holotype (NMW), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773559" httpUri="https://zenodo.org/record/13773559/files/figure.png" pageId="129" pageNumber="130">Fig. 62</figureCitation>
): phallobase slightly expanded proximally, PhL/PrL: 1.47–1.70 (1.55± 0.05, n=25); parameres evenly narrowed to apices, curved ventrad, apices narrowly rounded (lateral aspect); penis narrowly rounded apically in dorsal aspect, expanded and curved ventrad in lateral aspect; sclerotised fibula rather slender. Ovipositor: valvifers ca 1.90 × as long as coxites; right coxite ca 1.30 × as long as left one. Bursa copulatrix (
<figureCitation id="1319FD92DA89E7CEFD24F8A6FCC02376" box="[698,803,1820,1846]" captionStart="Fig" captionStartId="137.[189,232,1893,1919]" captionTargetBox="[200,1398,275,1860]" captionTargetId="figure-52@137.[189,1398,265,1868]" captionTargetPageId="137" captionText="Fig. 67. Bursa copulatrix and vagina. A. Elmomorphus simplipes sp. nov., paratype from type locality (CKB).B. E. donatus Kodada, Selnekovič &amp; Jäch sp. nov., holotype (NMW), lateral aspect. C. E. donatus, holotype (NMW), ventral aspect. D. E. paradonatus sp. nov., holotype (IAECAS). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773575" httpUri="https://zenodo.org/record/13773575/files/figure.png" pageId="129" pageNumber="130">Fig. 67A</figureCitation>
) with microsclerites scattered over dorsal portions, sometimes forming a regular row; spermatheca tubular with accessory gland.
</paragraph>
<paragraph id="8B9DE117DA89E7CEFF23F8C7FDD423D7" blockId="129.[189,567,1917,1944]" box="[189,567,1917,1944]" pageId="129" pageNumber="130">
<heading id="D0D5567BDA89E7CEFF23F8C7FDD423D7" bold="true" box="[189,567,1917,1944]" fontSize="11" level="3" pageId="129" pageNumber="130" reason="3">
<emphasis id="B9563D05DA89E7CEFF23F8C7FDD423D7" bold="true" box="[189,567,1917,1944]" pageId="129" pageNumber="130">Secondary sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DA89E7CEFF23F811FC7123A8" blockId="129.[189,1398,1963,2025]" pageId="129" pageNumber="130">Labrum, lateral rims of prosternal process, and median part of metaventrite in males with groups of long setae. Terminal ventrite in females with small apical excision.</paragraph>
</subSubSection>
<subSubSection id="C338B29CDA8AE7CDFF23FEB2FE5A2513" box="[189,441,264,339]" pageId="130" pageNumber="131" type="distribution">
<paragraph id="8B9DE117DA8AE7CDFF23FEB2FEB52562" blockId="130.[189,342,264,290]" box="[189,342,264,290]" pageId="130" pageNumber="131">
<heading id="D0D5567BDA8AE7CDFF23FEB2FEB52562" bold="true" box="[189,342,264,290]" fontSize="11" level="3" pageId="130" pageNumber="131" reason="3">
<emphasis id="B9563D05DA8AE7CDFF23FEB2FEB52562" bold="true" box="[189,342,264,290]" pageId="130" pageNumber="131">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DA8AE7CDFF23FE83FE5A2513" blockId="130.[189,441,313,339]" box="[189,441,313,339]" pageId="130" pageNumber="131">
<collectingCountry id="F335A187DA8AE7CDFF23FE83FEC32513" box="[189,288,313,339]" name="Vietnam" pageId="130" pageNumber="131">Vietnam</collectingCountry>
(
<figureCitation id="1319FD92DA8AE7CDFEAEFE83FE4F2513" box="[304,428,313,339]" captionStart="Fig" captionStartId="220.[189,232,1871,1897]" captionTargetBox="[254,1334,266,1829]" captionTargetId="figure-44@220.[189,1399,265,1830]" captionTargetPageId="220" captionText="Fig. 112. Distribution maps. A. Elmomorphus bispinosus sp. nov. B. E. nepalensis Satô, 1981. C. E. punctulatus sp. nov. D. E. simplipes sp. nov. E. E. donatus Kodada, Selnekovič &amp; Jäch sp. nov. F. E. paradonatus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13773718" httpUri="https://zenodo.org/record/13773718/files/figure.png" pageId="130" pageNumber="131">Fig. 112D</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>