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<document id="8133D0C836376B0EAD702EDCEC05D40E" ID-CLB-Dataset="4042" ID-DOI="10.5852/ejt.2021.772.1505" ID-GBIF-Dataset="b65bb5a9-bbe7-49a4-af44-4b4c03121288" ID-ISSN="2118-9773" ID-Zenodo-Dep="5536410" ID-ZooBank="9B82A32F-0A07-47E3-8684-FED7C8EBF1E9" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1632919496065" checkinUser="felipe" docAuthor="Miranda, Gustavo Silva de, Giupponi, Alessandro P. L., Prendini, Lorenzo &amp; Scharff, Nikolaj" docDate="2021" docId="8F431375FFE2FFAFA583FBCAFE71DBE9" docLanguage="en" docName="ejt-772_1-409.pdf" docOrigin="European Journal of Taxonomy 772" docSource="http://dx.doi.org/10.5852/ejt.2021.772.1505" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Charinus puri Miranda &amp; Giupponi &amp; Prendini &amp; Scharff 2021, sp. nov." docType="treatment" docUuid="58C93BF8-8C3F-4120-B389-61528D8C3811" docUuidSource="ZooBank" docVersion="14" lastPageNumber="125" masterDocId="737A6B0DFF99FFD3A710FFEFFFC6DC01" masterDocTitle="Systematic revision of the pantropical whip spider family Charinidae Quintero, 1986 (Arachnida, Amblypygi)" masterLastPageNumber="409" masterPageNumber="1" pageNumber="124" updateTime="1699416367315" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="127C4B799A254A349C4FD59EF113736B">Systematic revision of the pantropical whip spider family Charinidae Quintero, 1986 (Arachnida, Amblypygi)</mods:title>
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<mods:namePart id="C5CE666D77D60DBB56A72BB329C92BB9">Miranda, Gustavo Silva de</mods:namePart>
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<mods:affiliation id="42EA8A2C35C83457D8231689B5E5FA26">Center for Macroecology, Evolution and Climate, Natural History Museum of Denmark, University of Copenhagen, Copenhagen, Denmark. &amp; Entomology Department, National Museum of Natural History, Smithsonian Institution, Washington DC, USA.</mods:affiliation>
<mods:nameIdentifier id="0750F15467FF6EC31CCDED97AD4BD663" type="email">smiranda.gustavo@gmail.com</mods:nameIdentifier>
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<mods:namePart id="8A7E31ABC399CAC94153A00066D8F6BB">Giupponi, Alessandro P. L.</mods:namePart>
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<mods:affiliation id="0BE340E1B22F4D1EEF1F28F5423C4AA9">Laboratório de Referência Nacional em Vetores das Riquetsioses, LIRN-IOC-FIOCRUZ, Rio de Janeiro, RJ, Brazil.</mods:affiliation>
<mods:nameIdentifier id="78DF860B092BA310104B12617BDEDA27" type="email">agiupponi@gmail.com</mods:nameIdentifier>
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<mods:namePart id="D0A3B1C57490F820C8007B51AB99D4F4">Prendini, Lorenzo</mods:namePart>
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<mods:affiliation id="C9B204B02385A9C94F9C6D224AFBB6FB">Arachnology Lab, Division of Invertebrate Zoology, American Museum of Natural History, New York, NY, USA.</mods:affiliation>
<mods:nameIdentifier id="CB5E818B7412604B152EA815BD8BA4F0" type="email">lorenzo@amnh.org</mods:nameIdentifier>
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<mods:namePart id="4A692D1F011952B40455C160D8A4985A">Scharff, Nikolaj</mods:namePart>
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<mods:affiliation id="9ED7C313BD800121D1661DDF73F6E148">Center for Macroecology, Evolution and Climate, Natural History Museum of Denmark, University of Copenhagen, Copenhagen, Denmark. &amp; Entomology Department, National Museum of Natural History, Smithsonian Institution, Washington DC, USA. &amp; Zoology Section, Natural History Museum of Denmark, University of Copenhagen, Denmark.</mods:affiliation>
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<mods:title id="DAA93E82A92D4F26F3597390CB8E096F">European Journal of Taxonomy</mods:title>
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<mods:date id="017B97FBE7CE16E5CE4D328EDCDB9A1F">2021</mods:date>
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<treatment id="8F431375FFE2FFAFA583FBCAFE71DBE9" ID-DOI="http://doi.org/10.5281/zenodo.5536818" ID-GBIF-Taxon="188640151" ID-Zenodo-Dep="5536818" LSID="urn:lsid:zoobank.org:act:58C93BF8-8C3F-4120-B389-61528D8C3811" httpUri="http://treatment.plazi.org/id/8F431375FFE2FFAFA583FBCAFE71DBE9" lastPageId="124" lastPageNumber="125" pageId="123" pageNumber="124">
<subSubSection id="4FF0F1E8FFE2FFA8A583FBCAFC67D841" box="[659,929,1061,1088]" pageId="123" pageNumber="124" type="nomenclature">
<paragraph id="0755A263FFE2FFA8A583FBCAFC67D841" blockId="123.[376,1211,1061,1156]" box="[659,929,1061,1088]" pageId="123" pageNumber="124">
<heading id="5C1D150FFFE2FFA8A583FBCAFC67D841" box="[659,929,1061,1088]" centered="true" fontSize="11" level="2" pageId="123" pageNumber="124" reason="2">
<taxonomicName id="C0EAD9E0FFE2FFA8A583FBCAFCF9D83E" ID-CoL="8VNFR" authority="Miranda &amp; Giupponi &amp; Prendini &amp; Scharff, 2021" authorityName="Miranda &amp; Giupponi &amp; Prendini &amp; Scharff" authorityYear="2021" box="[659,831,1061,1088]" class="Arachnida" family="Charinidae" genus="Charinus" kingdom="Animalia" order="Amblypygi" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="puri" status="sp. nov.">
<emphasis id="359E7E71FFE2FFA8A583FBCAFCF9D83E" bold="true" box="[659,831,1061,1088]" italics="true" pageId="123" pageNumber="124">Charinus puri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2EADC30AFFE2FFA8A456FBC9FC67D841" box="[838,929,1062,1088]" pageId="123" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
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<subSubSection id="4FF0F1E8FFE2FFA8A668FBA8FC62D885" pageId="123" pageNumber="124" type="description">
<paragraph id="0755A263FFE2FFA8A668FBA8FB7DD863" blockId="123.[376,1211,1061,1156]" box="[376,1211,1095,1122]" pageId="123" pageNumber="124">
<uri id="737BAE61FFE2FFA8A668FBA8FB7DD863" box="[376,1211,1095,1122]" pageId="123" pageNumber="124">
urn:lsid:zoobank.org:act:
<uuid id="734C98B6FFE2FFA8A58BFBA8FB7DD863" box="[667,1211,1095,1122]" pageId="123" pageNumber="124">58C93BF8-8C3F-4120-B389-61528D8C3811</uuid>
</uri>
</paragraph>
<paragraph id="0755A263FFE2FFA8A59FFB86FC62D885" blockId="123.[376,1211,1061,1156]" box="[655,932,1129,1156]" pageId="123" pageNumber="124">
<figureCitation id="9FD1BEE6FFE2FFA8A59FFB86FD2FD885" box="[655,745,1129,1156]" captionStart="Fig" captionStartId="298.[189,232,1225,1251]" captionTargetBox="[189,1398,265,1185]" captionTargetId="figure-0@298.[189,1399,265,1185]" captionTargetPageId="298" captionText="Fig. 43. Map plotting known distributions of species of Charinus Simon, 1892 in eastern South America, with inset for southeastern Brazil." figureDoi="http://doi.org/10.5281/zenodo.5536508" httpUri="https://zenodo.org/record/5536508/files/figure.png" pageId="123" pageNumber="124">Figs 43</figureCitation>
,
<figureCitation id="9FD1BEE6FFE2FFA8A5E5FB85FC84D885" box="[757,834,1130,1156]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="326.[189,232,1718,1744]" captionStartId-1="327.[189,232,1445,1471]" captionTargetBox-0="[189,1399,265,1667]" captionTargetBox-1="[189,1398,265,1393]" captionTargetId-0="figure-47@326.[189,1399,265,1667]" captionTargetId-1="figure-42@327.[189,1399,265,1393]" captionTargetPageId-0="326" captionTargetPageId-1="327" captionText-0="Fig. 71. Charinus puri sp. nov. (MNRJ 9223), general morphology.A. Carapace, dorsal view.B. Sternum, ventral view. C. Pedipalp tarsus, frontal view. D. Pedipalp, dorsal view. E. Pedipalp, ventral view. Scale bars: A–B, D–E = 1 mm; C = 0.5 mm." captionText-1="Fig. 72. Charinus puri sp. nov. (MNRJ 9223), female gonopod and genital operculum. A. Sucker-like gonopod, dorsal view. B. Genital operculum, ventral view. C. Gonopods, posterior view. D. Posterior margin of genital operculum, ventral view. F. Cerotegument." figureDoi-0="http://doi.org/10.5281/zenodo.5536573" figureDoi-1="http://doi.org/10.5281/zenodo.5536575" httpUri-0="https://zenodo.org/record/5536573/files/figure.png" httpUri-1="https://zenodo.org/record/5536575/files/figure.png" pageId="123" pageNumber="124">71–72</figureCitation>
;
<tableCitation id="4A6897D8FFE2FFA8A45DFB86FC62D885" box="[845,932,1129,1156]" captionStart="Table 4" captionStartId="89.[181,251,552,581]" captionTargetBox="[185,1925,631,1740]" captionTargetId="graphics-20@89.[181,1932,659,1753]" captionTargetPageId="89" captionText="Table 4 (continued on next five pages). Measurements of all eastern South American species of Charinus Simon, 1892." httpUri="http://table.plazi.org/id/5395F2EBFFC0FF8AA7A5FDC7F9F2DE45" pageId="123" pageNumber="124" tableUuid="5395F2EBFFC0FF8AA7A5FDC7F9F2DE45">Table 4</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="4FF0F1E8FFE2FFA8A7ADFB45FC8DDA42" pageId="123" pageNumber="124" type="diagnosis">
<paragraph id="0755A263FFE2FFA8A7ADFB45FEF0D8C5" blockId="123.[189,310,1194,1220]" box="[189,310,1194,1220]" pageId="123" pageNumber="124">
<heading id="5C1D150FFFE2FFA8A7ADFB45FEF0D8C5" bold="true" box="[189,310,1194,1220]" fontSize="11" level="3" pageId="123" pageNumber="124" reason="3">
<emphasis id="359E7E71FFE2FFA8A7ADFB45FEF0D8C5" bold="true" box="[189,310,1194,1220]" pageId="123" pageNumber="124">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="0755A263FFE2FFA8A7ADFB38FB94D97B" blockId="123.[188,1398,1239,1403]" pageId="123" pageNumber="124">
This species may be separated from other
<taxonomicName id="C0EAD9E0FFE2FFA8A5B5FB38FCD4D8F0" authorityName="Simon" authorityYear="1892" box="[677,786,1239,1265]" class="Arachnida" family="Charinidae" genus="Charinus" kingdom="Animalia" order="Amblypygi" pageId="123" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="359E7E71FFE2FFA8A5B5FB38FCD4D8F0" box="[677,786,1239,1265]" italics="true" pageId="123" pageNumber="124">Charinus</emphasis>
</taxonomicName>
in eastern South America by means of the following combination of characters: cheliceral claw with eleven teeth; pedipalp coxa with three long setae on margin, and two distinct setae encircled by round carina; pedipalp tarsus with two subequal dorsal spines (
<figureCitation id="9FD1BEE6FFE2FFA8A7D4FAD1FEF7D959" box="[196,305,1342,1368]" captionStart="Fig" captionStartId="326.[189,232,1718,1744]" captionTargetBox="[189,1399,265,1667]" captionTargetId="figure-47@326.[189,1399,265,1667]" captionTargetPageId="326" captionText="Fig. 71. Charinus puri sp. nov. (MNRJ 9223), general morphology.A. Carapace, dorsal view.B. Sternum, ventral view. C. Pedipalp tarsus, frontal view. D. Pedipalp, dorsal view. E. Pedipalp, ventral view. Scale bars: A–B, D–E = 1 mm; C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5536573" httpUri="https://zenodo.org/record/5536573/files/figure.png" pageId="123" pageNumber="124">Fig. 71C</figureCitation>
); leg IV basitibia trichobothrium
<emphasis id="359E7E71FFE2FFA8A5AEFAD1FD13D959" box="[702,725,1342,1368]" italics="true" pageId="123" pageNumber="124">bt</emphasis>
situated proximal to submedial; distitibia trichobothrium
<emphasis id="359E7E71FFE2FFA8A7AEFA8FFF1CD97B" box="[190,218,1376,1402]" italics="true" pageId="123" pageNumber="124">bc</emphasis>
situated closer to
<emphasis id="359E7E71FFE2FFA8A6BEFA8FFE16D97B" box="[430,464,1376,1402]" italics="true" pageId="123" pageNumber="124">sbf</emphasis>
than to
<emphasis id="359E7E71FFE2FFA8A520FA8FFD81D97B" box="[560,583,1376,1402]" italics="true" pageId="123" pageNumber="124">bf</emphasis>
,
<emphasis id="359E7E71FFE2FFA8A545FA8EFDABD97B" box="[597,621,1377,1402]" italics="true" pageId="123" pageNumber="124">sc</emphasis>
and
<emphasis id="359E7E71FFE2FFA8A5B7FA8EFD79D97B" box="[679,703,1377,1402]" italics="true" pageId="123" pageNumber="124">sc</emphasis>
series each with six trichobothria.
</paragraph>
<paragraph id="0755A263FFE2FFA8A7ADFA4EFC8DDA42" blockId="123.[188,1399,1441,1603]" pageId="123" pageNumber="124">
This species resembles
<taxonomicName id="C0EAD9E0FFE2FFA8A6C1FA4EFDACD9BA" authority="Miranda &amp; Giupponi &amp; Prendini &amp; Scharff, 2021" authorityName="Miranda &amp; Giupponi &amp; Prendini &amp; Scharff" authorityYear="2021" box="[465,618,1441,1467]" class="Arachnida" family="Charinidae" genus="Charinus" kingdom="Animalia" order="Amblypygi" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="imperialis" status="sp. nov.">
<emphasis id="359E7E71FFE2FFA8A6C1FA4EFDACD9BA" box="[465,618,1441,1467]" italics="true" pageId="123" pageNumber="124">C. imperialis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2EADC30AFFE2FFA8A562FA4EFD0AD9BA" box="[626,716,1441,1467]" pageId="123" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
but possesses more spines on the pedipalp. One specimen (MNRJ 9098) exhibits a spine between dorsal spines 1 and 2 of the femur and a series of three or four small spines parallel to the primary series of dorsal spines, the first spine situated between spines 1 and 2, the second spine ventral to spine 2, and the third between spines 3 and 4. These spines vary in number and position in other individuals. The male is unknown.
</paragraph>
</subSubSection>
<subSubSection id="4FF0F1E8FFE2FFA8A7ADF985FE68DAD2" pageId="123" pageNumber="124" type="etymology">
<paragraph id="0755A263FFE2FFA8A7ADF985FE82DA85" blockId="123.[189,324,1642,1668]" box="[189,324,1642,1668]" pageId="123" pageNumber="124">
<heading id="5C1D150FFFE2FFA8A7ADF985FE82DA85" bold="true" box="[189,324,1642,1668]" fontSize="11" level="3" pageId="123" pageNumber="124" reason="3">
<emphasis id="359E7E71FFE2FFA8A7ADF985FE82DA85" bold="true" box="[189,324,1642,1668]" pageId="123" pageNumber="124">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="0755A263FFE2FFA8A7ADF977FE68DAD2" blockId="123.[189,1399,1687,1748]" pageId="123" pageNumber="124">
Noun in apposition honoring the Puri indigenous tribe who originally inhabited the area where the
<typeStatus id="D8511CC1FFE2FFA8A254F977FAB1DAB3" box="[1348,1399,1688,1714]" pageId="123" pageNumber="124">type</typeStatus>
series was collected.
</paragraph>
</subSubSection>
<subSubSection id="4FF0F1E8FFE2FFA8A7ADF915FB5ADBE9" pageId="123" pageNumber="124" type="materials_examined">
<paragraph id="0755A263FFE2FFA8A7ADF915FEAADB15" blockId="123.[189,364,1786,1812]" box="[189,364,1786,1812]" pageId="123" pageNumber="124">
<heading id="5C1D150FFFE2FFA8A7ADF915FEAADB15" bold="true" box="[189,364,1786,1812]" fontSize="11" level="3" pageId="123" pageNumber="124" reason="3">
<emphasis id="359E7E71FFE2FFA8A7ADF915FEAADB15" bold="true" box="[189,364,1786,1812]" pageId="123" pageNumber="124">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="0755A263FFE2FFA8A7FAF8C8FAB1DB65" blockId="123.[189,1399,1831,1892]" pageId="123" pageNumber="124">
<materialsCitation id="B782A83EFFE2FFA8A7FAF8C8FAB1DB65" ID-GBIF-Occurrence="3383565888" collectionCode="MNRJ" country="Brazil" latitude="-21.560993" location="Cambuci" longLatPrecision="1" longitude="-41.940445" pageId="123" pageNumber="124" specimenCode="MNRJ 9223" specimenCount="1" specimenCount-female="1" stateProvince="Rio de Janeiro" typeStatus="holotype">
<emphasis id="359E7E71FFE2FFA8A7FAF8C8FE9ADB40" bold="true" box="[234,348,1831,1857]" pageId="123" pageNumber="124">
<typeStatus id="D8511CC1FFE2FFA8A7FAF8C8FE9ADB40" box="[234,348,1831,1857]" pageId="123" pageNumber="124" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="7FFDE2F3FFE2FFA8A7ADF8A5FEE1DB65" box="[189,295,1866,1892]" name="Brazil" pageId="123" pageNumber="124">BRAZIL</collectingCountry>
•
<specimenCount id="11EC69EAFFE2FFA8A62EF8A5FEADDB65" box="[318,363,1866,1892]" pageId="123" pageNumber="124" type="female">1 ♀</specimenCount>
;
<collectingRegion id="C52E6C81FFE2FFA8A66AF8A6FDE0DB62" box="[378,550,1865,1891]" country="Brazil" name="Rio de Janeiro" pageId="123" pageNumber="124">Rio de Janeiro</collectingRegion>
,
<location id="0235F4B8FFE2FFA8A521F8A6FD58DB62" LSID="urn:lsid:plazi:treatment:8F431375FFE2FFAFA583FBCAFE71DBE9:0235F4B8FFE2FFA8A521F8A6FD58DB62" box="[561,670,1865,1891]" country="Brazil" latitude="-21.560993" longLatPrecision="1" longitude="-41.940445" name="Cambuci" pageId="123" pageNumber="124" stateProvince="Rio de Janeiro">Cambuci</location>
, unnamed cave;
<geoCoordinate id="62DEC4A4FFE2FFA8A471F8A7FBD3DB65" box="[865,1045,1864,1892]" degrees="21" direction="south" minutes="33" orientation="latitude" pageId="123" pageNumber="124" precision="1" seconds="39.58" value="-21.560993">21°33′39.58″ S</geoCoordinate>
,
<geoCoordinate id="62DEC4A4FFE2FFA8A332F8A7FB15DB65" box="[1058,1235,1864,1892]" degrees="41" direction="west" minutes="56" orientation="longitude" pageId="123" pageNumber="124" precision="1" seconds="25.6" value="-41.940445">41°56′25.6″ W</geoCoordinate>
;
<specimenCode id="574C0A18FFE2FFA8A3CFF8A5FAB4DB65" box="[1247,1394,1866,1892]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="123" pageNumber="124">MNRJ 9223</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="0755A263FFE2FFA8A7FAF865FB5ADBE9" blockId="123.[189,1398,1930,2025]" pageId="123" pageNumber="124">
<materialsCitation id="B782A83EFFE2FFA8A7FAF865FC18DBC7" ID-GBIF-Occurrence="3383565569" collectionCode="MNRJ" country="Brazil" latitude="-21.560993" location="BRAZIL" longLatPrecision="1" longitude="-41.940445" pageId="123" pageNumber="124" specimenCode="MNRJ 9223" specimenCount="4" specimenCount-female="4" stateProvince="Rio de Janeiro" typeStatus="paratype">
<emphasis id="359E7E71FFE2FFA8A7FAF865FEA1DBA5" bold="true" box="[234,359,1930,1956]" pageId="123" pageNumber="124">
<typeStatus id="D8511CC1FFE2FFA8A7FAF865FEA1DBA5" box="[234,359,1930,1956]" pageId="123" pageNumber="124" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="7FFDE2F3FFE2FFA8A7ADF843FEE1DBC7" box="[189,295,1964,1990]" name="Brazil" pageId="123" pageNumber="124">BRAZIL</collectingCountry>
•
<specimenCount id="11EC69EAFFE2FFA8A650F843FE42DBC6" box="[320,388,1964,1991]" pageId="123" pageNumber="124" type="female">4 ♀♀</specimenCount>
; same collection data as for holotype;
<specimenCode id="574C0A18FFE2FFA8A45DF842FC18DBC7" box="[845,990,1964,1991]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="123" pageNumber="124">MNRJ 9223</specimenCode>
</materialsCitation>
•
<materialsCitation id="B782A83EFFE2FFA8A4E8F843FB5EDBE9" ID-GBIF-Occurrence="3383565371" collectingDate="2005-11-12" collectionCode="MNRJ" collectorName="M. S. Silva &amp; R. L. Ferreira" country="Brazil" location="Gruta das Furnas" pageId="123" pageNumber="124" specimenCode="MNRJ 9098" specimenCount="2" specimenCount-female="2" stateProvince="Rio de Janeiro" typeStatus="paratype">
<specimenCount id="11EC69EAFFE2FFA8A4E8F843FBFDDBC6" box="[1016,1083,1964,1991]" pageId="123" pageNumber="124" type="female">2 ♀♀</specimenCount>
;
<collectingRegion id="C52E6C81FFE2FFA8A35BF843FB3FDBC7" box="[1099,1273,1964,1990]" country="Brazil" name="Rio de Janeiro" pageId="123" pageNumber="124">Rio de Janeiro</collectingRegion>
,
<location id="0235F4B8FFE2FFA8A215F843FECBDBE8" LSID="urn:lsid:plazi:treatment:8F431375FFE2FFAFA583FBCAFE71DBE9:0235F4B8FFE2FFA8A215F843FECBDBE8" country="Brazil" name="Gruta das Furnas" pageId="123" pageNumber="124" stateProvince="Rio de Janeiro">Gruta das Furnas</location>
or
<location id="0235F4B8FFE2FFA8A624F821FE7CDBE9" LSID="urn:lsid:plazi:treatment:8F431375FFE2FFAFA583FBCAFE71DBE9:0235F4B8FFE2FFA8A624F821FE7CDBE9" box="[308,442,1998,2024]" country="Brazil" name="Sumidouro" pageId="123" pageNumber="124" stateProvince="Rio de Janeiro">Sumidouro</location>
;
<date id="735484A3FFE2FFA8A6D6F821FDAEDBE8" box="[454,616,1998,2025]" pageId="123" pageNumber="124" value="2005-11-12">
<collectingDate id="63107D4BFFE2FFA8A6D6F821FDAEDBE8" box="[454,616,1998,2025]" pageId="123" pageNumber="124" value="2005-11-12">12 Nov. 2005</collectingDate>
</date>
;
<collectorName id="AA1FC7B5FFE2FFA8A564F821FD36DBE9" box="[628,752,1998,2024]" pageId="123" pageNumber="124">M.S. Silva</collectorName>
and
<collectorName id="AA1FC7B5FFE2FFA8A43AF820FC04DBE9" box="[810,962,1998,2025]" pageId="123" pageNumber="124">R.L. Ferreira</collectorName>
leg.;
<specimenCode id="574C0A18FFE2FFA8A314F820FB5EDBE9" box="[1028,1176,1998,2025]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="123" pageNumber="124">MNRJ 9098</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4FF0F1E8FFE5FFAFA7ADFEE7FE8ADAFE" pageId="124" pageNumber="125" type="description">
<paragraph id="0755A263FFE5FFAFA7ADFEE7FE88DD23" blockId="124.[189,334,264,290]" box="[189,334,264,290]" pageId="124" pageNumber="125">
<heading id="5C1D150FFFE5FFAFA7ADFEE7FE88DD23" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="124" pageNumber="125" reason="3">
<emphasis id="359E7E71FFE5FFAFA7ADFEE7FE88DD23" bold="true" box="[189,334,264,290]" pageId="124" pageNumber="125">Description</emphasis>
</heading>
</paragraph>
<paragraph id="0755A263FFE5FFAFA7ADFED6FCC5DDC2" blockId="124.[189,1398,313,451]" pageId="124" pageNumber="125">
CARAPACE. Six anterior setae (
<figureCitation id="9FD1BEE6FFE5FFAFA53AFED6FD5CDD52" box="[554,666,313,339]" captionStart="Fig" captionStartId="326.[189,232,1718,1744]" captionTargetBox="[189,1399,265,1667]" captionTargetId="figure-47@326.[189,1399,265,1667]" captionTargetPageId="326" captionText="Fig. 71. Charinus puri sp. nov. (MNRJ 9223), general morphology.A. Carapace, dorsal view.B. Sternum, ventral view. C. Pedipalp tarsus, frontal view. D. Pedipalp, dorsal view. E. Pedipalp, ventral view. Scale bars: A–B, D–E = 1 mm; C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5536573" httpUri="https://zenodo.org/record/5536573/files/figure.png" pageId="124" pageNumber="125">Fig. 71A</figureCitation>
); frontal process triangular. Small granules densely scattered between ocular triads and among sulci. Median eyes well developed (
<figureCitation id="9FD1BEE6FFE5FFAFA4F9FEB1FB90DD79" box="[1001,1110,350,376]" captionStart="Fig" captionStartId="326.[189,232,1718,1744]" captionTargetBox="[189,1399,265,1667]" captionTargetId="figure-47@326.[189,1399,265,1667]" captionTargetPageId="326" captionText="Fig. 71. Charinus puri sp. nov. (MNRJ 9223), general morphology.A. Carapace, dorsal view.B. Sternum, ventral view. C. Pedipalp tarsus, frontal view. D. Pedipalp, dorsal view. E. Pedipalp, ventral view. Scale bars: A–B, D–E = 1 mm; C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5536573" httpUri="https://zenodo.org/record/5536573/files/figure.png" pageId="124" pageNumber="125">Fig. 71A</figureCitation>
); median ocular tubercle reduced, with pair of setae; lateral eyes well developed, seta posterior to lateral ocular triad; lateral ocular triad well separated from carapace margin.
</paragraph>
<paragraph id="0755A263FFE5FFAFA7ADFE03FED7DE77" blockId="124.[186,1399,492,630]" pageId="124" pageNumber="125">
STERNUM. Tritosternum projected anteriorly with typical setation, long, surpassing base of pedipalp coxae (
<figureCitation id="9FD1BEE6FFE5FFAFA7D3FDFEFEF7DE2D" box="[195,305,529,556]" captionStart="Fig" captionStartId="326.[189,232,1718,1744]" captionTargetBox="[189,1399,265,1667]" captionTargetId="figure-47@326.[189,1399,265,1667]" captionTargetPageId="326" captionText="Fig. 71. Charinus puri sp. nov. (MNRJ 9223), general morphology.A. Carapace, dorsal view.B. Sternum, ventral view. C. Pedipalp tarsus, frontal view. D. Pedipalp, dorsal view. E. Pedipalp, ventral view. Scale bars: A–B, D–E = 1 mm; C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5536573" httpUri="https://zenodo.org/record/5536573/files/figure.png" pageId="124" pageNumber="125">Fig. 71B</figureCitation>
); other sternal platelets narrow and concave, with pair of setae anteriorly on plaque and some smaller setae posteriorly; pentasternum with eleven setae anteriorly and three setae near membranous region.
</paragraph>
<paragraph id="0755A263FFE5FFAFA7ADFD70FC92DEB8" blockId="124.[189,852,671,698]" box="[189,852,671,698]" pageId="124" pageNumber="125">OPISTHOSOMA. Ventral sacs and ventral sac cover absent.</paragraph>
<paragraph id="0755A263FFE5FFAFA7ADFD0CFE39DF6D" blockId="124.[187,1401,739,877]" pageId="124" pageNumber="125">
GENITALIA. Female genital operculum with prominent setae posteromedially and some smaller setae near margin (
<figureCitation id="9FD1BEE6FFE5FFAFA630FCE7FE72DF23" box="[288,436,776,802]" captionStart="Fig" captionStartId="327.[189,232,1445,1471]" captionTargetBox="[189,1398,265,1393]" captionTargetId="figure-42@327.[189,1399,265,1393]" captionTargetPageId="327" captionText="Fig. 72. Charinus puri sp. nov. (MNRJ 9223), female gonopod and genital operculum. A. Sucker-like gonopod, dorsal view. B. Genital operculum, ventral view. C. Gonopods, posterior view. D. Posterior margin of genital operculum, ventral view. F. Cerotegument." figureDoi="http://doi.org/10.5281/zenodo.5536575" httpUri="https://zenodo.org/record/5536575/files/figure.png" pageId="124" pageNumber="125">Fig. 72A–C</figureCitation>
); gonopod sucker-like and slightly sclerotized basally (
<figureCitation id="9FD1BEE6FFE5FFAFA35AFCE7FAC7DF22" box="[1098,1281,776,803]" captionStart="Fig" captionStartId="327.[189,232,1445,1471]" captionTargetBox="[189,1398,265,1393]" captionTargetId="figure-42@327.[189,1399,265,1393]" captionTargetPageId="327" captionText="Fig. 72. Charinus puri sp. nov. (MNRJ 9223), female gonopod and genital operculum. A. Sucker-like gonopod, dorsal view. B. Genital operculum, ventral view. C. Gonopods, posterior view. D. Posterior margin of genital operculum, ventral view. F. Cerotegument." figureDoi="http://doi.org/10.5281/zenodo.5536575" httpUri="https://zenodo.org/record/5536575/files/figure.png" pageId="124" pageNumber="125">Fig. 72A, D–E</figureCitation>
); conical, without constriction (
<figureCitation id="9FD1BEE6FFE5FFAFA6ACFCC2FDA9DF49" box="[444,623,813,840]" captionStart="Fig" captionStartId="327.[189,232,1445,1471]" captionTargetBox="[189,1398,265,1393]" captionTargetId="figure-42@327.[189,1399,265,1393]" captionTargetPageId="327" captionText="Fig. 72. Charinus puri sp. nov. (MNRJ 9223), female gonopod and genital operculum. A. Sucker-like gonopod, dorsal view. B. Genital operculum, ventral view. C. Gonopods, posterior view. D. Posterior margin of genital operculum, ventral view. F. Cerotegument." figureDoi="http://doi.org/10.5281/zenodo.5536575" httpUri="https://zenodo.org/record/5536575/files/figure.png" pageId="124" pageNumber="125">Fig. 72A, D–E</figureCitation>
); smooth surface between gonopods and distal border of gonopod (
<figureCitation id="9FD1BEE6FFE5FFAFA7D6FCBDFEF4DF6C" box="[198,306,850,877]" captionStart="Fig" captionStartId="327.[189,232,1445,1471]" captionTargetBox="[189,1398,265,1393]" captionTargetId="figure-42@327.[189,1399,265,1393]" captionTargetPageId="327" captionText="Fig. 72. Charinus puri sp. nov. (MNRJ 9223), female gonopod and genital operculum. A. Sucker-like gonopod, dorsal view. B. Genital operculum, ventral view. C. Gonopods, posterior view. D. Posterior margin of genital operculum, ventral view. F. Cerotegument." figureDoi="http://doi.org/10.5281/zenodo.5536575" httpUri="https://zenodo.org/record/5536575/files/figure.png" pageId="124" pageNumber="125">Fig. 72B</figureCitation>
). Male unknown.
</paragraph>
<paragraph id="0755A263FFE5FFAFA7ADFC79FE1ED821" blockId="124.[188,1399,918,1056]" pageId="124" pageNumber="125">CHELICERAE. Small tooth projecting from retrolateral surface of basal segment, opposite to bifid tooth; retrolateral surface of claw with continous row of setae, basally to medially; claw with eleven teeth; row of eighteen setae on prolateral surface of basal segment; bifid tooth on basal segment with dorsal cusp larger than ventral cusp.</paragraph>
<paragraph id="0755A263FFE5FFAFA7ADFBA6FDB9D9D9" blockId="124.[188,1399,1097,1496]" pageId="124" pageNumber="125">
PEDIPALPS. Coxal dorsal carina with two prominent setae encircled by round carina and three setae on margin. Femur with four dorsal spines and five ventral spines (
<figureCitation id="9FD1BEE6FFE5FFAFA48BFB81FBC1D889" box="[923,1031,1134,1160]" captionStart="Fig" captionStartId="326.[189,232,1718,1744]" captionTargetBox="[189,1399,265,1667]" captionTargetId="figure-47@326.[189,1399,265,1667]" captionTargetPageId="326" captionText="Fig. 71. Charinus puri sp. nov. (MNRJ 9223), general morphology.A. Carapace, dorsal view.B. Sternum, ventral view. C. Pedipalp tarsus, frontal view. D. Pedipalp, dorsal view. E. Pedipalp, ventral view. Scale bars: A–B, D–E = 1 mm; C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5536573" httpUri="https://zenodo.org/record/5536573/files/figure.png" pageId="124" pageNumber="125">Fig. 71D</figureCitation>
); spine between dorsal spines 2 and 3, and spines 3 and 4; four prominent setiferous tubercles between dorsal spine 1 and proximal margin; ventral spine between spine 1 and proximal margin (
<figureCitation id="9FD1BEE6FFE5FFAFA46CFB56FC20D8D2" box="[892,998,1209,1235]" captionStart="Fig" captionStartId="326.[189,232,1718,1744]" captionTargetBox="[189,1399,265,1667]" captionTargetId="figure-47@326.[189,1399,265,1667]" captionTargetPageId="326" captionText="Fig. 71. Charinus puri sp. nov. (MNRJ 9223), general morphology.A. Carapace, dorsal view.B. Sternum, ventral view. C. Pedipalp tarsus, frontal view. D. Pedipalp, dorsal view. E. Pedipalp, ventral view. Scale bars: A–B, D–E = 1 mm; C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5536573" httpUri="https://zenodo.org/record/5536573/files/figure.png" pageId="124" pageNumber="125">Fig. 71E</figureCitation>
); small spine ventral to extra spine proximal to spine 1, near articulation between femur and trochanter; spine between ventral spines 1 and 2, two spines between spines 2 and 3, and between spines 3 and 4. Patella with six dorsal spines in primary series (
<figureCitation id="9FD1BEE6FFE5FFAFA662FAC7FE1AD943" box="[370,476,1320,1346]" captionStart="Fig" captionStartId="326.[189,232,1718,1744]" captionTargetBox="[189,1399,265,1667]" captionTargetId="figure-47@326.[189,1399,265,1667]" captionTargetPageId="326" captionText="Fig. 71. Charinus puri sp. nov. (MNRJ 9223), general morphology.A. Carapace, dorsal view.B. Sternum, ventral view. C. Pedipalp tarsus, frontal view. D. Pedipalp, dorsal view. E. Pedipalp, ventral view. Scale bars: A–B, D–E = 1 mm; C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5536573" httpUri="https://zenodo.org/record/5536573/files/figure.png" pageId="124" pageNumber="125">Fig. 71E</figureCitation>
); prominent setiferous tubercle distal to spine I, one-third length of spine I; four or five ventral spines (
<figureCitation id="9FD1BEE6FFE5FFAFA6D1FAA2FDECD969" box="[449,554,1357,1384]" captionStart="Fig" captionStartId="326.[189,232,1718,1744]" captionTargetBox="[189,1399,265,1667]" captionTargetId="figure-47@326.[189,1399,265,1667]" captionTargetPageId="326" captionText="Fig. 71. Charinus puri sp. nov. (MNRJ 9223), general morphology.A. Carapace, dorsal view.B. Sternum, ventral view. C. Pedipalp tarsus, frontal view. D. Pedipalp, dorsal view. E. Pedipalp, ventral view. Scale bars: A–B, D–E = 1 mm; C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5536573" httpUri="https://zenodo.org/record/5536573/files/figure.png" pageId="124" pageNumber="125">Fig. 71F</figureCitation>
); prominent setiferous tubercle between spine I and distal margin; small spine between spines 2 and 3. Tibia with ventral spine distally and four setae between spine and distal margin. Tarsus with two dorsal spines, distal spine twice as long as proximal spine (
<figureCitation id="9FD1BEE6FFE5FFAFA383FA77FB39D9B3" box="[1171,1279,1432,1458]" captionStart="Fig" captionStartId="326.[189,232,1718,1744]" captionTargetBox="[189,1399,265,1667]" captionTargetId="figure-47@326.[189,1399,265,1667]" captionTargetPageId="326" captionText="Fig. 71. Charinus puri sp. nov. (MNRJ 9223), general morphology.A. Carapace, dorsal view.B. Sternum, ventral view. C. Pedipalp tarsus, frontal view. D. Pedipalp, dorsal view. E. Pedipalp, ventral view. Scale bars: A–B, D–E = 1 mm; C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5536573" httpUri="https://zenodo.org/record/5536573/files/figure.png" pageId="124" pageNumber="125">Fig. 71C</figureCitation>
); cleaning organ with 27–29 setae in ventral row.
</paragraph>
<paragraph id="0755A263FFE5FFAFA7ADF9EEFAC3DA8B" blockId="124.[189,1399,1537,1675]" pageId="124" pageNumber="125">
LEGS. Tibia of leg I with 23 articles; tarsus I with 41 articles; first tarsal article shorter than second article. Leg IV basitibia with four pseudo-articles, with sclerotized, denticulate margin projecting from apex of articles; basitibia trichobothrium
<emphasis id="359E7E71FFE5FFAFA5B4F9A4FD7DDA64" box="[676,699,1611,1637]" italics="true" pageId="124" pageNumber="125">bt</emphasis>
situated proximal to submedial on pseudo-article; distitibia trichobothrium
<emphasis id="359E7E71FFE5FFAFA664F99FFE56DA8B" box="[372,400,1648,1674]" italics="true" pageId="124" pageNumber="125">bc</emphasis>
situated closer to
<emphasis id="359E7E71FFE5FFAFA575F99FFD41DA8B" box="[613,647,1648,1674]" italics="true" pageId="124" pageNumber="125">sbf</emphasis>
than to
<emphasis id="359E7E71FFE5FFAFA5F7F99FFD38DA8B" box="[743,766,1648,1674]" italics="true" pageId="124" pageNumber="125">bf</emphasis>
,
<emphasis id="359E7E71FFE5FFAFA41CF99EFCE2DA8B" box="[780,804,1649,1674]" italics="true" pageId="124" pageNumber="125">sc</emphasis>
and
<emphasis id="359E7E71FFE5FFAFA44EF99FFCB4DA8B" box="[862,882,1648,1674]" italics="true" pageId="124" pageNumber="125">sf</emphasis>
series each with six trichobothria.
</paragraph>
<paragraph id="0755A263FFE5FFAFA7ADF95BFEB3DACF" blockId="124.[189,373,1716,1742]" box="[189,373,1716,1742]" pageId="124" pageNumber="125">
<heading id="5C1D150FFFE5FFAFA7ADF95BFEB3DACF" bold="true" box="[189,373,1716,1742]" fontSize="11" level="3" pageId="124" pageNumber="125" reason="3">
<emphasis id="359E7E71FFE5FFAFA7ADF95BFEB3DACF" bold="true" box="[189,373,1716,1742]" pageId="124" pageNumber="125">Measurements</emphasis>
</heading>
</paragraph>
<paragraph id="0755A263FFE5FFAFA7ADF90AFE8ADAFE" blockId="124.[189,332,1764,1791]" box="[189,332,1764,1791]" pageId="124" pageNumber="125">
See
<tableCitation id="4A6897D8FFE5FFAFA7FEF90BFE8EDAFE" box="[238,328,1764,1791]" captionStart="Table 4" captionStartId="89.[181,251,552,581]" captionTargetBox="[185,1925,631,1740]" captionTargetId="graphics-20@89.[181,1932,659,1753]" captionTargetPageId="89" captionText="Table 4 (continued on next five pages). Measurements of all eastern South American species of Charinus Simon, 1892." httpUri="http://table.plazi.org/id/5395F2EBFFC0FF8AA7A5FDC7F9F2DE45" pageId="124" pageNumber="125" tableUuid="5395F2EBFFC0FF8AA7A5FDC7F9F2DE45">Table 4</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4FF0F1E8FFE5FFAFA7ADF8C7FD9FDB73" box="[189,601,1832,1907]" pageId="124" pageNumber="125" type="distribution">
<paragraph id="0755A263FFE5FFAFA7ADF8C7FE90DB43" blockId="124.[189,342,1832,1858]" box="[189,342,1832,1858]" pageId="124" pageNumber="125">
<heading id="5C1D150FFFE5FFAFA7ADF8C7FE90DB43" bold="true" box="[189,342,1832,1858]" fontSize="11" level="3" pageId="124" pageNumber="125" reason="3">
<emphasis id="359E7E71FFE5FFAFA7ADF8C7FE90DB43" bold="true" box="[189,342,1832,1858]" pageId="124" pageNumber="125">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="0755A263FFE5FFAFA7ADF8B6FD9FDB73" blockId="124.[189,601,1880,1907]" box="[189,601,1880,1907]" pageId="124" pageNumber="125">
Known only from the
<typeStatus id="D8511CC1FFE5FFAFA6D2F8B6FE33DB72" box="[450,501,1881,1907]" pageId="124" pageNumber="125">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="4FF0F1E8FFE5FFAFA7ADF873FE71DBE9" pageId="124" pageNumber="125" type="biology_ecology">
<paragraph id="0755A263FFE5FFAFA7ADF873FEB8DBB7" blockId="124.[189,382,1948,1974]" box="[189,382,1948,1974]" pageId="124" pageNumber="125">
<heading id="5C1D150FFFE5FFAFA7ADF873FEB8DBB7" bold="true" box="[189,382,1948,1974]" fontSize="11" level="3" pageId="124" pageNumber="125" reason="3">
<emphasis id="359E7E71FFE5FFAFA7ADF873FEB8DBB7" bold="true" box="[189,382,1948,1974]" pageId="124" pageNumber="125">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="0755A263FFE5FFAFA7ADF821FE71DBE9" blockId="124.[189,439,1998,2025]" box="[189,439,1998,2025]" pageId="124" pageNumber="125">Found on cave walls.</paragraph>
</subSubSection>
</treatment>
</document>
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