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<document ID-CLB-Dataset="264788" ID-DOI="http://dx.doi.org/10.5091/plecevo.98847" ID-GBIF-Dataset="2aaa4e51-ec7b-4d32-ba88-5d690b8f2a01" ID-Pensoft-Pub="2032-3921-3-276" ID-Pensoft-UUID="BC5E6BD70022518ABF185FC3C18147CE" ModsDocID="2032-3921-156-3-276" checkinTime="1693470036859" checkinUser="pensoft" docAuthor="Rasaminirina, Fitiavana &amp; Larridon, Isabel" docDate="2023" docId="F5341F7C7CEE5ACBB49F46132881D09E" docLanguage="en" docName="PlantEcolEvol 156(3): 276-310" docOrigin="Plant Ecology and Evolution 156 (3)" docPubDate="2023-08-29" docSource="http://dx.doi.org/10.5091/plecevo.98847" docTitle="Machaerina Vahl (Vahl 1805" docType="treatment" docVersion="4" id="BC5E6BD70022518ABF185FC3C18147CE" lastPageNumber="276" masterDocId="BC5E6BD70022518ABF185FC3C18147CE" masterDocTitle="The genera of Cyperaceae of Madagascar" masterLastPageNumber="310" masterPageNumber="276" pageNumber="276" updateTime="1720463581714" updateUser="admin">
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<mods:titleInfo id="19A34AD29A2123577FEA45039B56B816">
<mods:title id="4D52C31A75A584BCF1F45207F8625B9C">The genera of Cyperaceae of Madagascar</mods:title>
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<mods:namePart id="EC02DE934F0A193AFB667EE15491FFB5">Rasaminirina, Fitiavana</mods:namePart>
<mods:nameIdentifier id="41FD93BA4F05CDC41D88CBD099DE5CC7" type="ORCID">https://orcid.org/0000-0003-0162-7975</mods:nameIdentifier>
<mods:affiliation id="D84F87B524E425390DFD19E6C04DA302">University of Antananarivo, Antananarivo, Madagascar</mods:affiliation>
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<mods:namePart id="844260C8622CEE296591EA38F32F580E">Larridon, Isabel</mods:namePart>
<mods:nameIdentifier id="C57E8E6DFF4664600DF8BEE80B289CB8" type="ORCID">https://orcid.org/0000-0003-0285-722X</mods:nameIdentifier>
<mods:affiliation id="F4744F2ADDCD346B3FC5FDB455B847A3">Kew Madagascar Conservation Centre, Antananarivo, Madagascar &amp; Royal Botanic Gardens, Kew, Richmond, UK</mods:affiliation>
<mods:nameIdentifier id="82186286B428DFB4F66BD1264E85E04D" type="email">i.larridon@kew.org</mods:nameIdentifier>
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<mods:title id="58B979D62CE7E3FB9EC868A0E1A1ABD3">Plant Ecology and Evolution</mods:title>
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<mods:date id="D21B6DB89A9EB3D0DD0299C9B4769110">2023</mods:date>
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<mods:number id="337F1C587C7122C435903FC1E6C9AA87">156</mods:number>
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<mods:identifier id="CCAA494F3CA1AEE160C36B6C262773D9" type="DOI">http://dx.doi.org/10.5091/plecevo.98847</mods:identifier>
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<treatment id="F5341F7C7CEE5ACBB49F46132881D09E" ID-GBIF-Taxon="212738608" LSID="urn:lsid:plazi:treatment:F5341F7C7CEE5ACBB49F46132881D09E" httpUri="http://treatment.plazi.org/id/F5341F7C7CEE5ACBB49F46132881D09E" lastPageNumber="276" pageId="0" pageNumber="276">
<subSubSection id="D3CEC1A8E0C6E33463A8C8E168FA2059" pageId="0" pageNumber="276" type="nomenclature">
<paragraph id="1F72BC5D1E6DCA105FED5A82C3676A21" pageId="0" pageNumber="276">
<taxonomicName id="3C239DDE19C4D79A6895A6BB0514D43C" ID-CoL="8VZ3W" LSID="F5341F7C-7CEE-5ACB-B49F-46132881D09E" authority="Vahl (Vahl 1805: 238)" authorityName="Vahl (Vahl" authorityYear="1805" class="Liliopsida" family="Cyperaceae" genus="Machaerina" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Machaerina" order="Poales" pageId="0" pageNumber="276" phylum="Tracheophyta" rank="genus">Machaerina Vahl (Vahl 1805: 238)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="FD4AF825A5CEE96AEB08578339A8649B" pageId="0" pageNumber="276" type="type species">
<paragraph id="37FFC0A57368D3D5002E66F49E52E487" pageId="0" pageNumber="276">Type species.</paragraph>
<paragraph id="302013A1D56E7E0CC5F5FE3631983EE8" pageId="0" pageNumber="276">
<taxonomicName id="297BCA26B41370657D8CF462E9D17ECF" class="Liliopsida" family="Cyperaceae" genus="Schoenus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Schoenus restioides" order="Poales" pageId="0" pageNumber="276" phylum="Magnoliophyta" rank="species" species="restioides">
<emphasis id="EA40EF7C832F33D2BD3B3FB1F462135F" italics="true" pageId="0" pageNumber="276">Schoenus restioides</emphasis>
</taxonomicName>
Sw. [=
<taxonomicName id="149DFB6A22ADC659E6A70CCAF0B6B5DA" class="Liliopsida" family="Cyperaceae" genus="Machaerina" kingdom="Plantae" lsidName="Machaerina restioides" order="Poales" pageId="0" pageNumber="276" phylum="Tracheophyta" rank="species" species="restioides">
<emphasis id="6FA8AEAFA2CF6511A388B58058B6DA36" italics="true" pageId="0" pageNumber="276">Machaerina restioides</emphasis>
</taxonomicName>
Vahl]
</paragraph>
</subSubSection>
<subSubSection id="3EE525116010186EAEF2289662C8C28E" pageId="0" pageNumber="276" type="description">
<paragraph id="08195487A20E16009C3DEAA556683CA6" pageId="0" pageNumber="276">Description of the genus.</paragraph>
<paragraph id="773C98A0173E58DADCDCACD0AB25498C" pageId="0" pageNumber="276">Medium-sized to tall creeping rhizomatous or stoloniferous perennials. Culms scapose or with few nodes, flattened, angular, compressed or (sub)terete, sometimes septate. Leaves distichous, eligulate, blade ensiform to (sub)terete, sometimes septate, rarely reduced to a sheath. Primary bracts often leaf-like, sheathing. Inflorescence sometimes more or less pseudolateral, paniculate, main axis often sinuous, partial inflorescences sometimes capitately contracted. Spikelets many. Glumes 2-10 distichous, long-persistent, of increasing length, with (1-)2-4(-5) florets subtended by the larger glumes and enclosed by the wings of the next glume. Lower 1-2 floret(s) usually bisexual, upper floret(s) usually functionally male. Bristles absent or poorly developed, or up to 6, delicate, shorter than, to as long as the fruit, deciduous with the fruit. Stamens 3; filaments sometimes conspicuously elongated after anthesis. Style 3-fid, base not distinct, thickened, persistent, sometimes scabrid. Nutlets ovoid to oblong, terete to triquetrous, more or less 3-ribbed to winged, stipitate or sessile, beaked, smooth to rugose.</paragraph>
</subSubSection>
<subSubSection id="47026C38E3C5EDF7D3BA58C5A208B14E" pageId="0" pageNumber="276" type="distribution">
<paragraph id="71672E7088E69A7518E822265FE5FF4D" pageId="0" pageNumber="276">Distribution and ecology.</paragraph>
<paragraph id="E34F7DAF4D50E8D7EFA1D54FF2802B6F" pageId="0" pageNumber="276">
<taxonomicName id="6AF3B0D04DD9385679BCFE1319A7CE89" authorityName="Vahl (Vahl" authorityYear="1805" class="Liliopsida" family="Cyperaceae" genus="Machaerina" kingdom="Plantae" lsidName="Machaerina" order="Poales" pageId="0" pageNumber="276" phylum="Tracheophyta" rank="genus">
<emphasis id="9EBC8E1CD5BAD71F3CE74BAB1C0A75A3" italics="true" pageId="0" pageNumber="276">Machaerina</emphasis>
</taxonomicName>
occurs in Tanzania, Western Indian Ocean to Pacific, Tropical America (
<bibRefCitation id="6DA74FC75ADF1B80274C21C83D6B1843" author="Persoon, CH" journalOrPublisher="3 e Aufl. O. R. Reisland, Leipzig" pageId="0" pageNumber="276" publicationUrl="http://www.plantsoftheworldonline.org/" refId="B39" refString="2022. . http://www.plantsoftheworldonline.org/" url="http://www.plantsoftheworldonline.org/" year="2022">POWO 2022</bibRefCitation>
). It grows in forest, shrubland, or grassland, along streams, in marshy areas or on rocks.
<taxonomicName id="50A4EAC5EA54A6397F5A1EA35653C4E6" authorityName="Vahl (Vahl" authorityYear="1805" class="Liliopsida" family="Cyperaceae" genus="Machaerina" kingdom="Plantae" lsidName="Machaerina" order="Poales" pageId="0" pageNumber="276" phylum="Tracheophyta" rank="genus">
<emphasis id="C635E3FAC448EC35FD254F41D6C2B2FD" italics="true" pageId="0" pageNumber="276">Machaerina</emphasis>
</taxonomicName>
occurs in northern, northeastern, and southeastern Madagascar; four species are known including e.g.
<taxonomicName id="4BFF81CEC609571DACFE739AE90F148F" class="Liliopsida" family="Cyperaceae" genus="Machaerina" kingdom="Plantae" lsidName="Machaerina flexuosa" order="Poales" pageId="0" pageNumber="276" phylum="Tracheophyta" rank="species" species="flexuosa">
<emphasis id="BDAE16736A27AB23FD9D43E68EB21888" italics="true" pageId="0" pageNumber="276">Machaerina flexuosa</emphasis>
</taxonomicName>
(Boeckeler) J.Kern (Fig.
<figureCitation id="35314329EE39DBD4048B60854845AB7A" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Machaerina flexuosa. A. Habit (x 2 / 3). B. Spikes from top of inflorescence (x 3). C - D. Glume, abaxial and side view (x 10). E. Flower (x 10). F. Young flower (x 8). G. Anther (x 16). H. Nutlet (x 16). A from Andriamahay &amp; Rakotoarison 1969; B-G from Nusbaumer et al. 1103; H from Ranirison 642. Drawn by Juliet Beentje, reproduced with permission from the artist, originally published in Hoenselaar et al. (2010)." figureDoi="10.5091/plecevo.98847.figure10" httpUri="https://binary.pensoft.net/fig/898161" pageId="0" pageNumber="276">10</figureCitation>
).
</paragraph>
<caption id="D3DCE0F65ABDD5111A53B8161DCBC13D" doi="10.5091/plecevo.98847.figure10" httpUri="https://binary.pensoft.net/fig/898161" pageId="0" pageNumber="276" start="Figure 10" startId="F10">
<paragraph id="5301A27063F9FC3FE32D6D7C1DF18D2C" pageId="0" pageNumber="276">
<emphasis id="001485CBC70283B653EE1B99DE42FF88" bold="true" pageId="0" pageNumber="276">Figure 10.</emphasis>
<taxonomicName id="1D0928465FD8AF12A3BAF1DA9538714C" class="Liliopsida" family="Cyperaceae" genus="Machaerina" kingdom="Plantae" lsidName="Machaerina flexuosa" order="Poales" pageId="0" pageNumber="276" phylum="Tracheophyta" rank="species" species="flexuosa">
<emphasis id="92561149357837F06E67E0BD6EC30DAD" italics="true" pageId="0" pageNumber="276">Machaerina flexuosa</emphasis>
</taxonomicName>
.
<emphasis id="95B69B268E42E880BB9086112EC57594" bold="true" pageId="0" pageNumber="276">A</emphasis>
. Habit (
<normalizedToken id="AB20D48E3D52006BA364B352633FB643" originalValue="×">x</normalizedToken>
2/3).
<emphasis id="9A28829C041474B84AF55E47E80501C7" bold="true" pageId="0" pageNumber="276">B</emphasis>
. Spikes from top of inflorescence (
<normalizedToken id="DB602852AF2A19F45F562F3526F70F9B" originalValue="×">x</normalizedToken>
3).
<emphasis id="6C25C0EB8E17EE8F7851826EA54BD883" bold="true" pageId="0" pageNumber="276">C</emphasis>
-
<emphasis id="9C56D73A2793350D08F48CF3BF23D08C" bold="true" pageId="0" pageNumber="276">D</emphasis>
. Glume, abaxial and side view (
<normalizedToken id="C73F3502FB0A9FBFBF6E5363EDBA0E8C" originalValue="×">x</normalizedToken>
10).
<emphasis id="BD77E01F089AE69FCD307FA1CF2CB8FB" bold="true" pageId="0" pageNumber="276">E</emphasis>
. Flower (
<normalizedToken id="B4DBF8062928863AC133D3E40E77DC18" originalValue="×">x</normalizedToken>
10).
<emphasis id="AD07A3F4B0B1BFA57D6D826A5A5A7F9E" bold="true" pageId="0" pageNumber="276">F</emphasis>
. Young flower (
<normalizedToken id="5E6A3A90546EBA4D74EAEB4E6CBD8C09" originalValue="×">x</normalizedToken>
8).
<emphasis id="F15428A06EB32EAB213605F44220599E" bold="true" pageId="0" pageNumber="276">G</emphasis>
. Anther (
<normalizedToken id="ABF424D1891398897E9B35611C2BD17F" originalValue="×">x</normalizedToken>
16).
<emphasis id="9B82306EC70E39267E8E24890640957B" bold="true" pageId="0" pageNumber="276">H</emphasis>
. Nutlet (
<normalizedToken id="0A15CC11692EAD2A3967C0E7A51C8A11" originalValue="×">x</normalizedToken>
16). A from
<emphasis id="95653BAE54D385D8DCF7005E9063DB92" italics="true" pageId="0" pageNumber="276">Andriamahay &amp; Rakotoarison 1969</emphasis>
; B-G from
<emphasis id="A44889572537243D9CED0A6C99A148B9" italics="true" pageId="0" pageNumber="276">Nusbaumer et al. 1103</emphasis>
; H from
<emphasis id="B2FC42CF187A72A6C1D29143B9A6FCB1" italics="true" pageId="0" pageNumber="276">Ranirison 642</emphasis>
. Drawn by Juliet Beentje, reproduced with permission from the artist, originally published in
<bibRefCitation id="26C77794864C56DF9178B38FD6266C6B" author="Hoenselaar, K" editor="Beentje, HJ" journalOrPublisher="Royal Botanic Gardens, Kew, Richmond" pageId="0" pageNumber="276" pagination="1 - 466" refId="B20" refString="Hoenselaar, K, Verdcourt, B, Beentje, HJ, 2010. Cyperaceae. In: Beentje, HJ, Ed., Flora of Tropical East Africa. Royal Botanic Gardens, Kew, Richmond: 1 - 466" title="Cyperaceae." volumeTitle="Flora of Tropical East Africa." year="2010">Hoenselaar et al. (2010)</bibRefCitation>
.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>