treatments-xml/data/03/E5/33/03E53311FFAAFFD2B62CFDC5FC46F923.xml
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<document id="7BF3077ED3BCBE678B2BED8315A7375B" ID-DOI="10.24823/EJB.2022.407" ID-ISSN="1747-0036" ID-Zenodo-Dep="10553332" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1704825216459" checkinUser="felipe" docAuthor="Moonlight, Peter &amp; Fuentes, Alfredo F." docDate="2022" docId="03E53311FFAAFFD2B62CFDC5FC46F923" docLanguage="en" docName="EdinbJBot.79.407.1-66.pdf" docOrigin="Edinburgh Journal of Botany 79 (407)" docSource="http://dx.doi.org/10.24823/ejb.2022.407" docStyle="DocumentStyle:73DE8CF451BB5BCBD336E07A8276E690.4:EdinbJBot.2021-.journal_article" docStyleId="73DE8CF451BB5BCBD336E07A8276E690" docStyleName="EdinbJBot.2021-.journal_article" docStyleVersion="4" docTitle="Begonia unduavensis Rusby, Descr. S. Amer. Pl." docType="treatment" docVersion="4" lastPageNumber="38" masterDocId="FFDC4B69FF8EFFF7B655FFBDFFD6FF96" masterDocTitle="AN UPDATED CHECKLIST AND KEY TO THE SPECIES OF BOLIVIAN BEGONIA, INCLUDING ONE NEW SPECIES" masterLastPageNumber="66" masterPageNumber="1" pageNumber="37" updateTime="1706008907290" updateUser="carolina" zenodo-license-document="CC-BY-4.0">
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<mods:title id="D362F69DA114343DAE95A221A4E4242B">AN UPDATED CHECKLIST AND KEY TO THE SPECIES OF BOLIVIAN BEGONIA, INCLUDING ONE NEW SPECIES</mods:title>
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<emphasis id="B9385E15FFAAFFD3B62CFDC5FF64FD07" bold="true" box="[121,178,632,657]" pageId="36" pageNumber="37">5.28.</emphasis>
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<emphasis id="B9385E15FFAAFFD3B6ECFDC5FE7AFD07" bold="true" box="[185,428,632,657]" pageId="36" pageNumber="37">Begonia unduavensis</emphasis>
Rusby, Descr. S. Amer. Pl.
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64 (1920).
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Department,
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677 (
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[
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by Smith, L.B. &amp; Schubert, B.G
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.;
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[
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).
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<bibCitation id="5E668A6BFFAAFFD3B6C2FC8FFC5DFCDD" author="L. B. Smith &amp; B. G. Schubert" box="[151,907,818,843]" issue="87" journal="Revista Univ. (Cuzco)" page="80" pageId="36" pageNumber="37" score="0.150130185271274" volume="33" year="1944">
L.B. Smith &amp; B.G. Schubert,
<bibCitation id="5E668A6BFFAAFFD3B79BFC8FFC5DFCDD" box="[462,907,818,843]" issue="87" journal="Revista Univ. (Cuzco)" page="80" pageId="36" pageNumber="37" score="0.150130185271274" volume="33" year="1944">Revista Univ. (Cuzco) 33(87): 80 (1944)</bibCitation>
</bibCitation>
; R.C. Foster, Contr. Gray Herb. 184: 137 (1958); D.C. Wasshausen
<emphasis id="B9385E15FFAAFFD3B4F5FCE5FD01FCE7" box="[672,727,856,881]" italics="true" pageId="36" pageNumber="37">et al.</emphasis>
in P.M. JØrgensen
<emphasis id="B9385E15FFAAFFD3B5F8FCE5FC35FCE7" box="[941,995,856,881]" italics="true" pageId="36" pageNumber="37">et al.</emphasis>
(eds), Cat.
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,
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.
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<taxonomicName id="4C4CF984FFAAFFD3B62CFC1FFD86FC2D" authority="Rusby, Descr. S. Amer. Pl." box="[121,592,930,955]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="species" species="lignosa">
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<emphasis id="B9385E15FFAAFFD3B62CFC1FFEFCFC2D" box="[121,298,930,955]" italics="true" pageId="36" pageNumber="37">Begonia lignosa</emphasis>
Rusby
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, Descr. S. Amer. Pl.
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65 (1920). Type:
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, [
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], Prov. Sud Yungas, Unduavi, [
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16
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19S
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,
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67
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54W
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2899 (
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NY [
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, first stage designated in: Revista
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by Smith, L.B. &amp; Schubert, B.G.;
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[
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,
<specimenCode id="DBEA2A7CFFAAFFD3B47CFB85FD6CFBC7" box="[553,698,1080,1105]" pageId="36" pageNumber="37">US00074725</specimenCode>
]
</materialsCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C356D18CFFAAFFD3B6C2FBE0FC0FFB46" pageId="36" pageNumber="37" type="reference_group">
<paragraph id="8BF38207FFAAFFD3B6C2FBE0FD76FBE0" blockId="36.[121,1135,669,1142]" box="[151,672,1117,1142]" pageId="36" pageNumber="37">R.C. Foster, Contr. Gray Herb. 184: 137 (1958).</paragraph>
<paragraph id="8BF38207FFAAFFD3B62CFB2FFC0FFB46" blockId="36.[121,1108,1170,1232]" pageId="36" pageNumber="37">
<treatmentCitationGroup id="AB5CA529FFAAFFD3B62CFB2FFC0FFB46" pageId="36" pageNumber="37">
<taxonomicName id="4C4CF984FFAAFFD3B62CFB2FFC92FB3D" ID-CoL="87YZ4" authorityName="Irmsch" box="[121,836,1170,1195]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="species" species="obtecticaulis">
<emphasis id="B9385E15FFAAFFD3B62CFB2FFEBEFB3D" box="[121,360,1170,1195]" italics="true" pageId="36" pageNumber="37">Begonia obtecticaulis</emphasis>
auct. non. Irmsch., D.C. Wasshausen
<emphasis id="B9385E15FFAAFFD3B544FB2FFC92FB3D" box="[785,836,1170,1195]" italics="true" pageId="36" pageNumber="37">et al</emphasis>
</taxonomicName>
<emphasis id="B9385E15FFAAFFD3B511FB2FFC9EFB3D" box="[836,840,1170,1195]" italics="true" pageId="36" pageNumber="37">.</emphasis>
in P.M. JØrgensen
<emphasis id="B9385E15FFAAFFD3B248FB2FFB82FB3D" box="[1053,1108,1170,1195]" italics="true" pageId="36" pageNumber="37">et al.</emphasis>
(eds), Cat.
<treatmentCitation id="0AEDA416FFAAFFD3B747FB0AFC00FB46" author="Bolivia" box="[274,982,1207,1232]" journal="Monogr. Syst. Bot. Missouri Bot. Gard." page="385" pageId="36" pageNumber="37" score="1124.4052770756598" volume="129" year="2013">
<bibCitation id="5E668A6BFFAAFFD3B747FB0AFC00FB46" author="Bolivia" box="[274,982,1207,1232]" journal="Monogr. Syst. Bot. Missouri Bot. Gard." page="385" pageId="36" pageNumber="37" score="1124.4052770756598" volume="129" year="2013">
<collectingCountry id="F35BC297FFAAFFD3B747FB0AFE89FB46" box="[274,351,1207,1232]" name="Bolivia" pageId="36" pageNumber="37">Bolivia</collectingCountry>
,
<bibCitation id="5E668A6BFFAAFFD3B73CFB0AFC00FB46" box="[361,982,1207,1232]" journal="Monogr. Syst. Bot. Missouri Bot. Gard." page="385" pageId="36" pageNumber="37" score="1222.4916941493243" volume="129" year="2013">Monogr. Syst. Bot. Missouri Bot. Gard. 129: 385 (2013)</bibCitation>
</bibCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C356D18CFFAAFFD2B62CFB56FD3FFCEE" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="description">
<paragraph id="8BF38207FFAAFFD2B62CFB56FD3FFCEE" blockId="36.[121,1165,1259,1695]" lastBlockId="37.[121,1162,266,888]" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
Caulescent herb, lacking a tuber or rhizome.
<emphasis id="B9385E15FFAAFFD3B425FB56FD7CFA92" box="[624,682,1259,1284]" italics="true" pageId="36" pageNumber="37">Stem</emphasis>
erect, to
<quantity id="4CB42FE2FFAAFFD3B547FB56FC8EFA92" box="[786,856,1259,1284]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="37" unit="cm" value="50.0">50 cm</quantity>
tall,
<quantity id="4CB42FE2FFAAFFD3B5D8FB56FBD4FA92" box="[909,1026,1259,1284]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="4.0" metricValueMin="1.5" pageId="36" pageNumber="37" unit="mm" value="2.75" valueMax="4.0" valueMin="1.5">1.54 mm</quantity>
in diameter, branching, internodes (5)
<quantity id="4CB42FE2FFAAFFD3B7CBFAACFDF4FABC" box="[414,546,1297,1322]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="5.0" metricValueMin="2.5" pageId="36" pageNumber="37" unit="mm" value="37.5" valueMax="50.0" valueMin="25.0">2550 mm</quantity>
, glabrous to sparsely pilose, red.
<emphasis id="B9385E15FFAAFFD3B5CCFAACFC24FABC" box="[921,1010,1297,1322]" italics="true" pageId="36" pageNumber="37">Stipules</emphasis>
persistent, triangular, 718 ×
<quantity id="4CB42FE2FFAAFFD3B710FA8BFE71FAD9" box="[325,423,1334,1359]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="36" pageNumber="37" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">24 mm</quantity>
, apex acuminate, mucronate, glabrous, light brown, margin entire, aciliate.
<emphasis id="B9385E15FFAAFFD3B771FAE6FEA4FAE2" box="[292,370,1371,1396]" italics="true" pageId="36" pageNumber="37">Leaves</emphasis>
alternate, basifixed;
<emphasis id="B9385E15FFAAFFD3B40DFAE6FD79FAE2" box="[600,687,1371,1396]" italics="true" pageId="36" pageNumber="37">petioles</emphasis>
joining blade at almost a right angle,
<quantity id="4CB42FE2FFAAFFD3B62CFA3CFF31FA0C" box="[121,231,1409,1434]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="2.8" metricValueMin="0.5" pageId="36" pageNumber="37" unit="mm" value="16.5" valueMax="28.0" valueMin="5.0">528 mm</quantity>
long, glabrous to sparsely pilose;
<emphasis id="B9385E15FFAAFFD3B432FA3CFD66FA0C" box="[615,688,1409,1434]" italics="true" pageId="36" pageNumber="37">blades</emphasis>
asymmetrical, transversely deltoid, 3095 ×
<quantity id="4CB42FE2FFAAFFD3B6DBFA1BFEDAFA29" box="[142,268,1446,1471]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.5" metricValueMin="1.5" pageId="36" pageNumber="37" unit="mm" value="25.0" valueMax="35.0" valueMin="15.0">1535 mm</quantity>
, apex acuminate, base cuneate on the narrow side of the blade, auriculate on the broad side of the blade, venation palmate-pinnate, 4 or 5 veins from the base, 3 or 4 veins on the broad side of the blade, 2 or 3 on the narrow side, upper surface dark green, sometimes with a red line around the margin, sparsely pilose, lower surface pale green to red, sparsely pilose, margin double-dentate, ciliate.
<emphasis id="B9385E15FFAAFFD3B4ECF986FC8DF9C2" box="[697,859,1595,1620]" italics="true" pageId="36" pageNumber="37">Inflorescences</emphasis>
13 per stem, axillary, erect, arising close to the apex of the stem, a dichasial cyme, protandrous, with up to 16 staminate flowers and 8 pistillate flowers;
<emphasis id="B9385E15FFAAFFD3B403F93BFD6DF909" box="[598,699,1670,1695]" italics="true" pageId="36" pageNumber="37">peduncle</emphasis>
<quantity id="4CB42FE2FFAAFFD3B497F93BFCE8F909" box="[706,830,1670,1695]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.3" metricValueMin="2.2" pageId="36" pageNumber="37" unit="mm" value="27.5" valueMax="33.0" valueMin="22.0">2233 mm</quantity>
long, glabrous, red;
<emphasis id="B9385E15FFAAFFD2B274F93BFE8BFEB5" italics="true" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">pedicels of staminate flowers</emphasis>
<quantity id="4CB42FE2FFABFFD2B731FEB7FE04FEB5" box="[356,466,266,291]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="2.2" metricValueMin="0.5" pageId="37" pageNumber="38" unit="mm" value="13.5" valueMax="22.0" valueMin="5.0">522 mm</quantity>
long, glabrous, red;
<emphasis id="B9385E15FFABFFD2B4E0FEB7FC3AFEB5" box="[693,1004,266,291]" italics="true" pageId="37" pageNumber="38">pedicels of pistillate flowers</emphasis>
<quantity id="4CB42FE2FFABFFD2B5A7FEB7FBB6FEB5" box="[1010,1120,266,291]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.6" metricValueMin="0.7" pageId="37" pageNumber="38" unit="mm" value="11.5" valueMax="16.0" valueMin="7.0">716 mm</quantity>
long, glabrous, red;
<emphasis id="B9385E15FFABFFD2B700FE92FE4CFEDE" box="[341,410,303,328]" italics="true" pageId="37" pageNumber="38">bracts</emphasis>
persistent, elliptic to lanceolate, 47 ×
<quantity id="4CB42FE2FFABFFD2B506FE92FC63FEDE" box="[851,949,303,328]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="37" pageNumber="38" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">13 mm</quantity>
, apex obtuse to acute, red to brown, glabrous, margin lacerate, ciliate.
<emphasis id="B9385E15FFABFFD2B48DFEE9FC76FEFB" box="[728,928,340,365]" italics="true" pageId="37" pageNumber="38">Staminate flowers</emphasis>
:
<emphasis id="B9385E15FFABFFD2B5FBFEE9FC24FEFB" box="[942,1010,340,365]" italics="true" pageId="37" pageNumber="38">tepals</emphasis>
2, spreading, white to pink, orbicular, 614 ×
<quantity id="4CB42FE2FFABFFD2B78FFEC7FD9CFE05" box="[474,586,378,403]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.4" metricValueMin="0.7" pageId="37" pageNumber="38" unit="mm" value="10.5" valueMax="14.0" valueMin="7.0">714 mm</quantity>
, apex rounded, base rounded, margin entire, glabrous;
<emphasis id="B9385E15FFABFFD2B6B2FE22FE91FE2E" box="[231,327,415,440]" italics="true" pageId="37" pageNumber="38">stamens</emphasis>
c.75, fused at the base, spreading, filaments
<quantity id="4CB42FE2FFABFFD2B513FE22FC6CFE2E" box="[838,954,415,440]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="37" pageNumber="38" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.52 mm</quantity>
long, anthers symmetrically basifixed, oblong,
<quantity id="4CB42FE2FFABFFD2B7B2FE79FDA7FE4B" box="[487,625,452,477]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="37" pageNumber="38" unit="mm" value="1.35" valueMax="1.5" valueMin="1.2">1.21.5 mm</quantity>
long, dehiscing via lateral slits, connectives extended
<quantity id="4CB42FE2FFABFFD2B6B2FE57FEA3FD95" box="[231,373,490,515]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="37" pageNumber="38" unit="mm" value="0.4" valueMax="0.5" valueMin="0.3">0.30.5 mm</quantity>
.
<emphasis id="B9385E15FFABFFD2B72AFE57FDE1FD95" box="[383,567,490,515]" italics="true" pageId="37" pageNumber="38">Pistillate flowers</emphasis>
:
<emphasis id="B9385E15FFABFFD2B411FE57FD6EFD95" box="[580,696,490,515]" italics="true" pageId="37" pageNumber="38">bracteoles</emphasis>
lacking;
<emphasis id="B9385E15FFABFFD2B54EFE57FC89FD95" box="[795,863,490,515]" italics="true" pageId="37" pageNumber="38">tepals</emphasis>
deciduous in fruit, 5, spreading, elliptic to ovate, 914 ×
<quantity id="4CB42FE2FFABFFD2B456FDB2FDB3FDBE" box="[515,613,527,552]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="37" pageNumber="38" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">36 mm</quantity>
, apex rounded, margin entire, aciliate, glabrous, white to pink;
<emphasis id="B9385E15FFABFFD2B740FD89FE58FDDB" box="[277,398,564,589]" italics="true" pageId="37" pageNumber="38">ovary body</emphasis>
ovoid, 4.55 ×
<quantity id="4CB42FE2FFABFFD2B469FD89FD65FDDB" box="[572,691,564,589]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="37" pageNumber="38" unit="mm" value="3.75" valueMax="4.0" valueMin="3.5">3.54 mm</quantity>
, pink, 3-winged, wings pink, one wing longer than the other two, rectangular, ascending, front edge truncate, apex truncate, lower edge slightly rounded, 69 ×
<quantity id="4CB42FE2FFABFFD2B7EAFDC2FDF9FD0E" box="[447,559,639,664]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.3" metricValueMin="0.7" pageId="37" pageNumber="38" unit="mm" value="10.0" valueMax="13.0" valueMin="7.0">713 mm</quantity>
, the shorter two wings semicircular to triangular, front edge truncate, apex rounded, lower edge gently curved, 4.55 ×
<quantity id="4CB42FE2FFABFFD2B5D0FD19FC31FD2B" box="[901,999,676,701]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="37" pageNumber="38" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">12 mm</quantity>
, 3-locular, placentas bifid, bearing ovules on both surfaces;
<emphasis id="B9385E15FFABFFD2B4F4FD77FD34FD75" box="[673,738,714,739]" italics="true" pageId="37" pageNumber="38">styles</emphasis>
3, yellow, free at base, irregularly multifid from around half their length, stigmatic papillae in a spirally twisted band.
<emphasis id="B9385E15FFABFFD2B24DFD52FF1EFCBB" italics="true" pageId="37" pageNumber="38">Fruiting pedicel</emphasis>
to
<quantity id="4CB42FE2FFABFFD2B6B9FCA9FEECFCBB" box="[236,314,788,813]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="37" pageNumber="38" unit="cm" value="3.2">3.2 cm</quantity>
long;
<emphasis id="B9385E15FFABFFD2B72AFCA9FE7AFCBB" box="[383,428,788,813]" italics="true" pageId="37" pageNumber="38">fruit</emphasis>
held erect, body ovoid, 9.512.5 ×
<quantity id="4CB42FE2FFABFFD2B561FCA9FC6FFCBB" box="[820,953,788,813]" metricMagnitude="-3" metricUnit="m" metricValue="9.75" metricValueMax="12.0" metricValueMin="7.5" pageId="37" pageNumber="38" unit="mm" value="9.75" valueMax="12.0" valueMin="7.5">7.512 mm</quantity>
, light brown, glabrous, wings same shape as in pistillate flower, light brown, longest wing expanding to 17 ×
<quantity id="4CB42FE2FFABFFD2B6D8FCE2FF09FCEE" box="[141,223,863,888]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="37" pageNumber="38" unit="mm" value="30.0">30 mm</quantity>
, shorter two wings expanding to 10.5 ×
<quantity id="4CB42FE2FFABFFD2B4F5FCE2FD33FCEE" box="[672,741,863,888]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="37" pageNumber="38" unit="mm" value="3.0">3 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C356D18CFFABFFD2B62CFC2EFE08FC3A" box="[121,478,915,940]" pageId="37" pageNumber="38" type="distribution">
<paragraph id="8BF38207FFABFFD2B62CFC2EFE08FC3A" blockId="37.[121,478,915,940]" box="[121,478,915,940]" pageId="37" pageNumber="38">
<emphasis id="B9385E15FFABFFD2B62CFC2EFF2EFC3A" box="[121,248,915,940]" italics="true" pageId="37" pageNumber="38">Distribution</emphasis>
. Endemic to
<collectingCountry id="F35BC297FFABFFD2B7D9FC2EFE0CFC3A" box="[396,474,915,940]" name="Bolivia" pageId="37" pageNumber="38">Bolivia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C356D18CFFABFFD2B62CFC75FC46F923" pageId="37" pageNumber="38" type="discussion">
<paragraph id="8BF38207FFABFFD2B62CFC75FE7DFB70" blockId="37.[121,1164,968,1441]" pageId="37" pageNumber="38">
<emphasis id="B9385E15FFABFFD2B62CFC75FEB6FC77" box="[121,352,968,993]" italics="true" pageId="37" pageNumber="38">Nomenclatural notes</emphasis>
. The name
<taxonomicName id="4C4CF984FFABFFD2B7B0FC75FCCAFC77" authority="Rusby" authorityName="Rusby" box="[485,796,968,993]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="37" pageNumber="38" phylum="Tracheophyta" rank="species" species="unduavensis">
<emphasis id="B9385E15FFABFFD2B7B0FC75FD06FC77" box="[485,720,968,993]" italics="true" pageId="37" pageNumber="38">Begonia unduavensis</emphasis>
Rusby
</taxonomicName>
was published based on the collection
<emphasis id="B9385E15FFABFFD2B6B8FC50FEBEFB90" box="[237,360,1005,1030]" italics="true" pageId="37" pageNumber="38">H.H. Rusby</emphasis>
677, but no herbaria were cited. Rusbys protologue cited material collected at an elevation of
<quantity id="4CB42FE2FFABFFD2B7E4FBAFFDD5FBBD" box="[433,515,1042,1067]" metricMagnitude="3" metricUnit="m" metricValue="2.4384" pageId="37" pageNumber="38" unit="ft" value="8000.0">8000 ft</quantity>
, and the known sheets of this material state either
<quantity id="4CB42FE2FFABFFD2B62CFB85FF1CFBC7" box="[121,202,1080,1105]" metricMagnitude="3" metricUnit="m" metricValue="2.4384" pageId="37" pageNumber="38" unit="ft" value="8000.0">
<elevation id="00616534FFABFFD2B62CFB85FF1CFBC7" box="[121,202,1080,1105]" metricMagnitude="3" metricUnit="m" metricValue="2.4384" pageId="37" pageNumber="38" unit="ft" value="8000.0">8000 ft</elevation>
</quantity>
or
<quantity id="4CB42FE2FFABFFD2B6BAFB85FE8FFBC7" box="[239,345,1080,1105]" metricMagnitude="3" metricUnit="m" metricValue="3.6576" pageId="37" pageNumber="38" unit="ft" value="12000.0">
<elevation id="00616534FFABFFD2B6BAFB85FE8FFBC7" box="[239,345,1080,1105]" metricMagnitude="3" metricUnit="m" metricValue="3.6576" pageId="37" pageNumber="38" unit="ft" value="12000.0">12,000 ft.</elevation>
</quantity>
<collectingCounty id="6292FA8BFFABFFD2B735FB85FE5DFBC7" box="[352,395,1080,1105]" pageId="37" pageNumber="38">The</collectingCounty>
two sheets that state
<quantity id="4CB42FE2FFABFFD2B4D2FB85FD0EFBC7" box="[647,728,1080,1105]" metricMagnitude="3" metricUnit="m" metricValue="2.4384" pageId="37" pageNumber="38" unit="ft" value="8000.0">
<elevation id="00616534FFABFFD2B4D2FB85FD0EFBC7" box="[647,728,1080,1105]" metricMagnitude="3" metricUnit="m" metricValue="2.4384" pageId="37" pageNumber="38" unit="ft" value="8000.0">8000 ft</elevation>
</quantity>
are one in the
<collectingRegion id="49884CE5FFABFFD2B52AFB85FC3DFBC7" box="[895,1003,1080,1105]" country="United States of America" name="New York" pageId="37" pageNumber="38">New York</collectingRegion>
<collectingMunicipality id="6B97187DFFABFFD2B5A7FB85FF1CFBE0" pageId="37" pageNumber="38">Botanical Garden</collectingMunicipality>
herbarium (
<specimenCode id="DBEA2A7CFFABFFD2B706FBE0FE30FBE0" box="[339,486,1117,1142]" collectionCode="NY" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="37" pageNumber="38" type="Herbarium">NY00118708</specimenCode>
) and one in the
<location id="8E93D4DCFFABFFD2B4C8FBE0FC6DFBE0" LSID="urn:lsid:plazi:treatment:03E53311FFAAFFD2B62CFDC5FC46F923:8E93D4DCFFABFFD2B4C8FBE0FC6DFBE0" box="[669,955,1117,1142]" county="The" municipality="Botanical Garden" name="Field Museum in Chicago" pageId="37" pageNumber="38" stateProvince="New York">Field Museum in Chicago</location>
(
<specimenCode id="DBEA2A7CFFABFFD2B59DFBE0FB95FBE0" box="[968,1091,1117,1142]" collectionCode="F" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="37" pageNumber="38" type="Herbarium">F0052644</specimenCode>
F).
<bibRefCitation id="EFDDFFF6FFABFFD2B62FFB3FFE4BFB0D" author="Smith LB &amp; Schubert BG" box="[122,413,1154,1179]" pageId="37" pageNumber="38" pagination="71 - 87" refId="ref41249" refString="Smith LB, Schubert BG. 1944 a. Revision de las especies Bolivianas de genero Begonia. Revista Universitat [Universidad Nacional de Cusco]. 87: 71 - 87." type="journal article" year="1944">Smith &amp; Schubert (1944a)</bibRefCitation>
cited the type of
<taxonomicName id="4C4CF984FFABFFD2B437FB3FFC9BFB0D" box="[610,845,1154,1179]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="37" pageNumber="38" phylum="Tracheophyta" rank="species" species="unduavensis">
<emphasis id="B9385E15FFABFFD2B437FB3FFC9BFB0D" box="[610,845,1154,1179]" italics="true" pageId="37" pageNumber="38">Begonia unduavensis</emphasis>
</taxonomicName>
in the
<collectingRegion id="49884CE5FFABFFD2B5CFFB3FFBD3FB0D" box="[922,1029,1154,1179]" country="United States of America" name="New York" pageId="37" pageNumber="38">New York</collectingRegion>
<location id="8E93D4DCFFABFFD2B259FB3FFF1CFB57" LSID="urn:lsid:plazi:treatment:03E53311FFAAFFD2B62CFDC5FC46F923:8E93D4DCFFABFFD2B259FB3FFF1CFB57" county="The" municipality="Botanical Garden" name="Botanical Garden" pageId="37" pageNumber="38" stateProvince="New York">Botanical Garden</location>
herbarium, which is an effective lectotypification.
<location id="8E93D4DCFFABFFD2B4AAFB15FCF5FB57" LSID="urn:lsid:plazi:treatment:03E53311FFAAFFD2B62CFDC5FC46F923:8E93D4DCFFABFFD2B4AAFB15FCF5FB57" box="[767,803,1192,1217]" county="The" municipality="Botanical Garden" name="We" pageId="37" pageNumber="38" stateProvince="New York">We</location>
cite as
<typeStatus id="54F73CA5FFABFFD2B52EFB15FC09FB57" box="[891,991,1192,1217]" pageId="37" pageNumber="38" type="syntype">syntypes</typeStatus>
all sheets that were collected at
<quantity id="4CB42FE2FFABFFD2B714FB70FE7DFB70" box="[321,427,1229,1254]" metricMagnitude="3" metricUnit="m" metricValue="3.6576" pageId="37" pageNumber="38" unit="ft" value="12000.0">
<elevation id="00616534FFABFFD2B714FB70FE7DFB70" box="[321,427,1229,1254]" metricMagnitude="3" metricUnit="m" metricValue="3.6576" pageId="37" pageNumber="38" unit="ft" value="12000.0">12,000 ft.</elevation>
</quantity>
</paragraph>
<paragraph id="8BF38207FFABFFD2B6CCFB4FFDFDFA37" blockId="37.[121,1164,968,1441]" pageId="37" pageNumber="38">
The protologue of
<taxonomicName id="4C4CF984FFABFFD2B73FFB4FFDB0FA9D" authority="Rusby" authorityName="Rusby" box="[362,614,1266,1291]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="37" pageNumber="38" phylum="Tracheophyta" rank="species" species="lignosa">
<emphasis id="B9385E15FFABFFD2B73FFB4FFDCCFA9D" box="[362,538,1266,1291]" italics="true" pageId="37" pageNumber="38">Begonia lignosa</emphasis>
Rusby
</taxonomicName>
cited the collection
<emphasis id="B9385E15FFABFFD2B51FFB4FFC1EFA9D" box="[842,968,1266,1291]" italics="true" pageId="37" pageNumber="38">O. Buchtien</emphasis>
2899, but no herbarium (
<bibRefCitation id="EFDDFFF6FFABFFD2B6ACFAA5FE55FAA7" author="Rusby HH" box="[249,387,1304,1329]" pageId="37" pageNumber="38" pagination="1 - 170" refId="ref41068" refString="Rusby HH. 1920. Three hundred new species of South American Plants. New York: published by the author. pp. 1 - 170." type="book chapter" year="1920">Rusby, 1920</bibRefCitation>
).
<bibRefCitation id="EFDDFFF6FFABFFD2B7CFFAA5FD68FAA7" author="Smith LB &amp; Schubert BG" box="[410,702,1304,1329]" pageId="37" pageNumber="38" pagination="71 - 87" refId="ref41249" refString="Smith LB, Schubert BG. 1944 a. Revision de las especies Bolivianas de genero Begonia. Revista Universitat [Universidad Nacional de Cusco]. 87: 71 - 87." type="journal article" year="1944">Smith &amp; Schubert (1944a)</bibRefCitation>
cited material in the
<collectingRegion id="49884CE5FFABFFD2B5FEFAA5FBC1FAA7" box="[939,1047,1304,1329]" country="United States of America" name="New York" pageId="37" pageNumber="38">New York</collectingRegion>
Botanical Garden herbarium as the type, which counts as an effective lectotypification because there are two matching sheets in
<collectingRegion id="49884CE5FFABFFD2B7E7FADFFDF7FAED" box="[434,545,1378,1403]" country="United States of America" name="New York" pageId="37" pageNumber="38">New York</collectingRegion>
. We select the duplicate NY00118625 as the
<typeStatus id="54F73CA5FFABFFD2B24AFADFFB5EFAED" box="[1055,1160,1378,1403]" pageId="37" pageNumber="38" type="lectotype">lectotype</typeStatus>
, because it has both flowers and fruits.
</paragraph>
<paragraph id="8BF38207FFABFFD2B62CFA01FC46F923" blockId="37.[121,1160,1468,1717]" pageId="37" pageNumber="38">
<emphasis id="B9385E15FFABFFD2B62CFA01FE98FA43" box="[121,334,1468,1493]" italics="true" pageId="37" pageNumber="38">Identification notes</emphasis>
.
<taxonomicName id="4C4CF984FFABFFD2B70EFA01FD91FA43" box="[347,583,1468,1493]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="37" pageNumber="38" phylum="Tracheophyta" rank="species" species="unduavensis">
<emphasis id="B9385E15FFABFFD2B70EFA01FD91FA43" box="[347,583,1468,1493]" italics="true" pageId="37" pageNumber="38">Begonia unduavensis</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C4CF984FFABFFD2B54FFA01FC2DFA43" authority="Rusby" authorityName="Rusby" box="[794,1019,1468,1493]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="37" pageNumber="38" phylum="Tracheophyta" rank="species" species="unilateralis">
<emphasis id="B9385E15FFABFFD2B54FFA01FC79FA43" box="[794,943,1468,1493]" italics="true" pageId="37" pageNumber="38">B. unilateralis</emphasis>
Rusby
</taxonomicName>
, with both species having relatively small (&lt;
<quantity id="4CB42FE2FFABFFD2B7A6FA5FFDFDFA6D" box="[499,555,1506,1531]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="37" pageNumber="38" unit="cm" value="8.0">8 cm</quantity>
long), obliquely lanceolate to ovate leaves that are truncate at the base. They are best distinguished by their indumentum.
<taxonomicName id="4C4CF984FFABFFD2B5C8F9BAFB5EF9B6" box="[925,1160,1543,1568]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="37" pageNumber="38" phylum="Tracheophyta" rank="species" species="unduavensis">
<emphasis id="B9385E15FFABFFD2B5C8F9BAFB5EF9B6" box="[925,1160,1543,1568]" italics="true" pageId="37" pageNumber="38">Begonia unduavensis</emphasis>
</taxonomicName>
has stems, petioles and leaf laminae that are sparsely to densely pilose, whereas
<taxonomicName id="4C4CF984FFABFFD2B62CF9EFFED9F9FD" box="[121,271,1618,1643]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="37" pageNumber="38" phylum="Tracheophyta" rank="species" species="unilateralis">
<emphasis id="B9385E15FFABFFD2B62CF9EFFED9F9FD" box="[121,271,1618,1643]" italics="true" pageId="37" pageNumber="38">B. unilateralis</emphasis>
</taxonomicName>
has an indumentum of peltate scales. This indumentum is densest on the stems, petioles, and veins on the leaf lower surface (although it can rub off); moderately dense on the lower leaf surface; and sparse on the upper leaf surface.
</paragraph>
</subSubSection>
</treatment>
</document>