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<document id="576F2B88128B3D33367A693672701E08" ID-CLB-Dataset="45583" ID-DOI="10.11646/zootaxa.3811.4.4" ID-GBIF-Dataset="2b8dd933-6926-4671-9b84-c3f5a520fe51" ID-ISSN="1175-5326" ID-Zenodo-Dep="249520" ID-ZooBank="AE22B01B-B3FF-4B60-9452-A94DDDB20C2B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460432306852" checkinUser="plazi" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I. &amp; Panha, Somsak" docDate="2014" docId="F53487E6FFD2D80AFF7EFA21FAB35C9C" docLanguage="en" docName="zt03811p514.pdf" docOrigin="Zootaxa 3811 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Enghoffosoma bispinum Likhitrakarn, Golovatch &amp; Panha, 2014, sp. n." docType="treatment" docVersion="11" lastPageNumber="511" masterDocId="090DFF9EFFC3D81EFFE9FFD7FFFD593F" masterDocTitle="Review of the Southeast Asian millipede genus Enghoffosoma Golovatch, 1993 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of new species" masterLastPageNumber="514" masterPageNumber="491" pageNumber="508" updateTime="1698331558507" updateUser="ExternalLinkService">
<mods:mods id="DE4CF778EDE995B1C07EE983DCFAC9B6" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="FB9B05857DCE2A5A47C45F9665EAC1B3">Review of the Southeast Asian millipede genus Enghoffosoma Golovatch, 1993 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of new species</mods:title>
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<mods:name id="7D26DF510196D1CEA0D7E52CB39E0193" type="personal">
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<mods:namePart id="633121929862AA0B0E608E9425ABD003">Likhitrakarn, Natdanai</mods:namePart>
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<mods:namePart id="0A7EA799DFFF1AB48D31646CADB18B1F">Golovatch, Sergei I.</mods:namePart>
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<mods:role id="8E6D7449F2034A41DA40D9E136141863">
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<mods:namePart id="3AB0E593BD4C92A7267506B7A7B0395B">Panha, Somsak</mods:namePart>
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<mods:title id="2542DBD858EE6D5254DF782B04699B26">Zootaxa</mods:title>
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<mods:part id="208ADF59A94912DC6B5FB0938583BFD7">
<mods:date id="7666A1AFE16C3CDCB36DD299AE7B6A16">2014</mods:date>
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<mods:number id="7F92D9AF71AD371C26D92C53CA717306">3811</mods:number>
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<mods:identifier id="E50EB63F1465F4EE37FB9C2DADB068BC" type="DOI">10.11646/zootaxa.3811.4.4</mods:identifier>
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<treatment id="F53487E6FFD2D80AFF7EFA21FAB35C9C" ID-DOI="http://doi.org/10.5281/zenodo.6136574" ID-GBIF-Taxon="119430655" ID-Zenodo-Dep="6136574" LSID="urn:lsid:plazi:treatment:F53487E6FFD2D80AFF7EFA21FAB35C9C" httpUri="http://treatment.plazi.org/id/F53487E6FFD2D80AFF7EFA21FAB35C9C" lastPageId="20" lastPageNumber="511" pageId="17" pageNumber="508">
<subSubSection id="3587657BFFD2D80FFF7EFA21FEE95F0E" pageId="17" pageNumber="508" type="nomenclature">
<paragraph id="7D2236F0FFD2D80FFF7EFA21FDF65F2F" blockId="17.[151,523,1526,1586]" box="[151,523,1526,1552]" pageId="17" pageNumber="508">
<heading id="266A819CFFD2D80FFF7EFA21FDF65F2F" bold="true" box="[151,523,1526,1552]" fontSize="11" level="1" pageId="17" pageNumber="508" reason="1">
<emphasis id="4FE9EAE2FFD2D80FFF7EFA21FDF65F2F" bold="true" box="[151,523,1526,1552]" pageId="17" pageNumber="508">
<taxonomicName id="BA9D4D73FFD2D80FFF7EFA21FE3C5F2F" ID-CoL="39T49" box="[151,449,1526,1552]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="508" phylum="Arthropoda" rank="species" species="bispinum" status="sp. nov.">
<emphasis id="4FE9EAE2FFD2D80FFF7EFA21FE3C5F2F" bold="true" box="[151,449,1526,1552]" italics="true" pageId="17" pageNumber="508">Enghoffosoma bispinum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="54DA5799FFD2D80FFE21FA21FDF65F2F" box="[456,523,1526,1552]" pageId="17" pageNumber="508" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="7D2236F0FFD2D80FFF7EF9CEFEE95F0E" blockId="17.[151,523,1526,1586]" box="[151,276,1561,1586]" pageId="17" pageNumber="508">
<figureCitation id="E5A62A75FFD2D80FFF7EF9CEFEE95F0E" box="[151,276,1561,1586]" captionStart-0="FIGURE 17" captionStart-1="FIGURE 18" captionStart-2="FIGURE 19" captionStartId-0="19.[151,250,1902,1924]" captionStartId-1="20.[151,250,1038,1060]" captionStartId-2="21.[151,250,1499,1521]" captionTargetBox-0="[151,1436,553,1880]" captionTargetBox-1="[304,1283,199,1012]" captionTargetBox-2="[151,1435,195,1476]" captionTargetId-0="figure@19.[151,1436,553,1881]" captionTargetId-1="figure@20.[298,1289,193,1017]" captionTargetId-2="figure@21.[151,1436,195,1477]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="21" captionText-0="FIGURE 17. Enghoffosoma bispinum sp. n., male paratype (A), holotype (B – J). A: habitus, live coloration. B, C: anterior part of body, dorsal and lateral views, respectively. D, E: segments 10 and 11, dorsal and lateral views, respectively. F – H: posterior part of body, dorsal, ventral and lateral views, respectively. I, J: sternal cones between coxae 4, subcaudal and sublateral views, respectively." captionText-1="FIGURE 18. Enghoffosoma bispinum sp. n., holotype. A – C: right gonopod, lateral, mesal and caudal views, respectively. Scale bar: 0.5 mm. Designations explained in text." captionText-2="FIGURE 19. Enghoffosoma bispinum sp. n., holotype, right gonopod. A, B: mesal and lateral views, respectively. C – F: distal part, mesal, lateral, subcaudal and suboral views, respectively. Scale bar: 0.2 mm." httpUri-0="https://zenodo.org/record/249537/files/figure.png" httpUri-1="https://zenodo.org/record/249538/files/figure.png" httpUri-2="https://zenodo.org/record/249539/files/figure.png" pageId="17" pageNumber="508">Figs 17–19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="3587657BFFD2D80FFF7EF9B6FEF95E12" pageId="17" pageNumber="508" type="materials_examined">
<paragraph id="7D2236F0FFD2D80FFF7EF9B6FC325FA2" blockId="17.[151,1436,1633,1982]" pageId="17" pageNumber="508">
<typeStatus id="A2268852FFD2D80FFF7EF9B6FEF95F45" box="[151,260,1633,1658]" pageId="17" pageNumber="508" type="holotype">
<emphasis id="4FE9EAE2FFD2D80FFF7EF9B6FEF95F45" bold="true" box="[151,260,1633,1658]" pageId="17" pageNumber="508">Holotype</emphasis>
</typeStatus>
male (
<collectionCode id="1B8CAE35FFD2D80FFEB7F9B6FE485F46" box="[350,437,1633,1657]" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="17" pageNumber="508">CUMZ</collectionCode>
),
<collectingCountry id="058A7660FFD2D80FFE26F9B6FDC85F45" box="[463,565,1633,1658]" name="Thailand" pageId="17" pageNumber="508">Thailand</collectingCountry>
, Rayong Province, Mueang Rayong District, Kaosab temple,
<quantity id="BA659B15FFD2D80FFAC8F9B6FAAE5F46" box="[1313,1363,1633,1657]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="17" pageNumber="508" unit="m" value="9.0">9 m</quantity>
a.s.l.,
<geoCoordinate id="18A95037FFD2D80FFF7EF953FEE05FA2" box="[151,285,1668,1693]" direction="north" orientation="latitude" pageId="17" pageNumber="508" precision="15" value="12.613334">12°36'48&quot;N</geoCoordinate>
,
<geoCoordinate id="18A95037FFD2D80FFEC0F952FE475FA2" box="[297,442,1669,1693]" direction="east" orientation="longitude" pageId="17" pageNumber="508" precision="15" value="101.38972">101°23'23&quot;E</geoCoordinate>
,
<date id="09231030FFD2D80FFE2CF952FDB85FA2" box="[453,581,1669,1693]" pageId="17" pageNumber="508" value="2013-10-11">10.11.2013</date>
, leg. S. Panha and N. Likhitrakarn.
</paragraph>
<paragraph id="7D2236F0FFD2D80FFF2EF97EFD145FDA" blockId="17.[151,1436,1633,1982]" pageId="17" pageNumber="508">
<emphasis id="4FE9EAE2FFD2D80FFF2EF97EFEBB5FFD" bold="true" box="[199,326,1705,1730]" pageId="17" pageNumber="508">
<typeStatus id="A2268852FFD2D80FFF2EF97EFEBC5FFD" box="[199,321,1705,1730]" pageId="17" pageNumber="508" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="6B9BFD79FFD2D80FFEA6F97EFE565FFD" box="[335,427,1705,1730]" pageId="17" pageNumber="508" type="male">3 males</specimenCount>
,
<specimenCount id="6B9BFD79FFD2D80FFE50F97EFDD55FFD" box="[441,552,1705,1730]" pageId="17" pageNumber="508" type="female">3 females</specimenCount>
(
<collectionCode id="1B8CAE35FFD2D80FFDD3F97EFD6C5FFE" box="[570,657,1705,1729]" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="17" pageNumber="508">CUMZ</collectionCode>
),
<specimenCount id="6B9BFD79FFD2D80FFD4FF97EFD0A5FFD" box="[678,759,1705,1730]" pageId="17" pageNumber="508" type="male">1 male</specimenCount>
,
<specimenCount id="6B9BFD79FFD2D80FFCEFF97EFC945FFD" box="[774,873,1705,1730]" pageId="17" pageNumber="508" type="female">1 female</specimenCount>
(
<collectionCode id="1B8CAE35FFD2D80FFC92F97EFC2F5FFE" box="[891,978,1705,1729]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="17" pageNumber="508">ZMUC</collectionCode>
),
<specimenCount id="6B9BFD79FFD2D80FFC0EF97EFBC55FFD" box="[999,1080,1705,1730]" pageId="17" pageNumber="508" type="male">1 male</specimenCount>
,
<specimenCount id="6B9BFD79FFD2D80FFBAEF97EFB565FFD" box="[1095,1195,1705,1730]" pageId="17" pageNumber="508" type="female">1 female</specimenCount>
(
<collectionCode id="1B8CAE35FFD2D80FFB55F97EFAE45FFE" box="[1212,1305,1705,1729]" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="17" pageNumber="508">ZMUM</collectionCode>
),
<specimenCount id="6B9BFD79FFD2D80FFAC6F97EFA7D5FFD" box="[1327,1408,1705,1730]" pageId="17" pageNumber="508" type="male">1 male</specimenCount>
,
<specimenCount id="6B9BFD79FFD2D80FFA67F97EFF1E5FDA" pageId="17" pageNumber="508" type="female">1 female</specimenCount>
(
<collectionCode id="1B8CAE35FFD2D80FFF1BF91AFEA95FDA" box="[242,340,1741,1765]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="17" pageNumber="508">NHMW</collectionCode>
), same data, together with
<typeStatus id="A2268852FFD2D80FFD96F91BFD185FDA" box="[639,741,1740,1765]" pageId="17" pageNumber="508" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="7D2236F0FFD2D80FFF2EF926FEF95E12" blockId="17.[151,1436,1633,1982]" pageId="17" pageNumber="508">
<emphasis id="4FE9EAE2FFD2D80FFF2EF926FEE95E35" bold="true" box="[199,276,1777,1802]" pageId="17" pageNumber="508">Name.</emphasis>
To emphasize both gonopod process
<emphasis id="4FE9EAE2FFD2D80FFD0BF926FD135E35" bold="true" box="[738,750,1777,1802]" pageId="17" pageNumber="508">e</emphasis>
and the solenomere (
<emphasis id="4FE9EAE2FFD2D80FFC13F926FBF05E35" bold="true" box="[1018,1037,1777,1802]" pageId="17" pageNumber="508">sl</emphasis>
) being prominent and spiniform; adjective.
</paragraph>
</subSubSection>
<subSubSection id="3587657BFFD2D80FFF2EF8EEFD115E49" pageId="17" pageNumber="508" type="diagnosis">
<paragraph id="7D2236F0FFD2D80FFF2EF8EEFD115E49" blockId="17.[151,1436,1633,1982]" pageId="17" pageNumber="508">
<emphasis id="4FE9EAE2FFD2D80FFF2EF8EEFEBC5E6D" bold="true" box="[199,321,1849,1874]" pageId="17" pageNumber="508">Diagnosis.</emphasis>
Differs by mostly strongly developed paraterga, coupled with process
<emphasis id="4FE9EAE2FFD2D80FFB9EF8EEFB7E5E6D" bold="true" box="[1143,1155,1849,1874]" pageId="17" pageNumber="508">e</emphasis>
and the solenomere (
<emphasis id="4FE9EAE2FFD2D80FFA69F8EEFA6E5E6D" bold="true" box="[1408,1427,1849,1874]" pageId="17" pageNumber="508">sl</emphasis>
) being prominent and spiniform (see also Key below).
</paragraph>
</subSubSection>
<subSubSection id="3587657BFFD2D80AFF2EF856FC3D5C64" lastPageId="20" lastPageNumber="511" pageId="17" pageNumber="508" type="description">
<paragraph id="7D2236F0FFD2D80FFF2EF856FC405E81" blockId="17.[151,1436,1633,1982]" pageId="17" pageNumber="508">
<emphasis id="4FE9EAE2FFD2D80FFF2EF856FEA45EA5" bold="true" box="[199,345,1921,1946]" pageId="17" pageNumber="508">Description.</emphasis>
Length
<quantity id="BA659B15FFD2D80FFE5FF856FDD25EA6" box="[438,559,1921,1945]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="2.8" metricValueMin="2.2" pageId="17" pageNumber="508" unit="mm" value="25.0" valueMax="28.0" valueMin="22.0">22–28 mm</quantity>
(male) or
<quantity id="BA659B15FFD2D80FFD4CF856FCE35EA6" box="[677,798,1921,1946]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" metricValueMax="3.3" metricValueMin="2.6" pageId="17" pageNumber="508" unit="mm" value="29.5" valueMax="33.0" valueMin="26.0">26–33 mm</quantity>
(female), width of midbody pro- and metazonae 2.7–3.1 and
<quantity id="BA659B15FFD2D80FFF2FF872FEB05E82" box="[198,333,1957,1981]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.1" metricValueMin="3.4" pageId="17" pageNumber="508" unit="mm" value="3.75" valueMax="4.1" valueMin="3.4">3.4–4.1 mm</quantity>
(male), 3.2–4.0 and 4.0–
<quantity id="BA659B15FFD2D80FFD8EF872FD415E82" box="[615,700,1957,1981]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="17" pageNumber="508" unit="mm" value="4.8">4.8 mm</quantity>
(female), respectively.
</paragraph>
<caption id="29E26678FFD1D80CFF7EFA12FC635CC6" httpUri="https://zenodo.org/record/249536/files/figure.png" pageId="18" pageNumber="509" targetBox="[151,1435,176,1454]" targetPageId="18">
<paragraph id="7D2236F0FFD1D80CFF7EFA12FC635CC6" blockId="18.[151,1436,1476,1529]" pageId="18" pageNumber="509">
<emphasis id="4FE9EAE2FFD1D80CFF7EFA12FEE25CE5" bold="true" box="[151,287,1477,1499]" pageId="18" pageNumber="509">FIGURE 16.</emphasis>
<taxonomicName id="BA9D4D73FFD1D80CFECDFA13FE0B5CE5" box="[292,502,1476,1498]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="18" pageNumber="509" phylum="Arthropoda" rank="species" species="funda" status="sp. nov.">
<emphasis id="4FE9EAE2FFD1D80CFECDFA13FE0B5CE5" box="[292,502,1476,1498]" italics="true" pageId="18" pageNumber="509">Enghoffosoma funda</emphasis>
</taxonomicName>
<emphasis id="4FE9EAE2FFD1D80CFE15FA12FDC75CE5" bold="true" box="[508,570,1477,1498]" pageId="18" pageNumber="509">
<taxonomicNameLabel id="54DA5799FFD1D80CFE15FA12FDC95CE5" box="[508,564,1477,1498]" pageId="18" pageNumber="509" rank="species">sp. n.</taxonomicNameLabel>
,
</emphasis>
holotype, right gonopod. A, B: mesal and lateral views, respectively; C–F: distal part, mesal, lateral, subcaudal and suboral views, respectively. Scale bar: 0.2 mm.
</paragraph>
</caption>
<paragraph id="7D2236F0FFD1D80CFF2EF9F2FB7B5F95" blockId="18.[151,1436,1573,2030]" pageId="18" pageNumber="509">
Coloration of live animals blackish to brown with a pattern of contrasting light brown to whitish paraterga and epiproct (
<figureCitation id="E5A62A75FFD1D80CFEE8F99DFEAF5F5D" box="[257,338,1610,1635]" captionStart="FIGURE 17" captionStartId="19.[151,250,1902,1924]" captionTargetBox="[151,1436,553,1880]" captionTargetId="figure@19.[151,1436,553,1881]" captionTargetPageId="19" captionText="FIGURE 17. Enghoffosoma bispinum sp. n., male paratype (A), holotype (B – J). A: habitus, live coloration. B, C: anterior part of body, dorsal and lateral views, respectively. D, E: segments 10 and 11, dorsal and lateral views, respectively. F – H: posterior part of body, dorsal, ventral and lateral views, respectively. I, J: sternal cones between coxae 4, subcaudal and sublateral views, respectively." httpUri="https://zenodo.org/record/249537/files/figure.png" pageId="18" pageNumber="509">Fig. 17</figureCitation>
A); coloration in alcohol, after half a year of preservation, blackish to light brown with a pattern of contrasting light brown to whitish paraterga and epiproct (
<figureCitation id="E5A62A75FFD1D80CFCDBF9BAFC7A5FB9" box="[818,903,1645,1670]" captionStart="FIGURE 17" captionStartId="19.[151,250,1902,1924]" captionTargetBox="[151,1436,553,1880]" captionTargetId="figure@19.[151,1436,553,1881]" captionTargetPageId="19" captionText="FIGURE 17. Enghoffosoma bispinum sp. n., male paratype (A), holotype (B – J). A: habitus, live coloration. B, C: anterior part of body, dorsal and lateral views, respectively. D, E: segments 10 and 11, dorsal and lateral views, respectively. F – H: posterior part of body, dorsal, ventral and lateral views, respectively. I, J: sternal cones between coxae 4, subcaudal and sublateral views, respectively." httpUri="https://zenodo.org/record/249537/files/figure.png" pageId="18" pageNumber="509">Figs 17</figureCitation>
B–F &amp; H); antennae and legs blackish to light brown, but venter and a few basal podomeres light brown to whitish (
<figureCitation id="E5A62A75FFD1D80CFC48F945FC0A5F95" box="[929,1015,1682,1707]" captionStart="FIGURE 17" captionStartId="19.[151,250,1902,1924]" captionTargetBox="[151,1436,553,1880]" captionTargetId="figure@19.[151,1436,553,1881]" captionTargetPageId="19" captionText="FIGURE 17. Enghoffosoma bispinum sp. n., male paratype (A), holotype (B – J). A: habitus, live coloration. B, C: anterior part of body, dorsal and lateral views, respectively. D, E: segments 10 and 11, dorsal and lateral views, respectively. F – H: posterior part of body, dorsal, ventral and lateral views, respectively. I, J: sternal cones between coxae 4, subcaudal and sublateral views, respectively." httpUri="https://zenodo.org/record/249537/files/figure.png" pageId="18" pageNumber="509">Figs 17</figureCitation>
B, C &amp; F–J).
</paragraph>
<paragraph id="7D2236F0FFD1D80CFF2EF962FCDC5FF1" blockId="18.[151,1436,1573,2030]" box="[199,801,1717,1742]" pageId="18" pageNumber="509">
All characters as in
<taxonomicName id="BA9D4D73FFD1D80CFE4CF960FDFF5FF1" box="[421,514,1718,1742]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="18" pageNumber="509" phylum="Arthropoda" rank="species" species="zebra" status="sp. nov.">
<emphasis id="4FE9EAE2FFD1D80CFE4CF960FDFF5FF1" box="[421,514,1718,1742]" italics="true" pageId="18" pageNumber="509">E. zebra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="54DA5799FFD1D80CFDE0F962FDB45FF1" box="[521,585,1717,1742]" pageId="18" pageNumber="509" rank="species">
<emphasis id="4FE9EAE2FFD1D80CFDE0F962FDB45FF1" bold="true" box="[521,585,1717,1742]" pageId="18" pageNumber="509">sp. n.</emphasis>
</taxonomicNameLabel>
, except as follows.
</paragraph>
<paragraph id="7D2236F0FFD1D80CFF2EF90DFA8D5E61" blockId="18.[151,1436,1573,2030]" pageId="18" pageNumber="509">
Antennae long (
<figureCitation id="E5A62A75FFD1D80CFE68F90DFE285FCD" box="[385,469,1754,1779]" captionStart="FIGURE 17" captionStartId="19.[151,250,1902,1924]" captionTargetBox="[151,1436,553,1880]" captionTargetId="figure@19.[151,1436,553,1881]" captionTargetPageId="19" captionText="FIGURE 17. Enghoffosoma bispinum sp. n., male paratype (A), holotype (B – J). A: habitus, live coloration. B, C: anterior part of body, dorsal and lateral views, respectively. D, E: segments 10 and 11, dorsal and lateral views, respectively. F – H: posterior part of body, dorsal, ventral and lateral views, respectively. I, J: sternal cones between coxae 4, subcaudal and sublateral views, respectively." httpUri="https://zenodo.org/record/249537/files/figure.png" pageId="18" pageNumber="509">Fig. 17</figureCitation>
A), clavate, extending behind body segment 4 (male) or 3 (female) when stretched dorsally. In width, head &lt;segment 3 = 4 &lt;collum &lt;segment 2 &lt;5–14 (both sexes), gently and gradually tapering thereafter. Collum with three transverse rows of setae: 4+
<quantity id="BA659B15FFD1D80CFCC0F8F5FCAB5E04" box="[809,854,1826,1851]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="18" pageNumber="509" unit="in" value="4.0">4 in</quantity>
anterior, 3+
<quantity id="BA659B15FFD1D80CFC0AF8F5FBF25E04" box="[995,1039,1826,1851]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="18" pageNumber="509" unit="in" value="3.0">3 in</quantity>
intermediate, and 5+
<quantity id="BA659B15FFD1D80CFAEAF8F5FACD5E04" box="[1283,1328,1826,1851]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="18" pageNumber="509" unit="in" value="5.0">5 in</quantity>
posterior row; caudal corner of paraterga very broadly rounded, declined, extending behind rear margin (
<figureCitation id="E5A62A75FFD1D80CFB28F892FAEB5E61" box="[1217,1302,1861,1886]" captionStart="FIGURE 17" captionStartId="19.[151,250,1902,1924]" captionTargetBox="[151,1436,553,1880]" captionTargetId="figure@19.[151,1436,553,1881]" captionTargetPageId="19" captionText="FIGURE 17. Enghoffosoma bispinum sp. n., male paratype (A), holotype (B – J). A: habitus, live coloration. B, C: anterior part of body, dorsal and lateral views, respectively. D, E: segments 10 and 11, dorsal and lateral views, respectively. F – H: posterior part of body, dorsal, ventral and lateral views, respectively. I, J: sternal cones between coxae 4, subcaudal and sublateral views, respectively." httpUri="https://zenodo.org/record/249537/files/figure.png" pageId="18" pageNumber="509">Figs 17</figureCitation>
B &amp; C).
</paragraph>
<paragraph id="7D2236F0FFD1D80DFF2EF8BCFE9358CB" blockId="18.[151,1436,1573,2030]" lastBlockId="19.[151,1436,151,501]" lastPageId="19" lastPageNumber="510" pageId="18" pageNumber="509">
Tegument smooth and leathery, prozonae finely shagreened, metaterga smooth and leathery, delicately rugulose; surface below paraterga finely microgranulate (
<figureCitation id="E5A62A75FFD1D80CFCD6F85AFC655E99" box="[831,920,1933,1958]" captionStart="FIGURE 17" captionStartId="19.[151,250,1902,1924]" captionTargetBox="[151,1436,553,1880]" captionTargetId="figure@19.[151,1436,553,1881]" captionTargetPageId="19" captionText="FIGURE 17. Enghoffosoma bispinum sp. n., male paratype (A), holotype (B – J). A: habitus, live coloration. B, C: anterior part of body, dorsal and lateral views, respectively. D, E: segments 10 and 11, dorsal and lateral views, respectively. F – H: posterior part of body, dorsal, ventral and lateral views, respectively. I, J: sternal cones between coxae 4, subcaudal and sublateral views, respectively." httpUri="https://zenodo.org/record/249537/files/figure.png" pageId="18" pageNumber="509">Figs 17</figureCitation>
B–F &amp; H). Postcollum metaterga with two transverse rows of setae traceable at least as insertion points when setae broken off: 2+
<quantity id="BA659B15FFD1D80CFB92F865FB5A5EF4" box="[1147,1191,1970,1995]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="18" pageNumber="509" unit="in" value="2.0">2 in</quantity>
anterior (pre-sulcus), 3+
<quantity id="BA659B15FFD1D80CFF5CF801FF225ED1" box="[181,223,2005,2030]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="18" pageNumber="509" unit="in" value="3.0">3 in</quantity>
posterior (post-sulcus) row, anterior row usually with retained setae, caudal row barely traceable as insertion points. Tergal setae long, strong, slender, about 1/3 as long as metaterga. Axial line visible on both pro- and metazonae. Paraterga very strongly developed (
<figureCitation id="E5A62A75FFD0D80DFD47FF6BFCFF59EB" box="[686,770,188,213]" captionStart="FIGURE 17" captionStartId="19.[151,250,1902,1924]" captionTargetBox="[151,1436,553,1880]" captionTargetId="figure@19.[151,1436,553,1881]" captionTargetPageId="19" captionText="FIGURE 17. Enghoffosoma bispinum sp. n., male paratype (A), holotype (B – J). A: habitus, live coloration. B, C: anterior part of body, dorsal and lateral views, respectively. D, E: segments 10 and 11, dorsal and lateral views, respectively. F – H: posterior part of body, dorsal, ventral and lateral views, respectively. I, J: sternal cones between coxae 4, subcaudal and sublateral views, respectively." httpUri="https://zenodo.org/record/249537/files/figure.png" pageId="19" pageNumber="510">Figs 17</figureCitation>
B–F &amp; H), especially so in male, mostly slightly upturned, all lying below dorsum, set at about upper 1/3 of body height, subhorizontal, caudal corner almost or fully pointed, largely produced beyond rear tergal margin until segment 15, increasingly rounded and smaller on following segments; posterior edge concave (
<figureCitation id="E5A62A75FFD0D80DFDCAFEF0FD84587F" box="[547,633,295,320]" captionStart="FIGURE 17" captionStartId="19.[151,250,1902,1924]" captionTargetBox="[151,1436,553,1880]" captionTargetId="figure@19.[151,1436,553,1881]" captionTargetPageId="19" captionText="FIGURE 17. Enghoffosoma bispinum sp. n., male paratype (A), holotype (B – J). A: habitus, live coloration. B, C: anterior part of body, dorsal and lateral views, respectively. D, E: segments 10 and 11, dorsal and lateral views, respectively. F – H: posterior part of body, dorsal, ventral and lateral views, respectively. I, J: sternal cones between coxae 4, subcaudal and sublateral views, respectively." httpUri="https://zenodo.org/record/249537/files/figure.png" pageId="19" pageNumber="510">Figs 17</figureCitation>
B–F). Paraterga 2 broad, anterior edge angular to rounded, lateral edge without incision (
<figureCitation id="E5A62A75FFD0D80DFEB3FE9BFE52585B" box="[346,431,332,357]" captionStart="FIGURE 17" captionStartId="19.[151,250,1902,1924]" captionTargetBox="[151,1436,553,1880]" captionTargetId="figure@19.[151,1436,553,1881]" captionTargetPageId="19" captionText="FIGURE 17. Enghoffosoma bispinum sp. n., male paratype (A), holotype (B – J). A: habitus, live coloration. B, C: anterior part of body, dorsal and lateral views, respectively. D, E: segments 10 and 11, dorsal and lateral views, respectively. F – H: posterior part of body, dorsal, ventral and lateral views, respectively. I, J: sternal cones between coxae 4, subcaudal and sublateral views, respectively." httpUri="https://zenodo.org/record/249537/files/figure.png" pageId="19" pageNumber="510">Figs 17</figureCitation>
B &amp; C). Transverse sulcus at most very faint and abbreviated (
<figureCitation id="E5A62A75FFD0D80DFB80FE9BFB43585B" box="[1129,1214,332,357]" captionStart="FIGURE 17" captionStartId="19.[151,250,1902,1924]" captionTargetBox="[151,1436,553,1880]" captionTargetId="figure@19.[151,1436,553,1881]" captionTargetPageId="19" captionText="FIGURE 17. Enghoffosoma bispinum sp. n., male paratype (A), holotype (B – J). A: habitus, live coloration. B, C: anterior part of body, dorsal and lateral views, respectively. D, E: segments 10 and 11, dorsal and lateral views, respectively. F – H: posterior part of body, dorsal, ventral and lateral views, respectively. I, J: sternal cones between coxae 4, subcaudal and sublateral views, respectively." httpUri="https://zenodo.org/record/249537/files/figure.png" pageId="19" pageNumber="510">Figs 17</figureCitation>
B, D &amp; F). Stricture between pro- and metazonae narrow, deep, line-shaped, clearly beaded at bottom down to base of paraterga (
<figureCitation id="E5A62A75FFD0D80DFA83FEB8FF4E5893" captionStart="FIGURE 17" captionStartId="19.[151,250,1902,1924]" captionTargetBox="[151,1436,553,1880]" captionTargetId="figure@19.[151,1436,553,1881]" captionTargetPageId="19" captionText="FIGURE 17. Enghoffosoma bispinum sp. n., male paratype (A), holotype (B – J). A: habitus, live coloration. B, C: anterior part of body, dorsal and lateral views, respectively. D, E: segments 10 and 11, dorsal and lateral views, respectively. F – H: posterior part of body, dorsal, ventral and lateral views, respectively. I, J: sternal cones between coxae 4, subcaudal and sublateral views, respectively." httpUri="https://zenodo.org/record/249537/files/figure.png" pageId="19" pageNumber="510">Figs 17</figureCitation>
B–F). Pleurosternal carinae strongly developed, complete crests with a very sharp caudal tooth in segments 2–13 (male) or 2–12 (female), a small sharp caudal tooth in segments 14–17 (male) or 13–16 (female), absent thereafter (
<figureCitation id="E5A62A75FFD0D80DFF76FE0BFF0E58CB" box="[159,243,476,501]" captionStart="FIGURE 17" captionStartId="19.[151,250,1902,1924]" captionTargetBox="[151,1436,553,1880]" captionTargetId="figure@19.[151,1436,553,1881]" captionTargetPageId="19" captionText="FIGURE 17. Enghoffosoma bispinum sp. n., male paratype (A), holotype (B – J). A: habitus, live coloration. B, C: anterior part of body, dorsal and lateral views, respectively. D, E: segments 10 and 11, dorsal and lateral views, respectively. F – H: posterior part of body, dorsal, ventral and lateral views, respectively. I, J: sternal cones between coxae 4, subcaudal and sublateral views, respectively." httpUri="https://zenodo.org/record/249537/files/figure.png" pageId="19" pageNumber="510">Figs 17</figureCitation>
C, E &amp; H).
</paragraph>
<caption id="29E26678FFD0D80DFF7EF8B9FEEA5EDF" httpUri="https://zenodo.org/record/249537/files/figure.png" pageId="19" pageNumber="510" targetBox="[151,1436,553,1880]" targetPageId="19">
<paragraph id="7D2236F0FFD0D80DFF7EF8B9FEEA5EDF" blockId="19.[151,1436,1902,2016]" pageId="19" pageNumber="510">
<emphasis id="4FE9EAE2FFD0D80DFF7EF8B9FEE25EBC" bold="true" box="[151,287,1902,1924]" pageId="19" pageNumber="510">FIGURE 17.</emphasis>
<taxonomicName id="BA9D4D73FFD0D80DFECCF8B9FDE65EBB" box="[293,539,1902,1924]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="19" pageNumber="510" phylum="Arthropoda" rank="species" species="bispinum" status="sp. nov.">
<emphasis id="4FE9EAE2FFD0D80DFECCF8B9FDE65EBB" box="[293,539,1902,1924]" italics="true" pageId="19" pageNumber="510">Enghoffosoma bispinum</emphasis>
</taxonomicName>
<emphasis id="4FE9EAE2FFD0D80DFDC8F8B9FD9C5EBC" bold="true" box="[545,609,1902,1923]" pageId="19" pageNumber="510">
<taxonomicNameLabel id="54DA5799FFD0D80DFDC8F8B9FDA75EBC" box="[545,602,1902,1923]" pageId="19" pageNumber="510" rank="species">sp. n.</taxonomicNameLabel>
,
</emphasis>
male paratype (A), holotype (B–J). A: habitus, live coloration. B, C: anterior part of body, dorsal and lateral views, respectively. D, E: segments 10 and 11, dorsal and lateral views, respectively. F–H: posterior part of body, dorsal, ventral and lateral views, respectively. I, J: sternal cones between coxae 4, subcaudal and sublateral views, respectively.
</paragraph>
</caption>
<caption id="29E26678FFD7D80AFF7EFBD9FD6B5D7D" httpUri="https://zenodo.org/record/249538/files/figure.png" pageId="20" pageNumber="511" targetBox="[304,1283,199,1012]" targetPageId="20">
<paragraph id="7D2236F0FFD7D80AFF7EFBD9FD6B5D7D" blockId="20.[151,1437,1038,1090]" pageId="20" pageNumber="511">
<emphasis id="4FE9EAE2FFD7D80AFF7EFBD9FEDF5D1C" bold="true" box="[151,290,1038,1060]" pageId="20" pageNumber="511">FIGURE 18.</emphasis>
<taxonomicName id="BA9D4D73FFD7D80AFEC5FBD9FDD85D1B" box="[300,549,1038,1060]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="20" pageNumber="511" phylum="Arthropoda" rank="species" species="bispinum" status="sp. nov.">
<emphasis id="4FE9EAE2FFD7D80AFEC5FBD9FDD85D1B" box="[300,549,1038,1060]" italics="true" pageId="20" pageNumber="511">Enghoffosoma bispinum</emphasis>
</taxonomicName>
<emphasis id="4FE9EAE2FFD7D80AFDC7FBD8FD8C5D1B" bold="true" box="[558,625,1039,1060]" pageId="20" pageNumber="511">
<taxonomicNameLabel id="54DA5799FFD7D80AFDC7FBD8FD965D1B" box="[558,619,1039,1060]" pageId="20" pageNumber="511" rank="species">sp. n.</taxonomicNameLabel>
,
</emphasis>
holotype. A–C: right gonopod, lateral, mesal and caudal views, respectively. Scale bar: 0.5 mm. Designations explained in text.
</paragraph>
</caption>
<paragraph id="7D2236F0FFD7D80AFF2EFBBCFB535DD0" blockId="20.[151,1436,1130,1443]" pageId="20" pageNumber="511">
Sterna very densely setose, with an evident cone caudally near each coxa, rear cones being a little stronger than front ones; a rather large, linguiform, densely setose sternal lobe between male coxae 4 (
<figureCitation id="E5A62A75FFD7D80AFB58FB59FAF45D98" box="[1201,1289,1166,1191]" captionStart="FIGURE 17" captionStartId="19.[151,250,1902,1924]" captionTargetBox="[151,1436,553,1880]" captionTargetId="figure@19.[151,1436,553,1881]" captionTargetPageId="19" captionText="FIGURE 17. Enghoffosoma bispinum sp. n., male paratype (A), holotype (B – J). A: habitus, live coloration. B, C: anterior part of body, dorsal and lateral views, respectively. D, E: segments 10 and 11, dorsal and lateral views, respectively. F – H: posterior part of body, dorsal, ventral and lateral views, respectively. I, J: sternal cones between coxae 4, subcaudal and sublateral views, respectively." httpUri="https://zenodo.org/record/249537/files/figure.png" pageId="20" pageNumber="511">
Figs
<date id="09231030FFD7D80AFB07FB58FAF45D98" box="[1262,1289,1167,1191]" pageId="20" pageNumber="511" value="1917-01">17</date>
</figureCitation>
I &amp; J). Legs moderately long and slender, midbody ones ca 1.1–1.2 (male) or 0.9–1.0 times (female) as long as body height; ventral brushes traceable on male tibiae and tarsi until legs of segment 8 and 10, respectively.
</paragraph>
<paragraph id="7D2236F0FFD7D80AFF2EFB2DFC3D5C64" blockId="20.[151,1436,1130,1443]" pageId="20" pageNumber="511">
Gonopods (
<figureCitation id="E5A62A75FFD7D80AFEA5FB2DFE5E5C2C" box="[332,419,1274,1299]" captionStart="FIGURE 18" captionStartId="20.[151,250,1038,1060]" captionTargetBox="[304,1283,199,1012]" captionTargetId="figure@20.[298,1289,193,1017]" captionTargetPageId="20" captionText="FIGURE 18. Enghoffosoma bispinum sp. n., holotype. A – C: right gonopod, lateral, mesal and caudal views, respectively. Scale bar: 0.5 mm. Designations explained in text." httpUri="https://zenodo.org/record/249538/files/figure.png" pageId="20" pageNumber="511">Figs 18</figureCitation>
&amp;
<figureCitation id="E5A62A75FFD7D80AFE25FB2CFE145C2C" box="[460,489,1275,1299]" captionStart="FIGURE 19" captionStartId="21.[151,250,1499,1521]" captionTargetBox="[151,1435,195,1476]" captionTargetId="figure@21.[151,1436,195,1477]" captionTargetPageId="21" captionText="FIGURE 19. Enghoffosoma bispinum sp. n., holotype, right gonopod. A, B: mesal and lateral views, respectively. C – F: distal part, mesal, lateral, subcaudal and suboral views, respectively. Scale bar: 0.2 mm." httpUri="https://zenodo.org/record/249539/files/figure.png" pageId="20" pageNumber="511">19</figureCitation>
) simple. Femorite slender and long, slightly curved, with seminal groove running mesally, but turning laterad already in distal half of femorite; apicoventral shelf (
<emphasis id="4FE9EAE2FFD7D80AFBD6FAC9FBB35C08" bold="true" box="[1087,1102,1310,1335]" pageId="20" pageNumber="511">b</emphasis>
) evident, both process
<emphasis id="4FE9EAE2FFD7D80AFAB4FAC9FA945C08" bold="true" box="[1373,1385,1310,1335]" pageId="20" pageNumber="511">e</emphasis>
and solenomere (
<emphasis id="4FE9EAE2FFD7D80AFEC1FA95FEC65C64" bold="true" box="[296,315,1346,1371]" pageId="20" pageNumber="511">sl</emphasis>
) large, spiniform, slightly curved, but
<emphasis id="4FE9EAE2FFD7D80AFD05FA95FD025C64" bold="true" box="[748,767,1346,1371]" pageId="20" pageNumber="511">sl</emphasis>
longer and bifid.
</paragraph>
</subSubSection>
<subSubSection id="3587657BFFD7D80AFF2EFAB1FAB35C9C" pageId="20" pageNumber="511" type="discussion">
<paragraph id="7D2236F0FFD7D80AFF2EFAB1FAB35C9C" blockId="20.[151,1436,1130,1443]" pageId="20" pageNumber="511">
<emphasis id="4FE9EAE2FFD7D80AFF2EFAB1FEC65C40" bold="true" box="[199,315,1382,1407]" pageId="20" pageNumber="511">Remarks.</emphasis>
The specimens were collected hidden under leaf litter, in a rotten log and even under roof tiles near a monk’s house in a temple. Some of the animals were still colourless and soft-bodied shortly after molting.
</paragraph>
</subSubSection>
</treatment>
</document>
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