treatments-xml/data/20/0B/EB/200BEB73FF92724D53B867B1FDB28230.xml

<document ID-DOI="http://doi.org/10.5281/zenodo.3791622" ID-GBIF-Dataset="02c11e5e-32c9-4316-80c8-084d87826982" ID-GBIF-Taxon="163748459" ID-ISSN="1313–2970" ID-Zenodo-Dep="3791622" ID-ZooBank="83634493-F32F-4AA7-8E38-39509131E706" LSID="urn:lsid:zoobank.org:act:0D4FFE8A-AA79-4F64-9420-C9AC902A612A" checkinTime="1587958249361" checkinUser="plazi" docAuthor="Arnedo, Miquel, Gasparo, Fulvio &amp; Opatova, Vera" docDate="2009" docId="200BEB73FF92724D53B867B1FDB28230" docLanguage="en" docName="ZK_article_2023.pdf" docOrigin="ZooKeys 16 (16)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@18c8ecf5" docStyleName="zookeys.2008.journal_article" docTitle="Dysdera shardana Opatova &amp; Arnedo 2009, sp. n." docType="treatment" docUuid="0D4FFE8A-AA79-4F64-9420-C9AC902A612A" docUuidSource="ZooBank" docVersion="9" lastPageId="21" lastPageNumber="340" masterDocId="DC32930BFF9D725853346331FF80837D" masterDocTitle="Systematics and phylogeography of the Dysdera erythrina species complex (Araneae, Dysderidae) in Sardinia" masterLastPageNumber="345" masterPageNumber="319" pageId="15" pageNumber="334" updateTime="1634720684126" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Systematics and phylogeography of the Dysdera erythrina species complex (Araneae, Dysderidae) in Sardinia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Arnedo, Miquel</mods:namePart>
<mods:affiliation>University of Barcelona ,, Spain</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gasparo, Fulvio</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Via Vittoria Colonna &amp; Czech Republic &amp; Aea- 42 Be- 8886 - 0 A</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Opatova, Vera</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Departament de Biologia Animal and Biodiversity Research Institute, Universitat de Barcelona, Av. Diagonal 645, Barcelona 08028, Spain &amp; B &amp; Department of Zoology, Faculty of Science, Charles University in Prague, Viničná</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-07-29</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>16</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>16</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>319</mods:start>
<mods:end>345</mods:end>
</mods:extent>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.16.128</mods:identifier>
<mods:identifier type="GBIF-Dataset">02c11e5e-32c9-4316-80c8-084d87826982</mods:identifier>
<mods:identifier type="ISSN">1313–2970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576489</mods:identifier>
<mods:identifier type="ZooBank">83634493-F32F-4AA7-8E38-39509131E706</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791622" ID-GBIF-Taxon="163748459" ID-Zenodo-Dep="3791622" LSID="urn:lsid:zoobank.org:act:0D4FFE8A-AA79-4F64-9420-C9AC902A612A" httpUri="http://treatment.plazi.org/id/200BEB73FF92724D53B867B1FDB28230" lastPageId="21" lastPageNumber="340" pageId="15" pageNumber="334">
<subSubSection box="[140,672,1152,1179]" pageId="15" pageNumber="334" type="nomenclature">
<paragraph blockId="15.[140,975,1152,1249]" box="[140,672,1152,1179]" pageId="15" pageNumber="334">
<heading box="[140,672,1152,1179]" fontSize="11" level="2" pageId="15" pageNumber="334" reason="3">
<emphasis bold="true" box="[140,672,1152,1179]" pageId="15" pageNumber="334">
<taxonomicName authority="Opatova &amp; Arnedo" authorityName="Opatova &amp; Arnedo" authorityYear="2009" box="[140,594,1152,1179]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="334" phylum="Arthropoda" rank="species" species="shardana" status="sp. n.">
<emphasis bold="true" box="[140,356,1152,1179]" italics="true" pageId="15" pageNumber="334">
<emphasis box="[140,238,1152,1179]" inLineHeading="true" pageId="15" pageNumber="334" reason="1">Dysdera</emphasis>
shardana
</emphasis>
Opatova &amp; Arnedo
</taxonomicName>
, 
<taxonomicNameLabel box="[607,672,1152,1178]" pageId="15" pageNumber="334" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection pageId="15" pageNumber="334" type="description">
<paragraph blockId="15.[140,975,1152,1249]" box="[140,975,1187,1213]" pageId="15" pageNumber="334">
<uri box="[140,975,1187,1213]" pageId="15" pageNumber="334">
urn:lsid:zoobank.org:act: 
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</uri>
</paragraph>
<paragraph blockId="15.[140,975,1152,1249]" box="[140,431,1222,1249]" pageId="15" pageNumber="334">
<figureCitation box="[140,238,1222,1249]" captionStart="Figures 4-7" captionStartId="13.[140,222,1622,1644]" captionTargetBox="[140,1108,1166,1602]" captionTargetId="figure@13.[140,1109,1166,1603]" captionTargetPageId="13" captionText="Figures 4-7. Carapace, dorsal view: 4 D. jana sp. n. holotype 5 D. shardana sp. n. holotype. Carapace, lateral view: 6 D. jana sp. n. holotype 7 D. shardana sp. n. holotype." figureDoi="http://doi.org/10.5281/zenodo.3813919" httpUri="https://zenodo.org/record/3813919/files/figure.png" pageId="15" pageNumber="334">Figs 5, 7</figureCitation>
, 
<figureCitation box="[251,268,1223,1249]" captionStart="Figures 8-9" captionStartId="15.[140,222,1051,1073]" captionTargetBox="[140,1108,165,1031]" captionTargetId="figure@15.[140,1108,165,1032]" captionTargetPageId="15" captionText="Figures 8-9. Left male palp, retrolateral view: 8 D. jana sp. n. holotype 9 D. shardana sp. n. holotype." figureDoi="http://doi.org/10.5281/zenodo.3813921" httpUri="https://zenodo.org/record/3813921/files/figure.png" pageId="15" pageNumber="334">9</figureCitation>
, 
<figureCitation box="[280,350,1223,1249]" captionStart="Figures 10-13" captionStartId="16.[140,222,1622,1644]" captionTargetBox="[140,1108,554,1602]" captionTargetId="figure@16.[140,1108,551,1604]" captionTargetPageId="16" captionText="Figures 10-13. D. jana sp. n. paratype female 10 vulva, dorsal view 11 vulva, ventral view. D. shardana sp. n. paratype female 12 vulva, dorsal view 13 vulva, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3813923" httpUri="https://zenodo.org/record/3813923/files/figure.png" pageId="15" pageNumber="334">12-13</figureCitation>
, 
<figureCitation box="[363,431,1223,1249]" captionStart="Figures 19-22" captionStartId="18.[140,222,1622,1644]" captionTargetBox="[140,1108,408,1603]" captionTargetId="figure@18.[140,1108,408,1603]" captionTargetPageId="18" captionText="Figures 19-22. D. shardana holotype, right male bulb 19 prolateral view, general 20 prolateral view, tip detail 21 anterior view 22 P, prolateral view." figureDoi="http://doi.org/10.5281/zenodo.3813927" httpUri="https://zenodo.org/record/3813927/files/figure.png" pageId="15" pageNumber="334">19-22</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="334" type="reference_group">
<paragraph blockId="15.[140,1108,1292,1356]" pageId="15" pageNumber="334">
<treatmentCitationGroup pageId="15" pageNumber="334">
<treatmentCitation author="Grasshoff M" box="[140,541,1293,1320]" page="219" pageId="15" pageNumber="334" year="1959">
<taxonomicName authority="Grasshoff, 1959: 219" authorityName="Grasshoff" authorityPageNumber="219" authorityYear="1959" box="[140,541,1293,1320]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="334" phylum="Arthropoda" rank="species" species="kollari">
<emphasis box="[140,298,1293,1319]" italics="true" pageId="15" pageNumber="334">Dysdera kollari</emphasis>
<bibRefCitation author="Grasshoff" box="[306,541,1293,1320]" firstAuthor="Grasshoff" pageId="15" pageNumber="334" pagination="209 - 220" refId="ref11561" refString="Grasshoff M (1959) Dysdera - Arten von Inseln der Mittelmeergebietes (Arachn., Araneae). Senckenbergiana biologica 40: 209 - 220." type="journal article" year="1959">Grasshoff, 1959: 219</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 6 [♁] (NO Torralba, SO Cuglieri, S Cuglieri). 
<treatmentCitation author="Schult J" box="[185,402,1328,1354]" page="69" pageId="15" pageNumber="334" year="1983">
<bibRefCitation author="Schult" box="[185,402,1328,1354]" firstAuthor="Schult" pageId="15" pageNumber="334" pagination="69 - 84" refId="ref12405" refString="Schult J (1983) Taster haplogyner Spinnen unter phylogenetischem Aspekt (Arachnida: Araneae). Verhandlungen des Naturwissenschaftlichen Vereins in Hamburg 26: 69 - 84." type="journal article" year="1983">Schult, 1983: 69-84</bibRefCitation>
</treatmentCitation>
, fig. 8 [♁] (same specimens as 
<bibRefCitation author="Grasshoff" box="[723,898,1328,1355]" firstAuthor="Grasshoff" pageId="15" pageNumber="334" pagination="209 - 220" refId="ref11561" refString="Grasshoff M (1959) Dysdera - Arten von Inseln der Mittelmeergebietes (Arachn., Araneae). Senckenbergiana biologica 40: 209 - 220." type="journal article" year="1959">Grasshoff, 1959</bibRefCitation>
). Misidentification.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection lastPageId="18" lastPageNumber="337" pageId="15" pageNumber="334" type="materials_examined">
<paragraph blockId="15.[140,1108,1398,1673]" pageId="15" pageNumber="334">
<emphasis bold="true" box="[140,305,1398,1425]" pageId="15" pageNumber="334">
<emphasis box="[140,198,1399,1425]" inLineHeading="true" pageId="15" pageNumber="334" reason="1">Type</emphasis>
material
</emphasis>
. 
<materialsCitation ID-GBIF-Occurrence="2610757305" collectingDate="2005-10-15" collectorName="M. Arnedo &amp; M. Mejia-Chang" country="Italy" county="Oristano" elevation="566" latitude="39.7606" location="Is Benas" longLatPrecision="5" longitude="8.76062" municipality="Morgongiori" pageId="15" pageNumber="334" specimenCount="1" stateProvince="Sardegna" typeStatus="holotype">
<typeStatus box="[321,432,1399,1425]" pageId="15" pageNumber="334">Holotype</typeStatus>
: male: 
<collectingCountry box="[514,565,1399,1425]" name="Italy" pageId="15" pageNumber="334">Italy</collectingCountry>
, 
<collectingRegion box="[578,674,1399,1425]" country="Italy" name="Sardegna" pageId="15" pageNumber="334">Sardinia</collectingRegion>
, 
<collectingCounty box="[686,789,1399,1425]" pageId="15" pageNumber="334">Oristano</collectingCounty>
, 
<collectingMunicipality box="[802,950,1399,1425]" pageId="15" pageNumber="334">Morgongiori</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:200BEB73FF92724D53B867B1FDB28230:AD7D0CBEFF92725750F76646FBA186EC" box="[963,1057,1399,1425]" country="Italy" county="Oristano" latitude="39.7606" longLatPrecision="5" longitude="8.76062" municipality="Morgongiori" name="Is Benas" pageId="15" pageNumber="334" stateProvince="Sardegna">Is Benas</location>
recreation area, 
<location LSID="urn:lsid:plazi:treatment:200BEB73FF92724D53B867B1FDB28230:AD7D0CBEFF927257522A66ABFE0286C9" box="[286,386,1434,1460]" country="Italy" county="Oristano" latitude="39.7606" longLatPrecision="5" longitude="8.76062" municipality="Morgongiori" name="Mt. Arci" pageId="15" pageNumber="334" stateProvince="Sardegna">Mt. Arci</location>
, 
<geoCoordinate box="[398,520,1434,1460]" degrees="39.7606" direction="north" orientation="latitude" pageId="15" pageNumber="334" precision="5" value="39.7606">39.7606N</geoCoordinate>
, 
<geoCoordinate box="[531,647,1434,1460]" degrees="8.76062" direction="east" orientation="longitude" pageId="15" pageNumber="334" precision="1" value="8.76062">8.76062E</geoCoordinate>
, 
<quantity box="[658,736,1434,1461]" metricMagnitude="2" metricUnit="m" metricValue="5.66" pageId="15" pageNumber="334" unit="m" value="566.0">
<elevation box="[658,736,1434,1461]" metricMagnitude="2" metricUnit="m" metricValue="5.66" pageId="15" pageNumber="334" unit="m" value="566.0">566 m</elevation>
</quantity>
, mixed 
<taxonomicName box="[825,886,1435,1460]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="15" pageNumber="334" phylum="Tracheophyta" rank="genus">
<emphasis box="[825,886,1435,1460]" italics="true" pageId="15" pageNumber="334">Pinus</emphasis>
</taxonomicName>
and 
<taxonomicName box="[943,1031,1434,1460]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="15" pageNumber="334" phylum="Tracheophyta" rank="genus">
<emphasis box="[943,1031,1434,1460]" italics="true" pageId="15" pageNumber="334">Quercus</emphasis>
</taxonomicName>
forest, 
<date box="[140,264,1469,1496]" pageId="15" pageNumber="334" value="2005-10-15">
<collectingDate box="[140,264,1469,1496]" pageId="15" pageNumber="334" value="2005-10-15">15.X.2005</collectingDate>
</date>
, 
<collectorName box="[276,401,1469,1496]" pageId="15" pageNumber="334">M. Arnedo</collectorName>
and 
<collectorName box="[459,650,1469,1496]" pageId="15" pageNumber="334">M. Mejía-Chang</collectorName>
leg. (CRBA001246).
</materialsCitation>
</paragraph>
<paragraph blockId="15.[140,1108,1398,1673]" pageId="15" pageNumber="334">
<typeStatus box="[185,295,1505,1531]" pageId="15" pageNumber="334">Paratypes</typeStatus>
: 
<materialsCitation ID-GBIF-Occurrence="2610757306" pageId="15" pageNumber="334">
<specimenCount box="[309,409,1504,1530]" pageId="15" pageNumber="334" type="female">1 female</specimenCount>
(
<collectionCode box="[427,507,1504,1531]" pageId="15" pageNumber="334">CRBA</collectionCode>
001246), 
<specimenCount box="[619,719,1504,1530]" pageId="15" pageNumber="334" type="female">1 female</specimenCount>
(
<collectionCode box="[737,817,1504,1530]" pageId="15" pageNumber="334">CRBA</collectionCode>
001247, DNA extraction K477), 2 juvs (
<collectionCode box="[307,387,1540,1566]" pageId="15" pageNumber="334">CRBA</collectionCode>
001713, 
<collectionCode box="[486,565,1540,1566]" pageId="15" pageNumber="334">CRBA</collectionCode>
001714, DNA extraction LB297) same data and collectors.
</materialsCitation>
</paragraph>
<paragraph blockId="15.[140,1108,1398,1673]" pageId="15" pageNumber="334">
<emphasis bold="true" box="[185,557,1609,1635]" pageId="15" pageNumber="334">
<emphasis box="[185,312,1609,1635]" inLineHeading="true" pageId="15" pageNumber="334" reason="1">Additional</emphasis>
material examined.
</emphasis>
<materialsCitation ID-GBIF-Occurrence="2608812171" collectorName="Fluminimaggiore" country="Italy" location="Is Arenas" pageId="15" pageNumber="334" specimenCount="1" stateProvince="Sardegna">
<emphasis bold="true" box="[569,741,1609,1636]" pageId="15" pageNumber="334">
<collectingCountry box="[569,626,1610,1636]" name="Italy" pageId="15" pageNumber="334">Italy</collectingCountry>
: 
<collectingRegion box="[642,741,1609,1635]" country="Italy" name="Sardegna" pageId="15" pageNumber="334">Sardinia</collectingRegion>
</emphasis>
: 
<emphasis bold="true" box="[761,856,1609,1635]" pageId="15" pageNumber="334">
<collectingRegion box="[761,856,1609,1635]" country="Italy" name="Cagliari" pageId="15" pageNumber="334">Cagliari</collectingRegion>
</emphasis>
: 
<collectorName box="[875,1072,1610,1636]" pageId="15" pageNumber="334">Fluminimaggiore</collectorName>
, 
<location LSID="urn:lsid:plazi:treatment:200BEB73FF92724D53B867B1FDB28230:AD7D0CBEFF9272575775657BFF5B85FA" country="Italy" name="Is Arenas" pageId="15" pageNumber="334" stateProvince="Sardegna">Is Arenas</location>
, under 
<taxonomicName box="[306,439,1645,1672]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="15" pageNumber="334" phylum="Tracheophyta" rank="species" species="ilex">
<emphasis box="[306,439,1645,1672]" italics="true" pageId="15" pageNumber="334">Quercus ilex</emphasis>
</taxonomicName>
</materialsCitation>
; 
<quantity box="[456,534,1645,1672]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="15" pageNumber="334" unit="m" value="800.0">800 m</quantity>
; 
<materialsCitation ID-GBIF-Occurrence="2608812182" box="[547,994,1645,1673]" collectingDate="1991-03-29" collectionCode="CG" collectorName="G. Gardini" country="Italy" pageId="15" pageNumber="334" specimenCount="1" stateProvince="Cagliari">
<collectorName box="[547,673,1645,1671]" pageId="15" pageNumber="334">G. Gardini</collectorName>
leg. 
<date box="[729,863,1645,1671]" pageId="15" pageNumber="334" value="1991-03-29">
<collectingDate box="[729,863,1645,1671]" pageId="15" pageNumber="334" value="1991-03-29">29.III.1991</collectingDate>
</date>
, 1 ♁ (
<collectionCode box="[940,986,1645,1671]" country="Brazil" name="Embrapa Collection of Fungi of Invertebrates" pageId="15" pageNumber="334">CG</collectionCode>
)
</materialsCitation>
; Flumini-
</paragraph>
<paragraph blockId="16.[140,1108,167,511]" pageId="16" pageNumber="335">
<materialsCitation ID-GBIF-Occurrence="2608812175" box="[140,652,167,193]" location="Tempio di Antas" pageId="16" pageNumber="335" specimenCount="1">
maggiore, 
<location LSID="urn:lsid:plazi:treatment:200BEB73FF92724D53B867B1FDB28230:AD7D0CBEFF8D724853CA6396FE3983BC" box="[254,441,167,193]" name="Tempio di Antas" pageId="16" pageNumber="335">Tempio di Antas</location>
, under 
<taxonomicName box="[522,652,167,193]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="16" pageNumber="335" phylum="Tracheophyta" rank="species" species="ilex">
<emphasis box="[522,652,167,193]" italics="true" pageId="16" pageNumber="335">Quercus ilex</emphasis>
</taxonomicName>
</materialsCitation>
; 
<quantity box="[665,740,167,193]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="16" pageNumber="335" unit="m" value="360.0">360 m</quantity>
; 
<materialsCitation ID-GBIF-Occurrence="2608812181" collectingDate="1991-03-29" collectionCode="CG" collectorName="G. Gardini" county="Mt. Gennargentu" location="Nuoro" municipality="Castagneto Geratzia" pageId="16" pageNumber="335" specimenCount="1" specimenCount-female="1">
<collectorName box="[749,872,167,193]" pageId="16" pageNumber="335">G. Gardini</collectorName>
leg. 
<date box="[921,1055,167,193]" pageId="16" pageNumber="335" value="1991-03-29">
<collectingDate box="[921,1055,167,193]" pageId="16" pageNumber="335" value="1991-03-29">29.III.1991</collectingDate>
</date>
, 
<specimenCount box="[1065,1100,167,193]" pageId="16" pageNumber="335" type="female">1 ♀</specimenCount>
(
<collectionCode box="[149,195,202,228]" country="Brazil" name="Embrapa Collection of Fungi of Invertebrates" pageId="16" pageNumber="335">CG</collectionCode>
). 
<emphasis bold="true" box="[216,294,202,228]" pageId="16" pageNumber="335">
<location LSID="urn:lsid:plazi:treatment:200BEB73FF92724D53B867B1FDB28230:AD7D0CBEFF8D724853EC63FBFEA68399" box="[216,294,202,228]" county="Mt. Gennargentu" municipality="Castagneto Geratzia" name="Nuoro" pageId="16" pageNumber="335">Nuoro</location>
</emphasis>
, 
<location LSID="urn:lsid:plazi:treatment:200BEB73FF92724D53B867B1FDB28230:AD7D0CBEFF8D7248520063FBFEFC8399" box="[308,380,202,228]" county="Mt. Gennargentu" municipality="Castagneto Geratzia" name="Aritzo" pageId="16" pageNumber="335">Aritzo</location>
, 
<collectingMunicipality box="[392,622,202,228]" pageId="16" pageNumber="335">Castagneto Geratzia</collectingMunicipality>
, 
<collectingCounty box="[634,836,202,228]" pageId="16" pageNumber="335">Mt. Gennargentu</collectingCounty>
</materialsCitation>
; 
<geoCoordinate box="[848,981,202,228]" degrees="39.96846" direction="north" orientation="latitude" pageId="16" pageNumber="335" precision="1" value="39.96846">39.96846N</geoCoordinate>
<geoCoordinate box="[988,1103,202,228]" degrees="9.19782" direction="east" orientation="longitude" pageId="16" pageNumber="335" precision="1" value="9.19782">9.19782E</geoCoordinate>
; 
<quantity box="[140,218,237,264]" metricMagnitude="2" metricUnit="m" metricValue="8.67" pageId="16" pageNumber="335" unit="m" value="867.0">867 m</quantity>
; 
<emphasis box="[231,338,238,264]" italics="true" pageId="16" pageNumber="335">Castaneus</emphasis>
and 
<taxonomicName box="[398,550,237,264]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="16" pageNumber="335" phylum="Tracheophyta" rank="species" species="suber">
<emphasis box="[398,550,237,264]" italics="true" pageId="16" pageNumber="335">Quercus suber</emphasis>
</taxonomicName>
forest; 
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; 
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<collectorName box="[773,903,237,264]" pageId="16" pageNumber="335">M. Arnedo</collectorName>
, 
<collectorName box="[916,1108,237,264]" pageId="16" pageNumber="335">M. Mejía-Chang</collectorName>
&amp; 
<collectorName box="[175,297,272,298]" pageId="16" pageNumber="335">G. Giribet</collectorName>
leg. (
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cryo-collection, DNA extraction LB303, 
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cryo-collection, DNA extraction LB304, 1♁ 
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cryocollection, DNA extraction LB305, 
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<specimenCode box="[593,770,343,369]" collectionCode="CRBA" pageId="16" pageNumber="335">CRBA0001711</specimenCode>
cryo-collection, DNA extraction LB306)
</materialsCitation>
; 
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, rd. to Fonni, 
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; 
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; 
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; 
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<emphasis box="[181,352,413,440]" italics="true" pageId="16" pageNumber="335">Quercus pubens</emphasis>
</taxonomicName>
forest; 
<date box="[443,566,413,440]" pageId="16" pageNumber="335" value="2005-10-17">17.X.2005</date>
; 
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<collectorName box="[582,714,413,440]" pageId="16" pageNumber="335">M. Arnedo</collectorName>
, 
<collectorName box="[729,923,413,440]" pageId="16" pageNumber="335">M. Mejía-Chang</collectorName>
and 
<collectorName box="[986,1107,413,439]" pageId="16" pageNumber="335">G. Giribet</collectorName>
leg. (1♁ 
<specimenCode box="[241,421,448,474]" collectionCode="CRBA" pageId="16" pageNumber="335">CRBA0001268</specimenCode>
, DNA extraction LB294, 1♁ 
<specimenCode box="[762,925,448,474]" collectionCode="CRBA" pageId="16" pageNumber="335">CRBA001270</specimenCode>
cryo-collection. DNA extraction LB307, 1♁ 
<specimenCode box="[469,632,484,510]" collectionCode="CRBA" pageId="16" pageNumber="335">CRBA001712</specimenCode>
cryo-collection. DNA extraction LB308)
</materialsCitation>
;
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3813923" ID-Zenodo-Dep="3813923" httpUri="https://zenodo.org/record/3813923/files/figure.png" pageId="16" pageNumber="335" startId="16.[140,222,1622,1644]" targetBox="[140,1108,554,1602]" targetPageId="16">
<paragraph blockId="16.[140,1108,1622,1676]" pageId="16" pageNumber="335">
<emphasis bold="true" box="[140,296,1622,1644]" pageId="16" pageNumber="335">Figures 10-13.</emphasis>
<taxonomicName authority="Arnedo &amp; Gasparo &amp; Opatova, 2009" authorityName="Arnedo &amp; Gasparo &amp; Opatova" authorityYear="2009" box="[301,371,1623,1644]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="335" phylum="Arthropoda" rank="species" species="jana" status="sp. n.">
<emphasis box="[301,371,1623,1644]" italics="true" pageId="16" pageNumber="335">D. jana</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[376,426,1623,1644]" pageId="16" pageNumber="335" rank="species">sp. n.</taxonomicNameLabel>
<typeStatus box="[432,511,1623,1644]" pageId="16" pageNumber="335">paratype</typeStatus>
female 
<emphasis bold="true" box="[583,609,1622,1644]" pageId="16" pageNumber="335">1 0</emphasis>
vulva, dorsal view 
<emphasis bold="true" box="[784,810,1622,1644]" pageId="16" pageNumber="335">11</emphasis>
vulva, ventral view. 
<taxonomicName authority="Arnedo &amp; Gasparo &amp; Opatova, 2009" authorityName="Arnedo &amp; Gasparo &amp; Opatova" authorityYear="2009" box="[997,1108,1622,1644]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="335" phylum="Arthropoda" rank="species" species="shardana" status="sp. n.">
<emphasis box="[997,1108,1622,1644]" italics="true" pageId="16" pageNumber="335">D. shardana</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[140,190,1655,1676]" pageId="16" pageNumber="335" rank="species">sp. n.</taxonomicNameLabel>
<typeStatus box="[196,275,1655,1676]" pageId="16" pageNumber="335">paratype</typeStatus>
female 
<emphasis bold="true" box="[348,374,1654,1676]" pageId="16" pageNumber="335">1 2</emphasis>
vulva, dorsal view 
<emphasis bold="true" box="[550,576,1654,1676]" pageId="16" pageNumber="335">1 3</emphasis>
vulva, ventral view.
</paragraph>
</caption>
<paragraph blockId="17.[140,1108,166,369]" pageId="17" pageNumber="336">
Macomer, Mt. S. Antonio, A. Vigna leg. 
<date box="[601,707,167,193]" pageId="17" pageNumber="336" value="1967-05-02">2.V.1967</date>
, 1 ♁ (
<collectionCode box="[778,870,167,193]" pageId="17" pageNumber="336">MCVR</collectionCode>
). 
<emphasis bold="true" box="[889,995,166,192]" pageId="17" pageNumber="336">Oristano</emphasis>
, Cuglieri, rd. to San Leonardo 7 Fuentes, Mt. Ferru; 
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; 
<quantity box="[907,986,202,229]" metricMagnitude="2" metricUnit="m" metricValue="5.87" pageId="17" pageNumber="336" unit="m" value="587.0">587 m</quantity>
; dry, open 
<taxonomicName box="[140,227,238,264]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="17" pageNumber="336" phylum="Tracheophyta" rank="genus">
<emphasis box="[140,227,238,264]" italics="true" pageId="17" pageNumber="336">Quercus</emphasis>
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forest; 
<date box="[313,437,237,264]" pageId="17" pageNumber="336" value="2005-10-14">14.X.2005</date>
; M. Arnedo and M. Mejía-Chang leg. (
<specimenCount box="[894,922,237,264]" pageId="17" pageNumber="336" type="female">1♀</specimenCount>
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001238, cryo-collection, DNA extraction LB295); nearby S. Leonardo 7 Fuentes, Mt. Ferru, 
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; 
<quantity box="[461,544,307,334]" metricMagnitude="2" metricUnit="m" metricValue="6.99" pageId="17" pageNumber="336" unit="m" value="699.0">699 m</quantity>
; old crops and open forest 
<taxonomicName box="[879,966,308,334]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="17" pageNumber="336" phylum="Tracheophyta" rank="genus">
<emphasis box="[879,966,308,334]" italics="true" pageId="17" pageNumber="336">Quercus</emphasis>
</taxonomicName>
and 
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<emphasis box="[1034,1100,307,333]" italics="true" pageId="17" pageNumber="336">Rubus</emphasis>
</taxonomicName>
; 
<date box="[140,263,342,369]" pageId="17" pageNumber="336" value="2005-10-14">14.X.2005</date>
; M. Arnedo &amp; M. Mejía-Chang leg. (
<specimenCount box="[715,743,343,369]" pageId="17" pageNumber="336" type="female">1♀</specimenCount>
<collectionCode box="[760,837,343,369]" pageId="17" pageNumber="336">CRBA</collectionCode>
001240 cryo-collection,
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3813925" ID-Zenodo-Dep="3813925" httpUri="https://zenodo.org/record/3813925/files/figure.png" pageId="17" pageNumber="336" startId="17.[140,222,1622,1644]" targetBox="[140,1108,411,1602]" targetPageId="17">
<paragraph blockId="17.[140,1107,1622,1676]" pageId="17" pageNumber="336">
<emphasis bold="true" box="[140,298,1622,1644]" pageId="17" pageNumber="336">Figures 14-18.</emphasis>
<taxonomicName authority="Arnedo &amp; Gasparo &amp; Opatova, 2009" authorityName="Arnedo &amp; Gasparo &amp; Opatova" authorityYear="2009" box="[305,377,1623,1644]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="336" phylum="Arthropoda" rank="species" species="jana" status="sp. n.">
<emphasis box="[305,377,1623,1644]" italics="true" pageId="17" pageNumber="336">D. jana</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[384,437,1623,1644]" pageId="17" pageNumber="336" rank="species">sp. n.</taxonomicNameLabel>
<typeStatus box="[444,528,1622,1644]" pageId="17" pageNumber="336">holotype</typeStatus>
, right male bulb 
<emphasis bold="true" box="[693,719,1622,1644]" pageId="17" pageNumber="336">1 4</emphasis>
prolateral view, general 
<emphasis bold="true" box="[950,977,1622,1644]" pageId="17" pageNumber="336">1 5</emphasis>
anterior view 
<emphasis bold="true" box="[140,166,1654,1676]" pageId="17" pageNumber="336">1 6</emphasis>
posterior view 
<emphasis bold="true" box="[309,335,1654,1676]" pageId="17" pageNumber="336">1 7</emphasis>
retrolateral view 
<emphasis bold="true" box="[495,521,1654,1676]" pageId="17" pageNumber="336">1 8</emphasis>
P, prolateral view.
</paragraph>
</caption>
<paragraph blockId="18.[140,1108,167,334]" pageId="18" pageNumber="337">
<materialsCitation ID-GBIF-Occurrence="2608812177" collectionCode="CRBA" pageId="18" pageNumber="337" specimenCode="CRBA001707, CRBA001708" specimenCount="2" specimenCount-female="1">
DNA extraction LB296, 
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cryo-collection, DNA extraction LB299, 
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<specimenCode box="[203,366,202,228]" collectionCode="CRBA" pageId="18" pageNumber="337">CRBA001708</specimenCode>
cryo-collection, DNA extraction LB300
</materialsCitation>
,); Morgongiori, Bruncu Mutzuis, 
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; 
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; open 
<taxonomicName box="[679,766,238,264]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="18" pageNumber="337" phylum="Tracheophyta" rank="genus">
<emphasis box="[679,766,238,264]" italics="true" pageId="18" pageNumber="337">Quercus</emphasis>
</taxonomicName>
forest w/ mosses; 
<date box="[979,1102,237,264]" pageId="18" pageNumber="337" value="2005-10-15">15.X.2005</date>
; 
<materialsCitation ID-GBIF-Occurrence="2608812180" collectionCode="CRBA" collectorName="M. Arnedo &amp; M. Mejia-Chang" pageId="18" pageNumber="337" specimenCode="CRBA001256" specimenCount="1">
<collectorName box="[140,266,272,299]" pageId="18" pageNumber="337">M. Arnedo</collectorName>
&amp; 
<collectorName box="[305,496,272,299]" pageId="18" pageNumber="337">M. Mejía-Chang</collectorName>
leg. (1sub♁ 
<specimenCode box="[643,808,272,299]" collectionCode="CRBA" pageId="18" pageNumber="337">CRBA001256</specimenCode>
, cryo-collection, DNA extraction LB301)
</materialsCitation>
</paragraph>
</subSubSection>
<paragraph pageId="18" pageNumber="337">1 9 20 P AR L AR 2 1 22 P L</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3813927" ID-Zenodo-Dep="3813927" httpUri="https://zenodo.org/record/3813927/files/figure.png" pageId="18" pageNumber="337" startId="18.[140,222,1622,1644]" targetBox="[140,1108,408,1603]" targetPageId="18">
<paragraph blockId="18.[140,1108,1622,1676]" pageId="18" pageNumber="337">
<emphasis bold="true" box="[140,295,1622,1644]" pageId="18" pageNumber="337">
<figureCitation box="[140,249,1622,1644]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figures 4-7" captionStart-4="Figures 8-9" captionStartId-0="2.[140,212,1201,1223]" captionStartId-1="9.[140,212,1430,1452]" captionStartId-2="10.[140,212,772,794]" captionStartId-3="13.[140,222,1622,1644]" captionStartId-4="15.[140,222,1051,1073]" captionTargetBox-1="[140,1083,888,1411]" captionTargetBox-2="[339,910,164,753]" captionTargetBox-3="[140,1108,1166,1602]" captionTargetBox-4="[140,1108,165,1031]" captionTargetId-1="graphics@9.[193,853,897,1388]" captionTargetId-2="graphics@10.[400,845,376,753]" captionTargetId-3="figure@13.[140,1109,1166,1603]" captionTargetId-4="figure@15.[140,1108,165,1032]" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="13" captionTargetPageId-4="15" captionText-0="Figure 1. Maps showing the location of Sardinia and known localities of the new species. Dysdera jana sp. n.: Nuoro: 1 Dorgali, 450 m, 2 2 km S of Dorgali, 500 m, 3 Supramonte di Oliena, 900 m, 4 Sa Oche, 7 km ESE of Oliena, 150 m. D. shardana sp. n.: Cagliari: 5 Fluminimaggiore, Is Arenas, 6 Flu- minimaggiore, Tempio di Antas, Nuoro: 7 Aritzo, Castagneto Geratzia, 867 m, 8 Desulo, rd. to Fonni, 1146 m, 9 Macomer, Mt. S. Antonio. Oristano: 10 Cuglieri, rd. to San Leonardo 7 Fuentes, 587 m, 11 S. Leonardo 7 Fuentes, 699 m, 12 Morgongiori, Is Benas recreation area, 566 m, 13 Morgongiori, Bruncu Mutzuis, 761 m, 14 Sassari: NW Torralba." captionText-1="Figure 2. Chronogram of the species sampled in the present study, based on the preferred Bayesian tree obtained by simultaneous analyses of the cox1, 16S, nad1, 28S and H3 partitions. Bars on branches denote support as follows: anterior bar refers to posterior probability, middle bar to maximum likelihood bootstrap support, posterior bar to parsimony jackknife support. Black bar: posterior probability&gt;0.95 or ML bootstrap, Parismony jackknife&gt; 70, grey bar: posterior probability &lt;0.95 or ML bootstrap, Parsimo- ny jackknife &lt;70, white bar: this particular clade was not recovered in the analyses. Numbers on branches are the age of the clade in millions of years before the present, estimated using penalized likelihood (set penalty=add, smoothing=3200), number in brackets refer to bootstrap confidence intervals." captionText-2="Figure 3. Haplotype network constructed using statistical parsimony (95%). Dots on branches refer to missing haplotypes. Node size proportional to number of haplotypes (1 or 2), nodes colour code and numbers inside refer to localities in Fig. 1." captionText-3="Figures 4-7. Carapace, dorsal view: 4 D. jana sp. n. holotype 5 D. shardana sp. n. holotype. Carapace, lateral view: 6 D. jana sp. n. holotype 7 D. shardana sp. n. holotype." captionText-4="Figures 8-9. Left male palp, retrolateral view: 8 D. jana sp. n. holotype 9 D. shardana sp. n. holotype." figureDoi-0="http://doi.org/10.5281/zenodo.3813913" figureDoi-1="http://doi.org/10.5281/zenodo.3813915" figureDoi-2="http://doi.org/10.5281/zenodo.3813917" figureDoi-3="http://doi.org/10.5281/zenodo.3813919" figureDoi-4="http://doi.org/10.5281/zenodo.3813921" httpUri-0="https://zenodo.org/record/3813913/files/figure.png" httpUri-1="https://zenodo.org/record/3813915/files/figure.png" httpUri-2="https://zenodo.org/record/3813917/files/figure.png" httpUri-3="https://zenodo.org/record/3813919/files/figure.png" httpUri-4="https://zenodo.org/record/3813921/files/figure.png" pageId="18" pageNumber="337">Figures 19</figureCitation>
-22.
</emphasis>
<taxonomicName authorityName="Arnedo &amp; Gasparo &amp; Opatova" authorityYear="2009" box="[301,411,1622,1644]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="337" phylum="Arthropoda" rank="species" species="shardana">
<emphasis box="[301,411,1622,1644]" italics="true" pageId="18" pageNumber="337">D. shardana</emphasis>
</taxonomicName>
<typeStatus box="[416,500,1622,1644]" pageId="18" pageNumber="337">holotype</typeStatus>
, right male bulb 
<emphasis bold="true" box="[656,682,1622,1644]" pageId="18" pageNumber="337">1 9</emphasis>
prolateral view, general 
<emphasis bold="true" box="[905,931,1622,1644]" pageId="18" pageNumber="337">20</emphasis>
prolateral view, tip detail 
<emphasis bold="true" box="[197,223,1654,1676]" pageId="18" pageNumber="337">2 1</emphasis>
anterior view 
<emphasis bold="true" box="[356,382,1654,1676]" pageId="18" pageNumber="337">22</emphasis>
P, prolateral view.
</paragraph>
</caption>
<subSubSection pageId="19" pageNumber="338" type="diagnosis">
<paragraph blockId="19.[140,1109,166,1672]" pageId="19" pageNumber="338">
<emphasis bold="true" box="[185,312,166,192]" pageId="19" pageNumber="338">
<emphasis box="[185,307,166,192]" inLineHeading="true" pageId="19" pageNumber="338" reason="1">Diagnosis</emphasis>
.
</emphasis>
<taxonomicName authority="Arnedo &amp; Gasparo &amp; Opatova, 2009" authorityName="Arnedo &amp; Gasparo &amp; Opatova" authorityYear="2009" box="[318,510,167,193]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="338" phylum="Arthropoda" rank="species" species="shardana" status="sp. n.">
<emphasis box="[318,510,167,193]" italics="true" pageId="19" pageNumber="338">Dysdera shardana</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[516,575,167,193]" pageId="19" pageNumber="338" rank="species">sp. n.</taxonomicNameLabel>
differs from continental species of the 
<taxonomicName authority="Linnæus, 1758" authorityName="Walckenaer" authorityYear="1802" box="[1008,1108,167,193]" class="Insecta" family="Blaberidae" genus="Aranea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="19" pageNumber="338" phylum="Arthropoda" rank="species" species="erythrina">
<emphasis box="[1008,1108,167,193]" italics="true" pageId="19" pageNumber="338">erythrina</emphasis>
</taxonomicName>
group in the shape of the posterior-distal part of the tegulum, which is expanded in the continental species. In addition, continental females of the 
<taxonomicName authority="Linnæus, 1758" authorityName="Walckenaer" authorityYear="1802" box="[849,949,237,263]" class="Insecta" family="Blaberidae" genus="Aranea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="19" pageNumber="338" phylum="Arthropoda" rank="species" species="erythrina">
<emphasis box="[849,949,237,263]" italics="true" pageId="19" pageNumber="338">erythrina</emphasis>
</taxonomicName>
group exhibit a characteristic hourglass-shaped spermatheca, which is very attenuated in 
<taxonomicName authority="Arnedo &amp; Gasparo &amp; Opatova, 2009" authorityName="Arnedo &amp; Gasparo &amp; Opatova" authorityYear="2009" box="[973,1108,272,298]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="338" phylum="Arthropoda" rank="species" species="shardana" status="sp. n.">
<emphasis box="[973,1108,272,298]" italics="true" pageId="19" pageNumber="338">D. shardana</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[140,200,308,334]" pageId="19" pageNumber="338" rank="species">sp. n.</taxonomicNameLabel>
It can also be distinguished from 
<taxonomicName authority="Linnæus, 1758" authorityName="Walckenaer" authorityYear="1802" box="[577,712,307,333]" class="Insecta" family="Blaberidae" genus="Aranea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="19" pageNumber="338" phylum="Arthropoda" rank="species" species="erythrina">
<emphasis box="[577,712,307,333]" italics="true" pageId="19" pageNumber="338">D. erythrina</emphasis>
</taxonomicName>
and 
<taxonomicName authority="Linnæus, 1758" authorityName="Linnæus" authorityYear="1758" box="[766,1032,307,333]" class="Insecta" family="Blaberidae" genus="Aranea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="19" pageNumber="338" phylum="Arthropoda" rank="subSpecies" species="erythrina" subSpecies="provincialis">
<emphasis box="[766,1032,307,333]" italics="true" pageId="19" pageNumber="338">D. erythrina provincialis</emphasis>
</taxonomicName>
by the lateral concave shape of the chelicera and a heavily wrinkled (foveate) carapace (
<figureCitation captionStart="Figures 4-7" captionStartId="13.[140,222,1622,1644]" captionTargetBox="[140,1108,1166,1602]" captionTargetId="figure@13.[140,1109,1166,1603]" captionTargetPageId="13" captionText="Figures 4-7. Carapace, dorsal view: 4 D. jana sp. n. holotype 5 D. shardana sp. n. holotype. Carapace, lateral view: 6 D. jana sp. n. holotype 7 D. shardana sp. n. holotype." figureDoi="http://doi.org/10.5281/zenodo.3813919" httpUri="https://zenodo.org/record/3813919/files/figure.png" pageId="19" pageNumber="338">Figs 5, 7</figureCitation>
). It differs from 
<taxonomicName authority="Arnedo &amp; Gasparo &amp; Opatova, 2009" authorityName="Arnedo &amp; Gasparo &amp; Opatova" authorityYear="2009" box="[341,425,379,404]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="338" phylum="Arthropoda" rank="species" species="jana" status="sp. n.">
<emphasis box="[341,425,379,404]" italics="true" pageId="19" pageNumber="338">D. jana</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[432,494,378,404]" pageId="19" pageNumber="338" rank="species">sp. n.</taxonomicNameLabel>
, also from 
<collectingRegion box="[618,714,378,404]" country="Italy" name="Sardegna" pageId="19" pageNumber="338">Sardinia</collectingRegion>
, in the presence of dorsal spines on tibia 4, the position and length of the finger-like process on the tip of the male bulb (compare 
<figureCitation box="[255,337,448,475]" captionStart="Figures 14-18" captionStartId="17.[140,222,1622,1644]" captionTargetBox="[140,1108,411,1602]" captionTargetId="figure@17.[140,1108,408,1603]" captionTargetPageId="17" captionText="Figures 14-18. D. jana sp. n. holotype, right male bulb 14 prolateral view, general 15 anterior view 16 posterior view 17 retrolateral view 18 P, prolateral view." figureDoi="http://doi.org/10.5281/zenodo.3813925" httpUri="https://zenodo.org/record/3813925/files/figure.png" pageId="19" pageNumber="338">Figs 15</figureCitation>
and 
<figureCitation box="[397,429,448,474]" captionStart="Figures 19-22" captionStartId="18.[140,222,1622,1644]" captionTargetBox="[140,1108,408,1603]" captionTargetId="figure@18.[140,1108,408,1603]" captionTargetPageId="18" captionText="Figures 19-22. D. shardana holotype, right male bulb 19 prolateral view, general 20 prolateral view, tip detail 21 anterior view 22 P, prolateral view." figureDoi="http://doi.org/10.5281/zenodo.3813927" httpUri="https://zenodo.org/record/3813927/files/figure.png" pageId="19" pageNumber="338">21</figureCitation>
), and by the fusion of the anterior sclerotised stripes of the vulva VA (compare 
<figureCitation box="[363,430,483,510]" captionStart="Figures 8-9" captionStartId="15.[140,222,1051,1073]" captionTargetBox="[140,1108,165,1031]" captionTargetId="figure@15.[140,1108,165,1032]" captionTargetPageId="15" captionText="Figures 8-9. Left male palp, retrolateral view: 8 D. jana sp. n. holotype 9 D. shardana sp. n. holotype." figureDoi="http://doi.org/10.5281/zenodo.3813921" httpUri="https://zenodo.org/record/3813921/files/figure.png" pageId="19" pageNumber="338">Figs 9</figureCitation>
and 
<figureCitation box="[487,518,484,510]" captionStart="Figures 10-13" captionStartId="16.[140,222,1622,1644]" captionTargetBox="[140,1108,554,1602]" captionTargetId="figure@16.[140,1108,551,1604]" captionTargetPageId="16" captionText="Figures 10-13. D. jana sp. n. paratype female 10 vulva, dorsal view 11 vulva, ventral view. D. shardana sp. n. paratype female 12 vulva, dorsal view 13 vulva, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3813923" httpUri="https://zenodo.org/record/3813923/files/figure.png" pageId="19" pageNumber="338">11</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="338" type="etymology">
<paragraph blockId="19.[140,1109,166,1672]" pageId="19" pageNumber="338">
<emphasis bold="true" box="[185,314,519,545]" inLineHeading="true" pageId="19" pageNumber="338" reason="1">Etymology</emphasis>
. The name in apposition refers to the Shardana people, one of several groups of “Sea Peoples” that appear in fragmentary historical records (Egyptian inscriptions) for the Mediterranean region in the second millennium B.C. According to some scholars they settled in 
<collectingRegion box="[399,490,624,650]" country="Italy" name="Sardegna" pageId="19" pageNumber="338">Sardinia</collectingRegion>
after the period of Ramses III (around 1180 B.C.).
</paragraph>
</subSubSection>
<subSubSection lastPageId="21" lastPageNumber="340" pageId="19" pageNumber="338" type="description">
<paragraph blockId="19.[140,1109,166,1672]" pageId="19" pageNumber="338">
<emphasis bold="true" box="[185,335,659,685]" pageId="19" pageNumber="338">
<emphasis box="[185,330,659,685]" inLineHeading="true" pageId="19" pageNumber="338" reason="1">Description</emphasis>
.
</emphasis>
<emphasis box="[342,496,659,685]" italics="true" pageId="19" pageNumber="338">
<typeStatus box="[342,437,659,685]" pageId="19" pageNumber="338">Holotype</typeStatus>
male
</emphasis>
num. CRBA 1246. 
<figureCitation box="[725,822,659,686]" captionStart="Figures 4-7" captionStartId="13.[140,222,1622,1644]" captionTargetBox="[140,1108,1166,1602]" captionTargetId="figure@13.[140,1109,1166,1603]" captionTargetPageId="13" captionText="Figures 4-7. Carapace, dorsal view: 4 D. jana sp. n. holotype 5 D. shardana sp. n. holotype. Carapace, lateral view: 6 D. jana sp. n. holotype 7 D. shardana sp. n. holotype." figureDoi="http://doi.org/10.5281/zenodo.3813919" httpUri="https://zenodo.org/record/3813919/files/figure.png" pageId="19" pageNumber="338">Figs 5, 7</figureCitation>
, 
<figureCitation box="[834,851,660,686]" captionStart="Figures 8-9" captionStartId="15.[140,222,1051,1073]" captionTargetBox="[140,1108,165,1031]" captionTargetId="figure@15.[140,1108,165,1032]" captionTargetPageId="15" captionText="Figures 8-9. Left male palp, retrolateral view: 8 D. jana sp. n. holotype 9 D. shardana sp. n. holotype." figureDoi="http://doi.org/10.5281/zenodo.3813921" httpUri="https://zenodo.org/record/3813921/files/figure.png" pageId="19" pageNumber="338">9</figureCitation>
, 
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. Carapace (
<figureCitation captionStart="Figures 4-7" captionStartId="13.[140,222,1622,1644]" captionTargetBox="[140,1108,1166,1602]" captionTargetId="figure@13.[140,1109,1166,1603]" captionTargetPageId="13" captionText="Figures 4-7. Carapace, dorsal view: 4 D. jana sp. n. holotype 5 D. shardana sp. n. holotype. Carapace, lateral view: 6 D. jana sp. n. holotype 7 D. shardana sp. n. holotype." figureDoi="http://doi.org/10.5281/zenodo.3813919" httpUri="https://zenodo.org/record/3813919/files/figure.png" pageId="19" pageNumber="338">Fig. 5</figureCitation>
) 
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long; maximum width 
<quantity box="[554,663,694,721]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" pageId="19" pageNumber="338" unit="mm" value="2.35">2.35 mm</quantity>
; minimum width 
<quantity box="[875,985,695,721]" metricMagnitude="-3" metricUnit="m" metricValue="1.68" pageId="19" pageNumber="338" unit="mm" value="1.68">1.68 mm</quantity>
. Dark red, darkened at borders; foveate at borders, slightly wrinkled in the middle, covered with small black grains. Frontal border roughly round, from 1/2 to 3/5 carapace length; anterior lateral borders parallel; rounded at maximum dorsal width, back lateral borders straight; back margin wide, straight. AME diameter 
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; PLE 
<quantity box="[924,1034,836,862]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="19" pageNumber="338" unit="mm" value="0.18">0.18 mm</quantity>
; PME 
<quantity box="[140,246,871,897]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="19" pageNumber="338" unit="mm" value="0.16">0.16 mm</quantity>
; AME slightly backwards from frontal border, separated from one another by about 2/3 diameter, close to PLE; PME close to each other, less than 1/4 PME diameter from PLE. Labium trapezoid-shaped, base wider than distal part; longer than wide at base; semicircular groove at tip. Sternum brownish orange, darkened on borders; wrinkled; uniformly covered in slender black hairs.
</paragraph>
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Chelicerae 
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long, about 2/5 of carapace length in dorsal view; fang long, 
<quantity box="[140,249,1082,1108]" metricMagnitude="-3" metricUnit="m" metricValue="1.22" pageId="19" pageNumber="338" unit="mm" value="1.22">1.22 mm</quantity>
; basal segment proximal dorsal, ventral side scantly covered with piligerous granulations; concave-shaped in lateral view (
<figureCitation box="[638,705,1117,1144]" captionStart="Figures 4-7" captionStartId="13.[140,222,1622,1644]" captionTargetBox="[140,1108,1166,1602]" captionTargetId="figure@13.[140,1109,1166,1603]" captionTargetPageId="13" captionText="Figures 4-7. Carapace, dorsal view: 4 D. jana sp. n. holotype 5 D. shardana sp. n. holotype. Carapace, lateral view: 6 D. jana sp. n. holotype 7 D. shardana sp. n. holotype." figureDoi="http://doi.org/10.5281/zenodo.3813919" httpUri="https://zenodo.org/record/3813919/files/figure.png" pageId="19" pageNumber="338">Fig. 7</figureCitation>
). Chelicera inner groove long, about 1/2 cheliceral length; armed with three teeth and lamina at base; B=M&gt;D; D triangular, slightly below groove midpoint; B close to basal lamina; M close to B. Legs yellow. Lengths of male described above: fe1 
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(all measurements in mm); pa1 1.3; ti1 1.84; me1 1.66; ta1 0.48; total 7.57; fe2 2.09; pa2 1.22; ti2 1.71; me2 1.58; ta2 0.51; total 7.11; fe3 1.58; pa3 0.87; ti3 1.07; me3 1.43; ta3 0.41; total 5.31; fe4 2.09; pa4 1.17; ti4 1.68; me4 1.89; ta4 0.51; total 7.34; fe Pdp 1.22; pa Pdp 0.66; ti Pdp 0.66; ta Pdp 0.71; total 3.26; relative length: 1&gt;4&gt;2&gt;3. Spination: leg1, leg2 spineless; fe3d spineless; pa3 spineless; tb3d spines arranged in two bands; proximal 1.0.0; distal 1.0.0; tb3v spines arranged in one band; proximal 0.1.0; with one prolateral spine on left leg, and two terminal on right leg. Fe4d spineless; pa4 spineless; tb4d spines arranged in two bands; proximal 1.0-1.1; distal 0.0.0-1; tb4v spines arranged in one band; proximal 0.1.0; with two terminal spines. Dorsal side of frontal legs, ventral side of palp covered with hairs. Claws with eight teeth or less; hardly larger than claw width.
</paragraph>
<paragraph blockId="19.[140,1109,166,1672]" pageId="19" pageNumber="338">
Abdomen 
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long; cream-coloured; cylindrical. Abdominal dorsal hairs 
<quantity box="[140,245,1645,1672]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="19" pageNumber="338" unit="mm" value="0.13">0.13 mm</quantity>
long; thin, curved, not compressed, pointed; uniformly thickly distributed.
</paragraph>
<paragraph blockId="20.[140,1108,167,1390]" pageId="20" pageNumber="339">
Male copulatory bulb (
<figureCitation box="[445,512,167,193]" captionStart="Figures 8-9" captionStartId="15.[140,222,1051,1073]" captionTargetBox="[140,1108,165,1031]" captionTargetId="figure@15.[140,1108,165,1032]" captionTargetPageId="15" captionText="Figures 8-9. Left male palp, retrolateral view: 8 D. jana sp. n. holotype 9 D. shardana sp. n. holotype." figureDoi="http://doi.org/10.5281/zenodo.3813921" httpUri="https://zenodo.org/record/3813921/files/figure.png" pageId="20" pageNumber="339">Fig. 9</figureCitation>
) T slightly shorter than DD; external, internal distal border sloped backwards. DD bent about 45 
<superScript attach="left" box="[642,650,201,215]" fontSize="6" pageId="20" pageNumber="339">o</superScript>
in lateral view; internal distal border not expanded. IS, ES equally developed; IS truncated at DD middle part. DD tip (
<figureCitation captionStart="Figures 19-22" captionStartId="18.[140,222,1622,1644]" captionTargetBox="[140,1108,408,1603]" captionTargetId="figure@18.[140,1108,408,1603]" captionTargetPageId="18" captionText="Figures 19-22. D. shardana holotype, right male bulb 19 prolateral view, general 20 prolateral view, tip detail 21 anterior view 22 P, prolateral view." figureDoi="http://doi.org/10.5281/zenodo.3813927" httpUri="https://zenodo.org/record/3813927/files/figure.png" pageId="20" pageNumber="339">Figs 19-21</figureCitation>
) straight in lateral view; AR present, parallel to DD; finger-like distal projection at AR retrolateral side, shorter than AR length. C absent. AC absent. LF absent. L reduced to distal part. LA absent. F absent. AL absent. P (
<figureCitation box="[805,887,343,369]" captionStart="Figures 19-22" captionStartId="18.[140,222,1622,1644]" captionTargetBox="[140,1108,408,1603]" captionTargetId="figure@18.[140,1108,408,1603]" captionTargetPageId="18" captionText="Figures 19-22. D. shardana holotype, right male bulb 19 prolateral view, general 20 prolateral view, tip detail 21 anterior view 22 P, prolateral view." figureDoi="http://doi.org/10.5281/zenodo.3813927" httpUri="https://zenodo.org/record/3813927/files/figure.png" pageId="20" pageNumber="339">Fig. 22</figureCitation>
) fused to T; sloped forming an angle of about 135° to T in lateral view; lateral length about 1/4 of T width; ridge present, parallel to T; not expanded, upper margin smooth; not distally projected; back margin slightly folded towards internal side.
</paragraph>
<paragraph blockId="20.[140,1108,167,1390]" pageId="20" pageNumber="339">
<emphasis box="[185,357,483,509]" italics="true" pageId="20" pageNumber="339">
<typeStatus box="[185,277,484,509]" pageId="20" pageNumber="339">Paratype</typeStatus>
female
</emphasis>
num. CRBA 1246. 
<figureCitation box="[598,723,483,510]" captionStart="Figures 10-13" captionStartId="16.[140,222,1622,1644]" captionTargetBox="[140,1108,554,1602]" captionTargetId="figure@16.[140,1108,551,1604]" captionTargetPageId="16" captionText="Figures 10-13. D. jana sp. n. paratype female 10 vulva, dorsal view 11 vulva, ventral view. D. shardana sp. n. paratype female 12 vulva, dorsal view 13 vulva, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3813923" httpUri="https://zenodo.org/record/3813923/files/figure.png" pageId="20" pageNumber="339">Figs 12-13</figureCitation>
. All characters as in male except: Carapace 
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long; maximum width 
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; minimum width 
<quantity box="[928,1036,519,545]" metricMagnitude="-3" metricUnit="m" metricValue="1.68" pageId="20" pageNumber="339" unit="mm" value="1.68">1.68 mm</quantity>
. AME diameter 
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; PLE 
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; PME 
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; AME slightly back from frontal border, separated from one another by about 2/3 diameter, touching PLE; PME very close to each other, less than 1/4 PME diameter from PLE. Sternum very slightly wrinkled, mainly between legs and frontal border.
</paragraph>
<paragraph blockId="20.[140,1108,167,1390]" pageId="20" pageNumber="339">
Chelicerae 
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long; fang 
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; basal segment proximal dorsal side scantly covered with piligerous granulations. Chelicera armed with three teeth and lamina at base; M&gt;B=D. Legs orange. Lengths of female described above: fe1 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="2.22" pageId="20" pageNumber="339" unit="mm" value="2.22">2.22 mm</quantity>
(all measurements in mm); pa1 1.35; ti1 1.73; me1 1.58; ta1 0.46; total 7.34; fe2 2.04; pa2 1.28; ti2 1.68; me2 1.53; ta2 0.48; total 7.01; fe3 1.66; pa3 0.92; ti3 1.15; me3 1.38; ta3 0.41; total 5.51; fe4 2.27; pa4 1.22; ti4 1.79; me4 1.89; ta4 0.48; total 7.65; fe Pdp 1.17; pa Pdp 0.61; ti Pdp 0.61; ta Pdp 0.71; total 3.11; relative length 4&gt;1&gt;2&gt;3. Spination: tb3d spines arranged in two bands; proximal 1.0.0; distal 1.0.0; tb3v with two terminal spines. Tb4d spines arranged in one band; proximal 1.0.1; tb4v spines arranged in one band; proximal 0.1.0; with two terminal spines.
</paragraph>
<paragraph blockId="20.[140,1108,167,1390]" pageId="20" pageNumber="339">
Abdomen 
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long. Abdominal dorsal hairs 
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long; medium-sized, curved, compressed, pointed; uniformly, thickly distributed. Vulva (
<figureCitation box="[922,1047,1082,1108]" captionStart="Figures 10-13" captionStartId="16.[140,222,1622,1644]" captionTargetBox="[140,1108,554,1602]" captionTargetId="figure@16.[140,1108,551,1604]" captionTargetPageId="16" captionText="Figures 10-13. D. jana sp. n. paratype female 10 vulva, dorsal view 11 vulva, ventral view. D. shardana sp. n. paratype female 12 vulva, dorsal view 13 vulva, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3813923" httpUri="https://zenodo.org/record/3813923/files/figure.png" pageId="20" pageNumber="339">Figs 12-13</figureCitation>
) DA clearly distinguishable from VA; DA slightly wider than long; DF wide in dorsal view. MF margins not fused, poorly developed, membranous (
<figureCitation box="[795,881,1152,1178]" captionStart="Figures 10-13" captionStartId="16.[140,222,1622,1644]" captionTargetBox="[140,1108,554,1602]" captionTargetId="figure@16.[140,1108,551,1604]" captionTargetPageId="16" captionText="Figures 10-13. D. jana sp. n. paratype female 10 vulva, dorsal view 11 vulva, ventral view. D. shardana sp. n. paratype female 12 vulva, dorsal view 13 vulva, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3813923" httpUri="https://zenodo.org/record/3813923/files/figure.png" pageId="20" pageNumber="339">Fig. 12</figureCitation>
). VA rectangle-like, pointed expansion at middle frontal part (
<figureCitation box="[627,711,1187,1214]" captionStart="Figures 10-13" captionStartId="16.[140,222,1622,1644]" captionTargetBox="[140,1108,554,1602]" captionTargetId="figure@16.[140,1108,551,1604]" captionTargetPageId="16" captionText="Figures 10-13. D. jana sp. n. paratype female 10 vulva, dorsal view 11 vulva, ventral view. D. shardana sp. n. paratype female 12 vulva, dorsal view 13 vulva, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3813923" httpUri="https://zenodo.org/record/3813923/files/figure.png" pageId="20" pageNumber="339">Fig. 13</figureCitation>
); frontal region completely sclerotized; posterior region internal margin sclerotized, forming two divergent stripes; AVD absent. S attachment not projected under VA; arms as long as DA, slightly curved; tips not projected; neck hardly visible. TB usual shape.
</paragraph>
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<emphasis bold="true" box="[185,304,1328,1354]" pageId="20" pageNumber="339">
<emphasis box="[185,298,1328,1354]" inLineHeading="true" pageId="20" pageNumber="339" reason="1">Variation</emphasis>
.
</emphasis>
Carapace ranges in length from 
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to 
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, males (n=7) average 2.79 (2.50-3.06), female (n=8) average 3.07 (2.68-3.42). In general male legs bear fewer spines that female legs, one male form Mt. S. Antonio representing an extreme case of reduced spination. Spination variability summarised in 
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.
</paragraph>
</subSubSection>
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<paragraph blockId="20.[140,733,1451,1473]" box="[140,733,1451,1473]" pageId="20" pageNumber="339">
<emphasis bold="true" box="[140,227,1451,1473]" pageId="20" pageNumber="339">Table 4.</emphasis>
Intraspecific spination variability in 
<taxonomicName authority="Arnedo &amp; Gasparo &amp; Opatova, 2009" authorityName="Arnedo &amp; Gasparo &amp; Opatova" authorityYear="2009" box="[566,677,1451,1473]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="339" phylum="Arthropoda" rank="species" species="shardana" status="sp. n.">
<emphasis box="[566,677,1451,1473]" italics="true" pageId="20" pageNumber="339">D. shardana</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[682,733,1452,1473]" pageId="20" pageNumber="339" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph pageId="20" pageNumber="339">
<table box="[147,1003,1506,1668]" colsContinueFrom="14.[147,987,1506,1668]" gridcols="5" gridrows="5" pageId="20" pageNumber="339">
<tr box="[147,1003,1506,1530]" gridrow="0" pageId="20" pageNumber="339" rowspan-0="1">
<th box="[341,440,1506,1530]" gridcol="1" gridrow="0" pageId="20" pageNumber="339">
<emphasis bold="true" box="[341,440,1506,1530]" pageId="20" pageNumber="339">Proximal</emphasis>
</th>
<th box="[534,867,1506,1530]" colspan="2" colspanRight="1" gridcol="2" gridrow="0" pageId="20" pageNumber="339">
<emphasis bold="true" box="[534,867,1506,1530]" pageId="20" pageNumber="339">Medio-proximal Medio-distal</emphasis>
</th>
<th box="[921,1003,1506,1530]" gridcol="4" gridrow="0" pageId="20" pageNumber="339">
<emphasis bold="true" box="[921,987,1506,1530]" pageId="20" pageNumber="339">Distal</emphasis>
</th>
</tr>
<tr box="[147,1003,1541,1565]" gridrow="1" pageId="20" pageNumber="339">
<th box="[147,298,1541,1565]" gridcol="0" gridrow="1" pageId="20" pageNumber="339">Tibia 3 dorsal</th>
<td box="[341,440,1541,1565]" gridcol="1" gridrow="1" pageId="20" pageNumber="339">1.0.0</td>
<td box="[534,547,1541,1565]" gridcol="2" gridrow="1" pageId="20" pageNumber="339">0</td>
<td box="[728,867,1541,1565]" gridcol="3" gridrow="1" pageId="20" pageNumber="339">0</td>
<td box="[921,1003,1541,1565]" gridcol="4" gridrow="1" pageId="20" pageNumber="339">1.0.0</td>
</tr>
<tr box="[147,1003,1575,1600]" gridrow="2" pageId="20" pageNumber="339">
<th box="[147,298,1575,1600]" gridcol="0" gridrow="2" pageId="20" pageNumber="339">Tibia 4 dorsal</th>
<td box="[341,440,1575,1600]" gridcol="1" gridrow="2" pageId="20" pageNumber="339">0-1.0-1.1</td>
<td box="[534,547,1575,1600]" gridcol="2" gridrow="2" pageId="20" pageNumber="339">0</td>
<td box="[728,867,1575,1600]" gridcol="3" gridrow="2" pageId="20" pageNumber="339">0</td>
<td box="[921,1003,1575,1600]" gridcol="4" gridrow="2" pageId="20" pageNumber="339">0.0.0-1</td>
</tr>
<tr box="[147,1003,1610,1634]" gridrow="3" pageId="20" pageNumber="339">
<th box="[147,298,1610,1634]" gridcol="0" gridrow="3" pageId="20" pageNumber="339">Tibia 3 ventral</th>
<td box="[341,440,1610,1634]" gridcol="1" gridrow="3" pageId="20" pageNumber="339">0. 0-1.0</td>
<td box="[534,547,1610,1634]" gridcol="2" gridrow="3" pageId="20" pageNumber="339">0</td>
<td box="[728,867,1610,1634]" gridcol="3" gridrow="3" pageId="20" pageNumber="339">0</td>
<td box="[921,1003,1610,1634]" gridcol="4" gridrow="3" pageId="20" pageNumber="339">1.0.0-1</td>
</tr>
<tr box="[147,1003,1644,1668]" gridrow="4" pageId="20" pageNumber="339">
<th box="[147,298,1644,1668]" gridcol="0" gridrow="4" pageId="20" pageNumber="339">Tibia 4 ventral</th>
<td box="[341,440,1644,1668]" gridcol="1" gridrow="4" pageId="20" pageNumber="339">0. 0-1.0</td>
<td box="[534,547,1644,1668]" gridcol="2" gridrow="4" pageId="20" pageNumber="339">0</td>
<td box="[728,867,1644,1668]" gridcol="3" gridrow="4" pageId="20" pageNumber="339">0</td>
<td box="[921,1003,1644,1668]" gridcol="4" gridrow="4" pageId="20" pageNumber="339">1.0. 0-1</td>
</tr>
</table>
</paragraph>
<subSubSection pageId="21" pageNumber="340" type="distribution">
<paragraph blockId="21.[140,1108,167,334]" pageId="21" pageNumber="340">
<emphasis bold="true" box="[185,344,237,263]" pageId="21" pageNumber="340">
<emphasis box="[185,339,237,263]" inLineHeading="true" pageId="21" pageNumber="340" reason="1">Distribution</emphasis>
.
</emphasis>
Known from several localities around the western half of 
<collectingRegion box="[1008,1104,237,263]" country="Italy" name="Sardegna" pageId="21" pageNumber="340">Sardinia</collectingRegion>
, and also in the Gennargentu region on the central-eastern side of the island, spanning from 360 to above 
<quantity box="[356,445,308,334]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="21" pageNumber="340" unit="m" value="1100.0">1100 m</quantity>
elevation.
</paragraph>
</subSubSection>
</treatment>
</document>