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<document ID-CLB-Dataset="9619" ID-DOI="http://dx.doi.org/10.3897/zookeys.988.56188" ID-GBIF-Dataset="a38f6648-1c49-486b-a7ee-a877315d4e09" ID-PMC="PMC7666111" ID-Pensoft-Pub="1313-2970-988-63" ID-Pensoft-UUID="D5BC323AA5F05894AE8FBB10A3F0960D" ID-PubMed="33223891" ID-ZooBank="6E64D69BDD734A7EAE2B3CD21247A5E3" ModsDocID="1313-2970-988-63" checkinTime="1604753996846" checkinUser="pensoft" docAuthor="Li, Ya, Lin, Yucheng &amp; Li, Shuqiang" docDate="2020" docId="E0EC47F4BC155C429D31AF5C685013FC" docLanguage="en" docName="ZooKeys 988: 63-128" docOrigin="ZooKeys 988" docSource="http://dx.doi.org/10.3897/zookeys.988.56188" docTitle="Crassignatha ertou Miller, Griswold &amp; Yin 2009" docType="treatment" docVersion="6" id="D5BC323AA5F05894AE8FBB10A3F0960D" lastPageNumber="63" masterDocId="D5BC323AA5F05894AE8FBB10A3F0960D" masterDocTitle="A review of Crassignatha (Araneae, Symphytognathidae)" masterLastPageNumber="128" masterPageNumber="63" pageNumber="63" updateTime="1732684104256" updateUser="ExternalLinkService">
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<mods:title id="0745B8D4C775F1BC53B1717C8ED9C1FD">A review of Crassignatha (Araneae, Symphytognathidae)</mods:title>
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<mods:namePart id="A1C602921B4C02B5085D49E779182A35">Li, Ya</mods:namePart>
<mods:affiliation id="6F050C3F2AEE920E0BC6B1F94F0C7B5C">Key Laboratory of Bio-resources and Eco-environment (Ministry of Education), College of Life Sciences, Sichuan University, Chengdu, Sichuan 610064, China &amp; The Sichuan Key Laboratory for Conservation Biology of Endangered Wildlife, Sichuan University, Chengdu, Sichuan 610064, China</mods:affiliation>
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<mods:namePart id="9C33419F16CD19AD22D5C98159C54D3A">Lin, Yucheng</mods:namePart>
<mods:affiliation id="8033BD55FDC017C458AC7FAF343F6AEF">Key Laboratory of Bio-resources and Eco-environment (Ministry of Education), College of Life Sciences, Sichuan University, Chengdu, Sichuan 610064, China &amp; The Sichuan Key Laboratory for Conservation Biology of Endangered Wildlife, Sichuan University, Chengdu, Sichuan 610064, China</mods:affiliation>
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<mods:namePart id="AFCA4E36B9C88C4822AD0092074DC813">Li, Shuqiang</mods:namePart>
<mods:affiliation id="3D7090573579630E47538CF120B88540">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<treatment id="E0EC47F4BC155C429D31AF5C685013FC" ID-GBIF-Taxon="168821875" LSID="urn:lsid:plazi:treatment:E0EC47F4BC155C429D31AF5C685013FC" httpUri="http://treatment.plazi.org/id/E0EC47F4BC155C429D31AF5C685013FC" lastPageNumber="63" pageId="0" pageNumber="63" scope_family="Symphytognathidae" scope_order="Araneae">
<subSubSection id="A8BF1F7A102A96E1E89E483072ACA674" pageId="0" pageNumber="63" type="nomenclature">
<paragraph id="9498A1F4BA4D21D48C007C225936CE34" pageId="0" pageNumber="63">
<taxonomicName id="B57AEC806053758D2FEDC309EAE8274F" ID-CoL="Z6KT" LSID="E0EC47F4-BC15-5C42-9D31-AF5C685013FC" authority="Miller, Griswold &amp; Yin, 2009" authorityName="Miller, Griswold &amp; Yin" authorityYear="2009" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crassignatha ertou" order="Araneae" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="ertou">Crassignatha ertou Miller, Griswold &amp; Yin, 2009</taxonomicName>
<figureCitation id="80B979CECACE17E86850E1CF5F6EFEFF" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Crassignatha ertou Miller, Griswold &amp; Yin, 2009 A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Scale bars: 0.50 mm (A-F)." figureDoi="10.3897/zookeys.988.56188.figure7" httpUri="https://binary.pensoft.net/fig/472592" pageId="0" pageNumber="63">Figs 7</figureCitation>
<figureCitation id="7D0E1359F7A066F97D4D7D4029F6D56B" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Crassignatha ertou Miller, Griswold &amp; Yin, 2009 A male palp, prolateral B male palp, ventral C male palp, retrolateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.10 mm (A-G)." figureDoi="10.3897/zookeys.988.56188.figure8" httpUri="https://binary.pensoft.net/fig/472593" pageId="0" pageNumber="63">, 8</figureCitation>
<figureCitation id="D7EF9059C9D7BBAF429392B6D69DAFDA" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Distribution records of Crassignatha spp. in Asia. 1 C. baihua sp. nov. 2 C. bangbie sp. nov. 3 C. bicorniventris 4 C. changyan sp. nov. 5 C. danaugirangensis 6 C. dongnai sp. nov. 7 C. ertou 8 C. gucheng sp. nov. 9 C. gudu 10 C. haeneli 11 C. mengla sp. nov. 12 C. nantou sp. nov. 13 C. nasalis sp. nov. 14 C. pianma 15 C. quadriventris 16 C. quanqu 17 C. rostriformis sp. nov. 18 C. shiluensis 19 C. shunani sp. nov. 20 C. si sp. nov. 21 C. thamphra sp. nov. 22 C. xichou sp. nov. 23 C. yamu 24 C. yinzhi." figureDoi="10.3897/zookeys.988.56188.figure38" httpUri="https://binary.pensoft.net/fig/472623" pageId="0" pageNumber="63">, 38</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DB8E8FE7E3A88D41D9E07D3B8B60B1EC" pageId="0" pageNumber="63" type="reference_group">
<paragraph id="E9024C3E20CC0C82FACB6AF623E7E1E4" pageId="0" pageNumber="63">
<taxonomicName id="505D798D77BB15F8C8B4EB84ED79B55A" authorityName="Miller, Griswold &amp; Yin" authorityYear="2009" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crassignatha ertou" order="Araneae" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="ertou">Crassignatha ertou</taxonomicName>
<bibRefCitation id="C55F41BEB19B61A2941D75166AC7300C" DOI="https://doi.org/10.3897/zookeys.11.160" author="Miller, JA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="63" pagination="9 - 195" refId="B15" refString="Miller, JA, Griswold, CE, Yin, CM, 2009. The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae, Araneoidea): systematics and diversity of micro-orbweavers. ZooKeys 11: 9 - 195, DOI: https://doi.org/10.3897/zookeys.11.160" title="The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae, Araneoidea): systematics and diversity of micro-orbweavers." url="https://doi.org/10.3897/zookeys.11.160" volume="11" year="2009">Miller et al., 2009</bibRefCitation>
: 74, figs 86D-F, 88A, B, 89A, B (♂♀).
</paragraph>
</subSubSection>
<subSubSection id="781D007C3DBB5FF14B551A8F3283708C" pageId="0" pageNumber="63" type="type material">
<paragraph id="0264D23B77CC82C690E5647D4645E066" pageId="0" pageNumber="63">Type materi al.</paragraph>
<paragraph id="835B163351D614895A55E75FC33D49BF" pageId="0" pageNumber="63">
<emphasis id="71581AB2CB7A8C957FD737D9BC5AB55B" bold="true" italics="true" pageId="0" pageNumber="63">Holotype</emphasis>
♂ (HNU-CASENT 9029324) and
<emphasis id="56D9B007ED479556D202B41186C94962" bold="true" italics="true" pageId="0" pageNumber="63">paratypes</emphasis>
3♀ (HNU-CASENT 9022397),
<emphasis id="1443676AE6280D05AB2C5E20D3633CDA" bold="true" pageId="0" pageNumber="63">China</emphasis>
: Yunnan Province, Longling County, Mangkuan Township, Zaotanghe at Baihualing Village, undisturbed subtropical broadleaf forest, dusting webs in understory (
<geoCoordinate id="33658855BCA18F277B645C4E33438BDE" degrees="25.30450" direction="north" orientation="latitude" precision="1" value="25.3045">25.30450°N</geoCoordinate>
,
<geoCoordinate id="01E15FEA104D82236FD67E7EF76DA242" degrees="98.80059" direction="east" orientation="longitude" precision="1" value="98.80059">98.80059°E</geoCoordinate>
; 1635 m), 2.VI.2005, C. Griswold leg. Examined.
</paragraph>
<caption id="F4B667D4C067DF47052B1659DA1E16D9" doi="10.3897/zookeys.988.56188.figure7" httpUri="https://binary.pensoft.net/fig/472592" pageId="0" pageNumber="63" start="Figure 7" startId="F7">
<paragraph id="09EAD9B05F313BED40864DA30D156CFB" pageId="0" pageNumber="63">
<emphasis id="B01FB6F63E52D51E876F0FFA452B18F9" bold="true" pageId="0" pageNumber="63">Figure 7.</emphasis>
<taxonomicName id="440D83842D065DD36150C4E0B1C7C63D" authorityName="Miller, Griswold &amp; Yin" authorityYear="2009" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crassignatha ertou" order="Araneae" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="ertou">
<emphasis id="AF193EE50A361EB7A5CB95EACC7351D3" italics="true" pageId="0" pageNumber="63">Crassignatha ertou</emphasis>
</taxonomicName>
Miller, Griswold &amp; Yin, 2009
<emphasis id="18DFEE7290C7306BF666C97C58C1700D" bold="true" pageId="0" pageNumber="63">A</emphasis>
male habitus, dorsal
<emphasis id="420576AAA6ABF3E50A73276D2C71F4DD" bold="true" pageId="0" pageNumber="63">B</emphasis>
male habitus, ventral
<emphasis id="8FF506F1C88C1F171AA39763F5E91A4D" bold="true" pageId="0" pageNumber="63">C</emphasis>
male habitus, lateral
<emphasis id="2C0AE46D81E84148DE750B21DDEC6E61" bold="true" pageId="0" pageNumber="63">D</emphasis>
female habitus, dorsal
<emphasis id="5C3CBE4B347183AD2529679611AD30AB" bold="true" pageId="0" pageNumber="63">E</emphasis>
female habitus, ventral
<emphasis id="B8423A75505E2F33623129576C206BFC" bold="true" pageId="0" pageNumber="63">F</emphasis>
female habitus, lateral. Scale bars: 0.50 mm (
<emphasis id="ADB8E11DE67B16F7E0E92DAD12269B06" bold="true" pageId="0" pageNumber="63">A-F</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="410CED950C08CD638CBDBDE59263F9DF" pageId="0" pageNumber="63" type="materials_examined">
<paragraph id="51E30B6AA3E2A40BE8413EC8A8BF24A9" pageId="0" pageNumber="63">Other material examined.</paragraph>
<paragraph id="EB334EB9639A98A8E2F77284FCEAA9EB" pageId="0" pageNumber="63">
1♂ 7♀ (NHMSU-HA108),
<emphasis id="504FC718565A7BD8BAB50F63125CC553" bold="true" pageId="0" pageNumber="63">China</emphasis>
: Yunnan Province, Baoshan City, Longyang District, Mangkuan Township, Zaotanghe at Baihualing Village, good subtropical broadleaf forest, in leaf litter (
<geoCoordinate id="5EBFC367359E97DACA0C323C515A25F7" degrees="25.30118" direction="north" orientation="latitude" precision="1" value="25.30118">25.30118°N</geoCoordinate>
,
<geoCoordinate id="F92E5CC3B50E9F2CEDDBFEB66F4B1ACF" degrees="98.79397" direction="east" orientation="longitude" precision="1" value="98.79397">98.79397°E</geoCoordinate>
; 1802 m), 18.VI.2016, Y. Li leg.; 1♀ (NHMSU-HA108) used for sequencing, GenBank: MT992005, same data as for preceding.
</paragraph>
</subSubSection>
<subSubSection id="2C95873194C31D0DF9445638F23680E9" pageId="0" pageNumber="63" type="diagnosis">
<paragraph id="FCAAF7150D04A9A6E4508D28FA7478E8" pageId="0" pageNumber="63">Diagnosis.</paragraph>
<paragraph id="CA9E38EB085CF1729649DD1ADC6FCEC9" pageId="0" pageNumber="63">
The male of
<taxonomicName id="323D6374B35C419505F288D8E551B9B9" lsidName="C. ertou" pageId="0" pageNumber="63" rank="species" species="ertou">
<emphasis id="AABD23CABDEB5E55E30FE1A7D73698CA" italics="true" pageId="0" pageNumber="63">C. ertou</emphasis>
</taxonomicName>
is similar to that of
<taxonomicName id="0A07EF91AF414C4385E8A196899A0751" lsidName="C. rostriformis" pageId="0" pageNumber="63" rank="species" species="rostriformis">
<emphasis id="9650C1B31F1C1AD07B2FE4A5F636BECB" italics="true" pageId="0" pageNumber="63">C. rostriformis</emphasis>
</taxonomicName>
sp. nov. but can be distinguished by the narrower embolic base, wider in the latter (Fig.
<figureCitation id="4CC3F7A9792D315E3AC511BAB8A8E161" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Crassignatha ertou Miller, Griswold &amp; Yin, 2009 A male palp, prolateral B male palp, ventral C male palp, retrolateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.10 mm (A-G)." figureDoi="10.3897/zookeys.988.56188.figure8" httpUri="https://binary.pensoft.net/fig/472593" pageId="0" pageNumber="63">8A</figureCitation>
). The female of
<taxonomicName id="D71E0FFF4C76602114F299AD4EE4F1A6" lsidName="C. ertou" pageId="0" pageNumber="63" rank="species" species="ertou">
<emphasis id="1F7F267FE4FA58D6F560788B51305354" italics="true" pageId="0" pageNumber="63">C. ertou</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="2B5C7D88DB54ECC99D10DFB74C1FAC9A" lsidName="C. yinzhi" pageId="0" pageNumber="63" rank="species" species="yinzhi">
<emphasis id="9AA6ADDF24AFB68DF4E69133600E0F5A" italics="true" pageId="0" pageNumber="63">C. yinzhi</emphasis>
</taxonomicName>
but differs by the copulatory ducts having four sharp turns, but only two in the latter (Fig.
<figureCitation id="6D0FC61215D3138BCE17B99A405B7467" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Crassignatha ertou Miller, Griswold &amp; Yin, 2009 A male palp, prolateral B male palp, ventral C male palp, retrolateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.10 mm (A-G)." figureDoi="10.3897/zookeys.988.56188.figure8" httpUri="https://binary.pensoft.net/fig/472593" pageId="0" pageNumber="63">8D-G</figureCitation>
).
</paragraph>
<caption id="B79E05AAD4219D939E0CF8730AEB678B" doi="10.3897/zookeys.988.56188.figure8" httpUri="https://binary.pensoft.net/fig/472593" pageId="0" pageNumber="63" start="Figure 8" startId="F8">
<paragraph id="10F4BCB9440C8B2F68C18C4660B5AF6D" pageId="0" pageNumber="63">
<emphasis id="C40AA1E2471BB4C6A5E43633562A2DD3" bold="true" pageId="0" pageNumber="63">Figure 8.</emphasis>
<taxonomicName id="CA490DCFD27054DB52E63EB1C305D3E8" authorityName="Miller, Griswold &amp; Yin" authorityYear="2009" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crassignatha ertou" order="Araneae" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="ertou">
<emphasis id="F3F7AB45B7C9013ADD552A8F83AEAB74" italics="true" pageId="0" pageNumber="63">Crassignatha ertou</emphasis>
</taxonomicName>
Miller, Griswold &amp; Yin, 2009
<emphasis id="AF62E78B7F9351A76087A8D9D15A5EF2" bold="true" pageId="0" pageNumber="63">A</emphasis>
male palp, prolateral
<emphasis id="6091E01222E2BC3B5E5899B846C20111" bold="true" pageId="0" pageNumber="63">B</emphasis>
male palp, ventral
<emphasis id="47E8386532E806B7D015117677F9B8D2" bold="true" pageId="0" pageNumber="63">C</emphasis>
male palp, retrolateral
<emphasis id="3289BF811C6B455D5024514B482C5F60" bold="true" pageId="0" pageNumber="63">D</emphasis>
epigyne, ventral
<emphasis id="8E3215E6927ACDEC2ACF2766E1BB3545" bold="true" pageId="0" pageNumber="63">E</emphasis>
epigyne, lateral
<emphasis id="992F252C483EED4766E439F9B783B3EF" bold="true" pageId="0" pageNumber="63">F</emphasis>
vulva, ventral
<emphasis id="B07E803F91FD282A2598CDA24F92A56B" bold="true" pageId="0" pageNumber="63">G</emphasis>
vulva, dorsal. Scale bars: 0.10 mm (
<emphasis id="7BA99CBD879466CA38F52EDBAA12BE58" bold="true" pageId="0" pageNumber="63">A-G</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="4F4196619DB8C58DBD466A7BE67CB44F" pageId="0" pageNumber="63" type="description">
<paragraph id="A010B57A2EC12723D744CD604049114B" pageId="0" pageNumber="63">Description.</paragraph>
<paragraph id="CA5541CB16D40EC675EBB7C29BC2D2C6" pageId="0" pageNumber="63">
See
<bibRefCitation id="C1AD435C5AA2BB621C4BACDEA2AE399C" DOI="https://doi.org/10.3897/zookeys.11.160" author="Miller, JA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="63" pagination="9 - 195" refId="B15" refString="Miller, JA, Griswold, CE, Yin, CM, 2009. The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae, Araneoidea): systematics and diversity of micro-orbweavers. ZooKeys 11: 9 - 195, DOI: https://doi.org/10.3897/zookeys.11.160" title="The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae, Araneoidea): systematics and diversity of micro-orbweavers." url="https://doi.org/10.3897/zookeys.11.160" volume="11" year="2009">Miller et al. (2009)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="777EB31DCB23F6D944B62BEEDFE10DE9" pageId="0" pageNumber="63" type="distribution">
<paragraph id="E700455CBBFA8EC0E417CB9960080070" pageId="0" pageNumber="63">Distribution.</paragraph>
<paragraph id="BF3E686D2D5FF639F16545521D29C13C" pageId="0" pageNumber="63">
China (Yunnan) (Fig.
<figureCitation id="94A54A0F3DCF22CA9A679A5AE1B3D993" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Distribution records of Crassignatha spp. in Asia. 1 C. baihua sp. nov. 2 C. bangbie sp. nov. 3 C. bicorniventris 4 C. changyan sp. nov. 5 C. danaugirangensis 6 C. dongnai sp. nov. 7 C. ertou 8 C. gucheng sp. nov. 9 C. gudu 10 C. haeneli 11 C. mengla sp. nov. 12 C. nantou sp. nov. 13 C. nasalis sp. nov. 14 C. pianma 15 C. quadriventris 16 C. quanqu 17 C. rostriformis sp. nov. 18 C. shiluensis 19 C. shunani sp. nov. 20 C. si sp. nov. 21 C. thamphra sp. nov. 22 C. xichou sp. nov. 23 C. yamu 24 C. yinzhi." figureDoi="10.3897/zookeys.988.56188.figure38" httpUri="https://binary.pensoft.net/fig/472623" pageId="0" pageNumber="63">38</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>