281 lines
36 KiB
XML
281 lines
36 KiB
XML
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<mods:title id="672A47C40C27363839C195D5BC6F5F26">The Frog-Biting Midges of the World (Corethrellidae: Diptera)</mods:title>
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<treatment id="03DF87D2FF24ABBC9EC815B243CB3EFC" ID-DOI="http://doi.org/10.5281/zenodo.5126435" ID-Zenodo-Dep="5126435" LSID="urn:lsid:plazi:treatment:03DF87D2FF24ABBC9EC815B243CB3EFC" httpUri="http://treatment.plazi.org/id/03DF87D2FF24ABBC9EC815B243CB3EFC" lastPageId="201" lastPageNumber="202" pageId="200" pageNumber="201">
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<subSubSection id="C36C654FFF24ABBD9EC815B242143D92" box="[151,1097,1431,1457]" pageId="200" pageNumber="201" type="nomenclature">
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<paragraph id="8BC936C4FF24ABBD9EC815B242143D92" blockId="200.[151,1097,1431,1457]" box="[151,1097,1431,1457]" pageId="200" pageNumber="201">
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<heading id="D08181A8FF24ABBD9EC815B242143D92" bold="true" box="[151,1097,1431,1457]" fontSize="11" level="1" pageId="200" pageNumber="201" reason="1">
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<taxonomicName id="4C764D47FF24ABBD9EC815B242143D92" ID-CoL="8BL6P" authority="Szadziewski, Krzeminski and Kutscher" authorityName="Szadziewski, Krzeminski and Kutscher" authorityYear="1994" box="[151,1097,1431,1457]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="200" pageNumber="201" phylum="Arthropoda" rank="species" species="miocaenica" subGenus="Corethrella">
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<emphasis id="B902EAD6FF24ABBD9EC815B242143D92" bold="true" box="[151,1097,1431,1457]" pageId="200" pageNumber="201">
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<emphasis id="B902EAD6FF24ABBD9EC815B2477C3D92" bold="true" box="[151,289,1431,1457]" italics="true" pageId="200" pageNumber="201">Corethrella</emphasis>
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(
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||
<emphasis id="B902EAD6FF24ABBD9F6D15B247E13D92" bold="true" box="[306,444,1431,1457]" italics="true" pageId="200" pageNumber="201">Corethrella</emphasis>
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)
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<emphasis id="B902EAD6FF24ABBD9F9115B244053D92" bold="true" box="[462,600,1431,1457]" italics="true" pageId="200" pageNumber="201">miocaenica</emphasis>
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Szadziewski, Krzeminski and Kutscher
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C36C654FFF24ABBD9EC815C743D43DDA" box="[151,1417,1506,1529]" pageId="200" pageNumber="201" type="reference_group">
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<paragraph id="8BC936C4FF24ABBD9EC815C743D43DDA" blockId="200.[151,1417,1506,1529]" box="[151,1417,1506,1529]" pageId="200" pageNumber="201">
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<treatmentCitationGroup id="AB6611EAFF24ABBD9EC815C743D43DDA" box="[151,1417,1506,1529]" pageId="200" pageNumber="201">
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<treatmentCitation id="0AD710D5FF24ABBD9EC815C745C03DDA" author="Szadziewski, R. & Krzeminski, W. & Kutscher, M." box="[151,925,1506,1529]" page="87" pageId="200" pageNumber="201" year="1994">
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||
<taxonomicName id="4C764D47FF24ABBD9EC815C745C03DDA" ID-CoL="8BL6P" authority="Szadziewski, Krzeminski and Kutscher 1994: 87" authorityName="Szadziewski, Krzeminski and Kutscher" authorityPageNumber="87" authorityYear="1994" box="[151,925,1506,1529]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="200" pageNumber="201" phylum="Arthropoda" rank="species" species="miocaenica">
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<emphasis id="B902EAD6FF24ABBD9EC815C747CE3DDA" box="[151,403,1506,1529]" italics="true" pageId="200" pageNumber="201">Corethrella miocaenica</emphasis>
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<materialsCitation id="3B1E3C99FF24ABBD9FC615C745C03DDA" box="[409,925,1506,1529]" collectionCode="ISZP" country="Germany" location="Germany" pageId="200" pageNumber="201" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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||
<bibRefCitation id="EFE74B35FF24ABBD9FC615C745C03DDA" author="Szadziewski, R. & Krzeminski, W. & Kutscher, M." box="[409,925,1506,1529]" pageId="200" pageNumber="201" pagination="87 - 90" refId="ref243119" refString="Szadziewski, R., Krzeminski, W. & Kutscher, M. (1994) A new species of Corethrella (Diptera, Corethrellidae) from Miocene Saxonian amber. Acta Zoologica Cracoviensia, 37, 87 - 90." type="journal article" year="1994">Szadziewski, Krzeminski and Kutscher 1994:87</bibRefCitation>
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</materialsCitation>
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||
</taxonomicName>
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||
</treatmentCitation>
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||
. Type-locality:
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<collectingCountry id="F3617654FF24ABBD9A1515C742F23DDA" box="[1098,1199,1506,1529]" name="Germany" pageId="200" pageNumber="201">Germany</collectingCountry>
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||
.
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<typeStatus id="54CD8866FF24ABBD9AE615C743413DDA" box="[1209,1308,1506,1529]" pageId="200" pageNumber="201" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D70FD4DFF24ABBD9B7C15C1436E3DD7" box="[1315,1331,1508,1524]" pageId="200" pageNumber="201" type="male">♂</specimenCount>
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(ISZP).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36C654FFF24ABBD9EC8160F45383EB7" pageId="200" pageNumber="201" type="diagnosis">
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<paragraph id="8BC936C4FF24ABBD9EC8160F45383EB7" blockId="200.[151,1436,1578,2009]" pageId="200" pageNumber="201">
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DIAGNOSIS:
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<emphasis id="B902EAD6FF24ABBD9F14160E47973E67" box="[331,458,1579,1604]" italics="true" pageId="200" pageNumber="201">Male adult</emphasis>
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: only fossil species of
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<taxonomicName id="4C764D47FF24ABBD9C8A160E45013E67" box="[725,860,1579,1604]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="200" pageNumber="201" phylum="Arthropoda" rank="genus">
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<emphasis id="B902EAD6FF24ABBD9C8A160E45013E67" box="[725,860,1579,1604]" italics="true" pageId="200" pageNumber="201">Corethrella</emphasis>
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</taxonomicName>
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with elongate, equal fore- and midleg claws with the anterior claw with a basal prong, without empodia (as in Fig. 75C), and a short aedeagus (not extending much beyond base of gonocoxites).
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<emphasis id="B902EAD6FF24ABBD9C1A165E44BD3EB7" box="[581,736,1659,1684]" italics="true" pageId="200" pageNumber="201">Female adult</emphasis>
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: unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36C654FFF24ABBC9E99168744F53A99" lastPageId="201" lastPageNumber="202" pageId="200" pageNumber="201" type="description">
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<paragraph id="8BC936C4FF24ABBC9E99168744253AB1" blockId="200.[151,1436,1578,2009]" lastBlockId="201.[151,1436,151,1759]" lastPageId="201" lastPageNumber="202" pageId="200" pageNumber="201">
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DESCRIPTION:
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<emphasis id="B902EAD6FF24ABBD9FC6168644443E9F" box="[409,537,1699,1724]" italics="true" pageId="200" pageNumber="201">Male adult</emphasis>
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. Habitus as in Fig. 112C. Descriptive statistics: see
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<tableCitation id="C6F4037FFF24ABBD9A201687435F3E9F" box="[1151,1282,1698,1724]" captionStart-0="TABLE 2" captionStart-1="TABLE 3" captionStart-2="TABLE 4" captionStart-3="TABLE 5" captionStartId-0="269.[151,241,151,175]" captionStartId-1="271.[151,241,153,177]" captionStartId-2="273.[151,241,151,175]" captionStartId-3="275.[151,241,151,175]" captionText-0="TABLE 2. Length of wing of males of species of Corethrella. Values are in mm." captionText-1="TABLE 3. Ratio of foreleg tarsomeres 3/4 of males of species of Corethrella." captionText-2="TABLE 4. Ratio of distance of dorsomedial seta from the gonocoxite base/length of the gonocoxite of males of species" captionText-3="TABLE 5. Ratio of distance of subbasal seta from the gonostylus base/ length of the gonostylus of males of species of Corethrella." pageId="200" pageNumber="201">Tables 2–5</tableCitation>
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.
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<emphasis id="B902EAD6FF24ABBD9B4B1687430A3E9F" bold="true" box="[1300,1367,1698,1724]" pageId="200" pageNumber="201">Head</emphasis>
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: Outline in anterior view uncertain, likely nearly circular (as in
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<figureCitation id="134D2A41FF24ABBD9D1316EF45F43EC7" box="[844,937,1738,1764]" captionStart="FIGURE 6" captionStartId="312.[151,255,1947,1971]" captionTargetBox="[177,1410,271,1917]" captionTargetId="figure-28@312.[177,1411,197,1921]" captionTargetPageId="312" captionText="FIGURE 6. Head capsules of female Corethrella, anterior view." pageId="200" pageNumber="201">Fig. 6A</figureCitation>
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). Number of large setae on frons between ventromedial area of ommatida uncertain. Antenna: pedicel without more elongate, stout, dorsal or dorsolateral setae; flagellomeres as in
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<figureCitation id="134D2A41FF24ABBD9FA3173F44343F17" box="[508,617,1818,1844]" captionStart="FIGURE 25" captionStartId="331.[151,255,1947,1971]" captionTargetBox="[187,1396,238,1921]" captionTargetId="figure-23@331.[177,1411,197,1921]" captionTargetPageId="331" captionText="FIGURE 25. Antennae of Corethrella. A-F. Male. G-L. Female." pageId="200" pageNumber="201">Fig. 25D</figureCitation>
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; sensilla coeloconica not visible; flagellomere 13 apex extended with 2 apical sensilla, not bifurcate. Clypeus with single elongate setae, numerous (at least 10) shorter setae. Palpus (
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<figureCitation id="134D2A41FF24ABBD9E8D174F47753FA7" box="[210,296,1898,1924]" captionStart="FIGURE 35" captionStartId="341.[151,255,1947,1971]" captionTargetBox="[177,1375,197,1921]" captionTargetId="figure-21@341.[177,1411,197,1921]" captionTargetPageId="341" captionText="FIGURE 35. Right palpi of female Corethrella." pageId="200" pageNumber="201">Fig. 35</figureCitation>
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AE) with segment 3 slightly swollen apically.
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<emphasis id="B902EAD6FF24ABBD9D3A174F459F3FA7" bold="true" box="[869,962,1898,1924]" pageId="200" pageNumber="201">Thorax</emphasis>
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: Posterior portion of dorsocentral row with 2 elongate setae situated somewhat lateral to one another. Prescutal suture uncertain. Details of shapes of pleura uncertain.
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<emphasis id="B902EAD6FF24ABBD9FD1179F478F3FF7" bold="true" box="[398,466,1978,2004]" pageId="200" pageNumber="201">Wing</emphasis>
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: Apex of R
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<subScript id="17F23481FF24ABBD9C3817EE44323FFA" attach="left" box="[615,623,1995,2009]" fontSize="6" pageId="200" pageNumber="201">2</subScript>
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basal to apex of M
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<subScript id="17F23481FF24ABBD9D0917EE45033FFA" attach="left" box="[854,862,1995,2009]" fontSize="6" pageId="200" pageNumber="201">1</subScript>
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. Veins (other than costa and wing margin) with very slender scales.
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<emphasis id="B902EAD6FF25ABBC9FCA10B247923892" bold="true" box="[405,463,151,177]" pageId="201" pageNumber="202">Legs</emphasis>
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: With only slender setae, lacking scales (except for some in patch of whip-like setae on posterior portion of hind tibia). Midleg without thick, subapical setae on each of at least tarsomeres 1–3. Apices of fore-, midleg fifth tarsomeres undivided, with claws slightly subapical to apical (as in Fig. 75C;
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<figureCitation id="134D2A41FF25ABBC9E89113547753909" box="[214,296,272,298]" captionStart="FIGURE 4" captionStartId="310.[151,255,1947,1971]" captionTargetBox="[181,1408,206,1903]" captionTargetId="figure-73@310.[177,1411,197,1921]" captionTargetPageId="310" captionText="FIGURE 4. A. Unidentified female Corethrella hovering and feeding on calling male of Bufo typhonius in Guyana (taken by Godfrey R. Bourne). B. At least 22 unidentified female Corethrella feeding on the head and back of a calling male of Hyla rosenbergi in Costa Rica (taken by Gernot Kunz (©) reproduced with his permission)." pageId="201" pageNumber="202">Figs. 4</figureCitation>
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,
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<figureCitation id="134D2A41FF25ABBC9F6B1135471E3909" box="[308,323,272,298]" captionStart="FIGURE 5" captionStartId="311.[151,255,1947,1971]" captionTargetBox="[195,1396,221,1913]" captionTargetId="figure-53@311.[177,1411,197,1921]" captionTargetPageId="311" captionText="FIGURE 5. A. Eggs of C. appendiculata floating on water. B. Live pupa of C. pallida in Costa Rica, floating at the water surface (A, part of a colony, taken by Sturgis McKeever; B, taken by L. Guillermo Chaverri)." pageId="201" pageNumber="202">5</figureCitation>
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in
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<bibRefCitation id="EFE74B35FF25ABBC9F38113544233909" author="Szadziewski, R. & Krzeminski, W. & Kutscher, M." box="[359,638,272,298]" pageId="201" pageNumber="202" pagination="87 - 90" refId="ref243119" refString="Szadziewski, R., Krzeminski, W. & Kutscher, M. (1994) A new species of Corethrella (Diptera, Corethrellidae) from Miocene Saxonian amber. Acta Zoologica Cracoviensia, 37, 87 - 90." type="journal article" year="1994">Szadziewski et al. 1994</bibRefCitation>
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). Claw of fore-, midleg long; those of hind leg claws missing. Each claw without inner tooth. Anterior claws of fore-, midleg with basal prong. All claws on given leg equal length. Foreleg third tarsomere longer than fourth tarsomere. Empodia absent.
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<emphasis id="B902EAD6FF25ABBC9A11117A429C395A" bold="true" box="[1102,1217,351,377]" pageId="201" pageNumber="202">Genitalia</emphasis>
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(
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<figureCitation id="134D2A41FF25ABBC9A8C1145431C3959" box="[1235,1345,352,378]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStart-3="FIGURE 9" captionStartId-0="312.[151,255,1947,1971]" captionStartId-1="313.[151,255,1982,2006]" captionStartId-2="314.[151,255,1947,1971]" captionStartId-3="315.[151,255,1947,1971]" captionTargetBox-0="[177,1410,271,1917]" captionTargetBox-1="[177,1408,249,1939]" captionTargetBox-2="[177,1410,267,1904]" captionTargetBox-3="[177,1410,239,1904]" captionTargetId-0="figure-28@312.[177,1411,197,1921]" captionTargetId-1="figure-23@313.[177,1411,232,1956]" captionTargetId-2="figure-28@314.[177,1411,197,1921]" captionTargetId-3="figure-23@315.[177,1411,197,1921]" captionTargetPageId-0="312" captionTargetPageId-1="313" captionTargetPageId-2="314" captionTargetPageId-3="315" captionText-0="FIGURE 6. Head capsules of female Corethrella, anterior view." captionText-1="FIGURE 7. Head capsules of female Corethrella, anterior view." captionText-2="FIGURE 8. Head capsules of female Corethrella, anterior view." captionText-3="FIGURE 9. Head capsules of female Corethrella, anterior view." pageId="201" pageNumber="202">
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Figs.
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<quantity id="4C8E9B21FF25ABBC9B4A1145431C3959" box="[1301,1345,352,378]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.286" metricValueMin="1.524" pageId="201" pageNumber="202" unit="in" value="7.5" valueMax="9.0" valueMin="6.0">6–9</quantity>
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||
</figureCitation>
|
||
in
|
||
<bibRefCitation id="EFE74B35FF25ABBC9B37114547D63981" author="Szadziewski, R. & Krzeminski, W. & Kutscher, M." pageId="201" pageNumber="202" pagination="87 - 90" refId="ref243119" refString="Szadziewski, R., Krzeminski, W. & Kutscher, M. (1994) A new species of Corethrella (Diptera, Corethrellidae) from Miocene Saxonian amber. Acta Zoologica Cracoviensia, 37, 87 - 90." type="journal article" year="1994">Szadziewski et al. 1994</bibRefCitation>
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||
): Gonocoxite gently tapering; anteromedial area with spicules similar in length to those elsewhere on gonocoxite; with well-defined dorsal row of setae of 4 setae, with seta 1 thinner than 3 posteriormost setae, with row curving posteromedially with posteriormost seta of row positioned dorsomedially. Without dorsomedial seta. Gonostylus (Fig. 99C) (in retracted position) straight, thick; expanding distally to truncate apex; one elongate, moderately thick subbasal seta, situated ventrally; with well-developed apical seta, possibly a peg (more clearly seen in other gonostylus drawn by
|
||
<bibRefCitation id="EFE74B35FF25ABBC9D821275434C3A49" author="Szadziewski, R. & Krzeminski, W. & Kutscher, M." box="[989,1297,592,618]" pageId="201" pageNumber="202" pagination="87 - 90" refId="ref243119" refString="Szadziewski, R., Krzeminski, W. & Kutscher, M. (1994) A new species of Corethrella (Diptera, Corethrellidae) from Miocene Saxonian amber. Acta Zoologica Cracoviensia, 37, 87 - 90." type="journal article" year="1994">Szadziewski et al. (1994)</bibRefCitation>
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. Aedeagus short, possibly bilobed, details uncertain.
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</paragraph>
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<paragraph id="8BC936C4FF25ABBC9E99128544F53A99" blockId="201.[151,1436,151,1759]" box="[198,680,672,698]" pageId="201" pageNumber="202">
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||
<emphasis id="B902EAD6FF25ABBC9E991285444A3A9A" box="[198,535,672,697]" italics="true" pageId="201" pageNumber="202">Female adult and immatures</emphasis>
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||
. Unknown.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36C654FFF25ABBC9E9912ED44F63D97" pageId="201" pageNumber="202" type="distribution">
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||
<paragraph id="8BC936C4FF25ABBC9E9912ED459A3B79" blockId="201.[151,1436,151,1759]" pageId="201" pageNumber="202">
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DISTRIBUTION AND BIONOMICS:
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<taxonomicName id="4C764D47FF25ABBC9CF012ED45943AC2" authorityName="Szadziewski, Krzeminski and Kutscher" authorityYear="1994" box="[687,969,712,737]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="species" species="miocaenica">
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<emphasis id="B902EAD6FF25ABBC9CF012ED45943AC2" box="[687,969,712,737]" italics="true" pageId="201" pageNumber="202">Corethrella miocaenica</emphasis>
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</taxonomicName>
|
||
is known from Saxonian amber (Fig. 139), dated at 44.1 Mya. This amber was previously stated to be 22 million years old but has since been discovered to be contemporaneous with Baltic amber (
|
||
<bibRefCitation id="EFE74B35FF25ABBC9CA1133D45E63B11" author="Weitschat, W." box="[766,955,792,818]" pageId="201" pageNumber="202" refId="ref243392" refString="Weitschat, W. (1997) Bitterfelder Bernstein - Ein eozaner Bernstein auf miozaner Lagerstatte." type="book" year="1997">Weitschat 1997</bibRefCitation>
|
||
). The amber piece bearing the
|
||
<typeStatus id="54CD8866FF25ABBC9B69133D43C13B11" box="[1334,1436,792,818]" pageId="201" pageNumber="202" type="holotype">holotype</typeStatus>
|
||
includes some trichomes and the tarsomeres of an unidentified insect.
|
||
</paragraph>
|
||
<paragraph id="8BC936C4FF25ABBC9E99134D470B3C34" blockId="201.[151,1436,151,1759]" pageId="201" pageNumber="202">
|
||
TAXONOMIC DISCUSSION: The Baltic amber specimen identified by
|
||
<bibRefCitation id="EFE74B35FF25ABBC9A7A134D42963BA1" author="Sontag E." box="[1061,1227,872,898]" pageId="201" pageNumber="202" pagination="341 - 342" refId="ref242922" refString="Sontag E. (2001) First record of Corethrellidae (Diptera) in Baltic amber. Polskie Pismo Entomologiczne, 70, 341 - 342." type="journal article" year="2001">Sontag (2001)</bibRefCitation>
|
||
as
|
||
<taxonomicName id="4C764D47FF25ABBC9AAC134D43C13BA2" authorityName="Szadziewski, Krzeminski and Kutscher" authorityYear="1994" box="[1267,1436,872,897]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="species" species="miocaenica">
|
||
<emphasis id="B902EAD6FF25ABBC9AAC134D43C13BA2" box="[1267,1436,872,897]" italics="true" pageId="201" pageNumber="202">C. miocaenica</emphasis>
|
||
</taxonomicName>
|
||
is here described below as
|
||
<taxonomicName id="4C764D47FF25ABBC9F8813A844163B85" authorityName="Borkent" authorityYear="2008" box="[471,587,909,934]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="species" species="baltica">
|
||
<emphasis id="B902EAD6FF25ABBC9F8813A844163B85" box="[471,587,909,934]" italics="true" pageId="201" pageNumber="202">C. baltica</emphasis>
|
||
</taxonomicName>
|
||
. The
|
||
<typeStatus id="54CD8866FF25ABBC9CC613A844A23B84" box="[665,767,909,935]" pageId="201" pageNumber="202" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C764D47FF25ABBC9D7613A8458E3B85" authorityName="Szadziewski, Krzeminski and Kutscher" authorityYear="1994" box="[809,979,909,934]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="species" species="miocaenica">
|
||
<emphasis id="B902EAD6FF25ABBC9D7613A8458E3B85" box="[809,979,909,934]" italics="true" pageId="201" pageNumber="202">C. miocaenica</emphasis>
|
||
</taxonomicName>
|
||
is in excellent condition with only the apices of the right wing, and the tarsomeres of the right foreleg, right midleg and both hind legs missing. The amber with the specimen was further ground and polished for this study, giving a somewhat enhanced view of some structures.
|
||
</paragraph>
|
||
<paragraph id="8BC936C4FF25ABBC9E991407455B3CF1" blockId="201.[151,1436,151,1759]" pageId="201" pageNumber="202">
|
||
<bibRefCitation id="EFE74B35FF25ABBC9E99140747B33C1F" author="Szadziewski, R. & Krzeminski, W. & Kutscher, M." box="[198,494,1058,1084]" pageId="201" pageNumber="202" pagination="87 - 90" refId="ref243119" refString="Szadziewski, R., Krzeminski, W. & Kutscher, M. (1994) A new species of Corethrella (Diptera, Corethrellidae) from Miocene Saxonian amber. Acta Zoologica Cracoviensia, 37, 87 - 90." type="journal article" year="1994">Szadziewski et al. (1994)</bibRefCitation>
|
||
recorded the wing length of the
|
||
<typeStatus id="54CD8866FF25ABBC9D341407458C3C1F" box="[875,977,1058,1084]" pageId="201" pageNumber="202" type="holotype">holotype</typeStatus>
|
||
as
|
||
<quantity id="4C8E9B21FF25ABBC9DA71407423E3C1F" box="[1016,1123,1058,1084]" metricMagnitude="-3" metricUnit="m" metricValue="1.54" pageId="201" pageNumber="202" unit="mm" value="1.54">1.54 mm</quantity>
|
||
, while my measurement is
|
||
<quantity id="4C8E9B21FF25ABBC9EC8146D47583C41" box="[151,261,1096,1122]" metricMagnitude="-3" metricUnit="m" metricValue="1.32" pageId="201" pageNumber="202" unit="mm" value="1.32">1.32 mm</quantity>
|
||
. The difference is due to the measurement here being taken from the arculus, rather than the very base.
|
||
<bibRefCitation id="EFE74B35FF25ABBC9EBF1448445A3CA4" author="Szadziewski, R. & Krzeminski, W. & Kutscher, M." box="[224,519,1133,1159]" pageId="201" pageNumber="202" pagination="87 - 90" refId="ref243119" refString="Szadziewski, R., Krzeminski, W. & Kutscher, M. (1994) A new species of Corethrella (Diptera, Corethrellidae) from Miocene Saxonian amber. Acta Zoologica Cracoviensia, 37, 87 - 90." type="journal article" year="1994">Szadziewski et al. (1994)</bibRefCitation>
|
||
also showed the bases of Sc and R to be fused but this was a misinterpretation based on the position of the wing. In fact, they are separate throughout their length. The subbasal seta on the gonostylus was visible and is therefore redrawn here.
|
||
</paragraph>
|
||
<paragraph id="8BC936C4FF25ABBC9E9914F844F63D97" blockId="201.[151,1436,151,1759]" pageId="201" pageNumber="202">
|
||
Although
|
||
<taxonomicName id="4C764D47FF25ABBC9F6214F847BB3CD5" authorityName="Szadziewski, Krzeminski and Kutscher" authorityYear="1994" box="[317,486,1245,1270]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="species" species="miocaenica">
|
||
<emphasis id="B902EAD6FF25ABBC9F6214F847BB3CD5" box="[317,486,1245,1270]" italics="true" pageId="201" pageNumber="202">C. miocaenica</emphasis>
|
||
</taxonomicName>
|
||
lacks many features of the subgenus
|
||
<taxonomicName id="4C764D47FF25ABBC9DFF14F8427A3CD5" box="[928,1063,1245,1270]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902EAD6FF25ABBC9DFF14F8427A3CD5" box="[928,1063,1245,1270]" italics="true" pageId="201" pageNumber="202">Corethrella</emphasis>
|
||
</taxonomicName>
|
||
, it is placed here because it has derived character states 14, 16' and 17 (male claws lacking teeth, Ta
|
||
<subScript id="17F23481FF25ABBC9D971537458D3D03" attach="left" box="[968,976,1298,1312]" fontSize="6" pageId="201" pageNumber="202">3</subScript>
|
||
/Ta
|
||
<subScript id="17F23481FF25ABBC9DAA153745A03D03" attach="left" box="[1013,1021,1298,1312]" fontSize="6" pageId="201" pageNumber="202">4</subScript>
|
||
of less than 1.60, gonocoxite with dorsal row of setae) (Fig. 143B).
|
||
<bibRefCitation id="EFE74B35FF25ABBC9C71150F45073D67" author="Szadziewski, R. & Krzeminski, W. & Kutscher, M." box="[558,858,1322,1348]" pageId="201" pageNumber="202" pagination="87 - 90" refId="ref243119" refString="Szadziewski, R., Krzeminski, W. & Kutscher, M. (1994) A new species of Corethrella (Diptera, Corethrellidae) from Miocene Saxonian amber. Acta Zoologica Cracoviensia, 37, 87 - 90." type="journal article" year="1994">Szadziewski et al. (1994)</bibRefCitation>
|
||
put this species in the subgenus
|
||
<taxonomicName id="4C764D47FF25ABBC9ABC150E43C13D67" box="[1251,1436,1323,1348]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="subGenus" subGenus="Notocorethrella">
|
||
<emphasis id="B902EAD6FF25ABBC9ABC150E43C13D67" box="[1251,1436,1323,1348]" italics="true" pageId="201" pageNumber="202">Notocorethrella</emphasis>
|
||
</taxonomicName>
|
||
because of its similarity to
|
||
<taxonomicName id="4C764D47FF25ABBC9F8F157544EB3D4A" authorityName="Tonnoir" authorityYear="1927" box="[464,694,1360,1385]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="species" species="novaezealandiae">
|
||
<emphasis id="B902EAD6FF25ABBC9F8F157544EB3D4A" box="[464,694,1360,1385]" italics="true" pageId="201" pageNumber="202">C. novaezealandiae</emphasis>
|
||
</taxonomicName>
|
||
(the only extant member of the subgenus). However, phylogenetic interpretation indicates that
|
||
<taxonomicName id="4C764D47FF25ABBC9C7B155344933DAC" authorityName="Szadziewski, Krzeminski and Kutscher" authorityYear="1994" box="[548,718,1398,1423]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="species" species="miocaenica">
|
||
<emphasis id="B902EAD6FF25ABBC9C7B155344933DAC" box="[548,718,1398,1423]" italics="true" pageId="201" pageNumber="202">C. miocaenica</emphasis>
|
||
</taxonomicName>
|
||
is more closely related to members of the subgenus
|
||
<taxonomicName id="4C764D47FF25ABBC9B1D155346933D97" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902EAD6FF25ABBC9B1D155346933D97" italics="true" pageId="201" pageNumber="202">Corethrella</emphasis>
|
||
</taxonomicName>
|
||
than to
|
||
<taxonomicName id="4C764D47FF25ABBC9F7315BE44493D97" authorityName="Tonnoir" authorityYear="1927" box="[300,532,1435,1460]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="species" species="novaezealandiae">
|
||
<emphasis id="B902EAD6FF25ABBC9F7315BE44493D97" box="[300,532,1435,1460]" italics="true" pageId="201" pageNumber="202">C. novaezealandiae</emphasis>
|
||
</taxonomicName>
|
||
(Fig. 143B).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C654FFF25ABBC9E9915E543CB3EFC" pageId="201" pageNumber="202" type="materials_examined">
|
||
<paragraph id="8BC936C4FF25ABBC9E9915E542653E99" blockId="201.[151,1436,151,1759]" pageId="201" pageNumber="202">
|
||
<materialsCitation id="3B1E3C99FF25ABBC9E9915E542653E99" collectionCode="ISZP" collectorName="Kutscher" county="Acta" location="Evol. of Animals" municipality="Miocene Saxonian" pageId="201" pageNumber="202" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
|
||
MATERIAL EXAMINED:
|
||
<typeStatus id="54CD8866FF25ABBC9C4315E544D13DF9" box="[540,652,1472,1498]" pageId="201" pageNumber="202" type="holotype">Holotype</typeStatus>
|
||
, adult male in amber on microscope slide, labeled “
|
||
<typeStatus id="54CD8866FF25ABBC9AA115E543C13DF9" box="[1278,1436,1472,1498]" pageId="201" pageNumber="202" type="holotype">HOLOTYPE</typeStatus>
|
||
<taxonomicName id="4C764D47FF25ABBC9EC815C345CE3DDC" authority="Szadziewski, Krzeminski and Kutscher" authorityName="Szadziewski, Krzeminski and Kutscher" authorityYear="1994" box="[151,915,1509,1535]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="species" species="miocaenica">
|
||
<emphasis id="B902EAD6FF25ABBC9EC815C347F23DDC" box="[151,431,1510,1535]" italics="true" pageId="201" pageNumber="202">Corethrella miocaenica</emphasis>
|
||
Szadziewski, Krzeminski and Kutscher
|
||
</taxonomicName>
|
||
”, “Saxonian amber coll.
|
||
<collectorName id="26835312FF25ABBC9A9D15C043733DDC" box="[1218,1326,1509,1535]" pageId="201" pageNumber="202">Kutscher</collectorName>
|
||
,
|
||
<taxonomicName id="4C764D47FF25ABBC9B6315C043C43DDC" baseAuthorityName="McAlpine" baseAuthorityYear="1981" box="[1340,1433,1509,1535]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="order">Diptera</taxonomicName>
|
||
:
|
||
<taxonomicName id="4C764D47FF25ABBC9EC8162F471F3E07" authorityName="Wood and Borkent" authorityYear="1986" box="[151,322,1546,1572]" class="Insecta" family="Corethrellidae" kingdom="Animalia" order="Diptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="family">Corethrellidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C764D47FF25ABBC9F11162E44383E07" authority="Published, 2008" authorityName="Published" authorityYear="2008" box="[334,613,1547,1572]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="species" species="miocaenica" status="sp. nov.">
|
||
<emphasis id="B902EAD6FF25ABBC9F11162E44383E07" box="[334,613,1547,1572]" italics="true" pageId="201" pageNumber="202">Corethrella miocaenica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23157ADFF25ABBC9C33162F44F33E07" box="[620,686,1546,1572]" pageId="201" pageNumber="202" rank="species">sp. n.</taxonomicNameLabel>
|
||
,
|
||
<typeStatus id="54CD8866FF25ABBC9CE3162F457F3E07" box="[700,802,1546,1572]" pageId="201" pageNumber="202" type="holotype">holotype</typeStatus>
|
||
<specimenCount id="9D70FD4DFF25ABBC9D74162845673E3E" box="[811,826,1549,1565]" pageId="201" pageNumber="202" type="male">♂</specimenCount>
|
||
”, “Collection:
|
||
<collectorName id="26835312FF25ABBC9DB1162F420A3E07" box="[1006,1111,1546,1572]" pageId="201" pageNumber="202">Kutscher</collectorName>
|
||
coll., Housed in: Inst. Systematic and
|
||
<location id="8EA9601FFF25ABBC9F75161547A43E69" LSID="urn:lsid:plazi:treatment:03DF87D2FF24ABBC9EC815B243CB3EFC:8EA9601FFF25ABBC9F75161547A43E69" box="[298,505,1584,1610]" county="Acta" municipality="Miocene Saxonian" name="Evol. of Animals" pageId="201" pageNumber="202">Evol. of Animals</location>
|
||
, Ordo:
|
||
<taxonomicName id="4C764D47FF25ABBC9C0A161544ED3E69" baseAuthorityName="McAlpine" baseAuthorityYear="1981" box="[597,688,1584,1610]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="order">Diptera</taxonomicName>
|
||
, Family (subfamily):
|
||
<taxonomicName id="4C764D47FF25ABBC9DEC161542003E69" authorityName="Wood and Borkent" authorityYear="1986" box="[947,1117,1584,1610]" class="Insecta" family="Corethrellidae" kingdom="Animalia" order="Diptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="family">Corethrellidae</taxonomicName>
|
||
, Species:
|
||
<taxonomicName id="4C764D47FF25ABBC9A89161545793E4C" authority="Szadziewski, Krzeminski and Kutscher (1994)" authorityName="Szadziewski, Krzeminski and Kutscher" authorityYear="1994" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="species" species="miocaenica">
|
||
<emphasis id="B902EAD6FF25ABBC9A89161546AF3E4C" italics="true" pageId="201" pageNumber="202">Corethrella miocaenica</emphasis>
|
||
<bibRefCitation id="EFE74B35FF25ABBC9EA4167045793E4C" author="Szadziewski, R. & Krzeminski, W. & Kutscher, M." box="[251,804,1621,1647]" pageId="201" pageNumber="202" pagination="87 - 90" refId="ref243119" refString="Szadziewski, R., Krzeminski, W. & Kutscher, M. (1994) A new species of Corethrella (Diptera, Corethrellidae) from Miocene Saxonian amber. Acta Zoologica Cracoviensia, 37, 87 - 90." type="journal article" year="1994">Szadziewski, Krzeminski and Kutscher (1994)</bibRefCitation>
|
||
</taxonomicName>
|
||
, Det.:, Specimen No. MPK 3/1/03 -
|
||
<specimenCount id="9D70FD4DFF25ABBC9A84167D42B73E4B" box="[1243,1258,1624,1640]" pageId="201" pageNumber="202" type="male">♂</specimenCount>
|
||
,
|
||
<typeStatus id="54CD8866FF25ABBC9AA5167043C53E4C" box="[1274,1432,1621,1647]" pageId="201" pageNumber="202" type="holotype">HOLOTYPE</typeStatus>
|
||
, Szadziewski, R., Krzeminski, W. and Kutscher, M. 1994, A new species of
|
||
<taxonomicName id="4C764D47FF25ABBC9A7C165E42F73EB7" box="[1059,1194,1659,1684]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902EAD6FF25ABBC9A7C165E42F73EB7" box="[1059,1194,1659,1684]" italics="true" pageId="201" pageNumber="202">Corethrella</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C764D47FF25ABBC9AE5165F434A3EB7" baseAuthorityName="McAlpine" baseAuthorityYear="1981" box="[1210,1303,1658,1684]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="order">Diptera</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C764D47FF25ABBC9B7B165F468F3E99" authorityName="Wood and Borkent" authorityYear="1986" class="Insecta" family="Corethrellidae" kingdom="Animalia" order="Diptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="family">Corethrellidae</taxonomicName>
|
||
) from
|
||
<collectingMunicipality id="6BADACBEFF25ABBC9F7E168547A13E99" box="[289,508,1696,1722]" pageId="201" pageNumber="202">Miocene Saxonian</collectingMunicipality>
|
||
amber.
|
||
<collectingCounty id="62A84E48FF25ABBC9C3E168544C53E99" box="[609,664,1696,1722]" pageId="201" pageNumber="202">Acta</collectingCounty>
|
||
zool. cracov. 37(2):87–90” (
|
||
<collectionCode id="ED67AE01FF25ABBC9DB6168542763E99" box="[1001,1067,1696,1722]" country="Poland" httpUri="http://grbio.org/cool/hv18-23j2" name="Institute of Systematic Zoology" pageId="201" pageNumber="202">ISZP</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BC936C4FF25ABBC9E9916E043CB3EFC" blockId="201.[151,1436,151,1759]" box="[198,1430,1733,1759]" pageId="201" pageNumber="202">
|
||
DERIVATION OF SPECIFIC EPITHET: The name
|
||
<taxonomicName id="4C764D47FF25ABBC9D6B16E345E63EFC" authorityName="Szadziewski, Krzeminski and Kutscher" authorityYear="1994" box="[820,955,1734,1759]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="species" species="miocaenica">
|
||
<emphasis id="B902EAD6FF25ABBC9D6B16E345E63EFC" box="[820,955,1734,1759]" italics="true" pageId="201" pageNumber="202">miocaenica</emphasis>
|
||
</taxonomicName>
|
||
refers to perceived age of the specimen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |