treatments-xml/data/92/28/4D/92284D549274FF8AFE09FEB21AEF553F.xml

<document id="B2F2F531C9A98B7C831EC7157A512C43" ID-DOI="10.37828/em.2023.69.9" ID-ISSN="2336-9744" ID-Zenodo-Dep="13247356" ID-ZooBank="urn:lsid:zoobank.org:pub:60F4EA09-B706-4181-9707-2DB0AD98F078" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722972770478" checkinUser="felipe" docAuthor="Afifeh, Bassam Abu, Aloufi, Abdulhadi, Al-Saraireh, Mohammad, Badry, Ahmed, Al-Qahtni, Abdulmani H. &amp; Amr, Zuhair S." docDate="2023" docId="92284D549274FF8AFE09FEB21AEF553F" docLanguage="en" docName="EcolMontenegrina.69.91-106.pdf" docOrigin="Ecologica Montenegrina 69" docSource="http://dx.doi.org/10.37828/em.2023.69.9" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.2:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="2" docTitle="Leiurus nigellus Afifeh, Aloufi, Al-Saraireh, Badry, Al-Qahtni &amp; Amr, 2023, sp. nov." docType="treatment" docUuid="F2C4226B-233D-4266-BF78-8FD6C8C5195B" docUuidSource="ZooBank" docVersion="2" lastPageNumber="101" masterDocId="6E11352C9276FF80FFB3FFDF1823533D" masterDocTitle="A new remarkable species of Leiurus Ehrenberg, 1828 from Saudi Arabia (Scorpiones: Buthidae)" masterLastPageNumber="106" masterPageNumber="91" pageNumber="93" updateTime="1722980527386" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="5C413F622754BFFCE3D080D9C45CA324">A new remarkable species of Leiurus Ehrenberg, 1828 from Saudi Arabia (Scorpiones: Buthidae)</mods:title>
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<mods:namePart id="3C660A1CAFE18DD64B266BBDB2EF1832">Afifeh, Bassam Abu</mods:namePart>
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<mods:affiliation id="2B45666D65958D31467FDD76EA10C68D">Oncology Department, Royal Medical Services, Amman, Jordan. https: // orcid. org / 0000 - 0002 - 0560 - 5469</mods:affiliation>
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<mods:affiliation id="CCF2E37D9E276A40FDB4CAA38F7D7061">Department of Zoology, Faculty of Science, Al-Azhar University, Cairo, Egypt. https: // orcid. org / 0000 - 0002 - 2610 - 2019 &amp; Department of Invertebrates, National Center for Wildlife, Saudi Arabia. https: // orcid. org / 0009 - 0002 - 6072 - 7773</mods:affiliation>
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<mods:affiliation id="AF8C6168F6E01F092C89DEFD2EE9E91C">Department of Invertebrates, National Center for Wildlife, Saudi Arabia. https: // orcid. org / 0009 - 0002 - 6072 - 7773</mods:affiliation>
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<mods:namePart id="7891F894AC989CCD8CBDD9FBBC0DECA7">Amr, Zuhair S.</mods:namePart>
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<mods:affiliation id="C1EF28A6332B720732636640491854FF">Department of Biology, Jordan University of Science &amp; Technology, Irbid, Jordan. E-mail: amrz @ just. edu. jo; https: // orcid. org / 0000 - 0002 - 4778 - 2308</mods:affiliation>
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<taxonomicName id="DD8187C19274FF82FE09FEB21A5B52BA" ID-CoL="CCHMF" authorityName="Afifeh &amp; Aloufi &amp; Al-Saraireh &amp; Badry &amp; Al-Qahtni &amp; Amr" authorityYear="2023" box="[442,632,365,391]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="nigellus" status="sp. nov.">
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Abu Afifeh, Aloufi &amp; Al-Saraireh
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; Tables 1 and 3)
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, 
<specimenCode id="4A2754399274FF82FB64FDEB1D4D5173" box="[1239,1390,564,590]" collectionCode="BAPC" pageId="2" pageNumber="93">BAPC 0254</specimenCode>
, 
<collectingCounty id="F35F84CE9274FF82FACCFDEB1925514D" pageId="2" pageNumber="93">Al Buriakah</collectingCounty>
, beside old Tabuk-Al 
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highway, 
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, 
<collectingRegion id="D84532A09274FF82FC73FD891C72514D" box="[960,1105,598,624]" country="Saudi Arabia" name="Al Madinah" pageId="2" pageNumber="93">Al Madinah</collectingRegion>
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, 
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a.s.l.
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, 
<date id="6E3FDA829274FF82FD5AFDA71BAA51AF" box="[745,905,632,658]" pageId="2" pageNumber="93" value="2023-07-29">
<collectingDate id="7E7B236A9274FF82FD5AFDA71BAA51AF" box="[745,905,632,658]" pageId="2" pageNumber="93" value="2023-07-29">29 July 2023</collectingDate>
</date>
, 
<emphasis id="28F520509274FF82FC24FDA71B9851AF" box="[919,955,632,658]" italics="true" pageId="2" pageNumber="93">leg</emphasis>
. 
<collectorName id="B77499949274FF82FC7EFDA71C6B51AF" box="[973,1096,632,658]" pageId="2" pageNumber="93">A. Aloufi.</collectorName>
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<materialsCitation id="AAE9F61F9274FF82FBE2FDA71D5C51AF" box="[1105,1407,632,658]" collectionCode="BAPC" country="Saudi Arabia" pageId="2" pageNumber="93" specimenCode="BAPC 0252-0253" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="0C8737CB9274FF82FBE2FDA71CB551AF" box="[1105,1174,632,658]" count="2" pageId="2" pageNumber="93" type="female">2 ♀♀</specimenCount>
, 
<specimenCode id="4A2754399274FF82FB14FDA71D5C51AF" box="[1191,1407,632,658]" collectionCode="BAPC" pageId="2" pageNumber="93">BAPC 0252-0253</specimenCode>
</materialsCitation>
, 
<materialsCitation id="AAE9F61F9274FF82FA3DFDA719E4518E" collectionCode="BAPC" country="Saudi Arabia" pageId="2" pageNumber="93" specimenCode="BAPC 0255" specimenCount="2" specimenCount-female="1" specimenCount-subadult="1" typeStatus="paratype">
<specimenCount id="0C8737CB9274FF82FA3DFDA718D8518E" count="1" pageId="2" pageNumber="93" type="subadult">1 subadult</specimenCount>
<specimenCount id="0C8737CB9274FF82FEB4FD46193E518E" box="[263,285,665,691]" count="1" pageId="2" pageNumber="93" type="female">♀</specimenCount>
, 
<specimenCode id="4A2754399274FF82FE83FD4619E4518E" box="[304,455,665,691]" collectionCode="BAPC" pageId="2" pageNumber="93">BAPC 0255</specimenCode>
</materialsCitation>
, 
<materialsCitation id="AAE9F61F9274FF82FE64FD461B2E518E" box="[471,781,665,691]" collectionCode="BAPC" country="Saudi Arabia" pageId="2" pageNumber="93" specimenCode="BAPC 0256-0258" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="0C8737CB9274FF82FE64FD461A02518E" box="[471,545,665,691]" count="3" pageId="2" pageNumber="93" type="male">3 ♂♂</specimenCount>
, 
<specimenCode id="4A2754399274FF82FD81FD461B2E518E" box="[562,781,665,691]" collectionCode="BAPC" pageId="2" pageNumber="93">BAPC 0256-0258</specimenCode>
</materialsCitation>
, 
<materialsCitation id="AAE9F61F9274FF82FCAFFD461B1F51CA" collectingDate="2023-08-05" collectionCode="BAPC" collectorName="A. Aloufi." country="Saudi Arabia" county="Al Buriakah" elevation="1109" latitude="27.331665" location="Al Ula Governorate" longLatPrecision="1" longitude="37.791973" municipality="Ula" pageId="2" pageNumber="93" specimenCode="BAPC 0259" specimenCount="2" specimenCount-male="1" specimenCount-subadult="1" stateProvince="Al Madinah" typeStatus="paratype">
<specimenCount id="0C8737CB9274FF82FCAFFD461BB8518E" box="[796,923,665,691]" count="1" pageId="2" pageNumber="93" type="subadult">1 subadult</specimenCount>
<specimenCount id="0C8737CB9274FF82FC14FD461B9C518E" box="[935,959,665,691]" count="1" pageId="2" pageNumber="93" type="male">♂</specimenCount>
, 
<specimenCode id="4A2754399274FF82FC62FD461C44518E" box="[977,1127,665,691]" collectionCode="BAPC" pageId="2" pageNumber="93">BAPC 0259</specimenCode>
, 
<collectingCounty id="F35F84CE9274FF82FBC4FD461D33518E" box="[1143,1296,665,691]" pageId="2" pageNumber="93">Al Buriakah</collectingCounty>
, beside old Tabuk-Al 
<collectingMunicipality id="FA5A66389274FF82FEA0FD64191D51E8" box="[275,318,699,725]" pageId="2" pageNumber="93">Ula</collectingMunicipality>
highway, 
<location id="1F5EAA999274FF82FE0FFD641A8D51E8" LSID="urn:lsid:plazi:treatment:92284D549274FF8AFE09FEB21AEF553F:1F5EAA999274FF82FE0FFD641A8D51E8" box="[444,686,699,725]" country="Saudi Arabia" county="Al Buriakah" latitude="27.331665" longLatPrecision="1" longitude="37.791973" municipality="Ula" name="Al Ula Governorate" pageId="2" pageNumber="93" stateProvince="Al Madinah">Al Ula Governorate</location>
, 
<collectingRegion id="D84532A09274FF82FD0FFD641B6851E8" box="[700,843,699,725]" country="Saudi Arabia" name="Al Madinah" pageId="2" pageNumber="93">Al Madinah</collectingRegion>
<location id="1F5EAA999274FF82FCE6FD641CB051E8" LSID="urn:lsid:plazi:treatment:92284D549274FF8AFE09FEB21AEF553F:1F5EAA999274FF82FCE6FD641CB051E8" box="[853,1171,699,725]" country="Saudi Arabia" county="Al Buriakah" latitude="27.331665" longLatPrecision="1" longitude="37.791973" municipality="Ula" name="Al Monawwarah Province" pageId="2" pageNumber="93" stateProvince="Al Madinah">Al Monawwarah Province</location>
, 
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, 
<geoCoordinate id="7FB59A859274FF82FAC0FD64190751CA" degrees="37" direction="east" minutes="47" orientation="longitude" pageId="2" pageNumber="93" precision="1" seconds="31.11" value="37.791973">37° 47' 31.11&quot;E</geoCoordinate>
, 
<elevation id="91AC1B719274FF82FE83FD0219EB51CA" box="[304,456,733,759]" metricMagnitude="3" metricUnit="m" metricValue="1.109" pageId="2" pageNumber="93" unit="m" value="1109.0">
<quantity id="DD7951A79274FF82FE83FD0219AA51CA" box="[304,393,733,759]" metricMagnitude="3" metricUnit="m" metricValue="1.109" pageId="2" pageNumber="93" unit="m" value="1109.0">1109 m</quantity>
a.s.l.
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, 
<date id="6E3FDA829274FF82FE66FD021AA651CA" box="[469,645,733,759]" pageId="2" pageNumber="93" value="2023-08-05">
<collectingDate id="7E7B236A9274FF82FE66FD021AA651CA" box="[469,645,733,759]" pageId="2" pageNumber="93" value="2023-08-05">5 August 2023</collectingDate>
</date>
, 
<emphasis id="28F520509274FF82FD22FD021A9651CA" box="[657,693,733,759]" italics="true" pageId="2" pageNumber="93">leg</emphasis>
. 
<collectorName id="B77499949274FF82FD77FD021B1F51CA" box="[708,828,733,759]" pageId="2" pageNumber="93">A. Aloufi.</collectorName>
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</paragraph>
<paragraph id="1A3EFC429274FF82FF24FCFF1A165007" blockId="2.[151,1437,800,1298]" box="[151,565,800,826]" pageId="2" pageNumber="93">
<emphasis id="28F520509274FF82FF24FCFF1A0F5007" bold="true" box="[151,556,800,826]" pageId="2" pageNumber="93">Comparative material examined</emphasis>
:
</paragraph>
<paragraph id="1A3EFC429274FF82FF24FC9D1A3C562F" blockId="2.[151,1437,800,1298]" pageId="2" pageNumber="93">
<emphasis id="28F520509274FF82FF24FC9D1A035061" bold="true" box="[151,544,834,860]" pageId="2" pageNumber="93">
<collectingCountry id="6296BCD29274FF82FF24FC9D19675061" box="[151,324,834,860]" name="Saudi Arabia" pageId="2" pageNumber="93">Saudi Arabia</collectingCountry>
: 
<taxonomicName id="DD8187C19274FF82FEE1FC9D1A035061" authorityName="Lowe, Yagmur &amp; Kovarik" authorityYear="2014" box="[338,544,834,860]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis id="28F520509274FF82FEE1FC9D1A035061" bold="true" box="[338,544,834,860]" italics="true" pageId="2" pageNumber="93">Leiurus arabicus</emphasis>
</taxonomicName>
</emphasis>
: 
<materialsCitation id="AAE9F61F9274FF82FD82FC9D1C9C5043" collectingDate="2019-03-21" collectionCode="BAPC" collectorName="A. Aloufi." country="Saudi Arabia" elevation="412" latitude="24.103634" location="Al Monawwarah Province" longLatPrecision="1" longitude="38.93004" municipality="Alaab" pageId="2" pageNumber="93" specimenCode="BAPC 0150, BAPC 0151" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Al Madinah">
<specimenCount id="0C8737CB9274FF82FD82FC9D1A435061" box="[561,608,834,860]" count="1" pageId="2" pageNumber="93" type="male">1 ♂</specimenCount>
, 
<specimenCode id="4A2754399274FF82FDDDFC9D1B225061" box="[622,769,834,860]" collectionCode="BAPC" pageId="2" pageNumber="93">BAPC 0150</specimenCode>
, 
<specimenCount id="0C8737CB9274FF82FCBEFC9D1B195061" box="[781,826,834,860]" count="1" pageId="2" pageNumber="93" type="female">1 ♀</specimenCount>
, 
<specimenCode id="4A2754399274FF82FCF9FC9D1BFE5061" box="[842,989,834,860]" collectionCode="BAPC" pageId="2" pageNumber="93">BAPC 0151</specimenCode>
, 
<collectingMunicipality id="FA5A66389274FF82FC5AFC9D1C105061" box="[1001,1075,834,860]" pageId="2" pageNumber="93">Alaab</collectingMunicipality>
, 
<collectingRegion id="D84532A09274FF82FB8CFC9D1CEE5061" box="[1087,1229,834,860]" country="Saudi Arabia" name="Al Madinah" pageId="2" pageNumber="93">Al Madinah</collectingRegion>
<location id="1F5EAA999274FF82FB65FC9D19205043" LSID="urn:lsid:plazi:treatment:92284D549274FF8AFE09FEB21AEF553F:1F5EAA999274FF82FB65FC9D19205043" country="Saudi Arabia" latitude="24.103634" longLatPrecision="1" longitude="38.93004" municipality="Alaab" name="Al Monawwarah Province" pageId="2" pageNumber="93" stateProvince="Al Madinah">Al Monawwarah Province</location>
, 
<geoCoordinate id="7FB59A859274FF82FEA3FCBB19F05043" box="[272,467,868,894]" degrees="24" direction="north" minutes="06" orientation="latitude" pageId="2" pageNumber="93" precision="1" seconds="13.08" value="24.103634">24° 06' 13.08&quot;N</geoCoordinate>
, 
<geoCoordinate id="7FB59A859274FF82FE52FCBB1A825043" box="[481,673,868,894]" degrees="38" direction="east" minutes="55" orientation="longitude" pageId="2" pageNumber="93" precision="1" seconds="48.13" value="38.93004">38° 55' 48.13&quot;E</geoCoordinate>
, 
<elevation id="91AC1B719274FF82FD1DFCBB1B185043" box="[686,827,868,894]" metricMagnitude="2" metricUnit="m" metricValue="4.12" pageId="2" pageNumber="93" unit="m" value="412.0">
<quantity id="DD7951A79274FF82FD1DFCBB1ADA5043" box="[686,761,868,894]" metricMagnitude="2" metricUnit="m" metricValue="4.12" pageId="2" pageNumber="93" unit="m" value="412.0">412 m</quantity>
a.s.l.
</elevation>
, 
<date id="6E3FDA829274FF82FCF9FCBB1C275043" box="[842,1028,868,894]" pageId="2" pageNumber="93" value="2019-03-21">
<collectingDate id="7E7B236A9274FF82FCF9FCBB1C275043" box="[842,1028,868,894]" pageId="2" pageNumber="93" value="2019-03-21">21 March 2019</collectingDate>
</date>
, 
<emphasis id="28F520509274FF82FBA2FCBB1C165043" box="[1041,1077,868,894]" italics="true" pageId="2" pageNumber="93">leg</emphasis>
. 
<collectorName id="B77499949274FF82FBF6FCBB1C9C5043" box="[1093,1215,868,894]" pageId="2" pageNumber="93">A. Aloufi.</collectorName>
</materialsCitation>
<materialsCitation id="AAE9F61F9274FF82FB7BFCBB19DF50FC" collectingDate="2017-08-06" collectionCode="BAPC" collectorName="A. Aloufi." country="Saudi Arabia" elevation="838" latitude="24.7405" location="Al Monawwarah Province" longLatPrecision="1" longitude="39.715916" municipality="Wadi Al Jala'b" pageId="2" pageNumber="93" specimenCode="BAPC 0149" specimenCount="1" specimenCount-female="1" stateProvince="Al Madinah">
<specimenCount id="0C8737CB9274FF82FB7BFCBB1CD65043" box="[1224,1269,868,894]" count="1" pageId="2" pageNumber="93" type="female">1 ♀</specimenCount>
, 
<specimenCode id="4A2754399274FF82FAB6FCBB1DBB5043" box="[1285,1432,868,894]" collectionCode="BAPC" pageId="2" pageNumber="93">BAPC 0149</specimenCode>
, 
<collectingMunicipality id="FA5A66389274FF82FF24FC5A196E50A2" box="[151,333,901,927]" pageId="2" pageNumber="93">Wadi Al Jala'b</collectingMunicipality>
, 
<collectingRegion id="D84532A09274FF82FEEFFC5A19CF50A2" box="[348,492,901,927]" country="Saudi Arabia" name="Al Madinah" pageId="2" pageNumber="93">Al Madinah</collectingRegion>
<location id="1F5EAA999274FF82FE44FC5A1B1550A2" LSID="urn:lsid:plazi:treatment:92284D549274FF8AFE09FEB21AEF553F:1F5EAA999274FF82FE44FC5A1B1550A2" box="[503,822,901,927]" country="Saudi Arabia" latitude="24.7405" longLatPrecision="1" longitude="39.715916" municipality="Wadi Al Jala'b" name="Al Monawwarah Province" pageId="2" pageNumber="93" stateProvince="Al Madinah">Al Monawwarah Province</location>
, 
<geoCoordinate id="7FB59A859274FF82FCF6FC5A1C2F50A2" box="[837,1036,901,927]" degrees="24" direction="north" minutes="44" orientation="latitude" pageId="2" pageNumber="93" precision="1" seconds="25.80" value="24.7405">24° 44' 25.80&quot;N</geoCoordinate>
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<geoCoordinate id="7FB59A859274FF82FBA8FC5A1CFD50A2" box="[1051,1246,901,927]" degrees="39" direction="east" minutes="42" orientation="longitude" pageId="2" pageNumber="93" precision="1" seconds="57.29" value="39.715916">39° 42' 57.29&quot;E</geoCoordinate>
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<elevation id="91AC1B719274FF82FB5EFC5A1D5E50A2" box="[1261,1405,901,927]" metricMagnitude="2" metricUnit="m" metricValue="8.379999999999999" pageId="2" pageNumber="93" unit="m" value="838.0">
<quantity id="DD7951A79274FF82FB5EFC5A1D1950A2" box="[1261,1338,901,927]" metricMagnitude="2" metricUnit="m" metricValue="8.379999999999999" pageId="2" pageNumber="93" unit="m" value="838.0">838 m</quantity>
a.s.l.
</elevation>
, 
<date id="6E3FDA829274FF82FA3DFC5A191550FC" pageId="2" pageNumber="93" value="2017-08-06">
<collectingDate id="7E7B236A9274FF82FA3DFC5A191550FC" pageId="2" pageNumber="93" value="2017-08-06">6 August 2017</collectingDate>
</date>
, 
<emphasis id="28F520509274FF82FEF4FC78194850FC" box="[327,363,935,961]" italics="true" pageId="2" pageNumber="93">leg</emphasis>
. 
<collectorName id="B77499949274FF82FECDFC7819DF50FC" box="[382,508,935,961]" pageId="2" pageNumber="93">A. Aloufi.</collectorName>
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<taxonomicName id="DD8187C19274FF82FDBBFC781AFE50FC" authorityName="Lowe, Yagmur &amp; Kovarik" authorityYear="2014" box="[520,733,935,961]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="haenggii">
<emphasis id="28F520509274FF82FDBBFC781AFE50FC" bold="true" box="[520,733,935,961]" italics="true" pageId="2" pageNumber="93">Leiurus haenggii</emphasis>
</taxonomicName>
: 
<materialsCitation id="AAE9F61F9274FF82FD42FC7819AC5738" collectingDate="2022-05-23" collectionCode="BAPC" collectorName="A. Aloufi." country="Saudi Arabia" county="Wadi Reem" elevation="1090" latitude="23.918472" location="Al Monawwarah Province" longLatPrecision="1" longitude="39.301525" municipality="Al" pageId="2" pageNumber="93" specimenCode="BAPC 0200, BAPC 02011" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Al Madinah">
<specimenCount id="0C8737CB9274FF82FD42FC781B0750FC" box="[753,804,935,961]" count="1" pageId="2" pageNumber="93" type="male">1 ♂</specimenCount>
, 
<specimenCode id="4A2754399274FF82FC85FC781BEE50FC" box="[822,973,935,961]" collectionCode="BAPC" pageId="2" pageNumber="93">BAPC 0200</specimenCode>
, 
<specimenCount id="0C8737CB9274FF82FC6DFC781C2C50FC" box="[990,1039,935,961]" count="1" pageId="2" pageNumber="93" type="female">1 ♀</specimenCount>
, 
<specimenCode id="4A2754399274FF82FB90FC781CE450FC" box="[1059,1223,935,961]" collectionCode="BAPC" pageId="2" pageNumber="93">BAPC 02011</specimenCode>
, 
<collectingCounty id="F35F84CE9274FF82FB6BFC781D4B50FC" box="[1240,1384,935,961]" pageId="2" pageNumber="93">Wadi Reem</collectingCounty>
(
<collectingMunicipality id="FA5A66389274FF82FACEFC781DBF50FC" box="[1405,1436,935,961]" pageId="2" pageNumber="93">Al</collectingMunicipality>
Aseel), 
<collectingRegion id="D84532A09274FF82FF45FC1619A450DE" box="[246,391,969,995]" country="Saudi Arabia" name="Al Madinah" pageId="2" pageNumber="93">Al Madinah</collectingRegion>
<location id="1F5EAA999274FF82FE21FC161AF150DE" LSID="urn:lsid:plazi:treatment:92284D549274FF8AFE09FEB21AEF553F:1F5EAA999274FF82FE21FC161AF150DE" box="[402,722,969,995]" country="Saudi Arabia" county="Wadi Reem" latitude="23.918472" longLatPrecision="1" longitude="39.301525" municipality="Al" name="Al Monawwarah Province" pageId="2" pageNumber="93" stateProvince="Al Madinah">Al Monawwarah Province</location>
, 
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, 
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, 
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<quantity id="DD7951A79274FF82FB3DFC161CC950DE" box="[1166,1258,969,995]" metricMagnitude="3" metricUnit="m" metricValue="1.09" pageId="2" pageNumber="93" unit="m" value="1090.0">1090 m</quantity>
a.s.l.
</elevation>
, 
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</date>
, 
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. 
<collectorName id="B77499949274FF82FEA6FC3419AC5738" box="[277,399,1003,1029]" pageId="2" pageNumber="93">A. Aloufi.</collectorName>
</materialsCitation>
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, 
<specimenCode id="4A2754399274FF82FE67FC341A575738" box="[468,628,1003,1029]" collectionCode="BAPC" pageId="2" pageNumber="93">BAPC 02021</specimenCode>
, 
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Fegrah (
<collectingMunicipality id="FA5A66389274FF82FCBAFC341B915738" box="[777,946,1003,1029]" pageId="2" pageNumber="93">Wadi Mzaber</collectingMunicipality>
), 
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<location id="1F5EAA999274FF82FBEEFC341DBB5738" LSID="urn:lsid:plazi:treatment:92284D549274FF8AFE09FEB21AEF553F:1F5EAA999274FF82FBEEFC341DBB5738" box="[1117,1432,1003,1029]" country="Saudi Arabia" county="Al" latitude="24.362316" longLatPrecision="1" longitude="38.962433" municipality="Wadi Mzaber" name="Al Monawwarah Province" pageId="2" pageNumber="93" stateProvince="Al Madinah">Al Monawwarah Province</location>
, 
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, 
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, 
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a.s.l.
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, 
<date id="6E3FDA829274FF82FCADFBD31BCE571B" box="[798,1005,1036,1062]" pageId="2" pageNumber="93" value="2022-08-28">
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</date>
, 
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. 
<collectorName id="B77499949274FF82FB8FFBD31C9E571B" box="[1084,1213,1036,1062]" pageId="2" pageNumber="93">A. Aloufi.</collectorName>
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<emphasis id="28F520509274FF82FB7FFBD31D05571B" bold="true" box="[1228,1318,1036,1062]" pageId="2" pageNumber="93">
<collectingCountry id="6296BCD29274FF82FB7FFBD31D05571B" box="[1228,1318,1036,1062]" name="Jordan" pageId="2" pageNumber="93">Jordan</collectingCountry>
</emphasis>
: 
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<emphasis id="28F520509274FF82FA8DFBD319005775" bold="true" italics="true" pageId="2" pageNumber="93">Leiurus jordanensis</emphasis>
</taxonomicName>
: 
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, 
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, 
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, 
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Mudawwarah, 
<collectingRegion id="D84532A09274FF82FC3FFBF11C565775" box="[908,1141,1070,1096]" country="Jordan" name="Ma'an" pageId="2" pageNumber="93">Ma'an Governorate</collectingRegion>
, 
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, 
<date id="6E3FDA829274FF82FF4CFB8F19EB5757" box="[255,456,1104,1130]" pageId="2" pageNumber="93" value="2020-10-24">
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</date>
, 
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. 
<collectorName id="B77499949274FF82FDB4FB8F1A935757" box="[519,688,1104,1130]" pageId="2" pageNumber="93">B. Abu Afifeh</collectorName>
&amp; 
<collectorName id="B77499949274FF82FD65FB8F1BB35757" box="[726,912,1104,1130]" pageId="2" pageNumber="93">M. Al-Saraireh.</collectorName>
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<materialsCitation id="AAE9F61F9274FF82FC24FB8F18C75790" collectingDate="2022-07-15" collectionCode="BAPC" collectorName="B. Abu Afifeh" country="Jordan" elevation="890" latitude="29.433361" location="Wadi Rum" longLatPrecision="1" longitude="35.682446" pageId="2" pageNumber="93" specimenCode="BAPC 0102-0105" specimenCount="4" specimenCount-female="4" stateProvince="Al 'Aqabah">
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, 
<specimenCode id="4A2754399274FF82FC52FB8F1C955757" box="[993,1206,1104,1130]" collectionCode="BAPC" pageId="2" pageNumber="93">BAPC 0102-0105</specimenCode>
, 
<location id="1F5EAA999274FF82FB71FB8F1D655757" LSID="urn:lsid:plazi:treatment:92284D549274FF8AFE09FEB21AEF553F:1F5EAA999274FF82FB71FB8F1D655757" box="[1218,1350,1104,1130]" country="Jordan" latitude="29.433361" longLatPrecision="1" longitude="35.682446" name="Wadi Rum" pageId="2" pageNumber="93" stateProvince="Al 'Aqabah">Wadi Rum</location>
, 
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SE Al Ghal
</locationDeviation>
, 
<collectingRegion id="D84532A09274FF82FE87FBAD1A0357B1" box="[308,544,1138,1164]" country="Jordan" name="Al 'Aqabah" pageId="2" pageNumber="93">Aqaba Governorate</collectingRegion>
, 
<geoCoordinate id="7FB59A859274FF82FD9DFBAD1ACE57B1" box="[558,749,1138,1164]" degrees="29" direction="north" minutes="26" orientation="latitude" pageId="2" pageNumber="93" precision="1" seconds="00.1" value="29.433361">29° 26' 00.1&quot; N</geoCoordinate>
, 
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, 
<elevation id="91AC1B719274FF82FC75FBAD1C7757B1" box="[966,1108,1138,1164]" metricMagnitude="2" metricUnit="m" metricValue="8.9" pageId="2" pageNumber="93" unit="m" value="890.0">
<quantity id="DD7951A79274FF82FC75FBAD1C3157B1" box="[966,1042,1138,1164]" metricMagnitude="2" metricUnit="m" metricValue="8.9" pageId="2" pageNumber="93" unit="m" value="890.0">890 m</quantity>
a.s.l.
</elevation>
, 
<date id="6E3FDA829274FF82FBD7FBAD1D2057B1" box="[1124,1283,1138,1164]" pageId="2" pageNumber="93" value="2022-07-15">
<collectingDate id="7E7B236A9274FF82FBD7FBAD1D2057B1" box="[1124,1283,1138,1164]" pageId="2" pageNumber="93" value="2022-07-15">15 July 2022</collectingDate>
</date>
, 
<emphasis id="28F520509274FF82FAA2FBAD1D1657B1" box="[1297,1333,1138,1164]" italics="true" pageId="2" pageNumber="93">leg</emphasis>
. 
<collectorName id="B77499949274FF82FAF5FBAD18C75790" pageId="2" pageNumber="93">B. Abu Afifeh</collectorName>
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&amp; 
<materialsCitation id="AAE9F61F9274FF82FEBEFB4C199D5790" box="[269,446,1171,1197]" collectionCode="R" collectorName="Abu Afifeh" country="Jordan" pageId="2" pageNumber="93" specimenCount="1">
<collectionCode id="7C9064879274FF82FEBEFB4C19075790" box="[269,292,1171,1197]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="93">R</collectionCode>
. 
<collectorName id="B77499949274FF82FE82FB4C199D5790" box="[305,446,1171,1197]" pageId="2" pageNumber="93">Abu Afifeh</collectorName>
</materialsCitation>
. 
<materialsCitation id="AAE9F61F9274FF82FE7AFB4C1B1E57F2" collectingDate="2023-07-07" collectionCode="BAPC" collectorName="B. Abu Afifeh" country="Jordan" elevation="879" latitude="29.713446" location="Batn El-Ghol" longLatPrecision="1" longitude="36.148224" pageId="2" pageNumber="93" specimenCode="BAPC 0106-0111" specimenCount="6" specimenCount-female="6" stateProvince="Ma'an">
<specimenCount id="0C8737CB9274FF82FE7AFB4C1A2D5790" box="[457,526,1171,1197]" count="6" pageId="2" pageNumber="93" type="female">6 ♀♀</specimenCount>
, 
<specimenCode id="4A2754399274FF82FDADFB4C1AD55790" box="[542,758,1171,1197]" collectionCode="BAPC" pageId="2" pageNumber="93">BAPC 0106-0111</specimenCode>
, 
<location id="1F5EAA999274FF82FCB0FB4C1B865790" LSID="urn:lsid:plazi:treatment:92284D549274FF8AFE09FEB21AEF553F:1F5EAA999274FF82FCB0FB4C1B865790" box="[771,933,1171,1197]" country="Jordan" latitude="29.713446" longLatPrecision="1" longitude="36.148224" name="Batn El-Ghol" pageId="2" pageNumber="93" stateProvince="Ma'an">Batn El-Ghol</location>
, 
<collectingRegion id="D84532A09274FF82FC00FB4C1CB95790" box="[947,1178,1171,1197]" country="Jordan" name="Ma'an" pageId="2" pageNumber="93">Ma'an Governorate</collectingRegion>
, 
<geoCoordinate id="7FB59A859274FF82FB14FB4C1D455790" box="[1191,1382,1171,1197]" degrees="29" direction="north" minutes="42" orientation="latitude" pageId="2" pageNumber="93" precision="1" seconds="48.4" value="29.713446">29° 42' 48.4&quot; N</geoCoordinate>
, 
<geoCoordinate id="7FB59A859274FF82FAC0FB4C193D57F2" degrees="36" direction="east" minutes="08" orientation="longitude" pageId="2" pageNumber="93" precision="1" seconds="53.6" value="36.148224">36° 08' 53.6&quot; E</geoCoordinate>
, 
<elevation id="91AC1B719274FF82FE98FB6A199457F2" box="[299,439,1205,1231]" metricMagnitude="2" metricUnit="m" metricValue="8.79" pageId="2" pageNumber="93" unit="m" value="879.0">
<quantity id="DD7951A79274FF82FE98FB6A195557F2" box="[299,374,1205,1231]" metricMagnitude="2" metricUnit="m" metricValue="8.79" pageId="2" pageNumber="93" unit="m" value="879.0">879 m</quantity>
a.s.l.
</elevation>
, 
<date id="6E3FDA829274FF82FE76FB6A1A7057F2" box="[453,595,1205,1231]" pageId="2" pageNumber="93" value="2023-07-07">
<collectingDate id="7E7B236A9274FF82FE76FB6A1A7057F2" box="[453,595,1205,1231]" pageId="2" pageNumber="93" value="2023-07-07">7 July 2023</collectingDate>
</date>
, 
<emphasis id="28F520509274FF82FDECFB6A1AA057F2" box="[607,643,1205,1231]" italics="true" pageId="2" pageNumber="93">leg</emphasis>
. 
<collectorName id="B77499949274FF82FD20FB6A1B1E57F2" box="[659,829,1205,1231]" pageId="2" pageNumber="93">B. Abu Afifeh</collectorName>
</materialsCitation>
&amp; 
<materialsCitation id="AAE9F61F9274FF82FCD6FB6A1CD557F2" box="[869,1270,1205,1231]" collectionCode="R" country="Jordan" location="Abu Afifeh." pageId="2" pageNumber="93" specimenCount="1">
<collectionCode id="7C9064879274FF82FCD6FB6A1B5F57F2" box="[869,892,1205,1231]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="93">R</collectionCode>
. 
<location id="1F5EAA999274FF82FC3BFB6A1C3B57F2" LSID="urn:lsid:plazi:treatment:92284D549274FF8AFE09FEB21AEF553F:1F5EAA999274FF82FC3BFB6A1C3B57F2" box="[904,1048,1205,1231]" country="Jordan" name="Abu Afifeh." pageId="2" pageNumber="93">Abu Afifeh.</location>
<taxonomicName id="DD8187C19274FF82FB93FB6A1CD557F2" baseAuthorityName="Birula" baseAuthorityYear="1908" box="[1056,1270,1205,1231]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="hebraeus">
<emphasis id="28F520509274FF82FB93FB6A1CD557F2" bold="true" box="[1056,1270,1205,1231]" italics="true" pageId="2" pageNumber="93">Leiurus hebraeus</emphasis>
</taxonomicName>
</materialsCitation>
: 
<materialsCitation id="AAE9F61F9274FF82FAB5FB6A1A3C562F" collectingDate="2023-07-20" collectionCode="BAPC" collectorName="B. Abu Afifeh &amp; M. Al-Saraireh." country="Jordan" elevation="5" latitude="31.93611" location="Wadi Shueib" longLatPrecision="1" longitude="35.66528" pageId="2" pageNumber="93" specimenCode="BAPC 0300-0301" specimenCount="2" specimenCount-female="2" stateProvince="Al Balqa'">
<specimenCount id="0C8737CB9274FF82FAB5FB6A1D6157F2" box="[1286,1346,1205,1231]" count="2" pageId="2" pageNumber="93" type="female">2♀♀</specimenCount>
, 
<specimenCode id="4A2754399274FF82FAE3FB6A193A57CC" collectionCode="BAPC" pageId="2" pageNumber="93">BAPC 0300-0301</specimenCode>
, 
<location id="1F5EAA999274FF82FE95FB0819E057CC" LSID="urn:lsid:plazi:treatment:92284D549274FF8AFE09FEB21AEF553F:1F5EAA999274FF82FE95FB0819E057CC" box="[294,451,1239,1265]" country="Jordan" latitude="31.93611" longLatPrecision="1" longitude="35.66528" name="Wadi Shueib" pageId="2" pageNumber="93" stateProvince="Al Balqa'">Wadi Shueib</location>
, 
<collectingRegion id="D84532A09274FF82FE62FB081A9657CC" box="[465,693,1239,1265]" country="Jordan" name="Al Balqa'" pageId="2" pageNumber="93">Balqa Governorate</collectingRegion>
, 
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<geoCoordinate id="7FB59A859274FF82FC35FB081C1457CC" box="[902,1079,1239,1265]" degrees="35" direction="east" minutes="39" orientation="longitude" pageId="2" pageNumber="93" precision="1" seconds="55.0" value="35.66528">35°39' 55.0&quot; E</geoCoordinate>
, - 
<elevation id="91AC1B719274FF82FBFDFB081C9E57CC" box="[1102,1213,1239,1265]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="93" unit="m" value="5.0">
<quantity id="DD7951A79274FF82FBFDFB081C5E57CC" box="[1102,1149,1239,1265]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="93" unit="m" value="5.0">5 m</quantity>
a.s.l.
</elevation>
<date id="6E3FDA829274FF82FB75FB081D4757CC" box="[1222,1380,1239,1265]" pageId="2" pageNumber="93" value="2023-07-20">
<collectingDate id="7E7B236A9274FF82FB75FB081D4757CC" box="[1222,1380,1239,1265]" pageId="2" pageNumber="93" value="2023-07-20">20 July 2023</collectingDate>
</date>
, 
<emphasis id="28F520509274FF82FAC2FB081DB657CC" box="[1393,1429,1239,1265]" italics="true" pageId="2" pageNumber="93">leg</emphasis>
. 
<collectorName id="B77499949274FF82FF24FB271963562F" box="[151,320,1272,1298]" pageId="2" pageNumber="93">B. Abu Afifeh</collectorName>
&amp; 
<collectorName id="B77499949274FF82FED6FB271A3C562F" box="[357,543,1272,1298]" pageId="2" pageNumber="93">M. Al-Saraireh.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="529BAFC99274FF82FF24FAE31BE45645" pageId="2" pageNumber="93" type="etymology">
<paragraph id="1A3EFC429274FF82FF24FAE31BE45645" blockId="2.[151,1437,1340,1400]" pageId="2" pageNumber="93">
<emphasis id="28F520509274FF82FF24FAE3193D566B" bold="true" box="[151,286,1340,1366]" pageId="2" pageNumber="93">Etymology</emphasis>
: The specific epithet from Latin 
<taxonomicName id="DD8187C19274FF82FD16FAE31B2D566B" authorityName="Afifeh &amp; Aloufi &amp; Al-Saraireh &amp; Badry &amp; Al-Qahtni &amp; Amr" authorityYear="2023" box="[677,782,1340,1366]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="nigellus">
“ 
<emphasis id="28F520509274FF82FD01FAE31B2D566B" box="[690,782,1340,1366]" italics="true" pageId="2" pageNumber="93">nigellus</emphasis>
</taxonomicName>
” is masculine adjective, diminutive of “ 
<emphasis id="28F520509274FF82FB59FAE31D0B566B" box="[1258,1320,1340,1366]" italics="true" pageId="2" pageNumber="93">niger</emphasis>
” (Black), which refers to the predominantly blackish colour of the new species.
</paragraph>
</subSubSection>
<subSubSection id="529BAFC99274FF82FF24FA7E19F054EA" pageId="2" pageNumber="93" type="diagnosis">
<paragraph id="1A3EFC429274FF82FF24FA7E19F054EA" blockId="2.[151,1437,1441,2007]" pageId="2" pageNumber="93">
<emphasis id="28F520509274FF82FF24FA7E19335686" bold="true" box="[151,272,1441,1467]" pageId="2" pageNumber="93">Diagnosis</emphasis>
: Large 
<taxonomicName id="DD8187C19274FF82FEC5FA7E19EC5686" authorityName="Ehrenberg" authorityYear="1828" box="[374,463,1441,1467]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="28F520509274FF82FEC5FA7E19EC5686" box="[374,463,1441,1467]" italics="true" pageId="2" pageNumber="93">Leiurus</emphasis>
</taxonomicName>
, adults 
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in total length, carapace of female reaching 
<quantity id="DD7951A79274FF82FB4FFA7E1D5B5686" box="[1276,1400,1441,1467]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="2" pageNumber="93" unit="mm" value="11.4">11.40 mm</quantity>
in length; body colour is predominantly black; carapace, mesosoma, metasoma and telson, pedipalps and proximal segments of legs black to blackish-brown; distal third of pedipalp chela fingers and the three most distal segments of legs dark yellow to yellowish-brown; anterior margin of carapace slightly concave; all carapace carinae strongly developed; central lateral and posterior median carinae fused to form lyre shaped row of granules. chelicerae with characteristic buthid dentition (Vachon, 1963); tergites I–II, VII pentacarinate, III–VI tricarinate; sternites III with weak to moderate median carinae in females; strong in males; pectinal tooth counts ranging from 
<quantity id="DD7951A79274FF82FD2EF9521B27559A" box="[669,772,1677,1703]" metricMagnitude="-1" metricUnit="m" metricValue="9.525" metricValueMax="9.906" metricValueMin="9.144" pageId="2" pageNumber="93" unit="in" value="37.5" valueMax="39.0" valueMin="36.0">36-39 in</quantity>
males and 
<quantity id="DD7951A79274FF82FC20F9521BDA559A" box="[915,1017,1677,1703]" metricMagnitude="-1" metricUnit="m" metricValue="7.874" metricValueMax="8.382" metricValueMin="7.366" pageId="2" pageNumber="93" unit="in" value="31.0" valueMax="33.0" valueMin="29.0">29-33 in</quantity>
females. Pedipalps slender; femur L/W 
<quantity id="DD7951A79274FF82FF65F970190955F4" box="[214,298,1711,1737]" metricMagnitude="-1" metricUnit="m" metricValue="1.0795" pageId="2" pageNumber="93" unit="in" value="4.25">4.25 in</quantity>
male, 
<quantity id="DD7951A79274FF82FEC8F9701A2F55F4" box="[379,524,1711,1737]" metricMagnitude="-2" metricUnit="m" metricValue="9.8552" metricValueMax="10.0584" metricValueMin="9.652" pageId="2" pageNumber="93" unit="in" value="3.88" valueMax="3.96" valueMin="3.8">3.80-3.96 in</quantity>
females; patella L/W 4.00 in male, 
<quantity id="DD7951A79274FF82FC0EF9701C6D55F4" box="[957,1102,1711,1737]" metricMagnitude="-2" metricUnit="m" metricValue="8.8265" metricValueMax="9.144" metricValueMin="8.509" pageId="2" pageNumber="93" unit="in" value="3.475" valueMax="3.6" valueMin="3.35">3.35-3.60 in</quantity>
females; chela L/W 
<quantity id="DD7951A79274FF82FAFAF9701DBE55F4" box="[1353,1437,1711,1737]" metricMagnitude="-1" metricUnit="m" metricValue="2.02184" pageId="2" pageNumber="93" unit="in" value="7.96">7.96 in</quantity>
male, 7.00- 
<quantity id="DD7951A79274FF82FEAFF90E195255D6" box="[284,369,1745,1771]" metricMagnitude="-1" metricUnit="m" metricValue="1.9304" pageId="2" pageNumber="93" unit="in" value="7.6">7.60 in</quantity>
females; fixed and movable finger of pedipalps with 12-13 rows of granules and marked accessory granules; trichobothrial pattern orthobothriotaxic, 
<typeStatus id="C53A42E09274FF82FCD4F92D1BB95431" box="[871,922,1778,1804]" pageId="2" pageNumber="93">type</typeStatus>
A-β (
<bibRefCitation id="7E1081B39274FF82FC51F92D1CAE5431" author="Vachon, M." box="[994,1165,1778,1804]" pageId="2" pageNumber="93" pagination="857 - 958" refId="ref8801" refString="Vachon, M. (1974) Etude des caracteres utilises pour classer les familles et les genres de Scorpions (Arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les Scorpions. Bulletin du Museum national d'Histoire naturelle, 3 e ser., n ° 140, Zoologie, 104, 857 - 958." type="journal article" year="1974">Vachon, 1974</bibRefCitation>
, 
<bibRefCitation id="7E1081B39274FF82FB28F92D1CF95431" author="Vachon, M." box="[1179,1242,1778,1804]" pageId="2" pageNumber="93" pagination="1597 - 1599" refId="ref8866" refString="Vachon, M. (1975) Sur l'utilisation de la trichobothriotaxie du bras des pedipalpes des Scorpions (Arachnides) dans le classement des genres de la famille des Buthidae Simon. Comptes Rendus des Seances de l'Academie des Sciences, 281 (D), 1597 - 1599." type="journal article" year="1975">1975</bibRefCitation>
); pedipalp chela fixed finger with trichobothrium 
<emphasis id="28F520509274FF82FD9EF8CB1A695413" box="[557,586,1812,1838]" italics="true" pageId="2" pageNumber="93">db</emphasis>
distal to 
<emphasis id="28F520509274FF82FD76F8CB1AC55413" box="[709,742,1812,1838]" italics="true" pageId="2" pageNumber="93">est</emphasis>
; metasoma I–III with 10 carinae, median lateral carinae complete on I, reduced on II–III; metasoma IV with 8 carinae; metasoma V with 7 carinae; metasoma slender, metasoma I L/W 
<quantity id="DD7951A79274FF82FE61F8871A04544F" box="[466,551,1880,1906]" metricMagnitude="-2" metricUnit="m" metricValue="3.4544" pageId="2" pageNumber="93" unit="in" value="1.36">1.36 in</quantity>
male, 
<quantity id="DD7951A79274FF82FDCFF8871B2E544F" box="[636,781,1880,1906]" metricMagnitude="-2" metricUnit="m" metricValue="3.2639" metricValueMax="3.302" metricValueMin="3.2258" pageId="2" pageNumber="93" unit="in" value="1.285" valueMax="1.3" valueMin="1.27">1.27-1.30 in</quantity>
females, metasoma II L/W 
<quantity id="DD7951A79274FF82FBD1F8871C94544F" box="[1122,1207,1880,1906]" metricMagnitude="-2" metricUnit="m" metricValue="4.8514" pageId="2" pageNumber="93" unit="in" value="1.91">1.91 in</quantity>
male, 
<quantity id="DD7951A79274FF82FAB8F8871DBF544F" box="[1291,1436,1880,1906]" metricMagnitude="-2" metricUnit="m" metricValue="4.6228" metricValueMax="4.7244" metricValueMin="4.5212" pageId="2" pageNumber="93" unit="in" value="1.82" valueMax="1.86" valueMin="1.78">1.78-1.86 in</quantity>
females; metasoma III L/W 
<quantity id="DD7951A79274FF82FE50F8A61A1554AE" box="[483,566,1913,1939]" metricMagnitude="-2" metricUnit="m" metricValue="5.4864" pageId="2" pageNumber="93" unit="in" value="2.16">2.16 in</quantity>
male, 
<quantity id="DD7951A79274FF82FD35F8A61B3654AE" box="[646,789,1913,1939]" metricMagnitude="-2" metricUnit="m" metricValue="5.1689" metricValueMax="5.334" metricValueMin="5.0038" pageId="2" pageNumber="93" unit="in" value="2.035" valueMax="2.1" valueMin="1.97">1.97-2.10 in</quantity>
females; metasoma IV L/W 
<quantity id="DD7951A79274FF82FBD8F8A61C9E54AE" box="[1131,1213,1913,1939]" metricMagnitude="-2" metricUnit="m" metricValue="7.1374" pageId="2" pageNumber="93" unit="in" value="2.81">2.81 in</quantity>
male, 
<quantity id="DD7951A79274FF82FABEF8A61DBE54AE" box="[1293,1437,1913,1939]" metricMagnitude="-2" metricUnit="m" metricValue="6.5278" metricValueMax="6.6548" metricValueMin="6.4008" pageId="2" pageNumber="93" unit="in" value="2.57" valueMax="2.62" valueMin="2.52">2.52-2.62 in</quantity>
females; metasoma V L/W 
<quantity id="DD7951A79274FF82FE43F8441A645488" box="[496,583,1947,1973]" metricMagnitude="-2" metricUnit="m" metricValue="8.636" pageId="2" pageNumber="93" unit="in" value="3.4">3.40 in</quantity>
male, 
<quantity id="DD7951A79274FF82FD12F8441B165488" box="[673,821,1947,1973]" metricMagnitude="-2" metricUnit="m" metricValue="7.7216" metricValueMax="7.8994" metricValueMin="7.5438" pageId="2" pageNumber="93" unit="in" value="3.04" valueMax="3.11" valueMin="2.97">2.97-3.11 in</quantity>
females. 
<emphasis id="28F520509274FF82FC02F8441BE05488" box="[945,963,1947,1973]" italics="true" pageId="2" pageNumber="93">F</emphasis>
s (slenderness factor) of adult females ranging from 27.78–32.32.
</paragraph>
</subSubSection>
<subSubSection id="529BAFC99275FF8AFF24FF1B1AEF553F" lastPageId="10" lastPageNumber="101" pageId="3" pageNumber="94" type="description">
<paragraph id="1A3EFC429275FF83FF24FF1B1CB053E3" blockId="3.[151,1171,196,222]" box="[151,1171,196,222]" pageId="3" pageNumber="94">
<emphasis id="28F520509275FF83FF24FF1B191153E3" bold="true" box="[151,306,196,222]" pageId="3" pageNumber="94">Description:</emphasis>
Based on female 
<typeStatus id="C53A42E09275FF83FDB6FF1B1A4853E3" box="[517,619,196,222]" pageId="3" pageNumber="94" type="holotype">holotype</typeStatus>
and 
<typeStatus id="C53A42E09275FF83FD17FF1B1B3B53E3" box="[676,792,196,222]" pageId="3" pageNumber="94" type="paratype">paratypes</typeStatus>
. Measurements are in Table (1).
</paragraph>
<paragraph id="1A3EFC429275FF83FF24FED719445133" blockId="3.[151,1437,264,526]" pageId="3" pageNumber="94">
<emphasis id="28F520509275FF83FF24FED7193C521F" bold="true" box="[151,287,264,290]" pageId="3" pageNumber="94">Coloration</emphasis>
. Generally black (
<figureCitation id="82BAE0C79275FF83FDB6FED71A78521F" box="[517,603,264,290]" captionStart="Figure 1" captionStartId="3.[151,227,1661,1684]" captionTargetBox="[151,1435,539,1643]" captionTargetId="figure-128@3.[151,1435,539,1643]" captionTargetPageId="3" captionText="Figure 1. Leiurus nigellus sp. nov., female paratype from Al Buriakah, Al Ula governorate, Saudi Arabia." figureDoi="http://doi.org/10.5281/zenodo.13247358" httpUri="https://zenodo.org/record/13247358/files/figure.png" pageId="3" pageNumber="94">Figs. 1</figureCitation>
&amp; 
<figureCitation id="82BAE0C79275FF83FD3AFED71AB9521F" box="[649,666,264,290]" captionStart="Figure 2" captionStartId="4.[151,227,1636,1659]" captionTargetBox="[169,1392,218,1607]" captionTargetId="figure-11@4.[152,1436,189,1617]" captionTargetPageId="4" captionText="Figure 2. Habitus of Leiurus nigellus sp. nov., male paratype and female holotype. A. male in dorsal view. B. male in ventral view. C. female in dorsal view. D. female in ventral view. Scale bar = 20 mm." figureDoi="http://doi.org/10.5281/zenodo.13247360" httpUri="https://zenodo.org/record/13247360/files/figure.png" pageId="3" pageNumber="94">2</figureCitation>
), carapace black to blackish-brown; tergites black to blackish-brown; venter yellowish-brown except sternite VII blackish-brown; metasomal segments black to blackish-brown; vesicle black to blackish-brown; aculeus dark reddish-brown at the base and dark black at its extremity; chelicerae retrodorsal surface dark yellow to yellowish-brown; prodorsal surface reticulated with dense dark spots; teeth blackish; pedipalps black to blackish-brown overall excepted for the chela fingers which are gradually turn to yellowish-brown on the distal third; rows of granules on the dentate margins of the fingers dark red; legs with black to blackish-brown proximal segments and the three most distal becoming lighter.
</paragraph>
<caption id="4EFEACCA9275FF83FF24F9A21D2055A9" ID-DOI="http://doi.org/10.5281/zenodo.13247358" ID-Zenodo-Dep="13247358" box="[151,1283,1661,1684]" httpUri="https://zenodo.org/record/13247358/files/figure.png" pageId="3" pageNumber="94" startId="3.[151,227,1661,1684]" targetBox="[151,1435,539,1643]" targetPageId="3" targetType="figure">
<paragraph id="1A3EFC429275FF83FF24F9A21D2055A9" blockId="3.[151,1283,1661,1684]" box="[151,1283,1661,1684]" pageId="3" pageNumber="94">
<emphasis id="28F520509275FF83FF24F9A218D555A9" bold="true" box="[151,246,1661,1684]" pageId="3" pageNumber="94">Figure 1</emphasis>
. 
<taxonomicName id="DD8187C19275FF83FEB7F9A2199355A9" authorityName="Afifeh &amp; Aloufi &amp; Al-Saraireh &amp; Badry &amp; Al-Qahtni &amp; Amr" authorityYear="2023" box="[260,432,1661,1684]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="nigellus" status="sp. nov.">
<emphasis id="28F520509275FF83FEB7F9A2199355A9" box="[260,432,1661,1684]" italics="true" pageId="3" pageNumber="94">Leiurus nigellus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="33C69D2B9275FF83FE04F9A21A2E55A9" box="[439,525,1661,1684]" pageId="3" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
, female paratype from Al Buriakah, Al Ula governorate, Saudi Arabia.
</paragraph>
</caption>
<paragraph id="1A3EFC429275FF83FF24F93F191155C7" blockId="3.[151,1437,1760,1988]" box="[151,306,1760,1786]" pageId="3" pageNumber="94">
<emphasis id="28F520509275FF83FF24F93F191155C7" bold="true" box="[151,306,1760,1786]" pageId="3" pageNumber="94">Morphology</emphasis>
</paragraph>
<paragraph id="1A3EFC429275FF83FF24F8DE1C8F54F9" blockId="3.[151,1437,1760,1988]" pageId="3" pageNumber="94">
<emphasis id="28F520509275FF83FF24F8DE19255426" bold="true" box="[151,262,1793,1819]" pageId="3" pageNumber="94">Prosoma</emphasis>
(
<figureCitation id="82BAE0C79275FF83FEACF8DE195A5426" box="[287,377,1793,1819]" captionStart="Figure 3" captionStartId="6.[151,227,1564,1587]" captionTargetBox="[155,1427,413,1540]" captionTargetId="figure-109@6.[151,1433,404,1545]" captionTargetPageId="6" captionText="Figure 3. Leiurus nigellus sp. nov., female holotype under UV light. A. carapace and mesosoma. B. sternopectinal area and ventral of mesosoma. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13247362" httpUri="https://zenodo.org/record/13247362/files/figure.png" pageId="3" pageNumber="94">Figs. 3</figureCitation>
&amp; 
<figureCitation id="82BAE0C79275FF83FE1CF8DE19E25426" box="[431,449,1793,1819]" captionStart="Figure 4" captionStartId="7.[151,227,1550,1573]" captionTargetBox="[155,1433,275,1532]" captionTargetId="figure-37@7.[151,1433,269,1532]" captionTargetPageId="7" captionText="Figure 4. Leiurus nigellus sp. nov., male paratype under UV light. A. carapace and mesosoma. B. sternopectinal area and ventral of mesosoma. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13247366" httpUri="https://zenodo.org/record/13247366/files/figure.png" pageId="3" pageNumber="94">4</figureCitation>
). Anterior margin of carapace slightly concave; all carapace carinae strongly developed, granulose, including central median, posterior median, anterior median, central lateral and central median; central lateral and posterior median carinae fused to form lyre shaped row of granules. Intercarinal spaces with very few irregular granules and almost smooth laterally and distally; median ocular tubercle only slightly anterior to the centre of the carapace, almost in a central position; median eyes separated by slightly more than two ocular diameters; five pairs of lateral eyes; (3 large, 2 small) on each side.
</paragraph>
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<paragraph id="1A3EFC429272FF84FF24F9BB1C0855A4" blockId="4.[151,1409,1636,1689]" pageId="4" pageNumber="95">
<emphasis id="28F520509272FF84FF24F9BB18D55546" bold="true" box="[151,246,1636,1659]" pageId="4" pageNumber="95">Figure 2</emphasis>
. Habitus of 
<taxonomicName id="DD8187C19272FF84FEC9F9BB1A045546" authorityName="Afifeh &amp; Aloufi &amp; Al-Saraireh &amp; Badry &amp; Al-Qahtni &amp; Amr" authorityYear="2023" box="[378,551,1636,1659]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="nigellus" status="sp. nov.">
<emphasis id="28F520509272FF84FEC9F9BB1A045546" box="[378,551,1636,1659]" italics="true" pageId="4" pageNumber="95">Leiurus nigellus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="33C69D2B9272FF84FD9DF9BB1AA75546" box="[558,644,1636,1659]" pageId="4" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
, male paratype and female holotype. 
<emphasis id="28F520509272FF84FBA0F9BB1C055546" bold="true" box="[1043,1062,1636,1659]" pageId="4" pageNumber="95">A</emphasis>
. male in dorsal view. 
<emphasis id="28F520509272FF84FAA1F9BB1D075546" bold="true" box="[1298,1316,1636,1659]" pageId="4" pageNumber="95">B</emphasis>
. male in ventral view. 
<emphasis id="28F520509272FF84FE9AF95D191F55A4" bold="true" box="[297,316,1666,1689]" pageId="4" pageNumber="95">C</emphasis>
. female in dorsal view. 
<emphasis id="28F520509272FF84FD8EF95D1A7355A4" bold="true" box="[573,592,1666,1689]" pageId="4" pageNumber="95">D</emphasis>
. female in ventral view. Scale bar = 20 mm.
</paragraph>
</caption>
<paragraph id="1A3EFC429272FF86FF24F8D81C0352BA" blockId="4.[151,1437,1799,1993]" lastBlockId="6.[151,1436,196,391]" lastPageId="6" lastPageNumber="97" pageId="4" pageNumber="95">
<emphasis id="28F520509272FF84FF24F8D81938541C" bold="true" box="[151,283,1799,1825]" pageId="4" pageNumber="95">Mesosoma</emphasis>
(
<figureCitation id="82BAE0C79272FF84FE83F8D819A4541C" box="[304,391,1799,1825]" captionStart="Figure 3" captionStartId="6.[151,227,1564,1587]" captionTargetBox="[155,1427,413,1540]" captionTargetId="figure-109@6.[151,1433,404,1545]" captionTargetPageId="6" captionText="Figure 3. Leiurus nigellus sp. nov., female holotype under UV light. A. carapace and mesosoma. B. sternopectinal area and ventral of mesosoma. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13247362" httpUri="https://zenodo.org/record/13247362/files/figure.png" pageId="4" pageNumber="95">Figs. 3</figureCitation>
&amp; 
<figureCitation id="82BAE0C79272FF84FE05F8D819EB541C" box="[438,456,1799,1825]" captionStart="Figure 4" captionStartId="7.[151,227,1550,1573]" captionTargetBox="[155,1433,275,1532]" captionTargetId="figure-37@7.[151,1433,269,1532]" captionTargetPageId="7" captionText="Figure 4. Leiurus nigellus sp. nov., male paratype under UV light. A. carapace and mesosoma. B. sternopectinal area and ventral of mesosoma. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13247366" httpUri="https://zenodo.org/record/13247366/files/figure.png" pageId="4" pageNumber="95">4</figureCitation>
). Tergites I, II and VII pentacarinate; III and IV tricarinate; all carinae strong, granular; median carina on I moderate to strong; on II-VI strong, crenulate; terminating distally on each segment with a spinoid process that extends very slightly beyond the posterior margin of the tergite; tergite VII with 5 strong, granular carinae; inner and outer lateral carinae joined anteriorly by transverse granule rows; median carina present on anterior one-half to 2/3 of the total length, moderate to strong. Intercarinal spaces weakly granular, almost smooth, excepted for the lateral margins of tergites III-VI which are strongly granulated; sternites: lateral carinae absent from sternite III; moderate to strong on sternites IV-VI; strong, crenulate on VII; median carinae on sternite III weak to moderate, irregularly granular; on IV weak; on V weak to obsolete; on VI moderate; on VII strong crenulate. Pectines long; reaching to or slightly beyond coxa-trochanter joint of leg IV in the female, well beyond this articulation in males; pectinal tooth count 
<quantity id="DD7951A79270FF86FAC6FEF618F05258" metricMagnitude="-1" metricUnit="m" metricValue="9.525" metricValueMax="9.906" metricValueMin="9.144" pageId="6" pageNumber="97" unit="in" value="37.5" valueMax="39.0" valueMin="36.0">36- 39 in</quantity>
males and 
<quantity id="DD7951A79270FF86FEEBFE9419995258" box="[344,442,331,357]" metricMagnitude="-1" metricUnit="m" metricValue="7.874" metricValueMax="8.382" metricValueMin="7.366" pageId="6" pageNumber="97" unit="in" value="31.0" valueMax="33.0" valueMin="29.0">29-33 in</quantity>
females. Pectines with 3 marginal lamellae, 9 middle lamellae, marginal and middle lamellae with dense cover of short reddish macrosetae; fulcra with 4–7 setae.
</paragraph>
<paragraph id="1A3EFC429273FF85FD2EFE811003570F" pageId="5" pageNumber="96">
<table id="68810EE29273007FFF3FFE821015570F" box="[140,2102,349,1074]" gridcols="11" gridrows="18" pageId="5" pageNumber="96">
<tr id="A4B1FE009273007FFF3FFE821015524C" box="[140,2102,349,369]" gridrow="0" pageId="5" pageNumber="96" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
<th id="E760977C9273007FFDBFFE821BC3524C" box="[524,992,349,369]" colspan="3" colspanRight="2" gridcol="2" gridrow="0" pageId="5" pageNumber="96">
<taxonomicName id="DD8187C19273FF85FD2EFE811B04524C" authorityName="Afifeh &amp; Aloufi &amp; Al-Saraireh &amp; Badry &amp; Al-Qahtni &amp; Amr" authorityYear="2023" box="[669,807,350,369]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="nigellus" status="sp. nov.">
<emphasis id="28F520509273FF85FD2EFE811B04524C" box="[669,807,350,369]" italics="true" pageId="5" pageNumber="96">Leiurus nigellus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="33C69D2B9273FF85FC9EFE811B52524C" box="[813,881,350,369]" pageId="5" pageNumber="96" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<th id="E760977C9273007FF9ABFE821EF1524C" box="[1560,1746,349,369]" gridcol="8" gridrow="0" pageId="5" pageNumber="96">
<taxonomicName id="DD8187C19273FF85F990FE821EF1524D" authorityName="Lourenco, Modry &amp; Amr" authorityYear="2002" box="[1571,1746,349,368]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis id="28F520509273FF85F990FE821EF1524D" box="[1571,1746,349,368]" italics="true" pageId="5" pageNumber="96">Leiurus jordanensis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="A4B1FE009273007FFF3FFE5B101552DC" box="[140,2102,388,481]" gridrow="1" pageId="5" pageNumber="96">
<th id="E760977C9273007FFF3FFE5B196C52DC" box="[140,335,388,481]" gridcol="0" gridrow="1" pageId="5" pageNumber="96">Measurements</th>
<td id="E760977C9273007FFED6FE5B19CF52DC" box="[357,492,388,481]" gridcol="1" gridrow="1" pageId="5" pageNumber="96">♂ paratype BAPC0251</td>
<td id="E760977C9273007FFDBFFE5B1AB052DC" box="[524,659,388,481]" gridcol="2" gridrow="1" pageId="5" pageNumber="96">♀ holotype BAPC0250</td>
<td id="E760977C9273007FFD00FE5B1B1952DC" box="[691,826,388,481]" gridcol="3" gridrow="1" pageId="5" pageNumber="96">♀ paratype BAPC0252</td>
<td id="E760977C9273007FFCE9FE5B1BC352DC" box="[858,992,388,481]" gridcol="4" gridrow="1" pageId="5" pageNumber="96">♀ paratype BAPC0253</td>
<td id="E760977C9273007FFBBAFE5B1CAC52DC" box="[1033,1167,388,481]" gridcol="5" gridrow="1" pageId="5" pageNumber="96">♀ paratype BAPC0254</td>
<td id="E760977C9273007FFB02FE5B1D1452DC" box="[1201,1335,388,481]" gridcol="6" gridrow="1" pageId="5" pageNumber="96">♂ MNHN</td>
<td id="E760977C9273007FFAE4FE5B1DFD52DC" box="[1367,1502,388,481]" gridcol="7" gridrow="1" pageId="5" pageNumber="96">♀ BAPC0100</td>
<td id="E760977C9273007FF9ABFE5B1EF152DC" box="[1560,1746,388,481]" gridcol="8" gridrow="1" pageId="5" pageNumber="96">♀ BAPC0101</td>
<td id="E760977C9273007FF95BFE5B1F4C52DC" box="[1768,1903,388,481]" gridcol="9" gridrow="1" pageId="5" pageNumber="96">♀ BAPC0102</td>
<td id="E760977C9273007FF81CFE5B101552DC" box="[1967,2102,388,481]" gridcol="10" gridrow="1" pageId="5" pageNumber="96">♀ BAPC0103</td>
</tr>
<tr id="A4B1FE009273007FFF3FFE2810155137" box="[140,2102,503,522]" gridrow="2" pageId="5" pageNumber="96">
<th id="E760977C9273007FFF3FFE28196C5137" box="[140,335,503,522]" gridcol="0" gridrow="2" pageId="5" pageNumber="96">Total length</th>
<td id="E760977C9273007FFED6FE2819CF5137" box="[357,492,503,522]" gridcol="1" gridrow="2" pageId="5" pageNumber="96">
<emphasis id="28F520509273FF85FE23FE2819E35137" bold="true" box="[400,448,503,522]" pageId="5" pageNumber="96">81.25</emphasis>
</td>
<td id="E760977C9273007FFDBFFE281AB05137" box="[524,659,503,522]" gridcol="2" gridrow="2" pageId="5" pageNumber="96">
<emphasis id="28F520509273FF85FD81FE281A4E5137" bold="true" box="[562,621,503,522]" pageId="5" pageNumber="96">104.21</emphasis>
</td>
<td id="E760977C9273007FFD00FE281B195137" box="[691,826,503,522]" gridcol="3" gridrow="2" pageId="5" pageNumber="96">
<emphasis id="28F520509273FF85FD6AFE281B375137" bold="true" box="[729,788,503,522]" pageId="5" pageNumber="96">100.90</emphasis>
</td>
<td id="E760977C9273007FFCE9FE281BC35137" box="[858,992,503,522]" gridcol="4" gridrow="2" pageId="5" pageNumber="96">
<emphasis id="28F520509273FF85FC36FE281B965137" bold="true" box="[901,949,503,522]" pageId="5" pageNumber="96">96.77</emphasis>
</td>
<td id="E760977C9273007FFBBAFE281CAC5137" box="[1033,1167,503,522]" gridcol="5" gridrow="2" pageId="5" pageNumber="96">
<emphasis id="28F520509273FF85FB87FE281C475137" bold="true" box="[1076,1124,503,522]" pageId="5" pageNumber="96">85.80</emphasis>
</td>
<td id="E760977C9273007FFB02FE281D145137" box="[1201,1335,503,522]" gridcol="6" gridrow="2" pageId="5" pageNumber="96">
<emphasis id="28F520509273FF85FB6FFE281D2F5137" bold="true" box="[1244,1292,503,522]" pageId="5" pageNumber="96">95.90</emphasis>
</td>
<td id="E760977C9273007FFAE4FE281DFD5137" box="[1367,1502,503,522]" gridcol="7" gridrow="2" pageId="5" pageNumber="96">
<emphasis id="28F520509273FF85FACEFE281D9B5137" bold="true" box="[1405,1464,503,522]" pageId="5" pageNumber="96">100.93</emphasis>
</td>
<td id="E760977C9273007FF9ABFE281EF15137" box="[1560,1746,503,522]" gridcol="8" gridrow="2" pageId="5" pageNumber="96">
<emphasis id="28F520509273FF85F98EFE281E5B5137" bold="true" box="[1597,1656,503,522]" pageId="5" pageNumber="96">100.34</emphasis>
</td>
<td id="E760977C9273007FF95BFE281F4C5137" box="[1768,1903,503,522]" gridcol="9" gridrow="2" pageId="5" pageNumber="96">
<emphasis id="28F520509273FF85F8A0FE281F605137" bold="true" box="[1811,1859,503,522]" pageId="5" pageNumber="96">98.65</emphasis>
</td>
<td id="E760977C9273007FF81CFE2810155137" box="[1967,2102,503,522]" gridcol="10" gridrow="2" pageId="5" pageNumber="96">
<emphasis id="28F520509273FF85F860FE2810205137" bold="true" box="[2003,2051,503,522]" pageId="5" pageNumber="96">97.09</emphasis>
</td>
</tr>
<tr id="A4B1FE009273007FFF3FFDC410155113" box="[140,2102,539,558]" gridrow="3" pageId="5" pageNumber="96">
<th id="E760977C9273007FFF3FFDC4196C5113" box="[140,335,539,558]" gridcol="0" gridrow="3" pageId="5" pageNumber="96">Carapace L</th>
<td id="E760977C9273007FFED6FDC419CF5113" box="[357,492,539,558]" gridcol="1" gridrow="3" pageId="5" pageNumber="96">8.76</td>
<td id="E760977C9273007FFDBFFDC41AB05113" box="[524,659,539,558]" gridcol="2" gridrow="3" pageId="5" pageNumber="96">11.40</td>
<td id="E760977C9273007FFD00FDC41B195113" box="[691,826,539,558]" gridcol="3" gridrow="3" pageId="5" pageNumber="96">11.16</td>
<td id="E760977C9273007FFCE9FDC41BC35113" box="[858,992,539,558]" gridcol="4" gridrow="3" pageId="5" pageNumber="96">11.16</td>
<td id="E760977C9273007FFBBAFDC41CAC5113" box="[1033,1167,539,558]" gridcol="5" gridrow="3" pageId="5" pageNumber="96">9.00</td>
<td id="E760977C9273007FFB02FDC41D145113" box="[1201,1335,539,558]" gridcol="6" gridrow="3" pageId="5" pageNumber="96">9.70</td>
<td id="E760977C9273007FFAE4FDC41DFD5113" box="[1367,1502,539,558]" gridcol="7" gridrow="3" pageId="5" pageNumber="96">11.34</td>
<td id="E760977C9273007FF9ABFDC41EF15113" box="[1560,1746,539,558]" gridcol="8" gridrow="3" pageId="5" pageNumber="96">11.40</td>
<td id="E760977C9273007FF95BFDC41F4C5113" box="[1768,1903,539,558]" gridcol="9" gridrow="3" pageId="5" pageNumber="96">10.44</td>
<td id="E760977C9273007FF81CFDC410155113" box="[1967,2102,539,558]" gridcol="10" gridrow="3" pageId="5" pageNumber="96">11.16</td>
</tr>
<tr id="A4B1FE009273007FFF3FFD9F1015516E" box="[140,2102,576,595]" gridrow="4" pageId="5" pageNumber="96">
<th id="E760977C9273007FFF3FFD9F196C516E" box="[140,335,576,595]" gridcol="0" gridrow="4" pageId="5" pageNumber="96">Carapace AW/PW</th>
<td id="E760977C9273007FFED6FD9F19CF516E" box="[357,492,576,595]" gridcol="1" gridrow="4" pageId="5" pageNumber="96">5.16/9.48</td>
<td id="E760977C9273007FFDBFFD9F1AB0516E" box="[524,659,576,595]" gridcol="2" gridrow="4" pageId="5" pageNumber="96">6.60/12.73</td>
<td id="E760977C9273007FFD00FD9F1B19516E" box="[691,826,576,595]" gridcol="3" gridrow="4" pageId="5" pageNumber="96">6.84/13.09</td>
<td id="E760977C9273007FFCE9FD9F1BC3516E" box="[858,992,576,595]" gridcol="4" gridrow="4" pageId="5" pageNumber="96">6.60/12.61</td>
<td id="E760977C9273007FFBBAFD9F1CAC516E" box="[1033,1167,576,595]" gridcol="5" gridrow="4" pageId="5" pageNumber="96">5.52/10.68</td>
<td id="E760977C9273007FFB02FD9F1D14516E" box="[1201,1335,576,595]" gridcol="6" gridrow="4" pageId="5" pageNumber="96">7.10/10.80</td>
<td id="E760977C9273007FFAE4FD9F1DFD516E" box="[1367,1502,576,595]" gridcol="7" gridrow="4" pageId="5" pageNumber="96">6.60/12.24</td>
<td id="E760977C9273007FF9ABFD9F1EF1516E" box="[1560,1746,576,595]" gridcol="8" gridrow="4" pageId="5" pageNumber="96">6.84/12.36</td>
<td id="E760977C9273007FF95BFD9F1F4C516E" box="[1768,1903,576,595]" gridcol="9" gridrow="4" pageId="5" pageNumber="96">6.72/11.40</td>
<td id="E760977C9273007FF81CFD9F1015516E" box="[1967,2102,576,595]" gridcol="10" gridrow="4" pageId="5" pageNumber="96">6.72/12.00</td>
</tr>
<tr id="A4B1FE009273007FFF3FFDBA10155145" box="[140,2102,613,632]" gridrow="5" pageId="5" pageNumber="96">
<th id="E760977C9273007FFF3FFDBA196C5145" box="[140,335,613,632]" gridcol="0" gridrow="5" pageId="5" pageNumber="96">Mesosoma L</th>
<td id="E760977C9273007FFED6FDBA19CF5145" box="[357,492,613,632]" gridcol="1" gridrow="5" pageId="5" pageNumber="96">18.37</td>
<td id="E760977C9273007FFDBFFDBA1AB05145" box="[524,659,613,632]" gridcol="2" gridrow="5" pageId="5" pageNumber="96">29.65</td>
<td id="E760977C9273007FFD00FDBA1B195145" box="[691,826,613,632]" gridcol="3" gridrow="5" pageId="5" pageNumber="96">24.01</td>
<td id="E760977C9273007FFCE9FDBA1BC35145" box="[858,992,613,632]" gridcol="4" gridrow="5" pageId="5" pageNumber="96">22.49</td>
<td id="E760977C9273007FFBBAFDBA1CAC5145" box="[1033,1167,613,632]" gridcol="5" gridrow="5" pageId="5" pageNumber="96">22.80</td>
<td id="E760977C9273007FFB02FDBA1D145145" box="[1201,1335,613,632]" gridcol="6" gridrow="5" pageId="5" pageNumber="96">24.80</td>
<td id="E760977C9273007FFAE4FDBA1DFD5145" box="[1367,1502,613,632]" gridcol="7" gridrow="5" pageId="5" pageNumber="96">25.09</td>
<td id="E760977C9273007FF9ABFDBA1EF15145" box="[1560,1746,613,632]" gridcol="8" gridrow="5" pageId="5" pageNumber="96">22.81</td>
<td id="E760977C9273007FF95BFDBA1F4C5145" box="[1768,1903,613,632]" gridcol="9" gridrow="5" pageId="5" pageNumber="96">26.77</td>
<td id="E760977C9273007FF81CFDBA10155145" box="[1967,2102,613,632]" gridcol="10" gridrow="5" pageId="5" pageNumber="96">21.85</td>
</tr>
<tr id="A4B1FE009273007FFF3FFD55101551A0" box="[140,2102,650,669]" gridrow="6" pageId="5" pageNumber="96">
<th id="E760977C9273007FFF3FFD55196C51A0" box="[140,335,650,669]" gridcol="0" gridrow="6" pageId="5" pageNumber="96">Metasoma I L/W</th>
<td id="E760977C9273007FFED6FD5519CF51A0" box="[357,492,650,669]" gridcol="1" gridrow="6" pageId="5" pageNumber="96">6.36/4.68</td>
<td id="E760977C9273007FFDBFFD551AB051A0" box="[524,659,650,669]" gridcol="2" gridrow="6" pageId="5" pageNumber="96">7.66/5.88</td>
<td id="E760977C9273007FFD00FD551B1951A0" box="[691,826,650,669]" gridcol="3" gridrow="6" pageId="5" pageNumber="96">7.80/6.00</td>
<td id="E760977C9273007FFCE9FD551BC351A0" box="[858,992,650,669]" gridcol="4" gridrow="6" pageId="5" pageNumber="96">7.44/5.76</td>
<td id="E760977C9273007FFBBAFD551CAC51A0" box="[1033,1167,650,669]" gridcol="5" gridrow="6" pageId="5" pageNumber="96">6.24/4.92</td>
<td id="E760977C9273007FFB02FD551D1451A0" box="[1201,1335,650,669]" gridcol="6" gridrow="6" pageId="5" pageNumber="96">7.80/5.40</td>
<td id="E760977C9273007FFAE4FD551DFD51A0" box="[1367,1502,650,669]" gridcol="7" gridrow="6" pageId="5" pageNumber="96">7.86/5.64</td>
<td id="E760977C9273007FF9ABFD551EF151A0" box="[1560,1746,650,669]" gridcol="8" gridrow="6" pageId="5" pageNumber="96">8.28/5.88</td>
<td id="E760977C9273007FF95BFD551F4C51A0" box="[1768,1903,650,669]" gridcol="9" gridrow="6" pageId="5" pageNumber="96">7.68/5.40</td>
<td id="E760977C9273007FF81CFD55101551A0" box="[1967,2102,650,669]" gridcol="10" gridrow="6" pageId="5" pageNumber="96">8.04/5.88</td>
</tr>
<tr id="A4B1FE009273007FFF3FFD70101551FF" box="[140,2102,687,706]" gridrow="7" pageId="5" pageNumber="96">
<th id="E760977C9273007FFF3FFD70196C51FF" box="[140,335,687,706]" gridcol="0" gridrow="7" pageId="5" pageNumber="96">Metasoma II L/W</th>
<td id="E760977C9273007FFED6FD7019CF51FF" box="[357,492,687,706]" gridcol="1" gridrow="7" pageId="5" pageNumber="96">7.80/4.08</td>
<td id="E760977C9273007FFDBFFD701AB051FF" box="[524,659,687,706]" gridcol="2" gridrow="7" pageId="5" pageNumber="96">9.18/5.04</td>
<td id="E760977C9273007FFD00FD701B1951FF" box="[691,826,687,706]" gridcol="3" gridrow="7" pageId="5" pageNumber="96">9.36/5.04</td>
<td id="E760977C9273007FFCE9FD701BC351FF" box="[858,992,687,706]" gridcol="4" gridrow="7" pageId="5" pageNumber="96">8.76/4.92</td>
<td id="E760977C9273007FFBBAFD701CAC51FF" box="[1033,1167,687,706]" gridcol="5" gridrow="7" pageId="5" pageNumber="96">7.44/4.02</td>
<td id="E760977C9273007FFB02FD701D1451FF" box="[1201,1335,687,706]" gridcol="6" gridrow="7" pageId="5" pageNumber="96">9.70/4.60</td>
<td id="E760977C9273007FFAE4FD701DFD51FF" box="[1367,1502,687,706]" gridcol="7" gridrow="7" pageId="5" pageNumber="96">9.24/4.68</td>
<td id="E760977C9273007FF9ABFD701EF151FF" box="[1560,1746,687,706]" gridcol="8" gridrow="7" pageId="5" pageNumber="96">9.48/4.80</td>
<td id="E760977C9273007FF95BFD701F4C51FF" box="[1768,1903,687,706]" gridcol="9" gridrow="7" pageId="5" pageNumber="96">8.76/4.56</td>
<td id="E760977C9273007FF81CFD70101551FF" box="[1967,2102,687,706]" gridcol="10" gridrow="7" pageId="5" pageNumber="96">9.36/4.86</td>
</tr>
<tr id="A4B1FE009273007FFF3FFD0C101551DB" box="[140,2102,723,742]" gridrow="8" pageId="5" pageNumber="96">
<th id="E760977C9273007FFF3FFD0C196C51DB" box="[140,335,723,742]" gridcol="0" gridrow="8" pageId="5" pageNumber="96">Metasoma III L/W</th>
<td id="E760977C9273007FFED6FD0C19CF51DB" box="[357,492,723,742]" gridcol="1" gridrow="8" pageId="5" pageNumber="96">8.04/3.72</td>
<td id="E760977C9273007FFDBFFD0C1AB051DB" box="[524,659,723,742]" gridcol="2" gridrow="8" pageId="5" pageNumber="96">9.60/4.62</td>
<td id="E760977C9273007FFD00FD0C1B1951DB" box="[691,826,723,742]" gridcol="3" gridrow="8" pageId="5" pageNumber="96">9.84/4.68</td>
<td id="E760977C9273007FFCE9FD0C1BC351DB" box="[858,992,723,742]" gridcol="4" gridrow="8" pageId="5" pageNumber="96">9.24/4.68</td>
<td id="E760977C9273007FFBBAFD0C1CAC51DB" box="[1033,1167,723,742]" gridcol="5" gridrow="8" pageId="5" pageNumber="96">7.80/3.72</td>
<td id="E760977C9273007FFB02FD0C1D1451DB" box="[1201,1335,723,742]" gridcol="6" gridrow="8" pageId="5" pageNumber="96">9.80/4.20</td>
<td id="E760977C9273007FFAE4FD0C1DFD51DB" box="[1367,1502,723,742]" gridcol="7" gridrow="8" pageId="5" pageNumber="96">9.60/4.32</td>
<td id="E760977C9273007FF9ABFD0C1EF151DB" box="[1560,1746,723,742]" gridcol="8" gridrow="8" pageId="5" pageNumber="96">9.96/4.44</td>
<td id="E760977C9273007FF95BFD0C1F4C51DB" box="[1768,1903,723,742]" gridcol="9" gridrow="8" pageId="5" pageNumber="96">9.36/4.26</td>
<td id="E760977C9273007FF81CFD0C101551DB" box="[1967,2102,723,742]" gridcol="10" gridrow="8" pageId="5" pageNumber="96">9.84/4.44</td>
</tr>
<tr id="A4B1FE009273007FFF3FFD2710155036" box="[140,2102,760,779]" gridrow="9" pageId="5" pageNumber="96">
<th id="E760977C9273007FFF3FFD27196C5036" box="[140,335,760,779]" gridcol="0" gridrow="9" pageId="5" pageNumber="96">Metasoma IV L/W</th>
<td id="E760977C9273007FFED6FD2719CF5036" box="[357,492,760,779]" gridcol="1" gridrow="9" pageId="5" pageNumber="96">9.12/3.24</td>
<td id="E760977C9273007FFDBFFD271AB05036" box="[524,659,760,779]" gridcol="2" gridrow="9" pageId="5" pageNumber="96">10.68/4.08</td>
<td id="E760977C9273007FFD00FD271B195036" box="[691,826,760,779]" gridcol="3" gridrow="9" pageId="5" pageNumber="96">10.92/4.20</td>
<td id="E760977C9273007FFCE9FD271BC35036" box="[858,992,760,779]" gridcol="4" gridrow="9" pageId="5" pageNumber="96">10.44/4.08</td>
<td id="E760977C9273007FFBBAFD271CAC5036" box="[1033,1167,760,779]" gridcol="5" gridrow="9" pageId="5" pageNumber="96">8.76/3.48</td>
<td id="E760977C9273007FFB02FD271D145036" box="[1201,1335,760,779]" gridcol="6" gridrow="9" pageId="5" pageNumber="96">11.10/3.80</td>
<td id="E760977C9273007FFAE4FD271DFD5036" box="[1367,1502,760,779]" gridcol="7" gridrow="9" pageId="5" pageNumber="96">11.04/4.08</td>
<td id="E760977C9273007FF9ABFD271EF15036" box="[1560,1746,760,779]" gridcol="8" gridrow="9" pageId="5" pageNumber="96">11.28/4.20</td>
<td id="E760977C9273007FF95BFD271F4C5036" box="[1768,1903,760,779]" gridcol="9" gridrow="9" pageId="5" pageNumber="96">10.44/3.84</td>
<td id="E760977C9273007FF81CFD2710155036" box="[1967,2102,760,779]" gridcol="10" gridrow="9" pageId="5" pageNumber="96">10.92/3.96</td>
</tr>
<tr id="A4B1FE009273007FFF3FFCC21015500D" box="[140,2102,797,816]" gridrow="10" pageId="5" pageNumber="96">
<th id="E760977C9273007FFF3FFCC2196C500D" box="[140,335,797,816]" gridcol="0" gridrow="10" pageId="5" pageNumber="96">Metasoma V L/W/D</th>
<td id="E760977C9273007FFED6FCC219CF500D" box="[357,492,797,816]" gridcol="1" gridrow="10" pageId="5" pageNumber="96">10.20/3.00/2.88</td>
<td id="E760977C9273007FFDBFFCC21AB0500D" box="[524,659,797,816]" gridcol="2" gridrow="10" pageId="5" pageNumber="96">12.24/3.96/3.72</td>
<td id="E760977C9273007FFD00FCC21B19500D" box="[691,826,797,816]" gridcol="3" gridrow="10" pageId="5" pageNumber="96">12.61/4.08/3.96</td>
<td id="E760977C9273007FFCE9FCC21BC3500D" box="[858,992,797,816]" gridcol="4" gridrow="10" pageId="5" pageNumber="96">12.12/4.08/3.60</td>
<td id="E760977C9273007FFBBAFCC21CAC500D" box="[1033,1167,797,816]" gridcol="5" gridrow="10" pageId="5" pageNumber="96">10.44/3.36/3.00</td>
<td id="E760977C9273007FFB02FCC21D14500D" box="[1201,1335,797,816]" gridcol="6" gridrow="10" pageId="5" pageNumber="96">12.70/3.40/3.20</td>
<td id="E760977C9273007FFAE4FCC21DFD500D" box="[1367,1502,797,816]" gridcol="7" gridrow="10" pageId="5" pageNumber="96">12.48/3.84/3.48</td>
<td id="E760977C9273007FF9ABFCC21EF1500D" box="[1560,1746,797,816]" gridcol="8" gridrow="10" pageId="5" pageNumber="96">12.61/3.96/3.72</td>
<td id="E760977C9273007FF95BFCC21F4C500D" box="[1768,1903,797,816]" gridcol="9" gridrow="10" pageId="5" pageNumber="96">11.64/3.72/3.36</td>
<td id="E760977C9273007FF81CFCC21015500D" box="[1967,2102,797,816]" gridcol="10" gridrow="10" pageId="5" pageNumber="96">12.12/3.66/3.48</td>
</tr>
<tr id="A4B1FE009273007FFF3FFC9D10155068" box="[140,2102,834,853]" gridrow="11" pageId="5" pageNumber="96">
<th id="E760977C9273007FFF3FFC9D196C5068" box="[140,335,834,853]" gridcol="0" gridrow="11" pageId="5" pageNumber="96">Telson L</th>
<td id="E760977C9273007FFED6FC9D19CF5068" box="[357,492,834,853]" gridcol="1" gridrow="11" pageId="5" pageNumber="96">9.00</td>
<td id="E760977C9273007FFDBFFC9D1AB05068" box="[524,659,834,853]" gridcol="2" gridrow="11" pageId="5" pageNumber="96">10.80</td>
<td id="E760977C9273007FFD00FC9D1B195068" box="[691,826,834,853]" gridcol="3" gridrow="11" pageId="5" pageNumber="96">11.16</td>
<td id="E760977C9273007FFCE9FC9D1BC35068" box="[858,992,834,853]" gridcol="4" gridrow="11" pageId="5" pageNumber="96">11.52</td>
<td id="E760977C9273007FFBBAFC9D1CAC5068" box="[1033,1167,834,853]" gridcol="5" gridrow="11" pageId="5" pageNumber="96">9.36</td>
<td id="E760977C9273007FFB02FC9D1D145068" box="[1201,1335,834,853]" gridcol="6" gridrow="11" pageId="5" pageNumber="96">10.30</td>
<td id="E760977C9273007FFAE4FC9D1DFD5068" box="[1367,1502,834,853]" gridcol="7" gridrow="11" pageId="5" pageNumber="96">11.28</td>
<td id="E760977C9273007FF9ABFC9D1EF15068" box="[1560,1746,834,853]" gridcol="8" gridrow="11" pageId="5" pageNumber="96">11.52</td>
<td id="E760977C9273007FF95BFC9D1F4C5068" box="[1768,1903,834,853]" gridcol="9" gridrow="11" pageId="5" pageNumber="96">10.56</td>
<td id="E760977C9273007FF81CFC9D10155068" box="[1967,2102,834,853]" gridcol="10" gridrow="11" pageId="5" pageNumber="96">10.80</td>
</tr>
<tr id="A4B1FE009273007FFF3FFCB810155047" box="[140,2102,871,890]" gridrow="12" pageId="5" pageNumber="96">
<th id="E760977C9273007FFF3FFCB8196C5047" box="[140,335,871,890]" gridcol="0" gridrow="12" pageId="5" pageNumber="96">Vesicle W/D</th>
<td id="E760977C9273007FFED6FCB819CF5047" box="[357,492,871,890]" gridcol="1" gridrow="12" pageId="5" pageNumber="96">3.12/3.00</td>
<td id="E760977C9273007FFDBFFCB81AB05047" box="[524,659,871,890]" gridcol="2" gridrow="12" pageId="5" pageNumber="96">4.08/3.96</td>
<td id="E760977C9273007FFD00FCB81B195047" box="[691,826,871,890]" gridcol="3" gridrow="12" pageId="5" pageNumber="96">3.96/3.84</td>
<td id="E760977C9273007FFCE9FCB81BC35047" box="[858,992,871,890]" gridcol="4" gridrow="12" pageId="5" pageNumber="96">4.44/3.96</td>
<td id="E760977C9273007FFBBAFCB81CAC5047" box="[1033,1167,871,890]" gridcol="5" gridrow="12" pageId="5" pageNumber="96">3.48/3.24</td>
<td id="E760977C9273007FFB02FCB81D145047" box="[1201,1335,871,890]" gridcol="6" gridrow="12" pageId="5" pageNumber="96">3.50/3.70</td>
<td id="E760977C9273007FFAE4FCB81DFD5047" box="[1367,1502,871,890]" gridcol="7" gridrow="12" pageId="5" pageNumber="96">3.96/3.84</td>
<td id="E760977C9273007FF9ABFCB81EF15047" box="[1560,1746,871,890]" gridcol="8" gridrow="12" pageId="5" pageNumber="96">4.44/4.20</td>
<td id="E760977C9273007FF95BFCB81F4C5047" box="[1768,1903,871,890]" gridcol="9" gridrow="12" pageId="5" pageNumber="96">3.84/3.66</td>
<td id="E760977C9273007FF81CFCB810155047" box="[1967,2102,871,890]" gridcol="10" gridrow="12" pageId="5" pageNumber="96">3.84/3.72</td>
</tr>
<tr id="A4B1FE009273007FFF3FFC54101550A3" box="[140,2102,907,926]" gridrow="13" pageId="5" pageNumber="96">
<th id="E760977C9273007FFF3FFC54196C50A3" box="[140,335,907,926]" gridcol="0" gridrow="13" pageId="5" pageNumber="96">Pedipalp Femur L/W</th>
<td id="E760977C9273007FFED6FC5419CF50A3" box="[357,492,907,926]" gridcol="1" gridrow="13" pageId="5" pageNumber="96">10.20/2.40</td>
<td id="E760977C9273007FFDBFFC541AB050A3" box="[524,659,907,926]" gridcol="2" gridrow="13" pageId="5" pageNumber="96">11.40/3.00</td>
<td id="E760977C9273007FFD00FC541B1950A3" box="[691,826,907,926]" gridcol="3" gridrow="13" pageId="5" pageNumber="96">12.30/3.12</td>
<td id="E760977C9273007FFCE9FC541BC350A3" box="[858,992,907,926]" gridcol="4" gridrow="13" pageId="5" pageNumber="96">11.64/2.94</td>
<td id="E760977C9273007FFBBAFC541CAC50A3" box="[1033,1167,907,926]" gridcol="5" gridrow="13" pageId="5" pageNumber="96">9.84/2.52</td>
<td id="E760977C9273007FFB02FC541D1450A3" box="[1201,1335,907,926]" gridcol="6" gridrow="13" pageId="5" pageNumber="96">11.40/2.40</td>
<td id="E760977C9273007FFAE4FC541DFD50A3" box="[1367,1502,907,926]" gridcol="7" gridrow="13" pageId="5" pageNumber="96">11.76/2.88</td>
<td id="E760977C9273007FF9ABFC541EF150A3" box="[1560,1746,907,926]" gridcol="8" gridrow="13" pageId="5" pageNumber="96">12.61/2.88</td>
<td id="E760977C9273007FF95BFC541F4C50A3" box="[1768,1903,907,926]" gridcol="9" gridrow="13" pageId="5" pageNumber="96">11.40/2.70</td>
<td id="E760977C9273007FF81CFC54101550A3" box="[1967,2102,907,926]" gridcol="10" gridrow="13" pageId="5" pageNumber="96">11.64/2.88</td>
</tr>
<tr id="A4B1FE009273007FFF3FFC6F101550FE" box="[140,2102,944,963]" gridrow="14" pageId="5" pageNumber="96">
<th id="E760977C9273007FFF3FFC6F196C50FE" box="[140,335,944,963]" gridcol="0" gridrow="14" pageId="5" pageNumber="96">Pedipalp Patella L/W</th>
<td id="E760977C9273007FFED6FC6F19CF50FE" box="[357,492,944,963]" gridcol="1" gridrow="14" pageId="5" pageNumber="96">11.04/2.76</td>
<td id="E760977C9273007FFDBFFC6F1AB050FE" box="[524,659,944,963]" gridcol="2" gridrow="14" pageId="5" pageNumber="96">12.73/3.60</td>
<td id="E760977C9273007FFD00FC6F1B1950FE" box="[691,826,944,963]" gridcol="3" gridrow="14" pageId="5" pageNumber="96">13.57/3.84</td>
<td id="E760977C9273007FFCE9FC6F1BC350FE" box="[858,992,944,963]" gridcol="4" gridrow="14" pageId="5" pageNumber="96">12.48/3.72</td>
<td id="E760977C9273007FFBBAFC6F1CAC50FE" box="[1033,1167,944,963]" gridcol="5" gridrow="14" pageId="5" pageNumber="96">10.80/3.00</td>
<td id="E760977C9273007FFB02FC6F1D1450FE" box="[1201,1335,944,963]" gridcol="6" gridrow="14" pageId="5" pageNumber="96">13.10/2.90</td>
<td id="E760977C9273007FFAE4FC6F1DFD50FE" box="[1367,1502,944,963]" gridcol="7" gridrow="14" pageId="5" pageNumber="96">13.09/3.36</td>
<td id="E760977C9273007FF9ABFC6F1EF150FE" box="[1560,1746,944,963]" gridcol="8" gridrow="14" pageId="5" pageNumber="96">13.81/3.48</td>
<td id="E760977C9273007FF95BFC6F1F4C50FE" box="[1768,1903,944,963]" gridcol="9" gridrow="14" pageId="5" pageNumber="96">12.48/3.36</td>
<td id="E760977C9273007FF81CFC6F101550FE" box="[1967,2102,944,963]" gridcol="10" gridrow="14" pageId="5" pageNumber="96">12.97/3.36</td>
</tr>
<tr id="A4B1FE009273007FFF3FFC0A101550D5" box="[140,2102,981,1000]" gridrow="15" pageId="5" pageNumber="96">
<th id="E760977C9273007FFF3FFC0A196C50D5" box="[140,335,981,1000]" gridcol="0" gridrow="15" pageId="5" pageNumber="96">Pedipalp Chela L/W/D</th>
<td id="E760977C9273007FFED6FC0A19CF50D5" box="[357,492,981,1000]" gridcol="1" gridrow="15" pageId="5" pageNumber="96">20.05/2.52/2.64</td>
<td id="E760977C9273007FFDBFFC0A1AB050D5" box="[524,659,981,1000]" gridcol="2" gridrow="15" pageId="5" pageNumber="96">23.65/3.18/3.78</td>
<td id="E760977C9273007FFD00FC0A1B1950D5" box="[691,826,981,1000]" gridcol="3" gridrow="15" pageId="5" pageNumber="96">24.61/3.24/3.84</td>
<td id="E760977C9273007FFCE9FC0A1BC350D5" box="[858,992,981,1000]" gridcol="4" gridrow="15" pageId="5" pageNumber="96">22.69/3.24/3.72</td>
<td id="E760977C9273007FFBBAFC0A1CAC50D5" box="[1033,1167,981,1000]" gridcol="5" gridrow="15" pageId="5" pageNumber="96">20.05/2.64/3.00</td>
<td id="E760977C9273007FFB02FC0A1D1450D5" box="[1201,1335,981,1000]" gridcol="6" gridrow="15" pageId="5" pageNumber="96">21.60/2.40/2.70</td>
<td id="E760977C9273007FFAE4FC0A1DFD50D5" box="[1367,1502,981,1000]" gridcol="7" gridrow="15" pageId="5" pageNumber="96">23.05/3.00/3.60</td>
<td id="E760977C9273007FF9ABFC0A1EF150D5" box="[1560,1746,981,1000]" gridcol="8" gridrow="15" pageId="5" pageNumber="96">24.25/3.12/3.60</td>
<td id="E760977C9273007FF95BFC0A1F4C50D5" box="[1768,1903,981,1000]" gridcol="9" gridrow="15" pageId="5" pageNumber="96">22.57/2.88/3.12</td>
<td id="E760977C9273007FF81CFC0A101550D5" box="[1967,2102,981,1000]" gridcol="10" gridrow="15" pageId="5" pageNumber="96">22.57/2.883.48</td>
</tr>
<tr id="A4B1FE009273007FFF3FFC2510155730" box="[140,2102,1018,1037]" gridrow="16" pageId="5" pageNumber="96">
<th id="E760977C9273007FFF3FFC25196C5730" box="[140,335,1018,1037]" gridcol="0" gridrow="16" pageId="5" pageNumber="96">Movable finger L</th>
<td id="E760977C9273007FFED6FC2519CF5730" box="[357,492,1018,1037]" gridcol="1" gridrow="16" pageId="5" pageNumber="96">13.45</td>
<td id="E760977C9273007FFDBFFC251AB05730" box="[524,659,1018,1037]" gridcol="2" gridrow="16" pageId="5" pageNumber="96">16.57</td>
<td id="E760977C9273007FFD00FC251B195730" box="[691,826,1018,1037]" gridcol="3" gridrow="16" pageId="5" pageNumber="96">16.69</td>
<td id="E760977C9273007FFCE9FC251BC35730" box="[858,992,1018,1037]" gridcol="4" gridrow="16" pageId="5" pageNumber="96">15.49</td>
<td id="E760977C9273007FFBBAFC251CAC5730" box="[1033,1167,1018,1037]" gridcol="5" gridrow="16" pageId="5" pageNumber="96">13.45</td>
<td id="E760977C9273007FFB02FC251D145730" box="[1201,1335,1018,1037]" gridcol="6" gridrow="16" pageId="5" pageNumber="96">15.20</td>
<td id="E760977C9273007FFAE4FC251DFD5730" box="[1367,1502,1018,1037]" gridcol="7" gridrow="16" pageId="5" pageNumber="96">15.37</td>
<td id="E760977C9273007FF9ABFC251EF15730" box="[1560,1746,1018,1037]" gridcol="8" gridrow="16" pageId="5" pageNumber="96">16.45</td>
<td id="E760977C9273007FF95BFC251F4C5730" box="[1768,1903,1018,1037]" gridcol="9" gridrow="16" pageId="5" pageNumber="96">15.37</td>
<td id="E760977C9273007FF81CFC2510155730" box="[1967,2102,1018,1037]" gridcol="10" gridrow="16" pageId="5" pageNumber="96">15.01</td>
</tr>
<tr id="A4B1FE009273007FFF3FFBC01015570F" box="[140,2102,1055,1074]" gridrow="17" pageId="5" pageNumber="96" rowspan-1="1" rowspan-6="1">
<th id="E760977C9273007FFF3FFBC0196C570F" box="[140,335,1055,1074]" gridcol="0" gridrow="17" pageId="5" pageNumber="96">Leg III patella L/D ♀</th>
<td id="E760977C9273007FFDBFFBC01AB0570F" box="[524,659,1055,1074]" gridcol="2" gridrow="17" pageId="5" pageNumber="96">9.60/2.28</td>
<td id="E760977C9273007FFD00FBC01B19570F" box="[691,826,1055,1074]" gridcol="3" gridrow="17" pageId="5" pageNumber="96">10.20/2.34</td>
<td id="E760977C9273007FFCE9FBC01BC3570F" box="[858,992,1055,1074]" gridcol="4" gridrow="17" pageId="5" pageNumber="96">9.60/2.28</td>
<td id="E760977C9273007FFBBAFBC01CAC570F" box="[1033,1167,1055,1074]" gridcol="5" gridrow="17" pageId="5" pageNumber="96">8.16/1.92</td>
<td id="E760977C9273007FFAE4FBC01DFD570F" box="[1367,1502,1055,1074]" gridcol="7" gridrow="17" pageId="5" pageNumber="96">10.08/2.22</td>
<td id="E760977C9273007FF9ABFBC01EF1570F" box="[1560,1746,1055,1074]" gridcol="8" gridrow="17" pageId="5" pageNumber="96">10.56/2.28</td>
<td id="E760977C9273007FF95BFBC01F4C570F" box="[1768,1903,1055,1074]" gridcol="9" gridrow="17" pageId="5" pageNumber="96">9.96/2.10</td>
<td id="E760977C9273007FF81CFBC01015570F" box="[1967,2102,1055,1074]" gridcol="10" gridrow="17" pageId="5" pageNumber="96">10.20/2.16</td>
</tr>
</table>
</paragraph>
<caption id="4EFEACCA9270FF86FF24F9C31A57556C" ID-DOI="http://doi.org/10.5281/zenodo.13247362" ID-Zenodo-Dep="13247362" httpUri="https://zenodo.org/record/13247362/files/figure.png" pageId="6" pageNumber="97" startId="6.[151,227,1564,1587]" targetBox="[155,1427,413,1540]" targetPageId="6" targetType="figure">
<paragraph id="1A3EFC429270FF86FF24F9C31A57556C" blockId="6.[151,1436,1564,1617]" pageId="6" pageNumber="97">
<emphasis id="28F520509270FF86FF24F9C318D4550E" bold="true" box="[151,247,1564,1587]" pageId="6" pageNumber="97">Figure 3</emphasis>
. 
<taxonomicName id="DD8187C19270FF86FEB6F9C31990550E" authorityName="Afifeh &amp; Aloufi &amp; Al-Saraireh &amp; Badry &amp; Al-Qahtni &amp; Amr" authorityYear="2023" box="[261,435,1564,1587]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="97" phylum="Arthropoda" rank="species" species="nigellus" status="sp. nov.">
<emphasis id="28F520509270FF86FEB6F9C31990550E" box="[261,435,1564,1587]" italics="true" pageId="6" pageNumber="97">Leiurus nigellus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="33C69D2B9270FF86FE09F9C31A32550E" box="[442,529,1564,1587]" pageId="6" pageNumber="97" rank="species">sp. nov.</taxonomicNameLabel>
, female holotype under UV light. 
<emphasis id="28F520509270FF86FC32F9C31BB7550E" bold="true" box="[897,916,1564,1587]" pageId="6" pageNumber="97">A</emphasis>
. carapace and mesosoma. 
<emphasis id="28F520509270FF86FB00F9C31CE6550E" bold="true" box="[1203,1221,1564,1587]" pageId="6" pageNumber="97">B</emphasis>
. sternopectinal area and ventral of mesosoma. Scale bar = 5 mm.
</paragraph>
</caption>
<paragraph id="1A3EFC429270FF87FF24F9421D13523D" blockId="6.[151,1437,1693,1989]" lastBlockId="7.[151,1436,196,256]" lastPageId="7" lastPageNumber="98" pageId="6" pageNumber="97">
<emphasis id="28F520509270FF86FF24F9421939558A" bold="true" box="[151,282,1693,1719]" pageId="6" pageNumber="97">Metasoma</emphasis>
(
<figureCitation id="82BAE0C79270FF86FE9DF9421958558A" box="[302,379,1693,1719]" captionStart="Figure 5" captionStartId="8.[151,227,1265,1288]" captionTargetBox="[152,1432,191,1244]" captionTargetId="figure-11@8.[151,1433,189,1246]" captionTargetPageId="8" captionText="Figure 5. Leiurus nigellus sp. nov. female holotype under UV light. A. metasoma and telson dorsal view. B. metasoma and telson ventral view. C. metasoma and telson lateral view. Male paratype under UV light. D. metasoma and telson dorsal view. E. metasoma and telson ventral view. F. metasoma and telson lateral view. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13247368" httpUri="https://zenodo.org/record/13247368/files/figure.png" pageId="6" pageNumber="97">Fig. 5</figureCitation>
). Metasomal segments I to III with 10 carinae, crenulate; median lateral carinae on I moderate to strong, crenulate; on II present on posterior one-third, crenulate; on III limited to posterior onefifth; IV with 8 carinae; dorsosubmedian carinae moderate on I–II, weak on III–IV; dorsolateral and ventrolateral carinae moderate on I–IV; ventromedian carinae moderate on segments I to IV. Segment V with 7 carinae; dorsolateral carinae very weak, faintly granulated, ventrolateral carinae strong with dentate granules increasing in size posteriorly, with several slightly enlarged subtriangular denticles, ventrosubmedian carinae marked by series of small to medium dentate granules on anterior 2/3 of segment, ventromedian carina strong, armed with small to large more spiniod than lobate granules; anal arch with 3 lateral spinoid lobes and 12-13 irregular, transverse crenulations on ventral margin. Dorsal furrows of all segments moderately to weakly developed with a thin granulation, almost smooth; intercarinal surfaces smooth on segment I, smooth to minutely shagreened on II-III, sparsely, finely shagreened on IV-V.
</paragraph>
<caption id="4EFEACCA9271FF87FF24F9D11A575579" ID-DOI="http://doi.org/10.5281/zenodo.13247366" ID-Zenodo-Dep="13247366" httpUri="https://zenodo.org/record/13247366/files/figure.png" pageId="7" pageNumber="98" startId="7.[151,227,1550,1573]" targetBox="[155,1433,275,1532]" targetPageId="7" targetType="figure">
<paragraph id="1A3EFC429271FF87FF24F9D11A575579" blockId="7.[151,1437,1550,1604]" pageId="7" pageNumber="98">
<emphasis id="28F520509271FF87FF24F9D118DB5518" bold="true" box="[151,248,1550,1573]" pageId="7" pageNumber="98">Figure 4</emphasis>
. 
<taxonomicName id="DD8187C19271FF87FEBBF9D119945518" authorityName="Afifeh &amp; Aloufi &amp; Al-Saraireh &amp; Badry &amp; Al-Qahtni &amp; Amr" authorityYear="2023" box="[264,439,1550,1573]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="98" phylum="Arthropoda" rank="species" species="nigellus" status="sp. nov.">
<emphasis id="28F520509271FF87FEBBF9D119945518" box="[264,439,1550,1573]" italics="true" pageId="7" pageNumber="98">Leiurus nigellus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="33C69D2B9271FF87FE0CF9D11A345518" box="[447,535,1550,1573]" pageId="7" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
, male paratype under UV light. 
<emphasis id="28F520509271FF87FCCAF9D11BAF5518" bold="true" box="[889,908,1550,1573]" pageId="7" pageNumber="98">A</emphasis>
. carapace and mesosoma. 
<emphasis id="28F520509271FF87FB02F9D11CE05518" bold="true" box="[1201,1219,1550,1573]" pageId="7" pageNumber="98">B</emphasis>
. sternopectinal area and ventral of mesosoma. Scale bar = 5 mm.
</paragraph>
</caption>
<paragraph id="1A3EFC429271FF87FF24F94F1C3A55F6" blockId="7.[151,1437,1680,1739]" pageId="7" pageNumber="98">
<emphasis id="28F520509271FF87FF24F94F18C85597" bold="true" box="[151,235,1680,1706]" pageId="7" pageNumber="98">Telson</emphasis>
(
<figureCitation id="82BAE0C79271FF87FF4DF94F19645597" box="[254,327,1680,1706]" captionStart="Figure 5" captionStartId="8.[151,227,1265,1288]" captionTargetBox="[152,1432,191,1244]" captionTargetId="figure-11@8.[151,1433,189,1246]" captionTargetPageId="8" captionText="Figure 5. Leiurus nigellus sp. nov. female holotype under UV light. A. metasoma and telson dorsal view. B. metasoma and telson ventral view. C. metasoma and telson lateral view. Male paratype under UV light. D. metasoma and telson dorsal view. E. metasoma and telson ventral view. F. metasoma and telson lateral view. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13247368" httpUri="https://zenodo.org/record/13247368/files/figure.png" pageId="7" pageNumber="98">Fig. 5</figureCitation>
). Vesicle smooth, bulbous; ventral surface with scattered fine microsetae and several short macrosetae; aculeus slightly shorter than vesicle, subaculear tubercle absent.
</paragraph>
<paragraph id="1A3EFC429271FF87FF24F92A1D1854E4" blockId="7.[151,1436,1781,2009]" pageId="7" pageNumber="98">
<emphasis id="28F520509271FF87FF24F92A19055432" bold="true" box="[151,294,1781,1807]" pageId="7" pageNumber="98">Chelicerae:</emphasis>
Dorsal surface of manus smooth, convex, prodorsal margin finely granular; retrodorsal surfaces smooth; prolateral and ventral surfaces densely setose; fingers with normal buthid dentition (Vachon, 1963), fixed finger dorsal and ventral surfaces densely setose, dorsal margin bears 4 teeth: distal, subdistal, median, and basal; ventral margin with basal and median denticles; movable finger dorsal surface smooth; ventral surface densely setose; dorsal margin bears 5 teeth: distal, subdistal, median, and a pair of basal denticles fused in bicusp; ventral margin with distal, median, and basal teeth. The movable finger always ends in two distal teeth–one dorsal and one ventral– between which is inserted the distal tooth of the fixed finger.
</paragraph>
<caption id="4EFEACCA927EFF88FF24FB2E1D2A5678" ID-DOI="http://doi.org/10.5281/zenodo.13247368" ID-Zenodo-Dep="13247368" httpUri="https://zenodo.org/record/13247368/files/figure.png" pageId="8" pageNumber="99" startId="8.[151,227,1265,1288]" targetBox="[152,1432,191,1244]" targetPageId="8" targetType="figure">
<paragraph id="1A3EFC42927EFF88FF24FB2E1D2A5678" blockId="8.[151,1436,1265,1349]" pageId="8" pageNumber="99">
<emphasis id="28F52050927EFF88FF24FB2E18DE5635" bold="true" box="[151,253,1265,1288]" pageId="8" pageNumber="99">Figure 5.</emphasis>
<taxonomicName id="DD8187C1927EFF88FEB6FB2E19915635" authorityName="Afifeh &amp; Aloufi &amp; Al-Saraireh &amp; Badry &amp; Al-Qahtni &amp; Amr" authorityYear="2023" box="[261,434,1265,1288]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="99" phylum="Arthropoda" rank="species" species="nigellus" status="sp. nov.">
<emphasis id="28F52050927EFF88FEB6FB2E19915635" box="[261,434,1265,1288]" italics="true" pageId="8" pageNumber="99">Leiurus nigellus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="33C69D2B927EFF88FE09FB2E1A2C5635" box="[442,527,1265,1288]" pageId="8" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
female holotype under UV light. 
<emphasis id="28F52050927EFF88FCCFFB2E1BAC5635" bold="true" box="[892,911,1265,1288]" pageId="8" pageNumber="99">A</emphasis>
. metasoma and telson dorsal view. 
<emphasis id="28F52050927EFF88FAA3FB2E1D015635" bold="true" box="[1296,1314,1265,1288]" pageId="8" pageNumber="99">B</emphasis>
. metasoma and telson ventral view. 
<emphasis id="28F52050927EFF88FE10FAD01995561B" bold="true" box="[419,438,1295,1318]" pageId="8" pageNumber="99">C</emphasis>
. metasoma and telson lateral view. Male paratype under UV light. 
<emphasis id="28F52050927EFF88FB24FAD01C89561B" bold="true" box="[1175,1194,1295,1318]" pageId="8" pageNumber="99">D</emphasis>
. metasoma and telson dorsal view. 
<emphasis id="28F52050927EFF88FE93FAF119125678" bold="true" box="[288,305,1326,1349]" pageId="8" pageNumber="99">E</emphasis>
. metasoma and telson ventral view. 
<emphasis id="28F52050927EFF88FD04FAF11AE45678" bold="true" box="[695,711,1326,1349]" pageId="8" pageNumber="99">F</emphasis>
. metasoma and telson lateral view. Scale bar = 5 mm.
</paragraph>
</caption>
<paragraph id="1A3EFC42927EFF88FF24FA4D19325691" blockId="8.[151,1437,1426,1586]" box="[151,273,1426,1452]" pageId="8" pageNumber="99">
<emphasis id="28F52050927EFF88FF24FA4D19325691" bold="true" box="[151,273,1426,1452]" pageId="8" pageNumber="99">Pedipalps</emphasis>
</paragraph>
<paragraph id="1A3EFC42927EFF88FF24FA6C1C71550F" blockId="8.[151,1437,1426,1586]" pageId="8" pageNumber="99">
<emphasis id="28F52050927EFF88FF24FA6C18CF56F0" bold="true" box="[151,236,1459,1485]" pageId="8" pageNumber="99">Femur</emphasis>
(
<figureCitation id="82BAE0C7927EFF88FEB4FA6C194056F0" box="[263,355,1459,1485]" captionStart="Figure 6" captionStartId="9.[151,227,816,839]" captionTargetBox="[154,1420,200,775]" captionTargetId="figure-9@9.[151,1435,189,798]" captionTargetPageId="9" captionText="Figure 6. Leiurus nigellus sp. nov., pedipalp segments, female holotype (A-F), A. femur internal view. B. femur dorsal view. C. patella dorsal view. D. patella external view. E. chela external view. F. chela ventral view; male paratype (G- L), G. femur internal view. H. femur dorsal view. I. patella dorsal view. J. patella external view. K. chela external view. L. chela ventral view. Trichobothrial pattern indicated by yellow circles. Scale = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13247370" httpUri="https://zenodo.org/record/13247370/files/figure.png" pageId="8" pageNumber="99">Figs. 6</figureCitation>
A-B, G-H) slender, L/W= 3.8; with five carinae; dorsoexternal, dorsointernal and ventrointernal carinae strong with regular large conical granules; internal carina strong, with irregular large granules; external carina obsolete, a smooth ridge with isolated large dentate granules; dorsal and internal surfaces finely, sparsely shagreened, ventral and external surfaces nearly smooth.
</paragraph>
<paragraph id="1A3EFC42927EFF88FF24F9841C9D55C1" blockId="8.[151,1437,1627,1788]" pageId="8" pageNumber="99">
<emphasis id="28F52050927EFF88FF24F98418CD5548" bold="true" box="[151,238,1627,1653]" pageId="8" pageNumber="99">Patella</emphasis>
(
<figureCitation id="82BAE0C7927EFF88FEB3F98419775548" box="[256,340,1627,1653]" captionStart="Figure 6" captionStartId="9.[151,227,816,839]" captionTargetBox="[154,1420,200,775]" captionTargetId="figure-9@9.[151,1435,189,798]" captionTargetPageId="9" captionText="Figure 6. Leiurus nigellus sp. nov., pedipalp segments, female holotype (A-F), A. femur internal view. B. femur dorsal view. C. patella dorsal view. D. patella external view. E. chela external view. F. chela ventral view; male paratype (G- L), G. femur internal view. H. femur dorsal view. I. patella dorsal view. J. patella external view. K. chela external view. L. chela ventral view. Trichobothrial pattern indicated by yellow circles. Scale = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13247370" httpUri="https://zenodo.org/record/13247370/files/figure.png" pageId="8" pageNumber="99">Figs. 6</figureCitation>
C-D, I-J) slender, L/W= 3.8; with seven carinae; dorsointernal carinae moderate to strong, with one conspicuous spinoid granule and several smaller granules; dorsomedian carina weak, finely granular; dorsoexternal, external and ventroexternal carinae weak, smooth; ventromedian carina weak, with fine granules; ventrointernal carina weak, with well-spaced medium to small granules and ventral patellar spur; internal carina moderate, with closely spaced small granules and dorsal patellar spur.
</paragraph>
<paragraph id="1A3EFC42927EFF88FF24F8F91A4954BE" blockId="8.[151,1436,1830,1923]" pageId="8" pageNumber="99">
<emphasis id="28F52050927EFF88FF24F8F918C2547D" bold="true" box="[151,225,1830,1856]" pageId="8" pageNumber="99">Chela</emphasis>
(
<figureCitation id="82BAE0C7927EFF88FF47F8F9196B547D" box="[244,328,1830,1856]" captionStart="Figure 6" captionStartId="9.[151,227,816,839]" captionTargetBox="[154,1420,200,775]" captionTargetId="figure-9@9.[151,1435,189,798]" captionTargetPageId="9" captionText="Figure 6. Leiurus nigellus sp. nov., pedipalp segments, female holotype (A-F), A. femur internal view. B. femur dorsal view. C. patella dorsal view. D. patella external view. E. chela external view. F. chela ventral view; male paratype (G- L), G. femur internal view. H. femur dorsal view. I. patella dorsal view. J. patella external view. K. chela external view. L. chela ventral view. Trichobothrial pattern indicated by yellow circles. Scale = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13247370" httpUri="https://zenodo.org/record/13247370/files/figure.png" pageId="8" pageNumber="99">Figs. 6</figureCitation>
E-F, K-L). slender, L/W=7.44; with elongated fingers; all carinae almost vestigial. Dentate margins of fixed and movable fingers composed of 12-13 almost linear rows of granules and conspicuous accessory granules (
<figureCitation id="82BAE0C7927EFF88FE37F8B619F454BE" box="[388,471,1897,1923]" captionStart="Figure 7" captionStartId="9.[151,227,1701,1724]" captionTargetBox="[151,1432,1178,1680]" captionTargetId="figure-146@9.[151,1435,1178,1683]" captionTargetPageId="9" captionText="Figure 7. Leiurus nigellus sp. nov., right tibia, basitarsus, and telotarsus, retrolateral view, female holotype, A. leg III. B. leg IV. C. pedipalp chela fixed finger. D. pedipalp chela movable finger; male paratype. E. leg III. F. leg IV. G. pedipalp chela fixed finger. H. pedipalp chela movable finger." figureDoi="http://doi.org/10.5281/zenodo.13247372" httpUri="https://zenodo.org/record/13247372/files/figure.png" pageId="8" pageNumber="99">Figs. 7</figureCitation>
, C-D, G-H).
</paragraph>
<paragraph id="1A3EFC42927EFF88FF24F8721D0054D5" blockId="8.[151,1437,1965,2024]" pageId="8" pageNumber="99">
<emphasis id="28F52050927EFF88FF24F8721AB954FA" bold="true" box="[151,666,1965,1991]" pageId="8" pageNumber="99">Trichobothrial pattern orthobothriotaxic</emphasis>
(
<figureCitation id="82BAE0C7927EFF88FD18F8721AD054FA" box="[683,755,1965,1991]" captionStart="Figure 6" captionStartId="9.[151,227,816,839]" captionTargetBox="[154,1420,200,775]" captionTargetId="figure-9@9.[151,1435,189,798]" captionTargetPageId="9" captionText="Figure 6. Leiurus nigellus sp. nov., pedipalp segments, female holotype (A-F), A. femur internal view. B. femur dorsal view. C. patella dorsal view. D. patella external view. E. chela external view. F. chela ventral view; male paratype (G- L), G. femur internal view. H. femur dorsal view. I. patella dorsal view. J. patella external view. K. chela external view. L. chela ventral view. Trichobothrial pattern indicated by yellow circles. Scale = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13247370" httpUri="https://zenodo.org/record/13247370/files/figure.png" pageId="8" pageNumber="99">Fig. 6</figureCitation>
). 
<typeStatus id="C53A42E0927EFF88FCB9F8721B1F54FA" box="[778,828,1965,1991]" pageId="8" pageNumber="99">type</typeStatus>
A (
<bibRefCitation id="7E1081B3927EFF88FCDAF8721C3754FA" author="Vachon, M." box="[873,1044,1965,1991]" pageId="8" pageNumber="99" pagination="857 - 958" refId="ref8801" refString="Vachon, M. (1974) Etude des caracteres utilises pour classer les familles et les genres de Scorpions (Arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les Scorpions. Bulletin du Museum national d'Histoire naturelle, 3 e ser., n ° 140, Zoologie, 104, 857 - 958." type="journal article" year="1974">Vachon, 1974</bibRefCitation>
); dorsal trichobothria of femur in beta configuration (
<bibRefCitation id="7E1081B3927EFF88FECEF8111A0454D5" author="Vachon, M." box="[381,551,1998,2024]" pageId="8" pageNumber="99" pagination="1597 - 1599" refId="ref8866" refString="Vachon, M. (1975) Sur l'utilisation de la trichobothriotaxie du bras des pedipalpes des Scorpions (Arachnides) dans le classement des genres de la famille des Buthidae Simon. Comptes Rendus des Seances de l'Academie des Sciences, 281 (D), 1597 - 1599." type="journal article" year="1975">Vachon, 1975</bibRefCitation>
); pedipalp chela fixed finger with trichobothrium 
<emphasis id="28F52050927EFF88FBC2F8111CAD54D5" box="[1137,1166,1998,2024]" italics="true" pageId="8" pageNumber="99">db</emphasis>
distal to 
<emphasis id="28F52050927EFF88FB4FF8111D3E54D5" box="[1276,1309,1998,2024]" italics="true" pageId="8" pageNumber="99">est</emphasis>
.
</paragraph>
<caption id="4EFEACCA927FFF89FF24FCEF1CA3509E" ID-DOI="http://doi.org/10.5281/zenodo.13247370" ID-Zenodo-Dep="13247370" httpUri="https://zenodo.org/record/13247370/files/figure.png" pageId="9" pageNumber="100" startId="9.[151,227,816,839]" targetBox="[154,1420,200,775]" targetPageId="9" targetType="figure">
<paragraph id="1A3EFC42927FFF89FF24FCEF1CA3509E" blockId="9.[151,1437,816,931]" pageId="9" pageNumber="100">
<emphasis id="28F52050927FFF89FF24FCEF18DE507A" bold="true" box="[151,253,816,839]" pageId="9" pageNumber="100">Figure 6.</emphasis>
<taxonomicName id="DD8187C1927FFF89FEB6FCEF1991507A" authorityName="Afifeh &amp; Aloufi &amp; Al-Saraireh &amp; Badry &amp; Al-Qahtni &amp; Amr" authorityYear="2023" box="[261,434,816,839]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="100" phylum="Arthropoda" rank="species" species="nigellus" status="sp. nov.">
<emphasis id="28F52050927FFF89FEB6FCEF1991507A" box="[261,434,816,839]" italics="true" pageId="9" pageNumber="100">Leiurus nigellus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="33C69D2B927FFF89FE09FCEF1A32507A" box="[442,529,816,839]" pageId="9" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
, pedipalp segments, female holotype (
<emphasis id="28F52050927FFF89FC1DFCEF1BF9507A" bold="true" box="[942,986,816,839]" pageId="9" pageNumber="100">A-F</emphasis>
), 
<emphasis id="28F52050927FFF89FC41FCEF1C26507A" bold="true" box="[1010,1029,816,839]" pageId="9" pageNumber="100">A</emphasis>
. femur internal view. 
<emphasis id="28F52050927FFF89FB47FCEF1D25507A" bold="true" box="[1268,1286,816,839]" pageId="9" pageNumber="100">B</emphasis>
. femur dorsal view. 
<emphasis id="28F52050927FFF89FF6AFC9018CF505B" bold="true" box="[217,236,847,870]" pageId="9" pageNumber="100">C</emphasis>
. patella dorsal view. 
<emphasis id="28F52050927FFF89FE67FC9019C4505B" bold="true" box="[468,487,847,870]" pageId="9" pageNumber="100">D</emphasis>
. patella external view. 
<emphasis id="28F52050927FFF89FD57FC901AD5505B" bold="true" box="[740,758,847,870]" pageId="9" pageNumber="100">E</emphasis>
. chela external view. 
<emphasis id="28F52050927FFF89FC57FC901BD7505B" bold="true" box="[996,1012,847,870]" pageId="9" pageNumber="100">F</emphasis>
. chela ventral view; male paratype (
<emphasis id="28F52050927FFF89FACCFC90188A50B8" bold="true" pageId="9" pageNumber="100">G- L</emphasis>
), 
<emphasis id="28F52050927FFF89FF71FCB118F550B8" bold="true" box="[194,214,878,901]" pageId="9" pageNumber="100">G</emphasis>
. femur internal view. 
<emphasis id="28F52050927FFF89FE62FCB119C550B8" bold="true" box="[465,486,878,901]" pageId="9" pageNumber="100">H</emphasis>
. femur dorsal view. 
<emphasis id="28F52050927FFF89FD63FCB11AF950B8" bold="true" box="[720,730,878,901]" pageId="9" pageNumber="100">I</emphasis>
. patella dorsal view. 
<emphasis id="28F52050927FFF89FC78FCB11BFB50B8" bold="true" box="[971,984,878,901]" pageId="9" pageNumber="100">J</emphasis>
. patella external view. 
<emphasis id="28F52050927FFF89FB6CFCB11CD750B8" bold="true" box="[1247,1268,878,901]" pageId="9" pageNumber="100">K</emphasis>
. chela external view. 
<emphasis id="28F52050927FFF89FF6BFC5318C9509E" bold="true" box="[216,234,908,931]" pageId="9" pageNumber="100">L</emphasis>
. chela ventral view. Trichobothrial pattern indicated by yellow circles. Scale = 5 mm.
</paragraph>
</caption>
<paragraph id="1A3EFC42927FFF89FF24FBF01BBC57B0" blockId="9.[151,1437,1071,1165]" pageId="9" pageNumber="100">
<emphasis id="28F52050927FFF89FF24FBF018F85774" bold="true" box="[151,219,1071,1097]" pageId="9" pageNumber="100">Legs:</emphasis>
(
<figureCitation id="82BAE0C7927FFF89FF5DFBF019675774" box="[238,324,1071,1097]" captionStart="Figure 7" captionStartId="9.[151,227,1701,1724]" captionTargetBox="[151,1432,1178,1680]" captionTargetId="figure-146@9.[151,1435,1178,1683]" captionTargetPageId="9" captionText="Figure 7. Leiurus nigellus sp. nov., right tibia, basitarsus, and telotarsus, retrolateral view, female holotype, A. leg III. B. leg IV. C. pedipalp chela fixed finger. D. pedipalp chela movable finger; male paratype. E. leg III. F. leg IV. G. pedipalp chela fixed finger. H. pedipalp chela movable finger." figureDoi="http://doi.org/10.5281/zenodo.13247372" httpUri="https://zenodo.org/record/13247372/files/figure.png" pageId="9" pageNumber="100">Figs. 7</figureCitation>
, A-B, E-F). with ventral aspect of tarsi presenting numerous thin setae not well arranged in rows. retrosuperior setae on basitarsus III counts 12-15. Tibial spurs present on legs III and IV strong. Pedal spurs present, strong on all legs, prolateral spurs basally bifurcate.
</paragraph>
<caption id="4EFEACCA927FFF89FF24F97A1B1355C7" ID-DOI="http://doi.org/10.5281/zenodo.13247372" ID-Zenodo-Dep="13247372" httpUri="https://zenodo.org/record/13247372/files/figure.png" pageId="9" pageNumber="100" startId="9.[151,227,1701,1724]" targetBox="[151,1432,1178,1680]" targetPageId="9" targetType="figure">
<paragraph id="1A3EFC42927FFF89FF24F97A1B1355C7" blockId="9.[151,1437,1701,1786]" pageId="9" pageNumber="100">
<emphasis id="28F52050927FFF89FF24F97A18DD5581" bold="true" box="[151,254,1701,1724]" pageId="9" pageNumber="100">Figure 7.</emphasis>
<taxonomicName id="DD8187C1927FFF89FEB4F97A19965581" authorityName="Afifeh &amp; Aloufi &amp; Al-Saraireh &amp; Badry &amp; Al-Qahtni &amp; Amr" authorityYear="2023" box="[263,437,1701,1724]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="100" phylum="Arthropoda" rank="species" species="nigellus" status="sp. nov.">
<emphasis id="28F52050927FFF89FEB4F97A19965581" box="[263,437,1701,1724]" italics="true" pageId="9" pageNumber="100">Leiurus nigellus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="33C69D2B927FFF89FE0FF97A1A375581" box="[444,532,1701,1724]" pageId="9" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
, right tibia, basitarsus, and telotarsus, retrolateral view, female holotype, 
<emphasis id="28F52050927FFF89FA83F97A1D605581" bold="true" box="[1328,1347,1701,1724]" pageId="9" pageNumber="100">A</emphasis>
. leg III. 
<emphasis id="28F52050927FFF89FF24F91B188A55E6" bold="true" box="[151,169,1732,1755]" pageId="9" pageNumber="100">B</emphasis>
. leg IV. 
<emphasis id="28F52050927FFF89FEA3F91B190055E6" bold="true" box="[272,291,1732,1755]" pageId="9" pageNumber="100">C</emphasis>
. pedipalp chela fixed finger. 
<emphasis id="28F52050927FFF89FDD9F91B1A5E55E6" bold="true" box="[618,637,1732,1755]" pageId="9" pageNumber="100">D</emphasis>
. pedipalp chela movable finger; male paratype. 
<emphasis id="28F52050927FFF89FB20F91B1C8655E6" bold="true" box="[1171,1189,1732,1755]" pageId="9" pageNumber="100">E</emphasis>
. leg III. 
<emphasis id="28F52050927FFF89FAB9F91B1D3955E6" bold="true" box="[1290,1306,1732,1755]" pageId="9" pageNumber="100">F</emphasis>
. leg IV. 
<emphasis id="28F52050927FFF89FA32F91B1DB655E6" bold="true" box="[1409,1429,1732,1755]" pageId="9" pageNumber="100">G</emphasis>
. pedipalp chela fixed finger. 
<emphasis id="28F52050927FFF89FE70F93C19FB55C7" bold="true" box="[451,472,1763,1786]" pageId="9" pageNumber="100">H</emphasis>
. pedipalp chela movable finger.
</paragraph>
</caption>
<paragraph id="1A3EFC42927FFF89FF24F8B719B654BF" blockId="9.[151,1436,1896,1989]" box="[151,405,1896,1922]" pageId="9" pageNumber="100">
<emphasis id="28F52050927FFF89FF24F8B719B654BF" bold="true" box="[151,405,1896,1922]" pageId="9" pageNumber="100">Sexual dimorphism:</emphasis>
</paragraph>
<paragraph id="1A3EFC42927FFF89FF24F8561BCB54F8" blockId="9.[151,1436,1896,1989]" pageId="9" pageNumber="100">Males differed from females as follows: Males with more robust carination on tergites and sternites III–V, more slender pedipalps and metasoma, longer pectines with larger teeth.</paragraph>
<paragraph id="1A3EFC42927CFF8AFF24FF1A192853E2" blockId="10.[151,1437,197,1200]" box="[151,267,197,223]" pageId="10" pageNumber="101">
<emphasis id="28F52050927CFF8AFF24FF1A192853E2" bold="true" box="[151,267,197,223]" pageId="10" pageNumber="101">Affinities</emphasis>
</paragraph>
<paragraph id="1A3EFC42927CFF8AFF24FF39190451A8" blockId="10.[151,1437,197,1200]" pageId="10" pageNumber="101">
<taxonomicName id="DD8187C1927CFF8AFF24FF391978523D" authorityName="Afifeh &amp; Aloufi &amp; Al-Saraireh &amp; Badry &amp; Al-Qahtni &amp; Amr" authorityYear="2023" box="[151,347,230,256]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="nigellus" status="sp. nov.">
<emphasis id="28F52050927CFF8AFF24FF391978523D" box="[151,347,230,256]" italics="true" pageId="10" pageNumber="101">Leiurus nigellus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="33C69D2B927CFF8AFEDAFF3919E8523D" box="[361,459,230,256]" pageId="10" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
is distinguished from most other species of 
<taxonomicName id="DD8187C1927CFF8AFBB1FF391C7F523D" authorityName="Ehrenberg" authorityYear="1828" box="[1026,1116,230,256]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="28F52050927CFF8AFBB1FF391C7F523D" box="[1026,1116,230,256]" italics="true" pageId="10" pageNumber="101">Leiurus</emphasis>
</taxonomicName>
known from the Arabian Peninsula by its distinctive predominantly blackish colour pattern, the new species differs from 
<taxonomicName id="DD8187C1927CFF8AFA36FED71903527E" authorityName="Lourenco, Modry &amp; Amr" authorityYear="2002" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis id="28F52050927CFF8AFA36FED71DB6521F" box="[1413,1429,264,290]" italics="true" pageId="10" pageNumber="101">L</emphasis>
. 
<emphasis id="28F52050927CFF8AFF24FEF61903527E" box="[151,288,297,323]" italics="true" pageId="10" pageNumber="101">jordanensis</emphasis>
</taxonomicName>
by having: 
<emphasis id="28F52050927CFF8AFE1FFEF619F6527E" bold="true" box="[428,469,297,323]" pageId="10" pageNumber="101">(a).</emphasis>
less slender body parts, including metasomal segments, legs and pedipalps (
<tableCitation id="5703C9F9927CFF8AFAEBFEF618845258" captionStart="Table 3" captionStartId="12.[410,475,314,337]" captionText="Table 3. Variation in morphometric ratios for Leiurus nigellus sp. nov., Leiurus jordanensis, Leiurus" pageId="10" pageNumber="101">Table 3</tableCitation>
); and by combining morphometric ratios describing slenderness of three segments (leg, pedipalp and metasomal segments) of adult females resulted in 
<emphasis id="28F52050927CFF8AFD40FEB21B2752BA" box="[755,772,365,391]" italics="true" pageId="10" pageNumber="101">F</emphasis>
s that ranged from 27.78–32.32 (n=4) in 
<taxonomicName id="DD8187C1927CFF8AFB47FEB21D5152BA" authorityName="Afifeh &amp; Aloufi &amp; Al-Saraireh &amp; Badry &amp; Al-Qahtni &amp; Amr" authorityYear="2023" box="[1268,1394,365,391]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="nigellus" status="sp. nov.">
<emphasis id="28F52050927CFF8AFB47FEB21D2752BA" box="[1268,1284,365,391]" italics="true" pageId="10" pageNumber="101">L</emphasis>
. 
<emphasis id="28F52050927CFF8AFAA6FEB21D5152BA" box="[1301,1394,365,391]" italics="true" pageId="10" pageNumber="101">nigellus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="33C69D2B927CFF8AFAC8FEB218EE5295" pageId="10" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
, 35.06–41.78 (n =12) in 
<taxonomicName id="DD8187C1927CFF8AFDA4FE511AE55295" authorityName="Lourenco, Modry &amp; Amr" authorityYear="2002" box="[535,710,398,424]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis id="28F52050927CFF8AFDA4FE511A045295" box="[535,551,398,424]" italics="true" pageId="10" pageNumber="101">L</emphasis>
. 
<emphasis id="28F52050927CFF8AFD8EFE511AE55295" box="[573,710,398,424]" italics="true" pageId="10" pageNumber="101">jordanensis</emphasis>
</taxonomicName>
; 
<emphasis id="28F52050927CFF8AFD6EFE511B2A5295" bold="true" box="[733,777,398,424]" pageId="10" pageNumber="101">(b).</emphasis>
sternite III with median carinae weak to moderate, irregularly granular, sternites IV–V with lateral carinae moderate, granular, median carinae weak in females of the new species, whereas sternite III with median carinae strong, granular, sternites IV–V with lateral carinae strong, granular, median carinae moderate in females 
<taxonomicName id="DD8187C1927CFF8AFC38FE2B1C155133" authorityName="Lourenco, Modry &amp; Amr" authorityYear="2002" box="[907,1078,500,526]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis id="28F52050927CFF8AFC38FE2B1BB85133" box="[907,923,500,526]" italics="true" pageId="10" pageNumber="101">L</emphasis>
. 
<emphasis id="28F52050927CFF8AFC1EFE2B1C155133" box="[941,1078,500,526]" italics="true" pageId="10" pageNumber="101">jordanensis</emphasis>
</taxonomicName>
; 
<emphasis id="28F52050927CFF8AFBF9FE2B1C515133" bold="true" box="[1098,1138,500,526]" pageId="10" pageNumber="101">(c).</emphasis>
different colour pattern, with vesicle of telson black to blackish brown in adults, and it is light black in juveniles, whereas the vesicle is yellow in adults and juveniles of 
<taxonomicName id="DD8187C1927CFF8AFDE6FDE81B22516C" authorityName="Lourenco, Modry &amp; Amr" authorityYear="2002" box="[597,769,567,593]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis id="28F52050927CFF8AFDE6FDE81A46516C" box="[597,613,567,593]" italics="true" pageId="10" pageNumber="101">L</emphasis>
. 
<emphasis id="28F52050927CFF8AFDCBFDE81B22516C" box="[632,769,567,593]" italics="true" pageId="10" pageNumber="101">jordanensis</emphasis>
</taxonomicName>
, also pedipalp fingers of the new species are black to blackish brown overall except for the distal third with yellowish-brown, but they are completely yellow in 
<taxonomicName id="DD8187C1927CFF8AFA36FD86190351A8" authorityName="Lourenco, Modry &amp; Amr" authorityYear="2002" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis id="28F52050927CFF8AFA36FD861DB6514E" box="[1413,1429,601,627]" italics="true" pageId="10" pageNumber="101">L</emphasis>
. 
<emphasis id="28F52050927CFF8AFF24FDA4190351A8" box="[151,288,635,661]" italics="true" pageId="10" pageNumber="101">jordanensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="1A3EFC42927CFF8AFF46FD421D085062" blockId="10.[151,1437,197,1200]" pageId="10" pageNumber="101">
<taxonomicName id="DD8187C1927CFF8AFF46FD4219E2518A" authorityName="Lowe, Yagmur &amp; Kovarik" authorityYear="2014" box="[245,449,669,695]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis id="28F52050927CFF8AFF46FD4219E2518A" box="[245,449,669,695]" italics="true" pageId="10" pageNumber="101">Leiurus arabicus</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName id="DD8187C1927CFF8AFD67FD421B70518A" authorityName="Afifeh &amp; Aloufi &amp; Al-Saraireh &amp; Badry &amp; Al-Qahtni &amp; Amr" authorityYear="2023" box="[724,851,669,695]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="nigellus" status="sp. nov.">
<emphasis id="28F52050927CFF8AFD67FD421AC7518A" box="[724,740,669,695]" italics="true" pageId="10" pageNumber="101">L</emphasis>
. 
<emphasis id="28F52050927CFF8AFD45FD421B70518A" box="[758,851,669,695]" italics="true" pageId="10" pageNumber="101">nigellus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="33C69D2B927CFF8AFCEDFD421B9D518A" box="[862,958,669,695]" pageId="10" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
by having: 
<emphasis id="28F52050927CFF8AFBE0FD421C5E518A" bold="true" box="[1107,1149,669,695]" pageId="10" pageNumber="101">(a).</emphasis>
less slender metasomal segments, and less slenderness factor in adult females (Table 2 &amp; 3); 
<emphasis id="28F52050927CFF8AFC78FD611BCB51E5" box="[971,1000,702,728]" italics="true" pageId="10" pageNumber="101">Fs</emphasis>
ranged from 
<quantity id="DD7951A7927CFF8AFB3BFD611D1951E5" box="[1160,1338,702,728]" metricMagnitude="-1" metricUnit="m" metricValue="6.668769999999999" metricValueMax="6.672580000000001" metricValueMin="6.664959999999999" pageId="10" pageNumber="101" unit="in" value="26.255" valueMax="26.27" valueMin="26.24">26.24–26.27 in</quantity>
females (n=2); 
<emphasis id="28F52050927CFF8AFF5BFD3F193051C7" bold="true" box="[232,275,736,762]" pageId="10" pageNumber="101">(b).</emphasis>
different colour pattern, base colour is yellow, with less or more dark reddish-brown pigmentation on carapace, tergites and metasoma V, in contrast the new species is black to blackish-brown overall; 
<emphasis id="28F52050927CFF8AFAC6FCDD1DBF5021" bold="true" box="[1397,1436,770,796]" pageId="10" pageNumber="101">(c).</emphasis>
ventrolateral carinae of metasoma V with enlarged triangular or subtriangular denticles, whereas in 
<taxonomicName id="DD8187C1927CFF8AFA36FCFC18D75062" authorityName="Afifeh &amp; Aloufi &amp; Al-Saraireh &amp; Badry &amp; Al-Qahtni &amp; Amr" authorityYear="2023" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="nigellus" status="sp. nov.">
<emphasis id="28F52050927CFF8AFA36FCFC1DB65000" box="[1413,1429,803,829]" italics="true" pageId="10" pageNumber="101">L</emphasis>
. 
<emphasis id="28F52050927CFF8AFF24FC9A18D75062" box="[151,244,837,863]" italics="true" pageId="10" pageNumber="101">nigellus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="33C69D2B927CFF8AFF4FFC9A197B5062" box="[252,344,837,863]" pageId="10" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
ventrolateral carinae of metasoma V with slightly enlarged subtriangular denticles.
</paragraph>
<paragraph id="1A3EFC42927CFF8AFF46FCB8192B578D" blockId="10.[151,1437,197,1200]" pageId="10" pageNumber="101">
<taxonomicName id="DD8187C1927CFF8AFF46FCB819E350BC" authorityName="Lowe, Yagmur &amp; Kovarik" authorityYear="2014" box="[245,448,871,897]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="haenggii">
<emphasis id="28F52050927CFF8AFF46FCB819E350BC" box="[245,448,871,897]" italics="true" pageId="10" pageNumber="101">Leiurus haenggii</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName id="DD8187C1927CFF8AFD60FCB81B7150BC" authorityName="Afifeh &amp; Aloufi &amp; Al-Saraireh &amp; Badry &amp; Al-Qahtni &amp; Amr" authorityYear="2023" box="[723,850,871,897]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="nigellus" status="sp. nov.">
<emphasis id="28F52050927CFF8AFD60FCB81AC050BC" box="[723,739,871,897]" italics="true" pageId="10" pageNumber="101">L</emphasis>
. 
<emphasis id="28F52050927CFF8AFD46FCB81B7150BC" box="[757,850,871,897]" italics="true" pageId="10" pageNumber="101">nigellus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="33C69D2B927CFF8AFCEEFCB81B9F50BC" box="[861,956,871,897]" pageId="10" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
by having 
<emphasis id="28F52050927CFF8AFBE0FCB81C5E50BC" bold="true" box="[1107,1149,871,897]" pageId="10" pageNumber="101">(a).</emphasis>
less slender metasomal segments, legs and pedipalps (Table 2 &amp; 3); 
<emphasis id="28F52050927CFF8AFD07FC571AF2509F" box="[692,721,904,930]" italics="true" pageId="10" pageNumber="101">Fs</emphasis>
ranged from 
<quantity id="DD7951A7927CFF8AFCC5FC571C0A509F" box="[886,1065,904,930]" metricMagnitude="-1" metricUnit="m" metricValue="5.23875" metricValueMax="5.3924199999999995" metricValueMin="5.08508" pageId="10" pageNumber="101" unit="in" value="20.625" valueMax="21.23" valueMin="20.02">20.02–21.23 in</quantity>
females (n=2); 
<emphasis id="28F52050927CFF8AFB58FC571D35509F" bold="true" box="[1259,1302,904,930]" pageId="10" pageNumber="101">(b).</emphasis>
sternite III with median carinae very weak, smooth, almost obsolete; sternites IV–V with weak, finely granulated lateral carinae, obsolete median carinae; whereas sternite III with median carinae weak to moderate, irregularly granular, sternites IV–V with lateral carinae moderate, granular, median carinae weak in females of the new species; 
<emphasis id="28F52050927CFF8AFEB5FBD0190D5714" bold="true" box="[262,302,1039,1065]" pageId="10" pageNumber="101">(c).</emphasis>
different colour pattern, base colour is yellow, with less or more dark reddish-brown pigmentation on carapace, tergites and metasoma V, in contrast 
<taxonomicName id="DD8187C1927CFF8AFC14FBEE1C045776" authorityName="Afifeh &amp; Aloufi &amp; Al-Saraireh &amp; Badry &amp; Al-Qahtni &amp; Amr" authorityYear="2023" box="[935,1063,1073,1099]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="nigellus" status="sp. nov.">
<emphasis id="28F52050927CFF8AFC14FBEE1B945776" box="[935,951,1073,1099]" italics="true" pageId="10" pageNumber="101">L</emphasis>
. 
<emphasis id="28F52050927CFF8AFC79FBEE1C045776" box="[970,1063,1073,1099]" italics="true" pageId="10" pageNumber="101">nigellus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="33C69D2B927CFF8AFB82FBEE1CB25776" box="[1073,1169,1073,1099]" pageId="10" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
is black to blackish-brown overall; 
<emphasis id="28F52050927CFF8AFEE6FB8C19A35750" bold="true" box="[341,384,1107,1133]" pageId="10" pageNumber="101">(d).</emphasis>
ventrolateral carinae of metasoma V with enlarged subtriangular or lobate denticles, whereas in 
<taxonomicName id="DD8187C1927CFF8AFE96FBAA198657B2" authorityName="Afifeh &amp; Aloufi &amp; Al-Saraireh &amp; Badry &amp; Al-Qahtni &amp; Amr" authorityYear="2023" box="[293,421,1141,1167]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="nigellus" status="sp. nov.">
<emphasis id="28F52050927CFF8AFE96FBAA191657B2" box="[293,309,1141,1167]" italics="true" pageId="10" pageNumber="101">L</emphasis>
. 
<emphasis id="28F52050927CFF8AFEFBFBAA198657B2" box="[328,421,1141,1167]" italics="true" pageId="10" pageNumber="101">nigellus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="33C69D2B927CFF8AFE03FBAA1A3257B2" box="[432,529,1141,1167]" pageId="10" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
ventrolateral carinae of metasoma V with slightly enlarged subtriangular denticles.
</paragraph>
<paragraph id="1A3EFC42927CFF8AFF24FB0518D457C9" blockId="10.[151,1437,1242,1538]" box="[151,247,1242,1268]" pageId="10" pageNumber="101">
<emphasis id="28F52050927CFF8AFF24FB0518D457C9" bold="true" box="[151,247,1242,1268]" pageId="10" pageNumber="101">Habitat</emphasis>
</paragraph>
<paragraph id="1A3EFC42927CFF8AFF24FB231AEF553F" blockId="10.[151,1437,1242,1538]" pageId="10" pageNumber="101">
The collection site is located in Al Buriakah village (
<quantity id="DD7951A7927CFF8AFCAEFB231B59562B" box="[797,890,1276,1302]" metricMagnitude="3" metricUnit="m" metricValue="1.109" pageId="10" pageNumber="101" unit="m" value="1109.0">1109 m</quantity>
a.s.l.), 
<quantity id="DD7951A7927CFF8AFC65FB231C11562B" box="[982,1074,1276,1302]" metricMagnitude="5" metricUnit="m" metricValue="1.3" pageId="10" pageNumber="101" unit="km" value="130.0">130 km</quantity>
north of Al Ula Governorate, north of 
<collectingRegion id="D84532A0927CFF8AFEBBFAC219B8560A" box="[264,411,1309,1335]" country="Saudi Arabia" name="Al Madinah" pageId="10" pageNumber="101">Al Madinah</collectingRegion>
Al Monawwarah Province. 
<taxonomicName id="DD8187C1927CFF8AFD4CFAC21BE0560A" authorityName="Afifeh &amp; Aloufi &amp; Al-Saraireh &amp; Badry &amp; Al-Qahtni &amp; Amr" authorityYear="2023" box="[767,963,1309,1335]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="nigellus" status="sp. nov.">
<emphasis id="28F52050927CFF8AFD4CFAC21BE0560A" box="[767,963,1309,1335]" italics="true" pageId="10" pageNumber="101">Leiurus nigellus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="33C69D2B927CFF8AFC63FAC21C10560A" box="[976,1075,1309,1335]" pageId="10" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
was collected from an arid environment with sparse vegetation. The habitat is a flat desert covered with thin layer of sand on top of gravel with scattered sandstone bed rocks of various sizes, with several deserted rodent burrows (
<figureCitation id="82BAE0C7927CFF8AFAF1FABE1DAE5646" box="[1346,1421,1377,1403]" captionStart="Figure 8" captionStartId="13.[151,227,1171,1194]" captionTargetBox="[151,1435,189,1152]" captionTargetId="figure-9@13.[151,1435,189,1152]" captionTargetPageId="13" captionText="Figure 8. Habitat of Leiurus nigellus sp. nov. in Al Buriakah village, Al Ula Governorate, Al Madinah Al Monawwarah Province, Saudi Arabia." figureDoi="http://doi.org/10.5281/zenodo.13247374" httpUri="https://zenodo.org/record/13247374/files/figure.png" pageId="10" pageNumber="101">Fig. 8</figureCitation>
). More to the west, sandstone deserts become more prominent interrupted by narrow wadi systems, with few vegetation of 
<taxonomicName id="DD8187C1927CFF8AFEF8FA7B1A4C5683" box="[331,623,1444,1470]" class="Magnoliopsida" family="Amaranthaceae" genus="Haloxylon" kingdom="Plantae" order="Caryophyllales" pageId="10" pageNumber="101" phylum="Tracheophyta" rank="species" species="salicornicum">
<emphasis id="28F52050927CFF8AFEF8FA7B1A4C5683" box="[331,623,1444,1470]" italics="true" pageId="10" pageNumber="101">Haloxylon salicornicum</emphasis>
</taxonomicName>
, 
<taxonomicName id="DD8187C1927CFF8AFD34FA7B1B665683" box="[647,837,1444,1470]" class="Magnoliopsida" family="Fabaceae" genus="Retama" kingdom="Plantae" order="Fabales" pageId="10" pageNumber="101" phylum="Tracheophyta" rank="species" species="raetam">
<emphasis id="28F52050927CFF8AFD34FA7B1B665683" box="[647,837,1444,1470]" italics="true" pageId="10" pageNumber="101">Retama raetam</emphasis>
</taxonomicName>
, 
<taxonomicName id="DD8187C1927CFF8AFCEEFA7B1BD45683" box="[861,1015,1444,1470]" class="Liliopsida" family="Poaceae" genus="Panicum" kingdom="Plantae" order="Poales" pageId="10" pageNumber="101" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="28F52050927CFF8AFCEEFA7B1BE55683" box="[861,966,1444,1470]" italics="true" pageId="10" pageNumber="101">Panicum</emphasis>
sp.
</taxonomicName>
and 
<taxonomicName id="DD8187C1927CFF8AFBF0FA7B1D615683" box="[1091,1346,1444,1470]" class="Magnoliopsida" family="Cucurbitaceae" genus="Citrullus" kingdom="Plantae" order="Cucurbitales" pageId="10" pageNumber="101" phylum="Tracheophyta" rank="species" species="colocynthis">
<emphasis id="28F52050927CFF8AFBF0FA7B1D615683" box="[1091,1346,1444,1470]" italics="true" pageId="10" pageNumber="101">Citrullus colocynthis</emphasis>
</taxonomicName>
. Other scorpions found in the same area include 
<taxonomicName id="DD8187C1927CFF8AFD37FA191C4F56DD" authority="(Olivier, 1807)" baseAuthorityName="Olivier" baseAuthorityYear="1807" box="[644,1132,1478,1504]" class="Arachnida" family="Buthidae" genus="Androctonus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="crassicauda">
<emphasis id="28F52050927CFF8AFD37FA191B9356DD" box="[644,944,1478,1504]" italics="true" pageId="10" pageNumber="101">Androctonus crassicauda</emphasis>
(Olivier, 1807)
</taxonomicName>
, 
<taxonomicName id="DD8187C1927CFF8AFBCAFA1919DB553F" authority="Levy, Amitai &amp; Shulov, 1973" authorityName="Levy, Amitai &amp; Shulov" authorityYear="1973" class="Arachnida" family="Buthidae" genus="Vachoniolus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="globimanus">
<emphasis id="28F52050927CFF8AFBCAFA191DBE56DD" box="[1145,1437,1478,1504]" italics="true" pageId="10" pageNumber="101">Vachoniolus globimanus</emphasis>
<bibRefCitation id="7E1081B3927CFF8AFF24FA3719DB553F" author="Levy, G. &amp; Amitai, P. &amp; Shulov, A." box="[151,504,1512,1538]" pageId="10" pageNumber="101" pagination="113 - 140" refId="ref8392" refString="Levy, G., Amitai, P. &amp; Shulov, A. (1973) New scorpions from Israel, Jordan and Arabia. Zoological Journal of the Linnaean Society, 52, 113 - 140." type="journal article" year="1973">Levy, Amitai &amp; Shulov, 1973</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="DD8187C1927CFF8AFD85FA371AEF553F" box="[566,716,1512,1538]" class="Arachnida" family="Buthidae" genus="Buthacus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="101" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="28F52050927CFF8AFD85FA371A80553F" box="[566,675,1512,1538]" italics="true" pageId="10" pageNumber="101">Buthacus</emphasis>
sp.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>