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<document id="05C881DE7866873B1A64B4AA3768CEB3" ID-CLB-Dataset="4042" ID-DOI="10.5852/ejt.2021.772.1505" ID-GBIF-Dataset="b65bb5a9-bbe7-49a4-af44-4b4c03121288" ID-ISSN="2118-9773" ID-Zenodo-Dep="5536410" ID-ZooBank="9B82A32F-0A07-47E3-8684-FED7C8EBF1E9" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1632919496065" checkinUser="felipe" docAuthor="Miranda, Gustavo Silva de, Giupponi, Alessandro P. L., Prendini, Lorenzo &amp; Scharff, Nikolaj" docDate="2021" docId="8F431375FF45FF0EA6B6FEE6FEDCDD6B" docLanguage="en" docName="ejt-772_1-409.pdf" docOrigin="European Journal of Taxonomy 772" docSource="http://dx.doi.org/10.5852/ejt.2021.772.1505" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Sarax dhofarensis Miranda &amp; Giupponi &amp; Prendini &amp; Scharff 2021, comb. nov." docType="treatment" docVersion="14" lastPageNumber="222" masterDocId="737A6B0DFF99FFD3A710FFEFFFC6DC01" masterDocTitle="Systematic revision of the pantropical whip spider family Charinidae Quintero, 1986 (Arachnida, Amblypygi)" masterLastPageNumber="409" masterPageNumber="1" pageNumber="221" updateTime="1699416367315" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="0AB38906939A57AB169EC591CB617C39">Systematic revision of the pantropical whip spider family Charinidae Quintero, 1986 (Arachnida, Amblypygi)</mods:title>
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<mods:namePart id="60ECB900DCC1E1D673493900116CD627">Miranda, Gustavo Silva de</mods:namePart>
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<mods:affiliation id="35ED5107586E831AC001378F8B7E42B4">Center for Macroecology, Evolution and Climate, Natural History Museum of Denmark, University of Copenhagen, Copenhagen, Denmark. &amp; Entomology Department, National Museum of Natural History, Smithsonian Institution, Washington DC, USA.</mods:affiliation>
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<mods:namePart id="2973FA0D508801D6E290F3056D66E363">Giupponi, Alessandro P. L.</mods:namePart>
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<mods:affiliation id="E7666023B316BFBF0840A1A222C095EB">Laboratório de Referência Nacional em Vetores das Riquetsioses, LIRN-IOC-FIOCRUZ, Rio de Janeiro, RJ, Brazil.</mods:affiliation>
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<mods:namePart id="F24EA7FE7A1C4B3B3B765339CCA81F08">Prendini, Lorenzo</mods:namePart>
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<mods:affiliation id="00B984784532BE756B8C1E34E059BE1F">Arachnology Lab, Division of Invertebrate Zoology, American Museum of Natural History, New York, NY, USA.</mods:affiliation>
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<mods:namePart id="C427CE091F370B6DFA593C28AC6998DF">Scharff, Nikolaj</mods:namePart>
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<mods:affiliation id="4426ADF05D54D156CBE715A9BDEFB6F5">Center for Macroecology, Evolution and Climate, Natural History Museum of Denmark, University of Copenhagen, Copenhagen, Denmark. &amp; Entomology Department, National Museum of Natural History, Smithsonian Institution, Washington DC, USA. &amp; Zoology Section, Natural History Museum of Denmark, University of Copenhagen, Denmark.</mods:affiliation>
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<mods:date id="E943E9CB134E287C9B823CBFD479D42C">2021</mods:date>
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<heading id="5C1D150FFF45FF0FA6B6FEE6FB48DD22" box="[422,1166,264,291]" centered="true" fontSize="11" level="2" pageId="220" pageNumber="221" reason="2">
<taxonomicName id="C0EAD9E0FF45FF0FA6B6FEE6FBC3DD22" ID-CoL="8VMX4" authority="(Weygoldt, Pohl &amp; Polak, 2002)" authorityName="Miranda &amp; Giupponi &amp; Prendini &amp; Scharff" authorityYear="2021" baseAuthorityName="Weygoldt, Pohl &amp; Polak" baseAuthorityYear="2002" box="[422,1029,264,291]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="dhofarensis" status="comb. nov.">
<emphasis id="359E7E71FF45FF0FA6B6FEE6FD45DD22" bold="true" box="[422,643,264,291]" italics="true" pageId="220" pageNumber="221">Sarax dhofarensis</emphasis>
(Weygoldt, Pohl &amp; Polak, 2002)
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<taxonomicNameLabel id="2EADC30AFF45FF0FA31CFEE6FB48DD22" box="[1036,1166,265,291]" pageId="220" pageNumber="221" rank="species">comb. nov.</taxonomicNameLabel>
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<figureCitation id="9FD1BEE6FF45FF0FA5A6FEC2FCDDDD49" box="[694,795,301,328]" captionStart="Fig" captionStartId="390.[189,232,1237,1263]" captionTargetBox="[192,1396,268,1195]" captionTargetId="figure-21@390.[189,1398,265,1198]" captionTargetPageId="390" captionText="Fig. 135. Map plotting known distributions of species of Sarax Simon, 1892 in Africa, the Middle East and South Asia." figureDoi="http://doi.org/10.5281/zenodo.5536776" httpUri="https://zenodo.org/record/5536776/files/figure.png" pageId="220" pageNumber="221">Fig. 135</figureCitation>
;
<tableCitation id="4A6897D8FF45FF0FA437FEC2FCB8DD49" box="[807,894,301,328]" captionStart="Table 9" captionStartId="213.[181,251,576,605]" captionTargetId="graphics-0@213.[181,1934,690,1746]" captionTargetPageId="213" captionText="Table 9 (continued on next page). Measurements of all African, Middle Eastern and South Asian Sarax Simon, 1892." pageId="220" pageNumber="221">Table 9</tableCitation>
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<treatmentCitationGroup id="27FA854DFF45FF0FA7ADFE9FFC60DD8B" box="[189,934,368,395]" pageId="220" pageNumber="221">
<treatmentCitation id="864B8472FF45FF0FA7ADFE9FFCF0DD8A" author="Weygoldt P. &amp; Hans P. &amp; Polak S." box="[189,822,368,395]" page="295" pageId="220" pageNumber="221" year="2002">
<taxonomicName id="C0EAD9E0FF45FF0FA7ADFE9FFCF0DD8A" ID-CoL="TNV3" authority="Weygoldt et al., 2002: 292 - 295" authorityName="Weygoldt et al." authorityPageNumber="292 - 295" authorityYear="2002" box="[189,822,368,395]" class="Arachnida" family="Charinidae" genus="Charinus" kingdom="Animalia" order="Amblypygi" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="dhofarensis">
<emphasis id="359E7E71FF45FF0FA7ADFE9FFE7CDD8B" box="[189,442,368,394]" italics="true" pageId="220" pageNumber="221">Charinus dhofarensis</emphasis>
<bibRefCitation id="637BDF92FF45FF0FA6D0FE9FFCF0DD8A" author="Weygoldt P. &amp; Hans P. &amp; Polak S." box="[448,822,368,395]" etAl="et al." firstAuthor="Weygoldt" journalOrPublisher="Fauna of Arabia" page="292 - 295" pageId="220" pageNumber="221" pagination="289 - 309" part="19" refId="ref156579" refString="Weygoldt P., Hans P. &amp; Polak S. 2002. Arabian whip spiders: four new species of the genera Charinus and Phrynichus (Chelicerata: Amblypygi) from Oman and Socotra. Fauna of Arabia 19: 289 - 309." title="Arabian whip spiders: four new species of the genera Charinus and Phrynichus (Chelicerata: Amblypygi) from Oman and Socotra" type="journal article" year="2002">
Weygoldt
<emphasis id="359E7E71FF45FF0FA529FE9EFDB2DD8B" box="[569,628,368,394]" italics="true" pageId="220" pageNumber="221">et al.</emphasis>
, 2002: 292–295
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, figs 1–7.
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<treatmentCitationGroup id="27FA854DFF45FF0FA7ADFE5CFD39DDCC" box="[189,767,435,462]" pageId="220" pageNumber="221">
<taxonomicName id="C0EAD9E0FF45FF0FA7ADFE5CFE7CDDCC" ID-CoL="TNV3" authorityName="Weygoldt et al." authorityYear="2002" box="[189,442,435,461]" class="Arachnida" family="Charinidae" genus="Charinus" kingdom="Animalia" order="Amblypygi" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="dhofarensis">
<emphasis id="359E7E71FF45FF0FA7ADFE5CFE7CDDCC" box="[189,442,435,461]" italics="true" pageId="220" pageNumber="221">Charinus dhofarensis</emphasis>
</taxonomicName>
–
<treatmentCitation id="864B8472FF45FF0FA6C7FE5CFD3ADDCC" author="Miranda G. S. &amp; Milleri-Pinto M. &amp; Goncalves-Souza T. &amp; Giupponi A. P. L. &amp; Scharff N." box="[471,764,435,461]" page="29" pageId="220" pageNumber="221" year="2016">
<bibRefCitation id="637BDF92FF45FF0FA6C7FE5CFD3ADDCC" author="Miranda G. S. &amp; Milleri-Pinto M. &amp; Goncalves-Souza T. &amp; Giupponi A. P. L. &amp; Scharff N." box="[471,764,435,461]" etAl="et al." firstAuthor="Miranda" journalOrPublisher="ZooKeys" page="29" pageId="220" pageNumber="221" pagination="15 - 36" part="621" refId="ref151278" refString="Miranda G. S., Milleri-Pinto M., Goncalves-Souza T., Giupponi A. P. L. &amp; Scharff N. 2016 c. A new species of Charinus Simon, 1892 (Amblypygi, Charinidae) from Brazil, with notes on behavior. ZooKeys 621: 15 - 36. https: // doi. org / 10.3897 / zookeys. 621.9980" title="A new species of Charinus Simon, 1892 (Amblypygi, Charinidae) from Brazil, with notes on behavior" type="journal article" year="2016" yearSuffix="c">
<collectingRegion id="C52E6C81FF45FF0FA6C7FE5CFDFCDDCC" box="[471,570,435,461]" country="Venezuela" name="Miranda" pageId="220" pageNumber="221">Miranda</collectingRegion>
<emphasis id="359E7E71FF45FF0FA551FE5BFDBADDCC" box="[577,636,435,461]" italics="true" pageId="220" pageNumber="221">et al.</emphasis>
2016c: 29
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</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="4FF0F1E8FF45FF0FA7ADFE1AFEEDDF1A" pageId="220" pageNumber="221" type="diagnosis">
<paragraph id="0755A263FF45FF0FA7ADFE1AFEF0DE0E" blockId="220.[189,310,501,527]" box="[189,310,501,527]" pageId="220" pageNumber="221">
<heading id="5C1D150FFF45FF0FA7ADFE1AFEF0DE0E" bold="true" box="[189,310,501,527]" fontSize="11" level="3" pageId="220" pageNumber="221" reason="3">
<emphasis id="359E7E71FF45FF0FA7ADFE1AFEF0DE0E" bold="true" box="[189,310,501,527]" pageId="220" pageNumber="221">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="0755A263FF45FF0FA7ADFDC9FEEDDF1A" blockId="220.[189,1400,550,795]" pageId="220" pageNumber="221">
Based on
<bibRefCitation id="637BDF92FF45FF0FA626FDC9FD8CDE41" author="Weygoldt P. &amp; Hans P. &amp; Polak S." box="[310,586,550,576]" etAl="et al." firstAuthor="Weygoldt" journalOrPublisher="Fauna of Arabia" pageId="220" pageNumber="221" pagination="289 - 309" part="19" refId="ref156579" refString="Weygoldt P., Hans P. &amp; Polak S. 2002. Arabian whip spiders: four new species of the genera Charinus and Phrynichus (Chelicerata: Amblypygi) from Oman and Socotra. Fauna of Arabia 19: 289 - 309." title="Arabian whip spiders: four new species of the genera Charinus and Phrynichus (Chelicerata: Amblypygi) from Oman and Socotra" type="journal article" year="2002">
Weygoldt
<emphasis id="359E7E71FF45FF0FA6A1FDC9FE29DE41" box="[433,495,550,576]" italics="true" pageId="220" pageNumber="221">et al.</emphasis>
(2002)
</bibRefCitation>
, this cavernicolous species may be separated from other species of
<taxonomicName id="C0EAD9E0FF45FF0FA7ADFDA4FEC7DE64" authorityName="Simon" authorityYear="1892" box="[189,257,587,613]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="220" pageNumber="221" phylum="Arthropoda" rank="genus">
<emphasis id="359E7E71FF45FF0FA7ADFDA4FEC7DE64" box="[189,257,587,613]" italics="true" pageId="220" pageNumber="221">Sarax</emphasis>
</taxonomicName>
in Africa, the Middle East and South Asia as follows: it differs from
<taxonomicName id="C0EAD9E0FF45FF0FA357FDA4FB25DE65" authorityName="Miranda &amp; Giupponi &amp; Prendini &amp; Scharff" authorityYear="2021" baseAuthorityName="Delle Cave, Gardner &amp; Weygoldt" baseAuthorityYear="2009" box="[1095,1251,587,613]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="omanensis" status="comb. nov.">
<emphasis id="359E7E71FF45FF0FA357FDA4FB25DE65" box="[1095,1251,587,613]" italics="true" pageId="220" pageNumber="221">S. omanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2EADC30AFF45FF0FA3FCFDA5FAB4DE64" box="[1260,1394,586,613]" pageId="220" pageNumber="221" rank="species">comb. nov.</taxonomicNameLabel>
,
<taxonomicName id="C0EAD9E0FF45FF0FA7ADFD80FE9BDE88" authorityName="Miranda &amp; Giupponi &amp; Prendini &amp; Scharff" authorityYear="2021" baseAuthorityName="Weygoldt, Pohl &amp; Polak" baseAuthorityYear="2002" box="[189,349,623,649]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="socotranus" status="comb. nov.">
<emphasis id="359E7E71FF45FF0FA7ADFD80FE9BDE88" box="[189,349,623,649]" italics="true" pageId="220" pageNumber="221">S. socotranus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2EADC30AFF45FF0FA676FD80FE2FDE88" box="[358,489,623,649]" pageId="220" pageNumber="221" rank="species">comb. nov.</taxonomicNameLabel>
and
<taxonomicName id="C0EAD9E0FF45FF0FA537FD80FD32DE88" authorityName="Miranda &amp; Giupponi &amp; Prendini &amp; Scharff" authorityYear="2021" baseAuthorityName="Weygoldt &amp; Van Damme" baseAuthorityYear="2004" box="[551,756,623,649]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="stygochthobius" status="comb. nov.">
<emphasis id="359E7E71FF45FF0FA537FD80FD32DE88" box="[551,756,623,649]" italics="true" pageId="220" pageNumber="221">S. stygochthobius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2EADC30AFF45FF0FA5EDFD80FC45DE88" box="[765,899,623,649]" pageId="220" pageNumber="221" rank="species">comb. nov.</taxonomicNameLabel>
, in having a plunger-like female gonopod and not possessing a slender, tubular female gonopod and the convex genital operculum, without an acuminate apex.
<taxonomicName id="C0EAD9E0FF45FF0FA691FD57FD93DED3" authorityName="Miranda &amp; Giupponi &amp; Prendini &amp; Scharff" authorityYear="2021" baseAuthorityName="Weygoldt, Pohl &amp; Polak" baseAuthorityYear="2002" box="[385,597,696,722]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="dhofarensis" status="comb. nov.">
<emphasis id="359E7E71FF45FF0FA691FD57FD93DED3" box="[385,597,696,722]" italics="true" pageId="220" pageNumber="221">Sarax dhofarensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2EADC30AFF45FF0FA54CFD57FD1BDED3" box="[604,733,696,722]" pageId="220" pageNumber="221" rank="species">comb. nov.</taxonomicNameLabel>
also differs from
<taxonomicName id="C0EAD9E0FF45FF0FA4BBFD57FB85DED3" authorityName="Miranda &amp; Giupponi &amp; Prendini &amp; Scharff" authorityYear="2021" baseAuthorityName="Delle Cave, Gardner &amp; Weygoldt" baseAuthorityYear="2009" box="[939,1091,696,722]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="omanensis" status="comb. nov.">
<emphasis id="359E7E71FF45FF0FA4BBFD57FB85DED3" box="[939,1091,696,722]" italics="true" pageId="220" pageNumber="221">S. omanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2EADC30AFF45FF0FA35AFD57FB08DED3" box="[1098,1230,696,722]" pageId="220" pageNumber="221" rank="species">comb. nov.</taxonomicNameLabel>
,
<taxonomicName id="C0EAD9E0FF45FF0FA3C9FD57FAB1DED3" authorityName="Miranda &amp; Giupponi &amp; Prendini &amp; Scharff" authorityYear="2021" baseAuthorityName="Weygoldt, Pohl &amp; Polak" baseAuthorityYear="2002" box="[1241,1399,696,722]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="socotranus">
<emphasis id="359E7E71FF45FF0FA3C9FD57FAB1DED3" box="[1241,1399,696,722]" italics="true" pageId="220" pageNumber="221">S. socotranus</emphasis>
</taxonomicName>
and
<taxonomicName id="C0EAD9E0FF45FF0FA7FEFD32FE7DDEF7" authorityName="Miranda &amp; Giupponi &amp; Prendini &amp; Scharff" authorityYear="2021" baseAuthorityName="Weygoldt &amp; Van Damme" baseAuthorityYear="2004" box="[238,443,732,759]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="stygochthobius" status="comb. nov.">
<emphasis id="359E7E71FF45FF0FA7FEFD32FE7DDEF7" box="[238,443,732,759]" italics="true" pageId="220" pageNumber="221">S. stygochthobius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2EADC30AFF45FF0FA6D2FD33FD82DEF6" box="[450,580,732,759]" pageId="220" pageNumber="221" rank="species">comb. nov.</taxonomicNameLabel>
in the curved carina between the lateral eyes and lateral margin of the carapace.
</paragraph>
</subSubSection>
<subSubSection id="4FF0F1E8FF45FF0FA7ADFCABFECFDFB3" pageId="220" pageNumber="221" type="etymology">
<paragraph id="0755A263FF45FF0FA7ADFCABFE82DF5F" blockId="220.[189,324,836,862]" box="[189,324,836,862]" pageId="220" pageNumber="221">
<heading id="5C1D150FFF45FF0FA7ADFCABFE82DF5F" bold="true" box="[189,324,836,862]" fontSize="11" level="3" pageId="220" pageNumber="221" reason="3">
<emphasis id="359E7E71FF45FF0FA7ADFCABFE82DF5F" bold="true" box="[189,324,836,862]" pageId="220" pageNumber="221">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="0755A263FF45FF0FA7ADFC9CFECFDFB3" blockId="220.[189,1399,883,946]" pageId="220" pageNumber="221">
Adjective referring to
<collectingRegion id="C52E6C81FF45FF0FA6AFFC9CFDD2DF8C" box="[447,532,883,909]" country="Oman" name="Zufar" pageId="220" pageNumber="221">Dhofar</collectingRegion>
, the southern province of
<collectingCountry id="7FFDE2F3FF45FF0FA450FC9BFC4DDF8F" box="[832,907,884,910]" name="Oman" pageId="220" pageNumber="221">Oman</collectingCountry>
, where the species occurs (
<bibRefCitation id="637BDF92FF45FF0FA3D3FC9CFF3BDFB3" author="Weygoldt P. &amp; Hans P. &amp; Polak S." etAl="et al." firstAuthor="Weygoldt" journalOrPublisher="Fauna of Arabia" pageId="220" pageNumber="221" pagination="289 - 309" part="19" refId="ref156579" refString="Weygoldt P., Hans P. &amp; Polak S. 2002. Arabian whip spiders: four new species of the genera Charinus and Phrynichus (Chelicerata: Amblypygi) from Oman and Socotra. Fauna of Arabia 19: 289 - 309." title="Arabian whip spiders: four new species of the genera Charinus and Phrynichus (Chelicerata: Amblypygi) from Oman and Socotra" type="journal article" year="2002">
Weygoldt
<emphasis id="359E7E71FF45FF0FA22DFC9BFAB1DF8C" box="[1341,1399,883,909]" italics="true" pageId="220" pageNumber="221">et al.</emphasis>
2002
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4FF0F1E8FF45FF0FA7ADFC34FC4AD966" pageId="220" pageNumber="221" type="materials_examined">
<paragraph id="0755A263FF45FF0FA7ADFC34FEAADFF5" blockId="220.[189,364,986,1013]" box="[189,364,986,1013]" pageId="220" pageNumber="221">
<heading id="5C1D150FFF45FF0FA7ADFC34FEAADFF5" bold="true" box="[189,364,986,1013]" fontSize="11" level="3" pageId="220" pageNumber="221" reason="3">
<emphasis id="359E7E71FF45FF0FA7ADFC34FEAADFF5" bold="true" box="[189,364,986,1013]" pageId="220" pageNumber="221">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="0755A263FF45FF0FA7FAFBE4FEB1D86F" blockId="220.[189,1399,1035,1134]" pageId="220" pageNumber="221">
<materialsCitation id="B782A83EFF45FF0FA7FAFBE4FEB1D86F" ID-GBIF-Occurrence="3383565364" collectionCode="ONHM" collectorName="S. Polak" country="Oman" latitude="17.112501" location="Tawi Atayr Waterhole" longLatPrecision="21" longitude="54.37222" pageId="220" pageNumber="221" specimenCode="ONHM 2969" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="359E7E71FF45FF0FA7FAFBE4FE9AD824" bold="true" box="[234,348,1035,1061]" pageId="220" pageNumber="221">
<typeStatus id="D8511CC1FF45FF0FA7FAFBE4FE9AD824" box="[234,348,1035,1061]" pageId="220" pageNumber="221" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="7FFDE2F3FF45FF0FA7ADFBC0FED1D848" box="[189,279,1071,1097]" name="Oman" pageId="220" pageNumber="221">OMAN</collectingCountry>
•
<specimenCount id="11EC69EAFF45FF0FA620FBDFFE81D84B" box="[304,327,1072,1098]" pageId="220" pageNumber="221" type="female">♀</specimenCount>
;
<location id="0235F4B8FF45FF0FA645FBC0FD9FD848" LSID="urn:lsid:plazi:treatment:8F431375FF45FF0EA6B6FEE6FEDCDD6B:0235F4B8FF45FF0FA645FBC0FD9FD848" box="[341,601,1071,1097]" country="Oman" latitude="17.112501" longLatPrecision="21" longitude="54.37222" name="Tawi Atayr Waterhole" pageId="220" pageNumber="221">Tawi Atayr Waterhole</location>
,
<location id="0235F4B8FF45FF0FA574FBC0FD2DD848" LSID="urn:lsid:plazi:treatment:8F431375FF45FF0EA6B6FEE6FEDCDD6B:0235F4B8FF45FF0FA574FBC0FD2DD848" box="[612,747,1071,1097]" country="Oman" latitude="17.112501" longLatPrecision="21" longitude="54.37222" name="Tawi Atayr" pageId="220" pageNumber="221">Tawi Atayr</location>
;
<geoCoordinate id="62DEC4A4FF45FF0FA5E6FBC1FC4AD84B" box="[758,908,1070,1098]" degrees="17" direction="north" minutes="06" orientation="latitude" pageId="220" pageNumber="221" precision="15" seconds="45" value="17.112501">17°06′45″ N</geoCoordinate>
,
<geoCoordinate id="62DEC4A4FF45FF0FA488FBC1FBEDD84B" box="[920,1067,1070,1098]" degrees="54" direction="east" minutes="22" orientation="longitude" pageId="220" pageNumber="221" precision="15" seconds="20" value="54.37222">54°22′20″ E</geoCoordinate>
;
<collectorName id="AA1FC7B5FF45FF0FA327FBC0FB5FD848" box="[1079,1177,1071,1097]" pageId="220" pageNumber="221">S. Polak</collectorName>
leg.;
<specimenCode id="574C0A18FF45FF0FA3CBFBC0FAB1D848" box="[1243,1399,1071,1097]" pageId="220" pageNumber="221">ONHM 2969</specimenCode>
[not examined].
</materialsCitation>
</paragraph>
<paragraph id="0755A263FF45FF0FA7FAFB78FC4AD966" blockId="220.[189,1399,1175,1384]" pageId="220" pageNumber="221">
<materialsCitation id="B782A83EFF45FF0FA7FAFB78FBA8D8D4" ID-GBIF-Occurrence="3383565668" collectionCode="ONHM" collectorName="S. Polak" country="Oman" latitude="17.112501" location="OMAN" longLatPrecision="21" longitude="54.37222" pageId="220" pageNumber="221" specimenCode="ONHM 2970" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="359E7E71FF45FF0FA7FAFB78FEA1D8B0" bold="true" box="[234,359,1175,1201]" pageId="220" pageNumber="221">
<typeStatus id="D8511CC1FF45FF0FA7FAFB78FEA1D8B0" box="[234,359,1175,1201]" pageId="220" pageNumber="221" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="7FFDE2F3FF45FF0FA7ADFB54FED1D8D4" box="[189,279,1211,1237]" name="Oman" pageId="220" pageNumber="221">OMAN</collectingCountry>
•
<specimenCount id="11EC69EAFF45FF0FA63EFB54FE9BD8D7" box="[302,349,1211,1238]" pageId="220" pageNumber="221" type="male">1 ♂</specimenCount>
; same collection data as for holotype;
<specimenCode id="574C0A18FF45FF0FA40BFB54FC70D8D4" box="[795,950,1211,1237]" pageId="220" pageNumber="221">ONHM 2970</specimenCode>
[not examined]
</materialsCitation>
•
<materialsCitation id="B782A83EFF45FF0FA396FB54FD01D8FB" ID-GBIF-Occurrence="3383565781" collectionCode="SMNS" collectorName="S. Polak" country="Oman" latitude="17.112501" location="OMAN" longLatPrecision="21" longitude="54.37222" pageId="220" pageNumber="221" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="11EC69EAFF45FF0FA396FB54FB74D8D7" box="[1158,1202,1211,1238]" pageId="220" pageNumber="221" type="female">1 ♀</specimenCount>
; same collection data as for holotype;
<collectionCode id="61FB3AA6FF45FF0FA6ABFB0FFDCDD8FB" box="[443,523,1248,1274]" collectionName="Germany, Stuttgart, Staatliches Museum fuer Naturkunde" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="220" pageNumber="221">SMNS</collectionCode>
[not examined]
</materialsCitation>
•
<materialsCitation id="B782A83EFF45FF0FA5F3FB0FFCC5D91F" ID-GBIF-Occurrence="3383565596" collectingDate="1997-11-02" collectionCode="ONHM" collectorName="S. Polak" country="Oman" latitude="17.112501" location="Tawi Atayr" longLatPrecision="21" longitude="54.558334" municipality="Pop Cave" pageId="220" pageNumber="221" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="11EC69EAFF45FF0FA5F3FB0FFCD4D8FB" box="[739,786,1248,1274]" pageId="220" pageNumber="221" type="female">1 ♀</specimenCount>
;
<location id="0235F4B8FF45FF0FA431FB30FC6ED8FB" LSID="urn:lsid:plazi:treatment:8F431375FF45FF0EA6B6FEE6FEDCDD6B:0235F4B8FF45FF0FA431FB30FC6ED8FB" box="[801,936,1247,1274]" country="Oman" latitude="17.112501" longLatPrecision="21" longitude="54.558334" municipality="Pop Cave" name="Tawi Atayr" pageId="220" pageNumber="221">Tawi Atayr</location>
,
<collectingMunicipality id="E7313819FF45FF0FA4A5FB0FFBEDD8FB" box="[949,1067,1248,1274]" pageId="220" pageNumber="221">Pop Cave</collectingMunicipality>
;
<geoCoordinate id="62DEC4A4FF45FF0FA329FB31FB16D8FB" box="[1081,1232,1246,1274]" degrees="17" direction="north" minutes="06" orientation="latitude" pageId="220" pageNumber="221" precision="15" seconds="45" value="17.112501">17°06′45″ N</geoCoordinate>
,
<geoCoordinate id="62DEC4A4FF45FF0FA3CDFB31FAB7D8FB" box="[1245,1393,1246,1274]" degrees="54" direction="east" minutes="33" orientation="longitude" pageId="220" pageNumber="221" precision="15" seconds="30" value="54.558334">54°33′30″ E</geoCoordinate>
;
<date id="735484A3FF45FF0FA7ADFAEBFE8AD91F" box="[189,332,1284,1311]" pageId="220" pageNumber="221" value="1997-11-02">
<collectingDate id="63107D4BFF45FF0FA7ADFAEBFE8AD91F" box="[189,332,1284,1311]" pageId="220" pageNumber="221" value="1997-11-02">2 Nov. 1997</collectingDate>
</date>
;
<collectorName id="AA1FC7B5FF45FF0FA646FAEBFE70D91F" box="[342,438,1284,1310]" pageId="220" pageNumber="221">S. Polak</collectorName>
leg.;
<collectionCode id="61FB3AA6FF45FF0FA6E4FAEBFD88D91F" box="[500,590,1284,1310]" pageId="220" pageNumber="221">ONHM</collectionCode>
[not examined]
</materialsCitation>
•
<materialsCitation id="B782A83EFF45FF0FA409FAEBFC76D942" ID-GBIF-Occurrence="3383565799" collectingDate="1997-11-03" collectionCode="SMNS" collectorName="S. Polak" country="Oman" latitude="17.112501" location="Quanaf" longLatPrecision="21" longitude="54.604164" municipality="Blown Gasket Cave" pageId="220" pageNumber="221" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="11EC69EAFF45FF0FA409FAEBFC85D91E" box="[793,835,1284,1311]" pageId="220" pageNumber="221" type="female">1 ♀</specimenCount>
,
<specimenCount id="11EC69EAFF45FF0FA45FFAEBFCBAD91E" box="[847,892,1284,1311]" pageId="220" pageNumber="221" type="male">1 ♂</specimenCount>
;
<location id="0235F4B8FF45FF0FA497FAEBFC26D91F" LSID="urn:lsid:plazi:treatment:8F431375FF45FF0EA6B6FEE6FEDCDD6B:0235F4B8FF45FF0FA497FAEBFC26D91F" box="[903,992,1284,1310]" country="Oman" latitude="17.112501" longLatPrecision="21" longitude="54.604164" municipality="Blown Gasket Cave" name="Quanaf" pageId="220" pageNumber="221">Quanaf</location>
,
<collectingMunicipality id="E7313819FF45FF0FA4FAFAEBFB12D91F" box="[1002,1236,1284,1310]" pageId="220" pageNumber="221">Blown Gasket Cave</collectingMunicipality>
;
<geoCoordinate id="62DEC4A4FF45FF0FA3CEFAEDFAB4D91E" box="[1246,1394,1282,1311]" degrees="17" direction="north" minutes="06" orientation="latitude" pageId="220" pageNumber="221" precision="15" seconds="45" value="17.112501">17°06′45″ N</geoCoordinate>
,
<geoCoordinate id="62DEC4A4FF45FF0FA7ADFAC8FE96D942" box="[189,336,1319,1347]" degrees="54" direction="east" minutes="36" orientation="longitude" pageId="220" pageNumber="221" precision="15" seconds="15" value="54.604164">54°36′15″ E</geoCoordinate>
;
<date id="735484A3FF45FF0FA64EFAC6FE34D942" box="[350,498,1321,1347]" pageId="220" pageNumber="221" value="1997-11-03">
<collectingDate id="63107D4BFF45FF0FA64EFAC6FE34D942" box="[350,498,1321,1347]" pageId="220" pageNumber="221" value="1997-11-03">3 Nov. 1997</collectingDate>
</date>
;
<collectorName id="AA1FC7B5FF45FF0FA6EFFAC6FDA4D942" box="[511,610,1321,1347]" pageId="220" pageNumber="221">S. Polak</collectorName>
leg.;
<collectionCode id="61FB3AA6FF45FF0FA5B5FAC6FD33D942" box="[677,757,1321,1347]" collectionName="Germany, Stuttgart, Staatliches Museum fuer Naturkunde" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="220" pageNumber="221">SMNS</collectionCode>
[not examined]
</materialsCitation>
•
<materialsCitation id="B782A83EFF45FF0FA4DBFAC6FC4ED966" ID-GBIF-Occurrence="3383565474" collectingDate="1997-11-03" collectionCode="BFUL" collectorName="S. Polak" country="Oman" latitude="17.112501" location="Quanaf" longLatPrecision="21" longitude="54.604164" municipality="Blown Gasket Cave" pageId="220" pageNumber="221" specimenCount="4" specimenCount-female="1" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="11EC69EAFF45FF0FA4DBFAC6FC3ED942" box="[971,1016,1321,1347]" pageId="220" pageNumber="221" type="female">1 ♀</specimenCount>
,
<specimenCount id="11EC69EAFF45FF0FA317FAC6FB8AD942" box="[1031,1100,1321,1347]" pageId="220" pageNumber="221" type="male">3 ♂♂</specimenCount>
, 1 protonymph exuvium; same collection data as for preceding;
<collectionCode id="61FB3AA6FF45FF0FA590FAA1FD0CD969" box="[640,714,1358,1384]" pageId="220" pageNumber="221">BFUL</collectionCode>
[not examined]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4FF0F1E8FF45FF0FA7ADFA7FFE8BD9DB" box="[189,373,1424,1498]" pageId="220" pageNumber="221" type="description">
<paragraph id="0755A263FF45FF0FA7ADFA7FFEB3D9AB" blockId="220.[189,373,1424,1450]" box="[189,373,1424,1450]" pageId="220" pageNumber="221">
<heading id="5C1D150FFF45FF0FA7ADFA7FFEB3D9AB" bold="true" box="[189,373,1424,1450]" fontSize="11" level="3" pageId="220" pageNumber="221" reason="3">
<emphasis id="359E7E71FF45FF0FA7ADFA7FFEB3D9AB" bold="true" box="[189,373,1424,1450]" pageId="220" pageNumber="221">Measurements</emphasis>
</heading>
</paragraph>
<paragraph id="0755A263FF45FF0FA7ADFA2FFE8BD9DB" blockId="220.[189,333,1472,1498]" box="[189,333,1472,1498]" pageId="220" pageNumber="221">
See
<tableCitation id="4A6897D8FF45FF0FA7FEFA2FFE81D9DB" box="[238,327,1472,1498]" captionStart="Table 9" captionStartId="213.[181,251,576,605]" captionTargetId="graphics-0@213.[181,1934,690,1746]" captionTargetPageId="213" captionText="Table 9 (continued on next page). Measurements of all African, Middle Eastern and South Asian Sarax Simon, 1892." pageId="220" pageNumber="221">Table 9</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4FF0F1E8FF45FF0FA7ADF9EDFC1BDA4C" box="[189,989,1538,1613]" pageId="220" pageNumber="221" type="distribution">
<paragraph id="0755A263FF45FF0FA7ADF9EDFE90DA1D" blockId="220.[189,342,1538,1564]" box="[189,342,1538,1564]" pageId="220" pageNumber="221">
<heading id="5C1D150FFF45FF0FA7ADF9EDFE90DA1D" bold="true" box="[189,342,1538,1564]" fontSize="11" level="3" pageId="220" pageNumber="221" reason="3">
<emphasis id="359E7E71FF45FF0FA7ADF9EDFE90DA1D" bold="true" box="[189,342,1538,1564]" pageId="220" pageNumber="221">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="0755A263FF45FF0FA7ADF9DCFC1BDA4C" blockId="220.[189,989,1586,1613]" box="[189,989,1586,1613]" pageId="220" pageNumber="221">
Known only from the
<typeStatus id="D8511CC1FF45FF0FA6D2F9DCFE33DA4C" box="[450,501,1587,1613]" pageId="220" pageNumber="221">type</typeStatus>
locality in the
<collectingRegion id="C52E6C81FF45FF0FA5B4F9DDFCA1DA4D" box="[676,871,1586,1612]" country="Oman" name="Zufar" pageId="220" pageNumber="221">Dhofar Province</collectingRegion>
of
<collectingCountry id="7FFDE2F3FF45FF0FA49EF9DCFC1CDA4C" box="[910,986,1587,1613]" name="Oman" pageId="220" pageNumber="221">Oman</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="4FF0F1E8FF45FF0FA7ADF99AFE6BDB77" pageId="220" pageNumber="221" type="biology_ecology">
<paragraph id="0755A263FF45FF0FA7ADF99AFEB8DA8E" blockId="220.[189,382,1653,1679]" box="[189,382,1653,1679]" pageId="220" pageNumber="221">
<heading id="5C1D150FFF45FF0FA7ADF99AFEB8DA8E" bold="true" box="[189,382,1653,1679]" fontSize="11" level="3" pageId="220" pageNumber="221" reason="3">
<emphasis id="359E7E71FF45FF0FA7ADF99AFEB8DA8E" bold="true" box="[189,382,1653,1679]" pageId="220" pageNumber="221">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="0755A263FF45FF0FA7ADF94AFE6BDB77" blockId="220.[188,1400,1701,1910]" pageId="220" pageNumber="221">
According to
<bibRefCitation id="637BDF92FF45FF0FA671F94AFDB6DABE" author="Weygoldt P. &amp; Hans P. &amp; Polak S." box="[353,624,1701,1727]" etAl="et al." firstAuthor="Weygoldt" journalOrPublisher="Fauna of Arabia" pageId="220" pageNumber="221" pagination="289 - 309" part="19" refId="ref156579" refString="Weygoldt P., Hans P. &amp; Polak S. 2002. Arabian whip spiders: four new species of the genera Charinus and Phrynichus (Chelicerata: Amblypygi) from Oman and Socotra. Fauna of Arabia 19: 289 - 309." title="Arabian whip spiders: four new species of the genera Charinus and Phrynichus (Chelicerata: Amblypygi) from Oman and Socotra" type="journal article" year="2002">
Weygoldt
<emphasis id="359E7E71FF45FF0FA6CCF949FDDEDABE" box="[476,536,1701,1727]" italics="true" pageId="220" pageNumber="221">et al.</emphasis>
(2002)
</bibRefCitation>
, the species occurs in three deep limestone caves in a large karst plateau. The caves are sinkholes of usually dry rivers that occasionally flow during the rainy monsoon season, and all have water in the deeper parts.
<taxonomicName id="C0EAD9E0FF45FF0FA5C5F901FC6EDB09" authorityName="Miranda &amp; Giupponi &amp; Prendini &amp; Scharff" authorityYear="2021" baseAuthorityName="Weygoldt, Pohl &amp; Polak" baseAuthorityYear="2002" box="[725,936,1774,1800]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="dhofarensis" status="comb. nov.">
<emphasis id="359E7E71FF45FF0FA5C5F901FC6EDB09" box="[725,936,1774,1800]" italics="true" pageId="220" pageNumber="221">Sarax dhofarensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2EADC30AFF45FF0FA4BFF901FBF6DB08" box="[943,1072,1774,1801]" pageId="220" pageNumber="221" rank="species">comb. nov.</taxonomicNameLabel>
is found in the aphotic zone in deeper parts of the caves, between 150 and
<quantity id="C0120F86FF45FF0FA5E0F8FCFC8ADB2C" box="[752,844,1811,1837]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="220" pageNumber="221" unit="m" value="1000.0">1000 m</quantity>
from the entrance. Individuals were observed sitting on the roof, on the cave walls or on bat guano deposits on the cave floor. The temperature in the caves is about 23°C.
</paragraph>
</subSubSection>
<subSubSection id="4FF0F1E8FF45FF0EA7ADF870FEDCDD6B" lastPageId="221" lastPageNumber="222" pageId="220" pageNumber="221" type="discussion">
<paragraph id="0755A263FF45FF0FA7ADF870FEE9DBB8" blockId="220.[189,303,1951,1977]" box="[189,303,1951,1977]" pageId="220" pageNumber="221">
<heading id="5C1D150FFF45FF0FA7ADF870FEE9DBB8" bold="true" box="[189,303,1951,1977]" fontSize="11" level="3" pageId="220" pageNumber="221" reason="3">
<emphasis id="359E7E71FF45FF0FA7ADF870FEE9DBB8" bold="true" box="[189,303,1951,1977]" pageId="220" pageNumber="221">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="0755A263FF45FF0FA7ADF821FCC8DBE9" blockId="220.[189,782,1998,2025]" box="[189,782,1998,2025]" pageId="220" pageNumber="221">The type material was not deposited in the SMNS.</paragraph>
<paragraph id="0755A263FF44FF0EA7ADFEE6FEDCDD6B" blockId="221.[188,1398,265,362]" pageId="221" pageNumber="222">
This species was originally described in the genus
<taxonomicName id="C0EAD9E0FF44FF0EA41AFEE6FCB1DD22" authorityName="Miranda &amp; Giupponi &amp; Prendini &amp; Scharff" authorityYear="2021" box="[778,887,265,291]" class="Arachnida" family="Charinidae" genus="Charinus" kingdom="Animalia" order="Amblypygi" pageId="221" pageNumber="222" phylum="Arthropoda" rank="genus">
<emphasis id="359E7E71FF44FF0EA41AFEE6FCB1DD22" box="[778,887,265,291]" italics="true" pageId="221" pageNumber="222">Charinus</emphasis>
</taxonomicName>
and is hereby transferred to
<taxonomicName id="C0EAD9E0FF44FF0EA3D4FEE6FACEDD22" authorityName="Simon" authorityYear="1892" box="[1220,1288,265,291]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="221" pageNumber="222" phylum="Arthropoda" rank="genus">
<emphasis id="359E7E71FF44FF0EA3D4FEE6FACEDD22" box="[1220,1288,265,291]" italics="true" pageId="221" pageNumber="222">Sarax</emphasis>
</taxonomicName>
based on the phylogenetic analyses of
<bibRefCitation id="637BDF92FF44FF0EA503FEC2FCD2DD46" author="Miranda G. S. &amp; Giupponi A. P. &amp; Scharff N. &amp; Prendini L." box="[531,788,301,327]" etAl="et al." firstAuthor="Miranda" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="221" pageNumber="222" refId="ref151478" refString="Miranda G. S., Giupponi A. P., Scharff N. &amp; Prendini L. 2021. Phylogeny and biogeography of the pantropical whip spider family Charinidae (Arachnida: Amblypygi). Zoological Journal of the Linnean Society. https: // doi. org / 10.1093 / zoolinnean / zlaa 101" title="Phylogeny and biogeography of the pantropical whip spider family Charinidae (Arachnida: Amblypygi)" type="book" year="2021">
<collectingRegion id="C52E6C81FF44FF0EA503FEC2FDB0DD46" box="[531,630,301,327]" country="Venezuela" name="Miranda" pageId="221" pageNumber="222">Miranda</collectingRegion>
<emphasis id="359E7E71FF44FF0EA56FFEC1FD7CDD46" box="[639,698,301,327]" italics="true" pageId="221" pageNumber="222">et al.</emphasis>
(2021)
</bibRefCitation>
. See
<bibRefCitation id="637BDF92FF44FF0EA441FEC2FB9DDD46" author="Weygoldt P. &amp; Hans P. &amp; Polak S." box="[849,1115,301,327]" etAl="et al." firstAuthor="Weygoldt" journalOrPublisher="Fauna of Arabia" pageId="221" pageNumber="222" pagination="289 - 309" part="19" refId="ref156579" refString="Weygoldt P., Hans P. &amp; Polak S. 2002. Arabian whip spiders: four new species of the genera Charinus and Phrynichus (Chelicerata: Amblypygi) from Oman and Socotra. Fauna of Arabia 19: 289 - 309." title="Arabian whip spiders: four new species of the genera Charinus and Phrynichus (Chelicerata: Amblypygi) from Oman and Socotra" type="journal article" year="2002">
Weygoldt
<emphasis id="359E7E71FF44FF0EA4DAFEC1FBC3DD46" box="[970,1029,301,327]" italics="true" pageId="221" pageNumber="222">et al.</emphasis>
(2002)
</bibRefCitation>
for a description of this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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