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<document id="20B1E2095001C93145F064A5E783F9EB" ID-DOI="10.11646/zootaxa.5447.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="11150249" ID-ZooBank="367106D8-6248-49DA-8ABE-3125FC46BE08" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1715181318044" checkinUser="plazi" docAuthor="Chen, Jia-Heng, Yin, Wen-Qi &amp; Shi, Hong-Liang" docDate="2024" docId="03FD87AC65409665FF1FF9A22E9556A2" docLanguage="en" docName="zootaxa.5447.4.1.pdf" docOrigin="Zootaxa 5447 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5360.3.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pterostichus (Wraseiellus) pseudandrewesi Chen &amp; Yin &amp; Shi 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="456" masterDocId="FFC4FFD465439660FF88FFCD2D25501F" masterDocTitle="On the Pterostichus subgenus Wraseiellus from China: descriptions of five new species and supplementary notes on taxonomy (Coleoptera: Carabidae)" masterLastPageNumber="472" masterPageNumber="451" pageNumber="454" updateTime="1715181786403" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="81CC1D4A9B066A43E6B4B5CE5AA62823" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="4B5779374B2F4178815C8DD7689103D1">On the Pterostichus subgenus Wraseiellus from China: descriptions of five new species and supplementary notes on taxonomy (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="AF3F08C3E159B1C9F43769BDE20959E7">Chen, Jia-Heng</mods:namePart>
<mods:affiliation id="74418601DB1AC01AD94E40A6909B01B0">College of Forestry, Beijing Forestry University, Beijing, 100083, P. R. China</mods:affiliation>
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<mods:namePart id="A291FC8EABF36DB55B926647C29790B7">Yin, Wen-Qi</mods:namePart>
<mods:affiliation id="E157645312F36E78229184E84F1AD840">College of Forestry, Beijing Forestry University, Beijing, 100083, P. R. China</mods:affiliation>
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<mods:namePart id="58E9B64E1F53CEDA7B75B867DDA02736">Shi, Hong-Liang</mods:namePart>
<mods:affiliation id="C056C9442C7D26EA648947A4D1162F83">College of Forestry, Beijing Forestry University, Beijing, 100083, P. R. China</mods:affiliation>
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<mods:title id="2FC6A117E04B13E4656C4FDDB53DA7F9">Zootaxa</mods:title>
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<mods:part id="B2CAC5AC86F5336D6F4649E65EBE16BC">
<mods:date id="0211CE0DF8E5E631BF51723A9ABDAC37">2024</mods:date>
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<mods:number id="DAAB26B00E39F14E22834AD7B21D50C1">2024-05-08</mods:number>
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<mods:classification id="79E8E58ACC94768AF1A65D821491B392">journal article</mods:classification>
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<mods:identifier id="47AABFC9785583E14708CDC609533220" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03FD87AC65409665FF1FF9A22E9556A2" ID-DOI="http://doi.org/10.5281/zenodo.11150261" ID-Zenodo-Dep="11150261" LSID="urn:lsid:plazi:treatment:03FD87AC65409665FF1FF9A22E9556A2" httpUri="http://treatment.plazi.org/id/03FD87AC65409665FF1FF9A22E9556A2" lastPageId="5" lastPageNumber="456" pageId="3" pageNumber="454">
<subSubSection id="C34E653165409663FF1FF9A22F4C56B3" pageId="3" pageNumber="454" type="nomenclature">
<paragraph id="8BEB36BA65409663FF1FF9A22E255695" blockId="3.[151,768,1647,1746]" box="[151,768,1647,1674]" pageId="3" pageNumber="454">
<heading id="D0A381D665409663FF1FF9A22E255695" bold="true" box="[151,768,1647,1674]" fontSize="11" level="1" pageId="3" pageNumber="454" reason="1">
<emphasis id="B920EAA865409663FF1FF9A22E255695" bold="true" box="[151,768,1647,1674]" pageId="3" pageNumber="454">
<taxonomicName id="4C544D3965409663FF1FF9A22FBE5695" authority="Chen &amp; Yin &amp; Shi, 2024" authorityName="Chen &amp; Yin &amp; Shi" authorityYear="2024" box="[151,667,1647,1674]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="pseudandrewesi" status="sp. nov." subGenus="Wraseiellus">
<emphasis id="B920EAA865409663FF1FF9A22C085695" bold="true" box="[151,301,1647,1674]" italics="true" pageId="3" pageNumber="454">Pterostichus</emphasis>
(
<emphasis id="B920EAA865409663FEB6F9A22CEF5695" bold="true" box="[318,458,1647,1674]" italics="true" pageId="3" pageNumber="454">Wraseiellus</emphasis>
)
<emphasis id="B920EAA865409663FE53F9A22FBE5695" bold="true" box="[475,667,1647,1674]" italics="true" pageId="3" pageNumber="454">pseudandrewesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357D365409663FD2BF9BD2E255695" box="[675,768,1648,1674]" pageId="3" pageNumber="454" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEB36BA65409663FF1FF9592F4C56B3" blockId="3.[151,768,1647,1746]" box="[151,617,1684,1710]" pageId="3" pageNumber="454">
Chinese vernacular name:
<emphasis id="B920EAA865409663FE49F9582F4C56B3" bold="true" box="[449,617,1685,1708]" pageId="3" pageNumber="454">fDzaeữfflm</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C34E653165409663FF1FF9752C4B56CD" box="[151,366,1720,1746]" pageId="3" pageNumber="454" type="description">
<paragraph id="8BEB36BA65409663FF1FF9752C4B56CD" blockId="3.[151,768,1647,1746]" box="[151,366,1720,1746]" pageId="3" pageNumber="454">
<figureCitation id="136F2A3F65409663FF1FF9752C2556CD" box="[151,256,1720,1746]" captionStart="FIGURE 1" captionStartId="4.[151,250,1820,1844]" captionTargetBox="[191,1407,479,1778]" captionTargetId="figure-128@4.[151,1436,450,1794]" captionTargetPageId="4" captionText="FIGURE 1. Holotype of Pterostichus pseudandrewesi sp. nov. (A) habitus; (B) median lobe of aedeagus, left lateral view; (C) median lobe of aedeagus, dorsal view; (D) right paramere. Scale bars = 5.0 mm for A, = 1.0 mm for B–D." figureDoi="http://doi.org/10.5281/zenodo.11150253" httpUri="https://zenodo.org/record/11150253/files/figure.png" pageId="3" pageNumber="454">Figures 1</figureCitation>
,
<figureCitation id="136F2A3F65409663FE86F9752C1656CD" box="[270,307,1720,1746]" captionStart="FIGURE 9" captionStartId="15.[151,250,1411,1435]" captionTargetBox="[151,1436,201,1388]" captionTargetId="figure-17@15.[151,1436,201,1388]" captionTargetPageId="15" captionText="FIGURE 9. Pronotum posterior angle and basal area of Pterostichus (Wraseiellus) spp. (A) P. pseudandrewesi sp. nov.; (B) P. leishanensis sp. nov.; (C) P. strumatus sp. nov.; (D) P. idiopterus sp. nov.; (E) P. hemichlorus sp. nov.." figureDoi="http://doi.org/10.5281/zenodo.11150285" httpUri="https://zenodo.org/record/11150285/files/figure.png" pageId="3" pageNumber="454">9A</figureCitation>
,
<figureCitation id="136F2A3F65409663FEB6F9752C4B56CD" box="[318,366,1720,1746]" captionStart="FIGURE 10" captionStartId="16.[151,250,1743,1767]" captionTargetBox="[153,1432,692,1720]" captionTargetId="figure-228@16.[151,1436,669,1720]" captionTargetPageId="16" captionText="FIGURE 10. Apical portion of aedeagus of Pterostichus (Wraseiellus) spp., left lateral view and ventral view. (A) P. pseudandrewesi sp. nov.; (B) P. leishanensis sp. nov.; (C) P. strumatus sp. nov.; (D) P. andrewesi Jedlička." figureDoi="http://doi.org/10.5281/zenodo.11150289" httpUri="https://zenodo.org/record/11150289/files/figure.png" pageId="3" pageNumber="454">10A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34E653165409663FF1FF8CD29DD57B4" pageId="3" pageNumber="454" type="materials_examined">
<paragraph id="8BEB36BA65409663FF1FF8CD29DD57B4" blockId="3.[151,1437,1792,2035]" pageId="3" pageNumber="454">
<emphasis id="B920EAA865409663FF1FF8CD2C6D5705" bold="true" box="[151,328,1792,1818]" pageId="3" pageNumber="454">Type materials</emphasis>
.
<materialsCitation id="3B3C3CE765409663FEDDF8CD2C2A5799" collectingDate="2015-07-25" collectionCode="IZAS" country="China" county="Butuo county" determinerName="Chen" elevation="3218" latitude="27.4661" location="Pukui Mt." longLatPrecision="6" longitude="102.7741" municipality="Juesa" pageId="3" pageNumber="454" specimenCount="2" stateProvince="Sichuan" typeStatus="holotype">
<typeStatus id="54EF881865409663FEDDF8CD2C985705" box="[341,445,1792,1818]" pageId="3" pageNumber="454" type="holotype">Holotype</typeStatus>
(
<collectionCode id="ED45AE7F65409663FE43F8CD2F355705" box="[459,528,1792,1818]" country="0" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="3" pageNumber="454">IZAS</collectionCode>
): male, body length =
<quantity id="4CAC9B5F65409663FC8CF8CD2E4C5705" box="[772,873,1792,1819]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="3" pageNumber="454" unit="mm" value="15.1">15.1 mm</quantity>
, “
<collectingCountry id="F343762A65409663FCF6F8CD2EF85705" box="[894,989,1792,1818]" name="China" pageId="3" pageNumber="454">CHINA</collectingCountry>
:
<collectingRegion id="4990F85865409663FC6EF8CD295A5705" box="[998,1151,1792,1818]" country="China" name="Sichuan" pageId="3" pageNumber="454">Sichuan prov.</collectingRegion>
<collectingCounty id="628A4E3665409663FB0DF8CD28395705" box="[1157,1308,1792,1818]" pageId="3" pageNumber="454">Butuo county</collectingCounty>
,
<collectingMunicipality id="6B8FACC065409663FAAEF8CD28465705" box="[1318,1379,1792,1818]" pageId="3" pageNumber="454">Juesa</collectingMunicipality>
vill.,
<location id="8E8B606165409663FF1FF8E92C2C5721" LSID="urn:lsid:plazi:treatment:03FD87AC65409665FF1FF9A22E9556A2:8E8B606165409663FF1FF8E92C2C5721" box="[151,265,1828,1854]" country="China" county="Butuo county" latitude="27.4661" longLatPrecision="6" longitude="102.7741" municipality="Juesa" name="Pukui Mt." pageId="3" pageNumber="454" stateProvince="Sichuan">Pukui Mt.</location>
, alpine meadow,
<quantity id="4CAC9B5F65409663FE42F8E92F3F5721" box="[458,538,1828,1854]" metricMagnitude="3" metricUnit="m" metricValue="3.218" pageId="3" pageNumber="454" unit="m" value="3218.0">
<elevation id="0079D18965409663FE42F8E92F3F5721" box="[458,538,1828,1854]" metricMagnitude="3" metricUnit="m" metricValue="3.218" pageId="3" pageNumber="454" unit="m" value="3218.0">3218m</elevation>
</quantity>
,
<geoCoordinate id="EE60507D65409663FDADF8E92FB15721" box="[549,660,1828,1854]" degrees="27.4661" direction="north" orientation="latitude" pageId="3" pageNumber="454" precision="5" value="27.4661">N27.4661</geoCoordinate>
<geoCoordinate id="EE60507D65409663FD12F8E92E305721" box="[666,789,1828,1854]" degrees="102.7741" direction="east" orientation="longitude" pageId="3" pageNumber="454" precision="5" value="102.7741">E102.7741</geoCoordinate>
”// “
<date id="FFEA107A65409663FCCAF8E92EEB5720" box="[834,974,1828,1855]" pageId="3" pageNumber="454" value="2015-07-25">
<collectingDate id="EFAEE99265409663FCCAF8E92EEB5720" box="[834,974,1828,1855]" pageId="3" pageNumber="454" value="2015-07-25">2015.VII.25</collectingDate>
</date>
D, under stone, Shi HL., Liu B &amp; Ma YL, lgt. BJFU exp, 2015”// “
<typeStatus id="54EF881865409663FE2FF8852F1B577D" box="[423,574,1864,1890]" pageId="3" pageNumber="454" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C544D3965409663FDCDF8852902577D" authority="Chen &amp; Yin &amp; Shi, 2024" authorityName="Chen &amp; Yin &amp; Shi" authorityYear="2024" box="[581,1063,1864,1890]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="pseudandrewesi" status="sp. nov." subGenus="Wraseiellus">
<emphasis id="B920EAA865409663FDCDF8852FF5577D" box="[581,720,1864,1890]" italics="true" pageId="3" pageNumber="454">Pterostichus</emphasis>
(
<emphasis id="B920EAA865409663FD69F8852E41577D" box="[737,868,1864,1890]" italics="true" pageId="3" pageNumber="454">Wraseiellus</emphasis>
)
<emphasis id="B920EAA865409663FCFCF8852902577D" box="[884,1063,1864,1890]" italics="true" pageId="3" pageNumber="454">pseudandrewesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357D365409663FBA7F88529AC577D" box="[1071,1161,1864,1890]" pageId="3" pageNumber="454" rank="species">sp. nov.</taxonomicNameLabel>
, det.
<determinerName id="7E37DA7E65409663FB4DF8852825577D" box="[1221,1280,1864,1890]" pageId="3" pageNumber="454">Chen</determinerName>
&amp; Shi. 2024” [red label]
</materialsCitation>
;
<materialsCitation id="3B3C3CE765409663FE92F8A129D157B4" collectingDate="2020-05-26" collectionCode="IZAS" country="China" county="Shimian County" elevation="1988" latitude="19.028383" location="Liziping National Natural Reserve" longLatPrecision="1" longitude="102.98288" municipality="Ya'an City" pageId="3" pageNumber="454" specimenCount="2" specimenCount-male="2" stateProvince="Sichuan" typeStatus="paratype">
<typeStatus id="54EF881865409663FE92F8A12CAE5799" box="[282,395,1900,1926]" pageId="3" pageNumber="454" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D52FD3365409663FE1EF8A12CC95799" box="[406,492,1900,1926]" count="2" pageId="3" pageNumber="454" type="male">2 males</specimenCount>
(
<collectionCode id="ED45AE7F65409663FE74F8A12F185799" box="[508,573,1900,1926]" country="0" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="3" pageNumber="454">IZAS</collectionCode>
), “
<collectingRegion id="4990F85865409663FDD5F8A12E025799" box="[605,807,1900,1926]" country="China" name="Sichuan" pageId="3" pageNumber="454">Sichuan Province</collectingRegion>
,
<collectingMunicipality id="6B8FACC065409663FCBAF8A12E8E5799" box="[818,939,1900,1926]" pageId="3" pageNumber="454">Ya’an City</collectingMunicipality>
,
<collectingCounty id="628A4E3665409663FC3FF8A129555799" box="[951,1136,1900,1926]" pageId="3" pageNumber="454">Shimian County</collectingCounty>
,
<location id="8E8B606165409663FBF2F8A12DD157B5" LSID="urn:lsid:plazi:treatment:03FD87AC65409665FF1FF9A22E9556A2:8E8B606165409663FBF2F8A12DD157B5" country="China" county="Shimian County" latitude="19.028383" longLatPrecision="1" longitude="102.98288" municipality="Ya'an City" name="Liziping National Natural Reserve" pageId="3" pageNumber="454" stateProvince="Sichuan">Liziping National Natural Reserve</location>
,
<geoCoordinate id="EE60507D65409663FF77F85D2CBD57B5" box="[255,408,1936,1962]" degrees="102.982879" direction="east" orientation="longitude" pageId="3" pageNumber="454" precision="1" value="102.98288">102.982879E</geoCoordinate>
,
<geoCoordinate id="EE60507D65409663FE2BF85D2F1757B5" box="[419,562,1936,1962]" degrees="19.028384" direction="north" orientation="latitude" pageId="3" pageNumber="454" precision="1" value="19.028383">19.028384N</geoCoordinate>
,
<quantity id="4CAC9B5F65409663FDB5F85D2FAB57B5" box="[573,654,1936,1962]" metricMagnitude="3" metricUnit="m" metricValue="1.988" pageId="3" pageNumber="454" unit="m" value="1988.0">
<elevation id="0079D18965409663FDB5F85D2FAB57B5" box="[573,654,1936,1962]" metricMagnitude="3" metricUnit="m" metricValue="1.988" pageId="3" pageNumber="454" unit="m" value="1988.0">1988m</elevation>
</quantity>
,
<location id="8E8B606165409663FD10F85D2E0A57B5" LSID="urn:lsid:plazi:treatment:03FD87AC65409665FF1FF9A22E9556A2:8E8B606165409663FD10F85D2E0A57B5" box="[664,815,1936,1962]" country="China" county="Shimian County" latitude="19.028383" longLatPrecision="1" longitude="102.98288" municipality="Ya'an City" name="Zhudong Liu" pageId="3" pageNumber="454" stateProvince="Sichuan">Zhudong Liu</location>
,
<location id="8E8B606165409663FCB1F85D2E9957B5" LSID="urn:lsid:plazi:treatment:03FD87AC65409665FF1FF9A22E9556A2:8E8B606165409663FCB1F85D2E9957B5" box="[825,956,1936,1962]" country="China" county="Shimian County" latitude="19.028383" longLatPrecision="1" longitude="102.98288" municipality="Ya'an City" name="Zhiming Li" pageId="3" pageNumber="454" stateProvince="Sichuan">Zhiming Li</location>
,
<location id="8E8B606165409663FC40F85D294C57B5" LSID="urn:lsid:plazi:treatment:03FD87AC65409665FF1FF9A22E9556A2:8E8B606165409663FC40F85D294C57B5" box="[968,1129,1936,1962]" country="China" county="Shimian County" latitude="19.028383" longLatPrecision="1" longitude="102.98288" municipality="Ya'an City" name="Pingzhou Zhu" pageId="3" pageNumber="454" stateProvince="Sichuan">Pingzhou Zhu</location>
,
<date id="FFEA107A65409663FBFCF85D29CF57B4" box="[1140,1258,1936,1963]" pageId="3" pageNumber="454" value="2020-05-26">
<collectingDate id="EFAEE99265409663FBFCF85D29CF57B4" box="[1140,1258,1936,1963]" pageId="3" pageNumber="454" value="2020-05-26">2020.5.26</collectingDate>
</date>
”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E653165409665FF4FF8792823527E" lastPageId="5" lastPageNumber="456" pageId="3" pageNumber="454" type="diagnosis">
<paragraph id="8BEB36BA65409664FF4FF8792FA45196" blockId="3.[151,1437,1792,2035]" lastBlockId="4.[151,1437,151,393]" lastPageId="4" lastPageNumber="455" pageId="3" pageNumber="454">
<emphasis id="B920EAA865409663FF4FF8792C1F57D1" bold="true" box="[199,314,1972,1998]" pageId="3" pageNumber="454">Diagnosis</emphasis>
. Frontal furrows shallow and short, reaching level of anterior margin of eyes; vertex finely and sparsely punctate; temporae well-developed (TL/EYL 0.59–0.63). Maximum width of pronotum a little before middle; lateral margins evidently sinuate before posterior angles; posterior angles distinct, nearly rectangular, forming a prominent denticle; pronotal base and lateral sides densely punctate, punctate area reach anterior threefifths of pronotum; area between inner and outer basal foveal grooves shallowly convex, ridge between outer groove and lateral margin distinct; three mid-lateral setae present on each side of pronotum. Apical lamella of aedeagus short (AL/AW 1.09); in lateral view, apical lamella thickened on extreme apex, turned dorsally; in dorsal view, apex truncated; ventral surface of apical lamella with sharp ridge along the apical and right margins (
<figureCitation id="136F2A3F65479664FB6EFE862874517A" box="[1254,1361,331,357]" captionStart="FIGURE 10" captionStartId="16.[151,250,1743,1767]" captionTargetBox="[153,1432,692,1720]" captionTargetId="figure-228@16.[151,1436,669,1720]" captionTargetPageId="16" captionText="FIGURE 10. Apical portion of aedeagus of Pterostichus (Wraseiellus) spp., left lateral view and ventral view. (A) P. pseudandrewesi sp. nov.; (B) P. leishanensis sp. nov.; (C) P. strumatus sp. nov.; (D) P. andrewesi Jedlička." figureDoi="http://doi.org/10.5281/zenodo.11150289" httpUri="https://zenodo.org/record/11150289/files/figure.png" pageId="4" pageNumber="455">Fig. 10A</figureCitation>
); right paramere short, laminar with rounded apex.
</paragraph>
<caption id="DF2B663265479664FF1FF8D129E25747" ID-DOI="http://doi.org/10.5281/zenodo.11150253" ID-Zenodo-Dep="11150253" httpUri="https://zenodo.org/record/11150253/files/figure.png" pageId="4" pageNumber="455" startId="4.[151,250,1820,1844]" targetBox="[191,1407,479,1778]" targetPageId="4" targetType="figure">
<paragraph id="8BEB36BA65479664FF1FF8D129E25747" blockId="4.[151,1436,1820,1881]" pageId="4" pageNumber="455">
<emphasis id="B920EAA865479664FF1FF8D12C36572B" bold="true" box="[151,275,1820,1844]" pageId="4" pageNumber="455">FIGURE 1.</emphasis>
Holotype of
<taxonomicName id="4C544D3965479664FE12F8D12FE5572B" authority="Chen &amp; Yin &amp; Shi, 2024" authorityName="Chen &amp; Yin &amp; Shi" authorityYear="2024" box="[410,704,1820,1844]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="pseudandrewesi" status="sp. nov.">
<emphasis id="B920EAA865479664FE12F8D12FE5572B" box="[410,704,1820,1844]" italics="true" pageId="4" pageNumber="455">Pterostichus pseudandrewesi</emphasis>
</taxonomicName>
<emphasis id="B920EAA865479664FD4EF8D12E32572B" bold="true" box="[710,791,1820,1844]" pageId="4" pageNumber="455">
<taxonomicNameLabel id="A21357D365479664FD4EF8D12E32572B" box="[710,791,1820,1844]" pageId="4" pageNumber="455" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) habitus; (B) median lobe of aedeagus, left lateral view; (C) median lobe of aedeagus, dorsal view; (D) right paramere. Scale bars = 5.0 mm for A, = 1.0 mm for B–D.
</paragraph>
</caption>
<paragraph id="8BEB36BA65479665FF4FF84B28645196" blockId="4.[151,1436,1926,2025]" lastBlockId="5.[151,1437,151,1725]" lastPageId="5" lastPageNumber="456" pageId="4" pageNumber="455">
<emphasis id="B920EAA865479664FF4FF84B2C4357BF" bold="true" box="[199,358,1926,1952]" pageId="4" pageNumber="455">Comparisons</emphasis>
. The new species is most similar to
<taxonomicName id="4C544D3965479664FD67F84A2E5E57BF" box="[751,891,1926,1952]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis id="B920EAA865479664FD67F84A2E5E57BF" box="[751,891,1926,1952]" italics="true" pageId="4" pageNumber="455">P. andrewesi</emphasis>
</taxonomicName>
. Both species are different from other
<taxonomicName id="4C544D3965479664FAA9F84B28B657BF" box="[1313,1427,1926,1952]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis id="B920EAA865479664FAA9F84B28B657BF" box="[1313,1427,1926,1952]" italics="true" pageId="4" pageNumber="455">andrewesi</emphasis>
</taxonomicName>
- group species by the combination of following characters: (1) frontal furrows relatively shallow and short, just reaching level of anterior margin of eyes; (2) pronotum strongly cordiform, posterior margin narrower than or subequal to the anterior margin; (3) area between inner and outer basal foveal grooves not convex; (4) median lobe of aedeagus with apical lamella short and apically truncate, not bent to the left in dorsal view; (5) right paramere short with rounded apex. Comparing to
<taxonomicName id="4C544D3965469665FDE7FF122E2550E6" box="[623,768,223,249]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis id="B920EAA865469665FDE7FF122E2550E6" box="[623,768,223,249]" italics="true" pageId="5" pageNumber="456">P. andrewesi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C544D3965469665FC9AFF122EC050E6" authority="Chen &amp; Yin &amp; Shi, 2024" authorityName="Chen &amp; Yin &amp; Shi" authorityYear="2024" box="[786,997,223,249]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="pseudandrewesi" status="sp. nov.">
<emphasis id="B920EAA865469665FC9AFF122EC050E6" box="[786,997,223,249]" italics="true" pageId="5" pageNumber="456">P. pseudandrewesi</emphasis>
</taxonomicName>
<emphasis id="B920EAA865469665FC78FF12296B50E6" bold="true" box="[1008,1102,223,249]" pageId="5" pageNumber="456">
<taxonomicNameLabel id="A21357D365469665FC78FF12296B50E6" box="[1008,1102,223,249]" pageId="5" pageNumber="456" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is different in the following important aspects: (1) pronotum with three mid-lateral setae on each side (versus only single seta in
<taxonomicName id="4C544D3965469665FB75FECE28AE5102" box="[1277,1419,259,285]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis id="B920EAA865469665FB75FECE28AE5102" box="[1277,1419,259,285]" italics="true" pageId="5" pageNumber="456">P. andrewesi</emphasis>
</taxonomicName>
); (2) pronotal lateral margins abruptly sinuate before posterior angles (versus more gradually sinuate before posterior angles in
<taxonomicName id="4C544D3965469665FE8EFE862CB3517A" box="[262,406,331,357]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis id="B920EAA865469665FE8EFE862CB3517A" box="[262,406,331,357]" italics="true" pageId="5" pageNumber="456">P. andrewesi</emphasis>
</taxonomicName>
); (3) ventral surface of apical lamella of aedeagus not concaved but present a sharp ridge along the apical and right margins (versus distinctly concaved, without a ridge in
<taxonomicName id="4C544D3965469665FBA0FEA229905196" box="[1064,1205,367,393]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis id="B920EAA865469665FBA0FEA229905196" box="[1064,1205,367,393]" italics="true" pageId="5" pageNumber="456">P. andrewesi</emphasis>
</taxonomicName>
, (
<figureCitation id="136F2A3F65469665FB43FEA228115196" box="[1227,1332,367,393]" captionStart="FIGURE 10" captionStartId="16.[151,250,1743,1767]" captionTargetBox="[153,1432,692,1720]" captionTargetId="figure-228@16.[151,1436,669,1720]" captionTargetPageId="16" captionText="FIGURE 10. Apical portion of aedeagus of Pterostichus (Wraseiellus) spp., left lateral view and ventral view. (A) P. pseudandrewesi sp. nov.; (B) P. leishanensis sp. nov.; (C) P. strumatus sp. nov.; (D) P. andrewesi Jedlička." figureDoi="http://doi.org/10.5281/zenodo.11150289" httpUri="https://zenodo.org/record/11150289/files/figure.png" pageId="5" pageNumber="456">Fig. 10D</figureCitation>
).
</paragraph>
<paragraph id="8BEB36BA65469665FF4FFE5E2823527E" blockId="5.[151,1437,151,1725]" pageId="5" pageNumber="456">
<taxonomicName id="4C544D3965469665FF4FFE5E2CB251B2" authority="Chen &amp; Yin &amp; Shi, 2024" authorityName="Chen &amp; Yin &amp; Shi" authorityYear="2024" box="[199,407,403,429]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="pseudandrewesi" status="sp. nov.">
<emphasis id="B920EAA865469665FF4FFE5E2CB251B2" box="[199,407,403,429]" italics="true" pageId="5" pageNumber="456">P. pseudandrewesi</emphasis>
</taxonomicName>
<emphasis id="B920EAA865469665FE28FE5E2CDE51B2" bold="true" box="[416,507,403,429]" pageId="5" pageNumber="456">
<taxonomicNameLabel id="A21357D365469665FE28FE5E2CDE51B2" box="[416,507,403,429]" pageId="5" pageNumber="456" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be easily reminiscent of
<taxonomicName id="4C544D3965469665FCC3FE5E2EE051B3" authorityName="Shi &amp; Sciaky" authorityYear="2013" box="[843,965,403,428]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="comatus">
<emphasis id="B920EAA865469665FCC3FE5E2EE051B3" box="[843,965,403,428]" italics="true" pageId="5" pageNumber="456">P. comatus</emphasis>
</taxonomicName>
, both of which have more than two mid-lateral setae on each side of pronotum. But the new species can be distinguished from the latter by: (1) tempora more developed, TL/EYL 0.59–0.63 (versus
<quantity id="4CAC9B5F65469665FD04FE162E3C51EA" box="[652,793,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="8.763" metricValueMax="9.652" metricValueMin="7.874" pageId="5" pageNumber="456" unit="in" value="0.345" valueMax="0.38" valueMin="0.31">0.31–0.38 in</quantity>
<taxonomicName id="4C544D3965469665FC97FE162EB251EB" authorityName="Shi &amp; Sciaky" authorityYear="2013" box="[799,919,475,500]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="comatus">
<emphasis id="B920EAA865469665FC97FE162EB251EB" box="[799,919,475,500]" italics="true" pageId="5" pageNumber="456">P. comatus</emphasis>
</taxonomicName>
); (2) supraorbital setae placed forwardly, near anterior and posterior margin of eyes respectively (versus the front one placed near mid-length of eyes, and the hind one behind posterior margin of eyes in
<taxonomicName id="4C544D3965469665FDCDFDEE2F995223" authorityName="Shi &amp; Sciaky" authorityYear="2013" box="[581,700,547,572]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="comatus">
<emphasis id="B920EAA865469665FDCDFDEE2F995223" box="[581,700,547,572]" italics="true" pageId="5" pageNumber="456">P. comatus</emphasis>
</taxonomicName>
); (3) apical lamella of aedeagus much shorter and broadened, only weakly bent to the left (versus apical lamella very slender and strongly bent to the left in
<taxonomicName id="4C544D3965469665FBF5FD8A29D0527F" authorityName="Shi &amp; Sciaky" authorityYear="2013" box="[1149,1269,583,608]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="comatus">
<emphasis id="B920EAA865469665FBF5FD8A29D0527F" box="[1149,1269,583,608]" italics="true" pageId="5" pageNumber="456">P. comatus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C34E653165469665FF4FFDA729D355FA" pageId="5" pageNumber="456" type="description">
<paragraph id="8BEB36BA65469665FF4FFDA729D355FA" blockId="5.[151,1437,151,1725]" pageId="5" pageNumber="456">
<emphasis id="B920EAA865469665FF4FFDA72C74529B" bold="true" box="[199,337,618,644]" pageId="5" pageNumber="456">Description</emphasis>
. BL
<quantity id="4CAC9B5F65469665FE06FDA72CCB529A" box="[398,494,618,645]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="5" pageNumber="456" unit="mm" value="15.1">15.1mm</quantity>
; dorsal surface dark brown, elytra somewhat reddish, without metallic luster (
<figureCitation id="136F2A3F65469665FAE7FDA62D9952B6" captionStart="FIGURE 1" captionStartId="4.[151,250,1820,1844]" captionTargetBox="[191,1407,479,1778]" captionTargetId="figure-128@4.[151,1436,450,1794]" captionTargetPageId="4" captionText="FIGURE 1. Holotype of Pterostichus pseudandrewesi sp. nov. (A) habitus; (B) median lobe of aedeagus, left lateral view; (C) median lobe of aedeagus, dorsal view; (D) right paramere. Scale bars = 5.0 mm for A, = 1.0 mm for B–D." figureDoi="http://doi.org/10.5281/zenodo.11150253" httpUri="https://zenodo.org/record/11150253/files/figure.png" pageId="5" pageNumber="456">Fig. 1A</figureCitation>
).
<emphasis id="B920EAA865469665FF5CFD422C3152B6" bold="true" box="[212,276,655,681]" pageId="5" pageNumber="456">Head</emphasis>
. Frons finely and sparsely punctate, relatively densely in frontal furrows; frontal furrows short and shallow, just reaching level of anterior margin of eyes; vertex finely and sparsely punctate; temporae well-developed (TL/EYL 0.59–0.63), distinctly swollen behind eyes; paraocular grooves deep, impunctate and nearly straight, reaching posterior margin of eyes; antennomere 3 without accessory setae.
<emphasis id="B920EAA865469665FC6EFD36297A530A" bold="true" box="[998,1119,763,789]" pageId="5" pageNumber="456">Pronotum</emphasis>
strongly cordiform (PW/PL 1.31); anterior and posterior margins almost straight, lateral margins evenly curved from anterior angles to posterior third, abruptly sinuate before posterior angles; anterior margin slightly wider than posterior margin; anterior angles obtusely rounded, not projecting; posterior angles nearly rectangular, apex sharp and forming a prominent denticle; discal area glabrous, with faint transverse wrinkles; basal fovea relatively short and deep, densely punctate, area between inner and outer grooves rugose and flat, mid-basal area between two basal foveae densely punctate, ridge between outer groove and lateral margin distinct; some fine and sparse punctures also present in lateral channel and pronotum side area; three mid-lateral setae and one basal-lateral seta present on each side.
<emphasis id="B920EAA865469665FB48FC3A2829540E" bold="true" box="[1216,1292,1015,1041]" pageId="5" pageNumber="456">Elytra</emphasis>
oblong (EL/ EW 1.49), lateral margins subparallel before middle, slightly expanded behind middle; humeral teeth very small; intervals slightly convex; striae moderately deep, without punctures; parascutellar stria joined to stria 1, angular base of stria 1 free; umbilicate series on interval 9 composed of approximately 16–19 pores, sparse in middle and dense in anterior and posterior areas; males with microsculpture typically transverse on basal half, gradually turned to isodiametric towards apex.
<emphasis id="B920EAA865469665FE61FB662F5D54DB" bold="true" box="[489,632,1194,1221]" pageId="5" pageNumber="456">Ventral side</emphasis>
. Proepisterna and metaepisterna densely and coarsely punctate; sternite VII of males with a lunate transverse ridge at about anterior third, median region of sternite VII shallowly depressed posterior to the ridge.
<emphasis id="B920EAA865469665FE07FB3E2CE25512" bold="true" box="[399,455,1267,1293]" pageId="5" pageNumber="456">Legs</emphasis>
. Tarsomere 5 with three or four pairs of setae beneath; apical half of metatibiae slightly curved in males.
<emphasis id="B920EAA865469665FED3FADA2F22552F" bold="true" box="[347,519,1302,1329]" pageId="5" pageNumber="456">Male genitalia</emphasis>
(
<figureCitation id="136F2A3F65469665FD9EFADA2FB3552E" box="[534,662,1303,1329]" captionStart="FIGURE 1" captionStartId="4.[151,250,1820,1844]" captionTargetBox="[191,1407,479,1778]" captionTargetId="figure-128@4.[151,1436,450,1794]" captionTargetPageId="4" captionText="FIGURE 1. Holotype of Pterostichus pseudandrewesi sp. nov. (A) habitus; (B) median lobe of aedeagus, left lateral view; (C) median lobe of aedeagus, dorsal view; (D) right paramere. Scale bars = 5.0 mm for A, = 1.0 mm for B–D." figureDoi="http://doi.org/10.5281/zenodo.11150253" httpUri="https://zenodo.org/record/11150253/files/figure.png" pageId="5" pageNumber="456">Figs 1B–D</figureCitation>
,
<figureCitation id="136F2A3F65469665FD2BFADA2FF2552E" box="[675,727,1303,1329]" captionStart="FIGURE 10" captionStartId="16.[151,250,1743,1767]" captionTargetBox="[153,1432,692,1720]" captionTargetId="figure-228@16.[151,1436,669,1720]" captionTargetPageId="16" captionText="FIGURE 10. Apical portion of aedeagus of Pterostichus (Wraseiellus) spp., left lateral view and ventral view. (A) P. pseudandrewesi sp. nov.; (B) P. leishanensis sp. nov.; (C) P. strumatus sp. nov.; (D) P. andrewesi Jedlička." figureDoi="http://doi.org/10.5281/zenodo.11150289" httpUri="https://zenodo.org/record/11150289/files/figure.png" pageId="5" pageNumber="456">10A</figureCitation>
). Median lobe of aedeagus relatively stout, tapered to apex; in lateral view, ventral margin almost straight near middle, apex not bent ventrally, apical orifice large, slightly turned to the left side, apical lamella thickened apically and turned dorsally; in dorsal view, apical lamella short and wide (AL /AW 1.09), apex truncated, not twisted or bent to the left; ventral surface of apical lamella with a sharp ridge along apical and right margins, surfaces of both sides of the ridge not concaved (
<figureCitation id="136F2A3F65469665FBBCFA6A29BB55DE" box="[1076,1182,1447,1473]" captionStart="FIGURE 10" captionStartId="16.[151,250,1743,1767]" captionTargetBox="[153,1432,692,1720]" captionTargetId="figure-228@16.[151,1436,669,1720]" captionTargetPageId="16" captionText="FIGURE 10. Apical portion of aedeagus of Pterostichus (Wraseiellus) spp., left lateral view and ventral view. (A) P. pseudandrewesi sp. nov.; (B) P. leishanensis sp. nov.; (C) P. strumatus sp. nov.; (D) P. andrewesi Jedlička." figureDoi="http://doi.org/10.5281/zenodo.11150289" httpUri="https://zenodo.org/record/11150289/files/figure.png" pageId="5" pageNumber="456">Fig. 10A</figureCitation>
). Right paramere very short, laminar, apex rounded. Endophallus of male genitalia not studied. Female genitalia unknown.
</paragraph>
</subSubSection>
<subSubSection id="C34E653165469665FF4FFA2229DC5632" pageId="5" pageNumber="456" type="etymology">
<paragraph id="8BEB36BA65469665FF4FFA2229DC5632" blockId="5.[151,1437,151,1725]" pageId="5" pageNumber="456">
<emphasis id="B920EAA865469665FF4FFA222C6D5616" bold="true" box="[199,328,1519,1545]" pageId="5" pageNumber="456">Etymology</emphasis>
. The name
<taxonomicName id="4C544D3965469665FE50FA222FB25616" authorityName="Chen &amp; Yin &amp; Shi" authorityYear="2024" box="[472,663,1519,1545]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="pseudandrewesi">
“
<emphasis id="B920EAA865469665FE6CFA222FB25616" box="[484,663,1519,1545]" italics="true" pageId="5" pageNumber="456">pseudandrewesi</emphasis>
</taxonomicName>
” is the combination of the Greek epithet “
<emphasis id="B920EAA865469665FB1AFA2229F65616" box="[1170,1235,1519,1545]" italics="true" pageId="5" pageNumber="456">pseud</emphasis>
-” [false] and the species name
<taxonomicName id="4C544D3965469665FEBBF9DE2C945632" box="[307,433,1555,1581]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="andrewesi">
“
<emphasis id="B920EAA865469665FEB7F9DE2C945632" box="[319,433,1555,1581]" italics="true" pageId="5" pageNumber="456">andrewesi</emphasis>
</taxonomicName>
”, referring to the similarity of this new species with
<taxonomicName id="4C544D3965469665FB89F9DE29DC5632" authority="Jedlicka." authorityName="Jedlicka." box="[1025,1273,1555,1581]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis id="B920EAA865469665FB89F9DE29AB5632" box="[1025,1166,1555,1581]" italics="true" pageId="5" pageNumber="456">P. andrewesi</emphasis>
Jedlička.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C34E653165469665FF4FF9FB2E9556A2" pageId="5" pageNumber="456" type="distribution">
<paragraph id="8BEB36BA65469665FF4FF9FB2C41566A" blockId="5.[151,1437,151,1725]" pageId="5" pageNumber="456">
<emphasis id="B920EAA865469665FF4FF9FB2C7C564F" bold="true" box="[199,345,1590,1616]" pageId="5" pageNumber="456">Distribution</emphasis>
. Only known from the
<typeStatus id="54EF881865469665FDF8F9FA2F85564E" box="[624,672,1591,1617]" pageId="5" pageNumber="456">type</typeStatus>
locality, Pukui Mountain, Butuo County, in the southwest part of
<collectingRegion id="4990F85865469665FF1FF9962C7A566A" box="[151,351,1627,1653]" country="China" name="Sichuan" pageId="5" pageNumber="456">Sichuan Province</collectingRegion>
.
</paragraph>
<paragraph id="8BEB36BA65469665FF4FF9B32E9556A2" blockId="5.[151,1437,151,1725]" pageId="5" pageNumber="456">
<emphasis id="B920EAA865469665FF4FF9B32C065687" bold="true" box="[199,291,1662,1688]" pageId="5" pageNumber="456">Habitat</emphasis>
. The
<typeStatus id="54EF881865469665FEE1F9B22CEE5686" box="[361,459,1663,1689]" pageId="5" pageNumber="456" type="holotype">holotype</typeStatus>
of the new species was collected under rock or dead wood in alpine grassland with sparse stumps. The grassland is probably formed by historical logging.
</paragraph>
</subSubSection>
</treatment>
</document>
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