257 lines
33 KiB
XML
257 lines
33 KiB
XML
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<mods:title id="BA4EA6C902A4CA2787EA10686F91684A">World revision of Iteaphila with unbranched radial vein (Diptera: Empidoidea: Iteaphilidae)</mods:title>
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<mods:name id="18A8675D3FAF7A4E995CFFE20588CD0F" type="personal">
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<mods:roleTerm id="45E64BAD3C0FAC6DB2D12D653FCFD2AC">Author</mods:roleTerm>
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<mods:namePart id="9D10106B4EE1220DEB6ED9167C9B6B07">Sinclair, Bradley J.</mods:namePart>
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<mods:nameIdentifier id="4BC4E1DB1A83837D82E0AFB19388530A" type="email">bradley.sinclair@canada.ca</mods:nameIdentifier>
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<mods:namePart id="D036919BB268C66DF758C6E49E65B49E">Shamshev, Igor V.</mods:namePart>
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<mods:nameIdentifier id="A5DE53629406F4068A14FDA767FECF42" type="email">bradley.sinclair@canada.ca</mods:nameIdentifier>
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<mods:title id="1CA9F52F25BD84ADCFCD03806F75FDD5">Zootaxa</mods:title>
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<mods:part id="EE3D51D695D2E062CE16F34739B8B781">
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<mods:date id="432D6DDF97AB9544755FBE1285CD808D">2021</mods:date>
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<mods:number id="D828B0511F0041F38514CE5B7AD13531">2021-05-07</mods:number>
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<mods:number id="6F733B854B6A30DAF6D20292D8C75C3C">4968</mods:number>
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<mods:number id="9932E11CC436A0EE2407434491AB5B4E">1</mods:number>
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<treatment id="0E6887D4FF89FFA64EBCFE81FE05B014" ID-DOI="http://doi.org/10.5281/zenodo.4814488" ID-GBIF-Taxon="182399095" ID-Zenodo-Dep="4814488" LSID="urn:lsid:zoobank.org:act:8DFD999B-AF58-4040-AA67-C0CE4038FF0A" httpUri="http://treatment.plazi.org/id/0E6887D4FF89FFA64EBCFE81FE05B014" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57">
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<subSubSection id="CEDB6549FF89FFA74EBCFE81FE50B368" box="[151,460,330,357]" pageId="56" pageNumber="57" type="nomenclature">
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<paragraph id="867E36C2FF89FFA74EBCFE81FE50B368" blockId="56.[151,985,330,429]" box="[151,460,330,357]" pageId="56" pageNumber="57">
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<heading id="DD3681AEFF89FFA74EBCFE81FE50B368" bold="true" box="[151,460,330,357]" fontSize="11" level="2" pageId="56" pageNumber="57" reason="4">
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<emphasis id="B4B5EAD0FF89FFA74EBCFE81FE50B368" bold="true" box="[151,460,330,357]" pageId="56" pageNumber="57">
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<taxonomicName id="41C14D41FF89FFA74EBCFE81FEFBB368" ID-CoL="9HXKC" authority="Sinclair & Shamshev, 2021" authorityName="Sinclair & Shamshev" authorityYear="2021" box="[151,359,330,357]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">
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<emphasis id="B4B5EAD0FF89FFA74EBCFE81FEFBB368" bold="true" box="[151,359,330,357]" italics="true" pageId="56" pageNumber="57">Iteaphila spinosa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="AF8657ABFF89FFA74F45FE80FE50B368" box="[366,460,331,357]" pageId="56" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="CEDB6549FF89FFA74EBCFEA4FC45B3A0" pageId="56" pageNumber="57" type="description">
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<paragraph id="867E36C2FF89FFA74EBCFEA4FECDB384" blockId="56.[151,985,330,429]" box="[151,337,366,393]" pageId="56" pageNumber="57">
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(
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<figureCitation id="1EFA2A47FF89FFA74EB4FEA4FEBCB384" box="[159,288,366,393]" captionStart="FIGURES 53–57" captionStartId="52.[151,264,1898,1922]" captionTargetBox="[160,1424,322,1839]" captionTargetId="figure-63@52.[151,1436,309,1875]" captionTargetPageId="52" captionText="FIGURES 53–57. Male terminalia of Iteaphila with unbranched R 4+5. 53, I. luteitibia sp. nov., lateral view; 54, I. nupta (Melander), lateral view; 55, I. sierrensis sp. nov., lateral view; 56, I. spinosa sp. nov., lateral view; 57, same, left cercus, dorsal view. Scale bar = 0.1 mm. Abbreviations: cerc—cercus; hypd—hypandrium; hyprct proc—hypoproct process; ph—phallus; ph gd—phallic guide; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745608" httpUri="https://zenodo.org/record/4745608/files/figure.png" pageId="56" pageNumber="57">Figs 56, 57</figureCitation>
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,
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<figureCitation id="1EFA2A47FF89FFA74F07FEA4FED5B384" box="[300,329,367,393]" captionStart="FIGURES 79–81" captionStartId="77.[151,264,1589,1613]" captionTargetBox="[151,1436,181,1565]" captionTargetId="figure-17@77.[151,1436,181,1565]" captionTargetPageId="77" captionText="FIGURES 79–81. Distribution of Iteaphila with unbranched R 4+5. 79, I. longiphallus sp. nov., I. recurvata sp. nov.; 80, I. falcata Sinclair, I. parastentor sp. nov., I. stentor (Melander); 81, I. atruensis sp. nov., I. gracilis sp. nov., I. kyrgyzstanensis sp. nov., I. sakhalinensis sp. nov., I. spinosa sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4745630" httpUri="https://zenodo.org/record/4745630/files/figure.png" pageId="56" pageNumber="57">81</figureCitation>
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)
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</paragraph>
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<uri id="F2503AC0FF89FFA74EBCFE58FC45B3A0" box="[151,985,402,429]" pageId="56" pageNumber="57">
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urn:lsid:zoobank.org:act:
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</subSubSection>
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<subSubSection id="CEDB6549FF89FFA74EBCFE10FCD2B0FC" pageId="56" pageNumber="57" type="materials_examined">
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<paragraph id="867E36C2FF89FFA74EBCFE10FB27B088" blockId="56.[151,1437,474,2013]" pageId="56" pageNumber="57">
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<emphasis id="B4B5EAD0FF89FFA74EBCFE10FED9B3F9" bold="true" box="[151,325,474,501]" pageId="56" pageNumber="57">Type material.</emphasis>
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<typeStatus id="597A8860FF89FFA74F67FE11FE77B3F9" box="[332,491,474,500]" pageId="56" pageNumber="57" type="holotype">HOLOTYPE</typeStatus>
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</emphasis>
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<specimenCount id="90C7FD4BFF89FFA74FD9FE10FD95B3F8" box="[498,521,475,501]" pageId="56" pageNumber="57" type="male">♂</specimenCount>
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, labelled: “(
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)/
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[
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]/
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/
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. Saigusa”; “
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/
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/ spinosa/ Sinclair & Shamshev [red label]” (
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<collectionCode id="E0D0AE07FF89FFA74A30FE34FBEEB014" box="[1051,1138,511,537]" country="Thailand" httpUri="http://grbio.org/cool/m1dy-7gjd" name="Kasetsart University Museum of Fisheries" pageId="56" pageNumber="57">KUMF</collectionCode>
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)
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</materialsCitation>
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.
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:
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.
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:
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</emphasis>
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[
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],
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<location id="831E6019FF89FFA74C97FDE8FCDDB030" LSID="urn:lsid:plazi:treatment:0E6887D4FF89FFA64EBCFE81FE05B014:831E6019FF89FFA74C97FDE8FCDDB030" box="[700,833,547,573]" country="Japan" latitude="43.7" longLatPrecision="1142" longitude="142.83333" municipality="Nagayamadake" name="Taisetsuzan" pageId="56" pageNumber="57" stateProvince="Hokkaido">Taisetsuzan</location>
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,
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<date id="F27F1002FF89FFA74D67FDE8FC52B030" box="[844,974,547,573]" pageId="56" pageNumber="57" value="1967-07-10">
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<collectingDate id="E23BE9EAFF89FFA74D67FDE8FC52B030" box="[844,974,547,573]" pageId="56" pageNumber="57" value="1967-07-10">10.vii.1967</collectingDate>
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</date>
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, TS (
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<specimenCount id="90C7FD4BFF89FFA74A2CFDE8FBA8B030" box="[1031,1076,547,573]" pageId="56" pageNumber="57" type="male">4 ♂</specimenCount>
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,
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<collectionCode id="E0D0AE07FF89FFA74A14FDE8FB09B030" box="[1087,1173,547,573]" country="Thailand" httpUri="http://grbio.org/cool/m1dy-7gjd" name="Kasetsart University Museum of Fisheries" pageId="56" pageNumber="57">KUMF</collectionCode>
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)
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</materialsCitation>
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;
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<materialsCitation id="36A93C9FFF89FFA74A83FDE8FEA8B06C" ID-GBIF-Occurrence="3118988301" collectingDate="1967-06-13" collectionCode="KUMF" country="Japan" elevation="675" latitude="43.35" location="Nukabira" longLatPrecision="1144" longitude="143.23334" pageId="56" pageNumber="57" specimenCount="4" specimenCount-female="4" stateProvince="Hokkaido" typeStatus="paratype">
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same data as holotype (
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,
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<collectionCode id="E0D0AE07FF89FFA74EFDFD8CFEB0B06C" box="[214,300,583,609]" country="Thailand" httpUri="http://grbio.org/cool/m1dy-7gjd" name="Kasetsart University Museum of Fisheries" pageId="56" pageNumber="57">KUMF</collectionCode>
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)
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</materialsCitation>
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;
|
||
<materialsCitation id="36A93C9FFF89FFA74F17FD8CFBF2B06C" ID-GBIF-Occurrence="3118988726" box="[316,1134,583,611]" collectingDate="1986-07-16" collectionCode="KUMF" country="Japan" latitude="43.716667" location="Taisetsuzan" longLatPrecision="1142" longitude="142.8" municipality="Aizankei" pageId="56" pageNumber="57" specimenCount="1" specimenCount-male="1" stateProvince="Hokkaido" typeStatus="paratype">
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<collectingMunicipality id="661AACB8FF89FFA74F17FD8CFE3DB06C" box="[316,417,583,609]" pageId="56" pageNumber="57">Aizankei</collectingMunicipality>
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[
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|
||
<geoCoordinate id="E3F55005FF89FFA74C38FD83FD1CB06E" box="[531,640,584,611]" degrees="142" direction="east" minutes="48" orientation="longitude" pageId="56" pageNumber="57" precision="925" value="142.8">142°48′E</geoCoordinate>
|
||
],
|
||
<location id="831E6019FF89FFA74CBAFD8CFC8AB06C" LSID="urn:lsid:plazi:treatment:0E6887D4FF89FFA64EBCFE81FE05B014:831E6019FF89FFA74CBAFD8CFC8AB06C" box="[657,790,583,609]" country="Japan" latitude="43.716667" longLatPrecision="1142" longitude="142.8" municipality="Aizankei" name="Taisetsuzan" pageId="56" pageNumber="57" stateProvince="Hokkaido">Taisetsuzan</location>
|
||
,
|
||
<date id="F27F1002FF89FFA74D0BFD8CFC3DB06C" box="[800,929,583,609]" pageId="56" pageNumber="57" value="1986-07-16">
|
||
<collectingDate id="E23BE9EAFF89FFA74D0BFD8CFC3DB06C" box="[800,929,583,609]" pageId="56" pageNumber="57" value="1986-07-16">16.vii.1986</collectingDate>
|
||
</date>
|
||
, TS (
|
||
<specimenCount id="90C7FD4BFF89FFA74DF6FD8CFB99B06C" box="[989,1029,583,609]" pageId="56" pageNumber="57" type="male">1 ♂</specimenCount>
|
||
,
|
||
<collectionCode id="E0D0AE07FF89FFA74A3BFD8CFBFAB06C" box="[1040,1126,583,609]" country="Thailand" httpUri="http://grbio.org/cool/m1dy-7gjd" name="Kasetsart University Museum of Fisheries" pageId="56" pageNumber="57">KUMF</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="36A93C9FFF89FFA74A53FD8CFEC2B088" ID-GBIF-Occurrence="3118988873" collectingDate="1986-07-17" collectionCode="KUMF" country="Japan" latitude="43.716667" location="Taisetsuzan" longLatPrecision="1142" longitude="142.8" municipality="Aizankei" pageId="56" pageNumber="57" specimenCount="1" specimenCount-female="1" stateProvince="Hokkaido" typeStatus="paratype">
|
||
same locality,
|
||
<date id="F27F1002FF89FFA74B3DFD8CFA04B06C" box="[1302,1432,583,609]" pageId="56" pageNumber="57" value="1986-07-17">
|
||
<collectingDate id="E23BE9EAFF89FFA74B3DFD8CFA04B06C" box="[1302,1432,583,609]" pageId="56" pageNumber="57" value="1986-07-17">17.vii.1986</collectingDate>
|
||
</date>
|
||
, TS (
|
||
<specimenCount id="90C7FD4BFF89FFA74EE1FDA0FF6EB088" box="[202,242,619,645]" pageId="56" pageNumber="57" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectionCode id="E0D0AE07FF89FFA74F2BFDA0FECAB088" box="[256,342,619,645]" country="Thailand" httpUri="http://grbio.org/cool/m1dy-7gjd" name="Kasetsart University Museum of Fisheries" pageId="56" pageNumber="57">KUMF</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="36A93C9FFF89FFA74F42FDA0FB2BB088" ID-GBIF-Occurrence="3118988871" box="[361,1207,619,647]" collectionCode="KUMF" country="Japan" latitude="43.35" location="Nukabira" longLatPrecision="1144" longitude="143.23334" municipality="Otofukegawa" pageId="56" pageNumber="57" specimenCount="1" specimenCount-male="1" stateProvince="Hokkaido" typeStatus="paratype">
|
||
<collectingMunicipality id="661AACB8FF89FFA74F42FDA0FD9FB088" box="[361,515,619,645]" pageId="56" pageNumber="57">Otofukegawa</collectingMunicipality>
|
||
,
|
||
<location id="831E6019FF89FFA74C25FDA0FDEAB088" LSID="urn:lsid:plazi:treatment:0E6887D4FF89FFA64EBCFE81FE05B014:831E6019FF89FFA74C25FDA0FDEAB088" box="[526,630,619,645]" country="Japan" latitude="43.35" longLatPrecision="1144" longitude="143.23334" municipality="Otofukegawa" name="Nukabira" pageId="56" pageNumber="57" stateProvince="Hokkaido">Nukabira</location>
|
||
[
|
||
<geoCoordinate id="E3F55005FF89FFA74CAFFDA7FD79B08A" box="[644,741,620,647]" degrees="43" direction="north" minutes="21" orientation="latitude" pageId="56" pageNumber="57" precision="925" value="43.35">43°21′N</geoCoordinate>
|
||
<geoCoordinate id="E3F55005FF89FFA74CC0FDA7FCC5B08A" box="[747,857,620,647]" degrees="143" direction="east" minutes="14" orientation="longitude" pageId="56" pageNumber="57" precision="925" value="143.23334">143°14′E</geoCoordinate>
|
||
],
|
||
<date id="F27F1002FF89FFA74D40FDA0FC79B088" box="[875,997,619,645]" pageId="56" pageNumber="57" value="1067-06-14">14.vi.1067</date>
|
||
, TS (
|
||
<specimenCount id="90C7FD4BFF89FFA74A08FDA0FBD1B088" box="[1059,1101,619,645]" pageId="56" pageNumber="57" type="male">1 ♂</specimenCount>
|
||
,
|
||
<collectionCode id="E0D0AE07FF89FFA74A72FDA0FB33B088" box="[1113,1199,619,645]" country="Thailand" httpUri="http://grbio.org/cool/m1dy-7gjd" name="Kasetsart University Museum of Fisheries" pageId="56" pageNumber="57">KUMF</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="867E36C2FF89FFA74EECFD45FCD2B0FC" blockId="56.[151,1437,474,2013]" pageId="56" pageNumber="57">
|
||
<emphasis id="B4B5EAD0FF89FFA74EECFD45FEC1B0A5" bold="true" box="[199,349,654,680]" pageId="56" pageNumber="57">Recognition.</emphasis>
|
||
This species is distinguished from other Japanese species by the holoptic males, pale brown tibiae and pale abdominal setae. Males have short cerci and hypoproct processes, apex of phallus hooked, shaft of phallus with spines and surstyli tapered and project dorsally (
|
||
<figureCitation id="1EFA2A47FF89FFA74CC6FD1CFCDDB0FC" box="[749,833,726,753]" captionStart="FIGURES 53–57" captionStartId="52.[151,264,1898,1922]" captionTargetBox="[160,1424,322,1839]" captionTargetId="figure-63@52.[151,1436,309,1875]" captionTargetPageId="52" captionText="FIGURES 53–57. Male terminalia of Iteaphila with unbranched R 4+5. 53, I. luteitibia sp. nov., lateral view; 54, I. nupta (Melander), lateral view; 55, I. sierrensis sp. nov., lateral view; 56, I. spinosa sp. nov., lateral view; 57, same, left cercus, dorsal view. Scale bar = 0.1 mm. Abbreviations: cerc—cercus; hypd—hypandrium; hyprct proc—hypoproct process; ph—phallus; ph gd—phallic guide; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745608" httpUri="https://zenodo.org/record/4745608/files/figure.png" pageId="56" pageNumber="57">Fig. 56</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CEDB6549FF89FFA64EECFD31FB71B3A0" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57" type="description">
|
||
<paragraph id="867E36C2FF89FFA74EECFD31FC40B638" blockId="56.[151,1437,474,2013]" pageId="56" pageNumber="57">
|
||
<emphasis id="B4B5EAD0FF89FFA74EECFD31FEC5B119" bold="true" box="[199,345,762,788]" pageId="56" pageNumber="57">Description.</emphasis>
|
||
Wing length
|
||
<quantity id="41399B27FF89FFA74FDEFD30FD1DB118" box="[501,641,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.2" metricValueMin="3.1" pageId="56" pageNumber="57" unit="mm" value="3.15" valueMax="3.2" valueMin="3.1">3.1–3.2 mm</quantity>
|
||
.
|
||
<emphasis id="B4B5EAD0FF89FFA74CA6FD30FD4DB118" bold="true" box="[653,721,763,789]" pageId="56" pageNumber="57">Male.</emphasis>
|
||
Head black, mostly with black setation, occiput finely greyish pollinose. Eyes holoptic, with upper ommatidia slightly enlarged. Frons represented by small triangular space just above antennae, bare, greyish pollinose. Ocellar triangle prominent, with 2 pairs of thin setae; anterior ocellars moderately long, posterior ocellars short. Postvertical and postocular setae similar to anterior ocellars; additionally, occiput covered with numerous brownish hairs in lower part.Antenna brown (sometimes scape and pedicel somewhat paler); scape short, subequal to globular pedicel, both with short setulae; postpedicel broad at base, nearly 2.5 times longer than wide, strongly tapered; stylus short, sensillum-tipped; segment 9 tubular, about twice as long as wide, apical sensillum about half length of segment 9. Proboscis long, projected obliquely; labium about length of head height; palpus projected parallel to labrum, somewhat shorter than labrum.
|
||
</paragraph>
|
||
<paragraph id="867E36C2FF89FFA74EECFBF4FA13B73C" blockId="56.[151,1437,474,2013]" pageId="56" pageNumber="57">Thorax black, mostly with black setation; scutum viewed dorsally subshiny, finely brownish pollinose, viewed anteriorly more distinctly pollinose; mesopleuron uniformly denser greyish pollinose. Proepisternum with several short yellowish hair-like setae. Postpronotal lobe with 1 long and several short setae (sometimes intermixed pale and black). Mesonotal setae prominent; acr moderately long, arranged in 2–3 irregular rows anteriorly, becoming biserial posteriorly, lacking on prescutellar depression, distance between rows shorter than length of setae; dc uniserial, offset from row anteriorly, mostly subequal to acr, 2–3 prescutellar pairs longer; 1 ph, 1–2 presut spal, 2–3 long npl (sometimes with additional shorter setae), 1 psut spal (sometimes with additional setulae), 1 pal, 4–5 pairs of sctl.</paragraph>
|
||
<paragraph id="867E36C2FF89FFA74EECFAF0FED1B420" blockId="56.[151,1437,474,2013]" pageId="56" pageNumber="57">Legs largely tawny yellow; coxae darkened (except apical part), tarsomere 2 apically and tarsomeres 3–5 entirely brownish (in darker specimens tarsi almost entirely brownish, except basal part of tarsomere 1). Coxae and trochanters with ordinary hair-like setae. Fore femur with row of thin posteroventral setae longer subapically. Mid femur with similar pattern of setation but subapical setae longer (somewhat longer than femur is deep) and stronger. Hind femur with long thin anteroventral and dorsal setae, shorter basally. Tibiae lacking prominent setae. Tarsomere 1 of mid and hindlegs with stiff ventral setae; tarsomere 5 on all legs somewhat flattened; pulvilli broad, shorter than tarsal claw.</paragraph>
|
||
<paragraph id="867E36C2FF89FFA74EECF9FCFD33B4EC" blockId="56.[151,1437,474,2013]" pageId="56" pageNumber="57">
|
||
Wing finely infuscate; basal costal seta absent; pterostigma distinct brownish, elliptical, overlapping apex of R
|
||
<subScript id="1A453487FF89FFA74E81F9A6FF2EB476" attach="left" box="[170,178,1645,1659]" fontSize="6" pageId="56" pageNumber="57">1</subScript>
|
||
; anal lobe very prominent, acute. Sc complete; R
|
||
<subScript id="1A453487FF89FFA74CFEF9A6FD72B476" attach="left" box="[725,750,1645,1659]" fontSize="6" pageId="56" pageNumber="57">2+3</subScript>
|
||
straight in apical part; R
|
||
<subScript id="1A453487FF89FFA74DD4F9A6FB85B476" attach="left" box="[1023,1049,1645,1659]" fontSize="6" pageId="56" pageNumber="57">4+5</subScript>
|
||
unbranched; cell dm broad, longer than basal cells, hardly produced apically, almost truncate; base of M
|
||
<subScript id="1A453487FF89FFA74DBAF95AFC05B492" attach="left" box="[913,921,1681,1695]" fontSize="6" pageId="56" pageNumber="57">2</subScript>
|
||
(crossvein) nearly one-fourth length of dm-m, M branches widely separated; dm-m crossvein somewhat concave. Apex of cell cua recurved, CuA+CuP long, faint, ending just short of wing margin. Halter brown.
|
||
</paragraph>
|
||
<paragraph id="867E36C2FF89FFA64EECF920FB48B310" blockId="56.[151,1437,474,2013]" lastBlockId="57.[151,1437,151,537]" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57">
|
||
Abdomen pale brown, subshiny, finely greyish pollinose, covered with long yellowish hair-like setae. Terminalia (
|
||
<figureCitation id="1EFA2A47FF89FFA74E8BF8C4FF72B524" box="[160,238,1806,1833]" captionStart="FIGURES 53–57" captionStartId="52.[151,264,1898,1922]" captionTargetBox="[160,1424,322,1839]" captionTargetId="figure-63@52.[151,1436,309,1875]" captionTargetPageId="52" captionText="FIGURES 53–57. Male terminalia of Iteaphila with unbranched R 4+5. 53, I. luteitibia sp. nov., lateral view; 54, I. nupta (Melander), lateral view; 55, I. sierrensis sp. nov., lateral view; 56, I. spinosa sp. nov., lateral view; 57, same, left cercus, dorsal view. Scale bar = 0.1 mm. Abbreviations: cerc—cercus; hypd—hypandrium; hyprct proc—hypoproct process; ph—phallus; ph gd—phallic guide; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745608" httpUri="https://zenodo.org/record/4745608/files/figure.png" pageId="56" pageNumber="57">Fig. 56</figureCitation>
|
||
) concolorous with abdomen, small. Hypandrium with several short setae, rounded, slightly upcurved, tapered to narrow apex in posterior view; gonocoxal apodeme long and broad; inner apodeme expanded and flattened, half length of gonocoxal apodeme; postgonite long, narrow, not extending beyond epandrium, apex divided into pair of rounded lobes. Phallic guide not extended beyond epandrium, parallel with phallus; apical section short, strongly curved. Epandrium somewhat inflated laterally; dorsal bridge moderately narrow medially, produced distally into shiny, strongly tapered, dorsally projecting surstylus. Phallus slender, arched beyond epandrium, apex arched, parallel-sided, base of shaft with small spiny projections; ejaculatory apodeme plate-like, rounded, slightly longer than length of gonocoxal apodeme. Cercus short, one-third length of epandrium; apex narrowly rounded, extended free from epandrium, anteromedial margin with flattened, rounded plate-like expansion (
|
||
<figureCitation id="1EFA2A47FF88FFA64A9DFF14FA8CB2F4" box="[1206,1296,222,249]" captionStart="FIGURES 53–57" captionStartId="52.[151,264,1898,1922]" captionTargetBox="[160,1424,322,1839]" captionTargetId="figure-63@52.[151,1436,309,1875]" captionTargetPageId="52" captionText="FIGURES 53–57. Male terminalia of Iteaphila with unbranched R 4+5. 53, I. luteitibia sp. nov., lateral view; 54, I. nupta (Melander), lateral view; 55, I. sierrensis sp. nov., lateral view; 56, I. spinosa sp. nov., lateral view; 57, same, left cercus, dorsal view. Scale bar = 0.1 mm. Abbreviations: cerc—cercus; hypd—hypandrium; hyprct proc—hypoproct process; ph—phallus; ph gd—phallic guide; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745608" httpUri="https://zenodo.org/record/4745608/files/figure.png" pageId="57" pageNumber="58">Fig. 57</figureCitation>
|
||
); hypoproct produced into pair of long, slender processes, appressed to cercus, projecting beyond epandrium.
|
||
</paragraph>
|
||
<paragraph id="867E36C2FF88FFA64EECFEECFB71B3A0" blockId="57.[151,1437,151,537]" pageId="57" pageNumber="58">
|
||
<emphasis id="B4B5EAD0FF88FFA64EECFEECFEB9B34C" bold="true" box="[199,293,295,321]" pageId="57" pageNumber="58">Female.</emphasis>
|
||
Similar to male, except as follows: Eyes dichoptic, ommatidia equally small. Frons very broad, somewhat widened toward ocellar triangle, shiny, with marginal setulae. Legs mostly yellow, progressively darker apically; fore and mid femora without prominent posteroventral setae. Halter yellow. Abdomen largely yellowish brown, with shorter setation, tergites 6 and 7 yellowish posteriorly. Cercus brownish, long, slender.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CEDB6549FF88FFA64EECFE7DFAD3B3DC" box="[199,1359,438,465]" pageId="57" pageNumber="58" type="distribution">
|
||
<paragraph id="867E36C2FF88FFA64EECFE7DFAD3B3DC" blockId="57.[151,1437,151,537]" box="[199,1359,438,465]" pageId="57" pageNumber="58">
|
||
<emphasis id="B4B5EAD0FF88FFA64EECFE7DFEFDB3DD" bold="true" box="[199,353,438,464]" pageId="57" pageNumber="58">Distribution.</emphasis>
|
||
Palaearctic:
|
||
<collectingCountry id="FED67652FF88FFA64FDAFE7CFDA8B3DC" box="[497,564,439,465]" name="Japan" pageId="57" pageNumber="58">Japan</collectingCountry>
|
||
. This species is restricted to the Japanese island of
|
||
<collectingRegion id="4405F820FF88FFA64A5BFE7CFB42B3DC" box="[1136,1246,439,465]" country="Japan" name="Hokkaido" pageId="57" pageNumber="58">Hokkaido</collectingRegion>
|
||
(
|
||
<figureCitation id="1EFA2A47FF88FFA64AC5FE7CFADEB3DC" box="[1262,1346,439,465]" captionStart="FIGURES 79–81" captionStartId="77.[151,264,1589,1613]" captionTargetBox="[151,1436,181,1565]" captionTargetId="figure-17@77.[151,1436,181,1565]" captionTargetPageId="77" captionText="FIGURES 79–81. Distribution of Iteaphila with unbranched R 4+5. 79, I. longiphallus sp. nov., I. recurvata sp. nov.; 80, I. falcata Sinclair, I. parastentor sp. nov., I. stentor (Melander); 81, I. atruensis sp. nov., I. gracilis sp. nov., I. kyrgyzstanensis sp. nov., I. sakhalinensis sp. nov., I. spinosa sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4745630" httpUri="https://zenodo.org/record/4745630/files/figure.png" pageId="57" pageNumber="58">Fig. 81</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CEDB6549FF88FFA64EECFE10FE05B014" pageId="57" pageNumber="58" type="etymology">
|
||
<paragraph id="867E36C2FF88FFA64EECFE10FE05B014" blockId="57.[151,1437,151,537]" pageId="57" pageNumber="58">
|
||
<emphasis id="B4B5EAD0FF88FFA64EECFE10FED2B3F8" bold="true" box="[199,334,475,501]" pageId="57" pageNumber="58">Etymology.</emphasis>
|
||
The specific name is from the Latin
|
||
<emphasis id="B4B5EAD0FF88FFA64CC3FE10FCD4B3F9" box="[744,840,475,500]" italics="true" pageId="57" pageNumber="58">spinosus</emphasis>
|
||
(thorny), in reference to the spine-like projections on the base of the phallus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |