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<document id="9987E4C953159DC56B7EF3709D3FE9AB" ID-CLB-Dataset="6423" ID-DOI="10.11646/zootaxa.4968.1.1" ID-GBIF-Dataset="8f948bf9-6e6b-4167-aa50-bf648409d0e5" ID-ISSN="1175-5326" ID-Zenodo-Dep="4745566" ID-ZooBank="09F4CC3C-879C-4FCD-94D5-9ADE4A81EFAC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1620638208101" checkinUser="plazi" docAuthor="Sinclair, Bradley J. &amp; Shamshev, Igor V." docDate="2021" docId="0E6887D4FFFEFFCF4EBCF91BFA00B76D" docLanguage="en" docName="zootaxa.4968.1.1.pdf" docOrigin="Zootaxa 4968 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Iteaphila Zetterstedt" docType="key" docVersion="6" lastPageNumber="81" masterDocId="F251FFACFFB1FF9F4E2BFFCBFF9CB20D" masterDocTitle="World revision of Iteaphila with unbranched radial vein (Diptera: Empidoidea: Iteaphilidae)" masterLastPageNumber="89" masterPageNumber="1" pageNumber="80" updateTime="1698938463478" updateUser="ExternalLinkService">
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<mods:title id="9F7976F0C8D70CE8788F99C7AEF90E73">World revision of Iteaphila with unbranched radial vein (Diptera: Empidoidea: Iteaphilidae)</mods:title>
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<mods:namePart id="A19282579D257B227BC369E7CD824D5E">Sinclair, Bradley J.</mods:namePart>
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<mods:namePart id="3CE0BB1E51C69EB6315CC66E9A3E2957">Shamshev, Igor V.</mods:namePart>
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<mods:title id="2CD525C1B5748DBFE821AAC660492E83">Zootaxa</mods:title>
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<treatment id="0E6887D4FFFEFFCF4EBCF91BFA00B76D" ID-DOI="http://doi.org/10.5281/zenodo.4814524" ID-GBIF-Taxon="182399053" ID-Zenodo-Dep="4814524" LSID="urn:lsid:plazi:treatment:0E6887D4FFFEFFCF4EBCF91BFA00B76D" httpUri="http://treatment.plazi.org/id/0E6887D4FFFEFFCF4EBCF91BFA00B76D" lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80">
<subSubSection id="CEDB6549FFFEFFCF4EBCF91BFA00B76D" lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80" type="key">
<subSubSection id="CEDB6549FFFEFFD04EBCF91BFBE6B4E7" box="[151,1146,1743,1777]" pageId="79" pageNumber="80" type="nomenclature">
<paragraph id="867E36C2FFFEFFD04EBCF91BFBE6B4E7" blockId="79.[151,1146,1743,1770]" box="[151,1146,1743,1777]" pageId="79" pageNumber="80">
<emphasis id="B4B5EAD0FFFEFFD04EBCF91BFBE6B4E7" bold="true" box="[151,1146,1743,1777]" pageId="79" pageNumber="80">
Key to
<taxonomicName id="41C14D41FFFEFFD04EDAF904FEC2B4E7" ID-CoL="55TZ" ID-ENA="240891" authority="Zetterstedt" authorityName="Zetterstedt" box="[241,350,1743,1770]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis id="B4B5EAD0FFFEFFD04EDAF904FEC2B4E7" bold="true" box="[241,350,1743,1770]" italics="true" pageId="79" pageNumber="80">Iteaphila</emphasis>
</taxonomicName>
with unbranched radial vein (R 4+5) from the Palaearctic Region
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="CEDB6549FFFEFFCF4EBCF8DEFA00B76D" lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80" type="key">
<key id="F25061F3FFFEFFCF4EBCF8DEFA00B76D" lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80">
<keyStep id="3D352E67FFFEFFD04EBCF8DEFA00B545" pageId="79" pageNumber="80">
<paragraph id="867E36C2FFFEFFD04EBCF8DEFA00B527" blockId="79.[151,1437,1812,2040]" box="[151,1436,1812,1835]" pageId="79" pageNumber="80">
<keyLead id="3D3095F7FFFEFFD04EBCF8DEFA00B527" box="[151,1436,1812,1835]" pageId="79" pageNumber="80">
<specimenCount id="90C7FD4BFFFEFFD04EBCF8DEFE8FB526" box="[151,275,1813,1835]" pageId="79" pageNumber="80" type="male">1 Male</specimenCount>
[unknown for
<taxonomicName id="41C14D41FFFEFFD04F89F8DDFE62B527" ID-CoL="5VBCP" authorityName="Becker" authorityYear="1891" box="[418,510,1813,1835]" class="Insecta" family="Ragadidae" genus="Anthepiscopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="caelebs">
<emphasis id="B4B5EAD0FFFEFFD04F89F8DDFE62B527" box="[418,510,1813,1835]" italics="true" pageId="79" pageNumber="80">I. caelebs</emphasis>
</taxonomicName>
].......................................................................... 2
</keyLead>
</paragraph>
<paragraph id="867E36C2FFFEFFD04EBCF8F8FA00B545" blockId="79.[151,1437,1812,2040]" box="[151,1436,1842,1864]" pageId="79" pageNumber="80">
<keyLead id="3D3095F7FFFEFFD04EBCF8F8FA00B545" box="[151,1436,1842,1864]" pageId="79" pageNumber="80">
- Female [unknown for
<taxonomicName id="41C14D41FFFEFFD04F93F8F8FE6BB54A" ID-CoL="9HX45" authority="Sinclair &amp; Shamshev, 2021" authorityName="Sinclair &amp; Shamshev" authorityYear="2021" box="[440,503,1842,1864]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="falki" status="sp. nov.">
<emphasis id="B4B5EAD0FFFEFFD04F93F8F8FE6BB54A" box="[440,503,1842,1864]" italics="true" pageId="79" pageNumber="80">I. falki</emphasis>
</taxonomicName>
<emphasis id="B4B5EAD0FFFEFFD04FD7F8F9FDD5B54A" bold="true" box="[508,585,1842,1863]" pageId="79" pageNumber="80">
<taxonomicNameLabel id="AF8657ABFFFEFFD04FD7F8F9FDD5B54A" box="[508,585,1842,1863]" pageId="79" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="41C14D41FFFEFFD04C7EF8F8FD63B54A" ID-CoL="9HTFH" authority="Sinclair &amp; Shamshev, 2021" authorityName="Sinclair &amp; Shamshev" authorityYear="2021" box="[597,767,1842,1864]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="kyrgyzstanensis" status="sp. nov.">
<emphasis id="B4B5EAD0FFFEFFD04C7EF8F8FD63B54A" box="[597,767,1842,1864]" italics="true" pageId="79" pageNumber="80">I. kyrgyzstanensis</emphasis>
</taxonomicName>
<emphasis id="B4B5EAD0FFFEFFD04D2EF8F9FCCDB54A" bold="true" box="[773,849,1842,1863]" pageId="79" pageNumber="80">
<taxonomicNameLabel id="AF8657ABFFFEFFD04D2EF8F9FCCDB54A" box="[773,849,1842,1863]" pageId="79" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="41C14D41FFFEFFD04D75F8F8FC6FB54A" ID-CoL="9HSFC" authority="Sinclair &amp; Shamshev, 2021" authorityName="Sinclair &amp; Shamshev" authorityYear="2021" box="[862,1011,1842,1864]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="sakhalinensis" status="sp. nov.">
<emphasis id="B4B5EAD0FFFEFFD04D75F8F8FC6FB54A" box="[862,1011,1842,1864]" italics="true" pageId="79" pageNumber="80">I. sakhalinensis</emphasis>
</taxonomicName>
<emphasis id="B4B5EAD0FFFEFFD04DD2F8F9FBD9B54A" bold="true" box="[1017,1093,1842,1863]" pageId="79" pageNumber="80">
<taxonomicNameLabel id="AF8657ABFFFEFFD04DD2F8F9FBD9B54A" box="[1017,1093,1842,1863]" pageId="79" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="41C14D41FFFEFFD04A79F8F8FB55B54A" ID-CoL="9HZ2V" authority="Sinclair &amp; Shamshev, 2021" authorityName="Sinclair &amp; Shamshev" authorityYear="2021" box="[1106,1225,1842,1864]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="aktruensis" status="sp. nov.">
<emphasis id="B4B5EAD0FFFEFFD04A79F8F8FB55B54A" box="[1106,1225,1842,1864]" italics="true" pageId="79" pageNumber="80">I. aktruensis</emphasis>
</taxonomicName>
<emphasis id="B4B5EAD0FFFEFFD04AE5F8F9FA87B54A" bold="true" box="[1230,1307,1842,1863]" pageId="79" pageNumber="80">
<taxonomicNameLabel id="AF8657ABFFFEFFD04AE5F8F9FA87B54A" box="[1230,1307,1842,1863]" pageId="79" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
]........ 16
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3D352E67FFFEFFD04EBCF89BFA00B58F" pageId="79" pageNumber="80">
<paragraph id="867E36C2FFFEFFD04EBCF89BFA00B568" blockId="79.[151,1437,1812,2040]" box="[151,1436,1872,1894]" pageId="79" pageNumber="80">
<keyLead id="3D3095F7FFFEFFD04EBCF89BFA00B568" box="[151,1436,1872,1894]" pageId="79" pageNumber="80">2 Eyes dichoptic....................................................................................... 3</keyLead>
</paragraph>
<paragraph id="867E36C2FFFEFFD04EBCF8A6FA00B58F" blockId="79.[151,1437,1812,2040]" box="[151,1436,1901,1923]" pageId="79" pageNumber="80">
<keyLead id="3D3095F7FFFEFFD04EBCF8A6FA00B58F" box="[151,1436,1901,1923]" pageId="79" pageNumber="80">- Eyes holoptic........................................................................................ 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="3D352E67FFFEFFD04EBCF841FA00B5B0" pageId="79" pageNumber="80">
<paragraph id="867E36C2FFFEFFD04EBCF841FA00B592" blockId="79.[151,1437,1812,2040]" box="[151,1436,1930,1952]" pageId="79" pageNumber="80">
<keyLead id="3D3095F7FFFEFFD04EBCF841FA00B592" box="[151,1436,1930,1952]" pageId="79" pageNumber="80">
3 Stylus not much longer than wide; terminalia almost entirely yellow (
<figureCitation id="1EFA2A47FFFEFFD04D56F841FC5EB5AD" box="[893,962,1930,1952]" captionStart="FIGURES 1114" captionStartId="13.[151,264,1463,1487]" captionTargetBox="[151,1435,199,1439]" captionTargetId="figure-17@13.[151,1435,199,1439]" captionTargetPageId="13" captionText="FIGURES 1114. Species of Iteaphila with unbranched R 4+5. 11, I. beringiensis sp. nov., male, lateral view, scale bar = 1 mm; 12, I. bartaki sp. nov., male, lateral view, scale bar = 1 mm; 13, I. dichoptica sp. nov., male holotype, lateral view, scale bar = 0.5 mm; 14, I. flavipilosa, female, dorsal view, lateral view, scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4745578" httpUri="https://zenodo.org/record/4745578/files/figure.png" pageId="79" pageNumber="80">Fig. 13</figureCitation>
)....................
<emphasis id="B4B5EAD0FFFEFFD04AE5F840FA00B592" bold="true" box="[1230,1436,1930,1952]" pageId="79" pageNumber="80">
<taxonomicName id="41C14D41FFFEFFD04AE5F840FAD6B5AD" ID-CoL="9HRZH" authority="Sinclair &amp; Shamshev, 2021" authorityName="Sinclair &amp; Shamshev" authorityYear="2021" box="[1230,1354,1930,1952]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="dichoptica" status="sp. nov.">
<emphasis id="B4B5EAD0FFFEFFD04AE5F840FAD6B5AD" bold="true" box="[1230,1354,1930,1952]" italics="true" pageId="79" pageNumber="80">I. dichoptica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657ABFFFEFFD04B7BF841FA00B592" box="[1360,1436,1930,1951]" pageId="79" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="867E36C2FFFEFFD04EBCF863FA00B5B0" blockId="79.[151,1437,1812,2040]" box="[151,1436,1959,1982]" pageId="79" pageNumber="80">
<keyLead id="3D3095F7FFFEFFD04EBCF863FA00B5B0" box="[151,1436,1959,1982]" pageId="79" pageNumber="80">
- Stylus much longer than wide; terminalia very dark.....................................
<emphasis id="B4B5EAD0FFFEFFD04AB2F863FA00B5B0" bold="true" box="[1177,1436,1959,1981]" pageId="79" pageNumber="80">
<taxonomicName id="41C14D41FFFEFFD04AB2F863FAD6B5B0" ID-CoL="9HTFH" authority="Sinclair &amp; Shamshev, 2021" authorityName="Sinclair &amp; Shamshev" authorityYear="2021" box="[1177,1354,1959,1981]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="kyrgyzstanensis" status="sp. nov.">
<emphasis id="B4B5EAD0FFFEFFD04AB2F863FAD6B5B0" bold="true" box="[1177,1354,1959,1981]" italics="true" pageId="79" pageNumber="80">I. kyrgyzstanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657ABFFFEFFD04B7BF863FA00B5B0" box="[1360,1436,1960,1981]" pageId="79" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3D352E67FFFEFFCF4EBCF80EFA00B2C6" lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80">
<paragraph id="867E36C2FFFEFFD04EBCF80EFA00B5FA" blockId="79.[151,1437,1812,2040]" pageId="79" pageNumber="80">
<keyLead id="3D3095F7FFFEFFD04EBCF80EFA00B5FA" pageId="79" pageNumber="80">
4 Epandrium oval, with broad dorsal bridge connecting lamellae and narrow, dorsally directed surstylus; hypandrium clothed with short shiny projections (
<figureCitation id="1EFA2A47FFFEFFD04FD1F829FDDCB5F5" box="[506,576,2018,2040]" captionStart="FIGURES 2125" captionStartId="19.[151,264,1787,1811]" captionTargetBox="[184,1429,195,1746]" captionTargetId="figure-17@19.[151,1436,181,1763]" captionTargetPageId="19" captionText="FIGURES 2125. Male terminalia of Iteaphila with unbranched R 4+5. 21, I. beringiensis sp. nov., lateral view; 22, I. bifida sp. nov., lateral view, with arrow indicating apex of phallus in posterior view; 23, I. recta sp. nov., lateral view; 24, I. tribulosa sp. nov., lateral view; 25, same, apex of hypandrium, posterior view. Scale bar = 0.1 mm. Abbreviations: cerc—cercus; epand— epandrium; epand brg—epandrial bridge; hypd—hypandrium; ph—phallus; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745586" httpUri="https://zenodo.org/record/4745586/files/figure.png" pageId="79" pageNumber="80">Fig. 21</figureCitation>
)....................................................
<emphasis id="B4B5EAD0FFFEFFD04A97F828FA00B5FA" bold="true" box="[1212,1436,2018,2040]" pageId="79" pageNumber="80">
<taxonomicName id="41C14D41FFFEFFD04A97F828FAD5B5F5" ID-CoL="9HVWM" authority="Sinclair &amp; Shamshev, 2021" authorityName="Sinclair &amp; Shamshev" authorityYear="2021" box="[1212,1353,2018,2040]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="beringiensis" status="sp. nov.">
<emphasis id="B4B5EAD0FFFEFFD04A97F828FAD5B5F5" bold="true" box="[1212,1353,2018,2040]" italics="true" pageId="79" pageNumber="80">I. beringiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657ABFFFEFFD04B7BF829FA00B5FA" box="[1360,1436,2018,2039]" pageId="79" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="867E36C2FFE1FFCF4EBCFF53FA00B2C6" blockId="80.[151,1437,151,1376]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFF53FA00B2C6" pageId="80" pageNumber="81">- Epandrium narrowed, longer than wide, prolonged posteriorly, with narrow dorsal bridge and posteriorly directed surstylus; hypandrium not clothed with short shiny projections......................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="3D352E67FFE1FFCF4EBCFF19FA00B308" pageId="80" pageNumber="81">
<paragraph id="867E36C2FFE1FFCF4EBCFF19FA00B2E5" blockId="80.[151,1437,151,1376]" box="[151,1436,210,232]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFF19FA00B2E5" box="[151,1436,210,232]" pageId="80" pageNumber="81">
5 Surstylus held erect to epandrium (
<figureCitation id="1EFA2A47FFE1FFCF4C1AFF19FDE0B2E5" box="[561,636,210,232]" captionStart="FIGURES 4548" captionStartId="44.[151,264,1686,1710]" captionTargetBox="[160,1433,194,1631]" captionTargetId="figure-19@44.[151,1436,181,1663]" captionTargetPageId="44" captionText="FIGURES 4548. Male terminalia of Iteaphila with unbranched R, lateral view. 45, I. gracilis sp. nov.; 46, I. grandis sp. 4+5 nov.; 47, I. lolo sp. nov.; 48, I. longipalpis (Melander). Scale bar = 0.1 mm. Abbreviations: cerc—cercus; epand brg—epandrial bridge; hyprct proc—hypoproct process; ph—phallus; ph gd—phallic guide; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745600" httpUri="https://zenodo.org/record/4745600/files/figure.png" pageId="80" pageNumber="81">Figs 45</figureCitation>
,
<figureCitation id="1EFA2A47FFE1FFCF4CADFF19FD3CB2E5" box="[646,672,210,232]" captionStart="FIGURES 5357" captionStartId="52.[151,264,1898,1922]" captionTargetBox="[160,1424,322,1839]" captionTargetId="figure-63@52.[151,1436,309,1875]" captionTargetPageId="52" captionText="FIGURES 5357. Male terminalia of Iteaphila with unbranched R 4+5. 53, I. luteitibia sp. nov., lateral view; 54, I. nupta (Melander), lateral view; 55, I. sierrensis sp. nov., lateral view; 56, I. spinosa sp. nov., lateral view; 57, same, left cercus, dorsal view. Scale bar = 0.1 mm. Abbreviations: cerc—cercus; hypd—hypandrium; hyprct proc—hypoproct process; ph—phallus; ph gd—phallic guide; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745608" httpUri="https://zenodo.org/record/4745608/files/figure.png" pageId="80" pageNumber="81">56</figureCitation>
,
<figureCitation id="1EFA2A47FFE1FFCF4C81FF19FD58B2E5" box="[682,708,210,232]" captionStart="FIGURES 5861" captionStartId="59.[151,264,1870,1894]" captionTargetBox="[176,1429,299,1808]" captionTargetId="figure-59@59.[151,1436,273,1847]" captionTargetPageId="59" captionText="FIGURES 5861. Male terminalia of Iteaphila with unbranched R, lateral view. 58, I. subnupta sp. nov.; 59, I. aktruensis 4+5 sp. nov.; 60, I. incus sp. nov.; 61, I. miranda sp. nov. Scale bar = 0.1 mm. Abbreviations: pgt—postgonite; ph—phallus; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745610" httpUri="https://zenodo.org/record/4745610/files/figure.png" pageId="80" pageNumber="81">61</figureCitation>
,
<figureCitation id="1EFA2A47FFE1FFCF4CE5FF19FD7BB2E5" box="[718,743,210,232]" captionStart="FIGURES 7578" captionStartId="74.[151,264,1749,1773]" captionTargetBox="[164,1419,315,1674]" captionTargetId="figure-62@74.[151,1436,273,1726]" captionTargetPageId="74" captionText="FIGURES 7578. Male terminalia of Iteaphila with unbranched R 4+5. 75, I. zontaki (Nowicki), lateral view; 76, same, phallus, dorsal view; 77, I. parastentor sp. nov., lateral view; 78, I. stentor (Melander), lateral view. Scale bar = 0.1 mm. Abbreviations: ph—phallus; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745626" httpUri="https://zenodo.org/record/4745626/files/figure.png" pageId="80" pageNumber="81">75</figureCitation>
)........................................................ 6
</keyLead>
</paragraph>
<paragraph id="867E36C2FFE1FFCF4EBCFF3BFA00B308" blockId="80.[151,1437,151,1376]" box="[151,1436,239,262]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFF3BFA00B308" box="[151,1436,239,262]" pageId="80" pageNumber="81">
- Surstylus directed posteriorly (
<figureCitation id="1EFA2A47FFE1FFCF4C22FF24FDEBB30B" box="[521,631,239,262]" captionStart="FIGURES 2932" captionStartId="24.[151,264,1697,1721]" captionTargetBox="[164,1432,193,1652]" captionTargetId="figure-19@24.[151,1436,181,1674]" captionTargetPageId="24" captionText="FIGURES 2932. Male terminalia of Iteaphila with unbranched R 4+5, lateral view. 29, I. bartaki sp. nov.; 30, I. falki sp. nov.; 31, I. kyrgyzstanensis sp. nov.; 32, I. ribesii (Becker). Scale bar = 0.1 mm. Abbreviations: cerc—cercus; ej apod—ejaculatory apodeme; goncx apod—gonocoxal apodeme; hypd—hypandrium; hyprct proc—hypoproct process; i apod—inner apodeme; ph—phallus; ph gd—phallic guide; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745590" httpUri="https://zenodo.org/record/4745590/files/figure.png" pageId="80" pageNumber="81">Figs 29, 30</figureCitation>
)................................................................. 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3D352E67FFE1FFCF4EBCFEC6FA00B332" pageId="80" pageNumber="81">
<paragraph id="867E36C2FFE1FFCF4EBCFEC6FA00B32F" blockId="80.[151,1437,151,1376]" box="[151,1436,269,291]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFEC6FA00B32F" box="[151,1436,269,291]" pageId="80" pageNumber="81">
6 Surstylus very narrow and shiny (
<figureCitation id="1EFA2A47FFE1FFCF4C0EFEC6FDECB32E" box="[549,624,269,291]" captionStart="FIGURES 5861" captionStartId="59.[151,264,1870,1894]" captionTargetBox="[176,1429,299,1808]" captionTargetId="figure-59@59.[151,1436,273,1847]" captionTargetPageId="59" captionText="FIGURES 5861. Male terminalia of Iteaphila with unbranched R, lateral view. 58, I. subnupta sp. nov.; 59, I. aktruensis 4+5 sp. nov.; 60, I. incus sp. nov.; 61, I. miranda sp. nov. Scale bar = 0.1 mm. Abbreviations: pgt—postgonite; ph—phallus; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745610" httpUri="https://zenodo.org/record/4745610/files/figure.png" pageId="80" pageNumber="81">Figs 61</figureCitation>
,
<figureCitation id="1EFA2A47FFE1FFCF4C51FEC6FD0FB32E" box="[634,659,269,291]" captionStart="FIGURES 7578" captionStartId="74.[151,264,1749,1773]" captionTargetBox="[164,1419,315,1674]" captionTargetId="figure-62@74.[151,1436,273,1726]" captionTargetPageId="74" captionText="FIGURES 7578. Male terminalia of Iteaphila with unbranched R 4+5. 75, I. zontaki (Nowicki), lateral view; 76, same, phallus, dorsal view; 77, I. parastentor sp. nov., lateral view; 78, I. stentor (Melander), lateral view. Scale bar = 0.1 mm. Abbreviations: ph—phallus; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745626" httpUri="https://zenodo.org/record/4745626/files/figure.png" pageId="80" pageNumber="81">75</figureCitation>
)............................................................... 7
</keyLead>
</paragraph>
<paragraph id="867E36C2FFE1FFCF4EBCFEE0FA00B332" blockId="80.[151,1437,151,1376]" box="[151,1436,298,320]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFEE0FA00B332" box="[151,1436,298,320]" pageId="80" pageNumber="81">
- Surstylus broader, round and narrow apically (
<figureCitation id="1EFA2A47FFE1FFCF4CB4FEE1FD75B34D" box="[671,745,298,320]" captionStart="FIGURES 4548" captionStartId="44.[151,264,1686,1710]" captionTargetBox="[160,1433,194,1631]" captionTargetId="figure-19@44.[151,1436,181,1663]" captionTargetPageId="44" captionText="FIGURES 4548. Male terminalia of Iteaphila with unbranched R, lateral view. 45, I. gracilis sp. nov.; 46, I. grandis sp. 4+5 nov.; 47, I. lolo sp. nov.; 48, I. longipalpis (Melander). Scale bar = 0.1 mm. Abbreviations: cerc—cercus; epand brg—epandrial bridge; hyprct proc—hypoproct process; ph—phallus; ph gd—phallic guide; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745600" httpUri="https://zenodo.org/record/4745600/files/figure.png" pageId="80" pageNumber="81">Figs 45</figureCitation>
,
<figureCitation id="1EFA2A47FFE1FFCF4CD8FEE1FC90B34D" box="[755,780,298,320]" captionStart="FIGURES 5357" captionStartId="52.[151,264,1898,1922]" captionTargetBox="[160,1424,322,1839]" captionTargetId="figure-63@52.[151,1436,309,1875]" captionTargetPageId="52" captionText="FIGURES 5357. Male terminalia of Iteaphila with unbranched R 4+5. 53, I. luteitibia sp. nov., lateral view; 54, I. nupta (Melander), lateral view; 55, I. sierrensis sp. nov., lateral view; 56, I. spinosa sp. nov., lateral view; 57, same, left cercus, dorsal view. Scale bar = 0.1 mm. Abbreviations: cerc—cercus; hypd—hypandrium; hyprct proc—hypoproct process; ph—phallus; ph gd—phallic guide; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745608" httpUri="https://zenodo.org/record/4745608/files/figure.png" pageId="80" pageNumber="81">56</figureCitation>
)..................................................... 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3D352E67FFE1FFCF4EBCFE83FA00B377" pageId="80" pageNumber="81">
<paragraph id="867E36C2FFE1FFCF4EBCFE83FA00B350" blockId="80.[151,1437,151,1376]" box="[151,1436,327,349]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFE83FA00B350" box="[151,1436,327,349]" pageId="80" pageNumber="81">
7 Surstylus hook-like, shorter than cercus (
<figureCitation id="1EFA2A47FFE1FFCF4C47FE8CFD2DB350" box="[620,689,327,349]" captionStart="FIGURES 7578" captionStartId="74.[151,264,1749,1773]" captionTargetBox="[164,1419,315,1674]" captionTargetId="figure-62@74.[151,1436,273,1726]" captionTargetPageId="74" captionText="FIGURES 7578. Male terminalia of Iteaphila with unbranched R 4+5. 75, I. zontaki (Nowicki), lateral view; 76, same, phallus, dorsal view; 77, I. parastentor sp. nov., lateral view; 78, I. stentor (Melander), lateral view. Scale bar = 0.1 mm. Abbreviations: ph—phallus; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745626" httpUri="https://zenodo.org/record/4745626/files/figure.png" pageId="80" pageNumber="81">Fig. 75</figureCitation>
)............................................
<taxonomicName id="41C14D41FFE1FFCF4AFDFE83FA00B350" authority="(Nowicki)" baseAuthorityName="Nowicki" box="[1238,1436,327,349]" class="Insecta" family="Dolichopodidae" genus="Microphorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="zontaki">
<emphasis id="B4B5EAD0FFE1FFCF4AFDFE83FAA8B350" bold="true" box="[1238,1332,327,349]" italics="true" pageId="80" pageNumber="81">I. zontaki</emphasis>
(Nowicki)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="867E36C2FFE1FFCF4EBCFEAEFA00B377" blockId="80.[151,1437,151,1376]" box="[151,1436,356,379]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFEAEFA00B377" box="[151,1436,356,379]" pageId="80" pageNumber="81">
- Surstylus elongate and straight, as long as cercus (
<figureCitation id="1EFA2A47FFE1FFCF4CEFFEAEFC96B376" box="[708,778,357,379]" captionStart="FIGURES 5861" captionStartId="59.[151,264,1870,1894]" captionTargetBox="[176,1429,299,1808]" captionTargetId="figure-59@59.[151,1436,273,1847]" captionTargetPageId="59" captionText="FIGURES 5861. Male terminalia of Iteaphila with unbranched R, lateral view. 58, I. subnupta sp. nov.; 59, I. aktruensis 4+5 sp. nov.; 60, I. incus sp. nov.; 61, I. miranda sp. nov. Scale bar = 0.1 mm. Abbreviations: pgt—postgonite; ph—phallus; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745610" httpUri="https://zenodo.org/record/4745610/files/figure.png" pageId="80" pageNumber="81">Fig. 61</figureCitation>
)......................................
<emphasis id="B4B5EAD0FFE1FFCF4ACBFEAEFA00B377" bold="true" box="[1248,1436,356,378]" pageId="80" pageNumber="81">
<taxonomicName id="41C14D41FFE1FFCF4ACBFEAEFAD5B377" ID-CoL="9HJCH" authority="Sinclair &amp; Shamshev, 2021" authorityName="Sinclair &amp; Shamshev" authorityYear="2021" box="[1248,1353,356,378]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="miranda" status="sp. nov.">
<emphasis id="B4B5EAD0FFE1FFCF4ACBFEAEFAD5B377" bold="true" box="[1248,1353,356,378]" italics="true" pageId="80" pageNumber="81">I. miranda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657ABFFE1FFCF4B7BFEAEFA00B377" box="[1360,1436,357,378]" pageId="80" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3D352E67FFE1FFCF4EBCFE49FA00B3E2" pageId="80" pageNumber="81">
<paragraph id="867E36C2FFE1FFCF4EBCFE49FA00B3B8" blockId="80.[151,1437,151,1376]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFE49FA00B3B8" pageId="80" pageNumber="81">
8 Surstylus tapered to narrow point (
<figureCitation id="1EFA2A47FFE1FFCF4C18FE49FDE0B395" box="[563,636,386,408]" captionStart="FIGURES 5357" captionStartId="52.[151,264,1898,1922]" captionTargetBox="[160,1424,322,1839]" captionTargetId="figure-63@52.[151,1436,309,1875]" captionTargetPageId="52" captionText="FIGURES 5357. Male terminalia of Iteaphila with unbranched R 4+5. 53, I. luteitibia sp. nov., lateral view; 54, I. nupta (Melander), lateral view; 55, I. sierrensis sp. nov., lateral view; 56, I. spinosa sp. nov., lateral view; 57, same, left cercus, dorsal view. Scale bar = 0.1 mm. Abbreviations: cerc—cercus; hypd—hypandrium; hyprct proc—hypoproct process; ph—phallus; ph gd—phallic guide; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745608" httpUri="https://zenodo.org/record/4745608/files/figure.png" pageId="80" pageNumber="81">Fig. 56</figureCitation>
); cercus broad with posterior, subapical lobe (
<figureCitation id="1EFA2A47FFE1FFCF4A17FE49FB19B395" box="[1084,1157,386,408]" captionStart="FIGURES 5357" captionStartId="52.[151,264,1898,1922]" captionTargetBox="[160,1424,322,1839]" captionTargetId="figure-63@52.[151,1436,309,1875]" captionTargetPageId="52" captionText="FIGURES 5357. Male terminalia of Iteaphila with unbranched R 4+5. 53, I. luteitibia sp. nov., lateral view; 54, I. nupta (Melander), lateral view; 55, I. sierrensis sp. nov., lateral view; 56, I. spinosa sp. nov., lateral view; 57, same, left cercus, dorsal view. Scale bar = 0.1 mm. Abbreviations: cerc—cercus; hypd—hypandrium; hyprct proc—hypoproct process; ph—phallus; ph gd—phallic guide; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745608" httpUri="https://zenodo.org/record/4745608/files/figure.png" pageId="80" pageNumber="81">Fig. 57</figureCitation>
); phallus slightly arched and broad at apex...........................................................................
<emphasis id="B4B5EAD0FFE1FFCF4AC2FE6BFA00B3B8" bold="true" box="[1257,1436,415,437]" pageId="80" pageNumber="81">
<taxonomicName id="41C14D41FFE1FFCF4AC2FE6BFAD6B3B9" ID-CoL="9HXKC" authority="Sinclair &amp; Shamshev, 2021" authorityName="Sinclair &amp; Shamshev" authorityYear="2021" box="[1257,1354,415,437]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">
<emphasis id="B4B5EAD0FFE1FFCF4AC2FE6BFAD6B3B9" bold="true" box="[1257,1354,415,437]" italics="true" pageId="80" pageNumber="81">I. spinosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657ABFFE1FFCF4B7BFE6BFA00B3B8" box="[1360,1436,416,437]" pageId="80" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="867E36C2FFE1FFCF4EBCFE76FA00B3E2" blockId="80.[151,1437,151,1376]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFE76FA00B3E2" pageId="80" pageNumber="81">
- Surstylus digitiform with rounded tip; cercus narrow without posterior, subapical lobe; phallus strongly arched and slender at apex (
<figureCitation id="1EFA2A47FFE1FFCF4F37FE11FEFEB3FD" box="[284,354,474,496]" captionStart="FIGURES 4548" captionStartId="44.[151,264,1686,1710]" captionTargetBox="[160,1433,194,1631]" captionTargetId="figure-19@44.[151,1436,181,1663]" captionTargetPageId="44" captionText="FIGURES 4548. Male terminalia of Iteaphila with unbranched R, lateral view. 45, I. gracilis sp. nov.; 46, I. grandis sp. 4+5 nov.; 47, I. lolo sp. nov.; 48, I. longipalpis (Melander). Scale bar = 0.1 mm. Abbreviations: cerc—cercus; epand brg—epandrial bridge; hyprct proc—hypoproct process; ph—phallus; ph gd—phallic guide; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745600" httpUri="https://zenodo.org/record/4745600/files/figure.png" pageId="80" pageNumber="81">Fig. 45</figureCitation>
)..........................................................................
<emphasis id="B4B5EAD0FFE1FFCF4AC0FE11FA00B3E2" bold="true" box="[1259,1436,474,496]" pageId="80" pageNumber="81">
<taxonomicName id="41C14D41FFE1FFCF4AC0FE11FAD5B3FD" ID-CoL="9HWD5" authority="Sinclair &amp; Shamshev, 2021" authorityName="Sinclair &amp; Shamshev" authorityYear="2021" box="[1259,1353,474,496]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="B4B5EAD0FFE1FFCF4AC0FE11FAD5B3FD" bold="true" box="[1259,1353,474,496]" italics="true" pageId="80" pageNumber="81">I. gracilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657ABFFE1FFCF4B7BFE11FA00B3E2" box="[1360,1436,474,495]" pageId="80" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3D352E67FFE1FFCF4EBCFE33FA00B027" pageId="80" pageNumber="81">
<paragraph id="867E36C2FFE1FFCF4EBCFE33FA00B000" blockId="80.[151,1437,151,1376]" box="[151,1436,503,525]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFE33FA00B000" box="[151,1436,503,525]" pageId="80" pageNumber="81">9 Anepisternum and katepisternum with shiny spot on middle.................................................. 10</keyLead>
</paragraph>
<paragraph id="867E36C2FFE1FFCF4EBCFDDEFA00B027" blockId="80.[151,1437,151,1376]" box="[151,1436,533,555]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFDDEFA00B027" box="[151,1436,533,555]" pageId="80" pageNumber="81">- Anepisternum and katepisternum greyish pollinose.......................................................... 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="3D352E67FFE1FFCF4EBCFDF9FA00B092" pageId="80" pageNumber="81">
<paragraph id="867E36C2FFE1FFCF4EBCFDF9FA00B068" blockId="80.[151,1437,151,1376]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFDF9FA00B068" pageId="80" pageNumber="81">
10 Apex of phallus narrowed, directed horizontally and slightly sinuous; hypoproct process elongate; hypandrium with subapical subtriangular projection (
<figureCitation id="1EFA2A47FFE1FFCF4FFEFD84FD87B06B" box="[469,539,591,614]" captionStart="FIGURES 2932" captionStartId="24.[151,264,1697,1721]" captionTargetBox="[164,1432,193,1652]" captionTargetId="figure-19@24.[151,1436,181,1674]" captionTargetPageId="24" captionText="FIGURES 2932. Male terminalia of Iteaphila with unbranched R 4+5, lateral view. 29, I. bartaki sp. nov.; 30, I. falki sp. nov.; 31, I. kyrgyzstanensis sp. nov.; 32, I. ribesii (Becker). Scale bar = 0.1 mm. Abbreviations: cerc—cercus; ej apod—ejaculatory apodeme; goncx apod—gonocoxal apodeme; hypd—hypandrium; hyprct proc—hypoproct process; i apod—inner apodeme; ph—phallus; ph gd—phallic guide; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745590" httpUri="https://zenodo.org/record/4745590/files/figure.png" pageId="80" pageNumber="81">Fig. 29</figureCitation>
)...........................................................
<emphasis id="B4B5EAD0FFE1FFCF4AC6FD9BFA00B068" bold="true" box="[1261,1436,591,613]" pageId="80" pageNumber="81">
<taxonomicName id="41C14D41FFE1FFCF4AC6FD9BFAD6B068" ID-CoL="9HXPD" authority="Sinclair &amp; Shamshev, 2021" authorityName="Sinclair &amp; Shamshev" authorityYear="2021" box="[1261,1354,591,613]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="bartaki" status="sp. nov.">
<emphasis id="B4B5EAD0FFE1FFCF4AC6FD9BFAD6B068" bold="true" box="[1261,1354,591,613]" italics="true" pageId="80" pageNumber="81">I. bartaki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657ABFFE1FFCF4B7BFD9BFA00B068" box="[1360,1436,592,613]" pageId="80" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="867E36C2FFE1FFCF4EBCFDA6FA00B092" blockId="80.[151,1437,151,1376]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFDA6FA00B092" pageId="80" pageNumber="81">
- Apex of phallus not narrowed, evenly curved ventrally; hypoproct process short; hypandrium evenly curved subapically (
<figureCitation id="1EFA2A47FFE1FFCF4B5EFDA6FF67B0AD" captionStart="FIGURES 2932" captionStartId="24.[151,264,1697,1721]" captionTargetBox="[164,1432,193,1652]" captionTargetId="figure-19@24.[151,1436,181,1674]" captionTargetPageId="24" captionText="FIGURES 2932. Male terminalia of Iteaphila with unbranched R 4+5, lateral view. 29, I. bartaki sp. nov.; 30, I. falki sp. nov.; 31, I. kyrgyzstanensis sp. nov.; 32, I. ribesii (Becker). Scale bar = 0.1 mm. Abbreviations: cerc—cercus; ej apod—ejaculatory apodeme; goncx apod—gonocoxal apodeme; hypd—hypandrium; hyprct proc—hypoproct process; i apod—inner apodeme; ph—phallus; ph gd—phallic guide; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745590" httpUri="https://zenodo.org/record/4745590/files/figure.png" pageId="80" pageNumber="81">Fig. 30</figureCitation>
).....................................................................................
<emphasis id="B4B5EAD0FFE1FFCF4B2CFD41FA00B092" bold="true" box="[1287,1436,650,672]" pageId="80" pageNumber="81">
<taxonomicName id="41C14D41FFE1FFCF4B2CFD41FAD5B0AD" ID-CoL="9HX45" authority="Sinclair &amp; Shamshev, 2021" authorityName="Sinclair &amp; Shamshev" authorityYear="2021" box="[1287,1353,650,672]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="falki" status="sp. nov.">
<emphasis id="B4B5EAD0FFE1FFCF4B2CFD41FAD5B0AD" bold="true" box="[1287,1353,650,672]" italics="true" pageId="80" pageNumber="81">I. falki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657ABFFE1FFCF4B7BFD41FA00B092" box="[1360,1436,650,671]" pageId="80" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3D352E67FFE1FFCF4EBCFD63FA00B0D7" pageId="80" pageNumber="81">
<paragraph id="867E36C2FFE1FFCF4EBCFD63FA00B0B0" blockId="80.[151,1437,151,1376]" box="[151,1436,679,701]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFD63FA00B0B0" box="[151,1436,679,701]" pageId="80" pageNumber="81">
11 Phallic guide long, produced beyond epandrium as far as phallus, well visible in situ (
<figureCitation id="1EFA2A47FFE1FFCF4A33FD6CFBFFB0B0" box="[1048,1123,679,701]" captionStart="FIGURES 2932" captionStartId="24.[151,264,1697,1721]" captionTargetBox="[164,1432,193,1652]" captionTargetId="figure-19@24.[151,1436,181,1674]" captionTargetPageId="24" captionText="FIGURES 2932. Male terminalia of Iteaphila with unbranched R 4+5, lateral view. 29, I. bartaki sp. nov.; 30, I. falki sp. nov.; 31, I. kyrgyzstanensis sp. nov.; 32, I. ribesii (Becker). Scale bar = 0.1 mm. Abbreviations: cerc—cercus; ej apod—ejaculatory apodeme; goncx apod—gonocoxal apodeme; hypd—hypandrium; hyprct proc—hypoproct process; i apod—inner apodeme; ph—phallus; ph gd—phallic guide; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745590" httpUri="https://zenodo.org/record/4745590/files/figure.png" pageId="80" pageNumber="81">Figs 32</figureCitation>
,
<figureCitation id="1EFA2A47FFE1FFCF4A46FD6CFB1AB0B0" box="[1133,1158,679,701]" captionStart="FIGURES 5357" captionStartId="52.[151,264,1898,1922]" captionTargetBox="[160,1424,322,1839]" captionTargetId="figure-63@52.[151,1436,309,1875]" captionTargetPageId="52" captionText="FIGURES 5357. Male terminalia of Iteaphila with unbranched R 4+5. 53, I. luteitibia sp. nov., lateral view; 54, I. nupta (Melander), lateral view; 55, I. sierrensis sp. nov., lateral view; 56, I. spinosa sp. nov., lateral view; 57, same, left cercus, dorsal view. Scale bar = 0.1 mm. Abbreviations: cerc—cercus; hypd—hypandrium; hyprct proc—hypoproct process; ph—phallus; ph gd—phallic guide; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745608" httpUri="https://zenodo.org/record/4745608/files/figure.png" pageId="80" pageNumber="81">53</figureCitation>
).................... 12
</keyLead>
</paragraph>
<paragraph id="867E36C2FFE1FFCF4EBCFD0EFA00B0D7" blockId="80.[151,1437,151,1376]" box="[151,1436,709,731]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFD0EFA00B0D7" box="[151,1436,709,731]" pageId="80" pageNumber="81">
- Phallic guide short, not produced beyond epandrium, not visible in situ (
<figureCitation id="1EFA2A47FFE1FFCF4DB3FD0EFC7FB0D6" box="[920,995,709,731]" captionStart="FIGURES 5861" captionStartId="59.[151,264,1870,1894]" captionTargetBox="[176,1429,299,1808]" captionTargetId="figure-59@59.[151,1436,273,1847]" captionTargetPageId="59" captionText="FIGURES 5861. Male terminalia of Iteaphila with unbranched R, lateral view. 58, I. subnupta sp. nov.; 59, I. aktruensis 4+5 sp. nov.; 60, I. incus sp. nov.; 61, I. miranda sp. nov. Scale bar = 0.1 mm. Abbreviations: pgt—postgonite; ph—phallus; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745610" httpUri="https://zenodo.org/record/4745610/files/figure.png" pageId="80" pageNumber="81">Figs 60</figureCitation>
,
<figureCitation id="1EFA2A47FFE1FFCF4DC6FD0EFB9AB0D6" box="[1005,1030,709,731]" captionStart="FIGURES 6670" captionStartId="66.[151,264,1859,1883]" captionTargetBox="[163,1430,249,1802]" captionTargetId="figure-45@66.[151,1436,237,1836]" captionTargetPageId="66" captionText="FIGURES 6670. Male terminalia of Iteaphila with unbranched R. 66, I. oedalina (Zetterstedt), posterior view; 67, same, 4+5 lateral view; 68, I. polygyna (Melander), lateral view; 69, I. recurvata sp. nov., lateral view; 70, I. sakhalinensis sp. nov., lateral view. Scale bar = 0.1 mm. Abbreviations: pgt—postgonite; ph—phallus; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745618" httpUri="https://zenodo.org/record/4745618/files/figure.png" pageId="80" pageNumber="81">67</figureCitation>
)............................... 13
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3D352E67FFE1FFCF4EBCFD29FA00B13E" pageId="80" pageNumber="81">
<paragraph id="867E36C2FFE1FFCF4EBCFD29FA00B118" blockId="80.[151,1437,151,1376]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFD29FA00B118" pageId="80" pageNumber="81">
12 Tibiae and usually halteres yellowish; cercus short, as long as surstylus; phallic guide subequal in width to phallus (
<figureCitation id="1EFA2A47FFE1FFCF4B1CFD29FAE1B0F5" box="[1335,1405,738,760]" captionStart="FIGURES 5357" captionStartId="52.[151,264,1898,1922]" captionTargetBox="[160,1424,322,1839]" captionTargetId="figure-63@52.[151,1436,309,1875]" captionTargetPageId="52" captionText="FIGURES 5357. Male terminalia of Iteaphila with unbranched R 4+5. 53, I. luteitibia sp. nov., lateral view; 54, I. nupta (Melander), lateral view; 55, I. sierrensis sp. nov., lateral view; 56, I. spinosa sp. nov., lateral view; 57, same, left cercus, dorsal view. Scale bar = 0.1 mm. Abbreviations: cerc—cercus; hypd—hypandrium; hyprct proc—hypoproct process; ph—phallus; ph gd—phallic guide; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745608" httpUri="https://zenodo.org/record/4745608/files/figure.png" pageId="80" pageNumber="81">Fig. 53</figureCitation>
).....................................................................................
<emphasis id="B4B5EAD0FFE1FFCF4AF7FCCBFA00B118" bold="true" box="[1244,1436,767,789]" pageId="80" pageNumber="81">
<taxonomicName id="41C14D41FFE1FFCF4AF7FCCBFAD5B118" ID-CoL="9HT9K" authority="Sinclair &amp; Shamshev, 2021" authorityName="Sinclair &amp; Shamshev" authorityYear="2021" box="[1244,1353,767,789]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="luteitibia" status="sp. nov.">
<emphasis id="B4B5EAD0FFE1FFCF4AF7FCCBFAD5B118" bold="true" box="[1244,1353,767,789]" italics="true" pageId="80" pageNumber="81">I. luteitibia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657ABFFE1FFCF4B7BFCCBFA00B118" box="[1360,1436,768,789]" pageId="80" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="867E36C2FFE1FFCF4EBCFCD6FA00B13E" blockId="80.[151,1437,151,1376]" box="[151,1436,796,819]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFCD6FA00B13E" box="[151,1436,796,819]" pageId="80" pageNumber="81">
- Tibiae and halteres dark; cercus long, longer than surstylus; phallic guide broader than phallus (
<figureCitation id="1EFA2A47FFE1FFCF4A5CFCD6FB21B13E" box="[1143,1213,797,819]" captionStart="FIGURES 2932" captionStartId="24.[151,264,1697,1721]" captionTargetBox="[164,1432,193,1652]" captionTargetId="figure-19@24.[151,1436,181,1674]" captionTargetPageId="24" captionText="FIGURES 2932. Male terminalia of Iteaphila with unbranched R 4+5, lateral view. 29, I. bartaki sp. nov.; 30, I. falki sp. nov.; 31, I. kyrgyzstanensis sp. nov.; 32, I. ribesii (Becker). Scale bar = 0.1 mm. Abbreviations: cerc—cercus; ej apod—ejaculatory apodeme; goncx apod—gonocoxal apodeme; hypd—hypandrium; hyprct proc—hypoproct process; i apod—inner apodeme; ph—phallus; ph gd—phallic guide; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745590" httpUri="https://zenodo.org/record/4745590/files/figure.png" pageId="80" pageNumber="81">Fig. 32</figureCitation>
)..
<taxonomicName id="41C14D41FFE1FFCF4ADBFCD6FA00B13E" ID-CoL="9HW8B" authority="(Becker)" authorityName="Sinclair &amp; Shamshev" authorityYear="2021" baseAuthorityName="Becker" box="[1264,1436,796,819]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="ribesii">
<emphasis id="B4B5EAD0FFE1FFCF4ADBFCD6FADEB13F" bold="true" box="[1264,1346,796,818]" italics="true" pageId="80" pageNumber="81">I. ribesii</emphasis>
(Becker)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3D352E67FFE1FFCF4EBCFCF1FA00B160" pageId="80" pageNumber="81">
<paragraph id="867E36C2FFE1FFCF4EBCFCF1FA00B15D" blockId="80.[151,1437,151,1376]" box="[151,1436,826,848]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFCF1FA00B15D" box="[151,1436,826,848]" pageId="80" pageNumber="81">
13 Phallus bent subapically nearly 90°, directed straight before arched apically (
<figureCitation id="1EFA2A47FFE1FFCF4DEFFCF1FB96B15D" box="[964,1034,826,848]" captionStart="FIGURES 6670" captionStartId="66.[151,264,1859,1883]" captionTargetBox="[163,1430,249,1802]" captionTargetId="figure-45@66.[151,1436,237,1836]" captionTargetPageId="66" captionText="FIGURES 6670. Male terminalia of Iteaphila with unbranched R. 66, I. oedalina (Zetterstedt), posterior view; 67, same, 4+5 lateral view; 68, I. polygyna (Melander), lateral view; 69, I. recurvata sp. nov., lateral view; 70, I. sakhalinensis sp. nov., lateral view. Scale bar = 0.1 mm. Abbreviations: pgt—postgonite; ph—phallus; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745618" httpUri="https://zenodo.org/record/4745618/files/figure.png" pageId="80" pageNumber="81">Fig. 67</figureCitation>
)............
<taxonomicName id="41C14D41FFE1FFCF4A9EFCF1FA00B15D" authority="(Zetterstedt)" baseAuthorityName="Zetterstedt" box="[1205,1436,826,848]" class="Insecta" family="Empididae" genus="Rhamphomyza" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="oedalina">
<emphasis id="B4B5EAD0FFE1FFCF4A9EFCF1FA83B15D" bold="true" box="[1205,1311,826,848]" italics="true" pageId="80" pageNumber="81">I. oedalina</emphasis>
(Zetterstedt)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="867E36C2FFE1FFCF4EBCFC93FA00B160" blockId="80.[151,1437,151,1376]" box="[151,1436,855,878]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFC93FA00B160" box="[151,1436,855,878]" pageId="80" pageNumber="81">
- Phallus not bent subapically nearly 90°, gradually arched and curved apically (
<figureCitation id="1EFA2A47FFE1FFCF4DFEFC9CFBDFB160" box="[981,1091,855,877]" captionStart="FIGURES 5861" captionStartId="59.[151,264,1870,1894]" captionTargetBox="[176,1429,299,1808]" captionTargetId="figure-59@59.[151,1436,273,1847]" captionTargetPageId="59" captionText="FIGURES 5861. Male terminalia of Iteaphila with unbranched R, lateral view. 58, I. subnupta sp. nov.; 59, I. aktruensis 4+5 sp. nov.; 60, I. incus sp. nov.; 61, I. miranda sp. nov. Scale bar = 0.1 mm. Abbreviations: pgt—postgonite; ph—phallus; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745610" httpUri="https://zenodo.org/record/4745610/files/figure.png" pageId="80" pageNumber="81">Figs 59, 60</figureCitation>
,
<figureCitation id="1EFA2A47FFE1FFCF4A66FC93FBFAB163" box="[1101,1126,856,878]" captionStart="FIGURES 6670" captionStartId="66.[151,264,1859,1883]" captionTargetBox="[163,1430,249,1802]" captionTargetId="figure-45@66.[151,1436,237,1836]" captionTargetPageId="66" captionText="FIGURES 6670. Male terminalia of Iteaphila with unbranched R. 66, I. oedalina (Zetterstedt), posterior view; 67, same, 4+5 lateral view; 68, I. polygyna (Melander), lateral view; 69, I. recurvata sp. nov., lateral view; 70, I. sakhalinensis sp. nov., lateral view. Scale bar = 0.1 mm. Abbreviations: pgt—postgonite; ph—phallus; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745618" httpUri="https://zenodo.org/record/4745618/files/figure.png" pageId="80" pageNumber="81">70</figureCitation>
)....................... 14
</keyLead>
</paragraph>
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<keyStep id="3D352E67FFE1FFCF4EBCFCBEFA00B1A5" pageId="80" pageNumber="81">
<paragraph id="867E36C2FFE1FFCF4EBCFCBEFA00B187" blockId="80.[151,1437,151,1376]" box="[151,1436,885,907]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFCBEFA00B187" box="[151,1436,885,907]" pageId="80" pageNumber="81">
14 Surstylus broad with rounded apex; phallus often projected well beyond epandrium (
<figureCitation id="1EFA2A47FFE1FFCF4A24FCBEFBC9B186" box="[1039,1109,885,907]" captionStart="FIGURES 5861" captionStartId="59.[151,264,1870,1894]" captionTargetBox="[176,1429,299,1808]" captionTargetId="figure-59@59.[151,1436,273,1847]" captionTargetPageId="59" captionText="FIGURES 5861. Male terminalia of Iteaphila with unbranched R, lateral view. 58, I. subnupta sp. nov.; 59, I. aktruensis 4+5 sp. nov.; 60, I. incus sp. nov.; 61, I. miranda sp. nov. Scale bar = 0.1 mm. Abbreviations: pgt—postgonite; ph—phallus; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745610" httpUri="https://zenodo.org/record/4745610/files/figure.png" pageId="80" pageNumber="81">Fig. 60</figureCitation>
)............
<emphasis id="B4B5EAD0FFE1FFCF4AD4FCBEFA00B187" bold="true" box="[1279,1436,885,906]" pageId="80" pageNumber="81">
<taxonomicName id="41C14D41FFE1FFCF4AD4FCBEFAD5B187" ID-CoL="9HVLP" authority="Sinclair &amp; Shamshev, 2021" authorityName="Sinclair &amp; Shamshev" authorityYear="2021" box="[1279,1353,885,906]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="incus" status="sp. nov.">
<emphasis id="B4B5EAD0FFE1FFCF4AD4FCBEFAD5B187" bold="true" box="[1279,1353,885,906]" italics="true" pageId="80" pageNumber="81">I. incus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657ABFFE1FFCF4B7BFCBEFA00B187" box="[1360,1436,885,906]" pageId="80" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="867E36C2FFE1FFCF4EBCFC58FA00B1A5" blockId="80.[151,1437,151,1376]" box="[151,1436,914,936]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFC58FA00B1A5" box="[151,1436,914,936]" pageId="80" pageNumber="81">
- Surstylus narrow, hook-like; phallus arched along margin of epandrium (
<figureCitation id="1EFA2A47FFE1FFCF4DB7FC59FC7AB1A5" box="[924,998,914,936]" captionStart="FIGURES 5861" captionStartId="59.[151,264,1870,1894]" captionTargetBox="[176,1429,299,1808]" captionTargetId="figure-59@59.[151,1436,273,1847]" captionTargetPageId="59" captionText="FIGURES 5861. Male terminalia of Iteaphila with unbranched R, lateral view. 58, I. subnupta sp. nov.; 59, I. aktruensis 4+5 sp. nov.; 60, I. incus sp. nov.; 61, I. miranda sp. nov. Scale bar = 0.1 mm. Abbreviations: pgt—postgonite; ph—phallus; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745610" httpUri="https://zenodo.org/record/4745610/files/figure.png" pageId="80" pageNumber="81">Figs 59</figureCitation>
,
<figureCitation id="1EFA2A47FFE1FFCF4DDBFC59FB95B1A5" box="[1008,1033,914,936]" captionStart="FIGURES 6670" captionStartId="66.[151,264,1859,1883]" captionTargetBox="[163,1430,249,1802]" captionTargetId="figure-45@66.[151,1436,237,1836]" captionTargetPageId="66" captionText="FIGURES 6670. Male terminalia of Iteaphila with unbranched R. 66, I. oedalina (Zetterstedt), posterior view; 67, same, 4+5 lateral view; 68, I. polygyna (Melander), lateral view; 69, I. recurvata sp. nov., lateral view; 70, I. sakhalinensis sp. nov., lateral view. Scale bar = 0.1 mm. Abbreviations: pgt—postgonite; ph—phallus; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745618" httpUri="https://zenodo.org/record/4745618/files/figure.png" pageId="80" pageNumber="81">70</figureCitation>
).............................. 15
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3D352E67FFE1FFCF4EBCFC64FA00B1EF" pageId="80" pageNumber="81">
<paragraph id="867E36C2FFE1FFCF4EBCFC64FA00B1C8" blockId="80.[151,1437,151,1376]" box="[151,1436,943,965]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFC64FA00B1C8" box="[151,1436,943,965]" pageId="80" pageNumber="81">
15 Phallus prolonged, arched to hypandrium (
<figureCitation id="1EFA2A47FFE1FFCF4C57FC64FD5DB1C8" box="[636,705,943,965]" captionStart="FIGURES 5861" captionStartId="59.[151,264,1870,1894]" captionTargetBox="[176,1429,299,1808]" captionTargetId="figure-59@59.[151,1436,273,1847]" captionTargetPageId="59" captionText="FIGURES 5861. Male terminalia of Iteaphila with unbranched R, lateral view. 58, I. subnupta sp. nov.; 59, I. aktruensis 4+5 sp. nov.; 60, I. incus sp. nov.; 61, I. miranda sp. nov. Scale bar = 0.1 mm. Abbreviations: pgt—postgonite; ph—phallus; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745610" httpUri="https://zenodo.org/record/4745610/files/figure.png" pageId="80" pageNumber="81">Fig. 59</figureCitation>
)..........................................
<emphasis id="B4B5EAD0FFE1FFCF4AE6FC7BFA00B1C8" bold="true" box="[1229,1436,943,965]" pageId="80" pageNumber="81">
<taxonomicName id="41C14D41FFE1FFCF4AE6FC7BFAD6B1C8" ID-CoL="9HZ2V" authority="Sinclair &amp; Shamshev, 2021" authorityName="Sinclair &amp; Shamshev" authorityYear="2021" box="[1229,1354,943,965]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="aktruensis" status="sp. nov.">
<emphasis id="B4B5EAD0FFE1FFCF4AE6FC7BFAD6B1C8" bold="true" box="[1229,1354,943,965]" italics="true" pageId="80" pageNumber="81">I. aktruensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657ABFFE1FFCF4B7BFC7BFA00B1C8" box="[1360,1436,944,965]" pageId="80" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="867E36C2FFE1FFCF4EBCFC06FA00B1EF" blockId="80.[151,1437,151,1376]" box="[151,1436,972,995]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFC06FA00B1EF" box="[151,1436,972,995]" pageId="80" pageNumber="81">
- Phallus not prolonged, arched to apex of epandrium (
<figureCitation id="1EFA2A47FFE1FFCF4CCBFC06FCBAB1EE" box="[736,806,973,995]" captionStart="FIGURES 6670" captionStartId="66.[151,264,1859,1883]" captionTargetBox="[163,1430,249,1802]" captionTargetId="figure-45@66.[151,1436,237,1836]" captionTargetPageId="66" captionText="FIGURES 6670. Male terminalia of Iteaphila with unbranched R. 66, I. oedalina (Zetterstedt), posterior view; 67, same, 4+5 lateral view; 68, I. polygyna (Melander), lateral view; 69, I. recurvata sp. nov., lateral view; 70, I. sakhalinensis sp. nov., lateral view. Scale bar = 0.1 mm. Abbreviations: pgt—postgonite; ph—phallus; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745618" httpUri="https://zenodo.org/record/4745618/files/figure.png" pageId="80" pageNumber="81">Fig. 70</figureCitation>
)...............................
<emphasis id="B4B5EAD0FFE1FFCF4A86FC06FA00B1EF" bold="true" box="[1197,1436,972,994]" pageId="80" pageNumber="81">
<taxonomicName id="41C14D41FFE1FFCF4A86FC06FAD6B1EF" ID-CoL="9HSFC" authority="Sinclair &amp; Shamshev, 2021" authorityName="Sinclair &amp; Shamshev" authorityYear="2021" box="[1197,1354,972,994]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="sakhalinensis" status="sp. nov.">
<emphasis id="B4B5EAD0FFE1FFCF4A86FC06FAD6B1EF" bold="true" box="[1197,1354,972,994]" italics="true" pageId="80" pageNumber="81">I. sakhalinensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657ABFFE1FFCF4B7BFC06FA00B1EF" box="[1360,1436,973,994]" pageId="80" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3D352E67FFE1FFCF4EBCFC21FA00B610" pageId="80" pageNumber="81">
<paragraph id="867E36C2FFE1FFCF4EBCFC21FA00B1F2" blockId="80.[151,1437,151,1376]" box="[151,1436,1002,1024]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFC21FA00B1F2" box="[151,1436,1002,1024]" pageId="80" pageNumber="81">16 Halteres pale........................................................................................ 17</keyLead>
</paragraph>
<paragraph id="867E36C2FFE1FFCF4EBCFBC3FA00B610" blockId="80.[151,1437,151,1376]" box="[151,1436,1031,1053]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFBC3FA00B610" box="[151,1436,1031,1053]" pageId="80" pageNumber="81">- Halteres dark........................................................................................ 19</keyLead>
</paragraph>
</keyStep>
<keyStep id="3D352E67FFE1FFCF4EBCFBEEFA00B65A" pageId="80" pageNumber="81">
<paragraph id="867E36C2FFE1FFCF4EBCFBEEFA00B637" blockId="80.[151,1437,151,1376]" box="[151,1436,1060,1083]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFBEEFA00B637" box="[151,1436,1060,1083]" pageId="80" pageNumber="81">
17 Antennal stylus (excluding apical sensillum) distinctly longer than wide...........
<emphasis id="B4B5EAD0FFE1FFCF4A36FBEEFB4CB637" bold="true" box="[1053,1232,1061,1082]" pageId="80" pageNumber="81">
<taxonomicName id="41C14D41FFE1FFCF4A36FBEEFBE2B637" ID-CoL="9HXKC" authority="Sinclair &amp; Shamshev, 2021" authorityName="Sinclair &amp; Shamshev" authorityYear="2021" box="[1053,1150,1061,1082]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">
<emphasis id="B4B5EAD0FFE1FFCF4A36FBEEFBE2B637" bold="true" box="[1053,1150,1061,1082]" italics="true" pageId="80" pageNumber="81">I. spinosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657ABFFE1FFCF4AA8FBEEFB4CB637" box="[1155,1232,1061,1082]" pageId="80" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="B4B5EAD0FFE1FFCF4AF7FBEEFA00B637" bold="true" box="[1244,1436,1060,1082]" pageId="80" pageNumber="81">
<taxonomicName id="41C14D41FFE1FFCF4AF7FBEEFAD5B637" ID-CoL="9HT9K" authority="Sinclair &amp; Shamshev, 2021" authorityName="Sinclair &amp; Shamshev" authorityYear="2021" box="[1244,1353,1060,1082]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="luteitibia" status="sp. nov.">
<emphasis id="B4B5EAD0FFE1FFCF4AF7FBEEFAD5B637" bold="true" box="[1244,1353,1060,1082]" italics="true" pageId="80" pageNumber="81">I. luteitibia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657ABFFE1FFCF4B7BFBEEFA00B637" box="[1360,1436,1061,1082]" pageId="80" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="867E36C2FFE1FFCF4EBCFB88FA00B65A" blockId="80.[151,1437,151,1376]" box="[151,1436,1090,1112]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFB88FA00B65A" box="[151,1436,1090,1112]" pageId="80" pageNumber="81">- Antennal stylus (excluding apical sensillum) about as long as wide............................................. 18</keyLead>
</paragraph>
</keyStep>
<keyStep id="3D352E67FFE1FFCF4EBCFBABFA00B69F" pageId="80" pageNumber="81">
<paragraph id="867E36C2FFE1FFCF4EBCFBABFA00B678" blockId="80.[151,1437,151,1376]" box="[151,1436,1119,1141]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFBABFA00B678" box="[151,1436,1119,1141]" pageId="80" pageNumber="81">
18 Scutum viewed anteriorly subshiny, finely pollinose.........................................
<emphasis id="B4B5EAD0FFE1FFCF4AE5FBABFA00B678" bold="true" box="[1230,1436,1119,1141]" pageId="80" pageNumber="81">
<taxonomicName id="41C14D41FFE1FFCF4AE5FBABFAD6B678" ID-CoL="9HRZH" authority="Sinclair &amp; Shamshev, 2021" authorityName="Sinclair &amp; Shamshev" authorityYear="2021" box="[1230,1354,1119,1141]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="dichoptica" status="sp. nov.">
<emphasis id="B4B5EAD0FFE1FFCF4AE5FBABFAD6B678" bold="true" box="[1230,1354,1119,1141]" italics="true" pageId="80" pageNumber="81">I. dichoptica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657ABFFE1FFCF4B7BFBABFA00B678" box="[1360,1436,1120,1141]" pageId="80" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="867E36C2FFE1FFCF4EBCFBB6FA00B69F" blockId="80.[151,1437,151,1376]" box="[151,1436,1148,1171]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFBB6FA00B69F" box="[151,1436,1148,1171]" pageId="80" pageNumber="81">
- Scutum viewed anteriorly densely pollinose......
<emphasis id="B4B5EAD0FFE1FFCF4CCBFBB6FC0EB69F" bold="true" box="[736,914,1148,1170]" pageId="80" pageNumber="81">
<taxonomicName id="41C14D41FFE1FFCF4CCBFBB6FCA3B69F" ID-CoL="9HWD5" authority="Sinclair &amp; Shamshev, 2021" authorityName="Sinclair &amp; Shamshev" authorityYear="2021" box="[736,831,1148,1170]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="B4B5EAD0FFE1FFCF4CCBFBB6FCA3B69F" bold="true" box="[736,831,1148,1170]" italics="true" pageId="80" pageNumber="81">I. gracilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657ABFFE1FFCF4D6EFBB6FC0EB69F" box="[837,914,1149,1170]" pageId="80" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="41C14D41FFE1FFCF4DB5FBB6FBC4B69E" ID-CoL="5VBCP" authority="(Becker)" baseAuthorityName="Becker" box="[926,1112,1148,1171]" class="Insecta" family="Ragadidae" genus="Anthepiscopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="caelebs">
<emphasis id="B4B5EAD0FFE1FFCF4DB5FBB6FC67B69F" bold="true" box="[926,1019,1148,1170]" pageId="80" pageNumber="81">
<emphasis id="B4B5EAD0FFE1FFCF4DB5FBB6FC3BB69F" bold="true" box="[926,935,1149,1170]" italics="true" pageId="80" pageNumber="81">I</emphasis>
.
<emphasis id="B4B5EAD0FFE1FFCF4D98FBB7FC67B69F" bold="true" box="[947,1019,1148,1170]" italics="true" pageId="80" pageNumber="81">caelebs</emphasis>
</emphasis>
(Becker)
</taxonomicName>
,
<taxonomicName id="41C14D41FFE1FFCF4A4AFBB6FAD4B69E" authority="(Zetterstedt)" baseAuthorityName="Zetterstedt" box="[1121,1352,1148,1171]" class="Insecta" family="Empididae" genus="Rhamphomyza" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="oedalina">
<emphasis id="B4B5EAD0FFE1FFCF4A4AFBB6FB57B69F" bold="true" box="[1121,1227,1148,1170]" italics="true" pageId="80" pageNumber="81">I. oedalina</emphasis>
(Zetterstedt)
</taxonomicName>
[in part]
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3D352E67FFE1FFCF4EBCFB51FA00B6C0" pageId="80" pageNumber="81">
<paragraph id="867E36C2FFE1FFCF4EBCFB51FA00B6A2" blockId="80.[151,1437,151,1376]" box="[151,1436,1178,1200]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFB51FA00B6A2" box="[151,1436,1178,1200]" pageId="80" pageNumber="81">19 Antennal stylus (excluding apical sensillum) about as long as wide............................................. 20</keyLead>
</paragraph>
<paragraph id="867E36C2FFE1FFCF4EBCFB73FA00B6C0" blockId="80.[151,1437,151,1376]" box="[151,1436,1207,1229]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFB73FA00B6C0" box="[151,1436,1207,1229]" pageId="80" pageNumber="81">- Antennal stylus (excluding apical sensillum) distinctly longer than wide......................................... 21</keyLead>
</paragraph>
</keyStep>
<keyStep id="3D352E67FFE1FFCF4EBCFB1EFA00B728" pageId="80" pageNumber="81">
<paragraph id="867E36C2FFE1FFCF4EBCFB1EFA00B6E7" blockId="80.[151,1437,151,1376]" box="[151,1436,1236,1259]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFB1EFA00B6E7" box="[151,1436,1236,1259]" pageId="80" pageNumber="81">
20 Anepisternum and katepisternum shiny......................................................
<emphasis id="B4B5EAD0FFE1FFCF4AC6FB1EFA00B6E7" bold="true" box="[1261,1436,1236,1258]" pageId="80" pageNumber="81">
<taxonomicName id="41C14D41FFE1FFCF4AC6FB1EFAD6B6E7" ID-CoL="9HXPD" authority="Sinclair &amp; Shamshev, 2021" authorityName="Sinclair &amp; Shamshev" authorityYear="2021" box="[1261,1354,1236,1258]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="bartaki" status="sp. nov.">
<emphasis id="B4B5EAD0FFE1FFCF4AC6FB1EFAD6B6E7" bold="true" box="[1261,1354,1236,1258]" italics="true" pageId="80" pageNumber="81">I. bartaki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657ABFFE1FFCF4B7BFB1EFA00B6E7" box="[1360,1436,1237,1258]" pageId="80" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="867E36C2FFE1FFCF4EBCFB38FA00B728" blockId="80.[151,1437,151,1376]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFB38FA00B728" pageId="80" pageNumber="81">
- Anepisternum and katepisternum dull with pruinescence.................................................................................................
<taxonomicName id="41C14D41FFE1FFCF4CCDFADBFC32B728" authority="(Nowicki)" baseAuthorityName="Nowicki" box="[742,942,1295,1317]" class="Insecta" family="Dolichopodidae" genus="Microphorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="zontaki">
<emphasis id="B4B5EAD0FFE1FFCF4CCDFADBFCDFB728" bold="true" box="[742,835,1295,1317]" italics="true" pageId="80" pageNumber="81">I. zontaki</emphasis>
(Nowicki)
</taxonomicName>
,
<taxonomicName id="41C14D41FFE1FFCF4D93FADBFB03B728" authority="(Zetterstedt)" baseAuthorityName="Zetterstedt" box="[952,1183,1295,1317]" class="Insecta" family="Empididae" genus="Rhamphomyza" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="oedalina">
<emphasis id="B4B5EAD0FFE1FFCF4D93FADBFBBEB728" bold="true" box="[952,1058,1295,1317]" italics="true" pageId="80" pageNumber="81">I. oedalina</emphasis>
(Zetterstedt)
</taxonomicName>
[in part],
<emphasis id="B4B5EAD0FFE1FFCF4AD4FADBFA00B728" bold="true" box="[1279,1436,1295,1317]" pageId="80" pageNumber="81">
<taxonomicName id="41C14D41FFE1FFCF4AD4FADBFAD5B729" ID-CoL="9HVLP" authority="Sinclair &amp; Shamshev, 2021" authorityName="Sinclair &amp; Shamshev" authorityYear="2021" box="[1279,1353,1295,1317]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="incus" status="sp. nov.">
<emphasis id="B4B5EAD0FFE1FFCF4AD4FADBFAD5B729" bold="true" box="[1279,1353,1295,1317]" italics="true" pageId="80" pageNumber="81">I. incus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657ABFFE1FFCF4B7BFADBFA00B728" box="[1360,1436,1296,1317]" pageId="80" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3D352E67FFE1FFCF4EBCFAE6FA00B76D" pageId="80" pageNumber="81">
<paragraph id="867E36C2FFE1FFCF4EBCFAE6FA00B74F" blockId="80.[151,1437,151,1376]" box="[151,1436,1324,1347]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFAE6FA00B74F" box="[151,1436,1324,1347]" pageId="80" pageNumber="81">
21 Sternite 8 shiny.....................................................................
<emphasis id="B4B5EAD0FFE1FFCF4A97FAE6FA00B74F" bold="true" box="[1212,1436,1324,1346]" pageId="80" pageNumber="81">
<taxonomicName id="41C14D41FFE1FFCF4A97FAE6FAD5B74F" ID-CoL="9HVWM" authority="Sinclair &amp; Shamshev, 2021" authorityName="Sinclair &amp; Shamshev" authorityYear="2021" box="[1212,1353,1324,1346]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="beringiensis" status="sp. nov.">
<emphasis id="B4B5EAD0FFE1FFCF4A97FAE6FAD5B74F" bold="true" box="[1212,1353,1324,1346]" italics="true" pageId="80" pageNumber="81">I. beringiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AF8657ABFFE1FFCF4B7BFAE6FA00B74F" box="[1360,1436,1325,1346]" pageId="80" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="867E36C2FFE1FFCF4EBCFA80FA00B76D" blockId="80.[151,1437,151,1376]" box="[151,1436,1354,1376]" pageId="80" pageNumber="81">
<keyLead id="3D3095F7FFE1FFCF4EBCFA80FA00B76D" box="[151,1436,1354,1376]" pageId="80" pageNumber="81">
- Sternite 8 with pruinescence...............................................................
<taxonomicName id="41C14D41FFE1FFCF4ADBFA81FA00B76D" ID-CoL="9HW8B" authority="(Becker)" authorityName="Sinclair &amp; Shamshev" authorityYear="2021" baseAuthorityName="Becker" box="[1264,1436,1354,1376]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="ribesii">
<emphasis id="B4B5EAD0FFE1FFCF4ADBFA81FADEB76D" bold="true" box="[1264,1346,1354,1376]" italics="true" pageId="80" pageNumber="81">I. ribesii</emphasis>
(Becker)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</subSubSection>
</treatment>
</document>