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<document id="A24BE6FC6E86745C464E36F0896668A3" ID-CLB-Dataset="299687" ID-DOI="10.11646/zootaxa.5476.1.14" ID-GBIF-Dataset="83be1d12-3615-41b3-9b8f-b1490f55488e" ID-ISSN="1175-5326" ID-Zenodo-Dep="12724044" ID-ZooBank="59712C82-A1A8-43F8-9B15-7FD4B43DC0E0" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1720374679655" checkinUser="plazi" docAuthor="Hsu, Jhih-Wei &amp; Shih, Hsi-Te" docDate="2024" docId="F5111D137568C644FF3BF89DFB974C5B" docLanguage="en" docName="zootaxa.5476.1.14.pdf" docOrigin="Zootaxa 5476 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5476.1.14" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parahelice undefined-1" docType="treatment" docVersion="4" lastPageNumber="144" masterDocId="0928656B756EC643FFACFFB7FFDC4D46" masterDocTitle="First records of Parahelice K. Sakai, Türkay &amp; Yang, 2006 and Pseudohelice K. Sakai, Türkay &amp; Yang, 2006 from Vanuatu, with description of a new species of Parahelice (Crustacea: Brachyura: Varunidae)" masterLastPageNumber="151" masterPageNumber="138" pageNumber="144" updateTime="1720709943798" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="EA8D01C7976AA3911C99747CDFEF48FC">First records of Parahelice K. Sakai, Türkay &amp; Yang, 2006 and Pseudohelice K. Sakai, Türkay &amp; Yang, 2006 from Vanuatu, with description of a new species of Parahelice (Crustacea: Brachyura: Varunidae)</mods:title>
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<mods:namePart id="F80F9041603CFAF283BD04EE5BDE4FCC">Hsu, Jhih-Wei</mods:namePart>
<mods:affiliation id="A4E2994860376D745D93C67767799C4B">National Chung Hsing University, Department of Life Science, 250 Kuo Kuang Road, Taichung 402, Taiwan.</mods:affiliation>
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<mods:namePart id="155795D20718007273E16147FD2B530A">Shih, Hsi-Te</mods:namePart>
<mods:affiliation id="6D0CA88CF8B8323B2555D9AE87D914CA">National Chung Hsing University, Department of Life Science and Global Change Biology Research Center, 250 Kuo Kuang Road, Taichung 402, Taiwan.</mods:affiliation>
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<taxonomicName id="BAB8D7867568C645FF3BF89DFE864A02" box="[151,346,1834,1861]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="4FCC70177568C645FF3BF89DFE864A02" bold="true" box="[151,346,1834,1861]" pageId="6" pageNumber="144">
<emphasis id="4FCC70177568C645FF3BF89DFEC54A03" bold="true" box="[151,281,1834,1861]" italics="true" pageId="6" pageNumber="144">Parahelice</emphasis>
sp. 1
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<paragraph id="7D07AC057568C645FF3BF8C5FC854AF7" blockId="6.[151,1436,1906,2041]" pageId="6" pageNumber="144">
<emphasis id="4FCC70177568C645FF3BF8C5FEA04ACA" bold="true" box="[151,380,1906,1932]" pageId="6" pageNumber="144">Material examined</emphasis>
.
<materialsCitation id="CDD0A6587568C645FE21F8C4FC894AF7" ID-GBIF-Occurrence="4905139304" collectingDate="2006-09-23" collectionCode="ZRC" country="Vanuatu" location="Palikulo Peninsula" municipality="Espiritu Santo Island" pageId="6" pageNumber="144" specimenCode="ZRC 2023.0245" specimenCount="2" specimenCount-female="2" stateProvince="Sanma">
<specimenCount id="6BBE678C7568C645FE21F8C4FE204ACB" box="[397,508,1907,1933]" count="2" pageId="6" pageNumber="144" type="female">2 females</specimenCount>
(21.3 × 17.2, 18.3 ×
<quantity id="BA4001E07568C645FD57F8C4FCBA4ACB" box="[763,870,1907,1933]" metricMagnitude="-2" metricUnit="m" metricValue="1.48" pageId="6" pageNumber="144" unit="mm" value="14.8">14.8 mm</quantity>
),
<specimenCode id="2D1E047E7568C645FCD0F8C4FBEA4ACB" box="[892,1078,1906,1933]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="6" pageNumber="144">ZRC 2023.0245</specimenCode>
, st. VM34,
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,
<collectingRegion id="BF7C62E77568C645FF3BF820FE8C4AF7" box="[151,336,1943,1969]" country="Vanuatu" name="Sanma" pageId="6" pageNumber="144">Sanma Province</collectingRegion>
,
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,
<collectingCountry id="05AFEC957568C645FDF8F820FD6F4AF7" box="[596,691,1943,1969]" name="Vanuatu" pageId="6" pageNumber="144">Vanuatu</collectingCountry>
,
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<collectingDate id="1942732D7568C645FD12F820FC894AF7" box="[702,853,1943,1969]" pageId="6" pageNumber="144" value="2006-09-23">23 Sep. 2006</collectingDate>
</date>
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.
</paragraph>
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<subSubSection id="35A2FF8E7568C644FF6BF80CFB974C5B" lastPageId="7" lastPageNumber="145" pageId="6" pageNumber="144" type="discussion">
<paragraph id="7D07AC057568C644FF6BF80CFB974C5B" blockId="6.[151,1436,1906,2041]" lastBlockId="7.[151,1437,150,285]" lastPageId="7" lastPageNumber="145" pageId="6" pageNumber="144">
<emphasis id="4FCC70177568C645FF6BF80CFEE84A93" bold="true" box="[199,308,1979,2005]" pageId="6" pageNumber="144">Remarks</emphasis>
. In the collection of
<taxonomicName id="BAB8D7867568C645FD9EF80CFD714A93" authorityName="K. Sakai, Turkay &amp; Yang" authorityYear="2006" box="[562,685,1979,2005]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="4FCC70177568C645FD9EF80CFD714A93" box="[562,685,1979,2005]" italics="true" pageId="6" pageNumber="144">Parahelice</emphasis>
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from
<collectingCountry id="05AFEC957568C645FD55F80CFC844A93" box="[761,856,1979,2005]" name="Vanuatu" pageId="6" pageNumber="144">Vanuatu</collectingCountry>
, there are an additional
<specimenCount id="6BBE678C7568C645FBDBF80CFADA4A93" box="[1143,1286,1979,2005]" count="3" pageId="6" pageNumber="144" type="generic">3 specimens</specimenCount>
superficially similar to
<taxonomicName id="BAB8D7867568C645FEA6F868FE644ABF" authorityName="Hsu &amp; Shih" authorityYear="2024" box="[266,440,2015,2041]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="ngankeeae">
<emphasis id="4FCC70177568C645FEA6F868FE644ABF" box="[266,440,2015,2041]" italics="true" pageId="6" pageNumber="144">Par. ngankeeae</emphasis>
</taxonomicName>
. The
<specimenCount id="6BBE678C7568C645FE55F868FD5F4ABF" box="[505,643,2015,2041]" count="3" pageId="6" pageNumber="144" type="generic">3 specimens</specimenCount>
form two closely related clades, sister to
<taxonomicName id="BAB8D7867568C645FBF6F868FAD44ABF" authorityName="Hsu &amp; Shih" authorityYear="2024" box="[1114,1288,2015,2041]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="ngankeeae">
<emphasis id="4FCC70177568C645FBF6F868FAD44ABF" box="[1114,1288,2015,2041]" italics="true" pageId="6" pageNumber="144">Par. ngankeeae</emphasis>
</taxonomicName>
based on the molecular analyses (
<figureCitation id="E583B0807569C644FE2DFF20FE1B4DF7" box="[385,455,150,177]" captionStart="FIGURE 5" captionStartId="11.[151,250,1683,1707]" captionTargetBox="[158,1429,676,1653]" captionTargetId="figure-291@11.[151,1436,669,1660]" captionTargetPageId="11" captionText="FIGURE 5. A Bayesian inference (BI) tree for species of Parahelice and Pseudohelice, based on the cytochrome c oxidase subunit I (COI) gene. Probability values at the nodes represent support values for BI and maximum likelihood (ML) analyses. For haplotypes from Vanuatu (shown in bold with “*”) obtained in this study, see Table 1. Other haplotypes from GenBank are shown as the accession numbers." figureDoi="http://doi.org/10.5281/zenodo.12724060" httpUri="https://zenodo.org/record/12724060/files/figure.png" pageId="7" pageNumber="145">Fig. 5</figureCitation>
;
<tableCitation id="303A99BE7569C644FE7AFF20FDF24DF7" box="[470,558,151,177]" captionStart="TABLE 2" captionStartId="8.[151,239,608,634]" captionTargetPageId="8" captionText="TABLE 2. Matrix of percentage pairwise nucleotide divergence with Kimura 2-parameter (K2P) distances (lower left) and mean numbers of differences (upper right) based on the" httpUri="http://table.plazi.org/id/29C7FC8D7566C64BFF3BFDD7F8294F3C" pageId="7" pageNumber="145" tableUuid="29C7FC8D7566C64BFF3BFDD7F8294F3C">Table 2</tableCitation>
). One clade (
<emphasis id="4FCC70177569C644FD66FF20FD284DF6" box="[714,756,151,176]" italics="true" pageId="7" pageNumber="145">Par</emphasis>
. sp. 1) has
<specimenCount id="6BBE678C7569C644FCD0FF20FC364DF7" box="[892,1002,151,177]" count="2" pageId="7" pageNumber="145" type="female">2 females</specimenCount>
(ZRC 2023.0245) that agree with the character of suborbital cristae of
<taxonomicName id="BAB8D7867569C644FDA6FF0CFD494D92" baseAuthorityName="Clark" baseAuthorityYear="1987" box="[522,661,187,212]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="145" phylum="Arthropoda" rank="species" species="georgei">
<emphasis id="4FCC70177569C644FDA6FF0CFD494D92" box="[522,661,187,212]" italics="true" pageId="7" pageNumber="145">Par. georgei</emphasis>
</taxonomicName>
described in K.
<bibRefCitation id="1929D1F47569C644FCE2FF0CFBFD4D93" author="Sakai, K. &amp; Turkay, M. &amp; Yang, S. - L." box="[846,1057,187,213]" pageId="7" pageNumber="145" pagination="1 - 76" refId="ref10458" refString="Sakai, K., Turkay, M. &amp; Yang, S. - L. (2006) Revision of the Helice / Chasmagnathus complex (Crustacea: Decapoda: Brachyura). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 565, 1 - 76." type="journal article" year="2006">
Sakai
<emphasis id="4FCC70177569C644FC38FF0CFC104D93" box="[916,972,187,213]" italics="true" pageId="7" pageNumber="145">et al.</emphasis>
(2006)
</bibRefCitation>
. Another clade (
<emphasis id="4FCC70177569C644FB76FF0CFAD44D92" box="[1242,1288,187,212]" italics="true" pageId="7" pageNumber="145">Par.</emphasis>
sp. 2) with
<specimenCount id="6BBE678C7569C644FA22FF0CFF114DBF" count="1" pageId="7" pageNumber="145" type="male">1 male</specimenCount>
(ZRC 2023.0246) also conforms to the male characters of suborbital cristae, chelipedal manus, and male G1s of
<taxonomicName id="BAB8D7867569C644FF1AFEB4FEE24C5A" baseAuthorityName="Clark" baseAuthorityYear="1987" box="[182,318,259,284]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="145" phylum="Arthropoda" rank="species" species="georgei">
<emphasis id="4FCC70177569C644FF1AFEB4FEE24C5A" box="[182,318,259,284]" italics="true" pageId="7" pageNumber="145">Par. georgei</emphasis>
</taxonomicName>
. Further study is necessary to confirm the identity of the two species.
</paragraph>
</subSubSection>
</treatment>
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