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<document id="8F117280C0E27B75DC69BFD35E20C3C1" ID-CLB-Dataset="53640" ID-DOI="10.11646/zootaxa.5124.4.3" ID-GBIF-Dataset="5b295616-4831-4d3c-8db2-16f0e24233a5" ID-ISSN="1175-5326" ID-Zenodo-Dep="6413668" ID-ZooBank="C88CB391-5086-4680-A06E-1DB457F629A2" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1649142558394" checkinUser="plazi" docAuthor="Bidzilya, Oleksiy, Blackstein, Hans, Budashkin, Yuriy, Šumpich, Jan &amp; Tsvetkov, Eugene" docDate="2022" docId="03DF1143FFAADC1AFF4FFE5DFC0B42D5" docLanguage="en" docName="zootaxa.5124.4.3.pdf" docOrigin="Zootaxa 5124 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thiodia klapperichi Bidzilya &amp; Blackstein &amp; Budashkin &amp; Šumpich &amp; Tsvetkov 2022, comb. nov." docType="treatment" docVersion="7" lastPageNumber="462" masterDocId="FFE6693BFFAEDC1EFFD8FF87FFBA4402" masterDocTitle="New synonyms and redescription of two little-known Thiodia species (Lepidoptera, Tortricidae): T. ursulana (Kennel, 1919) comb. nov. and T. klapperichi (Razowski, 1967) comb. nov." masterLastPageNumber="470" masterPageNumber="458" pageNumber="462" updateTime="1699319360148" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="13F311F001636BC7BEA07037E6DA087C">New synonyms and redescription of two little-known Thiodia species (Lepidoptera, Tortricidae): T. ursulana (Kennel, 1919) comb. nov. and T. klapperichi (Razowski, 1967) comb. nov.</mods:title>
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<mods:namePart id="E505DB90E8ECF76388B5B2A0B5285799">Bidzilya, Oleksiy</mods:namePart>
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<mods:namePart id="27BCA87864E59C91641DDA7C90DE8CDB">Blackstein, Hans</mods:namePart>
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<mods:namePart id="4918ACA226AC8842B079C432EB538B4C">Budashkin, Yuriy</mods:namePart>
<mods:affiliation id="760AAA80B2386A5BA0E56203A817431A">T. I. Vyazemsky Karadag Scientific Station-Nature Reserve-Branch of FIC “ A. O. Kovalevsky Institute of Biology of the South Seas ” RAS, 298188, Kurortnoye, Pheodosia, Crimea Republic.</mods:affiliation>
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<mods:affiliation id="986D919E86B604995E116C8323A69FC5">National Museum, Department of Entomology, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice, Czech Republic. jansumpich @ seznam. cz; https: // orcid. org / 0000 - 0002 - 0262 - 2941</mods:affiliation>
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<emphasis id="B9027C47FFAADC1AFF4FFE5DFE3C45F7" bold="true" box="[151,390,474,501]" italics="true" pageId="4" pageNumber="462">Thiodia klapperichi</emphasis>
(
<bibRefCitation id="EFE7DDA4FFAADC1AFE4DFE5DFDE045F6" author="Razowski, J." box="[405,602,474,500]" pageId="4" pageNumber="462" pagination="91 - 108" refId="ref7818" refString="Razowski, J. (1967) Afghanische Wickler-Arten (Lepidoptera: Tortricidae und Cochylidae). Beitrage zur naturkundlichen Forschung in Sudwestdeutschland, 26 (3), 91 - 108." type="journal article" year="1967">Razowski, 1967</bibRefCitation>
)
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<figureCitation id="134DBCD0FFAADC1AFF4FFE78FEBC461B" box="[151,262,511,537]" captionStart="FIGURES 613" captionStartId="5.[151,264,1938,1962]" captionTargetBox="[172,1416,181,1914]" captionTargetId="figure-24@5.[172,1416,181,1914]" captionTargetPageId="5" captionText="FIGURES 613. Thiodia klapperichi (Razowski, 1967). 610. Adults. 6. Holotype of E. klapperichi. 7. Holotype of E. buratana. 8. Male, Turkmenistan. 9. Female, Turkmenistan. 10. Fragment of forewing with bulging (indicated by arrow). 11. Female genitalia. 12, 13. Male genitalia. 12. Holotype of E. klapperichi. 13. Holotype of E. buratana." figureDoi="http://doi.org/10.5281/zenodo.6413674" httpUri="https://zenodo.org/record/6413674/files/figure.png" pageId="4" pageNumber="462">Figs 613</figureCitation>
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<emphasis id="B9027C47FFAADC1AFF4FFDC4FEC94659" box="[151,371,579,603]" italics="true" pageId="4" pageNumber="462">Eriopsela klapperichi</emphasis>
<bibRefCitation id="EFE7DDA4FFAADC1AFEA2FDC4FDFC4659" author="Razowski, J." box="[378,582,579,603]" pageId="4" pageNumber="462" pagination="91 - 108" refId="ref7818" refString="Razowski, J. (1967) Afghanische Wickler-Arten (Lepidoptera: Tortricidae und Cochylidae). Beitrage zur naturkundlichen Forschung in Sudwestdeutschland, 26 (3), 91 - 108." type="journal article" year="1967">Razowski, 1967: 95</bibRefCitation>
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.
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<emphasis id="B9027C47FFAADC1AFF4FFDE4FF124678" box="[151,168,611,634]" italics="true" pageId="4" pageNumber="462">=</emphasis>
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<emphasis id="B9027C47FFAADC1AFF77FDE4FED54678" box="[175,367,610,634]" italics="true" pageId="4" pageNumber="462">Eucosma buratana</emphasis>
<bibRefCitation id="EFE7DDA4FFAADC1AFEADFDE5FDF34678" author="Blackstein, H." box="[373,585,610,634]" pageId="4" pageNumber="462" pagination="65 - 69" refId="ref6344" refString="Blackstein, H. (2014) Olethreutinae aus der Sammlung Shchetkin des Museums fur Naturkunde Berlin (Lepidoptera, Tortricidae). Entomologische Nachrichten und Berichte, 58 (1 - 2), 65 - 69." type="journal article" year="2014">Blackstein, 2014: 66</bibRefCitation>
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<taxonomicNameLabel id="A231C13CFFAADC1AFD8BFDE5FD0A4678" box="[595,688,610,634]" pageId="4" pageNumber="462">syn. nov.</taxonomicNameLabel>
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<emphasis id="B9027C47FFAADC1AFF4FFD2FFEFC46C0" bold="true" box="[151,326,680,706]" pageId="4" pageNumber="462">Type material.</emphasis>
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<emphasis id="B9027C47FFAADC1AFE06FD2FFD6946C0" box="[478,723,680,706]" italics="true" pageId="4" pageNumber="462">Eriopsela klapperichi</emphasis>
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, labelled: “
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, Pagmangebirge,
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, 28.8.53, O.
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” | “
<typeStatus id="54CD1EF7FFAADC1AFEABFD4BFE5B46E4" box="[371,481,716,742]" pageId="4" pageNumber="462" type="holotype">Holotype</typeStatus>
” | “G.S.-5663
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,
<taxonomicName id="4C76DBD6FFAADC1AFD72FD4BFC5A46E5" authority="Raz." authorityName="Razowski" authorityYear="1967" box="[682,992,716,743]" class="Insecta" family="Tortricidae" genus="Eriopsela" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="462" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="B9027C47FFAADC1AFD72FD4BFC1B46E4" box="[682,929,716,742]" italics="true" pageId="4" pageNumber="462">Eriopsela klapperichi</emphasis>
Raz.
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, det.
<determinerName id="7E154C91FFAADC1AFBF9FD4AFB1446E4" box="[1057,1198,716,743]" pageId="4" pageNumber="462">J. Razowski</determinerName>
” | Prep. Nr. 4882
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” (
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).
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of
<taxonomicName id="4C76DBD6FFAADC1AFE4DFD76FDD34708" authorityName="Blackstein" authorityYear="2014" box="[405,617,752,778]" class="Insecta" family="Tortricidae" genus="Eucosma" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="462" phylum="Arthropoda" rank="species" species="buratana">
<emphasis id="B9027C47FFAADC1AFE4DFD76FDD34708" box="[405,617,752,778]" italics="true" pageId="4" pageNumber="462">Eucosma buratana</emphasis>
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,
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, labelled: “
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7.x.
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г., ТадЖикистан у подноЖЬя г. Буратау, на свет, Ю. Щеткин” [
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,
<collectingCountry id="F361E0C5FFAADC1AFE77FC93FD9A472C" box="[431,544,788,814]" name="Tajikistan" pageId="4" pageNumber="462">Tajikistan</collectingCountry>
, near the foot of
<location id="8EA9F68EFFAADC1AFD06FC92FCDE472D" LSID="urn:lsid:plazi:treatment:03DF1143FFAADC1AFF4FFE5DFC0B42D5:8EA9F68EFFAADC1AFD06FC92FCDE472D" box="[734,868,789,815]" country="Afghanistan" name="Buratau mt." pageId="4" pageNumber="462">Buratau mt.</location>
, on light,
<collectorName id="2683C583FFAADC1AFC0BFC92FBCC472C" box="[979,1142,787,815]" pageId="4" pageNumber="462">Yu. Shchetkin</collectorName>
”] (
<collectionCode id="ED673890FFAADC1AFB4DFC93FB68472D" box="[1173,1234,788,815]" country="Germany" httpUri="http://grbio.org/cool/jp8y-i5nv" name="Museum für Naturkunde" pageId="4" pageNumber="462" type="Museum">MfN</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BC9A055FFAADC1AFF4FFCBFFB0347BD" blockId="4.[151,1437,680,1751]" pageId="4" pageNumber="462">
<emphasis id="B9027C47FFAADC1AFF4FFCBFFDA64750" bold="true" box="[151,540,824,850]" pageId="4" pageNumber="462">Additional material examined.</emphasis>
<materialsCitation id="3B1EAA08FFAADC1AFDF7FCBEFA994750" ID-GBIF-Occurrence="3746894302" box="[559,1315,823,851]" collectingDate="1961-09-28" country="Turkmenistan" location="Kara-Kala" municipality="Ashhabad Province" pageId="4" pageNumber="462" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D706BDCFFAADC1AFDF7FCBEFDFC4751" box="[559,582,825,851]" pageId="4" pageNumber="462" type="male"></specimenCount>
, [
<collectingCountry id="F361E0C5FFAADC1AFDBCFCB0FCBF4750" box="[612,773,823,850]" name="Turkmenistan" pageId="4" pageNumber="462">Turkmenistan</collectingCountry>
]
<location id="8EA9F68EFFAADC1AFCC6FCBFFC284750" LSID="urn:lsid:plazi:treatment:03DF1143FFAADC1AFF4FFE5DFC0B42D5:8EA9F68EFFAADC1AFCC6FCBFFC284750" box="[798,914,824,850]" country="Turkmenistan" municipality="Ashhabad Province" name="Kara-Kala" pageId="4" pageNumber="462">Каrа-Каlа</location>
,
<collectingMunicipality id="6BAD3A2FFFAADC1AFC7FFCBFFB2A4750" box="[935,1168,824,850]" pageId="4" pageNumber="462">Ashhabad Province</collectingMunicipality>
,
<date id="FFC88695FFAADC1AFB70FCBFFA994750" box="[1192,1315,824,850]" pageId="4" pageNumber="462" value="1961-09-28">
<collectingDate id="EF8C7F7DFFAADC1AFB70FCBFFA994750" box="[1192,1315,824,850]" pageId="4" pageNumber="462" value="1961-09-28">28.ix.1961</collectingDate>
</date>
</materialsCitation>
, on light (Krasilnikova);
<materialsCitation id="3B1EAA08FFAADC1AFE95FCDAFCD84774" ID-GBIF-Occurrence="3746894306" box="[333,866,860,887]" collectingDate="1961-09-29" country="Turkmenistan" location="Kara-Kala" municipality="Ashhabad Province" pageId="4" pageNumber="462" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D706BDCFFAADC1AFE95FCDAFEDE4775" box="[333,356,861,887]" pageId="4" pageNumber="462" type="male"></specimenCount>
, same data as for preceding but
<date id="FFC88695FFAADC1AFD3FFCDBFCD84774" box="[743,866,860,886]" pageId="4" pageNumber="462" value="1961-09-29">
<collectingDate id="EF8C7F7DFFAADC1AFD3FFCDBFCD84774" box="[743,866,860,886]" pageId="4" pageNumber="462" value="1961-09-29">29.ix.1961</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1EAA08FFAADC1AFCAAFCDAFACB4775" ID-GBIF-Occurrence="3746894311" box="[882,1393,860,887]" collectingDate="1961-10-05" country="Turkmenistan" location="Kara-Kala" municipality="Ashhabad Province" pageId="4" pageNumber="462" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D706BDCFFAADC1AFCAAFCDAFC3D4775" box="[882,903,861,887]" pageId="4" pageNumber="462" type="female"></specimenCount>
, same data as for preceding but
<date id="FFC88695FFAADC1AFAD4FCDBFACB4775" box="[1292,1393,860,887]" pageId="4" pageNumber="462" value="1961-10-05">
<collectingDate id="EF8C7F7DFFAADC1AFAD4FCDBFACB4775" box="[1292,1393,860,887]" pageId="4" pageNumber="462" value="1961-10-05">5.x.1961</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B1EAA08FFAADC1AFA58FCDAFBA94799" ID-GBIF-Occurrence="3746894307" collectorName="Krasilnikova" country="Turkmenistan" location="Phiryuza" municipality="Ashhabad Province" pageId="4" pageNumber="462" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D706BDCFFAADC1AFA58FCDAFA2D4775" box="[1408,1431,861,887]" pageId="4" pageNumber="462" type="male"></specimenCount>
,
<location id="8EA9F68EFFAADC1AFF4FFC07FF414798" LSID="urn:lsid:plazi:treatment:03DF1143FFAADC1AFF4FFE5DFC0B42D5:8EA9F68EFFAADC1AFF4FFC07FF414798" box="[151,251,896,922]" country="Turkmenistan" municipality="Ashhabad Province" name="Phiryuza" pageId="4" pageNumber="462">Phiryuza</location>
env.,
<collectingMunicipality id="6BAD3A2FFFAADC1AFEE0FC07FDA94798" box="[312,531,896,922]" pageId="4" pageNumber="462">Ashhabad Province</collectingMunicipality>
, [no data].ix.[no year], on light (
<collectorName id="2683C583FFAADC1AFCACFC07FBB64799" box="[884,1036,896,923]" pageId="4" pageNumber="462">Krasilnikova</collectorName>
)
</materialsCitation>
;
<materialsCitation id="3B1EAA08FFAADC1AFBC4FC06FEBD47BD" ID-GBIF-Occurrence="3746894312" collectingDate="1980-10-03" collectorName="Pechen'" country="Turkmenistan" location="Badkhyz" municipality="Kyzyldzhar" pageId="4" pageNumber="462" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D706BDCFFAADC1AFBC4FC06FB894799" box="[1052,1075,897,923]" pageId="4" pageNumber="462" type="male"></specimenCount>
,
<location id="8EA9F68EFFAADC1AFBE5FC07FB194798" LSID="urn:lsid:plazi:treatment:03DF1143FFAADC1AFF4FFE5DFC0B42D5:8EA9F68EFFAADC1AFBE5FC07FB194798" box="[1085,1187,896,922]" country="Turkmenistan" municipality="Kyzyldzhar" name="Badkhyz" pageId="4" pageNumber="462">Badkhyz</location>
,
<collectingMunicipality id="6BAD3A2FFFAADC1AFB75FC07FA8B4798" box="[1197,1329,896,922]" pageId="4" pageNumber="462">Kyzyldzhar</collectingMunicipality>
,
<date id="FFC88695FFAADC1AFAE2FC07FA264798" box="[1338,1436,896,922]" pageId="4" pageNumber="462" value="1980-10-03">
<collectingDate id="EF8C7F7DFFAADC1AFAE2FC07FA264798" box="[1338,1436,896,922]" pageId="4" pageNumber="462" value="1980-10-03">3.x.1980</collectingDate>
</date>
(
<collectorName id="2683C583FFAADC1AFF47FC23FF4547BD" box="[159,255,932,959]" pageId="4" pageNumber="462">Pechen</collectorName>
)
</materialsCitation>
;
<materialsCitation id="3B1EAA08FFAADC1AFECBFC22FBFD47BD" ID-GBIF-Occurrence="3746894309" box="[275,1095,932,959]" collectingDate="1953-10-07" collectorName="Shchetkin" country="Tajikistan" location="Buratau mt." pageId="4" pageNumber="462" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D706BDCFFAADC1AFECBFC22FE9047BD" box="[275,298,933,959]" pageId="4" pageNumber="462" type="male"></specimenCount>
,
<collectingCountry id="F361E0C5FFAADC1AFEEDFC23FE1C47BC" box="[309,422,932,958]" name="Tajikistan" pageId="4" pageNumber="462">Tajikistan</collectingCountry>
, near the foot of
<location id="8EA9F68EFFAADC1AFDBCFC22FD5147BD" LSID="urn:lsid:plazi:treatment:03DF1143FFAADC1AFF4FFE5DFC0B42D5:8EA9F68EFFAADC1AFDBCFC22FD5147BD" box="[612,747,933,959]" country="Tajikistan" name="Buratau mt." pageId="4" pageNumber="462">Buratau mt.</location>
, on light,
<date id="FFC88695FFAADC1AFC83FC23FC0747BD" box="[859,957,932,959]" pageId="4" pageNumber="462" value="1953-10-07">
<collectingDate id="EF8C7F7DFFAADC1AFC83FC23FC0747BD" box="[859,957,932,959]" pageId="4" pageNumber="462" value="1953-10-07">7.x.1953</collectingDate>
</date>
(
<collectorName id="2683C583FFAADC1AFC13FC23FB8547BD" box="[971,1087,932,959]" pageId="4" pageNumber="462">Shchetkin</collectorName>
)
</materialsCitation>
(all
<collectionCode id="ED673890FFAADC1AFBA2FC22FB1747BD" box="[1146,1197,933,959]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="462" type="Museum">ZIN</collectionCode>
).
</paragraph>
<paragraph id="8BC9A055FFAADC1AFF1FFC4FFE5C4148" blockId="4.[151,1437,680,1751]" pageId="4" pageNumber="462">
<taxonomicName id="4C76DBD6FFAADC1AFF1FFC4FFD3947E0" authority="Razowski, 1967" authorityName="Razowski" authorityYear="1967" box="[199,643,968,994]" class="Insecta" family="Tortricidae" genus="Eriopsela" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="462" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="B9027C47FFAADC1AFF1FFC4FFE7A47E0" box="[199,448,968,994]" italics="true" pageId="4" pageNumber="462">Eriopsela klapperichi</emphasis>
<bibRefCitation id="EFE7DDA4FFAADC1AFE13FC4FFD3947E0" author="Razowski, J." box="[459,643,968,994]" pageId="4" pageNumber="462" pagination="91 - 108" refId="ref7818" refString="Razowski, J. (1967) Afghanische Wickler-Arten (Lepidoptera: Tortricidae und Cochylidae). Beitrage zur naturkundlichen Forschung in Sudwestdeutschland, 26 (3), 91 - 108." type="journal article" year="1967">Razowski, 1967</bibRefCitation>
</taxonomicName>
was described from the male
<typeStatus id="54CD1EF7FFAADC1AFC36FC4FFBEA47E0" box="[1006,1104,968,994]" pageId="4" pageNumber="462" type="holotype">holotype</typeStatus>
from Pagman Mountains in
<collectingCountry id="F361E0C5FFAADC1AFF4FFC6BFE9E4004" box="[151,292,1004,1030]" name="Afghanistan" pageId="4" pageNumber="462">Afghanistan</collectingCountry>
. The specimen is in rather poor condition (
<figureCitation id="134DBCD0FFAADC1AFCDBFC6BFCFD4005" box="[771,839,1004,1031]" captionStart="FIGURES 613" captionStartId="5.[151,264,1938,1962]" captionTargetBox="[172,1416,181,1914]" captionTargetId="figure-24@5.[172,1416,181,1914]" captionTargetPageId="5" captionText="FIGURES 613. Thiodia klapperichi (Razowski, 1967). 610. Adults. 6. Holotype of E. klapperichi. 7. Holotype of E. buratana. 8. Male, Turkmenistan. 9. Female, Turkmenistan. 10. Fragment of forewing with bulging (indicated by arrow). 11. Female genitalia. 12, 13. Male genitalia. 12. Holotype of E. klapperichi. 13. Holotype of E. buratana." figureDoi="http://doi.org/10.5281/zenodo.6413674" httpUri="https://zenodo.org/record/6413674/files/figure.png" pageId="4" pageNumber="462">Fig. 6</figureCitation>
), however the genitalia are characteristic by having a deep sub-ovate emargination on the ventral margin of the valva (
<figureCitation id="134DBCD0FFAADC1AFCAFFB97FC714028" box="[887,971,1040,1066]" captionStart="FIGURES 613" captionStartId="5.[151,264,1938,1962]" captionTargetBox="[172,1416,181,1914]" captionTargetId="figure-24@5.[172,1416,181,1914]" captionTargetPageId="5" captionText="FIGURES 613. Thiodia klapperichi (Razowski, 1967). 610. Adults. 6. Holotype of E. klapperichi. 7. Holotype of E. buratana. 8. Male, Turkmenistan. 9. Female, Turkmenistan. 10. Fragment of forewing with bulging (indicated by arrow). 11. Female genitalia. 12, 13. Male genitalia. 12. Holotype of E. klapperichi. 13. Holotype of E. buratana." figureDoi="http://doi.org/10.5281/zenodo.6413674" httpUri="https://zenodo.org/record/6413674/files/figure.png" pageId="4" pageNumber="462">Fig. 12</figureCitation>
).
<taxonomicName id="4C76DBD6FFAADC1AFC39FB96FB0D4028" authorityName="Blackstein" authorityYear="2014" box="[993,1207,1040,1066]" class="Insecta" family="Tortricidae" genus="Eucosma" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="462" phylum="Arthropoda" rank="species" species="buratana">
<emphasis id="B9027C47FFAADC1AFC39FB96FB0D4028" box="[993,1207,1040,1066]" italics="true" pageId="4" pageNumber="462">Eucosma buratana</emphasis>
</taxonomicName>
was described from
<specimenCount id="9D706BDCFFAADC1AFF4FFBB2FEB1404C" box="[151,267,1076,1103]" pageId="4" pageNumber="462" type="male">two males</specimenCount>
collected near the foot of Buratau Mountain in
<collectingCountry id="F361E0C5FFAADC1AFCEDFBB3FC7D404C" box="[821,967,1076,1102]" name="Tajikistan" pageId="4" pageNumber="462">Tadzhikistan</collectingCountry>
. The photographs of wings and the male genitalia in the original description (
<bibRefCitation id="EFE7DDA4FFAADC1AFDF5FBDFFD5D4070" author="Blackstein, H." box="[557,743,1112,1138]" pageId="4" pageNumber="462" pagination="65 - 69" refId="ref6344" refString="Blackstein, H. (2014) Olethreutinae aus der Sammlung Shchetkin des Museums fur Naturkunde Berlin (Lepidoptera, Tortricidae). Entomologische Nachrichten und Berichte, 58 (1 - 2), 65 - 69." type="journal article" year="2014">Blackstein 2014</bibRefCitation>
, figs 2, 3) fully agree with the
<typeStatus id="54CD1EF7FFAADC1AFBE5FBDFFB254070" box="[1085,1183,1112,1138]" pageId="4" pageNumber="462" type="holotype">holotype</typeStatus>
of
<emphasis id="B9027C47FFAADC1AFB1CFBDEFAD04070" box="[1220,1386,1112,1138]" italics="true" pageId="4" pageNumber="462">
<taxonomicName id="4C76DBD6FFAADC1AFB1CFBDEFADC4070" authorityName="Razowski" authorityYear="1967" box="[1220,1382,1112,1138]" class="Insecta" family="Tortricidae" genus="Eriopsela" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="462" phylum="Arthropoda" rank="species" species="klapperichi">E. klapperichi</taxonomicName>
.
</emphasis>
Our study of additional material (
<specimenCount id="9D706BDCFFAADC1AFE04FBFBFDE84094" box="[476,594,1148,1174]" pageId="4" pageNumber="462" type="male">five males</specimenCount>
and
<specimenCount id="9D706BDCFFAADC1AFD52FBFAFCBC4094" box="[650,774,1148,1175]" pageId="4" pageNumber="462" type="female">one female</specimenCount>
from
<collectingCountry id="F361E0C5FFAADC1AFC93FBFBFC5C4094" box="[843,998,1148,1174]" name="Turkmenistan" pageId="4" pageNumber="462">Turkmenistan</collectingCountry>
and
<specimenCount id="9D706BDCFFAADC1AFBC4FBFAFB384094" box="[1052,1154,1148,1175]" pageId="4" pageNumber="462" type="male">one male</specimenCount>
from the type locality of
<taxonomicName id="4C76DBD6FFAADC1AFF4FFB26FEA640B8" authorityName="Blackstein" authorityYear="2014" box="[151,284,1184,1210]" class="Insecta" family="Tortricidae" genus="Eucosma" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="462" phylum="Arthropoda" rank="species" species="buratana">
<emphasis id="B9027C47FFAADC1AFF4FFB26FEA640B8" box="[151,284,1184,1210]" italics="true" pageId="4" pageNumber="462">E. buratana</emphasis>
</taxonomicName>
in
<collectingCountry id="F361E0C5FFAADC1AFEE5FB27FE1740B9" box="[317,429,1184,1211]" name="Tajikistan" pageId="4" pageNumber="462">Tajikistan</collectingCountry>
) showed the presence of small rounded bloating on the base of the male forewing (
<figureCitation id="134DBCD0FFAADC1AFAE5FB27FA3540B8" box="[1341,1423,1184,1210]" captionStart="FIGURES 613" captionStartId="5.[151,264,1938,1962]" captionTargetBox="[172,1416,181,1914]" captionTargetId="figure-24@5.[172,1416,181,1914]" captionTargetPageId="5" captionText="FIGURES 613. Thiodia klapperichi (Razowski, 1967). 610. Adults. 6. Holotype of E. klapperichi. 7. Holotype of E. buratana. 8. Male, Turkmenistan. 9. Female, Turkmenistan. 10. Fragment of forewing with bulging (indicated by arrow). 11. Female genitalia. 12, 13. Male genitalia. 12. Holotype of E. klapperichi. 13. Holotype of E. buratana." figureDoi="http://doi.org/10.5281/zenodo.6413674" httpUri="https://zenodo.org/record/6413674/files/figure.png" pageId="4" pageNumber="462">Fig. 10</figureCitation>
). This character is an autapomorphy of the genus
<taxonomicName id="4C76DBD6FFAADC1AFD6BFB43FBA340DC" authority="(Kuznetzov 2001: 440)" baseAuthorityName="Kuznetzov" baseAuthorityPageNumber="440" baseAuthorityYear="2001" box="[691,1049,1220,1246]" class="Insecta" family="Tortricidae" genus="Thiodia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="462" phylum="Arthropoda" rank="genus">
<emphasis id="B9027C47FFAADC1AFD6BFB43FCB040DC" box="[691,778,1220,1246]" italics="true" pageId="4" pageNumber="462">Thiodia</emphasis>
(
<bibRefCitation id="EFE7DDA4FFAADC1AFCC1FB43FBAB40DC" author="Kuznetzov, V. I." box="[793,1041,1220,1246]" pageId="4" pageNumber="462" pagination="11 - 472" refId="ref7323" refString="Kuznetzov, V. I. (2001) 48. Sem. Tortricidae (Olethreutidae, Cochylidae) - listovyortki. In: Ponomarenko, M. G. (Ed.), Key to the Insects of Russian Far East. Vol. 5. Trichoptera and Lepidoptera, Part 3. Vladivostok, Dal'nauka, pp. 11 - 472. [in Russian]" type="book chapter" year="2001">Kuznetzov 2001: 440</bibRefCitation>
)
</taxonomicName>
. Taking into consideration that the genitalia of both sexes also agree in general with
<taxonomicName id="4C76DBD6FFAADC1AFD60FB6FFCB54100" baseAuthorityName="Kuznetzov" baseAuthorityYear="2001" box="[696,783,1256,1282]" class="Insecta" family="Tortricidae" genus="Thiodia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="462" phylum="Arthropoda" rank="genus">
<emphasis id="B9027C47FFAADC1AFD60FB6FFCB54100" box="[696,783,1256,1282]" italics="true" pageId="4" pageNumber="462">Thiodia</emphasis>
</taxonomicName>
(see remarks in Discussion), we transfer the species to this genus and we propose the new synonymy:
<taxonomicName id="4C76DBD6FFAADC1AFD5AFA8AFB9A4124" authority="Blackstein, 2014" authorityName="Blackstein" authorityYear="2014" box="[642,1056,1292,1318]" class="Insecta" family="Tortricidae" genus="Eucosma" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="462" phylum="Arthropoda" rank="species" species="buratana">
<emphasis id="B9027C47FFAADC1AFD5AFA8AFCE24124" box="[642,856,1292,1318]" italics="true" pageId="4" pageNumber="462">Eucosma buratana</emphasis>
<bibRefCitation id="EFE7DDA4FFAADC1AFCB8FA8BFB9A4124" author="Blackstein, H." box="[864,1056,1292,1318]" pageId="4" pageNumber="462" pagination="65 - 69" refId="ref6344" refString="Blackstein, H. (2014) Olethreutinae aus der Sammlung Shchetkin des Museums fur Naturkunde Berlin (Lepidoptera, Tortricidae). Entomologische Nachrichten und Berichte, 58 (1 - 2), 65 - 69." type="journal article" year="2014">Blackstein, 2014</bibRefCitation>
</taxonomicName>
,
<emphasis id="B9027C47FFAADC1AFBF5FA8BFB2F4124" bold="true" box="[1069,1173,1292,1318]" pageId="4" pageNumber="462">
<taxonomicNameLabel id="A231C13CFFAADC1AFBF5FA8BFB2F4124" box="[1069,1173,1292,1318]" pageId="4" pageNumber="462">syn. nov.</taxonomicNameLabel>
</emphasis>
of
<taxonomicName id="4C76DBD6FFAADC1AFB65FA8BFEE44148" authority="(Razowski, 1967)" authorityName="Bidzilya &amp; Blackstein &amp; Budashkin &amp; Šumpich &amp; Tsvetkov" authorityYear="2022" baseAuthorityName="Razowski" baseAuthorityYear="1967" class="Insecta" family="Tortricidae" genus="Thiodia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="462" phylum="Arthropoda" rank="species" species="klapperichi" status="comb. nov.">
<emphasis id="B9027C47FFAADC1AFB65FA8BFA274124" box="[1213,1437,1292,1318]" italics="true" pageId="4" pageNumber="462">Thiodia klapperichi</emphasis>
(
<bibRefCitation id="EFE7DDA4FFAADC1AFF47FAB7FEEF4148" author="Razowski, J." box="[159,341,1328,1354]" pageId="4" pageNumber="462" pagination="91 - 108" refId="ref7818" refString="Razowski, J. (1967) Afghanische Wickler-Arten (Lepidoptera: Tortricidae und Cochylidae). Beitrage zur naturkundlichen Forschung in Sudwestdeutschland, 26 (3), 91 - 108." type="journal article" year="1967">Razowski, 1967</bibRefCitation>
<emphasis id="B9027C47FFAADC1AFE8DFAB7FEE44148" bold="true" box="[341,350,1328,1354]" pageId="4" pageNumber="462">)</emphasis>
</taxonomicName>
<emphasis id="B9027C47FFAADC1AFEBDFAB7FE5C4148" bold="true" box="[357,486,1328,1354]" pageId="4" pageNumber="462">
<taxonomicNameLabel id="A231C13CFFAADC1AFEBDFAB7FE5C4148" box="[357,486,1328,1354]" pageId="4" pageNumber="462" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BC9A055FFAADC1AFF1FFAD3FF484190" blockId="4.[151,1437,680,1751]" pageId="4" pageNumber="462">
Below we provide a description of the female genitalia of
<taxonomicName id="4C76DBD6FFAADC1AFC99FAD2FC61416C" authorityName="Bidzilya &amp; Blackstein &amp; Budashkin &amp; Šumpich &amp; Tsvetkov" authorityYear="2022" baseAuthorityName="Razowski" baseAuthorityYear="1967" box="[833,987,1364,1390]" class="Insecta" family="Tortricidae" genus="Thiodia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="462" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="B9027C47FFAADC1AFC99FAD2FC61416C" box="[833,987,1364,1390]" italics="true" pageId="4" pageNumber="462">T. klapperichi</emphasis>
</taxonomicName>
based on a specimen from
<collectingCountry id="F361E0C5FFAADC1AFADAFAD3FA27416C" box="[1282,1437,1364,1390]" name="Turkmenistan" pageId="4" pageNumber="462">Turkmenistan</collectingCountry>
(
<figureCitation id="134DBCD0FFAADC1AFF78FAFFFF5E4190" box="[160,228,1400,1426]" captionStart="FIGURES 613" captionStartId="5.[151,264,1938,1962]" captionTargetBox="[172,1416,181,1914]" captionTargetId="figure-24@5.[172,1416,181,1914]" captionTargetPageId="5" captionText="FIGURES 613. Thiodia klapperichi (Razowski, 1967). 610. Adults. 6. Holotype of E. klapperichi. 7. Holotype of E. buratana. 8. Male, Turkmenistan. 9. Female, Turkmenistan. 10. Fragment of forewing with bulging (indicated by arrow). 11. Female genitalia. 12, 13. Male genitalia. 12. Holotype of E. klapperichi. 13. Holotype of E. buratana." figureDoi="http://doi.org/10.5281/zenodo.6413674" httpUri="https://zenodo.org/record/6413674/files/figure.png" pageId="4" pageNumber="462">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="8BC9A055FFAADC1AFF1FFA1BFA2142B0" blockId="4.[151,1437,680,1751]" pageId="4" pageNumber="462">
<emphasis id="B9027C47FFAADC1AFF1FFA1BFDB841B4" bold="true" box="[199,514,1436,1462]" pageId="4" pageNumber="462">
Female genitalia (
<figureCitation id="134DBCD0FFAADC1AFE46FA1BFE4941B4" box="[414,499,1436,1462]" captionStart="FIGURES 613" captionStartId="5.[151,264,1938,1962]" captionTargetBox="[172,1416,181,1914]" captionTargetId="figure-24@5.[172,1416,181,1914]" captionTargetPageId="5" captionText="FIGURES 613. Thiodia klapperichi (Razowski, 1967). 610. Adults. 6. Holotype of E. klapperichi. 7. Holotype of E. buratana. 8. Male, Turkmenistan. 9. Female, Turkmenistan. 10. Fragment of forewing with bulging (indicated by arrow). 11. Female genitalia. 12, 13. Male genitalia. 12. Holotype of E. klapperichi. 13. Holotype of E. buratana." figureDoi="http://doi.org/10.5281/zenodo.6413674" httpUri="https://zenodo.org/record/6413674/files/figure.png" pageId="4" pageNumber="462">Fig. 11</figureCitation>
).
</emphasis>
Papillae anales elongate, membranous, relatively small, covered by fine bristles. Posterior apophyses longer than anterior apophyses, strongly broadened in posterior 1/2, narrowing in middle, very thin in anterior 1/2. Anterior apophyses well sclerotised, slightly bent at their bases. Antrum cup-shaped. Ductus bursae membranous, cingulum long, distinctly sclerotised; long anterior portion of ductus rolled (shown ventrolaterally on fig. 11, so as bursa copulatrix), entirely enclosing the corpus bursae in its original state. Sternum VII broad and short, anterior margin straight, widely rounded on the sides; vane-like, sclerotised areas located posteriorly on the sides, covered by modified bristles (very small, oval). Corpus bursa spherical, membranous, with two horn-like signa located posteriorly opposite to each other; one of these signa very small and hardly noticeable.
</paragraph>
</subSubSection>
<subSubSection id="C36CF3DEFFAADC1AFF1FF93BFC0B42D5" box="[199,945,1724,1751]" pageId="4" pageNumber="462" type="distribution">
<paragraph id="8BC9A055FFAADC1AFF1FF93BFC0B42D5" blockId="4.[151,1437,680,1751]" box="[199,945,1724,1751]" pageId="4" pageNumber="462">
<emphasis id="B9027C47FFAADC1AFF1FF93BFEDB42D4" bold="true" box="[199,353,1724,1750]" pageId="4" pageNumber="462">Distribution.</emphasis>
<collectingCountry id="F361E0C5FFAADC1AFEBEF93BFE4842D4" box="[358,498,1724,1750]" name="Afghanistan" pageId="4" pageNumber="462">Afghanistan</collectingCountry>
,
<collectingCountry id="F361E0C5FFAADC1AFE24F93BFDD742D4" box="[508,621,1724,1750]" name="Tajikistan" pageId="4" pageNumber="462">Tajikistan</collectingCountry>
,
<collectingCountry id="F361E0C5FFAADC1AFDA0F93BFCA942D4" box="[632,787,1724,1750]" name="Turkmenistan" pageId="4" pageNumber="462">Turkmenistan</collectingCountry>
(new record).
</paragraph>
</subSubSection>
</treatment>
</document>