468 lines
58 KiB
XML
468 lines
58 KiB
XML
<document id="5B62AB5E56A6ADF8ABF2C4FD2E53CD2F" ID-CLB-Dataset="39078" ID-DOI="10.13102/sociobiology.v62i4.845" ID-GBIF-Dataset="2c55f0dc-857e-4db3-9b62-03524f872198" ID-Zenodo-Dep="269908" IM.bibliography_requiresApprovalFor="plazi" IM.metadata_requiresApprovalFor="plazi" IM.taxonomicNames_requiresApprovalFor="plazi" checkinTime="1463064616168" checkinUser="plazi" docAuthor="Ama JOma & Wp MacKay" docDate="2015" docId="100B5B7CFF62FF91FCBAE0A8FE0EFCA7" docLanguage="en" docName="joma__mackay_2015_sociobiology_revision_of_the_african_ants_of_the_bothroponera_pumicosa_species_complex_-hymenoptera_formicidae_ponerin.pdf" docOrigin="Sociobiology 62 (4)" docStyle="DocumentStyle{}" docTitle="Bothroponera berthoudi Forel" docType="treatment" docVersion="7" lastPageNumber="547" masterDocId="EC322304FF65FF98FF86E02AFFABFFAD" masterDocTitle="Revision of the African Ants of the Bothroponera pumicosa Species Complex (Hymenoptera: Formicidae: Ponerinae)" masterLastPageNumber="563" masterPageNumber="538" pageNumber="545" updateTime="1698662492046" updateUser="ExternalLinkService">
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<mods:mods id="912404DC966E63A271655039F45279CE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="7C992DC8D5AC671C774C04D12E49D354">
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<mods:title id="BAAE4ED49F731BCFE0E2A896083D1A0D">Revision of the African Ants of the Bothroponera pumicosa Species Complex (Hymenoptera: Formicidae: Ponerinae)</mods:title>
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<mods:name id="0EAD2599E287C6DC7444F4C4FF19B509" type="personal">
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<mods:role id="F1B0412718FD0567B791A42916C96BA0">
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<mods:roleTerm id="E5E479791CBA8C472773D72F55EF5916">Author</mods:roleTerm>
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<mods:namePart id="E495DF7272ACCCA0228F03AAB6DD3AEA">Ama JOma</mods:namePart>
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<mods:name id="7116B54D5C1B55024CABDC5256AE7AAA" type="personal">
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<mods:roleTerm id="95A1713AA73225D6B72F58B6FC9D0AA9">Author</mods:roleTerm>
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<mods:namePart id="40C806C01F4EE54EF4C502A68AE452DF">Wp MacKay</mods:namePart>
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<mods:typeOfResource id="BC5DC88EC5AE5FB3FE27A5C04C6AC1BB">text</mods:typeOfResource>
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<mods:titleInfo id="3475A1979BF24A406A497D5B4C809C27">
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<mods:title id="E18B76DE29E600B8BBAD56E39E331969">Sociobiology</mods:title>
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</mods:titleInfo>
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<mods:part id="5144C74EBCAF9CEFD9AFE1D000B2386B">
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<mods:date id="862038EFEE7BE1451C913A3B5EE0D31B">2015</mods:date>
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<mods:detail id="E81A15F0DD9AAF8F8EE9E2BAE63C519D" type="volume">
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<mods:number id="B5F8E58EB288F7D26F3C8CBA06D38849">62</mods:number>
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</mods:detail>
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<mods:detail id="12FC3B7E1C2C3B65203ED026AD12CE89" type="issue">
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<mods:number id="0D615305CDCD236800D757A2BE758A9D">4</mods:number>
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<mods:start id="2732B3138F0A285AA0F8FE831AD29B4D">538</mods:start>
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<mods:classification id="49CB5D7E29E29A029D42C410BF21C388">journal article</mods:classification>
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<mods:identifier id="0EC026E6BFBA52819D74EBD6DED4F6CE" type="CLB-Dataset">39078</mods:identifier>
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<mods:identifier id="9288AA130A3DFC8A2EC5709BA9186127" type="DOI">10.13102/sociobiology.v62i4.845</mods:identifier>
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<treatment id="100B5B7CFF62FF91FCBAE0A8FE0EFCA7" ID-DOI="http://doi.org/10.5281/zenodo.6102147" ID-GBIF-Taxon="120677596" ID-Zenodo-Dep="6102147" LSID="urn:lsid:plazi:treatment:100B5B7CFF62FF91FCBAE0A8FE0EFCA7" httpUri="http://treatment.plazi.org/id/100B5B7CFF62FF91FCBAE0A8FE0EFCA7" lastPageId="9" lastPageNumber="547" pageId="7" pageNumber="545">
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<subSubSection id="D0B8B9E1FF62FF9FFCBAE0A8FB37FF37" box="[828,1180,130,154]" pageId="7" pageNumber="545" type="nomenclature">
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<paragraph id="981DEA6AFF62FF9FFCBAE0A8FB37FF37" blockId="7.[828,1180,130,189]" box="[828,1180,130,154]" pageId="7" pageNumber="545">
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<heading id="C3555D06FF62FF9FFCBAE0A8FB37FF37" bold="true" box="[828,1180,130,154]" fontSize="10" level="1" pageId="7" pageNumber="545" reason="6">
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<taxonomicName id="5FA291E9FF62FF9FFCBAE0A8FB37FF37" authority="Forel" authorityName="Forel" box="[828,1180,130,154]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="545" phylum="Arthropoda" rank="species" species="berthoudi">
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||
<emphasis id="AAD63678FF62FF9FFCBAE0A8FB37FF37" bold="true" box="[828,1180,130,154]" pageId="7" pageNumber="545">
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<emphasis id="AAD63678FF62FF9FFCBAE0A8FBECFF37" bold="true" box="[828,1095,130,154]" italics="true" pageId="7" pageNumber="545">Bothroponera berthoudi</emphasis>
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(Forel)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="D0B8B9E1FF62FF9FFCBAE08FFE9BF863" pageId="7" pageNumber="545" type="description">
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<paragraph id="981DEA6AFF62FF9FFCBAE08FFE9BF863" blockId="7.[828,1180,130,189]" lastBlockId="7.[128,783,1904,1998]" pageId="7" pageNumber="545">
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Figures 6, 7 and
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<figureCitation id="0099F6EFFF62FF9FFC68E08FFB91FF10" box="[1006,1082,165,189]" captionStart="Plate 2" captionStartId="8.[805,857,1542,1563]" captionTargetBox="[805,1459,357,1519]" captionTargetId="figure@8.[805,1459,354,1521]" captionTargetPageId="8" captionText="Plate 2: Bothroponera berthoudi worker." httpUri="https://zenodo.org/record/269916/files/figure.png" pageId="7" pageNumber="545">Plate 2</figureCitation>
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;
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<figureCitation id="0099F6EFFF62FF9FFBC0E08CFB21FF10" box="[1094,1162,165,189]" captionStart="Map 3" captionStartId="9.[129,177,1982,2003]" captionTargetBox="[129,775,1383,1963]" captionTargetId="figure@9.[129,783,1378,1963]" captionTargetPageId="9" captionText="Map 3. The distribution of B. berthoudi." httpUri="https://zenodo.org/record/269918/files/figure.png" pageId="7" pageNumber="545">Map 3</figureCitation>
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The sculpturing of the head is shown only on the right or left side, to allow the illustration of the hairs on the other side of the head.
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</paragraph>
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</subSubSection>
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<subSubSection id="D0B8B9E1FF62FF9FFC01E0C1FAFAFD0F" pageId="7" pageNumber="545" type="reference_group">
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||
<paragraph id="981DEA6AFF62FF9FFC01E0C1FA1DFE58" blockId="7.[828,1483,235,1992]" pageId="7" pageNumber="545">
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||
<taxonomicName id="5FA291E9FF62FF9FFC01E0C1FA61FEAE" ID-CoL="4BQTC" authority="Forel, 1901: 344" authorityName="Forel" authorityPageNumber="344" authorityYear="1901" box="[903,1482,235,259]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="545" phylum="Arthropoda" rank="species" species="berthoudi" subGenus="Bothroponera">
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<emphasis id="AAD63678FF62FF9FFC01E0C1FBB3FEAE" box="[903,1048,235,259]" italics="true" pageId="7" pageNumber="545">Pachycondyla</emphasis>
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(
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<emphasis id="AAD63678FF62FF9FFBA2E0C1FB18FEAE" box="[1060,1203,235,259]" italics="true" pageId="7" pageNumber="545">Bothroponera</emphasis>
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)
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<emphasis id="AAD63678FF62FF9FFB39E0C1FA89FEAE" box="[1215,1314,235,259]" italics="true" pageId="7" pageNumber="545">berthoudi</emphasis>
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<bibRefCitation id="FC33979BFF62FF9FFAA0E0C1FA37FEAE" author="Forel, A." box="[1318,1436,235,259]" journalOrPublisher="Revue Suisse de Zoologie" pageId="7" pageNumber="545" pagination="325 - 353" part="9" refString="Forel, A. (1901). Nouvelles especes de Ponerinae. (Avec un nouveau sous-genre et une espece nouvelle d'Eciton). Revue Suisse de Zoologie, 9: 325 - 353." title="Nouvelles especes de Ponerinae. (Avec un nouveau sous-genre et une espece nouvelle d'Eciton)" type="journal article" year="1901">Forel, 1901</bibRefCitation>
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: 344
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</taxonomicName>
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(w), South Africa, Valdezia, Transvaal;
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<bibRefCitation id="FC33979BFF62FF9FFB53E124FAF6FE88" author="Emery, C." box="[1237,1373,269,294]" journalOrPublisher="Genera Insectorum" pageId="7" pageNumber="545" pagination="1 - 125" part="118" refString="Emery, C. (1911). Hymenoptera. Fam. Formicidae. Subfam. Ponerinae. Genera Insectorum, 118: 1 - 125." title="Hymenoptera. Fam. Formicidae. Subfam. Ponerinae" type="journal article" year="1911">Emery, 1911</bibRefCitation>
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: 76;
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<bibRefCitation id="FC33979BFF62FF9FFA0BE127FCD6FEE5" author="Forel, A." journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="7" pageNumber="545" pagination="108 - 147" part="57" refString="Forel, A. (1913 a). Fourmis de Rhodesie, etc. recoltees par M. G. Arnold, le Dr. H. Brauns et K. Fikendey. Annales de la Societe Entomologique de Belgique, 57: 108 - 147." title="Fourmis de Rhodesie, etc. recoltees par M. G. Arnold, le Dr. H. Brauns et K. Fikendey" type="journal article" year="1913" yearSuffix="a">Forel, 1913a</bibRefCitation>
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: 306 (m), Willwomore, colonie du Cap;
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<taxonomicName id="5FA291E9FF62FF9FFABCE11AFA39FEC6" ID-CoL="4BQTC" authority="Forel, 1913b: 109" authorityName="Forel" authorityPageNumber="109" authorityYear="1913" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="545" phylum="Arthropoda" rank="subSpecies" species="pumicosa" subGenus="Bothroponera" subSpecies="berthoudi">
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<emphasis id="AAD63678FF62FF9FFABCE11AFA62FEE5" box="[1338,1481,304,328]" italics="true" pageId="7" pageNumber="545">Pachycondyla</emphasis>
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(
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<emphasis id="AAD63678FF62FF9FFCC2E179FC79FEC6" box="[836,978,339,363]" italics="true" pageId="7" pageNumber="545">Bothroponera</emphasis>
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)
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<emphasis id="AAD63678FF62FF9FFC6EE179FB12FEC6" box="[1000,1209,339,363]" italics="true" pageId="7" pageNumber="545">pumicosa berthoudi</emphasis>
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:
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<bibRefCitation id="FC33979BFF62FF9FFB48E179FAF1FEC6" author="Forel, A." box="[1230,1370,339,363]" journalOrPublisher="Revue Zoologique Africaine (Brussels)" pageId="7" pageNumber="545" pagination="306 - 351" part="2" refString="Forel, A. (1913 b). Formicides du Congo Belge recoltes par MM. Bequaert, Luja, etc. Revue Zoologique Africaine (Brussels), 2: 306 - 351." title="Formicides du Congo Belge recoltes par MM. Bequaert, Luja, etc" type="journal article" year="1913" yearSuffix="b">Forel, 1913b</bibRefCitation>
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: 109
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</taxonomicName>
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(w), Willowmore [Willwomore], Cap;
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<taxonomicName id="5FA291E9FF62FF9FFB18E15FFA3DFE20" box="[1182,1430,373,397]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="545" phylum="Arthropoda" rank="species" species="pumicosa">
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<emphasis id="AAD63678FF62FF9FFB18E15FFA3DFE20" box="[1182,1430,373,397]" italics="true" pageId="7" pageNumber="545">Bothroponera pumicosa</emphasis>
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</taxonomicName>
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race
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<taxonomicName id="5FA291E9FF62FF9FFCBAE1B2FBC9FE1D" ID-CoL="4BQTC" authority="Arnold, 1952: 460" authorityName="Arnold" authorityPageNumber="460" authorityYear="1952" box="[828,1122,408,432]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="545" phylum="Arthropoda" rank="species" species="berthoudi">
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<emphasis id="AAD63678FF62FF9FFCBAE1B2FC37FE1D" box="[828,924,408,432]" italics="true" pageId="7" pageNumber="545">berthoudi</emphasis>
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:
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<bibRefCitation id="FC33979BFF62FF9FFC2FE1B2FB9AFE1D" author="Arnold, G." box="[937,1073,408,432]" journalOrPublisher="Occasional Papers of the National Museums of Southern Rhodesia" pageId="7" pageNumber="545" pagination="460 - 493" part="2" refString="Arnold, G. (1952). New species of African Hymenoptera. No. 10. Occasional Papers of the National Museums of Southern Rhodesia, 2: 460 - 493." title="New species of African Hymenoptera. No. 10" type="journal article" year="1952">Arnold, 1952</bibRefCitation>
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: 460
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</taxonomicName>
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, considered
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<taxonomicName id="5FA291E9FF62FF9FFB58E1B2FA96FE1D" ID-CoL="4BQTC" box="[1246,1341,408,432]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="545" phylum="Arthropoda" rank="species" species="berthoudi">
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<emphasis id="AAD63678FF62FF9FFB58E1B2FA96FE1D" box="[1246,1341,408,432]" italics="true" pageId="7" pageNumber="545">berthoudi</emphasis>
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</taxonomicName>
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to be a junior synonym of
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<taxonomicName id="5FA291E9FF62FF9FFC32E191FBBEFE7E" box="[948,1045,443,467]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="545" phylum="Arthropoda" rank="species" species="strigulosa">
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<emphasis id="AAD63678FF62FF9FFC32E191FBBEFE7E" box="[948,1045,443,467]" italics="true" pageId="7" pageNumber="545">strigulosa</emphasis>
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</taxonomicName>
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;
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<taxonomicName id="5FA291E9FF62FF9FFBA7E191FACFFE7F" ID-CoL="4BQTC" authority="Brown" authorityName="Brown" box="[1057,1380,443,467]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="545" phylum="Arthropoda" rank="species" species="berthoudi">
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<emphasis id="AAD63678FF62FF9FFBA7E191FABEFE7E" box="[1057,1301,443,467]" italics="true" pageId="7" pageNumber="545">Pachycondyla berthoudi</emphasis>
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: Brown
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</taxonomicName>
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in
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<bibRefCitation id="FC33979BFF62FF9FFA06E191FCDAFE58" author="Bolton, B." journalOrPublisher="Cambridge, Massachusetts, Harvard University Press, USA" pageId="7" pageNumber="545" pagination="504" refString="Bolton, B. (1995). A New General Catalogue of the Ants of the World (pp 504) Cambridge, Massachusetts, Harvard University Press, USA." title="A New General Catalogue of the Ants of the World (pp" type="book chapter" year="1995">Bolton, 1995</bibRefCitation>
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: 303;
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<taxonomicName id="5FA291E9FF62FF9FFC2DE1F7FA19FE58" authority="Joma and Mackay, 2013: 3" authorityName="Joma and Mackay" authorityPageNumber="3" authorityYear="2013" box="[939,1458,477,501]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="545" phylum="Arthropoda" rank="species" species="berthoudi">
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<emphasis id="AAD63678FF62FF9FFC2DE1F7FB31FE58" box="[939,1178,477,501]" italics="true" pageId="7" pageNumber="545">Bothroponera berthoudi</emphasis>
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:
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<bibRefCitation id="FC33979BFF62FF9FFB21E1F4FA30FE58" author="Joma, A. M. A. & Mackay, W. P." box="[1191,1435,477,501]" journalOrPublisher="Psyche" pageId="7" pageNumber="545" pagination="1 - 8" part="2013" publicationUrl="http://dx.doi.org/10.1155/2013/917847" refString="Joma, A. M. A. & Mackay, W. P. (2013). A new species of Afrotropical ants in the genus Bothroponera (Hymenoptera: Formicidae: Ponerinae). Psyche 2013: 1 - 8. doi: 10.1155 / 2013 / 917847" title="A new species of Afrotropical ants in the genus Bothroponera (Hymenoptera: Formicidae: Ponerinae)" type="journal article" year="2013">Joma and Mackay, 2013</bibRefCitation>
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: 3
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</taxonomicName>
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.
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</paragraph>
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<subSection id="F12DF381FF62FF9FFC01E22AFAFAFD0F" pageId="7" pageNumber="545" type="multiple">
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<paragraph id="981DEA6AFF62FF9FFC01E22AFAFAFD0F" blockId="7.[828,1483,235,1992]" pageId="7" pageNumber="545">
|
||
<taxonomicName id="5FA291E9FF62FF9FFC01E22AFAFBFDB5" authority="Arnold, 1947: 131" authorityName="Arnold" authorityPageNumber="131" authorityYear="1947" box="[903,1360,512,536]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="545" phylum="Arthropoda" rank="species" species="variolosa">
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<emphasis id="AAD63678FF62FF9FFC01E22AFB28FDB5" box="[903,1155,512,536]" italics="true" pageId="7" pageNumber="545">Bothroponera variolosa</emphasis>
|
||
<bibRefCitation id="FC33979BFF62FF9FFB0AE22AFAB7FDB5" author="Arnold, G." box="[1164,1308,512,536]" journalOrPublisher="Occasional Papers of the National Museums of Southern Rhodesia" pageId="7" pageNumber="545" pagination="131 - 167" part="2" refString="Arnold, G. (1947). New species of African Hymenoptera. No. 7. Occasional Papers of the National Museums of Southern Rhodesia, 2: 131 - 167." title="New species of African Hymenoptera. No. 7" type="journal article" year="1947">Arnold, 1947</bibRefCitation>
|
||
: 131
|
||
</taxonomicName>
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(w), South Africa, Mariepskop, Transvaal;
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<taxonomicName id="5FA291E9FF62FF9FFBFAE209FC50FDF0" authority="Arnold, 1960: 452" authorityName="Arnold" authorityPageNumber="452" authorityYear="1960" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="545" phylum="Arthropoda" rank="subSpecies" species="cariosa" subSpecies="variolosa">
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||
<emphasis id="AAD63678FF62FF9FFBFAE209FA69FD96" box="[1148,1474,547,571]" italics="true" pageId="7" pageNumber="545">Bothroponera cariosa variolosa</emphasis>
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||
:
|
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<bibRefCitation id="FC33979BFF62FF9FFCBAE26FFC63FDF0" author="Arnold, G." box="[828,968,581,605]" journalOrPublisher="Occasional Papers of the National Museum of Southern Rhodesia" pageId="7" pageNumber="545" pagination="452 - 488" part="24" refString="Arnold, G. (1960). New species of African Hymenoptera No. 15. Occasional Papers of the National Museum of Southern Rhodesia, 24 B: 452 - 488." title="New species of African Hymenoptera No. 15" type="journal article" year="1960">Arnold, 1960</bibRefCitation>
|
||
: 452
|
||
</taxonomicName>
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||
;
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||
<taxonomicName id="5FA291E9FF62FF9FFB82E26FFC3DFD2D" authority="Joma and Mackay, 2013: 3" authorityName="Joma and Mackay" authorityPageNumber="3" authorityYear="2013" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="545" phylum="Arthropoda" rank="species" species="variolosa">
|
||
<emphasis id="AAD63678FF62FF9FFB82E26FFB57FDF0" box="[1028,1276,581,605]" italics="true" pageId="7" pageNumber="545">Bothroponera variolosa</emphasis>
|
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:
|
||
<bibRefCitation id="FC33979BFF62FF9FFA8EE26CFCDFFD2D" author="Joma, A. M. A. & Mackay, W. P." journalOrPublisher="Psyche" pageId="7" pageNumber="545" pagination="1 - 8" part="2013" publicationUrl="http://dx.doi.org/10.1155/2013/917847" refString="Joma, A. M. A. & Mackay, W. P. (2013). A new species of Afrotropical ants in the genus Bothroponera (Hymenoptera: Formicidae: Ponerinae). Psyche 2013: 1 - 8. doi: 10.1155 / 2013 / 917847" title="A new species of Afrotropical ants in the genus Bothroponera (Hymenoptera: Formicidae: Ponerinae)" type="journal article" year="2013">Joma and Mackay, 2013</bibRefCitation>
|
||
: 3
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="FC33979BFF62FF9FFC2AE242FB59FD2D" author="Schmidt C. A. & Shattuck, S. O." box="[940,1266,616,640]" journalOrPublisher="Zootaxa, 3817" pageId="7" pageNumber="545" pagination="1 - 242" part="1" publicationUrl="http://dx.doi.org/10.11646/zootaxa.3647.2.1." refString="Schmidt C. A. & Shattuck, S. O. (2014). The higher classification of the ant subfamily Ponerinae (Hymenoptera: Formicidae), with a review of ponerine ecology and behavior. Zootaxa, 3817 (1): 1 - 242. doi: 10.11646 / zootaxa. 3647.2.1." title="The higher classification of the ant subfamily Ponerinae (Hymenoptera: Formicidae), with a review of ponerine ecology and behavior" type="journal article" year="2014">Schmidt and Shattuck, 2014</bibRefCitation>
|
||
: 77;
|
||
<taxonomicName id="5FA291E9FF62FF9FFAB1E242FC58FD0F" ID-CoL="4BQYT" authority="Brown" authorityName="Brown" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="545" phylum="Arthropoda" rank="species" species="variolosa">
|
||
<emphasis id="AAD63678FF62FF9FFAB1E242FC36FD0E" italics="true" pageId="7" pageNumber="545">Pachycondyla variolosa</emphasis>
|
||
: Brown
|
||
</taxonomicName>
|
||
, in
|
||
<bibRefCitation id="FC33979BFF62FF9FFB91E2A1FB0AFD0F" author="Bolton, B." box="[1047,1185,650,675]" journalOrPublisher="Cambridge, Massachusetts, Harvard University Press, USA" pageId="7" pageNumber="545" pagination="504" refString="Bolton, B. (1995). A New General Catalogue of the Ants of the World (pp 504) Cambridge, Massachusetts, Harvard University Press, USA." title="A New General Catalogue of the Ants of the World (pp" type="book chapter" year="1995">Bolton, 1995</bibRefCitation>
|
||
: 311 (
|
||
<emphasis id="AAD63678FF62FF9FFB67E2A1FAE2FD0F" bold="true" box="[1249,1353,650,675]" pageId="7" pageNumber="545">syn. nov.)</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSection>
|
||
</subSubSection>
|
||
<subSubSection id="D0B8B9E1FF62FF9FFC01E287FA07FC76" pageId="7" pageNumber="545" type="diagnosis">
|
||
<paragraph id="981DEA6AFF62FF9FFC01E287FA07FC76" blockId="7.[828,1483,235,1992]" pageId="7" pageNumber="545">
|
||
<emphasis id="AAD63678FF62FF9FFC01E287FC55FD68" bold="true" box="[903,1022,685,709]" pageId="7" pageNumber="545">Diagnosis:</emphasis>
|
||
The worker of
|
||
<taxonomicName id="5FA291E9FF62FF9FFB39E287FA61FD68" box="[1215,1482,685,709]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="545" phylum="Arthropoda" rank="species" species="berthoudi">
|
||
<emphasis id="AAD63678FF62FF9FFB39E287FA61FD68" box="[1215,1482,685,709]" italics="true" pageId="7" pageNumber="545">Bothroponera berthoudi</emphasis>
|
||
</taxonomicName>
|
||
is a relatively large ant (total length 9.60 – 12.00 mm). The mandibles are hairy and coarsely covered with punctures. The anterior medial margin of the clypeus is “u” shaped and slightly bent ventrally, with the disc has a raised smooth area with striae. The upper part of the raised area, between the frontal lobes is rough with a few punctulae, and with a small clypeal carinae. The clypeal wings are punctulate and obliquely striate. The scape barely reaches the posterior lateral corner of the head.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="CCDDBAE2FF62FF9FFF19E2C4FDA2FCA9" box="[159,521,750,772]" httpUri="https://zenodo.org/record/269914/files/figure.png" pageId="7" pageNumber="545" targetBox="[129,775,134,713]" targetPageId="7">
|
||
<paragraph id="981DEA6AFF62FF9FFF19E2C4FDA2FCA9" blockId="7.[159,521,750,772]" box="[159,521,750,772]" pageId="7" pageNumber="545">
|
||
<emphasis id="AAD63678FF62FF9FFF19E2C4FF4AFCAE" bold="true" box="[159,225,750,771]" pageId="7" pageNumber="545">Map 2</emphasis>
|
||
. The distribution of
|
||
<taxonomicName id="5FA291E9FF62FF9FFE21E2C5FDA9FCA9" box="[423,514,751,772]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="545" phylum="Arthropoda" rank="species" species="aspera">
|
||
<emphasis id="AAD63678FF62FF9FFE21E2C5FDA9FCA9" box="[423,514,751,772]" italics="true" pageId="7" pageNumber="545">B. aspera</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="D0B8B9E1FF62FF90FC01E3CFFE69FD2D" lastPageId="8" lastPageNumber="546" pageId="7" pageNumber="545" type="description">
|
||
<paragraph id="981DEA6AFF62FF90FC01E3CFFE69FD2D" blockId="7.[828,1483,235,1992]" lastBlockId="8.[106,760,131,1993]" lastPageId="8" lastPageNumber="546" pageId="7" pageNumber="545">
|
||
<emphasis id="AAD63678FF62FF9FFC01E3CFFBD7FC50" bold="true" box="[903,1148,997,1021]" pageId="7" pageNumber="545">Worker Description:</emphasis>
|
||
(n=8 for measurements), HL 2.00 - 2.61, HW 1.75 - 2.25, ML 1.15 - 1.45, EW 0.30 - 0.35, EL 0.30 - 0.45, SL 1.40 - 1.85, FL 2.10 - 2.60, WL 2.75 - 3.85, WPL 3.60 - 4.90, PL 0.90 - 1.30, PW 0.95 - 1.35, PH 1.30 - 1.60, CI 86.20 - 87.50, OI 17.14 - 20.00, MandI 57.50 - 55.56, SI 80.00 - 82.22, PetI 103.84 - 105.56. Mandibles smooth with about 7 teeth; head subquadrate; maximum transversal clypeal length 1.60 mm; compound eyes relatively large; length of malar space 0.40 mm; length from upper edge of eye to edge of posterior lobe 1.00 mm; maximum frontal lobes width 0.75 mm; surface of head, pronotum, mesonotum, mesopleuron, propodeum, lateropropodeum, metapleuron, densely foveolate, moderately shiny; petiolar and postpetiolar surfaces densely covered with larger foveolae than those of mesosoma, moderately shiny; cheek, sides of head, area posterior to eyes, frons covered with weakly defined striae; dorsum of fourth abdominal segment covered with shallow foveolae and striae; fifthseventh abdominal segments smooth, shiny; basalar sclerite oval in depressed surrounding area; pronotal shoulder rounded, lower margin straight (lateral view), anteroinferior pronotal process angled, inferior pronotal process rounded; mesometapleural suture developed; mesopleural-coxal excavation developed and continued with mesometapleural suture; antennae, legs, lower edges of frontal lobes, mandibles shiny; anterior face of petiolar node from dorsal view rounded, slightly narrowed anteriorly, posterior face vertical in side view, slightly concave with slight depression on medial upper margin; metapleuron rough, covered with striae, foveolae; posteropropodeum rough, slightly concave; dorsum of postpetiole densely covered with large foveolae and striae; surface of 4th abdominal segment rough, covered with large foveolae; 5th to 7th abdominal segments moderately shiny, covered with fine striae; head, dorsum of pronotum, mesonotum, propodeum covered with fine moderately long golden erect hairs; hairs moderately long (0.12 mm up to 0.25 mm) hairs on mandibles, legs, scapes; length of hairs on pronotum, mesonotum, propodeum slightly longer (0.20 - 0.25 mm); dorsum of petiole, dorsal and ventral surfaces of postpetiole, and 4th to 7th abdominal segments covered with longer erect golden hairs (0.30 - 0.35 mm); head, pronotum, mesonotum, mesopleuron, propodeum, petiole, postpetiole, entire gaster black; legs and antennae brownish black; mandibles reddish brown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="CCDDBAE2FF62FF9FFF07E6AFFE88F883" httpUri="https://zenodo.org/record/269915/files/figure.png" pageId="7" pageNumber="545" targetBox="[142,773,861,1629]" targetPageId="7">
|
||
<paragraph id="981DEA6AFF62FF9FFF07E6AFFE88F883" blockId="7.[129,783,1669,1838]" pageId="7" pageNumber="545">
|
||
<emphasis id="AAD63678FF62FF9FFF07E6AFFF18F937" bold="true" box="[129,179,1669,1691]" pageId="7" pageNumber="545">Fig 4</emphasis>
|
||
. The lateral view of a paratype worker of
|
||
<taxonomicName id="5FA291E9FF62FF9FFDC6E6AFFF6DF915" authority="Iziko" authorityName="Iziko" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="545" phylum="Arthropoda" rank="species" species="aspera">
|
||
<emphasis id="AAD63678FF62FF9FFDC6E6AFFCA5F936" box="[576,782,1669,1691]" italics="true" pageId="7" pageNumber="545">Bothroponera aspera</emphasis>
|
||
(Iziko)
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="AAD63678FF62FF9FFF48E688FEABF915" bold="true" box="[206,256,1698,1720]" pageId="7" pageNumber="545">Fig 5</emphasis>
|
||
. The head of the paratype worker of
|
||
<taxonomicName id="5FA291E9FF62FF9FFDE3E689FCA0F915" authority="Iziko" authorityName="Iziko" box="[613,779,1698,1720]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="545" phylum="Arthropoda" rank="species" species="aspera">
|
||
<emphasis id="AAD63678FF62FF9FFDE3E689FD6AF915" box="[613,705,1699,1720]" italics="true" pageId="7" pageNumber="545">B. aspera</emphasis>
|
||
(Iziko)
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="AAD63678FF62FF9FFF07E6EAFF1DF978" bold="true" box="[129,182,1728,1750]" pageId="7" pageNumber="545">Fig 6</emphasis>
|
||
. The head of the worker of
|
||
<taxonomicName id="5FA291E9FF62FF9FFE56E6EAFDE0F978" box="[464,587,1728,1749]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="545" phylum="Arthropoda" rank="species" species="berthoudi">
|
||
<emphasis id="AAD63678FF62FF9FFE56E6EAFDE0F978" box="[464,587,1728,1749]" italics="true" pageId="7" pageNumber="545">B. berthoudi</emphasis>
|
||
</taxonomicName>
|
||
from South Africa, Eastern Cape Province, Algoa-Bay (MfN).
|
||
<emphasis id="AAD63678FF62FF9FFDAAE6F7FDCAF95F" bold="true" box="[556,609,1757,1779]" pageId="7" pageNumber="545">Fig 7</emphasis>
|
||
. The lateral view of a worker of
|
||
<taxonomicName id="5FA291E9FF62FF9FFEACE6D1FE02F8A2" box="[298,425,1786,1808]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="545" phylum="Arthropoda" rank="species" species="berthoudi">
|
||
<emphasis id="AAD63678FF62FF9FFEACE6D1FE02F8A2" box="[298,425,1786,1808]" italics="true" pageId="7" pageNumber="545">B. berthoudi</emphasis>
|
||
</taxonomicName>
|
||
from South Africa, Eastern Cape Province (MfN).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="D0B8B9E1FF6DFF90FF33E2A1FB91FEE5" pageId="8" pageNumber="546" type="discussion">
|
||
<paragraph id="981DEA6AFF6DFF90FF33E2A1FD35FBEE" blockId="8.[106,760,131,1993]" pageId="8" pageNumber="546">
|
||
<emphasis id="AAD63678FF6DFF90FF33E2A1FEE0FD0E" bold="true" box="[181,331,651,675]" pageId="8" pageNumber="546">Comparison:</emphasis>
|
||
<bibRefCitation id="FC33979BFF6DFF90FED5E2A1FE45FD0E" author="Arnold, G." box="[339,494,651,675]" journalOrPublisher="Occasional Papers of the National Museums of Southern Rhodesia" pageId="8" pageNumber="546" pagination="460 - 493" part="2" refString="Arnold, G. (1952). New species of African Hymenoptera. No. 10. Occasional Papers of the National Museums of Southern Rhodesia, 2: 460 - 493." title="New species of African Hymenoptera. No. 10" type="journal article" year="1952">Arnold (1952)</bibRefCitation>
|
||
considered
|
||
<taxonomicName id="5FA291E9FF6DFF90FDF7E2A6FD53FD0E" box="[625,760,651,675]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="berthoudi">
|
||
<emphasis id="AAD63678FF6DFF90FDF7E2A6FD53FD0E" box="[625,760,651,675]" italics="true" pageId="8" pageNumber="546">B. berthoudi</emphasis>
|
||
</taxonomicName>
|
||
to be a race of
|
||
<taxonomicName id="5FA291E9FF6DFF90FE90E284FE37FD68" box="[278,412,686,709]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="pumicosa">
|
||
<emphasis id="AAD63678FF6DFF90FE90E284FE37FD68" box="[278,412,686,709]" italics="true" pageId="8" pageNumber="546">B. pumicosa</emphasis>
|
||
</taxonomicName>
|
||
, but it is clear that this species is different from
|
||
<taxonomicName id="5FA291E9FF6DFF90FEAFE2FBFE1BFD45" box="[297,432,721,744]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="pumicosa">
|
||
<emphasis id="AAD63678FF6DFF90FEAFE2FBFE1BFD45" box="[297,432,721,744]" italics="true" pageId="8" pageNumber="546">B. pumicosa</emphasis>
|
||
</taxonomicName>
|
||
in that
|
||
<taxonomicName id="5FA291E9FF6DFF90FD8CE2FBFD3AFD45" box="[522,657,721,744]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="pumicosa">
|
||
<emphasis id="AAD63678FF6DFF90FD8CE2FBFD3AFD45" box="[522,657,721,744]" italics="true" pageId="8" pageNumber="546">B. pumicosa</emphasis>
|
||
</taxonomicName>
|
||
has long hairs that cover the entire body, which is not the case in
|
||
<taxonomicName id="5FA291E9FF6DFF90FD67E2DEFF79FC83" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="berthoudi">
|
||
<emphasis id="AAD63678FF6DFF90FD67E2DEFF79FC83" italics="true" pageId="8" pageNumber="546">B. berthoudi</emphasis>
|
||
</taxonomicName>
|
||
. There are other slight differences between the two species. The anterior medial raised area of the clypeus of
|
||
<taxonomicName id="5FA291E9FF6DFF90FD67E313FF7BFCDE" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="pumicosa">
|
||
<emphasis id="AAD63678FF6DFF90FD67E313FF7BFCDE" italics="true" pageId="8" pageNumber="546">B. pumicosa</emphasis>
|
||
</taxonomicName>
|
||
forms a partial carina on the posterior part and a smooth narrowed area on the grooved anterior part. The same character is found in
|
||
<taxonomicName id="5FA291E9FF6DFF90FED3E38BFE76FC15" box="[341,477,928,952]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="berthoudi">
|
||
<emphasis id="AAD63678FF6DFF90FED3E38BFE76FC15" box="[341,477,928,952]" italics="true" pageId="8" pageNumber="546">B. berthoudi</emphasis>
|
||
</taxonomicName>
|
||
, but the upper part forms striae instead of a partial carina, the lower part is smooth but wider than that of
|
||
<taxonomicName id="5FA291E9FF6DFF90FEEFE3CCFE5BFC50" box="[361,496,998,1021]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="pumicosa">
|
||
<emphasis id="AAD63678FF6DFF90FEEFE3CCFE5BFC50" box="[361,496,998,1021]" italics="true" pageId="8" pageNumber="546">B. pumicosa</emphasis>
|
||
</taxonomicName>
|
||
. The mandibles have 7 teeth in
|
||
<taxonomicName id="5FA291E9FF6DFF90FF39E423FEEFFB8D" box="[191,324,1032,1056]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="berthoudi">
|
||
<emphasis id="AAD63678FF6DFF90FF39E423FEEFFB8D" box="[191,324,1032,1056]" italics="true" pageId="8" pageNumber="546">B. berthoudi</emphasis>
|
||
</taxonomicName>
|
||
, similar to the other
|
||
<taxonomicName id="5FA291E9FF6DFF90FDA7E423FD0FFB8D" box="[545,676,1033,1056]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="pumicosa">
|
||
<emphasis id="AAD63678FF6DFF90FDA7E423FD0FFB8D" box="[545,676,1033,1056]" italics="true" pageId="8" pageNumber="546">B. pumicosa</emphasis>
|
||
</taxonomicName>
|
||
species complex members, whereas
|
||
<taxonomicName id="5FA291E9FF6DFF90FE1FE406FDB6FBEE" box="[409,541,1068,1091]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="pumicosa">
|
||
<emphasis id="AAD63678FF6DFF90FE1FE406FDB6FBEE" box="[409,541,1068,1091]" italics="true" pageId="8" pageNumber="546">B. pumicosa</emphasis>
|
||
</taxonomicName>
|
||
has 8 teeth.
|
||
</paragraph>
|
||
<paragraph id="981DEA6AFF6DFF90FF33E464FD4EFB5D" blockId="8.[106,760,131,1993]" pageId="8" pageNumber="546">
|
||
<taxonomicName id="5FA291E9FF6DFF90FF33E464FE61FBCB" box="[181,458,1102,1126]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="laevissima">
|
||
<emphasis id="AAD63678FF6DFF90FF33E464FE61FBCB" box="[181,458,1102,1126]" italics="true" pageId="8" pageNumber="546">Bothroponera laevissima</emphasis>
|
||
</taxonomicName>
|
||
has the same “u” shaped clypeus as does
|
||
<taxonomicName id="5FA291E9FF6DFF90FEA1E45BFE19FB25" box="[295,434,1136,1160]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="berthoudi">
|
||
<emphasis id="AAD63678FF6DFF90FEA1E45BFE19FB25" box="[295,434,1136,1160]" italics="true" pageId="8" pageNumber="546">B. berthoudi</emphasis>
|
||
</taxonomicName>
|
||
, but is easily recognized by the unique sculpture: shiny with scattered punctures. The mandibles have 7 teeth in
|
||
<taxonomicName id="5FA291E9FF6DFF90FE1EE49CFD89FB63" box="[408,546,1206,1230]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="berthoudi">
|
||
<emphasis id="AAD63678FF6DFF90FE1EE49CFD89FB63" box="[408,546,1206,1230]" italics="true" pageId="8" pageNumber="546">B. berthoudi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="5FA291E9FF6DFF90FDD8E49CFD5AFB63" box="[606,753,1206,1230]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="laevissima">
|
||
<emphasis id="AAD63678FF6DFF90FDD8E49CFD5AFB63" box="[606,753,1206,1230]" italics="true" pageId="8" pageNumber="546">B. laevissima</emphasis>
|
||
</taxonomicName>
|
||
, similar to the other
|
||
<taxonomicName id="5FA291E9FF6DFF90FEBDE4F3FE15FB5D" box="[315,446,1241,1264]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="pumicosa">
|
||
<emphasis id="AAD63678FF6DFF90FEBDE4F3FE15FB5D" box="[315,446,1241,1264]" italics="true" pageId="8" pageNumber="546">B. pumicosa</emphasis>
|
||
</taxonomicName>
|
||
species complex members.
|
||
</paragraph>
|
||
<paragraph id="981DEA6AFF6DFF90FF33E4D1FD6EFA6D" blockId="8.[106,760,131,1993]" pageId="8" pageNumber="546">
|
||
The body surface of
|
||
<taxonomicName id="5FA291E9FF6DFF90FE15E4D6FDBCFABE" box="[403,535,1275,1299]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="berthoudi">
|
||
<emphasis id="AAD63678FF6DFF90FE15E4D6FDBCFABE" box="[403,535,1275,1299]" italics="true" pageId="8" pageNumber="546">B. berthoudi</emphasis>
|
||
</taxonomicName>
|
||
is densely foveolate, and the fourth abdominal segment is densely covered with foveolae and striae. Basically, the members of the type series consistently differ from the rest of
|
||
<taxonomicName id="5FA291E9FF6DFF90FE4CE54EFDE2FAD6" box="[458,585,1380,1403]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="pumicosa">
|
||
<emphasis id="AAD63678FF6DFF90FE4CE54EFDE2FAD6" box="[458,585,1380,1403]" italics="true" pageId="8" pageNumber="546">B. pumicosa</emphasis>
|
||
</taxonomicName>
|
||
species complex species in having long erect hairs on most surfaces (except the head) and in lacking a well-defined medial clypeal carina.
|
||
</paragraph>
|
||
<paragraph id="981DEA6AFF6DFF90FF33E5E1FDD6F82E" blockId="8.[106,760,131,1993]" pageId="8" pageNumber="546">
|
||
The specific epithet “
|
||
<taxonomicName id="5FA291E9FF6DFF90FE12E5E1FE57FA4E" box="[404,508,1483,1507]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="berthoudi">
|
||
<emphasis id="AAD63678FF6DFF90FE12E5E1FE57FA4E" box="[404,508,1483,1507]" italics="true" pageId="8" pageNumber="546">berthoudi</emphasis>
|
||
</taxonomicName>
|
||
” was first used by
|
||
<bibRefCitation id="FC33979BFF6DFF90FD39E5E1FF1AF9AB" author="Forel, A." journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="8" pageNumber="546" part="34" refString="Forel, A. (1890). Aenictus - Typhlatta decouverte de M. Wroughton. Nouveaux genres de Formicides. Annales de la Societe Entomologique de Belgique, 34: cii-cxiv." title="Aenictus - Typhlatta decouverte de M. Wroughton. Nouveaux genres de Formicides" type="journal volume" year="1890">Forel (1890)</bibRefCitation>
|
||
as a name for
|
||
<taxonomicName id="5FA291E9FF6DFF90FEDEE5C4FDDFF9AB" box="[344,628,1518,1542]" class="Insecta" family="Formicidae" genus="Ophthalmopone" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="berthoudi">
|
||
<emphasis id="AAD63678FF6DFF90FEDEE5C4FDDFF9AB" box="[344,628,1518,1542]" italics="true" pageId="8" pageNumber="546">Ophthalmopone berthoudi</emphasis>
|
||
</taxonomicName>
|
||
from South Africa. Later, he used it again to identify and describe the male of
|
||
<taxonomicName id="5FA291E9FF6DFF90FF0CE619FE03F9E6" box="[138,424,1587,1611]" class="Insecta" family="Formicidae" genus="Ophthalmopone" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="berthoudi">
|
||
<emphasis id="AAD63678FF6DFF90FF0CE619FE03F9E6" box="[138,424,1587,1611]" italics="true" pageId="8" pageNumber="546">Ophthalmopone berthoudi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="FC33979BFF6DFF90FE3EE619FD53F9E6" author="Wheeler, G. C. & Wheeler, J." box="[440,760,1587,1611]" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="8" pageNumber="546" pagination="386 - 394" part="73" refString="Wheeler, G. C. & Wheeler, J. (1971). The larvae of the ant genus Bothroponera (Hymenoptera: Formicidae). Proceedings of the Entomological Society of Washington, 73: 386 - 394." title="The larvae of the ant genus Bothroponera (Hymenoptera: Formicidae)" type="journal article" year="1971">Wheeler and Wheeler (1971)</bibRefCitation>
|
||
used the same name to identify larvae of
|
||
<taxonomicName id="5FA291E9FF6DFF90FD92E67CFD30F9C3" box="[532,667,1622,1646]" class="Insecta" family="Formicidae" genus="Ophthalmopone" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="berthoudi">
|
||
<emphasis id="AAD63678FF6DFF90FD92E67CFD30F9C3" box="[532,667,1622,1646]" italics="true" pageId="8" pageNumber="546">O. berthoudi</emphasis>
|
||
</taxonomicName>
|
||
. In 1901 Forel used the same specific epithet to describe
|
||
<taxonomicName id="5FA291E9FF6DFF90FDE7E652FE26F91E" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="berthoudi" subGenus="Bothroponera">
|
||
<emphasis id="AAD63678FF6DFF90FDE7E652FD53F93D" box="[609,760,1656,1680]" italics="true" pageId="8" pageNumber="546">Pachycondyla</emphasis>
|
||
(
|
||
<emphasis id="AAD63678FF6DFF90FFF5E6B1FEA2F91E" box="[115,265,1691,1715]" italics="true" pageId="8" pageNumber="546">Bothroponera</emphasis>
|
||
)
|
||
<emphasis id="AAD63678FF6DFF90FEA3E6B1FE26F91E" box="[293,397,1691,1715]" italics="true" pageId="8" pageNumber="546">berthoudi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="5FA291E9FF6DFF90FE20E6B1FF79F97B" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="berthoudi" subGenus="Bothroponera">
|
||
<emphasis id="AAD63678FF6DFF90FE20E6B1FD96F91E" box="[422,573,1691,1715]" italics="true" pageId="8" pageNumber="546">Pachycondyla</emphasis>
|
||
(
|
||
<emphasis id="AAD63678FF6DFF90FDDFE6B1FD44F91E" box="[601,751,1691,1715]" italics="true" pageId="8" pageNumber="546">Bothroponera</emphasis>
|
||
)
|
||
<emphasis id="AAD63678FF6DFF90FFECE694FF79F97B" box="[106,210,1726,1750]" italics="true" pageId="8" pageNumber="546">berthoudi</emphasis>
|
||
</taxonomicName>
|
||
has been considered to be a secondary homonym of
|
||
<taxonomicName id="5FA291E9FF6DFF90FFECE6CAFD99F955" box="[106,562,1760,1784]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="berthoudi" subGenus="Ophthalmopone">
|
||
<emphasis id="AAD63678FF6DFF90FFECE6CAFD99F955" box="[106,562,1760,1784]" italics="true" pageId="8" pageNumber="546">Pachycondyla (Ophthalmopone) berthoudi</emphasis>
|
||
</taxonomicName>
|
||
, as they were both placed in the same genus (
|
||
<bibRefCitation id="FC33979BFF6DFF90FE16E729FD88F8B6" author="Bolton, B." box="[400,547,1795,1819]" journalOrPublisher="Cambridge, Massachusetts, Harvard University Press, USA" pageId="8" pageNumber="546" pagination="504" refString="Bolton, B. (1995). A New General Catalogue of the Ants of the World (pp 504) Cambridge, Massachusetts, Harvard University Press, USA." title="A New General Catalogue of the Ants of the World (pp" type="book chapter" year="1995">Bolton, 1995</bibRefCitation>
|
||
), which has caused considerable confusion. In this project, we clearly distinguish between
|
||
<taxonomicName id="5FA291E9FF6DFF90FF4EE763FEE4F8CD" box="[200,335,1864,1888]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="berthoudi">
|
||
<emphasis id="AAD63678FF6DFF90FF4EE763FEE4F8CD" box="[200,335,1864,1888]" italics="true" pageId="8" pageNumber="546">O. berthoudi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="5FA291E9FF6DFF90FE06E763FDAEF8CD" box="[384,517,1864,1888]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="berthoudi">
|
||
<emphasis id="AAD63678FF6DFF90FE06E763FDAEF8CD" box="[384,517,1864,1888]" italics="true" pageId="8" pageNumber="546">B. berthoudi</emphasis>
|
||
</taxonomicName>
|
||
as these now belong to two different genera (
|
||
<bibRefCitation id="FC33979BFF6DFF90FED6E741FDDAF82E" author="Schmidt C. A. & Shattuck, S. O." box="[336,625,1899,1923]" journalOrPublisher="Zootaxa, 3817" pageId="8" pageNumber="546" pagination="1 - 242" part="1" publicationUrl="http://dx.doi.org/10.11646/zootaxa.3647.2.1." refString="Schmidt C. A. & Shattuck, S. O. (2014). The higher classification of the ant subfamily Ponerinae (Hymenoptera: Formicidae), with a review of ponerine ecology and behavior. Zootaxa, 3817 (1): 1 - 242. doi: 10.11646 / zootaxa. 3647.2.1." title="The higher classification of the ant subfamily Ponerinae (Hymenoptera: Formicidae), with a review of ponerine ecology and behavior" type="journal article" year="2014">Schmidt & Shattuck, 2014</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<subSection id="F12DF381FF6DFF90FCA3E62CFB18F9B1" box="[805,1203,1542,1564]" pageId="8" pageNumber="546" type="multiple">
|
||
<caption id="CCDDBAE2FF6DFF90FCA3E62CFB18F9B1" box="[805,1203,1542,1564]" httpUri="https://zenodo.org/record/269916/files/figure.png" pageId="8" pageNumber="546" targetBox="[805,1459,357,1519]" targetPageId="8">
|
||
<paragraph id="981DEA6AFF6DFF90FCA3E62CFB18F9B1" blockId="8.[805,1203,1542,1564]" box="[805,1203,1542,1564]" pageId="8" pageNumber="546">
|
||
<emphasis id="AAD63678FF6DFF90FCA3E62CFCC0F9B6" bold="true" box="[805,875,1542,1563]" pageId="8" pageNumber="546">Plate 2</emphasis>
|
||
:
|
||
<taxonomicName id="5FA291E9FF6DFF90FCFEE62CFBCAF9B6" box="[888,1121,1542,1563]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="berthoudi">
|
||
<emphasis id="AAD63678FF6DFF90FCFEE62CFBCAF9B6" box="[888,1121,1542,1563]" italics="true" pageId="8" pageNumber="546">Bothroponera berthoudi</emphasis>
|
||
</taxonomicName>
|
||
worker.
|
||
</paragraph>
|
||
</caption>
|
||
</subSection>
|
||
<subSection id="F12DF381FF6DFF90FF33E7A4FAA1FF13" pageId="8" pageNumber="546" type="multiple">
|
||
<paragraph id="981DEA6AFF6DFF90FF33E7A4FAA1FF13" blockId="8.[106,760,131,1993]" lastBlockId="8.[805,1459,131,328]" pageId="8" pageNumber="546">
|
||
The worker of
|
||
<taxonomicName id="5FA291E9FF6DFF90FEDFE7A4FE4AF80B" box="[345,481,1934,1958]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="berthoudi">
|
||
<emphasis id="AAD63678FF6DFF90FEDFE7A4FE4AF80B" box="[345,481,1934,1958]" italics="true" pageId="8" pageNumber="546">B. berthoudi</emphasis>
|
||
</taxonomicName>
|
||
is identical to the worker of
|
||
<taxonomicName id="5FA291E9FF6DFF90FF0FE79BFEA5F865" box="[137,270,1968,1992]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="variolosa">
|
||
<emphasis id="AAD63678FF6DFF90FF0FE79BFEA5F865" box="[137,270,1968,1992]" italics="true" pageId="8" pageNumber="546">B. variolosa</emphasis>
|
||
</taxonomicName>
|
||
, which is considered a synonym. They have the lower medial margin of the clypeus “u” shaped with a medial raised area and lack the sharp carinae.
|
||
</paragraph>
|
||
</subSection>
|
||
<paragraph id="981DEA6AFF6DFF90FCF7E0E2FB91FEE5" blockId="8.[805,1459,131,328]" pageId="8" pageNumber="546">
|
||
<taxonomicName id="5FA291E9FF6DFF90FCF7E0E2FBF7FF4D" box="[881,1116,200,224]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="cariosa">
|
||
<emphasis id="AAD63678FF6DFF90FCF7E0E2FBF7FF4D" box="[881,1116,200,224]" italics="true" pageId="8" pageNumber="546">Bothroponera cariosa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="5FA291E9FF6DFF90FB09E0E3FABCFF4D" box="[1167,1303,200,224]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="strigulosa">
|
||
<emphasis id="AAD63678FF6DFF90FB09E0E3FABCFF4D" box="[1167,1303,200,224]" italics="true" pageId="8" pageNumber="546">B. strigulosa</emphasis>
|
||
</taxonomicName>
|
||
have the same “u” shaped lower medial anterior margin of the clypeus, but this area forms a sharp clypeal carinae in both species, which is lacking in
|
||
<taxonomicName id="5FA291E9FF6DFF90FC28E11BFB9FFEE5" box="[942,1076,304,328]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="berthoudi">
|
||
<emphasis id="AAD63678FF6DFF90FC28E11BFB9FFEE5" box="[942,1076,304,328]" italics="true" pageId="8" pageNumber="546">B. berthoudi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D0B8B9E1FF6DFF91FCA3E66BFD95FF4D" lastPageId="9" lastPageNumber="547" pageId="8" pageNumber="546" type="materials_examined">
|
||
<subSection id="F12DF381FF6DFF90FCA3E66BFC56F9F4" box="[805,1021,1601,1625]" pageId="8" pageNumber="546" type="multiple">
|
||
<paragraph id="981DEA6AFF6DFF90FCA3E66BFC56F9F4" blockId="8.[805,1021,1601,1625]" box="[805,1021,1601,1625]" pageId="8" pageNumber="546">
|
||
<heading id="C3555D06FF6DFF90FCA3E66BFC56F9F4" bold="true" box="[805,1021,1601,1625]" fontSize="10" level="1" pageId="8" pageNumber="546" reason="2">
|
||
<emphasis id="AAD63678FF6DFF90FCA3E66BFC56F9F4" bold="true" box="[805,1021,1601,1625]" pageId="8" pageNumber="546">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSection>
|
||
<subSection id="F12DF381FF6DFF91FCF7E656FD95FF4D" lastPageId="9" lastPageNumber="547" pageId="8" pageNumber="546" type="multiple">
|
||
<paragraph id="981DEA6AFF6DFF91FCF7E656FD95FF4D" blockId="8.[805,1459,1660,1996]" lastBlockId="9.[128,789,131,1369]" lastPageId="9" lastPageNumber="547" pageId="8" pageNumber="546">
|
||
<emphasis id="AAD63678FF6DFF90FCF7E656FC3BF91A" bold="true" pageId="8" pageNumber="546">Type material: SOUTH AFRICA: Mpumalanga Province,</emphasis>
|
||
Salique, A. R. I. Pretoria A. C. U. P. 314 (forests), 24°36’0’’ S; 30°54’0’’ E, vii-1944, Donated by Univ. Pretoria, 14 Mar 1995,
|
||
<taxonomicName id="5FA291E9FF6DFF90FC4AE6CEFAD6F951" authority="Arnold, 1947" authorityName="Arnold" authorityYear="1947" box="[972,1405,1764,1788]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="variolosa">
|
||
<emphasis id="AAD63678FF6DFF90FC4AE6CEFB66F951" box="[972,1229,1764,1788]" italics="true" pageId="8" pageNumber="546">Bothroponera variolosa</emphasis>
|
||
(
|
||
<bibRefCitation id="FC33979BFF6DFF90FB64E6CEFADEF951" author="Arnold, G." box="[1250,1397,1764,1788]" journalOrPublisher="Occasional Papers of the National Museums of Southern Rhodesia" pageId="8" pageNumber="546" pagination="131 - 167" part="2" refString="Arnold, G. (1947). New species of African Hymenoptera. No. 7. Occasional Papers of the National Museums of Southern Rhodesia, 2: 131 - 167." title="New species of African Hymenoptera. No. 7" type="journal article" year="1947">Arnold, 1947</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(1w paratype # C008009, SAM-HYM [Iziko]);
|
||
<emphasis id="AAD63678FF6DFF90FB59E72DFA18F8B2" bold="true" box="[1247,1459,1799,1823]" pageId="8" pageNumber="546">Limpopo Province,</emphasis>
|
||
Marieps, A.R.I. Pretoria A. C. U.P. 367 (farmstead and forests), 24°35’0’’ S; 30°52’0’’ E, vii-1944,
|
||
<emphasis id="AAD63678FF6DFF90FB35E766FCCEF82A" italics="true" pageId="8" pageNumber="546">
|
||
<taxonomicName id="5FA291E9FF6DFF90FB35E766FA18F8C9" box="[1203,1459,1868,1892]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="variolosa" status="sp. nov.">Bothroponera variolosa</taxonomicName>
|
||
<taxonomicNameLabel id="B1E58B03FF6DFF90FCA3E75AFCCEF82A" box="[805,869,1904,1927]" pageId="8" pageNumber="546" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<bibRefCitation id="FC33979BFF6DFF90FCFDE745FBB8F82A" author="Arnold, G." box="[891,1043,1903,1927]" journalOrPublisher="Occasional Papers of the National Museums of Southern Rhodesia" pageId="8" pageNumber="546" pagination="131 - 167" part="2" refString="Arnold, G. (1947). New species of African Hymenoptera. No. 7. Occasional Papers of the National Museums of Southern Rhodesia, 2: 131 - 167." title="New species of African Hymenoptera. No. 7" type="journal article" year="1947">Arnold, 1947</bibRefCitation>
|
||
), (1w paratype, SAM-ENT [Iziko # 0011524]);
|
||
<emphasis id="AAD63678FF6DFF90FC26E7BBFB8CF804" bold="true" box="[928,1063,1937,1961]" pageId="8" pageNumber="546">Mariepskop</emphasis>
|
||
Transvaal, (farmstead), 24°35’0’’ S; 30°52’0’’ E, vii-1945, J. C. Faure,
|
||
<taxonomicName id="5FA291E9FF6DFF91FB28E79EFE89FF36" authority="Arnold, 1947" authorityName="Arnold" authorityYear="1947" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="9" lastPageNumber="547" order="Hymenoptera" pageId="8" pageNumber="546" phylum="Arthropoda" rank="species" species="variolosa">
|
||
<emphasis id="AAD63678FF6DFF90FB28E79EFA18F861" box="[1198,1459,1972,1996]" italics="true" pageId="8" pageNumber="546">Bothroponera variolosa</emphasis>
|
||
(
|
||
<bibRefCitation id="FC33979BFF6CFF91FF0EE0A9FEB1FF36" author="Arnold, G." box="[136,282,131,155]" journalOrPublisher="Occasional Papers of the National Museums of Southern Rhodesia" pageId="9" pageNumber="547" pagination="131 - 167" part="2" refString="Arnold, G. (1947). New species of African Hymenoptera. No. 7. Occasional Papers of the National Museums of Southern Rhodesia, 2: 131 - 167." title="New species of African Hymenoptera. No. 7" type="journal article" year="1947">Arnold, 1947</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, (2w # 0 0 11524 paratypes, SAM-ENT [Iziko], British Museum 1947-250, BMNH (E) 1015515), Antweb Casent 0 902470 (2w paratypes, BMNH).
|
||
</paragraph>
|
||
</subSection>
|
||
</subSubSection>
|
||
<subSection id="F12DF381FF6CFF91FF4AE0C1FE0EFCA7" pageId="9" pageNumber="547" type="multiple">
|
||
<subSubSection id="D0B8B9E1FF6CFF91FF4AE0C1FE0EFCA7" pageId="9" pageNumber="547" type="discussion">
|
||
<paragraph id="981DEA6AFF6CFF91FF4AE0C1FE0EFCA7" blockId="9.[128,789,131,1369]" pageId="9" pageNumber="547">
|
||
<emphasis id="AAD63678FF6CFF91FF4AE0C1FF4AFE88" bold="true" pageId="9" pageNumber="547">Non-type material: SOUTH AFRICA: Eastern Cape Province</emphasis>
|
||
, Algoa-Bay, Capland, 33°50’0’’ S; 25°50’0’’ E, Dr. H. Brauns, Forel det. 1922, that deposited in the Berlin Museum was designated by Forel, (labeled
|
||
<taxonomicName id="5FA291E9FF6CFF91FDF1E179FE76FE23" authority="Forel" authorityName="Forel" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="547" phylum="Arthropoda" rank="species" species="berthoudi" subGenus="Bothroponera">
|
||
<emphasis id="AAD63678FF6CFF91FDF1E179FCA5FEC6" box="[631,782,339,363]" italics="true" pageId="9" pageNumber="547">Pachycondyla</emphasis>
|
||
(
|
||
<emphasis id="AAD63678FF6CFF91FF0FE15CFEB4FE23" box="[137,287,374,398]" italics="true" pageId="9" pageNumber="547">Bothroponera</emphasis>
|
||
)
|
||
<emphasis id="AAD63678FF6CFF91FEB7E15CFE32FE23" box="[305,409,374,398]" italics="true" pageId="9" pageNumber="547">berthoudi</emphasis>
|
||
Forel
|
||
</taxonomicName>
|
||
, 1w # 6692, MfN), but this specimen is broken into two parts (head with the pronotum is one part and the second part includes the mesonotum, mesopleuron, propodeum, petiole, postpetiole and the 2nd to 5th gastral segments). It is not clear if this specimen is the type or not.
|
||
<emphasis id="AAD63678FF6CFF91FF79E209FE73FD96" bold="true" box="[255,472,547,571]" pageId="9" pageNumber="547">Limpopo Province,</emphasis>
|
||
Mariepskop, 4000 Transvaal (farm), 26°42’0’’ S; 29°53’0’’ E, vii-1944; South Africa Museum ex. National Museum Bulawayo 1981, (labeled
|
||
<taxonomicName id="5FA291E9FF6CFF91FF06E2A1FE21FD0E" box="[128,394,651,675]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="547" phylum="Arthropoda" rank="species" species="variolosa">
|
||
<emphasis id="AAD63678FF6CFF91FF06E2A1FE21FD0E" box="[128,394,651,675]" italics="true" pageId="9" pageNumber="547">Bothroponera variolosa</emphasis>
|
||
</taxonomicName>
|
||
, 1w # 11524 SAM-ENT); Some of the
|
||
<taxonomicName id="5FA291E9FF6CFF91FF54E284FEF5FD6B" box="[210,350,686,710]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="547" phylum="Arthropoda" rank="species" species="berthoudi">
|
||
<emphasis id="AAD63678FF6CFF91FF54E284FEF5FD6B" box="[210,350,686,710]" italics="true" pageId="9" pageNumber="547">B. berthoudi</emphasis>
|
||
</taxonomicName>
|
||
specimens that were collected by H. Brauns, Paul Berthoud and G. Arnold were considered to be subspecies of
|
||
<taxonomicName id="5FA291E9FF6CFF91FE90E2DEFE34FCA6" box="[278,415,755,779]" class="Insecta" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="547" phylum="Arthropoda" rank="species" species="strigulosa">
|
||
<emphasis id="AAD63678FF6CFF91FE90E2DEFE34FCA6" box="[278,415,755,779]" italics="true" pageId="9" pageNumber="547">B. strigulosa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</subSection>
|
||
</treatment>
|
||
</document> |