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<document id="8071A63CB6EA1027AFCDA98D1B3A07E7" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1627005272297" checkinUser="felipe" docAuthor="Published, First" docDate="2008" docId="03DF87D2FF80AB1B9EC8159A452D3D91" docLanguage="en" docName="zt01804p456part1.pdf" docOrigin="Zootaxa 1804" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Corethrella (Corethrella) douglasi Borkent 2008, new species" docType="treatment" docVersion="2" lastPageNumber="111" masterDocId="FFE6FFAAFFECAB759E5F1025465D3823" masterDocTitle="The Frog-Biting Midges of the World (Corethrellidae: Diptera)" masterLastPageNumber="456" masterPageNumber="1" pageNumber="109" updateTime="1699014769975" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="094F38E1E7DB6FB4E3D0817D80AAAE66">The Frog-Biting Midges of the World (Corethrellidae: Diptera)</mods:title>
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<mods:date id="C528915ABE9CBDD2B4CB910EBEEA777E">2008</mods:date>
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<treatment id="03DF87D2FF80AB1B9EC8159A452D3D91" LSID="urn:lsid:plazi:treatment:03DF87D2FF80AB1B9EC8159A452D3D91" httpUri="http://treatment.plazi.org/id/03DF87D2FF80AB1B9EC8159A452D3D91" lastPageId="110" lastPageNumber="111" pageId="108" pageNumber="109">
<subSubSection id="C36C654FFF80AB199EC8159A451D3DFA" box="[151,832,1471,1497]" pageId="108" pageNumber="109" type="nomenclature">
<paragraph id="8BC936C4FF80AB199EC8159A451D3DFA" blockId="108.[151,832,1471,1497]" box="[151,832,1471,1497]" pageId="108" pageNumber="109">
<heading id="D08181A8FF80AB199EC8159A451D3DFA" bold="true" box="[151,832,1471,1497]" fontSize="11" level="1" pageId="108" pageNumber="109" reason="1">
<emphasis id="B902EAD6FF80AB199EC8159A451D3DFA" bold="true" box="[151,832,1471,1497]" pageId="108" pageNumber="109">
<taxonomicName id="4C764D47FF80AB199EC8159A44F93DFA" ID-CoL="8BL5P" authority="Borkent" authorityName="Borkent" authorityYear="2008" box="[151,676,1471,1497]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="douglasi" status="sp. nov." subGenus="Corethrella">
<emphasis id="B902EAD6FF80AB199EC8159A477C3DFA" bold="true" box="[151,289,1471,1497]" italics="true" pageId="108" pageNumber="109">Corethrella</emphasis>
(
<emphasis id="B902EAD6FF80AB199F6D159A47E13DFA" bold="true" box="[306,444,1471,1497]" italics="true" pageId="108" pageNumber="109">Corethrella</emphasis>
)
<emphasis id="B902EAD6FF80AB199F91159A44693DFA" bold="true" box="[462,564,1471,1497]" italics="true" pageId="108" pageNumber="109">douglasi</emphasis>
Borkent
</taxonomicName>
,
<taxonomicNameLabel id="A23157ADFF80AB199CEE159A451D3DFA" box="[689,832,1471,1497]" pageId="108" pageNumber="109" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36C654FFF80AB199EC8162F43D63EBF" pageId="108" pageNumber="109" type="diagnosis">
<paragraph id="8BC936C4FF80AB199EC8162F43D63EBF" blockId="108.[151,1436,1546,2012]" pageId="108" pageNumber="109">
DIAGNOSIS:
<emphasis id="B902EAD6FF80AB199F10162E44343E07" box="[335,617,1547,1572]" italics="true" pageId="108" pageNumber="109">Male and female adults</emphasis>
: only extant species of
<taxonomicName id="4C764D47FF80AB199DD7162E42523E07" box="[904,1039,1547,1572]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis id="B902EAD6FF80AB199DD7162E42523E07" box="[904,1039,1547,1572]" italics="true" pageId="108" pageNumber="109">Corethrella</emphasis>
</taxonomicName>
in the New World with the wing with a distinct midlength band (Figs. 63D, 69C), with the katepisternum darker dorsally than ventrally (
<figureCitation id="134D2A41FF80AB199B32161746903E57" captionStart="FIGURE 48" captionStartId="354.[151,255,2014,2038]" captionTargetBox="[177,1406,203,1979]" captionTargetId="figure-28@354.[177,1410,197,1987]" captionTargetPageId="354" captionText="FIGURE 48. Thoraces and legs of female Corethrella, lateral view." pageId="108" pageNumber="109">Fig. 48A</figureCitation>
), with a uniformly light brown midfemur (some with very faint apical darker pigmentation) contrasting with the dark base of the hind femur, and with the hind tibia with discrete basal and apical bands (
<figureCitation id="134D2A41FF80AB199B4E16A743233EBF" box="[1297,1406,1666,1692]" captionStart="FIGURE 48" captionStartId="354.[151,255,2014,2038]" captionTargetBox="[177,1406,203,1979]" captionTargetId="figure-28@354.[177,1410,197,1987]" captionTargetPageId="354" captionText="FIGURE 48. Thoraces and legs of female Corethrella, lateral view." pageId="108" pageNumber="109">Fig. 48A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36C654FFF80AB189E99168F47DA3E2B" lastPageId="109" lastPageNumber="110" pageId="108" pageNumber="109" type="description">
<paragraph id="8BC936C4FF80AB189E99168F45E73C04" blockId="108.[151,1436,1546,2012]" lastBlockId="109.[151,1437,152,2027]" lastPageId="109" lastPageNumber="110" pageId="108" pageNumber="109">
DESCRIPTION:
<emphasis id="B902EAD6FF80AB199FC5168E44443EE7" box="[410,537,1707,1732]" italics="true" pageId="108" pageNumber="109">Male adult</emphasis>
. Descriptive statistics: see
<tableCitation id="C6F4037FFF80AB199D08168F45873EE7" box="[855,986,1706,1732]" captionStart-0="TABLE 2" captionStart-1="TABLE 3" captionStart-2="TABLE 4" captionStart-3="TABLE 5" captionStartId-0="269.[151,241,151,175]" captionStartId-1="271.[151,241,153,177]" captionStartId-2="273.[151,241,151,175]" captionStartId-3="275.[151,241,151,175]" captionText-0="TABLE 2. Length of wing of males of species of Corethrella. Values are in mm." captionText-1="TABLE 3. Ratio of foreleg tarsomeres 3/4 of males of species of Corethrella." captionText-2="TABLE 4. Ratio of distance of dorsomedial seta from the gonocoxite base/length of the gonocoxite of males of species" captionText-3="TABLE 5. Ratio of distance of subbasal seta from the gonostylus base/ length of the gonostylus of males of species of Corethrella." pageId="108" pageNumber="109">Tables 25</tableCitation>
.
<emphasis id="B902EAD6FF80AB199DB3168F42723EE7" bold="true" box="[1004,1071,1706,1732]" pageId="108" pageNumber="109">Head</emphasis>
: Outline in anterior view laterally elongate (as in
<figureCitation id="134D2A41FF80AB199FD716F747813ECF" box="[392,476,1746,1772]" captionStart="FIGURE 9" captionStartId="315.[151,255,1947,1971]" captionTargetBox="[177,1410,239,1904]" captionTargetId="figure-23@315.[177,1411,197,1921]" captionTargetPageId="315" captionText="FIGURE 9. Head capsules of female Corethrella, anterior view." pageId="108" pageNumber="109">Fig. 9I</figureCitation>
). Two large setae on frons between ventromedial area of ommatida (as in
<figureCitation id="134D2A41FF80AB199B3216F746973F37" captionStart="FIGURE 16" captionStartId="322.[151,255,1947,1971]" captionTargetBox="[177,1410,349,1917]" captionTargetId="figure-31@322.[177,1411,197,1921]" captionTargetPageId="322" captionText="FIGURE 16. Dorsal portion of head capsules of female Corethrella, anterior view." pageId="108" pageNumber="109">Fig. 16E</figureCitation>
). Antenna light brown; pedicel with at least one distinctive, more elongate, stout, dorsal or dorsolateral seta; flagellomeres as in
<figureCitation id="134D2A41FF80AB199FE0170744763F1F" box="[447,555,1826,1852]" captionStart="FIGURE 22" captionStartId="328.[151,255,1947,1971]" captionTargetBox="[208,1396,238,1921]" captionTargetId="figure-25@328.[177,1411,197,1921]" captionTargetPageId="328" captionText="FIGURE 22. Antennae of male Corethrella." pageId="108" pageNumber="109">Fig. 22A</figureCitation>
, sensilla coeloconica distributed as in
<tableCitation id="C6F4037FFF80AB199DA9170742123F1F" box="[1014,1103,1826,1852]" captionStart="TABLE 1" captionStartId="265.[151,241,151,175]" captionText="TABLE 1. Pattern of sensilla coeloconica (= s.c.) on antennal flagellomeres of species of Corethrella. Numbers refer to the flagellomere with sensilla coeloconica, with those in brackets referring to flagellomeres with or without sensilla coeloconica. Mis. (missing) refers to damaged specimens, unknown refers to missing stages, uncertain refers to specimens in which visibility was too poor to be certain of whether sensilla coeloconica were present or not. Those species names with a “†” are those of fossils. Corethrella librata, known only as immature stages, is not included." pageId="108" pageNumber="109">Table 1</tableCitation>
; flagellomere 13 with welldeveloped apical bifurcation. Palpus pale; segment 3 of constant width or slightly oval.
<emphasis id="B902EAD6FF80AB199AE2176F43473F47" bold="true" box="[1213,1306,1866,1892]" pageId="108" pageNumber="109">Thorax</emphasis>
(as in
<figureCitation id="134D2A41FF80AB199B32176F46923FAF" captionStart="FIGURE 48" captionStartId="354.[151,255,2014,2038]" captionTargetBox="[177,1406,203,1979]" captionTargetId="figure-28@354.[177,1410,197,1987]" captionTargetPageId="354" captionText="FIGURE 48. Thoraces and legs of female Corethrella, lateral view." pageId="108" pageNumber="109">Fig. 48A</figureCitation>
): Light brown with darker pigmentation in band from pronotum to anterior portion of anepimeron and dorsal portion of katepisternum. Posterior portion of dorsocentral row with group of about 11 elongate setae. Prescutal suture short, not extending more than half way to dorsocentral row of setae. Anterior anepisternum divided diagonally by sinuous suture, dorsal portion about equal to ventral portion. Ventral portion of posterior anepisternum triangular, uniformly brown, with anterodorsal margin thick.
<emphasis id="B902EAD6FF81AB189A61109A42DF38FA" bold="true" box="[1086,1154,191,217]" pageId="109" pageNumber="110">Wing</emphasis>
(Fig. 63D): Apex of R
<subScript id="17F23481FF81AB189BCB10F543C138FD" attach="left" box="[1428,1436,208,222]" fontSize="6" pageId="109" pageNumber="110">2</subScript>
basal to apex of M
<subScript id="17F23481FF81AB189F2E10DE4724392A" attach="left" box="[369,377,251,265]" fontSize="6" pageId="109" pageNumber="110">1</subScript>
. Anterior margin with differently, discretely pigmented scales (indicating anterior margin of midlength band), with midlength band; veins (other than costa and wing margin) with well-developed scales. Halter very light brown, lighter than scutellum.
<emphasis id="B902EAD6FF81AB189D65111845293974" bold="true" box="[826,884,317,343]" pageId="109" pageNumber="110">Legs</emphasis>
(as in
<figureCitation id="134D2A41FF81AB189D97111842653974" box="[968,1080,317,343]" captionStart="FIGURE 48" captionStartId="354.[151,255,2014,2038]" captionTargetBox="[177,1406,203,1979]" captionTargetId="figure-28@354.[177,1410,197,1987]" captionTargetPageId="354" captionText="FIGURE 48. Thoraces and legs of female Corethrella, lateral view." pageId="109" pageNumber="110">Fig. 48A</figureCitation>
): Light brown with mid-coxa, trochanter somewhat darker, apex of midfemur slightly pigmented in some; hind-coxa, trochanter, very base of hind femur dark brown; hind tibia with base, apex discrete dark brown, at least mid-, hind leg tarsomeres 2 4 with banding. With only slender setae, lacking scales (except for some in patch of whip-like setae on posterior portion of hind tibia). Midleg with thick, subapical setae on each of at least tarsomeres 13. Apices of fore-, midleg fifth tarsomeres undivided, with claws slightly subapical to apical (as in Fig. 75F). Claw of foreleg longer than those of mid-, hind leg. Each claw without inner tooth. Anterior claws of each leg without a basal prong. Foreleg claws unequal. Midleg claws equal. Foreleg third tarsomere shorter than fourth tarsomere. Empodia slender.
<emphasis id="B902EAD6FF81AB189F81125844053AB4" bold="true" box="[478,600,637,663]" pageId="109" pageNumber="110">Abdomen</emphasis>
(Fig. 78G): Tergites 17 uniformly pigmented light brown, with slight pigmentation on posterior margin of tergites 6, 7 in some. Sternites 12 medium brown, 37 light brown with posterior margins medium brown. Segments 8 medium brown. Segment 9 medium brown in some.
<emphasis id="B902EAD6FF81AB189B7712E843C63AC4" bold="true" box="[1320,1435,717,743]" pageId="109" pageNumber="110">Genitalia</emphasis>
(Fig. 90C): Gonocoxite light brown, strongly tapering; anteromedial area with spicules similar in length to those elsewhere on gonocoxite; with well-defined dorsal row of setae, with setae 1, 2 thicker than others, 2 thicker than 1, with row curving posteromedially with posteriormost seta of row positioned dorsomedially. With one dorsomedial stout seta, more or less of even thickness for most of length, tapering near apex. Gonostylus (partially extended) basally straight, evenly curved for apical 1/3; slender, somewhat thicker for basal 1/ 3, of more or less equal thickness for remainder, tapered apically; one elongate, slender subbasal seta, situated anteriorly or anteroventrally; apical seta slender, short, simple. Aedeagus slender, elongate, tapering gradually to apex, pointed apically, with lateral margins fused subapically.
</paragraph>
<paragraph id="8BC936C4FF81AB189E99141343C53D3B" blockId="109.[151,1437,152,2027]" pageId="109" pageNumber="110">
<emphasis id="B902EAD6FF81AB189E99141347393C6C" box="[198,356,1078,1103]" italics="true" pageId="109" pageNumber="110">Female adult</emphasis>
. Descriptive statistics: see
<tableCitation id="C6F4037FFF81AB189CEB1410451A3C6C" box="[692,839,1077,1103]" captionStart-0="TABLE 6" captionStart-1="TABLE 7" captionStart-2="TABLE 8" captionStart-3="TABLE 9" captionStart-4="TABLE 10" captionStart-5="TABLE 11" captionStartId-0="277.[151,241,153,177]" captionStartId-1="280.[151,241,153,177]" captionStartId-2="283.[151,241,153,177]" captionStartId-3="286.[151,241,151,175]" captionStartId-4="289.[151,239,153,177]" captionStartId-5="292.[151,241,153,177]" captionText-0="TABLE 6. Length of wing of females of species of Corethrella. Values are in mm." captionText-1="TABLE 7. Ratio of length of R1/wing length of females of species of Corethrella." captionText-2="TABLE 8. Ratio of head capsule width/height of females of species of Corethrella." captionText-3="TABLE 9. Number of setae on the clypeus of females of species of Corethrella. The number does not include those" captionText-4="TABLE 10. Number of posterior anepisternal setae of females of species of Corethrella." captionText-5="TABLE 11. Number of anepimeral setae of females of species of Corethrella." pageId="109" pageNumber="110">Tables 611</tableCitation>
. As for male, with following differences.
<emphasis id="B902EAD6FF81AB189B0E141043C93C6C" bold="true" box="[1361,1428,1077,1103]" pageId="109" pageNumber="110">Head</emphasis>
: Coronal suture short, not extending ventrally between ommatidia (as in
<figureCitation id="134D2A41FF81AB189D83147B42183C5B" box="[988,1093,1118,1144]" captionStart="FIGURE 16" captionStartId="322.[151,255,1947,1971]" captionTargetBox="[177,1410,349,1917]" captionTargetId="figure-31@322.[177,1411,197,1921]" captionTargetPageId="322" captionText="FIGURE 16. Dorsal portion of head capsules of female Corethrella, anterior view." pageId="109" pageNumber="110">Fig. 16E</figureCitation>
). Antenna flagellomeres 16 light brown, 713 somewhat darker; with flagellomeres as in
<figureCitation id="134D2A41FF81AB189D2E14A345803C83" box="[881,989,1158,1184]" captionStart="FIGURE 28" captionStartId="334.[151,255,1947,1971]" captionTargetBox="[208,1396,238,1917]" captionTargetId="figure-25@334.[177,1411,197,1921]" captionTargetPageId="334" captionText="FIGURE 28. Antennae of female Corethrella." pageId="109" pageNumber="110">Fig. 28K</figureCitation>
, sensilla coeloconica distributed as in
<tableCitation id="C6F4037FFF81AB189EC8148B46B23CEB" box="[151,239,1198,1224]" captionStart="TABLE 1" captionStartId="265.[151,241,151,175]" captionText="TABLE 1. Pattern of sensilla coeloconica (= s.c.) on antennal flagellomeres of species of Corethrella. Numbers refer to the flagellomere with sensilla coeloconica, with those in brackets referring to flagellomeres with or without sensilla coeloconica. Mis. (missing) refers to damaged specimens, unknown refers to missing stages, uncertain refers to specimens in which visibility was too poor to be certain of whether sensilla coeloconica were present or not. Those species names with a “†” are those of fossils. Corethrella librata, known only as immature stages, is not included." pageId="109" pageNumber="110">Table 1</tableCitation>
. Clypeus (
<figureCitation id="134D2A41FF81AB189F2A148B479A3CEB" box="[373,455,1198,1224]" captionStart="FIGURE 17" captionStartId="323.[151,255,1947,1971]" captionTargetBox="[177,1408,258,1907]" captionTargetId="figure-28@323.[177,1411,197,1921]" captionTargetPageId="323" captionText="FIGURE 17. Clypei of female Corethrella. Lateral setae, even if present, are not shown." pageId="109" pageNumber="110">Fig. 17</figureCitation>
AN) squarish. Mandible with large, apically somewhat squared teeth. Palpus as in
<figureCitation id="134D2A41FF81AB189EC814F346A53CD3" box="[151,248,1238,1264]" captionStart="FIGURE 34" captionStartId="340.[151,255,1947,1971]" captionText="FIGURE 34. Right palpi of female Corethrella." pageId="109" pageNumber="110">Fig. 34J</figureCitation>
.
<emphasis id="B902EAD6FF81AB189F5614F047103CCC" bold="true" box="[265,333,1237,1263]" pageId="109" pageNumber="110">Wing</emphasis>
(Fig. 69C).
<emphasis id="B902EAD6FF81AB189FBD14F044413CCC" bold="true" box="[482,540,1237,1263]" pageId="109" pageNumber="110">Legs</emphasis>
: Claws of each leg equal to those of others; equal on each leg, simple (without inner teeth).
<emphasis id="B902EAD6FF81AB189F3F14D847873D34" bold="true" box="[352,474,1277,1303]" pageId="109" pageNumber="110">Abdomen</emphasis>
: Light brown, with sternites 1, 2, segment 8 medium brown. Cercus light brown.
</paragraph>
<paragraph id="8BC936C4FF81AB189E99150346813D9B" blockId="109.[151,1437,152,2027]" pageId="109" pageNumber="110">
<emphasis id="B902EAD6FF81AB189E99150347593D1C" box="[198,260,1318,1343]" italics="true" pageId="109" pageNumber="110">Pupa</emphasis>
. Mostly undescribed.
<emphasis id="B902EAD6FF81AB189C76150044DB3D1C" bold="true" box="[553,646,1317,1343]" pageId="109" pageNumber="110">Thorax</emphasis>
: Scutum, metathorax each with spherical sensory pit (as in Fig. 100A). Respiratory organ (Fig. 102C): Tubular.
<emphasis id="B902EAD6FF81AB189CBD156845013D44" bold="true" box="[738,860,1357,1383]" pageId="109" pageNumber="110">Abdomen</emphasis>
(Fig. 106D): Segments 37 not expanded laterally. Paddle only moderately elongate; apicodorsal thick spine articulating; apicoventral seta longer than thick spine.
</paragraph>
<paragraph id="8BC936C4FF81AB189E9915E347E33DC3" blockId="109.[151,1437,152,2027]" box="[198,446,1478,1504]" pageId="109" pageNumber="110">
<emphasis id="B902EAD6FF81AB189E9915E347513DFC" box="[198,268,1478,1503]" italics="true" pageId="109" pageNumber="110">Larva</emphasis>
. Undescribed.
</paragraph>
<paragraph id="8BC936C4FF81AB189E9915CB47DA3E2B" blockId="109.[151,1437,152,2027]" box="[198,391,1518,1544]" pageId="109" pageNumber="110">
<emphasis id="B902EAD6FF81AB189E9915CB46A83E24" box="[198,245,1518,1543]" italics="true" pageId="109" pageNumber="110">Egg</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36C654FFF81AB1B9E99163347523971" lastPageId="110" lastPageNumber="111" pageId="109" pageNumber="110" type="distribution">
<paragraph id="8BC936C4FF81AB189E99163344883EF3" blockId="109.[151,1437,152,2027]" pageId="109" pageNumber="110">
DISTRIBUTION AND BIONOMICS:
<taxonomicName id="4C764D47FF81AB189CF7163345C33E0C" authorityName="Borkent" authorityYear="2008" box="[680,926,1558,1583]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="douglasi">
<emphasis id="B902EAD6FF81AB189CF7163345C33E0C" box="[680,926,1558,1583]" italics="true" pageId="109" pageNumber="110">Corethrella douglasi</emphasis>
</taxonomicName>
is known from
<collectingCountry id="F3617654FF81AB189A3D163342B83E13" box="[1122,1253,1558,1584]" name="Costa Rica" pageId="109" pageNumber="110">Costa Rica</collectingCountry>
(Fig. 120A) at altitudes ranging from
<quantity id="4C8E9B21FF81AB189FFA161B44703E7B" box="[421,557,1598,1624]" metricMagnitude="2" metricUnit="m" metricValue="5.85" metricValueMax="10.9" metricValueMin="0.8" pageId="109" pageNumber="110" unit="m" value="585.0" valueMax="1090.0" valueMin="80.0">801090 m</quantity>
, with most records from over 500 meters. Specimens have been collected using a sweep net, Malaise traps and by rearing. Reared specimens were collected as larvae or pupae from the leaf axils of
<taxonomicName id="4C764D47FF81AB189F6C16AB44333E84" box="[307,622,1678,1703]" class="Liliopsida" family="Araceae" genus="Dieffenbachia" kingdom="Plantae" order="Alismatales" pageId="109" pageNumber="110" phylum="Tracheophyta" rank="species" species="longispatha">
<emphasis id="B902EAD6FF81AB189F6C16AB44333E84" box="[307,622,1678,1703]" italics="true" pageId="109" pageNumber="110">Dieffenbachia longispatha</emphasis>
</taxonomicName>
in Volcán Tenorio National Park and from a bromeliad,
<taxonomicName id="4C764D47FF81AB189B7A16AB46823EEC" class="Liliopsida" family="Bromeliaceae" genus="Pitcairnia" kingdom="Plantae" order="Poales" pageId="109" pageNumber="110" phylum="Tracheophyta" rank="species" species="valeni">
<emphasis id="B902EAD6FF81AB189B7A16AB46823EEC" italics="true" pageId="109" pageNumber="110">Pitcairnia valeni</emphasis>
</taxonomicName>
, in Reserva Biológica Alberto Ml. Brenes.
</paragraph>
<paragraph id="8BC936C4FF81AB189E9916FB422A3FB8" blockId="109.[151,1437,152,2027]" pageId="109" pageNumber="110">
Three females were collected with an ABC/CO
<subScript id="17F23481FF81AB189D5B16C845513ED8" attach="right" box="[772,780,1773,1787]" fontSize="6" pageId="109" pageNumber="110">2</subScript>
/light trap at
<quantity id="4C8E9B21FF81AB189DF416FB45B43EDB" box="[939,1001,1758,1784]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="109" pageNumber="110" unit="km" value="5.0">5 km</quantity>
W.
<collectingRegion id="49B2F826FF81AB189A4016FB42293EDB" box="[1055,1140,1758,1784]" country="Puerto Rico" name="Rincon" pageId="109" pageNumber="110">Rincón</collectingRegion>
and
<specimenCount id="9D70FD4DFF81AB189AEC16FB436A3EDB" box="[1203,1335,1758,1784]" pageId="109" pageNumber="110" type="female">one female</specimenCount>
with an ABC/light/octenol trap at Los Planes Station in Corcovado National Park. It is uncertain what these latter specimens were attracted to. Generally
<taxonomicName id="4C764D47FF81AB189C2F171444AA3F69" box="[624,759,1841,1866]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="109" pageNumber="110" phylum="Arthropoda" rank="genus">
<emphasis id="B902EAD6FF81AB189C2F171444AA3F69" box="[624,759,1841,1866]" italics="true" pageId="109" pageNumber="110">Corethrella</emphasis>
</taxonomicName>
are rare in light traps and a frog-call trap was run nearby the ABC trap at
<quantity id="4C8E9B21FF81AB189F0A177C47CD3F50" box="[341,400,1881,1907]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="109" pageNumber="110" unit="km" value="5.0">5 km</quantity>
W.
<collectingRegion id="49B2F826FF81AB189FE1177C444F3F50" box="[446,530,1881,1907]" country="Puerto Rico" name="Rincon" pageId="109" pageNumber="110">Rincón</collectingRegion>
without collecting
<taxonomicName id="4C764D47FF81AB189CAB177C45263F51" authorityName="Borkent" authorityYear="2008" box="[756,891,1881,1906]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="douglasi">
<emphasis id="B902EAD6FF81AB189CAB177C45263F51" box="[756,891,1881,1906]" italics="true" pageId="109" pageNumber="110">C. douglasi</emphasis>
</taxonomicName>
, suggesting that it is not attracted to the call of
<taxonomicName id="4C764D47FF81AB189EC817A447413FB9" authorityName="Le Conte" authorityYear="1856" box="[151,284,1921,1946]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="109" pageNumber="110" phylum="Chordata" rank="species" species="gratiosa">
<emphasis id="B902EAD6FF81AB189EC817A447413FB9" box="[151,284,1921,1946]" italics="true" pageId="109" pageNumber="110">H. gratiosa</emphasis>
</taxonomicName>
. The female adults have serrate teeth indicating that they do indeed bite.
</paragraph>
<paragraph id="8BC936C4FF81AB1B9E99178C47523971" blockId="109.[151,1437,152,2027]" lastBlockId="110.[151,1436,152,1458]" lastPageId="110" lastPageNumber="111" pageId="109" pageNumber="110">
TAXONOMIC DISCUSSION: Males and females were associated through the shared presence of a distinctive pigmentation pattern and were reared together from a single collection of immatures. It is a striking feature of the distribution of this species that there appear to be two populations: one at upper elevations above 500 meters in most of
<collectingCountry id="F3617654FF82AB1B9FB110E5443238F9" box="[494,623,192,218]" name="Costa Rica" pageId="110" pageNumber="111">Costa Rica</collectingCountry>
and one in the Osa Peninsula in southwestern
<collectingCountry id="F3617654FF82AB1B9AC910E5434438F9" box="[1174,1305,192,218]" name="Costa Rica" pageId="110" pageNumber="111">Costa Rica</collectingCountry>
. This suggests the possibility of two species but I was unable to distinguish these on morphological grounds. Alternately, the lowland records in the Osa Peninsula may be specimens descending from higher elevations in the peninsula.
</paragraph>
</subSubSection>
<subSubSection id="C36C654FFF82AB1B9E991145452D3D91" pageId="110" pageNumber="111" type="description">
<paragraph id="8BC936C4FF82AB1B9E991145437D3CC9" blockId="110.[151,1436,152,1458]" pageId="110" pageNumber="111">
TYPES:
<typeStatus id="54CD8866FF82AB1B9F69114547FB3959" box="[310,422,352,378]" pageId="110" pageNumber="111" type="holotype">Holotype</typeStatus>
, male adult with pupal exuviae on microscope slide, labeled &quot;
<typeStatus id="54CD8866FF82AB1B9A2F114543523959" box="[1136,1295,352,378]" pageId="110" pageNumber="111" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C764D47FF82AB1B9B4A114547393981" authority="Borkent" authorityName="Borkent" authorityYear="2008" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="douglasi">
<emphasis id="B902EAD6FF82AB1B9B4A114546A13982" italics="true" pageId="110" pageNumber="111">Corethrella douglasi</emphasis>
Borkent
</taxonomicName>
, Costa Rica”, “
<collectingCountry id="F3617654FF82AB1B9C4B11AD44983981" box="[532,709,392,418]" name="Costa Rica" pageId="110" pageNumber="111">COSTA RICA</collectingCountry>
, Prov.
<collectingRegion id="49B2F826FF82AB1B9D4B11AD45273981" box="[788,890,392,418]" country="Costa Rica" name="Alajuela" pageId="110" pageNumber="111">Alajuela</collectingRegion>
, Upala, Bijaqua, ACA-T, P.N. Volcán Tenorio, Sendero al Volcán Tenorio, orilla del rio Roble,
<quantity id="4C8E9B21FF82AB1B9C8F1195454339E9" box="[720,798,432,458]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="110" pageNumber="111" unit="m" value="850.0">850 m</quantity>
,
<date id="FFC81004FF82AB1B9D741195459139E9" box="[811,972,432,458]" pageId="110" pageNumber="111" value="2004-06-27">27 JUN 2004</date>
, G. Chaverri. En pecidos de
<taxonomicName id="4C764D47FF82AB1B9B411195478139D1" class="Liliopsida" family="Araceae" genus="Dieffenbachia" kingdom="Plantae" order="Alismatales" pageId="110" pageNumber="111" phylum="Tracheophyta" rank="species" species="longispatha">
<emphasis id="B902EAD6FF82AB1B9B411195470039D2" italics="true" pageId="110" pageNumber="111">Dieffenbachia longispatha</emphasis>
(Araceae)
</taxonomicName>
, LN 297100 428543, LGCh-825.10” (INBC).
<typeStatus id="54CD8866FF82AB1B9A5711FD422C39D1" box="[1032,1137,472,498]" pageId="110" pageNumber="111" type="allotype">Allotype</typeStatus>
, female adult with larval and pupal exuviae on microscope slide, labeled as for
<typeStatus id="54CD8866FF82AB1B9D77122545CF3A39" box="[808,914,512,538]" pageId="110" pageNumber="111" type="holotype">holotype</typeStatus>
, but LGCh-825.04 (INBC).
<typeStatus id="54CD8866FF82AB1B9AB7122543023A39" box="[1256,1375,512,538]" pageId="110" pageNumber="111" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D70FD4DFF82AB1B9B33122543C93A30" box="[1388,1428,512,538]" pageId="110" pageNumber="111" type="male">6 ♂</specimenCount>
, each with pupal exuviae,
<specimenCount id="9D70FD4DFF82AB1B9F98120D47AD3A18" box="[455,496,552,578]" pageId="110" pageNumber="111" type="male">1 ♂</specimenCount>
with larval and pupal exuviae,
<specimenCount id="9D70FD4DFF82AB1B9D37120D45CD3A18" box="[872,912,552,578]" pageId="110" pageNumber="111" type="female">5 ♀</specimenCount>
each with pupal exuviae,
<materialsCitation id="3B1E3C99FF82AB1B9A97120D42503AB1" collectingDate="2005-02-09" collectionCode="CNCI, INBC" collectorName="Reserva Biologica Alberto Ml. Brenes &amp; Costa Rica" location="Angeles" municipality="Sendero Pajaro Sombrilla" pageId="110" pageNumber="111" specimenCount="14" specimenCount-female="2" specimenCount-larva="7" specimenCount-male="2" specimenCount-pupa="3" stateProvince="Alajuela">
<specimenCount id="9D70FD4DFF82AB1B9A97120D43743A61" box="[1224,1321,552,578]" pageId="110" pageNumber="111" type="pupa">3 pupae</specimenCount>
,
<specimenCount id="9D70FD4DFF82AB1B9B6A120D43CA3A61" box="[1333,1431,552,578]" pageId="110" pageNumber="111" type="larva">7 larvae</specimenCount>
,
<collectingMunicipality id="6BADACBEFF82AB1B9EC81275479B3A49" box="[151,454,592,618]" pageId="110" pageNumber="111">Sendero Pajaro Sombrilla</collectingMunicipality>
,
<collectorName id="26835312FF82AB1B9F8E127545C93A49" box="[465,916,592,618]" pageId="110" pageNumber="111">Reserva Biológica Alberto Ml. Brenes</collectorName>
,
<location id="8EA9601FFF82AB1B9DFF1275425E3A49" LSID="urn:lsid:plazi:treatment:03DF87D2FF80AB1B9EC8159A452D3D91:8EA9601FFF82AB1B9DFF1275425E3A49" box="[928,1027,592,618]" municipality="Sendero Pajaro Sombrilla" name="Angeles" pageId="110" pageNumber="111" stateProvince="Alajuela">Angeles</location>
, San Ramón,
<collectingRegion id="49B2F826FF82AB1B9AFB127543573A49" box="[1188,1290,592,618]" country="Costa Rica" name="Alajuela" pageId="110" pageNumber="111">Alajuela</collectingRegion>
,
<collectorName id="26835312FF82AB1B9B4A127543C53A49" box="[1301,1432,592,618]" pageId="110" pageNumber="111">Costa Rica</collectorName>
,
<date id="FFC81004FF82AB1B9EC8125D475A3AB1" box="[151,263,632,658]" pageId="110" pageNumber="111" value="2005-02-09">
<collectingDate id="EF8CE9ECFF82AB1B9EC8125D475A3AB1" box="[151,263,632,658]" pageId="110" pageNumber="111" value="2005-02-09">9-II-2005</collectingDate>
</date>
(
<specimenCount id="9D70FD4DFF82AB1B9F45125D471E3AA8" box="[282,323,632,658]" pageId="110" pageNumber="111" type="male">2 ♂</specimenCount>
,
<specimenCount id="9D70FD4DFF82AB1B9F0B125D47213AA8" box="[340,380,632,658]" pageId="110" pageNumber="111" type="female">2 ♀</specimenCount>
, each with pupal exuviae,
<collectionCode id="ED67AE01FF82AB1B9C9E125D45563AB1" box="[705,779,632,658]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="110" pageNumber="111">CNCI</collectionCode>
; remainder to
<collectionCode id="ED67AE01FF82AB1B9DE5125D42583AB1" box="[954,1029,632,658]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="110" pageNumber="111">INBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1E3C99FF82AB1B9A43125D456E3A99" collectingDate="2000-02-06" collectingDateMax="2000-02-10" collectingDateMin="2000-02-06" collectionCode="INBC" elevation="600" location="Costa Rica" municipality="Arenal National Park" pageId="110" pageNumber="111" specimenCount="1" specimenCount-female="1" stateProvince="Alajuela">
<specimenCount id="9D70FD4DFF82AB1B9A43125D42193AA8" box="[1052,1092,632,658]" pageId="110" pageNumber="111" type="female">1 ♀</specimenCount>
, Sector la Peninsula,
<collectingMunicipality id="6BADACBEFF82AB1B9B13125D47633A99" pageId="110" pageNumber="111">Arenal National Park</collectingMunicipality>
,
<location id="8EA9601FFF82AB1B9F141285478D3A99" LSID="urn:lsid:plazi:treatment:03DF87D2FF80AB1B9EC8159A452D3D91:8EA9601FFF82AB1B9F141285478D3A99" box="[331,464,672,698]" municipality="Arenal National Park" name="Costa Rica" pageId="110" pageNumber="111" stateProvince="Alajuela">Costa Rica</location>
,
<quantity id="4C8E9B21FF82AB1B9F821285447E3A99" box="[477,547,672,698]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="110" pageNumber="111" unit="m" value="600.0">
<elevation id="005BD1F7FF82AB1B9F821285447E3A99" box="[477,547,672,698]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="110" pageNumber="111" unit="m" value="600.0">600m</elevation>
</quantity>
,
<date id="FFC81004FF82AB1B9C6E128544903A99" box="[561,717,672,698]" pageId="110" pageNumber="111" value="2000-02-06" valueMax="2000-02-10" valueMin="2000-02-06">
<collectingDate id="EF8CE9ECFF82AB1B9C6E128544903A99" box="[561,717,672,698]" pageId="110" pageNumber="111" value="2000-02-06" valueMax="2000-02-10" valueMin="2000-02-06">610-II-2000</collectingDate>
</date>
(
<collectionCode id="ED67AE01FF82AB1B9C81128545773A99" box="[734,810,672,698]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="110" pageNumber="111">INBC</collectionCode>
)
</materialsCitation>
;
<specimenCount id="9D70FD4DFF82AB1B9D1E128545353A90" box="[833,872,672,698]" pageId="110" pageNumber="111" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D70FD4DFF82AB1B9D26128545FC3A90" box="[889,929,672,698]" pageId="110" pageNumber="111" type="female">3 ♀</specimenCount>
, each with pupal exuviae,
<materialsCitation id="3B1E3C99FF82AB1B9ABC128542443AC1" collectionCode="INBC" pageId="110" pageNumber="111" specimenCount="1" specimenCount-female="1" stateProvince="Alajuela" typeStatus="holotype">
<specimenCount id="9D70FD4DFF82AB1B9ABC128543563A90" box="[1251,1291,672,698]" pageId="110" pageNumber="111" type="female">1 ♀</specimenCount>
, with larval and pupal exuviae, 5 fourth instar larvae, labeled as for
<typeStatus id="54CD8866FF82AB1B9D1412ED45EC3AC1" box="[843,945,712,738]" pageId="110" pageNumber="111" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED67AE01FF82AB1B9D9A12ED424C3AC1" box="[965,1041,712,738]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="110" pageNumber="111">INBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1E3C99FF82AB1B9A7512ED452B3B29" collectingDate="2002-06-02" collectingDateMax="2002-07-31" collectingDateMin="2002-06-02" collectionCode="INBC" county="Cerro Rincon" elevation="745" location="Costa Rica" municipality="Corcovado National Park" pageId="110" pageNumber="111" specimenCount="1" specimenCount-male="1" stateProvince="Alajuela" typeStatus="holotype">
<specimenCount id="9D70FD4DFF82AB1B9A7512ED42093AF8" box="[1066,1108,712,738]" pageId="110" pageNumber="111" type="male">1 ♂</specimenCount>
,
<collectingCounty id="62A84E48FF82AB1B9A3912ED43563AC1" box="[1126,1291,712,738]" pageId="110" pageNumber="111">Cerro Rincón</collectingCounty>
,
<collectingMunicipality id="6BADACBEFF82AB1B9B4412ED47613B29" pageId="110" pageNumber="111">Corcovado National Park</collectingMunicipality>
,
<location id="8EA9601FFF82AB1B9F1512D547933B29" LSID="urn:lsid:plazi:treatment:03DF87D2FF80AB1B9EC8159A452D3D91:8EA9601FFF82AB1B9F1512D547933B29" box="[330,462,752,778]" county="Cerro Rincon" municipality="Corcovado National Park" name="Costa Rica" pageId="110" pageNumber="111" stateProvince="Alajuela">Costa Rica</location>
,
<quantity id="4C8E9B21FF82AB1B9F8412D544743B29" box="[475,553,752,778]" metricMagnitude="2" metricUnit="m" metricValue="7.45" pageId="110" pageNumber="111" unit="m" value="745.0">
<elevation id="005BD1F7FF82AB1B9F8412D544743B29" box="[475,553,752,778]" metricMagnitude="2" metricUnit="m" metricValue="7.45" pageId="110" pageNumber="111" unit="m" value="745.0">745 m</elevation>
</quantity>
,
<date id="FFC81004FF82AB1B9C6812D5454C3B29" box="[567,785,752,778]" pageId="110" pageNumber="111" value="2002-06-02" valueMax="2002-07-31" valueMin="2002-06-02">
<collectingDate id="EF8CE9ECFF82AB1B9C6812D5454C3B29" box="[567,785,752,778]" pageId="110" pageNumber="111" value="2002-06-02" valueMax="2002-07-31" valueMin="2002-06-02">2-VI31-VII-2002</collectingDate>
</date>
(
<collectionCode id="ED67AE01FF82AB1B9D7E12D545303B29" box="[801,877,752,778]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="110" pageNumber="111">INBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1E3C99FF82AB1B9DDC12D545DC3B11" collectingDate="2000-11-28" collectingDateMax="2001-02-03" collectingDateMin="2000-11-28" collectionCode="CNCI" county="Cerro Nicuesa" elevation="579" location="Golfito" municipality="Piedras Blancas" pageId="110" pageNumber="111" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="holotype">
<specimenCount id="9D70FD4DFF82AB1B9DDC12D545F63B20" box="[899,939,752,778]" pageId="110" pageNumber="111" type="male">1 ♂</specimenCount>
,
<collectingCounty id="62A84E48FF82AB1B9DE412D5423A3B29" box="[955,1127,752,778]" pageId="110" pageNumber="111">Cerro Nicuesa</collectingCounty>
,
<collectingMunicipality id="6BADACBEFF82AB1B9A2B12D5436E3B29" box="[1140,1331,752,778]" pageId="110" pageNumber="111">Piedras Blancas</collectingMunicipality>
,
<location id="8EA9601FFF82AB1B9B1F12D543C53B29" LSID="urn:lsid:plazi:treatment:03DF87D2FF80AB1B9EC8159A452D3D91:8EA9601FFF82AB1B9B1F12D543C53B29" box="[1344,1432,752,778]" county="Cerro Nicuesa" municipality="Piedras Blancas" name="Golfito" pageId="110" pageNumber="111" stateProvince="Puntarenas">Golfito</location>
,
<collectingRegion id="49B2F826FF82AB1B9EC8133D47463B11" box="[151,283,792,818]" country="Costa Rica" name="Puntarenas" pageId="110" pageNumber="111">Puntarenas</collectingRegion>
,
<location id="8EA9601FFF82AB1B9F78133D47F63B11" LSID="urn:lsid:plazi:treatment:03DF87D2FF80AB1B9EC8159A452D3D91:8EA9601FFF82AB1B9F78133D47F63B11" box="[295,427,792,818]" county="Cerro Nicuesa" municipality="Piedras Blancas" name="Costa Rica" pageId="110" pageNumber="111" stateProvince="Puntarenas">Costa Rica</location>
,
<quantity id="4C8E9B21FF82AB1B9FE7133D445B3B11" box="[440,518,792,818]" metricMagnitude="2" metricUnit="m" metricValue="5.79" pageId="110" pageNumber="111" unit="m" value="579.0">
<elevation id="005BD1F7FF82AB1B9FE7133D445B3B11" box="[440,518,792,818]" metricMagnitude="2" metricUnit="m" metricValue="5.79" pageId="110" pageNumber="111" unit="m" value="579.0">579 m</elevation>
</quantity>
,
<date id="FFC81004FF82AB1B9C4C133D44FD3B11" box="[531,672,792,818]" pageId="110" pageNumber="111" value="2000-11-28">
<collectingDate id="EF8CE9ECFF82AB1B9C4C133D44FD3B11" box="[531,672,792,818]" pageId="110" pageNumber="111" value="2000-11-28">28-XI-2000</collectingDate>
</date>
<date id="FFC81004FF82AB1B9CEF133D45413B11" box="[688,796,792,818]" pageId="110" pageNumber="111" value="2001-02-03">
<collectingDate id="EF8CE9ECFF82AB1B9CEF133D45413B11" box="[688,796,792,818]" pageId="110" pageNumber="111" value="2001-02-03">3-II-2001</collectingDate>
</date>
(
<collectionCode id="ED67AE01FF82AB1B9D72133D45243B11" box="[813,889,792,818]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="110" pageNumber="111">CNCI</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1E3C99FF82AB1B9DD0133D45CA3B79" collectingDate="1996-08-10" collectingDateMax="1996-08-20" collectingDateMin="1996-08-10" collectionCode="INBC" county="Send. Purruja" elevation="300" location="Rio Agujas" municipality="Est. Agujas" pageId="110" pageNumber="111" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="holotype">
<specimenCount id="9D70FD4DFF82AB1B9DD0133D45EB3B08" box="[911,950,792,818]" pageId="110" pageNumber="111" type="female">1 ♀</specimenCount>
,
<collectingCounty id="62A84E48FF82AB1B9D99133D42343B11" box="[966,1129,792,818]" pageId="110" pageNumber="111">Send. Purruja</collectingCounty>
,
<collectingMunicipality id="6BADACBEFF82AB1B9A28133D435F3B11" box="[1143,1282,792,818]" pageId="110" pageNumber="111">Est. Agujas</collectingMunicipality>
,
<location id="8EA9601FFF82AB1B9B50133D43C53B11" LSID="urn:lsid:plazi:treatment:03DF87D2FF80AB1B9EC8159A452D3D91:8EA9601FFF82AB1B9B50133D43C53B11" box="[1295,1432,792,818]" county="Send. Purruja" municipality="Est. Agujas" name="Rio Agujas" pageId="110" pageNumber="111" stateProvince="Puntarenas">Rio Agujas</location>
,
<location id="8EA9601FFF82AB1B9EC8136547353B79" LSID="urn:lsid:plazi:treatment:03DF87D2FF80AB1B9EC8159A452D3D91:8EA9601FFF82AB1B9EC8136547353B79" box="[151,360,832,858]" county="Send. Purruja" municipality="Est. Agujas" name="Peninsula de Osa" pageId="110" pageNumber="111" stateProvince="Puntarenas">Peninsula de Osa</location>
,
<location id="8EA9601FFF82AB1B9F29136547A63B79" LSID="urn:lsid:plazi:treatment:03DF87D2FF80AB1B9EC8159A452D3D91:8EA9601FFF82AB1B9F29136547A63B79" box="[374,507,832,858]" county="Send. Purruja" municipality="Est. Agujas" name="Costa Rica" pageId="110" pageNumber="111" stateProvince="Puntarenas">Costa Rica</location>
,
<quantity id="4C8E9B21FF82AB1B9C55136544053B79" box="[522,600,832,858]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="110" pageNumber="111" unit="m" value="300.0">
<elevation id="005BD1F7FF82AB1B9C55136544053B79" box="[522,600,832,858]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="110" pageNumber="111" unit="m" value="300.0">300 m</elevation>
</quantity>
,
<date id="FFC81004FF82AB1B9C381365456C3B79" box="[615,817,832,858]" pageId="110" pageNumber="111" value="1996-08-10" valueMax="1996-08-20" valueMin="1996-08-10">
<collectingDate id="EF8CE9ECFF82AB1B9C381365456C3B79" box="[615,817,832,858]" pageId="110" pageNumber="111" value="1996-08-10" valueMax="1996-08-20" valueMin="1996-08-10">1020-VIII-1996</collectingDate>
</date>
(
<collectionCode id="ED67AE01FF82AB1B9D1C136545D23B79" box="[835,911,832,858]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="110" pageNumber="111">INBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1E3C99FF82AB1B9DF9136544F43BA1" collectingDate="1999-07-15" collectingDateMax="1999-08-15" collectingDateMin="1999-07-15" collectionCode="INBC" county="Est. Agujas" elevation="300" location="Costa Rica" municipality="Peninsula de Osa" pageId="110" pageNumber="111" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="holotype">
<specimenCount id="9D70FD4DFF82AB1B9DF9136545933B70" box="[934,974,832,858]" pageId="110" pageNumber="111" type="female">1 ♀</specimenCount>
,
<collectingCounty id="62A84E48FF82AB1B9D80136542363B79" box="[991,1131,832,858]" pageId="110" pageNumber="111">Est. Agujas</collectingCounty>
,
<collectingMunicipality id="6BADACBEFF82AB1B9A25136543173B79" box="[1146,1354,832,858]" pageId="110" pageNumber="111">Peninsula de Osa</collectingMunicipality>
,
<location id="8EA9601FFF82AB1B9B061365468D3BA1" LSID="urn:lsid:plazi:treatment:03DF87D2FF80AB1B9EC8159A452D3D91:8EA9601FFF82AB1B9B061365468D3BA1" county="Est. Agujas" municipality="Peninsula de Osa" name="Costa Rica" pageId="110" pageNumber="111" stateProvince="Puntarenas">Costa Rica</location>
,
<quantity id="4C8E9B21FF82AB1B9EBF134D476F3BA1" box="[224,306,872,898]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="110" pageNumber="111" unit="m" value="300.0">
<elevation id="005BD1F7FF82AB1B9EBF134D476F3BA1" box="[224,306,872,898]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="110" pageNumber="111" unit="m" value="300.0">300 m</elevation>
</quantity>
,
<date id="FFC81004FF82AB1B9F1B134D441D3BA1" box="[324,576,872,898]" pageId="110" pageNumber="111" value="1999-07-15" valueMax="1999-08-15" valueMin="1999-07-15">
<collectingDate id="EF8CE9ECFF82AB1B9F1B134D441D3BA1" box="[324,576,872,898]" pageId="110" pageNumber="111" value="1999-07-15" valueMax="1999-08-15" valueMin="1999-07-15">15-VII15-VIII-1999</collectingDate>
</date>
(
<collectionCode id="ED67AE01FF82AB1B9C0A134D44FC3BA1" box="[597,673,872,898]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="110" pageNumber="111">INBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1E3C99FF82AB1B9CE4134D46AE3B89" collectingDate="1999-06-04" collectingDateMax="1999-06-20" collectingDateMin="1999-06-04" collectionCode="CNCI" elevation="300" pageId="110" pageNumber="111" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="holotype">
<specimenCount id="9D70FD4DFF82AB1B9CE4134D44BB3B58" box="[699,742,872,898]" pageId="110" pageNumber="111" type="female">1 ♀</specimenCount>
, as for previous locality but
<quantity id="4C8E9B21FF82AB1B9A09134D42BF3BA1" box="[1110,1250,872,898]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="110" pageNumber="111" unit="m" value="300.0" valueMax="350.0" valueMin="250.0">
<elevation id="005BD1F7FF82AB1B9A09134D42BF3BA1" box="[1110,1250,872,898]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="110" pageNumber="111" unit="m" value="300.0" valueMax="350.0" valueMin="250.0">250350 m</elevation>
</quantity>
,
<date id="FFC81004FF82AB1B9AAB134D43C63BA1" box="[1268,1435,872,898]" pageId="110" pageNumber="111" value="1999-06-04" valueMax="1999-06-20" valueMin="1999-06-04">
<collectingDate id="EF8CE9ECFF82AB1B9AAB134D43C63BA1" box="[1268,1435,872,898]" pageId="110" pageNumber="111" value="1999-06-04" valueMax="1999-06-20" valueMin="1999-06-04">420-VI-1999</collectingDate>
</date>
(
<collectionCode id="ED67AE01FF82AB1B9EC013B546B63B89" box="[159,235,912,938]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="110" pageNumber="111">CNCI</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1E3C99FF82AB1B9F5E13B545BF3B89" box="[257,994,912,938]" collectingDate="1999-07-15" collectingDateMax="1999-08-15" collectingDateMin="1999-07-15" collectionCode="INBC" pageId="110" pageNumber="111" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="holotype">
<specimenCount id="9D70FD4DFF82AB1B9F5E13B547743B80" box="[257,297,912,938]" pageId="110" pageNumber="111" type="female">1 ♀</specimenCount>
, as for previous locality but
<date id="FFC81004FF82AB1B9CDF13B545203B89" box="[640,893,912,938]" pageId="110" pageNumber="111" value="1999-07-15" valueMax="1999-08-15" valueMin="1999-07-15">
<collectingDate id="EF8CE9ECFF82AB1B9CDF13B545203B89" box="[640,893,912,938]" pageId="110" pageNumber="111" value="1999-07-15" valueMax="1999-08-15" valueMin="1999-07-15">15-VII15-VIII-1999</collectingDate>
</date>
(
<collectionCode id="ED67AE01FF82AB1B9DD113B545873B89" box="[910,986,912,938]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="110" pageNumber="111">INBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1E3C99FF82AB1B9DAF13B544C63BF1" collectingDate="1999-05-15" collectingDateMax="1999-06-15" collectingDateMin="1999-05-15" collectionCode="INBC" elevation="600" location="Los Charcos" pageId="110" pageNumber="111" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="holotype">
<specimenCount id="9D70FD4DFF82AB1B9DAF13B5424A3B80" box="[1008,1047,912,938]" pageId="110" pageNumber="111" type="female">1 ♀</specimenCount>
, as for previous locality but
<location id="8EA9601FFF82AB1B9B2F13B546A73BF1" LSID="urn:lsid:plazi:treatment:03DF87D2FF80AB1B9EC8159A452D3D91:8EA9601FFF82AB1B9B2F13B546A73BF1" name="Los Charcos" pageId="110" pageNumber="111" stateProvince="Puntarenas">Los Charcos</location>
,
<quantity id="4C8E9B21FF82AB1B9F59139D47093BF1" box="[262,340,952,978]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="110" pageNumber="111" unit="m" value="600.0">
<elevation id="005BD1F7FF82AB1B9F59139D47093BF1" box="[262,340,952,978]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="110" pageNumber="111" unit="m" value="600.0">600 m</elevation>
</quantity>
,
<date id="FFC81004FF82AB1B9F3E139D446B3BF1" box="[353,566,952,978]" pageId="110" pageNumber="111" value="1999-05-15" valueMax="1999-06-15" valueMin="1999-05-15">
<collectingDate id="EF8CE9ECFF82AB1B9F3E139D446B3BF1" box="[353,566,952,978]" pageId="110" pageNumber="111" value="1999-05-15" valueMax="1999-06-15" valueMin="1999-05-15">15-V15-VI-1999</collectingDate>
</date>
(
<collectionCode id="ED67AE01FF82AB1B9C18139D44CE3BF1" box="[583,659,952,978]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="110" pageNumber="111">INBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1E3C99FF82AB1B9CF7139D47823BD9" collectingDate="2002-07-23" collectingDateMax="2002-07-24" collectingDateMin="2002-07-23" collectionCode="INBC" county="Est. Los Planes" elevation="180" location="Costa Rica" municipality="Corcovado National Park" pageId="110" pageNumber="111" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="holotype">
<specimenCount id="9D70FD4DFF82AB1B9CF7139D44923BE8" box="[680,719,952,978]" pageId="110" pageNumber="111" type="female">1 ♀</specimenCount>
,
<collectingCounty id="62A84E48FF82AB1B9C80139D45CA3BF1" box="[735,919,952,978]" pageId="110" pageNumber="111">Est. Los Planes</collectingCounty>
,
<collectingMunicipality id="6BADACBEFF82AB1B9DFB139D428E3BF1" box="[932,1235,952,978]" pageId="110" pageNumber="111">Corcovado National Park</collectingMunicipality>
,
<location id="8EA9601FFF82AB1B9ABF139D433E3BF1" LSID="urn:lsid:plazi:treatment:03DF87D2FF80AB1B9EC8159A452D3D91:8EA9601FFF82AB1B9ABF139D433E3BF1" box="[1248,1379,952,978]" county="Est. Los Planes" municipality="Corcovado National Park" name="Costa Rica" pageId="110" pageNumber="111" stateProvince="Puntarenas">Costa Rica</location>
,
<quantity id="4C8E9B21FF82AB1B9B2F139D46ED3BD9" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="110" pageNumber="111" unit="m" value="180.0">
<elevation id="005BD1F7FF82AB1B9B2F139D46ED3BD9" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="110" pageNumber="111" unit="m" value="180.0">180 m</elevation>
</quantity>
,
<date id="FFC81004FF82AB1B9EE313C547263BD9" box="[188,379,992,1018]" pageId="110" pageNumber="111" value="2002-07-23" valueMax="2002-07-24" valueMin="2002-07-23">
<collectingDate id="EF8CE9ECFF82AB1B9EE313C547263BD9" box="[188,379,992,1018]" pageId="110" pageNumber="111" value="2002-07-23" valueMax="2002-07-24" valueMin="2002-07-23">2324-VII-2002</collectingDate>
</date>
(
<collectionCode id="ED67AE01FF82AB1B9FD413C5478A3BD9" box="[395,471,992,1018]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="110" pageNumber="111">INBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1E3C99FF82AB1B9FB413C547D33C01" collectingDate="2000-12-26" collectionCode="INBC" county="Send." elevation="100" location="Est. Los Patos" municipality="Sirena" pageId="110" pageNumber="111" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="holotype">
<specimenCount id="9D70FD4DFF82AB1B9FB413C5444D3BD0" box="[491,528,992,1018]" pageId="110" pageNumber="111" type="female">1 ♀</specimenCount>
,
<collectingCounty id="62A84E48FF82AB1B9C4113C5443D3BD9" box="[542,608,992,1018]" pageId="110" pageNumber="111">Send.</collectingCounty>
a
<collectingMunicipality id="6BADACBEFF82AB1B9C2413C544943BD9" box="[635,713,992,1018]" pageId="110" pageNumber="111">Sirena</collectingMunicipality>
,
<location id="8EA9601FFF82AB1B9C8B13C545213BD9" LSID="urn:lsid:plazi:treatment:03DF87D2FF80AB1B9EC8159A452D3D91:8EA9601FFF82AB1B9C8B13C545213BD9" box="[724,892,992,1018]" county="Send." municipality="Sirena" name="Est. Los Patos" pageId="110" pageNumber="111" stateProvince="Puntarenas">Est. Los Patos</location>
,
<location id="8EA9601FFF82AB1B9DD813C542EF3BD9" LSID="urn:lsid:plazi:treatment:03DF87D2FF80AB1B9EC8159A452D3D91:8EA9601FFF82AB1B9DD813C542EF3BD9" box="[903,1202,992,1018]" county="Send." municipality="Sirena" name="Corcovado National Park" pageId="110" pageNumber="111" stateProvince="Puntarenas">Corcovado National Park</location>
,
<location id="8EA9601FFF82AB1B9AE113C5431D3BD9" LSID="urn:lsid:plazi:treatment:03DF87D2FF80AB1B9EC8159A452D3D91:8EA9601FFF82AB1B9AE113C5431D3BD9" box="[1214,1344,992,1018]" county="Send." municipality="Sirena" name="Costa Rica" pageId="110" pageNumber="111" stateProvince="Puntarenas">Costa Rica</location>
,
<quantity id="4C8E9B21FF82AB1B9B1413C543CA3BD9" box="[1355,1431,992,1018]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="110" pageNumber="111" unit="m" value="100.0">
<elevation id="005BD1F7FF82AB1B9B1413C543CA3BD9" box="[1355,1431,992,1018]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="110" pageNumber="111" unit="m" value="100.0">100 m</elevation>
</quantity>
,
<date id="FFC81004FF82AB1B9EC8142D47773C01" box="[151,298,1032,1058]" pageId="110" pageNumber="111" value="2000-12-26">
<collectingDate id="EF8CE9ECFF82AB1B9EC8142D47773C01" box="[151,298,1032,1058]" pageId="110" pageNumber="111" value="2000-12-26">26-XII-2000</collectingDate>
</date>
(
<collectionCode id="ED67AE01FF82AB1B9F65142D47DB3C01" box="[314,390,1032,1058]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="110" pageNumber="111">INBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1E3C99FF82AB1B9FC4142D47943C69" collectingDate="2001-08-10" collectionCode="INBC, CNCI" county="Rincon" location="Sierpe" municipality="Aguabuena" pageId="110" pageNumber="111" specimenCount="4" specimenCount-female="4" stateProvince="Puntarenas" typeStatus="holotype">
<specimenCount id="9D70FD4DFF82AB1B9FC4142D479D3C38" box="[411,448,1032,1058]" pageId="110" pageNumber="111" type="female">4 ♀</specimenCount>
,
<quantity id="4C8E9B21FF82AB1B9F91142D44543C01" box="[462,521,1032,1058]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="110" pageNumber="111" unit="km" value="5.0">5 km</quantity>
W.
<collectingCounty id="62A84E48FF82AB1B9C67142D44D23C01" box="[568,655,1032,1058]" pageId="110" pageNumber="111">Rincón</collectingCounty>
,
<collectingMunicipality id="6BADACBEFF82AB1B9CC4142D457E3C01" box="[667,803,1032,1058]" pageId="110" pageNumber="111">Aguabuena</collectingMunicipality>
,
<location id="8EA9601FFF82AB1B9D71142D45213C01" LSID="urn:lsid:plazi:treatment:03DF87D2FF80AB1B9EC8159A452D3D91:8EA9601FFF82AB1B9D71142D45213C01" box="[814,892,1032,1058]" county="Rincon" municipality="Aguabuena" name="Sierpe" pageId="110" pageNumber="111" stateProvince="Puntarenas">Sierpe</location>
,
<location id="8EA9601FFF82AB1B9DD7142D42C23C01" LSID="urn:lsid:plazi:treatment:03DF87D2FF80AB1B9EC8159A452D3D91:8EA9601FFF82AB1B9DD7142D42C23C01" box="[904,1183,1032,1058]" county="Rincon" municipality="Aguabuena" name="Centro Juvenil Tropical" pageId="110" pageNumber="111" stateProvince="Puntarenas">Centro Juvenil Tropical</location>
,
<location id="8EA9601FFF82AB1B9AF5142D43703C01" LSID="urn:lsid:plazi:treatment:03DF87D2FF80AB1B9EC8159A452D3D91:8EA9601FFF82AB1B9AF5142D43703C01" box="[1194,1325,1032,1058]" county="Rincon" municipality="Aguabuena" name="Costa Rica" pageId="110" pageNumber="111" stateProvince="Puntarenas">Costa Rica</location>
,
<date id="FFC81004FF82AB1B9B67142D468C3C69" pageId="110" pageNumber="111" value="2001-08-10">
<collectingDate id="EF8CE9ECFF82AB1B9B67142D468C3C69" pageId="110" pageNumber="111" value="2001-08-10">10-VIII- 2001</collectingDate>
</date>
(2,
<collectionCode id="ED67AE01FF82AB1B9F5D141547113C69" box="[258,332,1072,1098]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="110" pageNumber="111">INBC</collectionCode>
; 2,
<collectionCode id="ED67AE01FF82AB1B9F291415479C3C69" box="[374,449,1072,1098]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="110" pageNumber="111">CNCI</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1E3C99FF82AB1B9F89141544DC3C51" collectingDate="2000-07-02" collectingDateMax="2000-08-19" collectingDateMin="2000-07-02" collectionCode="INBC" elevation="550" location="Costa Rica" pageId="110" pageNumber="111" specimenCount="1" specimenCount-female="1" stateProvince="Cartago" typeStatus="holotype">
<specimenCount id="9D70FD4DFF82AB1B9F89141547A03C60" box="[470,509,1072,1098]" pageId="110" pageNumber="111" type="female">1 ♀</specimenCount>
,
<locationDeviation id="9A7B2FE6FF82AB1B9C531415455A3C69" box="[524,775,1072,1098]" location="Estacion" pageId="110" pageNumber="111">
<quantity id="4C8E9B21FF82AB1B9C53141544153C69" box="[524,584,1072,1098]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="110" pageNumber="111" unit="km" value="2.0">2 km</quantity>
SE de
<locationDeviation id="9A7B2FE6FF82AB1B9CC01415455A3C69" box="[671,775,1072,1098]" pageId="110" pageNumber="111">Estación</locationDeviation>
</locationDeviation>
,
<locationDeviation id="9A7B2FE6FF82AB1B9D4C1415427D3C69" box="[787,1056,1072,1098]" pageId="110" pageNumber="111">Barbilla National Park</locationDeviation>
,
<locationDeviation id="9A7B2FE6FF82AB1B9A73141542C63C69" box="[1068,1179,1072,1098]" pageId="110" pageNumber="111">Turrialba</locationDeviation>
,
<collectingRegion id="49B2F826FF82AB1B9AF7141543553C69" box="[1192,1288,1072,1098]" country="Costa Rica" name="Cartago" pageId="110" pageNumber="111">Cartago</collectingRegion>
,
<location id="8EA9601FFF82AB1B9B4B141543C53C69" LSID="urn:lsid:plazi:treatment:03DF87D2FF80AB1B9EC8159A452D3D91:8EA9601FFF82AB1B9B4B141543C53C69" box="[1300,1432,1072,1098]" name="Costa Rica" pageId="110" pageNumber="111" stateProvince="Cartago">Costa Rica</location>
,
<quantity id="4C8E9B21FF82AB1B9EC8147D47423C51" box="[151,287,1112,1138]" metricMagnitude="2" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="110" pageNumber="111" unit="m" value="550.0" valueMax="600.0" valueMin="500.0">
<elevation id="005BD1F7FF82AB1B9EC8147D47423C51" box="[151,287,1112,1138]" metricMagnitude="2" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="110" pageNumber="111" unit="m" value="550.0" valueMax="600.0" valueMin="500.0">500600 m</elevation>
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<paragraph id="8BC936C4FF82AB1B9E9914DD452D3D91" blockId="110.[151,1436,152,1458]" pageId="110" pageNumber="111">DERIVATION OF SPECIFIC EPITHET: Douglas A. Craig was on the authors masters committee in 197578 and during that time and since has been a sounding board for my tenuous concepts, provided good advice, and shared a wealth of perspectives and stimulating ideas. It has always been a pleasure to work with him and this species is named to acknowledge his major contribution to my understanding of entomology, as well as recognizing his excellent contributions to Dipterology.</paragraph>
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