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<document id="C73DFBCD8AC55D5FC010B3552A22926E" ID-DOI="http://dx.doi.org/10.3897/zookeys.423.7536" ID-GBIF-Dataset="1f17876d-2290-4d36-aa1f-10b951701679" ID-PMC="PMC4106094" ID-Pensoft-Pub="1313-2970-423-1" ID-PubMed="25061393" ID-ZooBank="55B4F9C858934F88841660FF730E8872" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-423-1" ModsDocOrigin="ZooKeys 423" ModsDocTitle="A revision of the Neotropical species of Bolitogyrus Chevrolat, a geographically disjunct lineage of Staphylinini (Coleoptera, Staphylinidae)" checkinTime="1451245649604" checkinUser="pensoft" docAuthor="Brunke, Adam J. &amp; Solodovnikov, Alexey" docDate="2014" docId="B2A65EA2FB2A2FC5792901293B48EC78" docLanguage="en" docName="ZooKeys 423: 1-113" docOrigin="ZooKeys 423" docSource="http://dx.doi.org/10.3897/zookeys.423.7536" docTitle="Cyrtothorax nevermanni Scheerpeltz 1974, syn. n." docType="treatment" docVersion="4" lastPageNumber="24" masterDocId="FFAB12065F4CFFED925DFFB46A5FFFFB" masterDocTitle="A revision of the Neotropical species of Bolitogyrus Chevrolat, a geographically disjunct lineage of Staphylinini (Coleoptera, Staphylinidae)" masterLastPageNumber="113" masterPageNumber="1" pageNumber="21" updateTime="1701318337016" updateUser="plazi">
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<mods:title id="57D4D05FF8FC06DD94929C95FBF06AA6">A revision of the Neotropical species of Bolitogyrus Chevrolat, a geographically disjunct lineage of Staphylinini (Coleoptera, Staphylinidae)</mods:title>
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<mods:namePart id="1D14B5515D0058582176A78D8375025D">Brunke, Adam J.</mods:namePart>
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<mods:namePart id="22379BEBB55258C2E5A7EA898302992C">Solodovnikov, Alexey</mods:namePart>
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<treatment id="B2A65EA2FB2A2FC5792901293B48EC78" ID-GBIF-Taxon="152054090" LSID="urn:lsid:plazi:treatment:B2A65EA2FB2A2FC5792901293B48EC78" httpUri="http://treatment.plazi.org/id/B2A65EA2FB2A2FC5792901293B48EC78" lastPageId="23" lastPageNumber="24" pageId="20" pageNumber="21">
<subSubSection id="B6AEDADE39B513DAD905C127CFFEF1E4" pageId="20" pageNumber="21" type="multiple">
<paragraph id="D9F38CBB79098A9568E2AA1EC9A9CD42" pageId="20" pageNumber="21">Taxon classification Animalia Coleoptera Staphylinidae</paragraph>
</subSubSection>
<subSubSection id="83072EB549343AC1D7E0CE4FDA87FDF5" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="8706EE33C91E2B619BA7DC209928F9D1" pageId="20" pageNumber="21">
<taxonomicName id="1FF86058BCCF74295B38FD2D6B7D2AB0" ID-CoL="33NVF" authority="Scheerpeltz, 1974" authorityName="Scheerpeltz" authorityYear="1974" class="Insecta" family="Staphylinidae" genus="Cyrtothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtothorax nevermanni" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="nevermanni">Cyrtothorax nevermanni Scheerpeltz, 1974</taxonomicName>
<taxonomicNameLabel id="D384C10314C5E99FB68906EF8E6554C5" pageId="20" pageNumber="21">syn. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="202ECD0F7863850D9AC28620342A1A5E" pageId="20" pageNumber="21" type="holotype">
<paragraph id="B495084D0AEB44F9B8AF71BBD1A6985C" pageId="20" pageNumber="21">Holotype</paragraph>
<paragraph id="2B82263BA5F01AC723DCC096D659777E" pageId="20" pageNumber="21">
(♀, NMW): Costa Rica, Carpintera, leg. Nevermann [white hand printed label]/ ♀ [white printed card]/
<taxonomicName id="97A902E2BC9D3EE35B4CEAEC2AFF8CFA" class="Insecta" family="Staphylinidae" genus="Cyrtothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtothorax (Nevermanni)" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="subGenus" subGenus="Nevermanni">Cyrtothorax Nevermanni</taxonomicName>
, n. sp./ Allotypus [red printed label]/ Nevermanni, Scheerp. [green hand printed card]/ ex. coll. Scheerpeltz [blue printed label]/ TYPUS,
<taxonomicName id="E191109E11934403C1995DEA2B3E4628" class="Insecta" family="Staphylinidae" genus="Cyrtothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtothorax" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">Cyrtothorax</taxonomicName>
, nevermanni, O. Scheerpeltz [red hand printed label]/ Holotype ♀,
<taxonomicName id="A72900B3D5B5BE7D99473097B2D096C2" class="Insecta" family="Staphylinidae" genus="Cyrtothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtothorax nevermanni" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="nevermanni">Cyrtothorax nevermanni</taxonomicName>
Scheerpeltz, 1974, det. A. Brunke 2013 [red printed label] /
<taxonomicName id="721503BF54E8ACB0FA481702032036B8" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus costaricensis" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="costaricensis">Bolitogyrus costaricensis</taxonomicName>
(Wendeler) Det. A. Brunke 2012 [white printed label].
</paragraph>
<paragraph id="59B23EF695CB0D8375BBD0D586563C98" pageId="20" pageNumber="21">
The single specimen located in NMW is interpreted as the holotype as
<bibRefCitation id="65C5533C915C51F65CD39031B4DB6189" author="Scheerpeltz, O" journalOrPublisher="Reichenbachia" pageId="72" pageNumber="73" pagination="175 - 192" title="Studien an den Arten der Gattung Cyrtothorax Kraatz mit Beschreibung neuer Arten sowie einer Dichotomik aller bis heute bekannt gewordenen Arten dieser Gattung (Coleoptera, Staphylinindae, Staphylininae, Quediini)." volume="15" year="1974">Scheerpeltz (1974)</bibRefCitation>
stated that only one specimen was seen. Although this holotype is also female, it can be associated with the same species concept as the type of
<taxonomicName id="46E71305E79B96C7DFFF15E0A2527BDD" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus costaricensis" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="costaricensis">Bolitogyrus costaricensis</taxonomicName>
for the same reasons (above). Therefore,
<taxonomicName id="399EC41E3615D3307B3F7FDBF704FD31" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus nevermanni" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="nevermanni">Bolitogyrus nevermanni</taxonomicName>
(Scheerpeltz) becomes a synonym of
<taxonomicName id="29328A32756E69AD64AF07275ABB8128" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus costaricensis" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="costaricensis">Bolitogyrus costaricensis</taxonomicName>
(Wendeler).
</paragraph>
</subSubSection>
<subSubSection id="672DD73802DBC7E6F2412A06961AB9B6" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="other material">
<paragraph id="8C95FEA53A8902A02BB7FF71EF604601" pageId="21" pageNumber="22">
<pageBreakToken id="59DA487DC8069AC8FDC1B0FA87A940D0" pageId="21" pageNumber="22" start="start">Other</pageBreakToken>
material.
</paragraph>
<paragraph id="AA7121331191D890C35E57EFBD969399" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
GUATEMALA:
<pageBreakToken id="D131435C636569A31A66BADBDA785B1A" pageId="22" pageNumber="23" start="start">Baja</pageBreakToken>
Verapaz,
<normalizedToken id="59843C161C31BC11C95E7E7EE583C2F7" originalValue="Sinanjá">Sinanja</normalizedToken>
, Champion, &quot;Sharp Coll.1905.-313&quot;, 1 ♂ (BMNH). San Marcos: 2 km E de Tajomulco [=Tajumulco], Bosque
<normalizedToken id="CCABDC82D39D48E8C618AD56DCD99440" originalValue="mesófilo">mesofilo</normalizedToken>
Perturbado [=disturbed cloud forest], 2062m,
<geoCoordinate id="7B86DBABA2E5B14D2B8AE2BCE67FF86A" direction="north" orientation="latitude" precision="15" value="15.094167">15°04'99&quot;N</geoCoordinate>
, 91°55'563W, en hongos de reprisa [=in shelf fungus], 23.IX.1999, J.
<normalizedToken id="94D280FBDA5F40EE5EF39E66FA98E2F2" originalValue="Márquez">Marquez</normalizedToken>
, 1 ♂ (MZFC). HONDURAS:Comayagua: 13 km SE El Mochito, Sta. B, 22.VII.1977, 'CW &amp; L.',
<normalizedToken id="FC32A33F705F888A7DF9A72CCF71F558" originalValue="OBrien">O'Brien</normalizedToken>
&amp; Marshall, 1 ♂ (FMNH). El Paraiso: 9.4 km SE Zamaranom &amp; 9.4 km SE Galeras, 'Los
<normalizedToken id="F048085C21924BC5F8F1CDC9C5F47CE8" originalValue="Lavenderos">Lavenderos'</normalizedToken>
,
<geoCoordinate id="2D4B540C69DDDCF343C7D32F8C699FC7" direction="north" orientation="latitude" precision="925" value="13.4">13°24'N</geoCoordinate>
,
<geoCoordinate id="E1D7D6AE1A84DFCCBCF19AD81B56C987" direction="west" orientation="longitude" precision="925" value="-86.916664">86°55'W</geoCoordinate>
, 1450m, 11.VI.1994, ex. crustose fungi under logs, Ashe, Brooks, SM0037968, SM0037969, SM0037970, SM0037971, 1 ♂, 3 ♀ (SEMC). Francisco
<normalizedToken id="52B106589F54E430B85F149CFB8EC9B2" originalValue="Morazán">Morazan</normalizedToken>
: 10.4 km W Zamorano, Cerro Uyuca,
<geoCoordinate id="86BEFEC4ACB4121F9CAA1CAC63139E4C" direction="north" orientation="latitude" precision="925" value="14.033334">14°02'N</geoCoordinate>
,
<geoCoordinate id="4DFDF8852DB8240692C8B126B4225EF7" direction="west" orientation="longitude" precision="925" value="-87.066666">87°04'W</geoCoordinate>
, 1500m, 27.VI.1994, ex. crustose fungi under log, Ashe, Brooks, SM0037967, 1 ♂ (SEMC); 7.6 km N Guaimaca,
<geoCoordinate id="4D2C11AE14C6B4DC15CEB9E60894D460" direction="north" orientation="latitude" precision="925" value="14.6">14°36'N</geoCoordinate>
,
<geoCoordinate id="237255D6B1E9EE14151D85CFC8D4A498" direction="west" orientation="longitude" precision="925" value="-86.816666">86°49'W</geoCoordinate>
, 820m, 26.VI.1994, crustose fungi under logs, J. Ashe, R. Brooks, SM0037975, SM0037976, SM0037977, SM0037978, SM0037979, SM0037980, SM0037981, SM0037982, 6 ♂, 2 ♀ (SEMC); Lempira: 5.9 km SW Gracias, Mt. Celaque,
<geoCoordinate id="4530299769E3B61BD357BA7D87D9AEB6" direction="north" orientation="latitude" precision="925" value="14.55">14°33'N</geoCoordinate>
,
<geoCoordinate id="C2574189E3FAA8565DF240067BD53A3A" direction="west" orientation="longitude" precision="925" value="-88.63333">88°38'W</geoCoordinate>
, 1250m, 17.VI.1994, on/under fungusy log, SM0037965, SM0037966, 2 ♂ (SEMC); 13.1 km NE &amp; 7.3 km E Gracias, Mt. Puca,
<geoCoordinate id="9C8192168EBD3F2D5703EC1CB8500708" direction="north" orientation="latitude" precision="925" value="14.683333">14°41'N</geoCoordinate>
,
<geoCoordinate id="01AD151D20C1DD2C064069746EC28AEF" direction="west" orientation="longitude" precision="925" value="-88.51667">88°31'W</geoCoordinate>
, 1320m, 18.VI.1994, crustose fungi under logs, J. Ashe, R. Brooks, SM0037973 and SM0037974, 1 ♂, 1 ♀ (SEMC); same as previous except: under wood slabs, SM0037963, SM0037962, 1 ♂, 1 ♀ (SEMC). Santa Barbara: Santa Barbara, Mt. Santa Barbara, 11.5 km S &amp; 5.6 km W Pefia Blanca,
<geoCoordinate id="00BD8268FA4BA23CE62E188FBA93E9D0" direction="north" orientation="latitude" precision="925" value="14.95">14°57'N</geoCoordinate>
,
<geoCoordinate id="8C9F58018CF339EA2288E6CFBD9B7549" direction="west" orientation="longitude" precision="925" value="-88.083336">88°05'W</geoCoordinate>
', 1800m, 20.VI.1994, beating stick pile, R. Brooks, J. Ashe, SM0037972, 1 ♂ (SEMC). NICARAGUA:Grenada: Res. Nat.
<normalizedToken id="64CDB4312DE4CDA17BA1C6E30AAD7C32" originalValue="Volcán">Volcan</normalizedToken>
Mombacho, Bmm, 11°50'25N, 85°58'593W, 1039 m, ex. tronco podridos [=rotten logs], 2.II.2000, J.
<normalizedToken id="B8918031FCB83042AF4FB3C71CFA9BA7" originalValue="Márquez">Marquez</normalizedToken>
, 1 ♂, 1 ♀ (MZFC); same as previous except: 3.II.2000, 1 ♂ (MZFC); same as previous except: 4.II.2000, 2 ♂ (MZFC); Res. Nat. Volcan Mombacho, entrance road,
<geoCoordinate id="69A091C8AA4F911A8DC9FD75DBB463CA" direction="north" orientation="latitude" precision="9" value="11.834167">11°50.05'N</geoCoordinate>
,
<geoCoordinate id="01978E4F883E842438D7DE22B0E7C6F5" direction="west" orientation="longitude" precision="9" value="-85.9805">85°58.83'W</geoCoordinate>
, 1060m, flight intercept trap, 1 to 5.VI.2002, R. Brooks, Z. Falin, S. Chatzimanolis, SM0575457, SM0575458, SM0575459, 2 ♂, 1 ♀ (SEMC); same as previous except: 800-1000 m, pyrethrum fogging fungusy logs, NIC1BFC02 145, SM0413171, 1 ♂ (SEMC); Res. Nat. Volcan Mombacho, station,
<geoCoordinate id="8C17432FDCDDC7CACF6C8B1BA1978B3C" direction="north" orientation="latitude" precision="9" value="11.834167">11°50.05'N</geoCoordinate>
,
<geoCoordinate id="A462BE7B854EDBE65C524C44DBB31CE5" direction="west" orientation="longitude" precision="9" value="-85.9805">85°58.83'W</geoCoordinate>
, 1150m, mercury vapor/UV light, 2.VI.2002, R. Brooks, Z. Falin, S. Chatzimanolis, SM0533433, 1 ♂ (SEMC); Res. Nat. Volcan Mombacho,
<geoCoordinate id="DFACCDCE9020F22C56054D324BA42C18" direction="north" orientation="latitude" precision="9" value="11.834167">11°50.05'N</geoCoordinate>
,
<geoCoordinate id="39E99930088ED9121ABEBAE0F0703D27" direction="west" orientation="longitude" precision="9" value="-85.9805">85°58.83'W</geoCoordinate>
, 1150m, pyrethrum fogging fungusy logs, 2.VI.2002, R. Brooks, Z. Falin, S. Chatzimanolis, SM0533722, SM0413244, SM0413245, SM0413246, 3 ♂, 1 ♀ (SEMC); same as previous except: 1200m, ex. cut lumber, SM0413175, SM0413176, SM0413177, SM0413178, 4 ♂ (SEMC); same as previous except: pyrethrum fogging fungusy logs, SM0413179, SM0413180, 1 ♂, 1 ♀ (SEMC, ZMUC); Res. Nat. Volcan Mombacho,
<geoCoordinate id="8E4C12DE54B5B4D2BAA6FCBCAD423580" direction="north" orientation="latitude" precision="9" value="11.834167">11°50.05'N</geoCoordinate>
,
<geoCoordinate id="708018DBCBBD3377054A8605DD733223" direction="west" orientation="longitude" precision="9" value="-85.9805">85°58.83'W</geoCoordinate>
, 1150 m, pyrethrum fogging fungusy logs, 3.VI.2002, R. Brooks, Z. Falin, S. Chatzimanolis, SM0413225, SM0413226, SM0413228, SM0413230, SM0413231, SM0413233, SM0413235, SM0413236, SM0413237, SM0413238, SM0413239, SM0413240, SM0413241, 7 ♂, 6 ♀ (SEMC); same as previous except: 4.VI.2002, SM0413264, SM0413265, SM0413266, SM0413267, SM0413268, SM0413269, SM0413270, SM0413271, SM0413272, 5 ♂, 4 ♀ (SEMC); same as previous except: 5.VI.2002, SM0555363, SM0555364, SM0555365, SM0555366, 4 ♂ (SEMC, UTCI, ZMUC). Jinotega: 16 km N Matagalpa, Matagalpa-Jinotega road,
<geoCoordinate id="FB89655A68F929050F3CFC7B2FFF34B3" direction="north" orientation="latitude" precision="9" value="13.045">13°02.70'N</geoCoordinate>
,
<geoCoordinate id="6B65AAF9C0F5C0B3A4E7B16D8F6F741B" direction="west" orientation="longitude" precision="9" value="-85.935">85°56.10'W</geoCoordinate>
, 1385m, 22.V.2002, pyrethrum fogging fungusy logs, R. Brooks, Z. Falin, S. Chatzimanolis, SM0413166, SM0413167, SM0413168, SM0413169, 3 ♂, 1 ♀ (SEMC). Matagalpa: 6 km N Matagalpa, Selva Negra Hotel,
<geoCoordinate id="965C79D59D7BB03B5F9BAA0A1B3C5DC5" direction="north" orientation="latitude" precision="9" value="12.999833">12°59.99'N</geoCoordinate>
,
<geoCoordinate id="9B03CA6CE5911A3551CDB419779C9465" direction="west" orientation="longitude" precision="9" value="-85.90884">85°54.53'W</geoCoordinate>
, 1240m, flight intercept trap, Bavaria trail, 18 to 21.V.2002, S. Chatzimanolis, SM0413258, 1 ♂ (SEMC); same as previous except: 1300m, Cody trail, R. Brooks, Z. Falin, S. Chatzimanolis, SM0555396, SM0555397, 2 ♂ (SEMC); 6 km N Matagalpa, Selva Negra,
<geoCoordinate id="57F74B57B6600C957F8743F7F8D3795B" direction="north" orientation="latitude" precision="92" value="12.998333">12°59.9'N</geoCoordinate>
,
<geoCoordinate id="8243CED9D21CC830980E18EC6EE30DA3" direction="west" orientation="longitude" precision="92" value="-85.91">85°54.6'W</geoCoordinate>
, 1250m, flight intercept trap, 18 to 22.V.2002, S. Peck, SM0560604, SM0560605, SM0560610, SM0560623, SM0560627, SM0560630, SM0560632, SM0560635, SM0560636, SM0560638, SM0560639, SM0560645, SM0560647, SM0560652, 8 ♂ 5 ♀ (SEMC); 6 km N Matagalpa, Selva Negra Hotel,
<geoCoordinate id="6F897539C9D23E5AA5BDE4DC96C43763" direction="north" orientation="latitude" precision="9" value="12.999833">12°59.99'N</geoCoordinate>
,
<geoCoordinate id="E8D0710087397199A916743F7605B900" direction="west" orientation="longitude" precision="9" value="-85.90884">85°54.53'W</geoCoordinate>
, 1250m, 19.V.2002, pyrethrum fogging fungusy logs, R. Brooks, Z. Falin, S. Chatzimanolis, SM0575440, SM0575442, SM0575441, SM0575439, SM0558105, SM0558113, 3 ♂ 3 ♀ (SEMC); same as previous except: 1350m, crustose polypore, SM0540977, SM0540978, SM0540979, SM0540980, SM0540981, SM0540982, SM0541459, 1 ♂ 6 ♀ (SEMC); 6 km N Matagalpa, Selva Negra Hotel,
<geoCoordinate id="9B0D81F91B67A66B53FF137B25E066D2" direction="north" orientation="latitude" precision="9" value="12.999833">12°59.99'N</geoCoordinate>
,
<geoCoordinate id="FA638417F78B911F5A3E2E83D2CA6867" direction="west" orientation="longitude" precision="9" value="-85.90884">85°54.53'W</geoCoordinate>
, 1520m, 20.V.2002, pyrethrum fogging fungusy logs, R. Brooks, Z. Falin, S. Chatzimanolis, SM0542449, SM0542488, SM0575433, 2 ♂, 1 ♀ (SEMC); same as previous except: 1350m, ex. crustose polypore, SM0540939, SM0540940, SM05409941, SM0540942, SM0540943, SM0540944, SM540946, SM0540949, SM0540950, SM0540951, SM0540952, SM0540953, SM0540954, SM0540955, SM0540956, SM0540957, SM0540958, SM0540960 SM0540963, SM540964, 13 ♂, 7 ♀ (SEMC); same as previous except: 21.V.2002, SM0541899, SM0541903, SM0543437, SM0575530, SM0575531, SM0575532, SM0575533, SM0575534, SM0575535, SM0575536, SM0575537, SM0575538, SM0575539, SM0575540, SM0575541, SM0575542, SM0575543, SM0575544, SM0575545, SM0575546, SM0575547, SM0575548, SM0575549, SM0575550, SM0575551, SM0575552, SM0575553, SM0575554, SM0575555, SM0575556, 19 ♂, 11 ♀ (SEMC); 6 km N Matagalpa, Selva Negra Hotel,
<geoCoordinate id="AD10AF1BF48AEDC40C1D67D8F56F9259" direction="north" orientation="latitude" precision="9" value="12.999833">12°59.99'N</geoCoordinate>
,
<geoCoordinate id="D3106423CEA12F28304FA3D50A2725BE" direction="west" orientation="longitude" precision="9" value="-85.90884">85°54.53'W</geoCoordinate>
, 1350m, 21.V.2002, ex. small white bracket fungi, R. Brooks, Z. Falin, S. Chatzimanolis, SM0558419, 1 ♂ (SEMC); same as previous except: ex. cut lumber, SM0575557, SM0575558, SM0575560, SM0575563, 2 ♂, 2 ♀ (SEMC); same as previous except: on
<taxonomicName id="08A66FFB32DA17B460E55BA4AE68D7F0" genus="Xylariaceae" lsidName="Xylariaceae" pageId="22" pageNumber="23" rank="genus">Xylariaceae</taxonomicName>
, SM0575488, SM0575489, SM0575490, SM0575491, SM0575492, SM0575493, SM0575494, SM0575495, SM0575496, SM0575497, 8 ♂, 2 ♀ (SEMC). COSTA RICA: [locality illegible], 18.IV.1938, 1 ♂ (FMNH); country record only, unter loser Rinde [under loose bark], 9.XI.1936, 1 ♀ (USNM). Cartago: Carpintera, 16.VI.1940, 2 ♂, 2 ♀ (FMNH); Tres
<normalizedToken id="EBC9F6B5A799CCFF820F985FB77B5940" originalValue="Ríos">Rios</normalizedToken>
, 23.VI.1940, 4 ♂ (FMNH); Cerro Carpintera, Iztaru Scout Camp, 1670m, ex. fog dry log, 21.IV.2000, CR-1090, P.N. Thomas, 3 ♂, 4 ♀ (PTC); Santa Cruz de Turrialba, ex. sub bark log in pasture near road, 26.VIII.1994, P.N. Thomas, CR-432, 1 ♀ (PTC). Guanacaste: P. N. Guanacaste, Lado SO [=southwest side] vol. Cacao, Est. Cacao, 1000-1400 m, 21 to 29.V.1992, F. A. Quesada, CR1000378852, 1 ♂ (INBIO); same as previous except: R. Blanco &amp; C. Chavez, 1000086948, 1 ♂ (INBIO); P.N. Guanacaste, Lado O [=west side] Volcan Orosi, Estac. Maritza, 600 m, malaise, 1989, CR1000037210, 1 ♂ (INBIO); same as previous except: 1992, CR1000377704, 1 ♂ (INBIO); Rincon de la Vieja, Pailas Station, 740-790 m, ex. fog asst. logs, 1.XII.2000, CR-1147, 1 ♂ (PTC).
<normalizedToken id="5D66C4E914F403991955E1FE850FE807" originalValue="Limón">Limon</normalizedToken>
: Parismina, M. Valerio coll., 1 ♀ (USNM) [LOC. IN ERROR]. Puntarenas: Monte Verde, 1250m, 7.V.1989, J. Ashe, R. Brooks, R. Leschen, SM0037961, 1 ♂ (SEMC); Monte Verde, 1400m, ex. flight intercept trap, 21.V.1989, J. Ashe, R. Brooks, R. Leschen, SM0037960, 1 ♂ (SEMC). San Jose: La Caja, VIII.1938, 2 ♂ (FMNH); San Jose, 1935, 1 ♂ (FMNH), same except 1937, 1 ♀ (USNM);
<normalizedToken id="FEAEB8108A0D827235E4635E3112A932" originalValue="Río">Rio</normalizedToken>
Virilla, 15.V.1941, Bierig, 1 ♂ (FMNH); Rincon de Santa Maria South, 1650 m, ex. rotten stump, 17.XI.2003, CR-1481, P.N. Thomas, 3 ♂ (PTC); San Francisco de Leon Cortes, 1780 m, ex. fog dry logs, 3.IV.2000, CR-1119, P.N. Thomas, 3 ♂, 4 ♀ (PTC); San Marcos [in error] [San Lorenzo] de
<normalizedToken id="5F3759755C23B748887B08DFF2DFAFC2" originalValue="Tarrazú">Tarrazu</normalizedToken>
, Zapotal, 1500 m, ex. rotten oak logs, 4.I.1996, sample #65, P.N. Thomas, 3 ♂, 3 ♀ (PTC); same except: ex. logs with moss, 18.XII.1993, sample #267, 2 ♂, 3 ♀ (PTC); San Marcos, San Lorenzo de
<normalizedToken id="E790BA3F879918FAD7C0632049CEABB9" originalValue="Tarrazú">Tarrazu</normalizedToken>
, Cerro Zapotal, 1500 m, ex. dried mushrooms, pan, 16.XII.1993, CR-255, P.N. Thomas, 1 ♂ (PTC); same except: ex. encino and poro log, 23.VIII.1994, CR-417, 1 ♂ (PTC).
</paragraph>
</subSubSection>
<subSubSection id="49626A962C54DE2C4ED724DA05CFB8E6" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="ED307857698C2EAC9F1972B1E14F5537" pageId="22" pageNumber="23">Diagnosis.</paragraph>
<paragraph id="C952F8AEB4FABE84C4118F6A080C7BEF" pageId="22" pageNumber="23">
Head without central protuberance; antenna and legs distinctly bicolored (Fig. 5A; 11G); genital and abdominal segments VIII not distinctly paler than previous segments; abdominal sternites with fine transverse microsculpture, interspaces about as wide as lines (Fig. 13D); female procoxa completely dark; median lobe recurved at apex (Fig. 15F); paramere with convergent arms; paramere with fields of peg setae wide and circular at base of arms (Fig. 15
<normalizedToken id="57A60C0E9205B769AC5644E8EEA349DE" originalValue="GH">G-H</normalizedToken>
); female tergite X with raised disc converging strongly toward apex, strongly pigmented area not or weakly incised at apicolateral margin (Fig. 25
<normalizedToken id="DDD6E36E5C3EBE667830F16B3F72551E" originalValue="AD">A-D</normalizedToken>
); female secondary gonocoxite not distinctly swollen at base.
</paragraph>
</subSubSection>
<subSubSection id="840BBCFF81BBFC834BD9DF7A6B1278F3" pageId="22" pageNumber="23" type="redescription">
<paragraph id="5DD39A3371B8B50ADDAB22E7A9575B77" pageId="22" pageNumber="23">Redescription.</paragraph>
<paragraph id="F03FC925F8BF1E37AF890BF9DB361318" pageId="22" pageNumber="23">Measurements ♂ (n=5): HW/HL 1.54-1.62; PW/PL 1.29-1.67; EW/EL 1.24-1.26; ESut/PL 0.68-0.80; PW/HW 1.10-1.25; forebody length 4.1-4.9 mm.</paragraph>
<paragraph id="B79C72DA419D7916E90D28446A583077" pageId="22" pageNumber="23">Measurements ♀ (n=5): HW/HL 1.50-1.67; PW/PL 1.35-1.47; EW/EL 1.21-1.30; ESut/PL 0.75-0.78; forebody length - 4.5-5.1 mm.</paragraph>
<paragraph id="7C400595BA1DB4F8A8765900FB145675" pageId="22" pageNumber="23">
Morphology as in
<taxonomicName id="F0902FBCEB00C1CE9EE857BBAC41A8FD" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus buphthalmus" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="buphthalmus">Bolitogyrus buphthalmus</taxonomicName>
, differing only in the following: metallic reflections of body bronze to greenish or grayish-bronze or violet to violet-blue; median lobe in lateral view recurved dorsad (Fig. 15F); peg setae fields broad and circular at base of parameral lobe (Fig. 15
<normalizedToken id="C1CC027B8A0D600A2DA6ADE161135B4C" originalValue="GH">G-H</normalizedToken>
); male sternite IX with apical emargination deeper (Fig. 23B); female tergite X with strongly pigmented area not or weakly incised lateroapically (Fig. 25
<normalizedToken id="323BC11C971BC14583793A71325FA514" originalValue="AD">A-D</normalizedToken>
).
</paragraph>
</subSubSection>
<subSubSection id="18B69D9080F27EA078DB0457327FC665" pageId="22" pageNumber="23" type="distribution">
<paragraph id="510FD6B8AB63711C87C4796593F21863" pageId="22" pageNumber="23">Distribution.</paragraph>
<paragraph id="0849D8A53661B32C61EE23CE896F8991" pageId="22" pageNumber="23">
Figure 29
<normalizedToken id="8B928F46B95FCE0DF759E77F69E98E01" originalValue="AB">A-B</normalizedToken>
.
<taxonomicName id="823C4AC4045FC3AE6D885AD4CA25EC8B" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus costaricensis" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="costaricensis">Bolitogyrus costaricensis</taxonomicName>
is known from Guatemala, Honduras, Nicaragua and Costa Rica, west to the northern part of the Talamanca Cordillera. Two females from Belize were seen that might belong to this species or
<taxonomicName id="386B698BCCF627378BE83561C6ACA1A2" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus buphthalmus" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="buphthalmus">Bolitogyrus buphthalmus</taxonomicName>
. The specimen from
<normalizedToken id="E860B36042DD47E4D999BEE9B014DD36" originalValue="Limón">Limon</normalizedToken>
province in Costa Rica was almost certainly collected from elsewhere as Parismina is a city located on the Caribbean coast, situated at sea level, which is distinctly outside the elevational range of this species.
</paragraph>
</subSubSection>
<subSubSection id="AD490550459205EC5A0299B19899481A" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="bionomics">
<paragraph id="C72131FE19173373BB11C44B2B768804" pageId="22" pageNumber="23">Bionomics.</paragraph>
<paragraph id="07C1849853814DF2A275C351E43B41AD" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
Most specimens of
<taxonomicName id="35CDFB942669E12651C8A3224C32AF98" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus costaricensis" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="costaricensis">
<pageBreakToken id="4CD1EF042B8BBA8C8021495184514818" pageId="23" pageNumber="24" start="start">Bolitogyrus</pageBreakToken>
costaricensis
</taxonomicName>
have been collected by pyrethrum fogging of fungusy logs, while others were collected from bracket fungi, dried mushrooms, small crustose polypores, fungi in the family
<taxonomicName id="C8FAA59B8A3BDB9F1F8F3AF0860F6B73" genus="Xylariaceae" lsidName="Xylariaceae" pageId="23" pageNumber="24" rank="genus">Xylariaceae</taxonomicName>
, various woody debris and in FIT or Malaise traps; one slightly teneral specimen came to light.
<taxonomicName id="A3926B7EA4CCB72CB5CACB3076A22BA0" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus costaricensis" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="costaricensis">Bolitogyrus costaricensis</taxonomicName>
has been collected in forests at elevations ranging from 600-2062 m in every month of the year.
</paragraph>
</subSubSection>
<subSubSection id="952347F032F1A6B57D136F5191B7BA96" pageId="23" pageNumber="24" type="comments">
<paragraph id="7255AECEEE0CC4EBC24934EB70A14C30" pageId="23" pageNumber="24">Comments.</paragraph>
<paragraph id="B87ECADD34709A3E35379788B2F478A7" pageId="23" pageNumber="24">
<taxonomicName id="9EE93DF448A3487D200F32934D996C99" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus costaricensis" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="costaricensis">Bolitogyrus costaricensis</taxonomicName>
is most similar to
<taxonomicName id="41C973EF1183228ABB1E0C9B055973AE" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus buphthalmus" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="buphthalmus">Bolitogyrus buphthalmus</taxonomicName>
and can only be separated from it by characters on the aedeagus (see above).
</paragraph>
</subSubSection>
</treatment>
</document>