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<document id="0648E8864FCE8664DBBDD09F555577B1" ID-CLB-Dataset="8244" ID-DOI="10.11646/zootaxa.4860.1.4" ID-GBIF-Dataset="547a2201-b826-4c5d-a541-1139ad9ee6e7" ID-ISSN="1175-5326" ID-Zenodo-Dep="4413559" ID-ZooBank="904B90F0-65C6-4138-9680-B53B453FB5A7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609700737756" checkinUser="plazi" docAuthor="Silva, Vinícius De Assis, Salles, Frederico Falcão &amp; Pinto, Ângelo Parise" docDate="2020" docId="1558A064FFD187685BFFFB210577F952" docLanguage="en" docName="zootaxa.4860.1.4.pdf" docOrigin="Zootaxa 4860 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thraulodes marianoi Silva &amp; Salles &amp; Pinto 2020, sp. nov." docType="treatment" docUuid="DD0D51D8-12F3-4FC8-BDE4-AA7A07B1031B" docUuidSource="ZooBank" docVersion="8" lastPageNumber="93" masterDocId="E961D81CFFD087695B68FFD30161FFE5" masterDocTitle="Thraulodes marianoi sp. nov., a remarkable new species of mayfly (Ephemeroptera Leptophlebiidae) with dark wings from the southern Brazilian Atlantic Forest" masterLastPageNumber="100" masterPageNumber="92" pageNumber="93" updateTime="1698851851270" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="081250F54078D07C1E6D7A232A8166FF">Thraulodes marianoi sp. nov., a remarkable new species of mayfly (Ephemeroptera Leptophlebiidae) with dark wings from the southern Brazilian Atlantic Forest</mods:title>
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<mods:role id="24713921ACF246C90B5A2A86ABE87656">
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<mods:namePart id="403A71850D4485B77F0069CCE17C99C2">Silva, Vinícius De Assis</mods:namePart>
<mods:nameIdentifier id="0076A98ED51CFC15EBD678084312099C" type="ORCID">0000-0002-1650-5666</mods:nameIdentifier>
<mods:affiliation id="6FC4B6809A3F391B43BA09A69011FAFF">Programa de Pós-Graduação em Ciências Biológicas (Entomologia) - PPGEnto, Universidade Federal do Paraná, Curitiba, PR, Brazil. &amp; viniciusdeassisidl @ gmail. com; https: // orcid. org / 0000 - 0002 - 1002 - 1871 &amp; Laboratory of Systematics on Aquatic Insects (LABSIA), Departamento de Zoologia, Universidade Federal do Paraná, P. O. Box 19020, 81531 - 980, Curitiba, PR, Brazil. &amp; viniciusdeassisidl @ gmail. com; https: // orcid. org / 0000 - 0002 - 1002 - 1871 &amp; https: // orcid. org / 0000 - 0002 - 1650 - 5666</mods:affiliation>
<mods:nameIdentifier id="E0FF82414B947B561CF02F52DBEB01DE" type="email">viniciusdeassisidl@gmail.com</mods:nameIdentifier>
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<mods:role id="BAC824510019DAC7D7AF8CD8853D6175">
<mods:roleTerm id="FF3CBD8A2BC136AAB21E279F8127F9D7">Author</mods:roleTerm>
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<mods:namePart id="764584D78552DFB42AFA626D09EAC51A">Salles, Frederico Falcão</mods:namePart>
<mods:nameIdentifier id="2FAB1A50FBBEB9509056A0FC01B2BFB3" type="ORCID">0000-0001-8331-5929</mods:nameIdentifier>
<mods:affiliation id="AB7F0F09B91F174EA74904494826C13F">Museu de Entomologia, Departamento de Entomologia, Universidade Federal de Viçosa, MG, Brazil. &amp; frederico. salles @ ufv. br; https: // orcid. org / 0000 - 0001 - 8331 - 5929</mods:affiliation>
<mods:nameIdentifier id="2080E8C295408AAD7B1D00D59FC27C13" type="email">frederico.salles@ufv.br</mods:nameIdentifier>
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<mods:namePart id="40EB4015FC99FBB252492BB27CB2E215">Pinto, Ângelo Parise</mods:namePart>
<mods:nameIdentifier id="472D185D1011365E4DF727C90EB7976C" type="ORCID">0000-0002-1650-5666</mods:nameIdentifier>
<mods:affiliation id="EFDB5FB144117C3140614C1C2B482230">Laboratory of Systematics on Aquatic Insects (LABSIA), Departamento de Zoologia, Universidade Federal do Paraná, P. O. Box 19020, 81531 - 980, Curitiba, PR, Brazil. &amp; viniciusdeassisidl @ gmail. com; https: // orcid. org / 0000 - 0002 - 1002 - 1871 &amp; https: // orcid. org / 0000 - 0002 - 1650 - 5666</mods:affiliation>
<mods:nameIdentifier id="0A862C951D36585C120D680AC9710381" type="email">viniciusdeassisidl@gmail.com</mods:nameIdentifier>
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<mods:title id="365AC8CF5D2A7410DCE62DE8F363711A">Zootaxa</mods:title>
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<mods:date id="5A3E15BDA64C8AE37D80EEA25AE57B40">2020</mods:date>
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<mods:number id="2F0EA72E169A0C5E24C692F3A5CA5AB7">2020-10-09</mods:number>
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<mods:number id="F324FBA6D5BB35CA0B80EEDFBE0D4D11">4860</mods:number>
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<treatment id="1558A064FFD187685BFFFB210577F952" ID-DOI="http://doi.org/10.5281/zenodo.4413542" ID-GBIF-Taxon="177324037" ID-Zenodo-Dep="4413542" LSID="urn:lsid:zoobank.org:act:DD0D51D8-12F3-4FC8-BDE4-AA7A07B1031B" httpUri="http://treatment.plazi.org/id/1558A064FFD187685BFFFB210577F952" lastPageNumber="93" pageId="1" pageNumber="93">
<subSubSection id="D5EB42F9FFD187685BFFFB21009CFAE8" box="[151,509,1266,1293]" pageId="1" pageNumber="93" type="nomenclature">
<paragraph id="9D4E1172FFD187685BFFFB21009CFAE8" blockId="1.[151,911,1266,1359]" box="[151,509,1266,1293]" pageId="1" pageNumber="93">
<heading id="C606A61EFFD187685BFFFB21009CFAE8" bold="true" box="[151,509,1266,1293]" fontSize="11" level="1" pageId="1" pageNumber="93" reason="1">
<emphasis id="AF85CD60FFD187685BFFFB21009CFAE8" bold="true" box="[151,509,1266,1293]" pageId="1" pageNumber="93">
<taxonomicName id="5AF16AF1FFD187685BFFFB2100F9FAE8" authority="Silva &amp; Salles &amp; Pinto, 2020" authorityName="Silva &amp; Salles &amp; Pinto" authorityYear="2020" box="[151,408,1266,1293]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="93" phylum="Arthropoda" rank="species" species="marianoi" status="sp. nov.">
<emphasis id="AF85CD60FFD187685BFFFB2100F9FAE8" bold="true" box="[151,408,1266,1293]" italics="true" pageId="1" pageNumber="93">Thraulodes marianoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B4B6701BFFD187685AF7FB20009CFAE8" box="[415,509,1267,1293]" pageId="1" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D5EB42F9FFD187685BFFFAC402EEFAAA" pageId="1" pageNumber="93" type="description">
<paragraph id="9D4E1172FFD187685BFFFAC40028FAD4" blockId="1.[151,911,1266,1359]" box="[151,329,1303,1329]" pageId="1" pageNumber="93">
(
<figureCitation id="05CA0DF7FFD187685BF7FAC40079FAD4" box="[159,280,1303,1329]" captionStart-0="FIGURE 3–5" captionStart-1="FIGURES 6–8" captionStart-2="FIGURES 9–11" captionStartId-0="3.[151,250,1450,1474]" captionStartId-1="4.[151,264,1269,1293]" captionStartId-2="5.[151,264,1370,1394]" captionTargetBox-0="[177,1411,525,1427]" captionTargetBox-1="[195,1387,387,1239]" captionTargetBox-2="[189,1398,381,1347]" captionTargetId-0="figure-178@3.[177,1411,525,1427]" captionTargetId-1="figure-119@4.[189,1398,381,1246]" captionTargetId-2="figure-121@5.[189,1398,381,1347]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 3–5. Thraulodes marianoi sp. nov., holotype male (Brazil. PR: Pico do Marumbi State Park, DZUP 515215): (3) habitus in lateral view; (4) head and thorax in lateral view; (5) head and thorax in dorsal view." captionText-1="FIGURES 6–8. Thraulodes marianoi sp. nov. holotype male (Brazil. PR: Pico do Marumbi State Park, DZUP 515215): (6) forewing; (7) hindwing, (8) fore-, middle- and hind legs in lateral view." captionText-2="FIGURES 9–11. Genitalia of Thraulodes marianoi sp. nov. holotype male (Brazil. PR: Pico do Marumbi State Park, DZUP 515215): (9) dorsal view, (10) ventral view and (11) ventrolateral view. Figures 10–11 with the 9th segment removed." figureDoi-0="http://doi.org/10.5281/zenodo.4413563" figureDoi-1="http://doi.org/10.5281/zenodo.4413565" figureDoi-2="http://doi.org/10.5281/zenodo.4413567" httpUri-0="https://zenodo.org/record/4413563/files/figure.png" httpUri-1="https://zenodo.org/record/4413565/files/figure.png" httpUri-2="https://zenodo.org/record/4413567/files/figure.png" pageId="1" pageNumber="93">Figs. 3–11</figureCitation>
,
<figureCitation id="05CA0DF7FFD187685A4CFAC40021FAD4" box="[292,320,1303,1329]" captionStart="FIGURES 12–13" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[208,1379,181,1919]" captionTargetId="figure-19@6.[208,1379,181,1919]" captionTargetPageId="6" captionText="FIGURES 12–13. Outline of male genitalia of Thraulodes species in ventral view: (12) T. basimaculatus holotype (redrawn from Giordano &amp; Domínguez, 2005); (13) T. marianoi holotype." figureDoi="http://doi.org/10.5281/zenodo.4413569" httpUri="https://zenodo.org/record/4413569/files/figure.png" pageId="1" pageNumber="93">13</figureCitation>
)
</paragraph>
<paragraph id="9D4E1172FFD187685BFFFAE402EEFAAA" blockId="1.[151,911,1266,1359]" box="[151,911,1335,1359]" pageId="1" pageNumber="93">
<uri id="E9601D70FFD187685BFFFAE402EEFAAA" box="[151,911,1335,1359]" pageId="1" pageNumber="93">
urn:lsid:zoobank.org:act:
<uuid id="E9572BA7FFD187685AFCFAE402EEFAAA" box="[404,911,1335,1359]" pageId="1" pageNumber="93">DD0D51D8-12F3-4FC8-BDE4-AA7A07B1031B</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="D5EB42F9FFD187685BFFFAAE0402FA05" pageId="1" pageNumber="93" type="materials_examined">
<paragraph id="9D4E1172FFD187685BFFFAAE0402FA05" blockId="1.[151,1436,1405,1719]" pageId="1" pageNumber="93">
<emphasis id="AF85CD60FFD187685BFFFAAE001AFA72" bold="true" box="[151,379,1405,1431]" pageId="1" pageNumber="93">Material examined</emphasis>
.
<materialsCitation id="2D991B2FFFD187685AE2FAAE0402FA05" ID-GBIF-Occurrence="3034607301" collectingDate="2018-11-28" collectionCode="R" country="Brazil" county="Male" elevation="1021" latitude="-21.49639" location="Pico do Marumbi State Park" longLatPrecision="21" longitude="-48.981667" municipality="Piraquara" pageId="1" pageNumber="93" specimenCode="DZUP 515215" specimenCount="1" stateProvince="Parana" typeStatus="holotype">
<typeStatus id="424AAFD0FFD187685AE2FAAE0096FA72" box="[394,503,1405,1431]" pageId="1" pageNumber="93" type="holotype">Holotype</typeStatus>
:
<collectingCounty id="742F69FEFFD18768596BFAAE035CFA72" box="[515,573,1405,1431]" pageId="1" pageNumber="93">Male</collectingCounty>
imago (in alcohol),
<collectingCountry id="E5E651E2FFD18768584EFAAD02EFFA7D" box="[806,910,1406,1432]" name="Brazil" pageId="1" pageNumber="93">BRAZIL</collectingCountry>
,
<collectingRegion id="5F35DF90FFD1876858F3FAAE054AFA72" box="[923,1067,1405,1431]" country="Brazil" name="Parana" pageId="1" pageNumber="93">Paraná State</collectingRegion>
,
<collectingMunicipality id="7D2A8B08FFD187685F5FFAAE05C3FA72" box="[1079,1186,1405,1431]" pageId="1" pageNumber="93">Piraquara</collectingMunicipality>
municipality,
<location id="982E47A9FFD187685E2EFAAE001AFA5E" LSID="urn:lsid:plazi:treatment:1558A064FFD187685BFFFB210577F952:982E47A9FFD187685E2EFAAE001AFA5E" country="Brazil" county="Male" latitude="-21.49639" longLatPrecision="21" longitude="-48.981667" municipality="Piraquara" name="Pico do Marumbi State Park" pageId="1" pageNumber="93" stateProvince="Parana">Pico do Marumbi State Park</location>
,
<location id="982E47A9FFD187685AEEFA72030BFA5E" LSID="urn:lsid:plazi:treatment:1558A064FFD187685BFFFB210577F952:982E47A9FFD187685AEEFA72030BFA5E" box="[390,618,1441,1467]" country="Brazil" county="Male" latitude="-21.49639" longLatPrecision="21" longitude="-48.981667" municipality="Piraquara" name="Mananciais da Serra" pageId="1" pageNumber="93" stateProvince="Parana">
Mananciais
<collectorName id="300474A4FFD187685966FA72030BFA5E" box="[526,618,1441,1467]" pageId="1" pageNumber="93">da Serra</collectorName>
</location>
(SANEPAR),
<location id="982E47A9FFD187685867FA720550FA5E" LSID="urn:lsid:plazi:treatment:1558A064FFD187685BFFFB210577F952:982E47A9FFD187685867FA720550FA5E" box="[783,1073,1441,1467]" country="Brazil" county="Male" latitude="-21.49639" longLatPrecision="21" longitude="-48.981667" municipality="Piraquara" name="Reservatorio do Carvalho" pageId="1" pageNumber="93" stateProvince="Parana">Reservatório do Carvalho</location>
,
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,
<geoCoordinate id="F8C577B5FFD187685FA4FA720438FA59" box="[1228,1369,1441,1468]" degrees="48" direction="west" minutes="58" orientation="longitude" pageId="1" pageNumber="93" precision="15" seconds="54" value="-48.981667">48º58’54”W</geoCoordinate>
,
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<quantity id="5A09BC97FFD187685E0CFA7201CCFA05" metricMagnitude="3" metricUnit="m" metricValue="1.021" pageId="1" pageNumber="93" unit="m" value="1021.0">1021 m</quantity>
a.s.l.
</elevation>
, light sheet, lic. 04/18,
<date id="E94F37B2FFD187685A98FA160313FA3A" box="[496,626,1477,1503]" pageId="1" pageNumber="93" value="2018-11-28">
<collectingDate id="F90BCE5AFFD187685A98FA160313FA3A" box="[496,626,1477,1503]" pageId="1" pageNumber="93" value="2018-11-28">28.XI.2018</collectingDate>
</date>
,
<collectorName id="300474A4FFD187685914FA15038EFA3A" box="[636,751,1477,1504]" pageId="1" pageNumber="93">A.P. Pinto</collectorName>
, B.
<collectionCode id="FBE089B7FFD18768587BFA15024BFA05" box="[787,810,1478,1504]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="93">R</collectionCode>
.
<collectorName id="300474A4FFD18768585BFA1602E0FA3A" box="[819,897,1477,1503]" pageId="1" pageNumber="93">Araujo</collectorName>
&amp;
<collectorName id="300474A4FFD1876858CBFA16050FFA3A" box="[931,1134,1477,1503]" pageId="1" pageNumber="93">A.C. Domahovski</collectorName>
leg. (
<specimenCode id="CD57B909FFD187685FC5FA160436FA05" box="[1197,1367,1477,1504]" collectionCode="DZUP" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="1" pageNumber="93" type="University or college">DZUP 515215</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D5EB42F9FFD187685BAFFA3A0577F952" pageId="1" pageNumber="93" type="diagnosis">
<paragraph id="9D4E1172FFD187685BAFFA3A0577F952" blockId="1.[151,1436,1405,1719]" pageId="1" pageNumber="93">
<emphasis id="AF85CD60FFD187685BAFFA3A0023F9E6" bold="true" box="[199,322,1513,1539]" pageId="1" pageNumber="93">Diagnosis.</emphasis>
The new species can be distinguished from the other species of
<taxonomicName id="5AF16AF1FFD187685F74FA3A05FDF9E6" baseAuthorityName="Kluge" baseAuthorityYear="2020" box="[1052,1180,1513,1539]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="AF85CD60FFD187685F74FA3A05FDF9E6" box="[1052,1180,1513,1539]" italics="true" pageId="1" pageNumber="93">Thraulodes</emphasis>
</taxonomicName>
by the combination of the following characteristics: (1) four cross-veins basal to bulla in forewing; (2) brown area covering about of proximal half of the forewing; (3) pleura violet; (4) terga and sterna I–IV white washed with dark brown on posterior and lateral margin terga and sterna V–X dark brown; (5) penes short and wide with distinct lateral area “ear-like” and poorly developed lateral pouch; (6) styliger plate triangular, median projection short and rounded on the apex; (7) middle and posterior femora dark brown with yellowish hypodermal coloration.
</paragraph>
</subSubSection>
</treatment>
</document>
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