201 lines
20 KiB
XML
201 lines
20 KiB
XML
<document id="D55904162FFECD1CE454A6CC5C7AE36A" ID-DOI="10.11646/phytotaxa.344.3.6" ID-ISSN="1179-3163" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725659914456" checkinUser="felipe" docAuthor="Kessler, Michael & Smith, Alan R." docDate="2018" docId="F11187B0FFE0FFC2409AFE58952AFD56" docLanguage="en" docName="phytotaxa.344.3.6.pdf" docOrigin="Phytotaxa 344 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.344.3.6" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Asplenium achalense Hieron., Bot. Jahrb. Syst." docType="treatment" docVersion="1" lastPageNumber="280" masterDocId="0D28FFC8FFE4FFC74012F91D912DFFDB" masterDocTitle="Prodromus of a fern flora for Bolivia. XXIX. Aspleniaceae" masterLastPageNumber="280" masterPageNumber="259" pageNumber="259" updateTime="1725670605325" updateUser="GgImagineBatch" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="15936EDD60572FA4F161670C083E1B8F">
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<mods:title id="C89D5E15260C5B7DD3D2B91342A55D20">Prodromus of a fern flora for Bolivia. XXIX. Aspleniaceae</mods:title>
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</mods:titleInfo>
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<mods:name id="AF7BD4019B4C3891FC6E952E85D79AB8" type="personal">
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<mods:role id="92875072175557FEAB0E9B1A5588CB8E">
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<mods:roleTerm id="B14D9A0DC0DC37908B7AAC1E9F3390CB">Author</mods:roleTerm>
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<mods:namePart id="B2C8C3F0AFEC675301074C982B7E558F">Kessler, Michael</mods:namePart>
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<mods:affiliation id="C37DA3A07CDC9A64174F6F209D14497A">Systematic and Evolutionary Botany, University of Zurich, Zollikerstrasse 107, CH- 8008 Zurich, Switzerland</mods:affiliation>
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<mods:namePart id="174169197B9B6DE9010D622FD103993B">Smith, Alan R.</mods:namePart>
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<mods:affiliation id="9934924B2A4DE680EC69D22DB1F458DF">University Herbarium, 1001 Valley Life Sciences Bldg. # 2465, University of California, Berkeley, CA 94720 - 2465</mods:affiliation>
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<mods:typeOfResource id="409ACDD6458E5F4BD1F0B3DDC6D02C9F">text</mods:typeOfResource>
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<mods:title id="379DFDA94FEE8D914C7AF88943019A2F">Phytotaxa</mods:title>
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<mods:part id="44FE38164ED40347BA1203D0D0FDDF23">
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<mods:date id="1965A1675AC121C6C6773D08DA2F5353">2018</mods:date>
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<mods:number id="3B637E966DE092DF1757A16DD263678F">2018-03-16</mods:number>
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<mods:detail id="F8F8060454FE1B045247ED95AE95CCDF" type="volume">
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<mods:number id="EE4404F9BDD46B33366EEDFD432243E8">344</mods:number>
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<mods:start id="56B6CEA33A45C13846101B9553AC70EB">259</mods:start>
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<mods:url id="DC3F8662B74ED77BE192E92680666706">http://dx.doi.org/10.11646/phytotaxa.344.3.6</mods:url>
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<mods:classification id="8986CEE4CCB92BA84AF30E6CD36FB661">journal article</mods:classification>
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<mods:identifier id="3A2133AE35B6A712DE2788CB5A05EBB1" type="DOI">10.11646/phytotaxa.344.3.6</mods:identifier>
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<mods:identifier id="9A6B2A9E9DF67652393D02EAB975D44D" type="ISSN">1179-3163</mods:identifier>
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</mods:mods>
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<treatment id="F11187B0FFE0FFC2409AFE58952AFD56" LSID="urn:lsid:plazi:treatment:F11187B0FFE0FFC2409AFE58952AFD56" httpUri="http://treatment.plazi.org/id/F11187B0FFE0FFC2409AFE58952AFD56" lastPageId="5" pageId="4">
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<subSubSection id="31A2652DFFE0FFC3409AFE58929BF85F" pageId="4" type="nomenclature">
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<paragraph id="790736A6FFE0FFC3409AFE589267F8BB" blockId="4.[136,842,1861,1888]" box="[136,842,1861,1888]" pageId="4">
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<heading id="224F81CAFFE0FFC3409AFE589267F8BB" box="[136,842,1861,1888]" fontSize="11" level="2" pageId="4" reason="3">
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<treatmentCitation id="F81910B7FFE0FFC3409AFE58926BF8BB" author="Hieron." box="[136,838,1861,1888]" journal="Bot. Jahrb. Syst." page="378" pageId="4" score="144.29617767960897" volume="22" year="1896">
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<taxonomicName id="BEB84D25FFE0FFC3409AFE58938CF8BB" ID-CoL="686TL" authority="Hieron., Bot. Jahrb. Syst." box="[136,673,1861,1888]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="4" phylum="Tracheophyta" rank="species" species="achalense">
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<emphasis id="4BCCEAB4FFE0FFC3409AFE589052F8BB" bold="true" box="[136,383,1861,1888]" italics="true" pageId="4">Asplenium achalense</emphasis>
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Hieron., Bot. Jahrb. Syst.
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</taxonomicName>
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22: 378. 1896
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</treatmentCitation>
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.
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</heading>
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</paragraph>
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<paragraph id="790736A6FFE0FFC340AFFE76929BF85F" blockId="4.[136,1452,1899,2068]" box="[189,950,1899,1924]" pageId="4">
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<emphasis id="4BCCEAB4FFE0FFC340AFFE769002F85F" bold="true" box="[189,303,1899,1924]" pageId="4">Range:—</emphasis>
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<collectingCountry id="01AF7636FFE0FFC3413DFE7690ACF85F" box="[303,385,1899,1924]" name="Bolivia" pageId="4">Bolivia</collectingCountry>
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(
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<collectingCountry id="01AF7636FFE0FFC34183FE769091F85F" box="[401,444,1899,1924]" name="Switzerland" pageId="4">CH</collectingCountry>
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, CO, LP, SC, TA) to northwestern
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<collectingCountry id="01AF7636FFE0FFC3432DFE76929FF85F" box="[831,946,1899,1924]" name="Argentina" pageId="4">Argentina</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="31A2652DFFE0FFC240AFFE92952AFD56" lastPageId="5" pageId="4" type="biology_ecology">
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<paragraph id="790736A6FFE0FFC340AFFE929262F817" blockId="4.[136,1452,1899,2068]" pageId="4">
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<emphasis id="4BCCEAB4FFE0FFC340AFFE92906CF873" bold="true" box="[189,321,1935,1960]" pageId="4">Ecology:—</emphasis>
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Locally fairly common; epiphytic, terrestrial, and saxicolous in semihumid forests, especially with
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<taxonomicName id="BEB84D25FFE0FFC3409AFEAE903EF817" box="[136,275,1971,1996]" class="Magnoliopsida" family="Cunoniaceae" genus="Weinmannia" kingdom="Plantae" order="Oxalidales" pageId="4" phylum="Tracheophyta" rank="genus">
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<emphasis id="4BCCEAB4FFE0FFC3409AFEAE903EF817" box="[136,275,1971,1996]" italics="true" pageId="4">Weinmannia</emphasis>
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</taxonomicName>
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L. and
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<taxonomicName id="BEB84D25FFE0FFC3417BFEAE938FF817" authority="L. Her. ex Pers." authorityName="Pers." box="[361,674,1971,1996]" class="Magnoliopsida" family="Sapotaceae" genus="Podocarpus" kingdom="Plantae" order="Ericales" pageId="4" phylum="Tracheophyta" rank="genus">
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<emphasis id="4BCCEAB4FFE0FFC3417BFEAE90DDF817" box="[361,496,1971,1996]" italics="true" pageId="4">Podocarpus</emphasis>
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L.Her. ex Pers.
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</taxonomicName>
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;
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<quantity id="BE409B43FFE0FFC342BFFEAE9266F817" box="[685,843,1971,1996]" metricMagnitude="3" metricUnit="m" metricValue="2.175" metricValueMax="3.3" metricValueMin="1.05" pageId="4" unit="m" value="2175.0" valueMax="3300.0" valueMin="1050.0">1050–3300 m</quantity>
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.
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</paragraph>
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<paragraph id="790736A6FFE0FFC240AFFECA952AFD56" blockId="4.[136,1452,1899,2068]" lastBlockId="5.[136,1452,160,653]" lastPageId="5" pageId="4">
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<emphasis id="4BCCEAB4FFE0FFC340AFFECA900BF82B" bold="true" box="[189,294,2007,2032]" pageId="4">Notes:—</emphasis>
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The distinctions between
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<taxonomicName id="BEB84D25FFE0FFC34254FECA93F8F82B" box="[582,725,2007,2032]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="4" phylum="Tracheophyta" rank="species" species="achalense">
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<emphasis id="4BCCEAB4FFE0FFC34254FECA93F8F82B" box="[582,725,2007,2032]" italics="true" pageId="4">A. achalense</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="BEB84D25FFE0FFC3431FFECA924AF82B" box="[781,871,2007,2032]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="4" phylum="Tracheophyta" rank="species" species="serra">
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<emphasis id="4BCCEAB4FFE0FFC3431FFECA924AF82B" box="[781,871,2007,2032]" italics="true" pageId="4">A. serra</emphasis>
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</taxonomicName>
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, as well as between related species (
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<taxonomicName id="BEB84D25FFE0FFC34517FECA9488F82B" box="[1285,1445,2007,2032]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="4" phylum="Tracheophyta" rank="species" species="incurvatum">
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<emphasis id="4BCCEAB4FFE0FFC34517FECA9488F82B" box="[1285,1445,2007,2032]" italics="true" pageId="4">A. incurvatum</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="BEB84D25FFE0FFC3409AFEE69018F7CF" box="[136,309,2043,2068]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="4" phylum="Tracheophyta" rank="species" species="mosetenense">
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<emphasis id="4BCCEAB4FFE0FFC3409AFEE69018F7CF" box="[136,309,2043,2068]" italics="true" pageId="4">A. mosetenense</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="BEB84D25FFE0FFC34163FEE69333F7CF" box="[369,542,2043,2068]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="4" phylum="Tracheophyta" rank="species" species="tunquiniense">
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<emphasis id="4BCCEAB4FFE0FFC34163FEE69333F7CF" box="[369,542,2043,2068]" italics="true" pageId="4">A. tunquiniense</emphasis>
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</taxonomicName>
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), remain poorly understood (
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<bibRefCitation id="1D294B57FFE0FFC34376FEE6954EF7CF" author="Kessler, M. & Smith, A. R." box="[868,1123,2043,2068]" pageId="4" pagination="305 - 313" refId="ref16525" refString="Kessler, M. & Smith, A. R. (2006) Five new species of Asplenium (Aspleniaceae) from Bolivia. Candollea 61: 305 - 313." type="journal article" year="2006">Kessler & Smith 2006</bibRefCitation>
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). In addition to the characters presented in the key,
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<taxonomicName id="BEB84D25FFE1FFC24167F9BD9328FF62" box="[373,517,160,185]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="5" phylum="Tracheophyta" rank="species" species="achalense">
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<emphasis id="4BCCEAB4FFE1FFC24167F9BD9328FF62" box="[373,517,160,185]" italics="true" pageId="5">A. achalense</emphasis>
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</taxonomicName>
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tends to have blades with more strongly reduced distal pinnae (distal pinnae <50% as long as longest pinnae
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<emphasis id="4BCCEAB4FFE1FFC241BCF9D990E8FF06" box="[430,453,196,221]" italics="true" pageId="5">vs</emphasis>
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.>50% in
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<taxonomicName id="BEB84D25FFE1FFC2422DF9D993B6FF06" box="[575,667,196,221]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="5" phylum="Tracheophyta" rank="species" species="serra">
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<emphasis id="4BCCEAB4FFE1FFC2422DF9D993B6FF06" box="[575,667,196,221]" italics="true" pageId="5">A. serra</emphasis>
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</taxonomicName>
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) and more strongly biserrate pinna margins (
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<emphasis id="4BCCEAB4FFE1FFC2448DF9D9959BFF06" box="[1183,1206,196,221]" italics="true" pageId="5">vs</emphasis>
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. margins more finely and regularly serrulate). Juvenile plants of
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<taxonomicName id="BEB84D25FFE1FFC24278F9F593D7FEDA" box="[618,762,232,257]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="5" phylum="Tracheophyta" rank="species" species="achalense">
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<emphasis id="4BCCEAB4FFE1FFC24278F9F593D7FEDA" box="[618,762,232,257]" italics="true" pageId="5">A. achalense</emphasis>
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</taxonomicName>
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have strongly dissected leaves that are often up to 1-pinnate-pinnatisect in earliest leaves and become gradually less dissected as plants grow. In contrast, the earliest leaves of
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<taxonomicName id="BEB84D25FFE1FFC2409AF82D91C9FE92" box="[136,228,304,329]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="5" phylum="Tracheophyta" rank="species" species="serra">
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<emphasis id="4BCCEAB4FFE1FFC2409AF82D91C9FE92" box="[136,228,304,329]" italics="true" pageId="5">A. serra</emphasis>
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</taxonomicName>
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are typically undivided, resembling the terminal pinna of the mature plants, and only with age and size do the leaves attain additional lateral pinnae. Distinctions between
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<taxonomicName id="BEB84D25FFE1FFC2437BF84992D6FEB6" box="[873,1019,340,365]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="5" phylum="Tracheophyta" rank="species" species="achalense">
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<emphasis id="4BCCEAB4FFE1FFC2437BF84992D6FEB6" box="[873,1019,340,365]" italics="true" pageId="5">A. achalense</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="BEB84D25FFE1FFC24425F84995B9FEB6" box="[1079,1172,340,365]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="5" phylum="Tracheophyta" rank="species" species="serra">
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<emphasis id="4BCCEAB4FFE1FFC24425F84995B9FEB6" box="[1079,1172,340,365]" italics="true" pageId="5">A. serra</emphasis>
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</taxonomicName>
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are also supported by a molecular and morphometric study (
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<bibRefCitation id="1D294B57FFE1FFC2423EF865939FFE4A" author="Jonas, R." box="[556,690,376,401]" pageId="5" refId="ref16203" refString="Jonas, R. (2011) Revision and biogeographic analysis of the Asplenium serra - complex (Aspleniaceae) of Bolivia. M. Sc. thesis, University of Gottingen, Germany." type="book" year="2011">Jonas 2011</bibRefCitation>
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). However, three other species recognized here and by
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<bibRefCitation id="1D294B57FFE1FFC2452AF8659036FE6E" author="Kessler, M. & Smith, A. R." pageId="5" pagination="305 - 313" refId="ref16525" refString="Kessler, M. & Smith, A. R. (2006) Five new species of Asplenium (Aspleniaceae) from Bolivia. Candollea 61: 305 - 313." type="journal article" year="2006">Kessler & Smith (2006</bibRefCitation>
|
||
;
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<taxonomicName id="BEB84D25FFE1FFC2413BF88190E1FE6E" box="[297,460,412,437]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="5" phylum="Tracheophyta" rank="species" species="incurvatum">
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<emphasis id="4BCCEAB4FFE1FFC2413BF88190E1FE6E" box="[297,460,412,437]" italics="true" pageId="5">A. incurvatum</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="BEB84D25FFE1FFC241CFF88193A3FE6E" box="[477,654,412,437]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="5" phylum="Tracheophyta" rank="species" species="mosetenense">
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<emphasis id="4BCCEAB4FFE1FFC241CFF88193A3FE6E" box="[477,654,412,437]" italics="true" pageId="5">A. mosetenense</emphasis>
|
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</taxonomicName>
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, and
|
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<taxonomicName id="BEB84D25FFE1FFC242C0F88192A9FE6E" box="[722,900,412,437]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="5" phylum="Tracheophyta" rank="species" species="tunquiniense">
|
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<emphasis id="4BCCEAB4FFE1FFC242C0F88192A9FE6E" box="[722,900,412,437]" italics="true" pageId="5">A. tunquiniense</emphasis>
|
||
</taxonomicName>
|
||
) were found by
|
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<bibRefCitation id="1D294B57FFE1FFC2445EF88195F3FE6E" author="Jonas, R." box="[1100,1246,412,437]" pageId="5" refId="ref16203" refString="Jonas, R. (2011) Revision and biogeographic analysis of the Asplenium serra - complex (Aspleniaceae) of Bolivia. M. Sc. thesis, University of Gottingen, Germany." type="book" year="2011">Jonas (2011)</bibRefCitation>
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||
to nest within
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<taxonomicName id="BEB84D25FFE1FFC24581F88191D5FE02" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="5" phylum="Tracheophyta" rank="species" species="achalense">
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<emphasis id="4BCCEAB4FFE1FFC24581F88191D5FE02" italics="true" pageId="5">A. achalense</emphasis>
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</taxonomicName>
|
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, using molecular and morphometric data, and can be recognized only by the rhizome scale characters used in our key. To complicate matters, “
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<taxonomicName id="BEB84D25FFE1FFC2420BF8F99359FE26" box="[537,628,484,509]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="5" phylum="Tracheophyta" rank="species" species="serra">
|
||
<emphasis id="4BCCEAB4FFE1FFC2420BF8F99359FE26" box="[537,628,484,509]" italics="true" pageId="5">A. serra</emphasis>
|
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</taxonomicName>
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”, as described in some Argentinean floras (
|
||
<bibRefCitation id="1D294B57FFE1FFC2444FF8F9943DFE26" author="de la Sota, E. R." box="[1117,1296,484,509]" pageId="5" pagination="1 - 275" refId="ref15630" refString="de la Sota, E. R. (1977) Pteridophyta. In: Caberara, A. (Ed.) Flora de la Provincia de Jujuy. Collecion Cientifica del Instituto Nacional de Tecnologia Agropecuaria 13: 1 - 275." type="journal article" year="1977">de la Sota 1977</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="1D294B57FFE1FFC24509F8F991E9FDFA" author="Ganem, M. A. & Giudice, G. E. & Luna, M. L. & de la Sota, E. R." pageId="5" pagination="149 - 156" refId="ref15963" refString="Ganem, M. A., Giudice, G. E., Luna, M. L. & de la Sota, E. R. (2007) Revision of the group " Asplenium squamosum " in America. Candollea 62: 149 - 156." type="journal article" year="2007">
|
||
Ganem
|
||
<emphasis id="4BCCEAB4FFE1FFC24561F8F99488FE26" box="[1395,1445,484,509]" italics="true" pageId="5">et al</emphasis>
|
||
. 2007
|
||
</bibRefCitation>
|
||
), is not identical to
|
||
<taxonomicName id="BEB84D25FFE1FFC241B8FB15932BFDFA" box="[426,518,520,545]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="5" phylum="Tracheophyta" rank="species" species="serra">
|
||
<emphasis id="4BCCEAB4FFE1FFC241B8FB15932BFDFA" box="[426,518,520,545]" italics="true" pageId="5">A. serra</emphasis>
|
||
</taxonomicName>
|
||
as recognized here and actually represents a variant of
|
||
<taxonomicName id="BEB84D25FFE1FFC24495FB159435FDFA" box="[1159,1304,520,545]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="5" phylum="Tracheophyta" rank="species" species="achalense">
|
||
<emphasis id="4BCCEAB4FFE1FFC24495FB159435FDFA" box="[1159,1304,520,545]" italics="true" pageId="5">A. achalense</emphasis>
|
||
</taxonomicName>
|
||
as delimited by
|
||
<bibRefCitation id="1D294B57FFE1FFC240BEFB319096FD9E" author="Kessler, M. & Smith, A. R." box="[172,443,556,581]" pageId="5" pagination="305 - 313" refId="ref16525" refString="Kessler, M. & Smith, A. R. (2006) Five new species of Asplenium (Aspleniaceae) from Bolivia. Candollea 61: 305 - 313." type="journal article" year="2006">Kessler & Smith (2006)</bibRefCitation>
|
||
and here; it is possible that true
|
||
<taxonomicName id="BEB84D25FFE1FFC2433CFB3192A4FD9E" box="[814,905,556,581]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="5" phylum="Tracheophyta" rank="species" species="serra">
|
||
<emphasis id="4BCCEAB4FFE1FFC2433CFB3192A4FD9E" box="[814,905,556,581]" italics="true" pageId="5">A. serra</emphasis>
|
||
</taxonomicName>
|
||
does not occur in northeastern
|
||
<collectingCountry id="01AF7636FFE1FFC244FEFB319476FD9E" box="[1260,1371,556,581]" name="Argentina" pageId="5">Argentina</collectingCountry>
|
||
(
|
||
<bibRefCitation id="1D294B57FFE1FFC24579FB3191E9FDB2" author="Jonas, R." pageId="5" refId="ref16203" refString="Jonas, R. (2011) Revision and biogeographic analysis of the Asplenium serra - complex (Aspleniaceae) of Bolivia. M. Sc. thesis, University of Gottingen, Germany." type="book" year="2011">Jonas 2011</bibRefCitation>
|
||
).
|
||
<taxonomicName id="BEB84D25FFE1FFC240C4FB4D90EEFDB2" box="[214,451,592,617]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="5" phylum="Tracheophyta" rank="species" species="achalense">
|
||
<emphasis id="4BCCEAB4FFE1FFC240C4FB4D90EEFDB2" box="[214,451,592,617]" italics="true" pageId="5">Asplenium achalense</emphasis>
|
||
</taxonomicName>
|
||
may thus best be interpreted as a species complex; pending further research we choose to maintain five separate species, in order to call attention to the taxonomic issues.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |