226 lines
22 KiB
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226 lines
22 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1126.90916" ID-Pensoft-Pub="1313-2970-1126-131" ID-Pensoft-UUID="236ADC4AFF7954779FD983E0DA8705F7" ID-ZooBank="CCCFB66B5138426FBE11824811AC9916" ModsDocID="1313-2970-1126-131" checkinTime="1667357846417" checkinUser="pensoft" docAuthor="Yao, Junli, van Achterberg, Cornelis, Yaakop, Salmah, Long, Khuat Dang, Sharkey, Michael J. & Chapman, Eric G." docDate="2022" docId="19E88865296F5792B0D3B1211383DDB2" docLanguage="en" docName="ZooKeys 1126: 131-154" docOrigin="ZooKeys 1126" docPubDate="2022-11-01" docSource="http://dx.doi.org/10.3897/zookeys.1126.90916" docTitle="Anamalysia vanhengstumi van Achterberg 2022, sp. nov." docType="treatment" docUuid="74022A0F-DE25-42EB-811D-E60E50365907" docUuidSource="ZooBank" docVersion="1" id="236ADC4AFF7954779FD983E0DA8705F7" lastPageNumber="131" masterDocId="236ADC4AFF7954779FD983E0DA8705F7" masterDocTitle="A new genus Anamalysia van Achterberg (Hymenoptera, Braconidae, Alysiinae), six new species, and two new combinations from India, Indonesia, Malaysia, Singapore, Thailand, and Vietnam" masterLastPageNumber="154" masterPageNumber="131" pageNumber="131" updateTime="1667357846417" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new genus Anamalysia van Achterberg (Hymenoptera, Braconidae, Alysiinae), six new species, and two new combinations from India, Indonesia, Malaysia, Singapore, Thailand, and Vietnam</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yao, Junli</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6064-1315</mods:nameIdentifier>
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<mods:affiliation>Biological Control Research Institute, Fujian Agriculture & Forestry University, Fuzhou, Fujian 350002, China</mods:affiliation>
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<mods:nameIdentifier type="email">rjuneyao@outlook.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>van Achterberg, Cornelis</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6495-4853</mods:nameIdentifier>
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<mods:affiliation>Naturalis Biodiversity Centre, Postbus 9517, 2300 RA Leiden, Netherlands</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yaakop, Salmah</mods:namePart>
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<mods:affiliation>Department of Biological Sciences and Biotechnology, Faculty of Science and Technology, 43600 National University of Malaysia, Bangi, Selangor, Malaysia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Long, Khuat Dang</mods:namePart>
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<mods:affiliation>Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, Cau Giay, Hanoi, Vietnam</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sharkey, Michael J.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6201-7340</mods:nameIdentifier>
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<mods:affiliation>Department of Entomology, University of Kentucky, Lexington KY 40546 - 0091, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Chapman, Eric G.</mods:namePart>
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<mods:affiliation>Department of Entomology, University of Kentucky, Lexington KY 40546 - 0091, USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-11-01</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1126</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>131</mods:start>
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<mods:end>154</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1126.90916</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1126.90916</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1126-131</mods:identifier>
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<mods:identifier type="ZooBank">CCCFB66B5138426FBE11824811AC9916</mods:identifier>
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<mods:identifier type="Pensoft-UUID">236ADC4AFF7954779FD983E0DA8705F7</mods:identifier>
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</mods:mods>
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<subSection pageId="0" pageNumber="131" type="multiple">
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<treatment LSID="urn:lsid:zoobank.org:act:74022A0F-DE25-42EB-811D-E60E50365907" httpUri="http://treatment.plazi.org/id/19E88865296F5792B0D3B1211383DDB2" lastPageNumber="131" pageId="0" pageNumber="131">
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<subSubSection pageId="0" pageNumber="131" type="nomenclature">
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<paragraph pageId="0" pageNumber="131">
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<taxonomicName LSID="https://zoobank.org/74022A0F-DE25-42EB-811D-E60E50365907" authority="van Achterberg" authorityName="van Achterberg" authorityYear="2022" family="Braconidae" genus="Anamalysia" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Anamalysia vanhengstumi" order="Hymenoptera" pageId="0" pageNumber="131" rank="species" species="vanhengstumi" status="sp. nov.">Anamalysia vanhengstumi van Achterberg</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="131">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="131" type="description">
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<paragraph pageId="0" pageNumber="131">
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Anamalysia vanhengstumi sp. nov., ♂, holotype A habitus, lateral aspect B detail of basal antennal segments." figureDoi="10.3897/zookeys.1126.90916.figure7" httpUri="https://binary.pensoft.net/fig/764274" pageId="0" pageNumber="131">Figs 7A, B</figureCitation>
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Anamalysia vanhengstumi sp. nov., ♂, holotype A wings B head, anterior aspect C mesosoma, lateral aspect D head and mesosoma, dorsal aspect E head, dorsal aspect F propodeum and first metasomal tergite, dorsal aspect G mandible, full sight on first tooth H mandible, with full sight on third tooth I hind leg." figureDoi="10.3897/zookeys.1126.90916.figure8" httpUri="https://binary.pensoft.net/fig/764275" pageId="0" pageNumber="131">, 8A-I</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="131" type="type material">
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<paragraph pageId="0" pageNumber="131">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="131">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="131">Holotype</emphasis>
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, ♂ (RMNH), "Alysi. 070", VN [= Vietnam], Ha Giang, Vi Xuyen, Cao Bo Rung TS, 400 m, 10.v.2007, K.D. Long.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="131" type="description">
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<paragraph pageId="0" pageNumber="131">Description.</paragraph>
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<paragraph pageId="0" pageNumber="131">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="131">Holotype</emphasis>
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, ♂, length of body 3.9 mm, length of fore wing 3.3 mm.
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</paragraph>
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<paragraph pageId="0" pageNumber="131">
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<emphasis bold="true" pageId="0" pageNumber="131">
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<emphasis italics="true" pageId="0" pageNumber="131">Head</emphasis>
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.
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</emphasis>
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Width of head 2.4 times its median length, deeply depressed medially and largely glabrous dorsally; antenna incomplete, with 29+ segments, setae short and adpressed and seven basal segments strongly shiny, length of third segment 0.9 times as long as fourth segment, length of third and fourth segments 3.9 and 4.6 times their width, respectively (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Anamalysia vanhengstumi sp. nov., ♂, holotype A habitus, lateral aspect B detail of basal antennal segments." figureDoi="10.3897/zookeys.1126.90916.figure7" httpUri="https://binary.pensoft.net/fig/764274" pageId="0" pageNumber="131">7B</figureCitation>
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); length of maxillary palp 1.2 times height of head; eye in dorsal view 1.1 times as long as temple; temple in dorsal view strongly widened behind eyes (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Anamalysia vanhengstumi sp. nov., ♂, holotype A wings B head, anterior aspect C mesosoma, lateral aspect D head and mesosoma, dorsal aspect E head, dorsal aspect F propodeum and first metasomal tergite, dorsal aspect G mandible, full sight on first tooth H mandible, with full sight on third tooth I hind leg." figureDoi="10.3897/zookeys.1126.90916.figure8" httpUri="https://binary.pensoft.net/fig/764275" pageId="0" pageNumber="131">8E</figureCitation>
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); OOL:diameter of ocellus: POL = 15:4:3; minimum width of face 0.6 times maximum width of head and 1.8 times its height, coarsely punctate, moderately convex, with long setae and medio-ventrally narrowly smooth (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Anamalysia vanhengstumi sp. nov., ♂, holotype A wings B head, anterior aspect C mesosoma, lateral aspect D head and mesosoma, dorsal aspect E head, dorsal aspect F propodeum and first metasomal tergite, dorsal aspect G mandible, full sight on first tooth H mandible, with full sight on third tooth I hind leg." figureDoi="10.3897/zookeys.1126.90916.figure8" httpUri="https://binary.pensoft.net/fig/764275" pageId="0" pageNumber="131">8B</figureCitation>
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); with oblique groove from antennal socket to eye; clypeus rather robust and largely smooth (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Anamalysia vanhengstumi sp. nov., ♂, holotype A wings B head, anterior aspect C mesosoma, lateral aspect D head and mesosoma, dorsal aspect E head, dorsal aspect F propodeum and first metasomal tergite, dorsal aspect G mandible, full sight on first tooth H mandible, with full sight on third tooth I hind leg." figureDoi="10.3897/zookeys.1126.90916.figure8" httpUri="https://binary.pensoft.net/fig/764275" pageId="0" pageNumber="131">8B</figureCitation>
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); vertex strongly shiny and weakly convex and strongly depressed behind stemmaticum; anterior tentorial pit covered by mandible; length of malar space 0.1 times basal width of mandible; mandible strongly widened, 1.4 times as long as wide, middle tooth dorsally connected to wide sinuate and up curved lamella from upper corner of mandible, medially coarsely rugose, first tooth part of apical lamella and third tooth medium-sized (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Anamalysia vanhengstumi sp. nov., ♂, holotype A wings B head, anterior aspect C mesosoma, lateral aspect D head and mesosoma, dorsal aspect E head, dorsal aspect F propodeum and first metasomal tergite, dorsal aspect G mandible, full sight on first tooth H mandible, with full sight on third tooth I hind leg." figureDoi="10.3897/zookeys.1126.90916.figure8" httpUri="https://binary.pensoft.net/fig/764275" pageId="0" pageNumber="131">8G, H</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.1126.90916.figure7" httpUri="https://binary.pensoft.net/fig/764274" pageId="0" pageNumber="131" start="Figure 7" startId="F7">
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<paragraph pageId="0" pageNumber="131">
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<emphasis bold="true" pageId="0" pageNumber="131">Figure 7.</emphasis>
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<taxonomicName authorityName="van Achterberg" authorityYear="2022" family="Braconidae" genus="Anamalysia" kingdom="Animalia" lsidName="Anamalysia vanhengstumi" order="Hymenoptera" pageId="0" pageNumber="131" rank="species" species="vanhengstumi">
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<emphasis italics="true" pageId="0" pageNumber="131">Anamalysia vanhengstumi</emphasis>
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</taxonomicName>
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sp. nov., ♂, holotype
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<emphasis bold="true" pageId="0" pageNumber="131">A</emphasis>
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habitus, lateral aspect
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<emphasis bold="true" pageId="0" pageNumber="131">B</emphasis>
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detail of basal antennal segments.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="131">
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<emphasis bold="true" pageId="0" pageNumber="131">
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<emphasis italics="true" pageId="0" pageNumber="131">Mesosoma</emphasis>
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.
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</emphasis>
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Length of mesosoma 1.7 times its height; pronotum with large pronope; antescutal depression absent; side of pronotum largely crenulate medially (except subposteriorly) and posteriorly; epicnemial area with few crenulae; precoxal sulcus absent posteriorly, wide and coarsely crenulate (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Anamalysia vanhengstumi sp. nov., ♂, holotype A wings B head, anterior aspect C mesosoma, lateral aspect D head and mesosoma, dorsal aspect E head, dorsal aspect F propodeum and first metasomal tergite, dorsal aspect G mandible, full sight on first tooth H mandible, with full sight on third tooth I hind leg." figureDoi="10.3897/zookeys.1126.90916.figure8" httpUri="https://binary.pensoft.net/fig/764275" pageId="0" pageNumber="131">8C</figureCitation>
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); remainder of mesopleuron smooth; episternal scrobe elongate and medium-sized; pleural sulcus finely crenulate dorsally and distinctly crenulate ventrally; mesosternal sulcus wide and coarsely crenulate; metapleuron largely smooth but rugose ventrally; notauli complete, deeply impressed, rather narrow, and smooth; medio-posterior depression absent (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Anamalysia vanhengstumi sp. nov., ♂, holotype A wings B head, anterior aspect C mesosoma, lateral aspect D head and mesosoma, dorsal aspect E head, dorsal aspect F propodeum and first metasomal tergite, dorsal aspect G mandible, full sight on first tooth H mandible, with full sight on third tooth I hind leg." figureDoi="10.3897/zookeys.1126.90916.figure8" httpUri="https://binary.pensoft.net/fig/764275" pageId="0" pageNumber="131">8D</figureCitation>
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); mesoscutum smooth, glabrous, and its lateral carina complete in front of tegulae; scutellar sulcus deep, about 2.5 times as wide as long, with one carina, 0.3 times as long as scutellum; scutellum slightly convex, smooth except a few punctures; metanotum with complete median carina and rather protruding dorsally; propodeum smooth antero-laterally, reticulate medially and areolate posteriorly, with posteriorly narrowed pentagonal areola medially (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Anamalysia vanhengstumi sp. nov., ♂, holotype A wings B head, anterior aspect C mesosoma, lateral aspect D head and mesosoma, dorsal aspect E head, dorsal aspect F propodeum and first metasomal tergite, dorsal aspect G mandible, full sight on first tooth H mandible, with full sight on third tooth I hind leg." figureDoi="10.3897/zookeys.1126.90916.figure8" httpUri="https://binary.pensoft.net/fig/764275" pageId="0" pageNumber="131">8F</figureCitation>
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); propodeal spiracle small, round, and submedially in propodeum.
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</paragraph>
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<caption doi="10.3897/zookeys.1126.90916.figure8" httpUri="https://binary.pensoft.net/fig/764275" pageId="0" pageNumber="131" start="Figure 8" startId="F8">
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<paragraph pageId="0" pageNumber="131">
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<emphasis bold="true" pageId="0" pageNumber="131">Figure 8.</emphasis>
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<taxonomicName authorityName="van Achterberg" authorityYear="2022" family="Braconidae" genus="Anamalysia" kingdom="Animalia" lsidName="Anamalysia vanhengstumi" order="Hymenoptera" pageId="0" pageNumber="131" rank="species" species="vanhengstumi">
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<emphasis italics="true" pageId="0" pageNumber="131">Anamalysia vanhengstumi</emphasis>
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</taxonomicName>
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sp. nov., ♂, holotype
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<emphasis bold="true" pageId="0" pageNumber="131">A</emphasis>
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wings
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<emphasis bold="true" pageId="0" pageNumber="131">B</emphasis>
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head, anterior aspect
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<emphasis bold="true" pageId="0" pageNumber="131">C</emphasis>
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mesosoma, lateral aspect
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<emphasis bold="true" pageId="0" pageNumber="131">D</emphasis>
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head and mesosoma, dorsal aspect
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<emphasis bold="true" pageId="0" pageNumber="131">E</emphasis>
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head, dorsal aspect
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<emphasis bold="true" pageId="0" pageNumber="131">F</emphasis>
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propodeum and first metasomal tergite, dorsal aspect
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<emphasis bold="true" pageId="0" pageNumber="131">G</emphasis>
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mandible, full sight on first tooth
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<emphasis bold="true" pageId="0" pageNumber="131">H</emphasis>
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mandible, with full sight on third tooth
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<emphasis bold="true" pageId="0" pageNumber="131">I</emphasis>
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hind leg.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="131">
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<emphasis bold="true" pageId="0" pageNumber="131">
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<emphasis italics="true" pageId="0" pageNumber="131">Wings</emphasis>
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.
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</emphasis>
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Pterostigma elongate subtriangular, its posterior margin straight or slightly convex; vein r issued from basal 0.6 of pterostigma and oblique; 1-R1 narrow; r:3-SR:SR1 = 5:22:42; 1- SR+M narrow and nearly straight; SR1 straight; cu-a medium-sized, postfurcal; 1-CU1:2-CU1 = 1:11; 2-SR:3-SR:r-m = 20:22:9; r-m weakly inclivous; m-cu subinterstitial and slightly curved, converging to 1-M; 3-CU1 much longer than CU1b and narrow (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Anamalysia vanhengstumi sp. nov., ♂, holotype A wings B head, anterior aspect C mesosoma, lateral aspect D head and mesosoma, dorsal aspect E head, dorsal aspect F propodeum and first metasomal tergite, dorsal aspect G mandible, full sight on first tooth H mandible, with full sight on third tooth I hind leg." figureDoi="10.3897/zookeys.1126.90916.figure8" httpUri="https://binary.pensoft.net/fig/764275" pageId="0" pageNumber="131">8A</figureCitation>
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). Hind wing: M+CU:1-M (up to m-cu):1r-m = 20:18:9; m-cu distinct, largely unsclerotised (except basally) and interstitial (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Anamalysia vanhengstumi sp. nov., ♂, holotype A wings B head, anterior aspect C mesosoma, lateral aspect D head and mesosoma, dorsal aspect E head, dorsal aspect F propodeum and first metasomal tergite, dorsal aspect G mandible, full sight on first tooth H mandible, with full sight on third tooth I hind leg." figureDoi="10.3897/zookeys.1126.90916.figure8" httpUri="https://binary.pensoft.net/fig/764275" pageId="0" pageNumber="131">8A</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="131">
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<emphasis bold="true" pageId="0" pageNumber="131">
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<emphasis italics="true" pageId="0" pageNumber="131">Legs</emphasis>
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.
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</emphasis>
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Hind coxa smooth, baso-ventrally rounded, and not protruding; tarsal claws rather robust; length of femur, tibia, and basitarsus of hind leg 4.2, 11.0, and 9.0 times their width, respectively; hind tibia and basitarsus with numerous rather long setae dorsally; hind tibia densely setose and comb at inner apex of tibia absent; fore tarsus 1.4 times as long as fore tibia (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Anamalysia vanhengstumi sp. nov., ♂, holotype A wings B head, anterior aspect C mesosoma, lateral aspect D head and mesosoma, dorsal aspect E head, dorsal aspect F propodeum and first metasomal tergite, dorsal aspect G mandible, full sight on first tooth H mandible, with full sight on third tooth I hind leg." figureDoi="10.3897/zookeys.1126.90916.figure8" httpUri="https://binary.pensoft.net/fig/764275" pageId="0" pageNumber="131">8I</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="131">
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<emphasis bold="true" pageId="0" pageNumber="131">
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<emphasis italics="true" pageId="0" pageNumber="131">Metasoma</emphasis>
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.
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</emphasis>
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Length of first tergite 1.4 times its apical width, its surface longitudinally costate, its dorsal carinae united submedially (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Anamalysia vanhengstumi sp. nov., ♂, holotype A wings B head, anterior aspect C mesosoma, lateral aspect D head and mesosoma, dorsal aspect E head, dorsal aspect F propodeum and first metasomal tergite, dorsal aspect G mandible, full sight on first tooth H mandible, with full sight on third tooth I hind leg." figureDoi="10.3897/zookeys.1126.90916.figure8" httpUri="https://binary.pensoft.net/fig/764275" pageId="0" pageNumber="131">8F</figureCitation>
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); laterope deep and large; dorsope large, deep, and pointing dorsally; remainder of metasoma smooth and depressed; hypopygium truncate posteriorly (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Anamalysia vanhengstumi sp. nov., ♂, holotype A habitus, lateral aspect B detail of basal antennal segments." figureDoi="10.3897/zookeys.1126.90916.figure7" httpUri="https://binary.pensoft.net/fig/764274" pageId="0" pageNumber="131">7A</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="131">
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<emphasis bold="true" pageId="0" pageNumber="131">
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<emphasis italics="true" pageId="0" pageNumber="131">Colour</emphasis>
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.
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</emphasis>
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Black; scapus, pedicellus, mandible, and legs yellow; remainder of antenna dark brown; temple and vertex and metasoma laterally (except first tergite) chestnut brown; palpi ivory; tegulae mainly yellowish brown; pterostigma and most veins brown; wing membrane faintly infuscated.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="131" type="distribution">
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<paragraph pageId="0" pageNumber="131">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="131">Northern Vietnam.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="131" type="etymology">
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<paragraph pageId="0" pageNumber="131">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="131">Named after the former director of the National Museum of Natural History (Naturalis) Ronald van Hengstum (1952-2007), who tragically died after a short swim in the North Sea near The Hague. He visited Vietnam during one of the RMNH-IEBR expeditions and was strongly in favour of cooperation with our Vietnamese counterparts.</paragraph>
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</subSubSection>
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</treatment>
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</subSection>
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</document> |