373 lines
41 KiB
XML
373 lines
41 KiB
XML
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<mods:title id="5F618CA9EE10AA2B89089ECFEE0F71C9">First records of Parahelice K. Sakai, Türkay & Yang, 2006 and Pseudohelice K. Sakai, Türkay & Yang, 2006 from Vanuatu, with description of a new species of Parahelice (Crustacea: Brachyura: Varunidae)</mods:title>
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<mods:namePart id="037A4CB9393B257F98DC2B0C479D0B6D">Shih, Hsi-Te</mods:namePart>
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<mods:affiliation id="BA382822389C2ABD54D6433CFA482C63">National Chung Hsing University, Department of Life Science and Global Change Biology Research Center, 250 Kuo Kuang Road, Taichung 402, Taiwan.</mods:affiliation>
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<subSubSection id="35A2FF8E7567C64AFF3BFC41FDA04956" box="[151,636,1014,1041]" pageId="9" pageNumber="147" type="nomenclature">
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<paragraph id="7D07AC057567C64AFF3BFC41FDA04956" blockId="9.[151,636,1014,1077]" box="[151,636,1014,1041]" pageId="9" pageNumber="147">
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<heading id="264F1B697567C64AFF3BFC41FDA04956" bold="true" box="[151,636,1014,1041]" fontSize="11" level="1" pageId="9" pageNumber="147" reason="1">
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<taxonomicName id="BAB8D7867567C64AFF3BFC41FDA04956" ID-CoL="4NY2C" authority="(Dana, 1851)" baseAuthorityName="Dana" baseAuthorityYear="1851" box="[151,636,1014,1041]" class="Malacostraca" family="Varunidae" genus="Pseudohelice" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="147" phylum="Arthropoda" rank="species" species="subquadrata">
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<emphasis id="4FCC70177567C64AFF3BFC41FDA04956" bold="true" box="[151,636,1014,1041]" pageId="9" pageNumber="147">
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<emphasis id="4FCC70177567C64AFF3BFC41FE0A4957" bold="true" box="[151,470,1014,1041]" italics="true" pageId="9" pageNumber="147">Pseudohelice subquadrata</emphasis>
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(
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<bibRefCitation id="1929D1F47567C64AFE49FC41FDA84956" author="Dana, J. D." box="[485,628,1014,1040]" pageId="9" pageNumber="147" pagination="247 - 254" refId="ref8812" refString="Dana, J. D. (1851) Conspectus Crustaceorum quae in Orbis Terrarum circumnavigatione, Carolo Wilkes e classe Reipublicae Foeratae Duce, lexit et descripsit. Crustacea Grapsoidea, (Cyclometopa, Edwardsii). Proceedings of the Academy of Natural Sciences of Philadelphia, 5, 247 - 254." type="journal article" year="1851">Dana, 1851</bibRefCitation>
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)
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</emphasis>
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||
</taxonomicName>
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||
</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="35A2FF8E7567C64AFF3BFBACFF374973" box="[151,235,1051,1077]" pageId="9" pageNumber="147" type="description">
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<paragraph id="7D07AC057567C64AFF3BFBACFF374973" blockId="9.[151,636,1014,1077]" box="[151,235,1051,1077]" pageId="9" pageNumber="147">
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(
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<figureCitation id="E583B0807567C64AFF33FBACFF3F4973" box="[159,227,1051,1077]" captionStart="FIGURE 4" captionStartId="10.[151,250,1467,1491]" captionTargetBox="[159,1427,345,1431]" captionTargetId="figure-84@10.[159,1427,345,1444]" captionTargetPageId="10" captionText="FIGURE 4. Pseudohelice subquadrata (Dana, 1851). A–F, male (20.2 × 18.1 mm, ZRC 2023.0248). A–F, male (11.2 × 9.0 mm). A, dorsal view; B, ventral view; C, outer view of left male cheliped; D, left male suborbital crista; E, F, dorsal view of male right G1." figureDoi="http://doi.org/10.5281/zenodo.12724058" httpUri="https://zenodo.org/record/12724058/files/figure.png" pageId="9" pageNumber="147">Fig. 4</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="35A2FF8E7567C64AFF3BFBE8FC454808" pageId="9" pageNumber="147" type="reference_group">
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<paragraph id="7D07AC057567C64AFF3BFBE8FB464931" blockId="9.[151,1342,1119,1358]" box="[151,1178,1119,1143]" pageId="9" pageNumber="147">
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<treatmentCitationGroup id="5DA88B2B7567C64AFF3BFBE8FB464931" box="[151,1178,1119,1143]" pageId="9" pageNumber="147">
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<treatmentCitation id="FC198A147567C64AFF3BFBE8FDA54931" author="Dana, J. D." box="[151,633,1119,1143]" page="251" pageId="9" pageNumber="147" year="1851">
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<taxonomicName id="BAB8D7867567C64AFF3BFBE8FDA54931" ID-CoL="TPZX" authority="Dana, 1851: 251" authorityName="Dana" authorityPageNumber="251" authorityYear="1851" box="[151,633,1119,1143]" class="Malacostraca" family="Varunidae" genus="Chasmagnathus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="147" phylum="Arthropoda" rank="species" species="subquadratus">
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<emphasis id="4FCC70177567C64AFF3BFBE8FE174931" box="[151,459,1119,1143]" italics="true" pageId="9" pageNumber="147">Chasmagnathus subquadratus</emphasis>
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<materialsCitation id="CDD0A6587567C64AFE7DFBD7FDA54931" ID-GBIF-Occurrence="4905139311" box="[465,633,1119,1143]" country="Australia" location="New South Wales" pageId="9" pageNumber="147" specimenCount="1" stateProvince="New South Wales">
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<bibRefCitation id="1929D1F47567C64AFE7DFBD7FDA54931" author="Dana, J. D." box="[465,633,1119,1143]" pageId="9" pageNumber="147" pagination="247 - 254" refId="ref8812" refString="Dana, J. D. (1851) Conspectus Crustaceorum quae in Orbis Terrarum circumnavigatione, Carolo Wilkes e classe Reipublicae Foeratae Duce, lexit et descripsit. Crustacea Grapsoidea, (Cyclometopa, Edwardsii). Proceedings of the Academy of Natural Sciences of Philadelphia, 5, 247 - 254." type="journal article" year="1851">Dana, 1851: 251</bibRefCitation>
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</materialsCitation>
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</taxonomicName>
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(
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<typeStatus id="A20312A77567C64AFD2BFBE8FD684931" box="[647,692,1119,1143]" pageId="9" pageNumber="147">type</typeStatus>
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locality: possibly
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<collectingRegion id="BF7C62E77567C64AFCC2FBD7FBF94931" box="[878,1061,1119,1143]" country="Australia" name="New South Wales" pageId="9" pageNumber="147">New South Wales</collectingRegion>
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,
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<collectingCountry id="05AFEC957567C64AFB82FBE8FB4C4931" box="[1070,1168,1119,1143]" name="Australia" pageId="9" pageNumber="147">Australia</collectingCountry>
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).
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||
</treatmentCitationGroup>
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</paragraph>
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||
<paragraph id="7D07AC057567C64AFF3BFBC9FBA949D0" blockId="9.[151,1342,1119,1358]" box="[151,1141,1150,1174]" pageId="9" pageNumber="147">
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<treatmentCitationGroup id="5DA88B2B7567C64AFF3BFBC9FBA949D0" box="[151,1141,1150,1174]" pageId="9" pageNumber="147">
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<emphasis id="4FCC70177567C64AFF3BFBC9FE9D49D0" box="[151,321,1150,1174]" italics="true" pageId="9" pageNumber="147">
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||
<taxonomicName id="BAB8D7867567C64AFF3BFBC9FEF849D0" ID-CoL="3K69C" authorityName="Hess" authorityYear="1865" box="[151,292,1150,1174]" class="Malacostraca" family="Varunidae" genus="Helice" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="147" phylum="Arthropoda" rank="species" species="leachii">Helice leachii</taxonomicName>
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.—
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</emphasis>
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K.
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<treatmentCitation id="FC198A147567C64AFECCFBC9FDDB49D0" author="Sakai, T." box="[352,519,1150,1174]" page="698" pageId="9" pageNumber="147" year="1939">
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||
<bibRefCitation id="1929D1F47567C64AFECCFBC9FDDB49D0" author="Sakai, T." box="[352,519,1150,1174]" pageId="9" pageNumber="147" pagination="42 - 111" refId="ref10377" refString="Sakai, T. (1939) n. k. In: Studies on the Crabs of Japan. IV. Brachygnatha, Brachyrhyncha. Vol. 3. Yokendo, Tokyo, pp. 365 - 741 + 1 - 23 (index), pls. 42 - 111." type="book chapter" year="1939">Sakai 1939: 698</bibRefCitation>
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</treatmentCitation>
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||
, text-fig. 126.—
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<treatmentCitation id="FC198A147567C64AFD01FBC9FCA749D0" author="Dai, A. - Y. & Yang, S. - L. & Song, Y. - Z. & Chen, G. - X." box="[685,891,1150,1174]" page="505" pageId="9" pageNumber="147" year="1986">
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||
<bibRefCitation id="1929D1F47567C64AFD01FBC9FCA749D0" author="Dai, A. - Y. & Yang, S. - L. & Song, Y. - Z. & Chen, G. - X." box="[685,891,1150,1174]" pageId="9" pageNumber="147" refId="ref8753" refString="Dai, A. - Y., Yang, S. - L., Song, Y. - Z. & Chen, G. - X. (1986) Crabs of the China Seas. China Ocean Press, Beijing, 17 + 568 pp., 74 pls. [in Chinese]" type="book" year="1986">
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Dai
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||
<emphasis id="4FCC70177567C64AFD76FBC8FCD149D0" box="[730,781,1150,1174]" italics="true" pageId="9" pageNumber="147">et al.</emphasis>
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1986: 505
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</bibRefCitation>
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||
</treatmentCitation>
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||
, fig. 286(5–6), pl. 72(1).
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||
</treatmentCitationGroup>
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</paragraph>
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||
<paragraph id="7D07AC057567C64AFF3BFB2BFCC049F3" blockId="9.[151,1342,1119,1358]" box="[151,796,1180,1205]" pageId="9" pageNumber="147">
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<treatmentCitationGroup id="5DA88B2B7567C64AFF3BFB2BFCC049F3" box="[151,796,1180,1205]" pageId="9" pageNumber="147">
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<emphasis id="4FCC70177567C64AFF3BFB2BFEE649F2" box="[151,314,1180,1204]" italics="true" pageId="9" pageNumber="147">
|
||
<taxonomicName id="BAB8D7867567C64AFF3BFB2BFEC149F2" ID-CoL="7WKD2" authorityName="Hess" authorityYear="1865" box="[151,285,1180,1204]" class="Malacostraca" family="Varunidae" genus="Helice" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="147" phylum="Arthropoda" rank="species" species="leachi">Helice leachi</taxonomicName>
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||
.—
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</emphasis>
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||
K.
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||
<treatmentCitation id="FC198A147567C64AFEF5FB2BFDDC49F2" author="Sakai, T." box="[345,512,1180,1204]" page="672" pageId="9" pageNumber="147" year="1976">
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<bibRefCitation id="1929D1F47567C64AFEF5FB2BFDDC49F2" author="Sakai, T." box="[345,512,1180,1204]" pageId="9" pageNumber="147" refId="ref10430" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Kodansha Ltd., Tokyo, 773 pp., 251 pls." type="book" year="1976">Sakai 1976: 672</bibRefCitation>
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</treatmentCitation>
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||
, textfig. 370a, b, pl. 228(2).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="7D07AC057567C64AFF3BFB0CFCA04995" blockId="9.[151,1342,1119,1358]" box="[151,892,1211,1236]" pageId="9" pageNumber="147">
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<treatmentCitationGroup id="5DA88B2B7567C64AFF3BFB0CFCA04995" box="[151,892,1211,1236]" pageId="9" pageNumber="147">
|
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<taxonomicName id="BAB8D7867567C64AFF3BFB0CFEA14995" ID-CoL="3K69C" authorityName="Hess" authorityYear="1865" box="[151,381,1211,1235]" class="Malacostraca" family="Varunidae" genus="Helice" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="147" phylum="Arthropoda" rank="species" species="leachii" subGenus="Helice">
|
||
<emphasis id="4FCC70177567C64AFF3BFB0CFF054995" box="[151,217,1211,1235]" italics="true" pageId="9" pageNumber="147">Helice</emphasis>
|
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(
|
||
<emphasis id="4FCC70177567C64AFF44FB0CFEF64995" box="[232,298,1211,1235]" italics="true" pageId="9" pageNumber="147">Helice</emphasis>
|
||
)
|
||
<emphasis id="4FCC70177567C64AFE95FB0CFEA14995" box="[313,381,1211,1235]" italics="true" pageId="9" pageNumber="147">leachii</emphasis>
|
||
</taxonomicName>
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||
<emphasis id="4FCC70177567C64AFED1FB0CFE474995" box="[381,411,1211,1235]" italics="true" pageId="9" pageNumber="147">.—</emphasis>
|
||
<treatmentCitation id="FC198A147567C64AFE37FB0CFD5D4995" author="Dai, A. - Y. & Yang, S. - L." box="[411,641,1211,1236]" page="553" pageId="9" pageNumber="147" year="1991">
|
||
<bibRefCitation id="1929D1F47567C64AFE37FB0CFD5D4995" author="Dai, A. - Y. & Yang, S. - L." box="[411,641,1211,1236]" pageId="9" pageNumber="147" refId="ref8714" refString="Dai, A. - Y. & Yang, S. - L. (1991) Crabs of the China Seas. China Ocean Press, Beijing, 21 + 608 pp., 74 pls." type="book" year="1991">Dai & Yang 1991: 553</bibRefCitation>
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||
</treatmentCitation>
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||
, fig. 286(5–6), pl. 72(1).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="7D07AC057567C64AFF3BFB6DFBC849B4" blockId="9.[151,1342,1119,1358]" box="[151,1044,1242,1266]" pageId="9" pageNumber="147">
|
||
<treatmentCitationGroup id="5DA88B2B7567C64AFF3BFB6DFBC849B4" box="[151,1044,1242,1266]" pageId="9" pageNumber="147">
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<taxonomicName id="BAB8D7867567C64AFF3BFB6DFD9E49B4" ID-CoL="4NY2C" baseAuthorityName="Dana" baseAuthorityYear="1851" box="[151,578,1242,1266]" class="Malacostraca" family="Varunidae" genus="Pseudohelice" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="147" phylum="Arthropoda" rank="species" species="subquadrata" subGenus="Pseudohelice">
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||
<emphasis id="4FCC70177567C64AFF3BFB6DFEC249B4" box="[151,286,1242,1266]" italics="true" pageId="9" pageNumber="147">Pseudohelice</emphasis>
|
||
(
|
||
<emphasis id="4FCC70177567C64AFE81FB6DFE6849B4" box="[301,436,1242,1266]" italics="true" pageId="9" pageNumber="147">Pseudohelice</emphasis>
|
||
)
|
||
<emphasis id="4FCC70177567C64AFE6FFB6DFD9E49B4" box="[451,578,1242,1266]" italics="true" pageId="9" pageNumber="147">subquadrata</emphasis>
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||
</taxonomicName>
|
||
<emphasis id="4FCC70177567C64AFDEEFB6DFDBC49B4" box="[578,608,1242,1266]" italics="true" pageId="9" pageNumber="147">.—</emphasis>
|
||
K.
|
||
<treatmentCitation id="FC198A147567C64AFDD3FB6DFC8D49B4" author="Sakai, K. & Turkay, M. & Yang, S. - L." box="[639,849,1242,1266]" page="37" pageId="9" pageNumber="147" year="2006">
|
||
<bibRefCitation id="1929D1F47567C64AFDD3FB6DFC8D49B4" author="Sakai, K. & Turkay, M. & Yang, S. - L." box="[639,849,1242,1266]" pageId="9" pageNumber="147" pagination="1 - 76" refId="ref10458" refString="Sakai, K., Turkay, M. & Yang, S. - L. (2006) Revision of the Helice / Chasmagnathus complex (Crustacea: Decapoda: Brachyura). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 565, 1 - 76." type="journal article" year="2006">
|
||
Sakai
|
||
<emphasis id="4FCC70177567C64AFD11FB6CFD2D49B4" box="[701,753,1242,1266]" italics="true" pageId="9" pageNumber="147">et al.</emphasis>
|
||
2006: 37
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
, figs. 53–59 (part).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="7D07AC057567C64AFF3BFB4FFD0A4869" blockId="9.[151,1342,1119,1358]" pageId="9" pageNumber="147">
|
||
<treatmentCitationGroup id="5DA88B2B7567C64AFF3BFB4FFD0A4869" pageId="9" pageNumber="147">
|
||
<emphasis id="4FCC70177567C64AFF3BFB4FFE1E4856" box="[151,450,1272,1296]" italics="true" pageId="9" pageNumber="147">
|
||
<taxonomicName id="BAB8D7867567C64AFF3BFB4FFE784856" ID-CoL="4NY2C" baseAuthorityName="Dana" baseAuthorityYear="1851" box="[151,420,1272,1296]" class="Malacostraca" family="Varunidae" genus="Pseudohelice" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="147" phylum="Arthropoda" rank="species" species="subquadrata">Pseudohelice subquadrata</taxonomicName>
|
||
.—
|
||
</emphasis>
|
||
<treatmentCitation id="FC198A147567C64AFE6EFB4EFD594857" author="Ng, P. K. L. & Shih, H. - T. & Ho, P. - H. & Wang, C. - H." box="[450,645,1272,1297]" page="110" pageId="9" pageNumber="147" year="2017">
|
||
<bibRefCitation id="1929D1F47567C64AFE6EFB4EFD594857" author="Ng, P. K. L. & Shih, H. - T. & Ho, P. - H. & Wang, C. - H." box="[450,645,1272,1297]" pageId="9" pageNumber="147" pagination="1 - 208" refId="ref9984" refString="Ng, P. K. L., Shih, H. - T., Ho, P. - H. & Wang, C. - H. (2017) An updated annotated checklist of brachyuran crabs from Taiwan (Crustacea: Decapoda). Journal of the National Taiwan Museum, 70, 1 - 208. https: // doi. org / 10.6532 / JNTM. 201712 _ 70 (3; 4). 01" type="journal article" year="2017">
|
||
Ng
|
||
<emphasis id="4FCC70177567C64AFE4BFB4EFDC64856" box="[487,538,1272,1296]" italics="true" pageId="9" pageNumber="147">et al.</emphasis>
|
||
2017: 110
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
(list).—
|
||
<treatmentCitation id="FC198A147567C64AFD75FB4FFC434856" author="Hsu, J. - W. & Shih, H. - T. & Innocenti, G." box="[729,927,1272,1297]" httpUri="http://treatment.plazi.org/id/CF1687A678616F29FC0B5E8EFEC4F924" page="96" pageId="9" pageNumber="147" year="2022">
|
||
<bibRefCitation id="1929D1F47567C64AFD75FB4FFC434856" author="Hsu, J. - W. & Shih, H. - T. & Innocenti, G." box="[729,927,1272,1297]" pageId="9" pageNumber="147" pagination="94 - 107" refId="ref9196" refString="Hsu, J. - W., Shih, H. - T. & Innocenti, G. (2022) A review of the mud crab genus Pseudohelice Sakai, Turkay & Yang, 2006 (Crustacea: Brachyura: Varunidae), with redescription of Cyclograpsus latreillii H. Milne Edwards, 1837, from the western Indian Ocean. Raffles Bulletin of Zoology, 70, 94 - 107. https: // doi. org / 10.26107 / RBZ- 2022 - 0007" type="journal article" year="2022">
|
||
Hsu
|
||
<emphasis id="4FCC70177567C64AFCA6FB4EFCE14856" box="[778,829,1272,1296]" italics="true" pageId="9" pageNumber="147">et al.</emphasis>
|
||
2022: 96
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
, figs. 1, 2.—
|
||
<treatmentCitation id="FC198A147567C64AFBB2FB4EFB264857" author="Prema, M. & Hsu, J. - W. & Shih, H. - T. & Ravichandran, S." box="[1054,1274,1272,1297]" page="13" pageId="9" pageNumber="147" year="2022">
|
||
<bibRefCitation id="1929D1F47567C64AFBB2FB4EFB264857" author="Prema, M. & Hsu, J. - W. & Shih, H. - T. & Ravichandran, S." box="[1054,1274,1272,1297]" pageId="9" pageNumber="147" refId="ref10142" refString="Prema, M., Hsu, J. - W., Shih, H. - T. & Ravichandran, S. (2022) First record of the genus Pseudohelice Sakai, Turkay & Yang, 2006 from India and description of a new pseudocryptic species (Crustacea: Brachyura: Varunidae). Zoological Studies, 61, 56. https: // doi. org / 10.6620 / ZS. 2022.61 - 56" type="journal volume" year="2022">
|
||
Prema
|
||
<emphasis id="4FCC70177567C64AFBC4FB4EFB404856" box="[1128,1180,1272,1296]" italics="true" pageId="9" pageNumber="147">et al.</emphasis>
|
||
2022: 13
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
(p. 3).
|
||
<taxonomicName id="BAB8D7867567C64AFF3BFAA0FE5E4869" box="[151,386,1303,1327]" class="Malacostraca" family="Varunidae" genus="Pseudohelice" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="147" phylum="Arthropoda" rank="species" species="quadrata">
|
||
<emphasis id="4FCC70177567C64AFF3BFAA0FE5E4869" box="[151,386,1303,1327]" italics="true" pageId="9" pageNumber="147">Pseudohelice quadrata</emphasis>
|
||
</taxonomicName>
|
||
[sic].—
|
||
<treatmentCitation id="FC198A147567C64AFE7EFAA0FD454869" author="Ng, P. K. L. & Guinot, D. & Davie, P. J. F." box="[466,665,1303,1327]" page="227" pageId="9" pageNumber="147" year="2008">
|
||
<bibRefCitation id="1929D1F47567C64AFE7EFAA0FD454869" author="Ng, P. K. L. & Guinot, D. & Davie, P. J. F." box="[466,665,1303,1327]" pageId="9" pageNumber="147" pagination="1 - 296" refId="ref9930" refString="Ng, P. K. L., Guinot, D. & Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 296." type="journal article" year="2008">
|
||
Ng
|
||
<emphasis id="4FCC70177567C64AFE56FAAFFDF14869" box="[506,557,1303,1327]" italics="true" pageId="9" pageNumber="147">et al.</emphasis>
|
||
2008: 227
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
(list).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="7D07AC057567C64AFF3BFA81FC454808" blockId="9.[151,1342,1119,1358]" box="[151,921,1334,1358]" pageId="9" pageNumber="147">
|
||
For synonyms before 2023, see K.
|
||
<bibRefCitation id="1929D1F47567C64AFE56FA81FD694808" author="Sakai, K. & Turkay, M. & Yang, S. - L." box="[506,693,1334,1358]" pageId="9" pageNumber="147" pagination="1 - 76" refId="ref10458" refString="Sakai, K., Turkay, M. & Yang, S. - L. (2006) Revision of the Helice / Chasmagnathus complex (Crustacea: Decapoda: Brachyura). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 565, 1 - 76." type="journal article" year="2006">
|
||
Sakai
|
||
<emphasis id="4FCC70177567C64AFD94FA80FDB04808" box="[568,620,1334,1358]" italics="true" pageId="9" pageNumber="147">et al.</emphasis>
|
||
(2006)
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="1929D1F47567C64AFD4AFA81FC494808" author="Hsu, J. - W. & Shih, H. - T. & Innocenti, G." box="[742,917,1334,1358]" pageId="9" pageNumber="147" pagination="94 - 107" refId="ref9196" refString="Hsu, J. - W., Shih, H. - T. & Innocenti, G. (2022) A review of the mud crab genus Pseudohelice Sakai, Turkay & Yang, 2006 (Crustacea: Brachyura: Varunidae), with redescription of Cyclograpsus latreillii H. Milne Edwards, 1837, from the western Indian Ocean. Raffles Bulletin of Zoology, 70, 94 - 107. https: // doi. org / 10.26107 / RBZ- 2022 - 0007" type="journal article" year="2022">
|
||
Hsu
|
||
<emphasis id="4FCC70177567C64AFCB9FA80FC954808" box="[789,841,1334,1358]" italics="true" pageId="9" pageNumber="147">et al.</emphasis>
|
||
(2022)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="35A2FF8E7567C64AFF3BFACBFE314B44" pageId="9" pageNumber="147" type="materials_examined">
|
||
<paragraph id="7D07AC057567C64AFF3BFACBFE314B44" blockId="9.[151,1437,1404,2043]" pageId="9" pageNumber="147">
|
||
<emphasis id="4FCC70177567C64AFF3BFACBFEA748D0" bold="true" box="[151,379,1404,1430]" pageId="9" pageNumber="147">Material examined</emphasis>
|
||
.
|
||
<materialsCitation id="CDD0A6587567C64AFE27FACBFD4948FC" ID-GBIF-Occurrence="4905139313" collectingDate="2006-09-23" collectionCode="ZRC" country="Vanuatu" location="Palikulo Peninsula" municipality="Espiritu Santo Island" pageId="9" pageNumber="147" specimenCode="ZRC 2023.0248" specimenCount="1" specimenCount-male="1" stateProvince="Sanma">
|
||
<specimenCount id="6BBE678C7567C64AFE27FACBFE0B48D0" box="[395,471,1404,1430]" count="1" pageId="9" pageNumber="147" type="male">1 male</specimenCount>
|
||
(20.2 ×
|
||
<quantity id="BA4001E07567C64AFD90FACBFD7A48D0" box="[572,678,1404,1430]" metricMagnitude="-2" metricUnit="m" metricValue="1.81" pageId="9" pageNumber="147" unit="mm" value="18.1">18.1 mm</quantity>
|
||
),
|
||
<specimenCode id="2D1E047E7567C64AFD16FACBFCAF48D0" box="[698,883,1404,1430]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="147">ZRC 2023.0248</specimenCode>
|
||
, st. VM34,
|
||
<location id="7867FADE7567C64AFC56FACBFB0C48D0" LSID="urn:lsid:plazi:treatment:F5111D137567C649FF3BFC41FE204C07:7867FADE7567C64AFC56FACBFB0C48D0" box="[1018,1232,1404,1430]" country="Vanuatu" municipality="Espiritu Santo Island" name="Palikulo Peninsula" pageId="9" pageNumber="147" stateProvince="Sanma">Palikulo Peninsula</location>
|
||
,
|
||
<collectingRegion id="BF7C62E77567C64AFB71FACBFA4448D0" box="[1245,1432,1404,1430]" country="Vanuatu" name="Sanma" pageId="9" pageNumber="147">Sanma Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="9D63367F7567C64AFF3BFA17FE5A48FC" box="[151,390,1440,1466]" pageId="9" pageNumber="147">Espiritu Santo Island</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="05AFEC957567C64AFE3EFA16FE2D48FD" box="[402,497,1441,1467]" name="Vanuatu" pageId="9" pageNumber="147">Vanuatu</collectingCountry>
|
||
,
|
||
<date id="09068AC57567C64AFE50FA17FD4948FC" box="[508,661,1440,1466]" pageId="9" pageNumber="147" value="2006-09-23">
|
||
<collectingDate id="1942732D7567C64AFE50FA17FD4948FC" box="[508,661,1440,1466]" pageId="9" pageNumber="147" value="2006-09-23">23 Sep. 2006</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<specimenCount id="6BBE678C7567C64AFD0CFA17FD2B48FC" box="[672,759,1440,1466]" count="2" pageId="9" pageNumber="147" type="male">2 males</specimenCount>
|
||
(16.1 × 14.3, 13.9 × 12.0 mm),
|
||
<materialsCitation id="CDD0A6587567C64AFBC8FA17FE364B44" ID-GBIF-Occurrence="4905139310" collectingDate="2006-09-23" collectionCode="ZRC" country="Vanuatu" location="Palikulo Peninsula" municipality="Espiritu Santo Island" pageId="9" pageNumber="147" specimenCode="ZRC 2023.0249" specimenCount="1" specimenCount-female="1" stateProvince="Sanma">
|
||
<specimenCount id="6BBE678C7567C64AFBC8FA17FB1A48FC" box="[1124,1222,1440,1466]" count="1" pageId="9" pageNumber="147" type="female">1 female</specimenCount>
|
||
(14.4 ×
|
||
<quantity id="BA4001E07567C64AFA84FA17FA4C48FC" box="[1320,1424,1440,1466]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="9" pageNumber="147" unit="mm" value="12.3">12.3 mm</quantity>
|
||
), 1 ovig. female (16.7 ×
|
||
<quantity id="BA4001E07567C64AFE37FA73FDD84898" box="[411,516,1476,1502]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="9" pageNumber="147" unit="mm" value="14.3">14.3 mm</quantity>
|
||
),
|
||
<specimenCode id="2D1E047E7567C64AFDB4FA73FD0C4898" box="[536,720,1476,1502]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="147">ZRC 2023.0249</specimenCode>
|
||
, st. VM34,
|
||
<location id="7867FADE7567C64AFCF9FA73FBF64898" LSID="urn:lsid:plazi:treatment:F5111D137567C649FF3BFC41FE204C07:7867FADE7567C64AFCF9FA73FBF64898" box="[853,1066,1476,1502]" country="Vanuatu" municipality="Espiritu Santo Island" name="Palikulo Peninsula" pageId="9" pageNumber="147" stateProvince="Sanma">Palikulo Peninsula</location>
|
||
,
|
||
<collectingRegion id="BF7C62E77567C64AFB9AFA73FB2C4898" box="[1078,1264,1476,1502]" country="Vanuatu" name="Sanma" pageId="9" pageNumber="147">Sanma Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="9D63367F7567C64AFB50FA73FF024B44" pageId="9" pageNumber="147">Espiritu Santo Island</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="05AFEC957567C64AFF45FA5EFE944B45" box="[233,328,1513,1539]" name="Vanuatu" pageId="9" pageNumber="147">Vanuatu</collectingCountry>
|
||
,
|
||
<date id="09068AC57567C64AFEFFFA5FFE364B44" box="[339,490,1512,1538]" pageId="9" pageNumber="147" value="2006-09-23">
|
||
<collectingDate id="1942732D7567C64AFEFFFA5FFE364B44" box="[339,490,1512,1538]" pageId="9" pageNumber="147" value="2006-09-23">23 Sep. 2006</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="35A2FF8E7567C64AFF6BF9BBFD684AF5" pageId="9" pageNumber="147" type="diagnosis">
|
||
<paragraph id="7D07AC057567C64AFF6BF9BBFD684AF5" blockId="9.[151,1437,1404,2043]" pageId="9" pageNumber="147">
|
||
<emphasis id="4FCC70177567C64AFF6BF9BBFE9E4B60" bold="true" box="[199,322,1548,1574]" pageId="9" pageNumber="147">Diagnosis.</emphasis>
|
||
Carapace (
|
||
<figureCitation id="E583B0807567C64AFE6AF9BBFDFC4B60" box="[454,544,1548,1574]" captionStart="FIGURE 4" captionStartId="10.[151,250,1467,1491]" captionTargetBox="[159,1427,345,1431]" captionTargetId="figure-84@10.[159,1427,345,1444]" captionTargetPageId="10" captionText="FIGURE 4. Pseudohelice subquadrata (Dana, 1851). A–F, male (20.2 × 18.1 mm, ZRC 2023.0248). A–F, male (11.2 × 9.0 mm). A, dorsal view; B, ventral view; C, outer view of left male cheliped; D, left male suborbital crista; E, F, dorsal view of male right G1." figureDoi="http://doi.org/10.5281/zenodo.12724058" httpUri="https://zenodo.org/record/12724058/files/figure.png" pageId="9" pageNumber="147">Fig. 4A</figureCitation>
|
||
) quadrate, slightly broader than long, ca. 1.2 times as broad as long; surface convex, irregularly punctate and granulate, with noticeable epigastric groove. Frontal margin slightly concave. Anterolateral margins with 3 teeth including orbital tooth. Suborbital crista sexually heteromorphic; in male (
|
||
<figureCitation id="E583B0807567C64AFF33F9CFFF274BD4" box="[159,251,1656,1682]" captionStart="FIGURE 4" captionStartId="10.[151,250,1467,1491]" captionTargetBox="[159,1427,345,1431]" captionTargetId="figure-84@10.[159,1427,345,1444]" captionTargetPageId="10" captionText="FIGURE 4. Pseudohelice subquadrata (Dana, 1851). A–F, male (20.2 × 18.1 mm, ZRC 2023.0248). A–F, male (11.2 × 9.0 mm). A, dorsal view; B, ventral view; C, outer view of left male cheliped; D, left male suborbital crista; E, F, dorsal view of male right G1." figureDoi="http://doi.org/10.5281/zenodo.12724058" httpUri="https://zenodo.org/record/12724058/files/figure.png" pageId="9" pageNumber="147">Fig. 4D</figureCitation>
|
||
), mesial part with several small rounded or elongated tubercles, followed by well-spaced several large, heteromorphic, elongated and less convex tubercles, lateral part with 1 significantly largest, elliptical and very convex tubercle, and 2–5 large convex tubercles; in female, mesial part with several dense small rounded tubercles, followed by closely spaced several larger, elongated, less convex and mostly unfused tubercles, lateral part with 1 largest and 2–5 larger convex tubercles; tubercles in lateral part more convex in male, less convex in female. Chelipedal palm stout, almost glabrous; usually unequal in male, equal in adult female. Ambulatory legs (
|
||
<figureCitation id="E583B0807567C64AFA90F89BFA484A00" box="[1340,1428,1836,1862]" captionStart="FIGURE 4" captionStartId="10.[151,250,1467,1491]" captionTargetBox="[159,1427,345,1431]" captionTargetId="figure-84@10.[159,1427,345,1444]" captionTargetPageId="10" captionText="FIGURE 4. Pseudohelice subquadrata (Dana, 1851). A–F, male (20.2 × 18.1 mm, ZRC 2023.0248). A–F, male (11.2 × 9.0 mm). A, dorsal view; B, ventral view; C, outer view of left male cheliped; D, left male suborbital crista; E, F, dorsal view of male right G1." figureDoi="http://doi.org/10.5281/zenodo.12724058" httpUri="https://zenodo.org/record/12724058/files/figure.png" pageId="9" pageNumber="147">Fig. 4A</figureCitation>
|
||
) slender, anterior margins of merus, carpus, and propodus covered with dense short setae. G1 (
|
||
<figureCitation id="E583B0807567C64AFB74F8E7FA914A2C" box="[1240,1357,1872,1898]" captionStart="FIGURE 4" captionStartId="10.[151,250,1467,1491]" captionTargetBox="[159,1427,345,1431]" captionTargetId="figure-84@10.[159,1427,345,1444]" captionTargetPageId="10" captionText="FIGURE 4. Pseudohelice subquadrata (Dana, 1851). A–F, male (20.2 × 18.1 mm, ZRC 2023.0248). A–F, male (11.2 × 9.0 mm). A, dorsal view; B, ventral view; C, outer view of left male cheliped; D, left male suborbital crista; E, F, dorsal view of male right G1." figureDoi="http://doi.org/10.5281/zenodo.12724058" httpUri="https://zenodo.org/record/12724058/files/figure.png" pageId="9" pageNumber="147">Fig. 4E–F</figureCitation>
|
||
) stout, distal part relatively flat with small chitinous beak-like structure distally, blunter; female vulvae with elongated semicircular sternal cover; sunken on inner part.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="35A2FF8E7567C649FF6BF80BFDC94D93" lastPageId="10" lastPageNumber="148" pageId="9" pageNumber="147" type="distribution">
|
||
<paragraph id="7D07AC057567C649FF6BF80BFDC94D93" blockId="9.[151,1437,1404,2043]" lastBlockId="10.[151,1436,151,321]" lastPageId="10" lastPageNumber="148" pageId="9" pageNumber="147">
|
||
<emphasis id="4FCC70177567C64AFF6BF80BFEBD4A90" bold="true" box="[199,353,1980,2006]" pageId="9" pageNumber="147">Distribution.</emphasis>
|
||
Western Pacific, from
|
||
<collectingCountry id="05AFEC957567C64AFDF3F80BFCFE4A90" box="[607,802,1980,2006]" name="French Polynesia" pageId="9" pageNumber="147">French Polynesia</collectingCountry>
|
||
,
|
||
<collectingCountry id="05AFEC957567C64AFC80F80BFCA74A90" box="[812,891,1980,2006]" name="Samoa" pageId="9" pageNumber="147">Samoa</collectingCountry>
|
||
,
|
||
<collectingCountry id="05AFEC957567C64AFC29F80BFC734A90" box="[901,943,1980,2006]" name="Fiji" pageId="9" pageNumber="147">Fiji</collectingCountry>
|
||
,
|
||
<collectingCountry id="05AFEC957567C64AFC15F80AFBC44A91" box="[953,1048,1981,2007]" name="Vanuatu" pageId="9" pageNumber="147">Vanuatu</collectingCountry>
|
||
,
|
||
<collectingCountry id="05AFEC957567C64AFB8DF80AFB0C4A90" box="[1057,1232,1980,2007]" name="New Caledonia" pageId="9" pageNumber="147">New Caledonia</collectingCountry>
|
||
, eastern
|
||
<collectingCountry id="05AFEC957567C64AFA83F80BFA454A90" box="[1327,1433,1980,2006]" name="Australia" pageId="9" pageNumber="147">Australia</collectingCountry>
|
||
, Caroline Is.,
|
||
<collectingCountry id="05AFEC957567C64AFE9FF857FEA64ABC" box="[307,378,2016,2042]" name="Guam" pageId="9" pageNumber="147">Guam</collectingCountry>
|
||
,
|
||
<collectingCountry id="05AFEC957567C64AFE27F856FE174ABD" box="[395,459,2017,2043]" name="Japan" pageId="9" pageNumber="147">Japan</collectingCountry>
|
||
(including Ryukyus and Ogasawara Is.),
|
||
<collectingCountry id="05AFEC957567C64AFC11F856FBDD4ABD" box="[957,1025,2017,2043]" name="South Korea" pageId="9" pageNumber="147">Korea</collectingCountry>
|
||
(
|
||
<collectingRegion id="BF7C62E77567C64AFBB9F857FBB84ABC" box="[1045,1124,2016,2042]" country="South Korea" name="Jeju" pageId="9" pageNumber="147">Jejudo</collectingRegion>
|
||
),
|
||
<collectingCountry id="05AFEC957567C64AFBD1F857FB0E4ABC" box="[1149,1234,2016,2042]" name="Taiwan" pageId="9" pageNumber="147">Taiwan</collectingCountry>
|
||
,
|
||
<collectingCountry id="05AFEC957567C64AFB4EF857FAF94ABC" box="[1250,1317,2016,2042]" name="China" pageId="9" pageNumber="147">China</collectingCountry>
|
||
(
|
||
<collectingRegion id="BF7C62E77567C64AFA96F857FA4D4ABC" box="[1338,1425,2016,2042]" country="China" name="Hainan" pageId="9" pageNumber="147">Hainan</collectingRegion>
|
||
),
|
||
<collectingCountry id="05AFEC957564C649FF3BFF20FEC44DF7" box="[151,280,151,177]" name="Philippines" pageId="10" pageNumber="148">Philippines</collectingCountry>
|
||
,
|
||
<collectingCountry id="05AFEC957564C649FE89FF20FE4E4DF7" box="[293,402,151,177]" name="Indonesia" pageId="10" pageNumber="148">Indonesia</collectingCountry>
|
||
(
|
||
<collectingRegion id="BF7C62E77564C649FE0EFF20FDD74DF7" box="[418,523,151,177]" country="Indonesia" name="Sulawesi Utara" pageId="10" pageNumber="148">Sulawesi</collectingRegion>
|
||
,
|
||
<collectingRegion id="BF7C62E77564C649FDB4FF20FD954DF7" box="[536,585,151,177]" country="Indonesia" name="Bali" pageId="10" pageNumber="148">Bali</collectingRegion>
|
||
, and Lombok), to western
|
||
<collectingCountry id="05AFEC957564C649FCD1FF20FC3D4DF7" box="[893,993,151,177]" name="Thailand" pageId="10" pageNumber="148">Thailand</collectingCountry>
|
||
(
|
||
<collectingRegion id="BF7C62E77564C649FC5DFF20FBF34DF7" box="[1009,1071,151,177]" country="Thailand" name="Surin" pageId="10" pageNumber="148">Surin</collectingRegion>
|
||
I.,
|
||
<collectingRegion id="BF7C62E77564C649FBF4FF20FB0B4DF7" box="[1112,1239,151,177]" country="Thailand" name="Phangnga" pageId="10" pageNumber="148">Phang Nga</collectingRegion>
|
||
) (K.
|
||
<bibRefCitation id="1929D1F47564C649FAB9FF20FF0F4D93" author="Sakai, K. & Turkay, M. & Yang, S. - L." pageId="10" pageNumber="148" pagination="1 - 76" refId="ref10458" refString="Sakai, K., Turkay, M. & Yang, S. - L. (2006) Revision of the Helice / Chasmagnathus complex (Crustacea: Decapoda: Brachyura). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 565, 1 - 76." type="journal article" year="2006">
|
||
Sakai
|
||
<emphasis id="4FCC70177564C649FAF0FF20FA524DF7" box="[1372,1422,151,177]" italics="true" pageId="10" pageNumber="148">et al</emphasis>
|
||
., 2006
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="1929D1F47564C649FF72FF0CFE514D93" author="Hsu, J. - W. & Shih, H. - T. & Innocenti, G." box="[222,397,187,213]" pageId="10" pageNumber="148" pagination="94 - 107" refId="ref9196" refString="Hsu, J. - W., Shih, H. - T. & Innocenti, G. (2022) A review of the mud crab genus Pseudohelice Sakai, Turkay & Yang, 2006 (Crustacea: Brachyura: Varunidae), with redescription of Cyclograpsus latreillii H. Milne Edwards, 1837, from the western Indian Ocean. Raffles Bulletin of Zoology, 70, 94 - 107. https: // doi. org / 10.26107 / RBZ- 2022 - 0007" type="journal article" year="2022">
|
||
Hsu
|
||
<emphasis id="4FCC70177564C649FEBEFF0CFE9F4D93" box="[274,323,187,213]" italics="true" pageId="10" pageNumber="148">et al</emphasis>
|
||
. 2022
|
||
</bibRefCitation>
|
||
; this study).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="35A2FF8E7564C649FF6BFF68FE204C07" pageId="10" pageNumber="148" type="discussion">
|
||
<paragraph id="7D07AC057564C649FF6BFF68FE204C07" blockId="10.[151,1436,151,321]" pageId="10" pageNumber="148">
|
||
<emphasis id="4FCC70177564C649FF6BFF68FEE74DBF" bold="true" box="[199,315,223,249]" pageId="10" pageNumber="148">Remarks.</emphasis>
|
||
For the morphological comparison among the three species in
|
||
<taxonomicName id="BAB8D7867564C649FBA6FF68FB434DBF" authorityName="K. Sakai, Turkay & Yang" authorityYear="2006" box="[1034,1183,223,249]" class="Malacostraca" family="Varunidae" genus="Pseudohelice" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="148" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4FCC70177564C649FBA6FF68FB434DBF" box="[1034,1183,223,249]" italics="true" pageId="10" pageNumber="148">Pseudohelice</emphasis>
|
||
</taxonomicName>
|
||
, see
|
||
<bibRefCitation id="1929D1F47564C649FB77FF68FA404DBF" author="Hsu, J. - W. & Shih, H. - T. & Innocenti, G." box="[1243,1436,223,249]" pageId="10" pageNumber="148" pagination="94 - 107" refId="ref9196" refString="Hsu, J. - W., Shih, H. - T. & Innocenti, G. (2022) A review of the mud crab genus Pseudohelice Sakai, Turkay & Yang, 2006 (Crustacea: Brachyura: Varunidae), with redescription of Cyclograpsus latreillii H. Milne Edwards, 1837, from the western Indian Ocean. Raffles Bulletin of Zoology, 70, 94 - 107. https: // doi. org / 10.26107 / RBZ- 2022 - 0007" type="journal article" year="2022">
|
||
Hsu
|
||
<emphasis id="4FCC70177564C649FABCFF68FA964DBF" box="[1296,1354,223,249]" italics="true" pageId="10" pageNumber="148">et al.</emphasis>
|
||
(2022)
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="1929D1F47564C649FF6BFEB4FE784C5B" author="Prema, M. & Hsu, J. - W. & Shih, H. - T. & Ravichandran, S." box="[199,420,259,285]" pageId="10" pageNumber="148" refId="ref10142" refString="Prema, M., Hsu, J. - W., Shih, H. - T. & Ravichandran, S. (2022) First record of the genus Pseudohelice Sakai, Turkay & Yang, 2006 from India and description of a new pseudocryptic species (Crustacea: Brachyura: Varunidae). Zoological Studies, 61, 56. https: // doi. org / 10.6620 / ZS. 2022.61 - 56" type="journal volume" year="2022">
|
||
Prema
|
||
<emphasis id="4FCC70177564C649FEBAFEB4FE934C5B" box="[278,335,259,285]" italics="true" pageId="10" pageNumber="148">et al.</emphasis>
|
||
(2022)
|
||
</bibRefCitation>
|
||
. This is a newly recorded species to
|
||
<collectingCountry id="05AFEC957564C649FCECFEB4FC434C5B" box="[832,927,259,285]" name="Vanuatu" pageId="10" pageNumber="148">Vanuatu</collectingCountry>
|
||
, within its distributional range in the western Pacific (see above distribution).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |