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<document id="1AB15B5DE4EBBF225E36D1DC9E061FBD" ID-DOI="10.11646/zootaxa.3955.4.3" ID-GBIF-Dataset="ac08f47a-d395-406a-8461-b1a65d65fc38" ID-ISSN="1175-5326" ID-Zenodo-Dep="244417" ID-ZooBank="B4E21D99-203B-4818-904E-BBA145E5CD33" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461052425946" checkinUser="plazi" docAuthor="Silva, Fernando A. B. &amp; Vaz-De-Mello, Fernando" docDate="2015" docId="03D8893BB172CC352390F9054FE9FD9C" docLanguage="en" docName="zt03955p520.pdf" docOrigin="Zootaxa 3955 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aphengium Harold" docType="key" docVersion="8" lastPageNumber="508" masterDocId="FFE1F143B170CC362307FFA14A72FFA5" masterDocTitle="A revision of the genus Aphengium Harold, 1868 (Coleoptera: Scarabaeidae: Scarabaeinae: Ateuchini)" masterLastPageNumber="520" masterPageNumber="505" pageNumber="507" updateTime="1698579553482" updateUser="plazi">
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<mods:title id="DF8F9D3C55AA5C3EDAD333459AC80512">A revision of the genus Aphengium Harold, 1868 (Coleoptera: Scarabaeidae: Scarabaeinae: Ateuchini)</mods:title>
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<treatment id="03D8893BB172CC352390F9054FE9FD9C" ID-DOI="http://doi.org/10.5281/zenodo.6122144" ID-GBIF-Taxon="119612284" ID-Zenodo-Dep="6122144" LSID="urn:lsid:plazi:treatment:03D8893BB172CC352390F9054FE9FD9C" httpUri="http://treatment.plazi.org/id/03D8893BB172CC352390F9054FE9FD9C" lastPageId="3" lastPageNumber="508" pageId="2" pageNumber="507">
<subSubSection id="C36B6BA6B172CC342390F9054825F91B" box="[151,599,1700,1726]" pageId="2" pageNumber="507" type="nomenclature">
<paragraph id="8BCE382DB172CC342390F9054825F91B" blockId="2.[151,599,1700,1726]" box="[151,599,1700,1726]" pageId="2" pageNumber="507">
<heading id="D0868F41B172CC342390F9054825F91B" bold="true" box="[151,599,1700,1726]" fontSize="11" level="1" pageId="2" pageNumber="507" reason="1">
<emphasis id="B905E43FB172CC342390F9054825F91B" bold="true" box="[151,599,1700,1726]" pageId="2" pageNumber="507">
Key to males of
<taxonomicName id="4C7143AEB172CC342263F9054825F91B" ID-CoL="62D9J" authority="Harold" authorityName="Harold" box="[356,599,1700,1726]" class="Insecta" family="Scarabaeidae" genus="Aphengium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="507" phylum="Arthropoda" rank="genus">
<emphasis id="B905E43FB172CC342263F9054B87F91B" bold="true" box="[356,501,1700,1726]" italics="true" pageId="2" pageNumber="507">Aphengium</emphasis>
Harold
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36B6BA6B172CC352390F94A4FE9FD9C" lastPageId="3" lastPageNumber="508" pageId="2" pageNumber="507" type="key">
<keyStep id="30852088B172CC342390F94A4FEEF8C8" pageId="2" pageNumber="507">
<paragraph id="8BCE382DB172CC342390F94A4FE9F880" blockId="2.[151,1436,1771,1901]" pageId="2" pageNumber="507">
<keyLead id="30809B18B172CC342390F94A4FE9F880" pageId="2" pageNumber="507">
1. Dorsal surface of head with dense foveiform punctures deeply impressed and often rounded (
<figureCitation id="134A24A8B172CC342770F94A4ECFF8A4" box="[1143,1213,1771,1793]" captionStart="FIGURES 7 12" captionStartId="8.[151,264,1954,1976]" captionTargetBox="[171,1409,207,1927]" captionTargetId="figure@8.[165,1422,193,1931]" captionTargetPageId="8" captionText="FIGURES 7 12. Aphengium sordidum Harold; 7. Male (dorsal view); 8. Female (dorsal view); 9. Male head (arrow shows foveiform punctures along midline); 10. Female head (arrow shows foveiform punctures along midline); 11. Male pronotum (arrow shows punctures on central portion); 12. Female pronotum (arrow shows punctures on central portion)." httpUri="https://zenodo.org/record/244419/files/figure.png" pageId="2" pageNumber="507">Figs. 9</figureCitation>
,
<figureCitation id="134A24A8B172CC3427CFF94A4E90F8A4" box="[1224,1250,1771,1793]" captionStart="FIGURES 13 18" captionStartId="9.[151,264,1955,1977]" captionTargetBox="[183,1402,209,1920]" captionTargetId="figure@9.[164,1422,193,1934]" captionTargetPageId="9" captionText="FIGURES 13 18. Aphengium cupreum Shipp; 13. Male (dorsal view); 14. Female (dorsal view); 15. Male head; 16. Female head; 17. Male pronotum; 18. Female pronotum." httpUri="https://zenodo.org/record/244420/files/figure.png" pageId="2" pageNumber="507">15</figureCitation>
,
<figureCitation id="134A24A8B172CC3427EAF94A4F7AF8A4" box="[1261,1288,1771,1793]" captionStart="FIGURES 19 24" captionStartId="10.[151,264,1947,1969]" captionTargetBox="[180,1405,198,1922]" captionTargetId="figure@10.[173,1414,186,1923]" captionTargetPageId="10" captionText="FIGURES 19 24. Aphengium ibateguara sp. nov.; 19. Male (dorsal view); 20. Female (dorsal view); 21. Male head; 22. Female head; 23. Male pronotum; 24. Female pronotum." httpUri="https://zenodo.org/record/244421/files/figure.png" pageId="2" pageNumber="507">21</figureCitation>
). Anterolateral portion of pronotum lacking sleek callus............................................................... 2
</keyLead>
</paragraph>
<paragraph id="8BCE382DB172CC342390F8954FEEF8C8" blockId="2.[151,1436,1771,1901]" pageId="2" pageNumber="507">
<keyLead id="30809B18B172CC342390F8954FEEF8C8" pageId="2" pageNumber="507">
- Dorsal surface of head smooth and sleek or punctures almost inconspicuous (
<figureCitation id="134A24A8B172CC3420EEF8924E47F8EC" box="[1001,1077,1843,1865]" captionStart="FIGURES 25 30" captionStartId="11.[151,264,1936,1958]" captionTargetBox="[171,1414,195,1924]" captionTargetId="figure@11.[162,1424,186,1925]" captionTargetPageId="11" captionText="FIGURES 25 30. Aphengium curtum sp. nov.; 25. Male (dorsal view); 26. Female (dorsal view); 27. Male head; 28. Female head; 29. Male pronotum (arrow shows anterolateral portion of pronotum with a longitudinal and sleek callus); 30. Female pronotum." httpUri="https://zenodo.org/record/244422/files/figure.png" pageId="2" pageNumber="507">Fig. 27</figureCitation>
). Anterolateral portion of pronotum with a longitudinal and sleek callus (
<figureCitation id="134A24A8B172CC34214DF8F648E6F8C8" box="[586,660,1879,1901]" captionStart="FIGURES 25 30" captionStartId="11.[151,264,1936,1958]" captionTargetBox="[171,1414,195,1924]" captionTargetId="figure@11.[162,1424,186,1925]" captionTargetPageId="11" captionText="FIGURES 25 30. Aphengium curtum sp. nov.; 25. Male (dorsal view); 26. Female (dorsal view); 27. Male head; 28. Female head; 29. Male pronotum (arrow shows anterolateral portion of pronotum with a longitudinal and sleek callus); 30. Female pronotum." httpUri="https://zenodo.org/record/244422/files/figure.png" pageId="2" pageNumber="507">Fig. 29</figureCitation>
). Male genitalia figs. 34, 40.
<collectingCountry id="F36678BDB172CC3420B4F8F6499DF8C8" box="[947,1007,1879,1901]" name="Brazil" pageId="2" pageNumber="507">Brazil</collectingCountry>
(Bahia).....
<taxonomicName id="4C7143AEB172CC342784F8F64E96F8C9" ID-CoL="FD4D" box="[1155,1252,1879,1900]" class="Insecta" family="Scarabaeidae" genus="Aphengium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="507" phylum="Arthropoda" rank="species" species="curtum" status="sp. nov.">
<emphasis id="B905E43FB172CC342784F8F64E96F8C9" box="[1155,1252,1879,1900]" italics="true" pageId="2" pageNumber="507">A. curtum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2365944B172CC3427ECF8F64F4BF8C8" box="[1259,1337,1879,1901]" pageId="2" pageNumber="507" rank="species">
<emphasis id="B905E43FB172CC3427ECF8F64F4BF8C8" bold="true" box="[1259,1337,1879,1901]" pageId="2" pageNumber="507">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="134A24A8B172CC342641F8F64FE2F8C8" box="[1350,1424,1879,1901]" captionStart="FIGURES 25 30" captionStartId="11.[151,264,1936,1958]" captionTargetBox="[171,1414,195,1924]" captionTargetId="figure@11.[162,1424,186,1925]" captionTargetPageId="11" captionText="FIGURES 25 30. Aphengium curtum sp. nov.; 25. Male (dorsal view); 26. Female (dorsal view); 27. Male head; 28. Female head; 29. Male pronotum (arrow shows anterolateral portion of pronotum with a longitudinal and sleek callus); 30. Female pronotum." httpUri="https://zenodo.org/record/244422/files/figure.png" pageId="2" pageNumber="507">Fig. 25</figureCitation>
).
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30852088B173CC352390FF364FE9FE98" pageId="3" pageNumber="508">
<paragraph id="8BCE382DB173CC352390FF364FE7FF74" blockId="3.[151,1436,151,569]" pageId="3" pageNumber="508">
<keyLead id="30809B18B173CC352390FF364FE7FF74" pageId="3" pageNumber="508">
2. Dorsal surface of head with contiguous foveiform punctures over its central portion (
<figureCitation id="134A24A8B173CC352736FF364E0AFF08" box="[1073,1144,151,173]" captionStart="FIGURES 7 12" captionStartId="8.[151,264,1954,1976]" captionTargetBox="[171,1409,207,1927]" captionTargetId="figure@8.[165,1422,193,1931]" captionTargetPageId="8" captionText="FIGURES 7 12. Aphengium sordidum Harold; 7. Male (dorsal view); 8. Female (dorsal view); 9. Male head (arrow shows foveiform punctures along midline); 10. Female head (arrow shows foveiform punctures along midline); 11. Male pronotum (arrow shows punctures on central portion); 12. Female pronotum (arrow shows punctures on central portion)." httpUri="https://zenodo.org/record/244419/files/figure.png" pageId="3" pageNumber="508">Figs. 9</figureCitation>
,
<figureCitation id="134A24A8B173CC352784FF364EECFF08" box="[1155,1182,151,173]" captionStart="FIGURES 19 24" captionStartId="10.[151,264,1947,1969]" captionTargetBox="[180,1405,198,1922]" captionTargetId="figure@10.[173,1414,186,1923]" captionTargetPageId="10" captionText="FIGURES 19 24. Aphengium ibateguara sp. nov.; 19. Male (dorsal view); 20. Female (dorsal view); 21. Male head; 22. Female head; 23. Male pronotum; 24. Female pronotum." httpUri="https://zenodo.org/record/244421/files/figure.png" pageId="3" pageNumber="508">21</figureCitation>
). Anterolateral margin of pronotum explanate (
<figureCitation id="134A24A8B173CC3522B6FF1A4870FF74" box="[433,514,187,209]" captionStart="FIGURES 7 12" captionStartId="8.[151,264,1954,1976]" captionTargetBox="[171,1409,207,1927]" captionTargetId="figure@8.[165,1422,193,1931]" captionTargetPageId="8" captionText="FIGURES 7 12. Aphengium sordidum Harold; 7. Male (dorsal view); 8. Female (dorsal view); 9. Male head (arrow shows foveiform punctures along midline); 10. Female head (arrow shows foveiform punctures along midline); 11. Male pronotum (arrow shows punctures on central portion); 12. Female pronotum (arrow shows punctures on central portion)." httpUri="https://zenodo.org/record/244419/files/figure.png" pageId="3" pageNumber="508">Figs. 11</figureCitation>
,
<figureCitation id="134A24A8B173CC35210AFF1A4854FF74" box="[525,550,187,209]" captionStart="FIGURES 19 24" captionStartId="10.[151,264,1947,1969]" captionTargetBox="[180,1405,198,1922]" captionTargetId="figure@10.[173,1414,186,1923]" captionTargetPageId="10" captionText="FIGURES 19 24. Aphengium ibateguara sp. nov.; 19. Male (dorsal view); 20. Female (dorsal view); 21. Male head; 22. Female head; 23. Male pronotum; 24. Female pronotum." httpUri="https://zenodo.org/record/244421/files/figure.png" pageId="3" pageNumber="508">23</figureCitation>
).................................................................... 3
</keyLead>
</paragraph>
<paragraph id="8BCE382DB173CC352390FF7E4FE9FE98" blockId="3.[151,1436,151,569]" pageId="3" pageNumber="508">
<keyLead id="30809B18B173CC352390FF7E4FE9FE98" pageId="3" pageNumber="508">
- Dorsal surface of head with foveiform punctures separated by distance equivalent to at least two times puncture diameter along most of midline (
<figureCitation id="134A24A8B173CC3522CEFEA24861FEBC" box="[457,531,259,281]" captionStart="FIGURES 13 18" captionStartId="9.[151,264,1955,1977]" captionTargetBox="[183,1402,209,1920]" captionTargetId="figure@9.[164,1422,193,1934]" captionTargetPageId="9" captionText="FIGURES 13 18. Aphengium cupreum Shipp; 13. Male (dorsal view); 14. Female (dorsal view); 15. Male head; 16. Female head; 17. Male pronotum; 18. Female pronotum." httpUri="https://zenodo.org/record/244420/files/figure.png" pageId="3" pageNumber="508">Fig. 15</figureCitation>
). Anterolateral margin of pronotum not explanate (
<figureCitation id="134A24A8B173CC352709FEA24E2AFEBC" box="[1038,1112,259,281]" captionStart="FIGURES 13 18" captionStartId="9.[151,264,1955,1977]" captionTargetBox="[183,1402,209,1920]" captionTargetId="figure@9.[164,1422,193,1934]" captionTargetPageId="9" captionText="FIGURES 13 18. Aphengium cupreum Shipp; 13. Male (dorsal view); 14. Female (dorsal view); 15. Male head; 16. Female head; 17. Male pronotum; 18. Female pronotum." httpUri="https://zenodo.org/record/244420/files/figure.png" pageId="3" pageNumber="508">Fig. 17</figureCitation>
). Male genitalia figs. 32, 36, 38.
<collectingCountry id="F36678BDB173CC3523E6FE864B6FFE98" box="[225,285,295,317]" name="Brazil" pageId="3" pageNumber="508">Brazil</collectingCountry>
(Espírito Santo and Minas Gerais)..........................................
<taxonomicName id="4C7143AEB173CC35278EFE864F4AFE98" ID-CoL="FD4C" authority="Shipp" authorityName="Shipp" box="[1161,1336,295,317]" class="Insecta" family="Scarabaeidae" genus="Aphengium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="cupreum">
<emphasis id="B905E43FB173CC35278EFE864E8AFE99" box="[1161,1272,295,316]" italics="true" pageId="3" pageNumber="508">A. cupreum</emphasis>
Shipp
</taxonomicName>
(
<figureCitation id="134A24A8B173CC352641FE864FFDFE98" box="[1350,1423,295,317]" captionStart="FIGURES 13 18" captionStartId="9.[151,264,1955,1977]" captionTargetBox="[183,1402,209,1920]" captionTargetId="figure@9.[164,1422,193,1934]" captionTargetPageId="9" captionText="FIGURES 13 18. Aphengium cupreum Shipp; 13. Male (dorsal view); 14. Female (dorsal view); 15. Male head; 16. Female head; 17. Male pronotum; 18. Female pronotum." httpUri="https://zenodo.org/record/244420/files/figure.png" pageId="3" pageNumber="508">Fig. 13</figureCitation>
).
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30852088B173CC352390FEEA4FE9FD9C" pageId="3" pageNumber="508">
<paragraph id="8BCE382DB173CC352390FEEA4FE7FE0D" blockId="3.[151,1436,151,569]" pageId="3" pageNumber="508">
<keyLead id="30809B18B173CC352390FEEA4FE7FE0D" pageId="3" pageNumber="508">
3. Pronotum with larger and contiguous foveiform punctures on lateral and posterior portions (
<figureCitation id="134A24A8B173CC352774FEEA4ECEFEC4" box="[1139,1212,331,353]" captionStart="FIGURES 7 12" captionStartId="8.[151,264,1954,1976]" captionTargetBox="[171,1409,207,1927]" captionTargetId="figure@8.[165,1422,193,1931]" captionTargetPageId="8" captionText="FIGURES 7 12. Aphengium sordidum Harold; 7. Male (dorsal view); 8. Female (dorsal view); 9. Male head (arrow shows foveiform punctures along midline); 10. Female head (arrow shows foveiform punctures along midline); 11. Male pronotum (arrow shows punctures on central portion); 12. Female pronotum (arrow shows punctures on central portion)." httpUri="https://zenodo.org/record/244419/files/figure.png" pageId="3" pageNumber="508">Fig. 11</figureCitation>
). Length of parameres one-half that of phallobase (
<figureCitation id="134A24A8B173CC3522F2FECE484CFE20" box="[501,574,367,389]" captionStart="FIGURES 31 34" captionStartId="12.[151,264,1136,1158]" captionTargetBox="[165,1436,193,1114]" captionTargetId="figure@12.[150,1436,193,1115]" captionTargetPageId="12" captionText="FIGURES 31 34. Aedeagus of Aphengium; 31. A. sordidum (arrow shows dorsal margin of parameres curved inward at its apical third); 32. A. cupreum; 33. A. ibateguara sp. nov.; 34. A. curtum sp. nov." httpUri="https://zenodo.org/record/244423/files/figure.png" pageId="3" pageNumber="508">Fig. 31</figureCitation>
); apical third of parameres slightly narrowed (
<figureCitation id="134A24A8B173CC352701FECE4E3DFE20" box="[1030,1103,367,389]" captionStart="FIGURES 31 34" captionStartId="12.[151,264,1136,1158]" captionTargetBox="[165,1436,193,1114]" captionTargetId="figure@12.[150,1436,193,1115]" captionTargetPageId="12" captionText="FIGURES 31 34. Aedeagus of Aphengium; 31. A. sordidum (arrow shows dorsal margin of parameres curved inward at its apical third); 32. A. cupreum; 33. A. ibateguara sp. nov.; 34. A. curtum sp. nov." httpUri="https://zenodo.org/record/244423/files/figure.png" pageId="3" pageNumber="508">Fig. 31</figureCitation>
); fronto-lateral peripheral sclerite (FLP) as in figure 37.
<collectingCountry id="F36678BDB173CC3522BCFE324B85FE0C" box="[443,503,403,425]" name="Brazil" pageId="3" pageNumber="508">Brazil</collectingCountry>
(Rio de Janeiro and São Paulo)..........................
<taxonomicName id="4C7143AEB173CC35277AFE324F4DFE0C" ID-CoL="FD4G" authority="Harold" authorityName="Harold" box="[1149,1343,403,425]" class="Insecta" family="Scarabaeidae" genus="Aphengium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="sordidum">
<emphasis id="B905E43FB173CC35277AFE324E86FE0D" box="[1149,1268,403,424]" italics="true" pageId="3" pageNumber="508">A. sordidum</emphasis>
Harold
</taxonomicName>
(
<figureCitation id="134A24A8B173CC35264BFE324FFAFE0D" box="[1356,1416,403,425]" captionStart="FIGURES 7 12" captionStartId="8.[151,264,1954,1976]" captionTargetBox="[171,1409,207,1927]" captionTargetId="figure@8.[165,1422,193,1931]" captionTargetPageId="8" captionText="FIGURES 7 12. Aphengium sordidum Harold; 7. Male (dorsal view); 8. Female (dorsal view); 9. Male head (arrow shows foveiform punctures along midline); 10. Female head (arrow shows foveiform punctures along midline); 11. Male pronotum (arrow shows punctures on central portion); 12. Female pronotum (arrow shows punctures on central portion)." httpUri="https://zenodo.org/record/244419/files/figure.png" pageId="3" pageNumber="508">Fig. 7</figureCitation>
).
</keyLead>
</paragraph>
<paragraph id="8BCE382DB173CC352390FE164FE9FD9C" blockId="3.[151,1436,151,569]" pageId="3" pageNumber="508">
<keyLead id="30809B18B173CC352390FE164FE9FD9C" pageId="3" pageNumber="508">
- Foveiform punctures of lateral and posterior portions of pronotum not contiguous, usually separated by distance equivalent to less than one puncture diameter (
<figureCitation id="134A24A8B173CC35213CFE7A48F7FE54" box="[571,645,475,497]" captionStart="FIGURES 19 24" captionStartId="10.[151,264,1947,1969]" captionTargetBox="[180,1405,198,1922]" captionTargetId="figure@10.[173,1414,186,1923]" captionTargetPageId="10" captionText="FIGURES 19 24. Aphengium ibateguara sp. nov.; 19. Male (dorsal view); 20. Female (dorsal view); 21. Male head; 22. Female head; 23. Male pronotum; 24. Female pronotum." httpUri="https://zenodo.org/record/244421/files/figure.png" pageId="3" pageNumber="508">Fig. 23</figureCitation>
). Length of parameres less than one-half that of phallobase (
<figureCitation id="134A24A8B173CC3527D5FE7A4F69FE54" box="[1234,1307,475,497]" captionStart="FIGURES 31 34" captionStartId="12.[151,264,1136,1158]" captionTargetBox="[165,1436,193,1114]" captionTargetId="figure@12.[150,1436,193,1115]" captionTargetPageId="12" captionText="FIGURES 31 34. Aedeagus of Aphengium; 31. A. sordidum (arrow shows dorsal margin of parameres curved inward at its apical third); 32. A. cupreum; 33. A. ibateguara sp. nov.; 34. A. curtum sp. nov." httpUri="https://zenodo.org/record/244423/files/figure.png" pageId="3" pageNumber="508">Fig. 33</figureCitation>
); apical third of parameres abruptly narrowed (
<figureCitation id="134A24A8B173CC35212FFE5E4803FDB0" box="[552,625,511,533]" captionStart="FIGURES 31 34" captionStartId="12.[151,264,1136,1158]" captionTargetBox="[165,1436,193,1114]" captionTargetId="figure@12.[150,1436,193,1115]" captionTargetPageId="12" captionText="FIGURES 31 34. Aedeagus of Aphengium; 31. A. sordidum (arrow shows dorsal margin of parameres curved inward at its apical third); 32. A. cupreum; 33. A. ibateguara sp. nov.; 34. A. curtum sp. nov." httpUri="https://zenodo.org/record/244423/files/figure.png" pageId="3" pageNumber="508">Fig. 33</figureCitation>
); fronto-lateral peripheral sclerite (FLP) as in figure 39.
<collectingCountry id="F36678BDB173CC352799FE5E4EA8FDB0" box="[1182,1242,511,533]" name="Brazil" pageId="3" pageNumber="508">Brazil</collectingCountry>
(Alagoas).............................................................................
<taxonomicName id="4C7143AEB173CC35275AFD824E96FD9D" ID-CoL="67LY8" box="[1117,1252,547,568]" class="Insecta" family="Scarabaeidae" genus="Aphengium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="ibateguara" status="sp. nov.">
<emphasis id="B905E43FB173CC35275AFD824E96FD9D" box="[1117,1252,547,568]" italics="true" pageId="3" pageNumber="508">A. ibateguara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2365944B173CC3527EDFD824F4BFD9C" box="[1258,1337,547,569]" pageId="3" pageNumber="508" rank="species">
<emphasis id="B905E43FB173CC3527EDFD824F4BFD9C" bold="true" box="[1258,1337,547,569]" pageId="3" pageNumber="508">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="134A24A8B173CC352641FD824FFDFD9C" box="[1350,1423,547,569]" captionStart="FIGURES 19 24" captionStartId="10.[151,264,1947,1969]" captionTargetBox="[180,1405,198,1922]" captionTargetId="figure@10.[173,1414,186,1923]" captionTargetPageId="10" captionText="FIGURES 19 24. Aphengium ibateguara sp. nov.; 19. Male (dorsal view); 20. Female (dorsal view); 21. Male head; 22. Female head; 23. Male pronotum; 24. Female pronotum." httpUri="https://zenodo.org/record/244421/files/figure.png" pageId="3" pageNumber="508">Fig. 19</figureCitation>
).
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>