979 lines
172 KiB
XML
979 lines
172 KiB
XML
<document id="6E2A4F363CD114EF119E9C244105E721" ID-CLB-Dataset="31806" ID-DOI="10.5281/zenodo.893555" ID-GBIF-Dataset="4a2efe36-c48b-456b-9223-7b5793025a72" ID-ISSN="0035-418" ID-Zenodo-Dep="893555" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1508361162557" checkinUser="plazi" docAuthor="Peter J. Schwendinger" docDate="2017" docId="4C30A452FFE0FFFDBA9AFAC53979F872" docLanguage="en" docName="rsz_124_2_11_Schwendinger.pdf" docOrigin="Revue suisse de Zoologie 124 (2)" docStyle="DocumentStyle:225056F0A23C51DDC06AC27EAEA15D5B.2:RevSuisseZool.2015-2019.journal_article" docStyleId="225056F0A23C51DDC06AC27EAEA15D5B" docStyleName="RevSuisseZool.2015-2019.journal_article" docStyleVersion="2" docTitle="Liphistius malayanus Abraham 1923" docType="treatment" docVersion="9" lastPageNumber="401" masterDocId="B009DC2AFFE4FFF7BA6FFFF13818FFE3" masterDocTitle="A revision of the trapdoor spider genus Liphistius (Mesothelae: Liphistiidae) in peninsular Malaysia; part 1" masterLastPageNumber="445" masterPageNumber="391" pageNumber="395" updateTime="1698469770161" updateUser="ExternalLinkService">
|
||
<mods:mods id="B93421B07EE6C0D602C11ED282A80F45" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="3286B6E357A5042CB5F62636148A81C8">
|
||
<mods:title id="4B45ECDB67B7F0A362C1867425BF295C">A revision of the trapdoor spider genus Liphistius (Mesothelae: Liphistiidae) in peninsular Malaysia; part 1</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="A53CED9A82C689075D6D36EE4BC8F229" type="personal">
|
||
<mods:role id="0D032A7E5D327FB1796544A0128BF814">
|
||
<mods:roleTerm id="7C31C1AF3DC81DFC68B5CCD8BE50271E">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="ADD810E7560C521BD3F86E16486286B5">Peter J. Schwendinger</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="23B2982571C871E83E59B05E5131CE55">text</mods:typeOfResource>
|
||
<mods:relatedItem id="16BE560844EBBA85E95C21437D1EAED5" type="host">
|
||
<mods:titleInfo id="B22C76E6BCFCF5EAD4A5437C7353712A">
|
||
<mods:title id="23E286E9619DF8BDA5EDFCC78045289B">Revue suisse de Zoologie</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="81F156B2FDE3DC0DBDCFE064AFBFBB92">
|
||
<mods:date id="40C569588284EFBCFE2F8F770CA67398">2017</mods:date>
|
||
<mods:detail id="F9F411C64F1E63D3CF5E6C1696C29A20" type="pubDate">
|
||
<mods:number id="8FDE5CC5D7198FEFC13906D0694BE533">2017-09-30</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="4E988435E4E6918ABDE201F17EF91C1B" type="volume">
|
||
<mods:number id="86CAEF52949BB2DD23E8A0DC7ADC69A7">124</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="4378B2F17D2FAD7D45E2DD2FDF4146A4" type="issue">
|
||
<mods:number id="27F01C1D65DD651DBDDAA691265DC499">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="37360E8D536527EFBB19E55F428A45F8" unit="page">
|
||
<mods:start id="42BEE7BAAF3BDBA1F122F39D26525E3E">391</mods:start>
|
||
<mods:end id="025026B2C8D6BEF316F082CE168CA9BF">445</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="63554FDCF7B29155F5CB50ADB34BEA18">journal article</mods:classification>
|
||
<mods:identifier id="71F6634732BA0317DF419FE1D6669976" type="CLB-Dataset">31806</mods:identifier>
|
||
<mods:identifier id="5AC8C341081D4A13021B251FA0D17445" type="DOI">10.5281/zenodo.893555</mods:identifier>
|
||
<mods:identifier id="67E9984D7EE097DD7E59FC6752192DE2" type="GBIF-Dataset">4a2efe36-c48b-456b-9223-7b5793025a72</mods:identifier>
|
||
<mods:identifier id="88D696BD9193F7E548944534CFB2A264" type="ISSN">0035-418</mods:identifier>
|
||
<mods:identifier id="E855896FA005CD0DE54237E313ED5CA4" type="Zenodo-Dep">893555</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="4C30A452FFE0FFFDBA9AFAC53979F872" ID-DOI="http://doi.org/10.5281/zenodo.6042349" ID-GBIF-Taxon="135450781" ID-Zenodo-Dep="6042349" LSID="urn:lsid:plazi:treatment:4C30A452FFE0FFFDBA9AFAC53979F872" httpUri="http://treatment.plazi.org/id/4C30A452FFE0FFFDBA9AFAC53979F872" lastPageId="10" lastPageNumber="401" pageId="4" pageNumber="395">
|
||
<subSubSection id="8C8346CFFFE0FFF3BA9AFAC53A2EFA88" pageId="4" pageNumber="395" type="nomenclature">
|
||
<paragraph id="C4261544FFE0FFF3BA9AFAC53A8EFAA8" blockId="4.[245,662,1332,1389]" box="[245,662,1332,1355]" pageId="4" pageNumber="395">
|
||
<heading id="9F6EA228FFE0FFF3BA9AFAC53A8EFAA8" bold="true" box="[245,662,1332,1355]" centered="true" fontSize="10" level="3" pageId="4" pageNumber="395" reason="4">
|
||
<taxonomicName id="03996EC7FFE0FFF3BA9AFAC53A8EFAA8" ID-CoL="72J5G" authority="Abraham, 1923" authorityName="Abraham" authorityYear="1923" box="[245,662,1332,1355]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="395" phylum="Arthropoda" rank="species" species="malayanus">
|
||
<emphasis id="F6EDC956FFE0FFF3BA9AFAC53A8EFAA8" bold="true" box="[245,662,1332,1355]" pageId="4" pageNumber="395">
|
||
<emphasis id="F6EDC956FFE0FFF3BA9AFAC539F9FAA8" bold="true" box="[245,481,1332,1355]" italics="true" pageId="4" pageNumber="395">Liphistius malayanus</emphasis>
|
||
<bibRefCitation id="A00868B5FFE0FFF3BB89FAC53A8EFAA8" author="Abraham H. C." box="[486,662,1332,1355]" journalOrPublisher="Journal of the Malayan Branch of the Royal Asiatic Society" pageId="4" pageNumber="395" pagination="13 - 21" part="1" refString="Abraham H. C. 1923 a. A new spider of the genus Liphistius. Journal of the Malayan Branch of the Royal Asiatic Society 1: 13 - 21, pl. 1." title="A new spider of the genus Liphistius" type="journal article" year="1923">Abraham, 1923</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="C4261544FFE0FFF3BB3AFAA53A2EFA88" blockId="4.[245,662,1332,1389]" box="[341,566,1364,1389]" pageId="4" pageNumber="395">
|
||
<figureCitation id="5CA209C1FFE0FFF3BB3AFAA53A1FFA8F" box="[341,519,1364,1389]" captionStart="Fig. 3" captionStartId="5.[151,186,1940,1961]" captionTargetBox="[261,1329,261,1882]" captionTargetId="figure@5.[255,1334,257,1886]" captionTargetPageId="5" captionText="Fig. 3. SEM-micrographs of embolus proper of left palpal organ of two males: Liphistius desultor from Penang Hill (A, C, E, G) and Liphistius malayanus from Fraser’s Hill (B, D, F, H). (A) Retroventral-distal view. (B) Dorsal-distal view. (C) Distal view (dorsal side to the right). (D) Distal view (prolateral side to the right). (E) Retrodorsal view. (F) Prodorsal view. (G) Proventral view. (H) Prolateral view. Abbreviations: f - major embolic fold; m - membranous part of embolus proper; s - sclerotised part of embolus proper." httpUri="https://zenodo.org/record/1021943/files/figure.png" pageId="4" pageNumber="395">Figs 3B, D, F, H</figureCitation>
|
||
;
|
||
<figureCitation id="5CA209C1FFE0FFF3B87DFAA53A39FA88" box="[530,545,1364,1387]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="4" pageNumber="395">4</figureCitation>
|
||
-
|
||
<figureCitation id="5CA209C1FFE0FFF3B846FAA53A2EFA88" box="[553,566,1364,1387]" captionStart="Fig. 5" captionStartId="8.[151,186,1809,1830]" captionTargetBox="[154,1439,260,1762]" captionTargetId="figure@8.[148,1439,254,1779]" captionTargetPageId="8" captionText="Fig. 5. Liphistius malayanus, vulval plate of eight females: paratype of L. m. cameroni from Berinchang, Cameron Highlands (A- B); exuvia of female from Ringlet, Cameron Highlands (C); female (SMF 64093) from Fraser’s Hill (D); female (moulted 24. VI. 2002) from Fraser’s Hill; poreplate slightly deformed by brief immersion in cold KOH (E); female (SMF 56206) from Fraser’s Hill (F); female (SMF 60037) from Genting Highlands (G); female (moulted 8. XII. 2005) from Genting Highlands (H); exuvia of female from Templer Park (I-J). (A, C-I) Dorsal view. (B, J) Ventral view. Scale lines: 1.0 mm (A-B; C, F; D; E, G; H; I-J)." httpUri="https://zenodo.org/record/1021947/files/figure.png" pageId="4" pageNumber="395">5</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8C8346CFFFE0FFF3BAF8FA733AE9F9A7" pageId="4" pageNumber="395" type="reference_group">
|
||
<paragraph id="C4261544FFE0FFF3BAF8FA733AA1F9E8" blockId="4.[151,756,1410,1604]" pageId="4" pageNumber="395">
|
||
<treatmentCitationGroup id="E489326AFFE0FFF3BAF8FA733AA1F9E8" infered="true" pageId="4" pageNumber="395">
|
||
<taxonomicName id="03996EC7FFE0FFF3BAF8FA733A70FA7B" ID-CoL="72J5G" authority="Abraham, 1923 b: 770" authorityName="Abraham" authorityPageNumber="770" authorityYear="1923" box="[151,616,1410,1432]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="395" phylum="Arthropoda" rank="species" species="malayanus">
|
||
<emphasis id="F6EDC956FFE0FFF3BAF8FA733977FA74" box="[151,367,1410,1431]" italics="true" pageId="4" pageNumber="395">Liphistius malayanus</emphasis>
|
||
<treatmentCitation id="45383355FFE0FFF3BB11FA733A70FA7B" author="Abraham H. C." box="[382,616,1410,1432]" page="770" pageId="4" pageNumber="395" year="1923">
|
||
<bibRefCitation id="A00868B5FFE0FFF3BB11FA733A36FA74" author="Abraham H. C." box="[382,558,1410,1431]" journalOrPublisher="Proceedings of the Zoological Society of London 1923" pageId="4" pageNumber="395" pagination="769 - 774" refString="Abraham H. C. 1923 b. A new spider of the genus Liphistius from the Malay Peninsula, and some observations on its habits. Proceedings of the Zoological Society of London 1923: 769 - 774, pl. 1." title="A new spider of the genus Liphistius from the Malay Peninsula, and some observations on its habits" type="proceedings paper" year="1923" yearSuffix="b">Abraham, 1923b</bibRefCitation>
|
||
: 770
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
-774, text-fig. 1a-b (description of female). –
|
||
<bibRefCitation id="A00868B5FFE0FFF3B872FA6E3905FA0E" author="Abraham H. C." journalOrPublisher="Proceedings of the Zoological Society of London 1929" pageId="4" pageNumber="395" pagination="671 - 677" refString="Abraham H. C. 1929. The male of the spider Liphistius malayanus H. C. Abr., with further notes on the female and on its habits. Proceedings of the Zoological Society of London 1929: 671 - 677, pls 1 - 3." title="The male of the spider Liphistius malayanus H. C. Abr., with further notes on the female and on its habits" type="proceedings paper" year="1929">Abraham (1929: 674- 676, pl. 1, figs 3-4, pl. 2, figs 11-16; first description of male)</bibRefCitation>
|
||
. For synonymy and other taxonomically relevant publications see the
|
||
<bibRefCitation id="A00868B5FFE0FFF3BBC9FA043AADF9E8" author="World Spider Catalog" box="[422,693,1525,1547]" journalOrPublisher="Natural History Museum Bern. Available from http: // wsc. nmbe. ch (accessed in July" pageId="4" pageNumber="395" pagination="2017" refString="World Spider Catalog 2017. World Spider Catalog. Version 18.5. Natural History Museum Bern. Available from http: // wsc. nmbe. ch (accessed in July 2017)." title="World Spider Catalog. Version 18.5" type="book chapter" year="2017">World Spider Catalog, 2017</bibRefCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="C4261544FFE0FFF3BAF8F9E33AE9F9A7" blockId="4.[151,756,1410,1604]" pageId="4" pageNumber="395">
|
||
<treatmentCitationGroup id="E489326AFFE0FFF3BAF8F9E33AE9F9A7" pageId="4" pageNumber="395">
|
||
<taxonomicName id="03996EC7FFE0FFF3BAF8F9E33A63F9CB" ID-CoL="7K9NR" authority="Haupt, 1983: 282" authorityName="Haupt" authorityPageNumber="282" authorityYear="1983" box="[151,635,1554,1576]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="395" phylum="Arthropoda" rank="subSpecies" species="malayanus" subSpecies="cameroni">
|
||
<emphasis id="F6EDC956FFE0FFF3BAF8F9E339DFF9CB" box="[151,455,1554,1576]" italics="true" pageId="4" pageNumber="395">Liphistius malayanus cameroni</emphasis>
|
||
<treatmentCitation id="45383355FFE0FFF3BBA0F9E23A63F9CB" author="Haupt J." box="[463,635,1555,1576]" page="282" pageId="4" pageNumber="395" year="1983">
|
||
<bibRefCitation id="A00868B5FFE0FFF3BBA0F9E23A53F9CB" author="Haupt J." box="[463,587,1555,1576]" journalOrPublisher="Zeitschrift fur zoologische Systematik und Evolutionsforschung" pageId="4" pageNumber="395" pagination="275 - 293" part="21" refString="Haupt J. 1983. Vergleichende Morphologie der Genitalorgane und Phylogenie der liphistiomorphen Webspinnen (Araneae: Mesothelae). I. Revision der bisher bekannten Arten. Zeitschrift fur zoologische Systematik und Evolutionsforschung 21: 275 - 293." title="Vergleichende Morphologie der Genitalorgane und Phylogenie der liphistiomorphen Webspinnen (Araneae: Mesothelae). I. Revision der bisher bekannten Arten" type="journal article" year="1983">Haupt, 1983</bibRefCitation>
|
||
: 282
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
, figs 3e, 4d, 5f, 6f (description of male and female).
|
||
<emphasis id="F6EDC956FFE0FFF3B80DF9DE3AE9F9A7" bold="true" box="[610,753,1583,1604]" pageId="4" pageNumber="395">New synonym</emphasis>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8C8346CFFFE0FFF3BAF8F9AD3983F8CC" pageId="4" pageNumber="395" type="materials_examined">
|
||
<paragraph id="C4261544FFE0FFF3BAF8F9AD3983F8CC" blockId="4.[151,756,1628,1841]" pageId="4" pageNumber="395">
|
||
<emphasis id="F6EDC956FFE0FFF3BAF8F9AD394BF990" bold="true" box="[151,339,1628,1652]" pageId="4" pageNumber="395">Type material:</emphasis>
|
||
BMNH; female holotype of
|
||
<taxonomicName id="03996EC7FFE0FFF3B8B1F9AC3914F971" authorityName="Abraham" authorityYear="1923" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="395" phylum="Arthropoda" rank="species" species="malayanus">
|
||
<emphasis id="F6EDC956FFE0FFF3B8B1F9AC3914F971" italics="true" pageId="4" pageNumber="395">L. malayanus</emphasis>
|
||
</taxonomicName>
|
||
(not examined);
|
||
<materialsCitation id="74F11F19FFE0FFF3BBB6F98A3AE8F913" ID-GBIF-Occurrence="1675315678" collectingDate="1922-12" collectionCode="ZMH" collectorName="F. de la Mare Norris." location="Seremban" municipality="Gunung Angsi" pageId="4" pageNumber="395" specimenCount="2" stateProvince="Negri Sembilan" typeStatus="holotype">
|
||
<collectingRegion id="065DDBA6FFE0FFF3BBB6F98A3A94F971" box="[473,652,1659,1682]" country="Malaysia" name="Negeri Sembilan" pageId="4" pageNumber="395">Negri Sembilan</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="24428F3EFFE0FFF3B8F2F98D38CFF951" pageId="4" pageNumber="395">Gunung Angsi</collectingMunicipality>
|
||
near
|
||
<location id="C146439FFFE0FFF3BB4DF96A3989F951" LSID="urn:lsid:plazi:treatment:4C30A452FFE0FFFDBA9AFAC53979F872:C146439FFFE0FFF3BB4DF96A3989F951" box="[290,401,1691,1714]" municipality="Gunung Angsi" name="Seremban" pageId="4" pageNumber="395" stateProvince="Negri Sembilan">Seremban</location>
|
||
;
|
||
<collectingDate id="A063CA6CFFE0FFF3BBCCF96A3A11F950" box="[419,521,1691,1715]" pageId="4" pageNumber="395" value="1922-12">
|
||
<date id="B0273384FFE0FFF3BBCCF96A3A11F950" box="[419,521,1691,1715]" pageId="4" pageNumber="395" value="1922-12">XII.1922</date>
|
||
</collectingDate>
|
||
; leg.
|
||
<collectorName id="696C7092FFE0FFF3B83EF96D38FCF932" pageId="4" pageNumber="395">F. de la Mare Norris.</collectorName>
|
||
–
|
||
<collectionCode id="A2888D81FFE0FFF3BB65F94A395EF931" box="[266,326,1723,1746]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="4" pageNumber="395">ZMH</collectionCode>
|
||
(A16/84); male
|
||
<typeStatus id="1B22ABE6FFE0FFF3B863F94B3A72F932" box="[524,618,1722,1745]" pageId="4" pageNumber="395" type="holotype">holotype</typeStatus>
|
||
and female
|
||
<typeStatus id="1B22ABE6FFE0FFF3BAF8F92A38EDF910" box="[151,245,1755,1779]" pageId="4" pageNumber="395" type="paratype">paratype</typeStatus>
|
||
of
|
||
<taxonomicName id="03996EC7FFE0FFF3BB56F92A3A41F911" authorityName="Haupt" authorityYear="1983" box="[313,601,1753,1778]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="395" phylum="Arthropoda" rank="subSpecies" species="malayanus" subSpecies="cameroni">
|
||
<emphasis id="F6EDC956FFE0FFF3BB56F92A3A41F911" box="[313,601,1753,1778]" italics="true" pageId="4" pageNumber="395">L. malayanus cameroni</emphasis>
|
||
</taxonomicName>
|
||
(examined)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="74F11F19FFE0FFF3BAF8F9083983F8CC" ID-GBIF-Occurrence="1675315687" collectingDate="1981-03-04" collectorName="J. Haupt & Dach." location="Berinchang" pageId="4" pageNumber="395" specimenCount="1" stateProvince="Pahang" typeStatus="holotype">
|
||
<collectingRegion id="065DDBA6FFE0FFF3BAF8F90838F1F8F3" box="[151,233,1785,1808]" country="Malaysia" name="Pahang" pageId="4" pageNumber="395">Pahang</collectingRegion>
|
||
, Cameron
|
||
<collectingRegion id="065DDBA6FFE0FFF3BB03F90839C6F8F3" box="[364,478,1785,1808]" country="Czech Republic" name="Vysocina" pageId="4" pageNumber="395">Highlands</collectingRegion>
|
||
,
|
||
<location id="C146439FFFE0FFF3BB80F9083A77F8F3" LSID="urn:lsid:plazi:treatment:4C30A452FFE0FFFDBA9AFAC53979F872:C146439FFFE0FFF3BB80F9083A77F8F3" box="[495,623,1785,1808]" name="Berinchang" pageId="4" pageNumber="395" stateProvince="Pahang">Berinchang</location>
|
||
;
|
||
<collectingDate id="A063CA6CFFE0FFF3B8EEF90B3AE8F8F1" box="[641,752,1786,1810]" pageId="4" pageNumber="395" value="1981-03-04">
|
||
<date id="B0273384FFE0FFF3B8EEF90B3AE8F8F1" box="[641,752,1786,1810]" pageId="4" pageNumber="395" value="1981-03-04">4.III.1981</date>
|
||
</collectingDate>
|
||
; leg.
|
||
<collectorName id="696C7092FFE0FFF3BAAAF8EB3906F8D2" box="[197,286,1818,1841]" pageId="4" pageNumber="395">J. Haupt</collectorName>
|
||
& T.
|
||
<collectorName id="696C7092FFE0FFF3BB34F8E93983F8CC" box="[347,411,1816,1839]" pageId="4" pageNumber="395">Dach.</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8C8346CFFFE0FFF3BAF8F8B939B5F7F8" pageId="4" pageNumber="395" type="discussion">
|
||
<paragraph id="C4261544FFE0FFF3BAF8F8B939B5F7F8" blockId="4.[151,756,1863,2075]" pageId="4" pageNumber="395">
|
||
<emphasis id="F6EDC956FFE0FFF3BAF8F8B938E6F883" bold="true" box="[151,254,1864,1888]" pageId="4" pageNumber="395">Remark:</emphasis>
|
||
A
|
||
<taxonomicName id="03996EC7FFE0FFF3BB48F8B939A6F8BD" authorityName="Abraham" authorityYear="1923" box="[295,446,1863,1887]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="395" phylum="Arthropoda" rank="species" species="malayanus">
|
||
<emphasis id="F6EDC956FFE0FFF3BB48F8B939A6F8BD" box="[295,446,1863,1887]" italics="true" pageId="4" pageNumber="395">L. malayanus</emphasis>
|
||
</taxonomicName>
|
||
male (not examined) from Fraser’s Hill was described by
|
||
<bibRefCitation id="A00868B5FFE0FFF3B869F8973AD8F89C" author="Abraham H. C." box="[518,704,1894,1919]" journalOrPublisher="Proceedings of the Zoological Society of London 1929" pageId="4" pageNumber="395" pagination="671 - 677" refString="Abraham H. C. 1929. The male of the spider Liphistius malayanus H. C. Abr., with further notes on the female and on its habits. Proceedings of the Zoological Society of London 1929: 671 - 677, pls 1 - 3." title="The male of the spider Liphistius malayanus H. C. Abr., with further notes on the female and on its habits" type="proceedings paper" year="1929">Abraham (1929)</bibRefCitation>
|
||
and deposited in the BMNH.
|
||
<bibRefCitation id="A00868B5FFE0FFF3BBAFF8763A4AF87C" author="Haupt J." box="[448,594,1927,1951]" journalOrPublisher="Zeitschrift fur zoologische Systematik und Evolutionsforschung" pageId="4" pageNumber="395" pagination="275 - 293" part="21" refString="Haupt J. 1983. Vergleichende Morphologie der Genitalorgane und Phylogenie der liphistiomorphen Webspinnen (Araneae: Mesothelae). I. Revision der bisher bekannten Arten. Zeitschrift fur zoologische Systematik und Evolutionsforschung 21: 275 - 293." title="Vergleichende Morphologie der Genitalorgane und Phylogenie der liphistiomorphen Webspinnen (Araneae: Mesothelae). I. Revision der bisher bekannten Arten" type="journal article" year="1983">Haupt (1983</bibRefCitation>
|
||
: 281, figs 3d, 4c) re-described this specimen and referred to it as the
|
||
<typeStatus id="1B22ABE6FFE0FFF3BAF8F83738EEF83D" box="[151,246,1990,2014]" pageId="4" pageNumber="395" type="paratype">paratype</typeStatus>
|
||
, but it has no
|
||
<typeStatus id="1B22ABE6FFE0FFF3BBE3F83739A2F83D" box="[396,442,1990,2014]" pageId="4" pageNumber="395">type</typeStatus>
|
||
status, having been collected in
|
||
<date id="B0273384FFE0FFF3BADAF815397FF81E" box="[181,359,2020,2045]" pageId="4" pageNumber="395" value="1928-11">November 1928</date>
|
||
, after the publication of the original description of the species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8C8346CFFFE0FFF0B950FBA53A68FD31" lastPageId="7" lastPageNumber="398" pageId="4" pageNumber="395" type="materials_examined">
|
||
<paragraph id="C4261544FFE0FFF0B950FBA53A68FD31" blockId="4.[831,1437,1108,2079]" lastBlockId="7.[151,757,258,724]" lastPageId="7" lastPageNumber="398" pageId="4" pageNumber="395">
|
||
<materialsCitation id="74F11F19FFE0FFF3B950FBA53BA7FAAC" ID-GBIF-Occurrence="1675315692" collectingDate="1981-07" collectionCode="ZMH" collectorName="Cameron Highlands & Berinchang" location="Material" pageId="4" pageNumber="395" specimenCount="2" stateProvince="Pahang" typeStatus="holotype">
|
||
<emphasis id="F6EDC956FFE0FFF3B950FBA53C32FB88" bold="true" box="[831,1066,1108,1131]" pageId="4" pageNumber="395">
|
||
<location id="C146439FFFE0FFF3B950FBA53BBBFB88" LSID="urn:lsid:plazi:treatment:4C30A452FFE0FFFDBA9AFAC53979F872:C146439FFFE0FFF3B950FBA53BBBFB88" box="[831,931,1108,1131]" name="Material" pageId="4" pageNumber="395" stateProvince="Pahang">Material</location>
|
||
examined:
|
||
</emphasis>
|
||
<collectionCode id="A2888D81FFE0FFF3BE57FBA43C6CFB8F" box="[1080,1140,1109,1132]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="4" pageNumber="395">ZMH</collectionCode>
|
||
(A16/84); male
|
||
<typeStatus id="1B22ABE6FFE0FFF3BF51FBA53D84FB88" box="[1342,1436,1108,1131]" pageId="4" pageNumber="395" type="holotype">holotype</typeStatus>
|
||
[left palp detached, macerated and partly collapsed, cymbium and hematodocha cut open to expose tendons inside, all strong spines deformed; right palp without tegulum, contrategulum and embolus complex] (matured
|
||
<collectingDate id="A063CA6CFFE0FFF3B9C4FB093C17FAF3" box="[939,1039,1272,1296]" pageId="4" pageNumber="395" value="1981-07">
|
||
<date id="B0273384FFE0FFF3B9C4FB093C17FAF3" box="[939,1039,1272,1296]" pageId="4" pageNumber="395" value="1981-07">VII.1981</date>
|
||
</collectingDate>
|
||
) and female
|
||
<typeStatus id="1B22ABE6FFE0FFF3BEC7FB093D1DFAF3" box="[1192,1285,1272,1296]" pageId="4" pageNumber="395" type="paratype">paratype</typeStatus>
|
||
of
|
||
<taxonomicName id="03996EC7FFE0FFF3BF5DFB063C35FAD3" authorityName="Haupt" authorityYear="1983" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="395" phylum="Arthropoda" rank="subSpecies" species="malayanus" subSpecies="cameroni">
|
||
<emphasis id="F6EDC956FFE0FFF3BF5DFB063C35FAD3" italics="true" pageId="4" pageNumber="395">Liphistius malayanus cameroni</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<collectingRegion id="065DDBA6FFE0FFF3BE28FAE93C82FACC" box="[1095,1178,1304,1327]" country="Malaysia" name="Pahang" pageId="4" pageNumber="395">Pahang</collectingRegion>
|
||
,
|
||
<collectorName id="696C7092FFE0FFF3BEDFFAE83D80FACC" box="[1200,1432,1304,1329]" pageId="4" pageNumber="395">
|
||
Cameron
|
||
<collectingRegion id="065DDBA6FFE0FFF3BF49FAE93D80FACC" box="[1318,1432,1304,1327]" country="Czech Republic" name="Vysocina" pageId="4" pageNumber="395">Highlands</collectingRegion>
|
||
</collectorName>
|
||
,
|
||
<collectorName id="696C7092FFE0FFF3B950FAC93BA7FAAC" box="[831,959,1336,1359]" pageId="4" pageNumber="395">Berinchang</collectorName>
|
||
</materialsCitation>
|
||
;
|
||
<date id="B0273384FFE0FFF3B9BDFAC83C59FAB2" box="[978,1089,1337,1361]" pageId="4" pageNumber="395" value="1981-03-04">4.III.1981</date>
|
||
; leg. J. Haupt & T. Dach. –
|
||
<materialsCitation id="74F11F19FFE0FFF3B950FAA83BD1FA37" ID-GBIF-Occurrence="1675315685" collectingDate="2014-10-05" collectorName="N. Bay." county="Collection of Joseph Koh" latitude="4.473889" location="Taman Tringkap" longLatPrecision="21" longitude="101.38278" municipality="Cameron Highlands" pageId="4" pageNumber="395" specimenCount="1" specimenCount-male="1" stateProvince="Pahang" typeStatus="holotype">
|
||
<collectingCounty id="2D476DC8FFE0FFF3B950FAA83C6EFA93" box="[831,1142,1369,1392]" pageId="4" pageNumber="395">Collection of Joseph Koh</collectingCounty>
|
||
, n° JK.14.10.05.0001;
|
||
<specimenCount id="D29FDECDFFE0FFF3BFE0FAAB3B6FFA72" pageId="4" pageNumber="395" type="male">1 male</specimenCount>
|
||
;
|
||
<collectingRegion id="065DDBA6FFE0FFF3B9E5FA8B3BC4FA72" box="[906,988,1402,1425]" country="Malaysia" name="Pahang" pageId="4" pageNumber="395">Pahang</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="24428F3EFFE0FFF3B980FA8B3CCAFA72" box="[1007,1234,1402,1426]" pageId="4" pageNumber="395">
|
||
Cameron
|
||
<collectingRegion id="065DDBA6FFE0FFF3BE0FFA8B3CCAFA72" box="[1120,1234,1402,1425]" country="Czech Republic" name="Vysocina" pageId="4" pageNumber="395">Highlands</collectingRegion>
|
||
</collectingMunicipality>
|
||
,
|
||
<location id="C146439FFFE0FFF3BE8BFA8A3D84FA72" LSID="urn:lsid:plazi:treatment:4C30A452FFE0FFFDBA9AFAC53979F872:C146439FFFE0FFF3BE8BFA8A3D84FA72" box="[1252,1436,1402,1426]" county="Collection of Joseph Koh" latitude="4.473889" longLatPrecision="21" longitude="101.38278" municipality="Cameron Highlands" name="Taman Tringkap" pageId="4" pageNumber="395" stateProvince="Pahang">Taman Tringkap</location>
|
||
NE of
|
||
<location id="C146439FFFE0FFF3B9E2FA6B3BE7FA52" LSID="urn:lsid:plazi:treatment:4C30A452FFE0FFFDBA9AFAC53979F872:C146439FFFE0FFF3B9E2FA6B3BE7FA52" box="[909,1023,1434,1457]" county="Collection of Joseph Koh" latitude="4.473889" longLatPrecision="21" longitude="101.38278" municipality="Cameron Highlands" name="Brinchang" pageId="4" pageNumber="395" stateProvince="Pahang">Brinchang</location>
|
||
,
|
||
<geoCoordinate id="A1AD7383FFE0FFF3BE62FA6B3C9DFA52" box="[1037,1157,1434,1457]" direction="north" orientation="latitude" pageId="4" pageNumber="395" precision="15" value="4.473889">4°28’26”N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A1AD7383FFE0FFF3BEFCFA6B3D3BFA52" box="[1171,1315,1434,1457]" direction="east" orientation="longitude" pageId="4" pageNumber="395" precision="15" value="101.38278">101°22’58”E</geoCoordinate>
|
||
;
|
||
<collectingDate id="A063CA6CFFE0FFF3BF5DFA6B3D80FA52" box="[1330,1432,1434,1457]" pageId="4" pageNumber="395" value="2014-10-05">
|
||
<date id="B0273384FFE0FFF3BF5DFA6B3D80FA52" box="[1330,1432,1434,1457]" pageId="4" pageNumber="395" value="2014-10-05">5.X.2014</date>
|
||
</collectingDate>
|
||
; leg.
|
||
<collectorName id="696C7092FFE0FFF3B91CFA4D3BD1FA37" box="[883,969,1468,1492]" pageId="4" pageNumber="395">N. Bay.</collectorName>
|
||
</materialsCitation>
|
||
–
|
||
<materialsCitation id="74F11F19FFE0FFF3B982FA4D3CD9F9F0" ID-GBIF-Occurrence="1675315681" collectingDate="1995-01-21" collectionCode="MHNG" collectorName="P. J. Schwendinger." elevation="1060" location="Ringlet" municipality="Cameron Highlands" pageId="4" pageNumber="395" specimenCount="2" specimenCount-female="1" stateProvince="Perak" typeStatus="holotype">
|
||
<collectionCode id="A2888D81FFE0FFF3B982FA4D3C5AFA37" box="[1005,1090,1468,1492]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="4" pageNumber="395">MHNG</collectionCode>
|
||
;
|
||
<specimenCount id="D29FDECDFFE0FFF3BE3EFA4D3CAEFA31" box="[1105,1206,1467,1492]" pageId="4" pageNumber="395" type="female">1 female</specimenCount>
|
||
,
|
||
<specimenCount id="D29FDECDFFE0FFF3BEA9FA4D3D1FFA31" box="[1222,1287,1467,1492]" pageId="4" pageNumber="395" type="generic">1 juv.</specimenCount>
|
||
male;
|
||
<collectingRegion id="065DDBA6FFE0FFF3BF37FA4A3D80FA31" box="[1368,1432,1467,1490]" country="Malaysia" name="Perak" pageId="4" pageNumber="395">Perak</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="24428F3EFFE0FFF3B950FA2D3C04FA11" box="[831,1052,1499,1524]" pageId="4" pageNumber="395">
|
||
Cameron
|
||
<collectingRegion id="065DDBA6FFE0FFF3B9C4FA2A3C04FA11" box="[939,1052,1499,1522]" country="Czech Republic" name="Vysocina" pageId="4" pageNumber="395">Highlands</collectingRegion>
|
||
</collectingMunicipality>
|
||
, ca
|
||
<quantity id="0361B8A1FFE0FFF3BE23FA2D3C9DFA11" box="[1100,1157,1499,1524]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="395" unit="km" value="1.0">1 km</quantity>
|
||
SW of
|
||
<location id="C146439FFFE0FFF3BE8EFA2A3D2AFA11" LSID="urn:lsid:plazi:treatment:4C30A452FFE0FFFDBA9AFAC53979F872:C146439FFFE0FFF3BE8EFA2A3D2AFA11" box="[1249,1330,1499,1522]" municipality="Cameron Highlands" name="Ringlet" pageId="4" pageNumber="395" stateProvince="Perak">Ringlet</location>
|
||
,
|
||
<elevation id="4FB4F277FFE0FFF3BF2EFA2A3D80FA17" box="[1345,1432,1499,1524]" metricMagnitude="3" metricUnit="m" metricValue="1.06" pageId="4" pageNumber="395" unit="m" value="1060.0">
|
||
<quantity id="0361B8A1FFE0FFF3BF2EFA2A3D80FA17" box="[1345,1432,1499,1524]" metricMagnitude="3" metricUnit="m" metricValue="1.06" pageId="4" pageNumber="395" unit="m" value="1060.0">1060 m</quantity>
|
||
</elevation>
|
||
;
|
||
<collectingDate id="A063CA6CFFE0FFF3B950FA0D3BB1F9F0" box="[831,937,1532,1555]" pageId="4" pageNumber="395" value="1995-01-21">
|
||
<date id="B0273384FFE0FFF3B950FA0D3BB1F9F0" box="[831,937,1532,1555]" pageId="4" pageNumber="395" value="1995-01-21">21.I.1995</date>
|
||
</collectingDate>
|
||
; leg.
|
||
<collectorName id="696C7092FFE0FFF3B99FFA0C3CD9F9F0" box="[1008,1217,1532,1556]" pageId="4" pageNumber="395">P.J. Schwendinger.</collectorName>
|
||
</materialsCitation>
|
||
–
|
||
<materialsCitation id="74F11F19FFE0FFF3BE87FA0C3CF0F996" ID-GBIF-Occurrence="1675315691" collectingDate="2001-09-29" collectionCode="MHNG" collectorName="L. Monod." elevation="1210" latitude="4.4603333" location="Tanah Rata" longLatPrecision="1" longitude="101.39" municipality="Cameron Highlands" pageId="4" pageNumber="395" specimenCount="1" stateProvince="Pahang" typeStatus="holotype">
|
||
<collectionCode id="A2888D81FFE0FFF3BE87FA0C3D22F9F6" box="[1256,1338,1533,1557]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="4" pageNumber="395">MHNG</collectionCode>
|
||
(sample TM-15);
|
||
<specimenCount id="D29FDECDFFE0FFF3B9C3F9EC3BEAF9D7" box="[940,1010,1565,1589]" pageId="4" pageNumber="395" type="generic">1 juv.</specimenCount>
|
||
male;
|
||
<collectingRegion id="065DDBA6FFE0FFF3BE22F9EC3C87F9D7" box="[1101,1183,1565,1588]" country="Malaysia" name="Pahang" pageId="4" pageNumber="395">Pahang</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="24428F3EFFE0FFF3BEDBF9EC3D80F9D7" box="[1204,1432,1565,1589]" pageId="4" pageNumber="395">
|
||
Cameron
|
||
<collectingRegion id="065DDBA6FFE0FFF3BF49F9EC3D80F9D7" box="[1318,1432,1565,1588]" country="Czech Republic" name="Vysocina" pageId="4" pageNumber="395">Highlands</collectingRegion>
|
||
</collectingMunicipality>
|
||
,
|
||
<location id="C146439FFFE0FFF3B950F9CC3BDEF9B6" LSID="urn:lsid:plazi:treatment:4C30A452FFE0FFFDBA9AFAC53979F872:C146439FFFE0FFF3B950F9CC3BDEF9B6" box="[831,966,1597,1621]" latitude="4.4603333" longLatPrecision="1" longitude="101.39" municipality="Cameron Highlands" name="Tanah Rata" pageId="4" pageNumber="395" stateProvince="Pahang">Tanah Rata</location>
|
||
, trail n° 9,
|
||
<geoCoordinate id="A1AD7383FFE0FFF3BE08F9CC3CF1F9B7" box="[1127,1257,1597,1620]" direction="north" orientation="latitude" pageId="4" pageNumber="395" precision="1" value="4.4603333">4°27.620’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A1AD7383FFE0FFF3BE91F9CF3D80F9B5" box="[1278,1432,1598,1622]" direction="east" orientation="longitude" pageId="4" pageNumber="395" precision="1" value="101.39">101°23.400’E</geoCoordinate>
|
||
,
|
||
<elevation id="4FB4F277FFE0FFF3B950F9AE3B82F995" box="[831,922,1631,1655]" metricMagnitude="3" metricUnit="m" metricValue="1.21" pageId="4" pageNumber="395" unit="m" value="1210.0">
|
||
<quantity id="0361B8A1FFE0FFF3B950F9AE3B82F995" box="[831,922,1631,1655]" metricMagnitude="3" metricUnit="m" metricValue="1.21" pageId="4" pageNumber="395" unit="m" value="1210.0">1210 m</quantity>
|
||
</elevation>
|
||
;
|
||
<collectingDate id="A063CA6CFFE0FFF3B9C4F9AE3C31F994" box="[939,1065,1631,1655]" pageId="4" pageNumber="395" value="2001-09-29">
|
||
<date id="B0273384FFE0FFF3B9C4F9AE3C31F994" box="[939,1065,1631,1655]" pageId="4" pageNumber="395" value="2001-09-29">29.IX.2001</date>
|
||
</collectingDate>
|
||
; leg.
|
||
<collectorName id="696C7092FFE0FFF3BE00F9AE3CF0F996" box="[1135,1256,1630,1654]" pageId="4" pageNumber="395">L. Monod.</collectorName>
|
||
</materialsCitation>
|
||
–
|
||
<materialsCitation id="74F11F19FFE0FFF3BF60F9AE3C30F914" ID-GBIF-Occurrence="1675315680" collectionCode="SMF" collectorName="C. F. Roewer & Ginting Kial Highlands & Cameron Highlands" determinerName="Roewer" location="Pahang" pageId="4" pageNumber="395" specimenCode="SMF 7425" specimenCount="4" specimenCount-female="1" specimenCount-male="1" stateProvince="Pahang" typeStatus="holotype">
|
||
<specimenCode id="943FBD3FFFE0FFF3BF60F9AE3D9FF996" box="[1295,1415,1630,1655]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="4" pageNumber="395">SMF 7425</specimenCode>
|
||
/2 (ex coll.
|
||
<collectorName id="696C7092FFE0FFF3B9CAF98E3C26F974" box="[933,1086,1663,1687]" pageId="4" pageNumber="395">C. F. Roewer</collectorName>
|
||
);
|
||
<specimenCount id="D29FDECDFFE0FFF3BE3BF98E3C8BF976" box="[1108,1171,1662,1687]" pageId="4" pageNumber="395" type="generic">2 juv.</specimenCount>
|
||
females (labelled as “
|
||
<specimenCount id="D29FDECDFFE0FFF3BFFEF98E3B6CF955" pageId="4" pageNumber="395" type="male">1 male</specimenCount>
|
||
and
|
||
<specimenCount id="D29FDECDFFE0FFF3B9D2F9513C31F955" box="[957,1065,1695,1720]" pageId="4" pageNumber="395" type="female">1 female</specimenCount>
|
||
, det.
|
||
<determinerName id="31FAF980FFE0FFF3BE18F9513CD5F954" box="[1143,1229,1696,1719]" pageId="4" pageNumber="395">Roewer</determinerName>
|
||
, 1935”);
|
||
<collectingRegion id="065DDBA6FFE0FFF3BF29F96E3D80F955" box="[1350,1432,1695,1718]" country="Malaysia" name="Pahang" pageId="4" pageNumber="395">Pahang</collectingRegion>
|
||
, “
|
||
<collectorName id="696C7092FFE0FFF3B926F9313C79F934" box="[841,1121,1728,1751]" pageId="4" pageNumber="395">
|
||
Ginting Kial
|
||
<collectingRegion id="065DDBA6FFE0FFF3B980F9313C79F934" box="[1007,1121,1728,1751]" country="Czech Republic" name="Vysocina" pageId="4" pageNumber="395">Highlands</collectingRegion>
|
||
</collectorName>
|
||
” [probably a peak in the
|
||
<collectorName id="696C7092FFE0FFF3B950F9103C3AF914" box="[831,1058,1760,1785]" pageId="4" pageNumber="395">
|
||
Cameron
|
||
<collectingRegion id="065DDBA6FFE0FFF3B9C2F9113C3AF914" box="[941,1058,1760,1783]" country="Czech Republic" name="Vysocina" pageId="4" pageNumber="395">Highlands</collectingRegion>
|
||
</collectorName>
|
||
]
|
||
</materialsCitation>
|
||
. –
|
||
<materialsCitation id="74F11F19FFE0FFF3BE3FF9103BF4F8DB" ID-GBIF-Occurrence="1675315698" collectingDate="2001-12-19" collectionCode="SMF" collectorName="S. Huber." location="Jalan Girdle" municipality="Fraser's Hill" pageId="4" pageNumber="395" specimenCount="1" specimenCount-male="1" stateProvince="Pahang" typeStatus="holotype">
|
||
<collectionCode id="A2888D81FFE0FFF3BE3FF9103C9EF91A" box="[1104,1158,1761,1785]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="4" pageNumber="395">SMF</collectionCode>
|
||
40602;
|
||
<specimenCount id="D29FDECDFFE0FFF3BE88F9103D2FF914" box="[1255,1335,1760,1785]" pageId="4" pageNumber="395" type="male">1 male</specimenCount>
|
||
;
|
||
<collectingRegion id="065DDBA6FFE0FFF3BF29F9113D80F914" box="[1350,1432,1760,1783]" country="Malaysia" name="Pahang" pageId="4" pageNumber="395">Pahang</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="24428F3EFFE0FFF3B950F8F33BCAF8FB" box="[831,978,1793,1818]" pageId="4" pageNumber="395">Fraser’s Hill</collectingMunicipality>
|
||
, roadside at
|
||
<location id="C146439FFFE0FFF3BE1FF8F03CE7F8FB" LSID="urn:lsid:plazi:treatment:4C30A452FFE0FFFDBA9AFAC53979F872:C146439FFFE0FFF3BE1FF8F03CE7F8FB" box="[1136,1279,1793,1816]" municipality="Fraser's Hill" name="Jalan Girdle" pageId="4" pageNumber="395" stateProvince="Pahang">Jalan Girdle</location>
|
||
;
|
||
<collectingDate id="A063CA6CFFE0FFF3BF7EF8F33D80F8F9" box="[1297,1432,1794,1818]" pageId="4" pageNumber="395" value="2001-12-19">
|
||
<date id="B0273384FFE0FFF3BF7EF8F33D80F8F9" box="[1297,1432,1794,1818]" pageId="4" pageNumber="395" value="2001-12-19">19.XII.2001</date>
|
||
</collectingDate>
|
||
; leg.
|
||
<collectorName id="696C7092FFE0FFF3B915F8D33BF4F8DB" box="[890,1004,1825,1850]" pageId="4" pageNumber="395">S. Huber.</collectorName>
|
||
</materialsCitation>
|
||
–
|
||
<materialsCitation id="74F11F19FFE0FFF3BE72F8D33BFAF899" ID-GBIF-Occurrence="1675315696" collectingDate="2001-12-19" collectionCode="SMF" collectorName="S. Huber." location="Jalan Girdle" municipality="Fraser's Hill" pageId="4" pageNumber="395" specimenCount="1" specimenCount-female="1" stateProvince="Pahang" typeStatus="holotype">
|
||
<collectionCode id="A2888D81FFE0FFF3BE72F8D33C4BF8D9" box="[1053,1107,1826,1850]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="4" pageNumber="395">SMF</collectionCode>
|
||
56206;
|
||
<specimenCount id="D29FDECDFFE0FFF3BEADF8D33D28F8DB" box="[1218,1328,1825,1850]" pageId="4" pageNumber="395" type="female">1 female</specimenCount>
|
||
;
|
||
<collectingRegion id="065DDBA6FFE0FFF3BF29F8D03D81F8DB" box="[1350,1433,1825,1848]" country="Malaysia" name="Pahang" pageId="4" pageNumber="395">Pahang</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="24428F3EFFE0FFF3B950F8B23BCAF8BA" box="[831,978,1858,1883]" pageId="4" pageNumber="395">Fraser’s Hill</collectingMunicipality>
|
||
, roadside at
|
||
<location id="C146439FFFE0FFF3BE1FF8B33CE7F8BA" LSID="urn:lsid:plazi:treatment:4C30A452FFE0FFFDBA9AFAC53979F872:C146439FFFE0FFF3BE1FF8B33CE7F8BA" box="[1136,1279,1858,1881]" municipality="Fraser's Hill" name="Jalan Girdle" pageId="4" pageNumber="395" stateProvince="Pahang">Jalan Girdle</location>
|
||
;
|
||
<collectingDate id="A063CA6CFFE0FFF3BF7EF8B23D80F8B8" box="[1297,1432,1859,1883]" pageId="4" pageNumber="395" value="2001-12-19">
|
||
<date id="B0273384FFE0FFF3BF7EF8B23D80F8B8" box="[1297,1432,1859,1883]" pageId="4" pageNumber="395" value="2001-12-19">19.XII.2001</date>
|
||
</collectingDate>
|
||
; leg.
|
||
<collectorName id="696C7092FFE0FFF3B91AF8923BFAF899" box="[885,994,1891,1915]" pageId="4" pageNumber="395">S. Huber.</collectorName>
|
||
</materialsCitation>
|
||
– MHNG (ex coll. S. Huber, sample O-4, C); 1 male (matured end
|
||
<date id="B0273384FFE0FFF3BEF4F8723D3DF87F" box="[1179,1317,1923,1948]" pageId="4" pageNumber="395" value="2002-03">March 2002</date>
|
||
),
|
||
<materialsCitation id="74F11F19FFE0FFF3BF54F8753CEDF838" ID-GBIF-Occurrence="1675315693" collectingDate="2001-12-18" collectingDateMax="2002-06-24" collectingDateMin="2001-12-18" collectorName="S. Huber." location="Jalan Guillemard" municipality="Fraser's Hill" pageId="4" pageNumber="395" specimenCount="1" specimenCount-female="1" stateProvince="Pahang" typeStatus="holotype">
|
||
<specimenCount id="D29FDECDFFE0FFF3BF54F8753D84F879" box="[1339,1436,1923,1948]" pageId="4" pageNumber="395" type="female">1 female</specimenCount>
|
||
(moulted
|
||
<collectingDate id="A063CA6CFFE0FFF3B9DCF8543C2CF85E" box="[947,1076,1957,1981]" pageId="4" pageNumber="395" value="2002-06-24">
|
||
<date id="B0273384FFE0FFF3B9DCF8543C2CF85E" box="[947,1076,1957,1981]" pageId="4" pageNumber="395" value="2002-06-24">24.VI.2002</date>
|
||
</collectingDate>
|
||
);
|
||
<collectingRegion id="065DDBA6FFE0FFF3BE3EF8553CBCF858" box="[1105,1188,1956,1979]" country="Malaysia" name="Pahang" pageId="4" pageNumber="395">Pahang</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="24428F3EFFE0FFF3BED6F8543D57F858" box="[1209,1359,1956,1981]" pageId="4" pageNumber="395">Fraser’s Hill</collectingMunicipality>
|
||
,
|
||
<location id="C146439FFFE0FFF3BF0AF8553BA4F838" LSID="urn:lsid:plazi:treatment:4C30A452FFE0FFFDBA9AFAC53979F872:C146439FFFE0FFF3BF0AF8553BA4F838" municipality="Fraser's Hill" name="Jalan Guillemard" pageId="4" pageNumber="395" stateProvince="Pahang">Jalan Guillemard</location>
|
||
;
|
||
<collectingDate id="A063CA6CFFE0FFF3B9A5F8343C57F83E" box="[970,1103,1989,2013]" pageId="4" pageNumber="395" value="2001-12-18">
|
||
<date id="B0273384FFE0FFF3B9A5F8343C57F83E" box="[970,1103,1989,2013]" pageId="4" pageNumber="395" value="2001-12-18">18.XII.2001</date>
|
||
</collectingDate>
|
||
; leg.
|
||
<collectorName id="696C7092FFE0FFF3BEE2F8343CEDF838" box="[1165,1269,1988,2013]" pageId="4" pageNumber="395">S. Huber.</collectorName>
|
||
</materialsCitation>
|
||
–
|
||
<materialsCitation id="74F11F19FFE0FFF3BF7BF8343D05F7FC" ID-GBIF-Occurrence="1675315695" collectingDate="2013-06-17" collectionCode="SMF" collectorName="P. Jager." elevation="1300" latitude="3.7184167" location="Fraser's Hill" longLatPrecision="1" longitude="101.75273" pageId="4" pageNumber="395" specimenCount="1" specimenCount-female="1" stateProvince="Pahang" typeStatus="holotype">
|
||
<collectionCode id="A2888D81FFE0FFF3BF7BF8343D51F83E" box="[1300,1353,1989,2013]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="4" pageNumber="395">SMF</collectionCode>
|
||
64093;
|
||
<specimenCount id="D29FDECDFFE0FFF3B950F8173BBDF81F" box="[831,933,2021,2046]" pageId="4" pageNumber="395" type="female">1 female</specimenCount>
|
||
;
|
||
<collectingRegion id="065DDBA6FFE0FFF3B9DCF8143C1DF81F" box="[947,1029,2021,2044]" country="Malaysia" name="Pahang" pageId="4" pageNumber="395">Pahang</collectingRegion>
|
||
,
|
||
<location id="C146439FFFE0FFF3BE7BF8173CB9F81F" LSID="urn:lsid:plazi:treatment:4C30A452FFE0FFFDBA9AFAC53979F872:C146439FFFE0FFF3BE7BF8173CB9F81F" box="[1044,1185,2021,2046]" latitude="3.7184167" longLatPrecision="1" longitude="101.75273" name="Fraser's Hill" pageId="4" pageNumber="395" stateProvince="Pahang">Fraser’s Hill</location>
|
||
,
|
||
<elevation id="4FB4F277FFE0FFF3BEDFF8173D10F81E" box="[1200,1288,2022,2046]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="395" unit="m" value="1300.0">
|
||
<quantity id="0361B8A1FFE0FFF3BEDFF8173D10F81E" box="[1200,1288,2022,2046]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="395" unit="m" value="1300.0">1300 m</quantity>
|
||
</elevation>
|
||
,
|
||
<geoCoordinate id="A1AD7383FFE0FFF3BF78F8143D80F81F" box="[1303,1432,2021,2044]" direction="north" orientation="latitude" pageId="4" pageNumber="395" precision="1" value="3.7184167">3°43.105’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A1AD7383FFE0FFF3B950F7F73BC2F7FE" box="[831,986,2054,2077]" direction="east" orientation="longitude" pageId="4" pageNumber="395" precision="1" value="101.75273">101°45.164’E</geoCoordinate>
|
||
;
|
||
<collectingDate id="A063CA6CFFE0FFF3B980F7F73C76F7FD" box="[1007,1134,2054,2078]" pageId="4" pageNumber="395" value="2013-06-17">
|
||
<date id="B0273384FFE0FFF3B980F7F73C76F7FD" box="[1007,1134,2054,2078]" pageId="4" pageNumber="395" value="2013-06-17">17.VI.2013</date>
|
||
</collectingDate>
|
||
; leg.
|
||
<collectorName id="696C7092FFE0FFF3BED5F7F63D05F7FC" box="[1210,1309,2055,2079]" pageId="4" pageNumber="395">P. Jäger.</collectorName>
|
||
</materialsCitation>
|
||
– MHNG (sample MAL-04/02); 1 male (matured
|
||
<date id="B0273384FFE3FFF0B806FEF23AF0FEF8" box="[617,744,259,283]" pageId="7" pageNumber="398" value="2004-09-18">18.IX.2004</date>
|
||
), 2 females (moulted
|
||
<date id="B0273384FFE3FFF0BBCEFED33A32FED9" box="[417,554,290,314]" pageId="7" pageNumber="398" value="2004-07-30">30.VII.2004</date>
|
||
, 24.I.,
|
||
<date id="B0273384FFE3FFF0B8FAFED33AE8FED9" box="[661,752,290,314]" pageId="7" pageNumber="398" value="2005-08">28.VIII.</date>
|
||
,
|
||
<date id="B0273384FFE3FFF0BAF8FEB0390BFEBB" box="[151,275,321,344]" pageId="7" pageNumber="398" value="2005-12-08">8.XII.2005</date>
|
||
;
|
||
<date id="B0273384FFE3FFF0BB43FEB339BAFEB9" box="[300,418,322,346]" pageId="7" pageNumber="398" value="2004-10-19">19.X.2004</date>
|
||
,
|
||
<date id="B0273384FFE3FFF0BBD3FEB03A5DFEBB" box="[444,581,321,344]" pageId="7" pageNumber="398" value="2005-07-30">30.VII.2005</date>
|
||
),
|
||
<materialsCitation id="74F11F19FFE3FFF0B808FEB33AB8FE74" ID-GBIF-Occurrence="1675315679" collectingDate="2004-05-18" collectingDateMax="2004-05-19" collectingDateMin="2004-05-18" collectorName="P. J. Schwendinger." elevation="1650" latitude="3.4283335" location="Genting Highlands" longLatPrecision="21" longitude="101.79472" pageId="7" pageNumber="398" specimenCount="4" specimenCount-juvenile="4" stateProvince="Pahang" typeStatus="holotype">
|
||
<specimenCount id="D29FDECDFFE3FFF0B808FEB33AE8FEBB" box="[615,752,321,346]" pageId="7" pageNumber="398" type="juvenile">4 juveniles</specimenCount>
|
||
;
|
||
<collectingRegion id="065DDBA6FFE3FFF0BAF8FE9138F1FE94" box="[151,233,352,375]" country="Malaysia" name="Pahang" pageId="7" pageNumber="398">Pahang</collectingRegion>
|
||
,
|
||
<location id="C146439FFFE3FFF0BA98FE9139D1FE94" LSID="urn:lsid:plazi:treatment:4C30A452FFE0FFFDBA9AFAC53979F872:C146439FFFE3FFF0BA98FE9139D1FE94" box="[247,457,352,375]" latitude="3.4283335" longLatPrecision="21" longitude="101.79472" name="Genting Highlands" pageId="7" pageNumber="398" stateProvince="Pahang">
|
||
Genting
|
||
<collectingRegion id="065DDBA6FFE3FFF0BB37FE9139D1FE94" box="[344,457,352,375]" country="Czech Republic" name="Vysocina" pageId="7" pageNumber="398">Highlands</collectingRegion>
|
||
</location>
|
||
,
|
||
<geoCoordinate id="A1AD7383FFE3FFF0BBB7FE913A48FE94" box="[472,592,352,375]" direction="north" orientation="latitude" pageId="7" pageNumber="398" precision="15" value="3.4283335">3°25’42”N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A1AD7383FFE3FFF0B830FE903AE8FE9A" box="[607,752,353,377]" direction="east" orientation="longitude" pageId="7" pageNumber="398" precision="15" value="101.79472">101°47’41”E</geoCoordinate>
|
||
,
|
||
<elevation id="4FB4F277FFE3FFF0BAF8FE7138F6FE7B" box="[151,238,384,408]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="7" pageNumber="398" unit="m" value="1650.0">
|
||
<quantity id="0361B8A1FFE3FFF0BAF8FE7138F6FE7B" box="[151,238,384,408]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="7" pageNumber="398" unit="m" value="1650.0">1650 m</quantity>
|
||
</elevation>
|
||
;
|
||
<collectingDate id="A063CA6CFFE3FFF0BA94FE70398DFE7A" box="[251,405,385,409]" pageId="7" pageNumber="398" value="2004-05-18" valueMax="2004-05-19" valueMin="2004-05-18">
|
||
<date id="B0273384FFE3FFF0BA94FE70398DFE7A" box="[251,405,385,409]" pageId="7" pageNumber="398" value="2004-05-18" valueMax="2004-05-19" valueMin="2004-05-18">18.-19.V.2004</date>
|
||
</collectingDate>
|
||
; leg.
|
||
<collectorName id="696C7092FFE3FFF0BBBBFE703AB8FE74" box="[468,672,384,408]" pageId="7" pageNumber="398">P.J. Schwendinger.</collectorName>
|
||
</materialsCitation>
|
||
– SMF 21946/1; 1 female; without locality data;
|
||
<date id="B0273384FFE3FFF0B81DFE513AF7FE5B" box="[626,751,416,440]" pageId="7" pageNumber="398" value="1933-11-10">10.XI.1933</date>
|
||
; ex coll. Wiehle, don. W. S. Bristowe. –
|
||
<materialsCitation id="74F11F19FFE3FFF0B807FE313917FDD6" ID-GBIF-Occurrence="1675315675" collectingDate="1989-02-03" collectionCode="SMF" collectorName="U. Maschwitz & H. Steiner." elevation="1800" location="Genting Highlands" pageId="7" pageNumber="398" specimenCount="1" specimenCount-female="1" stateProvince="Selangor" typeStatus="holotype">
|
||
<collectionCode id="A2888D81FFE3FFF0B807FE313A86FE3B" box="[616,670,448,472]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="7" pageNumber="398">SMF</collectionCode>
|
||
60037;
|
||
<specimenCount id="D29FDECDFFE3FFF0BAF8FE2E391AFE15" box="[151,258,479,503]" pageId="7" pageNumber="398" type="female">1 female</specimenCount>
|
||
; “
|
||
<collectingRegion id="065DDBA6FFE3FFF0BB4EFE2E399EFE15" box="[289,390,479,502]" country="Malaysia" name="Selangor" pageId="7" pageNumber="398">Selangor</collectingRegion>
|
||
” [should be
|
||
<collectingRegion id="065DDBA6FFE3FFF0B842FE2E3A9AFE15" box="[557,642,479,502]" country="Malaysia" name="Pahang" pageId="7" pageNumber="398">Pahang</collectingRegion>
|
||
],
|
||
<location id="C146439FFFE3FFF0B8F2FE2E391FFDF6" LSID="urn:lsid:plazi:treatment:4C30A452FFE0FFFDBA9AFAC53979F872:C146439FFFE3FFF0B8F2FE2E391FFDF6" name="Genting Highlands" pageId="7" pageNumber="398" stateProvince="Selangor">
|
||
Genting
|
||
<collectingRegion id="065DDBA6FFE3FFF0BAF8FE0F391FFDF6" box="[151,263,510,533]" country="Czech Republic" name="Vysocina" pageId="7" pageNumber="398">Highlands</collectingRegion>
|
||
</location>
|
||
,
|
||
<elevation id="4FB4F277FFE3FFF0BB7AFE0E3974FDF5" box="[277,364,511,535]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="7" pageNumber="398" unit="m" value="1800.0">
|
||
<quantity id="0361B8A1FFE3FFF0BB7AFE0E3974FDF5" box="[277,364,511,535]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="7" pageNumber="398" unit="m" value="1800.0">1800 m</quantity>
|
||
</elevation>
|
||
;
|
||
<collectingDate id="A063CA6CFFE3FFF0BB16FE0E39C6FDF4" box="[377,478,511,535]" pageId="7" pageNumber="398" value="1989-02-03">
|
||
<date id="B0273384FFE3FFF0BB16FE0E39C6FDF4" box="[377,478,511,535]" pageId="7" pageNumber="398" value="1989-02-03">3.II.1989</date>
|
||
</collectingDate>
|
||
; leg.
|
||
<collectorName id="696C7092FFE3FFF0B872FE0E3AA0FDF6" box="[541,696,510,534]" pageId="7" pageNumber="398">U. Maschwitz</collectorName>
|
||
, don.
|
||
<collectorName id="696C7092FFE3FFF0BAF8FDEE3917FDD6" box="[151,271,542,566]" pageId="7" pageNumber="398">H. Steiner.</collectorName>
|
||
</materialsCitation>
|
||
–
|
||
<materialsCitation id="74F11F19FFE3FFF0BB5DFDEF3AC0FDB6" ID-GBIF-Occurrence="1675315697" collectingDate="2001-07-27" collectionCode="SMF" collectorName="A. Kovac." elevation="1200" location="Genting Highlands" pageId="7" pageNumber="398" specimenCount="1" specimenCount-female="1" stateProvince="Pahang" typeStatus="holotype">
|
||
<collectionCode id="A2888D81FFE3FFF0BB5DFDEF3970FDD5" box="[306,360,542,566]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="7" pageNumber="398">SMF</collectionCode>
|
||
40016;
|
||
<specimenCount id="D29FDECDFFE3FFF0BBA7FDEF3A36FDD6" box="[456,558,542,566]" pageId="7" pageNumber="398" type="female">1 female</specimenCount>
|
||
;
|
||
<collectingRegion id="065DDBA6FFE3FFF0B853FDEF3A96FDD6" box="[572,654,542,565]" country="Malaysia" name="Pahang" pageId="7" pageNumber="398">Pahang</collectingRegion>
|
||
,
|
||
<location id="C146439FFFE3FFF0B8F2FDEF3911FDB7" LSID="urn:lsid:plazi:treatment:4C30A452FFE0FFFDBA9AFAC53979F872:C146439FFFE3FFF0B8F2FDEF3911FDB7" name="Genting Highlands" pageId="7" pageNumber="398" stateProvince="Pahang">
|
||
Genting
|
||
<collectingRegion id="065DDBA6FFE3FFF0BAF8FDCC3911FDB7" box="[151,265,573,596]" country="Czech Republic" name="Vysocina" pageId="7" pageNumber="398">Highlands</collectingRegion>
|
||
</location>
|
||
,
|
||
<elevation id="4FB4F277FFE3FFF0BB72FDCF3962FDB6" box="[285,378,574,598]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="398" unit="m" value="1200.0">
|
||
<quantity id="0361B8A1FFE3FFF0BB72FDCF3962FDB6" box="[285,378,574,598]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="398" unit="m" value="1200.0">1200 m</quantity>
|
||
</elevation>
|
||
;
|
||
<collectingDate id="A063CA6CFFE3FFF0BBE1FDCF3A0EFDB5" box="[398,534,574,598]" pageId="7" pageNumber="398" value="2001-07-27">
|
||
<date id="B0273384FFE3FFF0BBE1FDCF3A0EFDB5" box="[398,534,574,598]" pageId="7" pageNumber="398" value="2001-07-27">27.VII.2001</date>
|
||
</collectingDate>
|
||
; leg.
|
||
<collectorName id="696C7092FFE3FFF0B80FFDCF3AC0FDB6" box="[608,728,574,598]" pageId="7" pageNumber="398">A. Kovac.</collectorName>
|
||
</materialsCitation>
|
||
– MHNG (sample SIM-01/14); 1 male (matured end
|
||
<date id="B0273384FFE3FFF0BAF8FD8C391DFD76" box="[151,261,637,661]" pageId="7" pageNumber="398" value="2003-08">VIII.2003</date>
|
||
), 1 female (moulted
|
||
<date id="B0273384FFE3FFF0BB88FD8C3A75FD76" box="[487,621,637,661]" pageId="7" pageNumber="398" value="2001-12-31">31.XII.2001</date>
|
||
),
|
||
<materialsCitation id="74F11F19FFE3FFF0B8EEFD8C3A68FD31" ID-GBIF-Occurrence="1675315683" collectingDate="2001-07-13" collectorName="P. J. Schwendinger." elevation="230" latitude="3.2986112" location="Templer Park" longLatPrecision="21" longitude="101.62028" pageId="7" pageNumber="398" specimenCount="1" specimenCount-juvenile="1" stateProvince="Selangor" typeStatus="holotype">
|
||
<specimenCount id="D29FDECDFFE3FFF0B8EEFD8C3AE8FD70" box="[641,752,636,661]" pageId="7" pageNumber="398" type="juvenile">1 juvenile</specimenCount>
|
||
;
|
||
<collectingRegion id="065DDBA6FFE3FFF0BAF8FD6D38E2FD50" box="[151,250,668,691]" country="Malaysia" name="Selangor" pageId="7" pageNumber="398">Selangor</collectingRegion>
|
||
,
|
||
<location id="C146439FFFE3FFF0BB7EFD6D39ACFD50" LSID="urn:lsid:plazi:treatment:4C30A452FFE0FFFDBA9AFAC53979F872:C146439FFFE3FFF0BB7EFD6D39ACFD50" box="[273,436,668,691]" latitude="3.2986112" longLatPrecision="21" longitude="101.62028" name="Templer Park" pageId="7" pageNumber="398" stateProvince="Selangor">Templer Park</location>
|
||
,
|
||
<geoCoordinate id="A1AD7383FFE3FFF0BBA4FD6D3A5EFD50" box="[459,582,668,691]" direction="north" orientation="latitude" pageId="7" pageNumber="398" precision="15" value="3.2986112">3°17’55”N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A1AD7383FFE3FFF0B832FD6C3AE8FD56" box="[605,752,669,693]" direction="east" orientation="longitude" pageId="7" pageNumber="398" precision="15" value="101.62028">101°37’13”E</geoCoordinate>
|
||
,
|
||
<elevation id="4FB4F277FFE3FFF0BAF8FD4D38C6FD30" box="[151,222,700,724]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="7" pageNumber="398" unit="m" value="230.0">
|
||
<quantity id="0361B8A1FFE3FFF0BAF8FD4D38C6FD30" box="[151,222,700,724]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="7" pageNumber="398" unit="m" value="230.0">230 m</quantity>
|
||
</elevation>
|
||
;
|
||
<collectingDate id="A063CA6CFFE3FFF0BA86FD4D3976FD37" box="[233,366,700,724]" pageId="7" pageNumber="398" value="2001-07-13">
|
||
<date id="B0273384FFE3FFF0BA86FD4D3976FD37" box="[233,366,700,724]" pageId="7" pageNumber="398" value="2001-07-13">13.VII.2001</date>
|
||
</collectingDate>
|
||
; leg.
|
||
<collectorName id="696C7092FFE3FFF0BBC8FD4D3A68FD31" box="[423,624,699,723]" pageId="7" pageNumber="398">P.J. Schwendinger.</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="90E645CCFFE1FFF2BAF8F8653981F7FE" httpUri="https://zenodo.org/record/1021943/files/figure.png" pageId="5" pageNumber="396" targetBox="[261,1329,261,1882]" targetPageId="5">
|
||
<paragraph id="C4261544FFE1FFF2BAF8F8653981F7FE" blockId="5.[151,1437,1940,2077]" pageId="5" pageNumber="396">
|
||
Fig. 3. SEM-micrographs of embolus proper of left palpal organ of two males:
|
||
<taxonomicName id="03996EC7FFE1FFF2B9F8F8653C53F84A" authorityName="Schiodte" authorityYear="1849" box="[919,1099,1940,1961]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="396" phylum="Arthropoda" rank="species" species="desultor">
|
||
<emphasis id="F6EDC956FFE1FFF2B9F8F8653C53F84A" box="[919,1099,1940,1961]" italics="true" pageId="5" pageNumber="396">Liphistius desultor</emphasis>
|
||
</taxonomicName>
|
||
from Penang Hill (A, C, E, G) and
|
||
<taxonomicName id="03996EC7FFE1FFF2BA8CF84039AAF825" authorityName="Abraham" authorityYear="1923" box="[227,434,1969,1990]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="396" phylum="Arthropoda" rank="species" species="malayanus">
|
||
<emphasis id="F6EDC956FFE1FFF2BA8CF84039AAF825" box="[227,434,1969,1990]" italics="true" pageId="5" pageNumber="396">Liphistius malayanus</emphasis>
|
||
</taxonomicName>
|
||
from Fraser’s Hill (B, D, F, H). (A) Retroventral-distal view. (B) Dorsal-distal view. (C) Distal view (dorsal side to the right). (D) Distal view (prolateral side to the right). (E) Retrodorsal view. (F) Prodorsal view. (G) Proventral view. (H) Prolateral view. Abbreviations: f - major embolic fold; m - membranous part of embolus proper; s - sclerotised part of embolus proper.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="90E645CCFFE2FFF1BAF8F9403B27F872" httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="6" pageNumber="397" targetBox="[156,1432,265,1656]" targetPageId="6">
|
||
<paragraph id="C4261544FFE2FFF1BAF8F9403B27F872" blockId="6.[151,1437,1713,1937]" pageId="6" pageNumber="397">
|
||
Fig. 4.
|
||
<taxonomicName id="03996EC7FFE2FFF1BA8CF94039B7F925" authorityName="Abraham" authorityYear="1923" box="[227,431,1713,1734]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="397" phylum="Arthropoda" rank="species" species="malayanus">
|
||
<emphasis id="F6EDC956FFE2FFF1BA8CF94039B7F925" box="[227,431,1713,1734]" italics="true" pageId="6" pageNumber="397">Liphistius malayanus</emphasis>
|
||
</taxonomicName>
|
||
, details of palp of four males: Templer Park (A, C-D, H-I); holotype of
|
||
<taxonomicName id="03996EC7FFE2FFF1BE36F9433D22F924" authorityName="Haupt" authorityYear="1983" box="[1113,1338,1713,1735]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="397" phylum="Arthropoda" rank="subSpecies" species="malayanus" subSpecies="cameroni">
|
||
<emphasis id="F6EDC956FFE2FFF1BE36F9433D22F924" box="[1113,1338,1713,1735]" italics="true" pageId="6" pageNumber="397">L. malayanus cameroni</emphasis>
|
||
</taxonomicName>
|
||
(B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="8C8346CFFFE3FFF0BAF8FD1B3991FA25" pageId="7" pageNumber="398" type="diagnosis">
|
||
<paragraph id="C4261544FFE3FFF0BAF8FD1B3AA8FA94" blockId="7.[151,757,746,1401]" pageId="7" pageNumber="398">
|
||
<emphasis id="F6EDC956FFE3FFF0BAF8FD1B3916FCE2" bold="true" box="[151,270,746,769]" pageId="7" pageNumber="398">Diagnosis:</emphasis>
|
||
Large, dark-coloured species in both sexes. Males distinguished from those of other species in the
|
||
<taxonomicName id="03996EC7FFE3FFF0BAF8FCD83913FCA3" authorityName="Abraham" authorityYear="1923" box="[151,267,809,832]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="398" phylum="Arthropoda" rank="species" species="malayanus">
|
||
<emphasis id="F6EDC956FFE3FFF0BAF8FCD83913FCA3" box="[151,267,809,832]" italics="true" pageId="7" pageNumber="398">malayanus</emphasis>
|
||
</taxonomicName>
|
||
-group by tibial apophysis situated distinctly lower (more proximal) than distal margin of palpal tibia (
|
||
<figureCitation id="5CA209C1FFE3FFF0BAB7FC993936FC62" box="[216,302,872,897]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="7" pageNumber="398">Fig. 4B</figureCitation>
|
||
); distal edge of contrategulum carrying a series of small denticles (
|
||
<figureCitation id="5CA209C1FFE3FFF0BBC7FC7939F1FC43" box="[424,489,904,928]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="7" pageNumber="398">Fig. 4</figureCitation>
|
||
G-L), those at proventral end arranged in a V-shaped row (
|
||
<figureCitation id="5CA209C1FFE3FFF0B85DFC563A89FC23" box="[562,657,935,960]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="7" pageNumber="398">Fig. 4C</figureCitation>
|
||
), 1-2 at prodorsal end isolated and slightly enlarged; dorsal apex of contrategulum large, wide, tounge-shaped and asymmetrical, its prolateral margin more strongly arched than retrolateral margin (
|
||
<figureCitation id="5CA209C1FFE3FFF0BB99FBD43A20FBDD" box="[502,568,1061,1086]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="7" pageNumber="398">Fig. 4</figureCitation>
|
||
G-L); dorsal wall of embolus proper as wide as ventral wall (
|
||
<figureCitation id="5CA209C1FFE3FFF0B81CFBB43AADFBBE" box="[627,693,1093,1117]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="7" pageNumber="398">Fig. 4</figureCitation>
|
||
G-H); membranous part of embolus proper distally narrower than proximally (
|
||
<figureCitation id="5CA209C1FFE3FFF0BB3FFB7539DDFB7F" box="[336,453,1156,1181]" captionStart="Fig. 3" captionStartId="5.[151,186,1940,1961]" captionTargetBox="[261,1329,261,1882]" captionTargetId="figure@5.[255,1334,257,1886]" captionTargetPageId="5" captionText="Fig. 3. SEM-micrographs of embolus proper of left palpal organ of two males: Liphistius desultor from Penang Hill (A, C, E, G) and Liphistius malayanus from Fraser’s Hill (B, D, F, H). (A) Retroventral-distal view. (B) Dorsal-distal view. (C) Distal view (dorsal side to the right). (D) Distal view (prolateral side to the right). (E) Retrodorsal view. (F) Prodorsal view. (G) Proventral view. (H) Prolateral view. Abbreviations: f - major embolic fold; m - membranous part of embolus proper; s - sclerotised part of embolus proper." httpUri="https://zenodo.org/record/1021943/files/figure.png" pageId="7" pageNumber="398">Figs 3F, H</figureCitation>
|
||
,
|
||
<figureCitation id="5CA209C1FFE3FFF0BBBFFB7439EBFB7E" box="[464,499,1157,1181]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="7" pageNumber="398">4C</figureCitation>
|
||
). Females distinguished by vulval plate (
|
||
<figureCitation id="5CA209C1FFE3FFF0BB3EFB52398DFB59" box="[337,405,1187,1210]" captionStart="Fig. 5" captionStartId="8.[151,186,1809,1830]" captionTargetBox="[154,1439,260,1762]" captionTargetId="figure@8.[148,1439,254,1779]" captionTargetPageId="8" captionText="Fig. 5. Liphistius malayanus, vulval plate of eight females: paratype of L. m. cameroni from Berinchang, Cameron Highlands (A- B); exuvia of female from Ringlet, Cameron Highlands (C); female (SMF 64093) from Fraser’s Hill (D); female (moulted 24. VI. 2002) from Fraser’s Hill; poreplate slightly deformed by brief immersion in cold KOH (E); female (SMF 56206) from Fraser’s Hill (F); female (SMF 60037) from Genting Highlands (G); female (moulted 8. XII. 2005) from Genting Highlands (H); exuvia of female from Templer Park (I-J). (A, C-I) Dorsal view. (B, J) Ventral view. Scale lines: 1.0 mm (A-B; C, F; D; E, G; H; I-J)." httpUri="https://zenodo.org/record/1021947/files/figure.png" pageId="7" pageNumber="398">Fig. 5</figureCitation>
|
||
) with widely trapezium-shaped (posteriorly widest) posterior stalk not connected to pigmented lateral patches in genital atrium; poreplate with distinct anterolateral lobes; CDO large, very variable in shape, mostly longer than wide; receptacular cluster very large, in most cases protruding beyond straight or procurved anterior margin of poreplate.
|
||
</paragraph>
|
||
<paragraph id="C4261544FFE3FFF0BAF8FA7E3991FA25" blockId="7.[151,757,1423,2078]" pageId="7" pageNumber="398">
|
||
<emphasis id="F6EDC956FFE3FFF0BAF8FA7E3A5FFA4B" bold="true" box="[151,583,1423,1448]" pageId="7" pageNumber="398">Additions to description of male:</emphasis>
|
||
<emphasis id="F6EDC956FFE3FFF0B835FA7E3AAAFA45" box="[602,690,1423,1446]" italics="true" pageId="7" pageNumber="398">Scopula</emphasis>
|
||
: See paragraph “Variation”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8C8346CFFFE3FFFDBAF8FA3F38E5FD9D" lastPageId="10" lastPageNumber="401" pageId="7" pageNumber="398" type="description">
|
||
<paragraph id="C4261544FFE3FFF0BAF8FA3F3D8EFD79" blockId="7.[151,757,1423,2078]" lastBlockId="7.[831,1437,258,668]" pageId="7" pageNumber="398">
|
||
<emphasis id="F6EDC956FFE3FFF0BAF8FA3F38D1FA06" box="[151,201,1486,1509]" italics="true" pageId="7" pageNumber="398">Palp</emphasis>
|
||
: Both apical lobes of cymbium equally short, dorsal one usually slightly more pointed than ventral one. Retrolateral apophysis of palpal tibia entire, situated at a clearly more proximal level than distal margin of article (
|
||
<figureCitation id="5CA209C1FFE3FFF0BA83F9BD395BF986" box="[236,323,1612,1637]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="7" pageNumber="398">Fig. 4B</figureCitation>
|
||
), carrying four long apical megaspines (
|
||
<figureCitation id="5CA209C1FFE3FFF0BAF0F99D38F8F966" box="[159,224,1644,1669]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="7" pageNumber="398">Fig. 4</figureCitation>
|
||
A-B). Paracymbium relatively small and shallow, about as deep as cymbium in retroventral view (
|
||
<figureCitation id="5CA209C1FFE3FFF0B8A6F97A38A2F927" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="7" pageNumber="398">Fig. 4A</figureCitation>
|
||
;
|
||
<bibRefCitation id="A00868B5FFE3FFF0BAA7F95A39E0F927" author="Platnick N. & Sedgwick W. C." box="[200,504,1707,1732]" journalOrPublisher="American Museum Novitates" pageId="7" pageNumber="398" pagination="1 - 31" part="2781" refString="Platnick N. I., Sedgwick W. C. 1984. A revision of the spider genus Liphistius (Araneae, Mesothelae). American Museum Novitates 2781: 1 - 31." title="A revision of the spider genus Liphistius (Araneae, Mesothelae)" type="journal article" year="1984">Platnick & Sedgwick, 1984</bibRefCitation>
|
||
: fig. 64); cumulus low, carrying a compact group of moderately long thick bristles. Subtegulum without apophysis. Tegulum with quite few small teeth only in retrodorsal portion of proximal edge, distal margin not drawn into an edge (
|
||
<figureCitation id="5CA209C1FFE3FFF0BAF0F8B938F8F882" box="[159,224,1864,1889]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="7" pageNumber="398">Fig. 4</figureCitation>
|
||
D-F). Contrategulum with very indistinct, widely rounded ventral process (
|
||
<figureCitation id="5CA209C1FFE3FFF0BBC5F89939F4F862" box="[426,492,1896,1921]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="7" pageNumber="398">Fig. 4</figureCitation>
|
||
G-H); distal edge with a series of small denticles, those at proventral end arranged in a V-shaped row (
|
||
<figureCitation id="5CA209C1FFE3FFF0BB03F85639D8F823" box="[364,448,1959,1984]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="7" pageNumber="398">Fig. 4C</figureCitation>
|
||
), 1-2 enlarged denticles or a sharp keel in prolateral to prodorsal section of distal edge (
|
||
<figureCitation id="5CA209C1FFE3FFF0BAF0F81738F8F81C" box="[159,224,2022,2047]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="7" pageNumber="398">Fig. 4</figureCitation>
|
||
G-L); dorsal apex of contrategulum developed as a large, linguiform, strongly projecting horizontal plate, its prolateral margin being more strongly arched than its retrolateral margin (
|
||
<figureCitation id="5CA209C1FFE3FFF0BE75FED33C44FED8" box="[1050,1116,290,315]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="7" pageNumber="398">Fig. 4</figureCitation>
|
||
G-L). Embolus complex with para-embolic plate developed as a more or less distinctly elevated, rounded edge (
|
||
<figureCitation id="5CA209C1FFE3FFF0BE20FE933C8BFE98" box="[1103,1171,354,379]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="7" pageNumber="398">Fig. 4</figureCitation>
|
||
D-F); below it a second small edge present in some cases (
|
||
<figureCitation id="5CA209C1FFE3FFF0BED3FE733D0CFE78" box="[1212,1300,386,411]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="7" pageNumber="398">Fig. 4D</figureCitation>
|
||
); sclerotised and membranous part of embolus proper in contact for almost their entire length (
|
||
<figureCitation id="5CA209C1FFE3FFF0BE38FE333CFFFE38" box="[1111,1255,450,475]" captionStart="Fig. 3" captionStartId="5.[151,186,1940,1961]" captionTargetBox="[261,1329,261,1882]" captionTargetId="figure@5.[255,1334,257,1886]" captionTargetPageId="5" captionText="Fig. 3. SEM-micrographs of embolus proper of left palpal organ of two males: Liphistius desultor from Penang Hill (A, C, E, G) and Liphistius malayanus from Fraser’s Hill (B, D, F, H). (A) Retroventral-distal view. (B) Dorsal-distal view. (C) Distal view (dorsal side to the right). (D) Distal view (prolateral side to the right). (E) Retrodorsal view. (F) Prodorsal view. (G) Proventral view. (H) Prolateral view. Abbreviations: f - major embolic fold; m - membranous part of embolus proper; s - sclerotised part of embolus proper." httpUri="https://zenodo.org/record/1021943/files/figure.png" pageId="7" pageNumber="398">Fig. 3D, F, H</figureCitation>
|
||
); sclerotized part with numerous longitudinal ribs carrying tiny denticles distally (
|
||
<figureCitation id="5CA209C1FFE3FFF0B9CFFDF23C43FDF8" box="[928,1115,515,539]" captionStart="Fig. 3" captionStartId="5.[151,186,1940,1961]" captionTargetBox="[261,1329,261,1882]" captionTargetId="figure@5.[255,1334,257,1886]" captionTargetPageId="5" captionText="Fig. 3. SEM-micrographs of embolus proper of left palpal organ of two males: Liphistius desultor from Penang Hill (A, C, E, G) and Liphistius malayanus from Fraser’s Hill (B, D, F, H). (A) Retroventral-distal view. (B) Dorsal-distal view. (C) Distal view (dorsal side to the right). (D) Distal view (prolateral side to the right). (E) Retrodorsal view. (F) Prodorsal view. (G) Proventral view. (H) Prolateral view. Abbreviations: f - major embolic fold; m - membranous part of embolus proper; s - sclerotised part of embolus proper." httpUri="https://zenodo.org/record/1021943/files/figure.png" pageId="7" pageNumber="398">Fig. 3B, D, F, H</figureCitation>
|
||
), its dorsal and ventral walls equally wide and parallel to each other (
|
||
<figureCitation id="5CA209C1FFE3FFF0BE84FDD23D5CFDDF" box="[1259,1348,547,572]" captionStart="Fig. 3" captionStartId="5.[151,186,1940,1961]" captionTargetBox="[261,1329,261,1882]" captionTargetId="figure@5.[255,1334,257,1886]" captionTargetPageId="5" captionText="Fig. 3. SEM-micrographs of embolus proper of left palpal organ of two males: Liphistius desultor from Penang Hill (A, C, E, G) and Liphistius malayanus from Fraser’s Hill (B, D, F, H). (A) Retroventral-distal view. (B) Dorsal-distal view. (C) Distal view (dorsal side to the right). (D) Distal view (prolateral side to the right). (E) Retrodorsal view. (F) Prodorsal view. (G) Proventral view. (H) Prolateral view. Abbreviations: f - major embolic fold; m - membranous part of embolus proper; s - sclerotised part of embolus proper." httpUri="https://zenodo.org/record/1021943/files/figure.png" pageId="7" pageNumber="398">Figs 3D</figureCitation>
|
||
; 4G-H), dorsal wall ending in indistinct rounded lobe (
|
||
<figureCitation id="5CA209C1FFE3FFF0BF4BFDB23D7BFDBF" box="[1316,1379,579,604]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="7" pageNumber="398">Fig. 4</figureCitation>
|
||
C-F); membranous part distally narrower than proximally (
|
||
<figureCitation id="5CA209C1FFE3FFF0BF1EFD923B85FD7F" captionStart="Fig. 3" captionStartId="5.[151,186,1940,1961]" captionTargetBox="[261,1329,261,1882]" captionTargetId="figure@5.[255,1334,257,1886]" captionTargetPageId="5" captionText="Fig. 3. SEM-micrographs of embolus proper of left palpal organ of two males: Liphistius desultor from Penang Hill (A, C, E, G) and Liphistius malayanus from Fraser’s Hill (B, D, F, H). (A) Retroventral-distal view. (B) Dorsal-distal view. (C) Distal view (dorsal side to the right). (D) Distal view (prolateral side to the right). (E) Retrodorsal view. (F) Prodorsal view. (G) Proventral view. (H) Prolateral view. Abbreviations: f - major embolic fold; m - membranous part of embolus proper; s - sclerotised part of embolus proper." httpUri="https://zenodo.org/record/1021943/files/figure.png" pageId="7" pageNumber="398">Fig. 3D, F, H</figureCitation>
|
||
), its proximal portion essentially unpigmented.
|
||
</paragraph>
|
||
<paragraph id="C4261544FFE3FFF0B950FD433C90FB48" blockId="7.[831,1437,690,1196]" pageId="7" pageNumber="398">
|
||
<emphasis id="F6EDC956FFE3FFF0B950FD433CE5FD2A" bold="true" box="[831,1277,690,715]" pageId="7" pageNumber="398">Additions to description of female:</emphasis>
|
||
Vulval plate (
|
||
<figureCitation id="5CA209C1FFE3FFF0B927FD233B97FD0A" box="[840,911,722,745]" captionStart="Fig. 5" captionStartId="8.[151,186,1809,1830]" captionTargetBox="[154,1439,260,1762]" captionTargetId="figure@8.[148,1439,254,1779]" captionTargetPageId="8" captionText="Fig. 5. Liphistius malayanus, vulval plate of eight females: paratype of L. m. cameroni from Berinchang, Cameron Highlands (A- B); exuvia of female from Ringlet, Cameron Highlands (C); female (SMF 64093) from Fraser’s Hill (D); female (moulted 24. VI. 2002) from Fraser’s Hill; poreplate slightly deformed by brief immersion in cold KOH (E); female (SMF 56206) from Fraser’s Hill (F); female (SMF 60037) from Genting Highlands (G); female (moulted 8. XII. 2005) from Genting Highlands (H); exuvia of female from Templer Park (I-J). (A, C-I) Dorsal view. (B, J) Ventral view. Scale lines: 1.0 mm (A-B; C, F; D; E, G; H; I-J)." httpUri="https://zenodo.org/record/1021947/files/figure.png" pageId="7" pageNumber="398">Fig. 5</figureCitation>
|
||
) always with few to many hairs laterally in genital atrium, rarely also posteromedially (
|
||
<figureCitation id="5CA209C1FFE3FFF0BF53FD023D97FCE9" box="[1340,1423,755,778]" captionStart="Fig. 5" captionStartId="8.[151,186,1809,1830]" captionTargetBox="[154,1439,260,1762]" captionTargetId="figure@8.[148,1439,254,1779]" captionTargetPageId="8" captionText="Fig. 5. Liphistius malayanus, vulval plate of eight females: paratype of L. m. cameroni from Berinchang, Cameron Highlands (A- B); exuvia of female from Ringlet, Cameron Highlands (C); female (SMF 64093) from Fraser’s Hill (D); female (moulted 24. VI. 2002) from Fraser’s Hill; poreplate slightly deformed by brief immersion in cold KOH (E); female (SMF 56206) from Fraser’s Hill (F); female (SMF 60037) from Genting Highlands (G); female (moulted 8. XII. 2005) from Genting Highlands (H); exuvia of female from Templer Park (I-J). (A, C-I) Dorsal view. (B, J) Ventral view. Scale lines: 1.0 mm (A-B; C, F; D; E, G; H; I-J)." httpUri="https://zenodo.org/record/1021947/files/figure.png" pageId="7" pageNumber="398">
|
||
Fig.
|
||
<date id="B0273384FFE3FFF0BF19FD023D97FCE9" box="[1398,1423,755,778]" pageId="7" pageNumber="398">5I</date>
|
||
</figureCitation>
|
||
). Posterior stalk more or less distinctly trapezium-shaped (posteriorly wider than anteriorly), densely pitted in posterior portion. Poreplate wider than long, with a pair of anterolateral lobes projecting anteriad and slightly bent ventrad; several of these lobes more or less distinctly constricted at base (
|
||
<figureCitation id="5CA209C1FFE3FFF0BEFAFC423CC1FC29" box="[1173,1241,947,970]" captionStart="Fig. 5" captionStartId="8.[151,186,1809,1830]" captionTargetBox="[154,1439,260,1762]" captionTargetId="figure@8.[148,1439,254,1779]" captionTargetPageId="8" captionText="Fig. 5. Liphistius malayanus, vulval plate of eight females: paratype of L. m. cameroni from Berinchang, Cameron Highlands (A- B); exuvia of female from Ringlet, Cameron Highlands (C); female (SMF 64093) from Fraser’s Hill (D); female (moulted 24. VI. 2002) from Fraser’s Hill; poreplate slightly deformed by brief immersion in cold KOH (E); female (SMF 56206) from Fraser’s Hill (F); female (SMF 60037) from Genting Highlands (G); female (moulted 8. XII. 2005) from Genting Highlands (H); exuvia of female from Templer Park (I-J). (A, C-I) Dorsal view. (B, J) Ventral view. Scale lines: 1.0 mm (A-B; C, F; D; E, G; H; I-J)." httpUri="https://zenodo.org/record/1021947/files/figure.png" pageId="7" pageNumber="398">Fig. 5</figureCitation>
|
||
A-C, E); anterior margin of poreplate straight to procurved; lateral and posterolateral margins on ventral side of pore plate bulging (
|
||
<figureCitation id="5CA209C1FFE3FFF0B9CDFBE23C0BFBCF" box="[930,1043,1043,1068]" captionStart="Fig. 5" captionStartId="8.[151,186,1809,1830]" captionTargetBox="[154,1439,260,1762]" captionTargetId="figure@8.[148,1439,254,1779]" captionTargetPageId="8" captionText="Fig. 5. Liphistius malayanus, vulval plate of eight females: paratype of L. m. cameroni from Berinchang, Cameron Highlands (A- B); exuvia of female from Ringlet, Cameron Highlands (C); female (SMF 64093) from Fraser’s Hill (D); female (moulted 24. VI. 2002) from Fraser’s Hill; poreplate slightly deformed by brief immersion in cold KOH (E); female (SMF 56206) from Fraser’s Hill (F); female (SMF 60037) from Genting Highlands (G); female (moulted 8. XII. 2005) from Genting Highlands (H); exuvia of female from Templer Park (I-J). (A, C-I) Dorsal view. (B, J) Ventral view. Scale lines: 1.0 mm (A-B; C, F; D; E, G; H; I-J)." httpUri="https://zenodo.org/record/1021947/files/figure.png" pageId="7" pageNumber="398">Fig. 5B, J</figureCitation>
|
||
). CDO large and of variable shape, mostly longer than wide, rarely wider than long (
|
||
<figureCitation id="5CA209C1FFE3FFF0BF1EFBC23B79FB88" captionStart="Fig. 5" captionStartId="8.[151,186,1809,1830]" captionTargetBox="[154,1439,260,1762]" captionTargetId="figure@8.[148,1439,254,1779]" captionTargetPageId="8" captionText="Fig. 5. Liphistius malayanus, vulval plate of eight females: paratype of L. m. cameroni from Berinchang, Cameron Highlands (A- B); exuvia of female from Ringlet, Cameron Highlands (C); female (SMF 64093) from Fraser’s Hill (D); female (moulted 24. VI. 2002) from Fraser’s Hill; poreplate slightly deformed by brief immersion in cold KOH (E); female (SMF 56206) from Fraser’s Hill (F); female (SMF 60037) from Genting Highlands (G); female (moulted 8. XII. 2005) from Genting Highlands (H); exuvia of female from Templer Park (I-J). (A, C-I) Dorsal view. (B, J) Ventral view. Scale lines: 1.0 mm (A-B; C, F; D; E, G; H; I-J)." httpUri="https://zenodo.org/record/1021947/files/figure.png" pageId="7" pageNumber="398">Fig. 5E</figureCitation>
|
||
). Receptacular cluster large and strongly racemose, in most cases protruding beyond anterior margin of poreplate, rarely not (
|
||
<figureCitation id="5CA209C1FFE3FFF0BE49FB653C63FB48" box="[1062,1147,1172,1195]" captionStart="Fig. 5" captionStartId="8.[151,186,1809,1830]" captionTargetBox="[154,1439,260,1762]" captionTargetId="figure@8.[148,1439,254,1779]" captionTargetPageId="8" captionText="Fig. 5. Liphistius malayanus, vulval plate of eight females: paratype of L. m. cameroni from Berinchang, Cameron Highlands (A- B); exuvia of female from Ringlet, Cameron Highlands (C); female (SMF 64093) from Fraser’s Hill (D); female (moulted 24. VI. 2002) from Fraser’s Hill; poreplate slightly deformed by brief immersion in cold KOH (E); female (SMF 56206) from Fraser’s Hill (F); female (SMF 60037) from Genting Highlands (G); female (moulted 8. XII. 2005) from Genting Highlands (H); exuvia of female from Templer Park (I-J). (A, C-I) Dorsal view. (B, J) Ventral view. Scale lines: 1.0 mm (A-B; C, F; D; E, G; H; I-J)." httpUri="https://zenodo.org/record/1021947/files/figure.png" pageId="7" pageNumber="398">Fig. 5G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C4261544FFE3FFF0B950FB323C7BF7FD" blockId="7.[831,1437,1219,2078]" pageId="7" pageNumber="398">
|
||
<emphasis id="F6EDC956FFE3FFF0B950FB323C2CFB3F" bold="true" box="[831,1076,1219,1244]" pageId="7" pageNumber="398">Taxonomic remarks:</emphasis>
|
||
<taxonomicName id="03996EC7FFE3FFF0BE2FFB323D85FB38" authorityName="Haupt" authorityYear="1983" box="[1088,1437,1219,1243]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="398" phylum="Arthropoda" rank="subSpecies" species="malayanus" subSpecies="cameroni">
|
||
<emphasis id="F6EDC956FFE3FFF0BE2FFB323D85FB38" box="[1088,1437,1219,1243]" italics="true" pageId="7" pageNumber="398">Liphistius malayanus cameroni</emphasis>
|
||
</taxonomicName>
|
||
has been the only subspecies in the genus
|
||
<taxonomicName id="03996EC7FFE3FFF0BF44FB123D8DFB19" authorityName="Schiodte" authorityYear="1849" box="[1323,1429,1251,1274]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="398" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F6EDC956FFE3FFF0BF44FB123D8DFB19" box="[1323,1429,1251,1274]" italics="true" pageId="7" pageNumber="398">Liphistius</emphasis>
|
||
</taxonomicName>
|
||
. As specimens later collected near the
|
||
<typeStatus id="1B22ABE6FFE3FFF0BF62FAF53D24FAFF" box="[1293,1340,1284,1308]" pageId="7" pageNumber="398">type</typeStatus>
|
||
locality show no relevant differences from specimens of the nominal subspecies, this case became doubtful. Thus a re-examination of the
|
||
<typeStatus id="1B22ABE6FFE3FFF0BE20FA943C90FA9E" box="[1103,1160,1381,1405]" pageId="7" pageNumber="398">types</typeStatus>
|
||
of
|
||
<taxonomicName id="03996EC7FFE3FFF0BEDBFA923BBBFA7F" authorityName="Haupt" authorityYear="1983" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="398" phylum="Arthropoda" rank="subSpecies" species="malayanus" subSpecies="cameroni">
|
||
<emphasis id="F6EDC956FFE3FFF0BEDBFA923BBBFA7F" italics="true" pageId="7" pageNumber="398">Liphistius malayanus cameroni</emphasis>
|
||
</taxonomicName>
|
||
was necessary, and it revealed that they differ considerably from the illustrations and explanations given in the original description of this subspecies. The vulva of the female
|
||
<typeStatus id="1B22ABE6FFE3FFF0BE4FFA143C64FA1E" box="[1056,1148,1509,1533]" pageId="7" pageNumber="398" type="paratype">paratype</typeStatus>
|
||
was cleared of tissue, but the ventral wall of the genital region (with bristles still attached) was not removed, thus not allowing a clear view of the ventral side of the poreplate. It therefore appears that Haupt’s (1983) illustrations were made with the help of a compound microscope without using a drawing tube, which would explain why the poreplate in the original illustrations is wider than in reality and why the outlines of the posterior stalk are too rounded. The same is probably also the case in the dorsal view of the same poreplate, it being too wide, with a triangular CDO leading to the receptacular cluster and with a peculiar double posterior margin of the posterior stalk (see
|
||
<bibRefCitation id="A00868B5FFE3FFF0B91AF8773C19F87D" author="Haupt J." box="[885,1025,1926,1950]" journalOrPublisher="Zeitschrift fur zoologische Systematik und Evolutionsforschung" pageId="7" pageNumber="398" pagination="275 - 293" part="21" refString="Haupt J. 1983. Vergleichende Morphologie der Genitalorgane und Phylogenie der liphistiomorphen Webspinnen (Araneae: Mesothelae). I. Revision der bisher bekannten Arten. Zeitschrift fur zoologische Systematik und Evolutionsforschung 21: 275 - 293." title="Vergleichende Morphologie der Genitalorgane und Phylogenie der liphistiomorphen Webspinnen (Araneae: Mesothelae). I. Revision der bisher bekannten Arten" type="journal article" year="1983">Haupt, 1983</bibRefCitation>
|
||
: figs 5f, 6f). In fact the vulva of the
|
||
<typeStatus id="1B22ABE6FFE3FFF0B950F8573B83F85D" box="[831,923,1958,1982]" pageId="7" pageNumber="398" type="paratype">paratype</typeStatus>
|
||
of
|
||
<taxonomicName id="03996EC7FFE3FFF0B9AEF8563CA5F85D" authorityName="Haupt" authorityYear="1983" box="[961,1213,1957,1982]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="398" phylum="Arthropoda" rank="subSpecies" species="malayanus" subSpecies="cameroni">
|
||
<emphasis id="F6EDC956FFE3FFF0B9AEF8563CA5F85D" box="[961,1213,1957,1982]" italics="true" pageId="7" pageNumber="398">L. malayanus cameroni</emphasis>
|
||
</taxonomicName>
|
||
does not differ from the vulvae of the other
|
||
<taxonomicName id="03996EC7FFE3FFF0BE50F8363CC9F83F" authorityName="Abraham" authorityYear="1923" box="[1087,1233,1989,2014]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="398" phylum="Arthropoda" rank="species" species="malayanus">
|
||
<emphasis id="F6EDC956FFE3FFF0BE50F8363CC9F83F" box="[1087,1233,1989,2014]" italics="true" pageId="7" pageNumber="398">L. malayanus</emphasis>
|
||
</taxonomicName>
|
||
females examined (
|
||
<figureCitation id="5CA209C1FFE3FFF0B928F8143B9EF81F" box="[839,902,2021,2044]" captionStart="Fig. 5" captionStartId="8.[151,186,1809,1830]" captionTargetBox="[154,1439,260,1762]" captionTargetId="figure@8.[148,1439,254,1779]" captionTargetPageId="8" captionText="Fig. 5. Liphistius malayanus, vulval plate of eight females: paratype of L. m. cameroni from Berinchang, Cameron Highlands (A- B); exuvia of female from Ringlet, Cameron Highlands (C); female (SMF 64093) from Fraser’s Hill (D); female (moulted 24. VI. 2002) from Fraser’s Hill; poreplate slightly deformed by brief immersion in cold KOH (E); female (SMF 56206) from Fraser’s Hill (F); female (SMF 60037) from Genting Highlands (G); female (moulted 8. XII. 2005) from Genting Highlands (H); exuvia of female from Templer Park (I-J). (A, C-I) Dorsal view. (B, J) Ventral view. Scale lines: 1.0 mm (A-B; C, F; D; E, G; H; I-J)." httpUri="https://zenodo.org/record/1021947/files/figure.png" pageId="7" pageNumber="398">Fig. 5</figureCitation>
|
||
A-B cf.
|
||
<figureCitation id="5CA209C1FFE3FFF0B9B0F8143C05F81F" box="[991,1053,2021,2044]" captionStart="Fig. 5" captionStartId="8.[151,186,1809,1830]" captionTargetBox="[154,1439,260,1762]" captionTargetId="figure@8.[148,1439,254,1779]" captionTargetPageId="8" captionText="Fig. 5. Liphistius malayanus, vulval plate of eight females: paratype of L. m. cameroni from Berinchang, Cameron Highlands (A- B); exuvia of female from Ringlet, Cameron Highlands (C); female (SMF 64093) from Fraser’s Hill (D); female (moulted 24. VI. 2002) from Fraser’s Hill; poreplate slightly deformed by brief immersion in cold KOH (E); female (SMF 56206) from Fraser’s Hill (F); female (SMF 60037) from Genting Highlands (G); female (moulted 8. XII. 2005) from Genting Highlands (H); exuvia of female from Templer Park (I-J). (A, C-I) Dorsal view. (B, J) Ventral view. Scale lines: 1.0 mm (A-B; C, F; D; E, G; H; I-J)." httpUri="https://zenodo.org/record/1021947/files/figure.png" pageId="7" pageNumber="398">Fig. 5</figureCitation>
|
||
C-J) in any way that would warrant a distinct taxonomic status.
|
||
</paragraph>
|
||
<caption id="90E645CCFFECFFFFBAF8F8E03931F854" httpUri="https://zenodo.org/record/1021947/files/figure.png" pageId="8" pageNumber="399" targetBox="[154,1439,260,1762]" targetPageId="8">
|
||
<paragraph id="C4261544FFECFFFFBAF8F8E03931F854" blockId="8.[151,1437,1809,1975]" pageId="8" pageNumber="399">
|
||
Fig. 5.
|
||
<taxonomicName id="03996EC7FFECFFFFBA8CF8E039AAF8C5" authorityName="Abraham" authorityYear="1923" box="[227,434,1809,1830]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="399" phylum="Arthropoda" rank="species" species="malayanus">
|
||
<emphasis id="F6EDC956FFECFFFFBA8CF8E039AAF8C5" box="[227,434,1809,1830]" italics="true" pageId="8" pageNumber="399">Liphistius malayanus</emphasis>
|
||
</taxonomicName>
|
||
, vulval plate of eight females: paratype of
|
||
<taxonomicName id="03996EC7FFECFFFFB930F8E23BECF8CB" authorityName="Haupt" authorityYear="1983" box="[863,1012,1811,1832]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="399" phylum="Arthropoda" rank="subSpecies" species="malayanus" subSpecies="cameroni">
|
||
<emphasis id="F6EDC956FFECFFFFB930F8E23BECF8CB" box="[863,1012,1811,1832]" italics="true" pageId="8" pageNumber="399">L. m. cameroni</emphasis>
|
||
</taxonomicName>
|
||
from Berinchang, Cameron Highlands (A- B); exuvia of female from Ringlet, Cameron Highlands (C); female (SMF 64093) from Fraser’s Hill (D); female (moulted 24.VI.2002) from Fraser’s Hill; poreplate slightly deformed by brief immersion in cold KOH (E); female (SMF 56206) from Fraser’s Hill (F); female (SMF 60037) from Genting Highlands (G); female (moulted 8.XII.2005) from Genting Highlands (H); exuvia of female from Templer Park (I-J). (A, C-I) Dorsal view. (B, J) Ventral view. Scale lines: 1.0 mm (A-B; C, F; D; E, G; H; I-J).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C4261544FFEDFFFEBAF8FEF33A47FAF1" blockId="9.[151,757,258,1298]" pageId="9" pageNumber="400">
|
||
The illustrated differences between the male
|
||
<typeStatus id="1B22ABE6FFEDFFFEB8F8FEF33AECFEFA" box="[663,756,258,281]" pageId="9" pageNumber="400" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="03996EC7FFEDFFFEBAD3FED239DFFED9" authorityName="Haupt" authorityYear="1983" box="[188,455,290,314]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="400" phylum="Arthropoda" rank="subSpecies" species="malayanus" subSpecies="cameroni">
|
||
<emphasis id="F6EDC956FFEDFFFEBAD3FED239DFFED9" box="[188,455,290,314]" italics="true" pageId="9" pageNumber="400">L. malayanus cameroni</emphasis>
|
||
</taxonomicName>
|
||
and the male that Haupt incorrectly referred to as a
|
||
<typeStatus id="1B22ABE6FFEDFFFEBBBBFEB23A28FEB8" box="[468,560,323,347]" pageId="9" pageNumber="400" type="paratype">paratype</typeStatus>
|
||
of
|
||
<taxonomicName id="03996EC7FFEDFFFEB831FEB23AECFEBB" authorityName="Abraham" authorityYear="1923" box="[606,756,321,346]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="400" phylum="Arthropoda" rank="species" species="malayanus">
|
||
<emphasis id="F6EDC956FFEDFFFEB831FEB23AECFEBB" box="[606,756,321,346]" italics="true" pageId="9" pageNumber="400">L. malayanus</emphasis>
|
||
</taxonomicName>
|
||
(in BMNH, not examined) are mostly due to a slightly different view of the palp and due to the fact that the palpal organ of the BMNH male is expanded (turned clockwise by about 90°) (
|
||
<bibRefCitation id="A00868B5FFEDFFFEBBDAFE30390EFE1A" author="Haupt J." journalOrPublisher="Zeitschrift fur zoologische Systematik und Evolutionsforschung" pageId="9" pageNumber="400" pagination="275 - 293" part="21" refString="Haupt J. 1983. Vergleichende Morphologie der Genitalorgane und Phylogenie der liphistiomorphen Webspinnen (Araneae: Mesothelae). I. Revision der bisher bekannten Arten. Zeitschrift fur zoologische Systematik und Evolutionsforschung 21: 275 - 293." title="Vergleichende Morphologie der Genitalorgane und Phylogenie der liphistiomorphen Webspinnen (Araneae: Mesothelae). I. Revision der bisher bekannten Arten" type="journal article" year="1983">Haupt, 1983: figs 3D, 4C cf. figs 3E, 4D</bibRefCitation>
|
||
). None of the
|
||
<taxonomicName id="03996EC7FFEDFFFEBBC0FE133A58FE14" authorityName="Abraham" authorityYear="1923" box="[431,576,480,505]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="400" phylum="Arthropoda" rank="species" species="malayanus">
|
||
<emphasis id="F6EDC956FFEDFFFEBBC0FE133A58FE14" box="[431,576,480,505]" italics="true" pageId="9" pageNumber="400">L. malayanus</emphasis>
|
||
</taxonomicName>
|
||
males examined has the tibial apophysis of the male palp situated as far distally as illustrated by
|
||
<bibRefCitation id="A00868B5FFEDFFFEBBA3FDD03AF7FDDA" author="Haupt J." box="[460,751,544,569]" journalOrPublisher="Zeitschrift fur zoologische Systematik und Evolutionsforschung" pageId="9" pageNumber="400" pagination="275 - 293" part="21" refString="Haupt J. 1983. Vergleichende Morphologie der Genitalorgane und Phylogenie der liphistiomorphen Webspinnen (Araneae: Mesothelae). I. Revision der bisher bekannten Arten. Zeitschrift fur zoologische Systematik und Evolutionsforschung 21: 275 - 293." title="Vergleichende Morphologie der Genitalorgane und Phylogenie der liphistiomorphen Webspinnen (Araneae: Mesothelae). I. Revision der bisher bekannten Arten" type="journal article" year="1983">Haupt (1983: figs 3D, 4C)</bibRefCitation>
|
||
, therefore this may also be due to a different view. The embolus proper of the left palp of the
|
||
<taxonomicName id="03996EC7FFEDFFFEB824FD903AECFD9B" authorityName="Haupt" authorityYear="1983" box="[587,756,609,632]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="400" phylum="Arthropoda" rank="subSpecies" species="malayanus" subSpecies="cameroni">
|
||
<emphasis id="F6EDC956FFEDFFFEB824FD903AECFD9B" box="[587,756,609,632]" italics="true" pageId="9" pageNumber="400">L. m. cameroni</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="1B22ABE6FFEDFFFEBAF8FD8E38ECFD75" box="[151,244,639,662]" pageId="9" pageNumber="400" type="holotype">holotype</typeStatus>
|
||
(that of the right palp is missing, together with tegulum and contrategulum) is slightly narrower than that of the second male from the Cameron
|
||
<collectingRegion id="065DDBA6FFEDFFFEB835FD4E3AD0FD35" box="[602,712,703,726]" country="Czech Republic" name="Vysocina" pageId="9" pageNumber="400">Highlands</collectingRegion>
|
||
and that of other conspecific males examined. This difference is presumably connected to size: the
|
||
<taxonomicName id="03996EC7FFEDFFFEB829FCF13AECFCF4" authorityName="Haupt" authorityYear="1983" box="[582,756,768,791]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="400" phylum="Arthropoda" rank="subSpecies" species="malayanus" subSpecies="cameroni">
|
||
<emphasis id="F6EDC956FFEDFFFEB829FCF13AECFCF4" box="[582,756,768,791]" italics="true" pageId="9" pageNumber="400">L. m. cameroni</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="1B22ABE6FFEDFFFEBAF8FCEF38ECFCD6" box="[151,244,798,821]" pageId="9" pageNumber="400" type="holotype">holotype</typeStatus>
|
||
is the smallest among the
|
||
<taxonomicName id="03996EC7FFEDFFFEB874FCD13AB6FCD6" authorityName="Abraham" authorityYear="1923" box="[539,686,798,823]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="400" phylum="Arthropoda" rank="species" species="malayanus">
|
||
<emphasis id="F6EDC956FFEDFFFEB874FCD13AB6FCD6" box="[539,686,798,823]" italics="true" pageId="9" pageNumber="400">L. malayanus</emphasis>
|
||
</taxonomicName>
|
||
males examined. The character in which the
|
||
<typeStatus id="1B22ABE6FFEDFFFEB857FCCF3A8DFCB6" box="[568,661,830,853]" pageId="9" pageNumber="400" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="03996EC7FFEDFFFEB8D3FCCE38E3FC95" authorityName="Haupt" authorityYear="1983" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="400" phylum="Arthropoda" rank="subSpecies" species="malayanus" subSpecies="cameroni">
|
||
<emphasis id="F6EDC956FFEDFFFEB8D3FCCE38E3FC95" italics="true" pageId="9" pageNumber="400">L. m. cameroni</emphasis>
|
||
</taxonomicName>
|
||
most visibly differs from other conspecific males is a slightly elevated para-embolic plate (
|
||
<figureCitation id="5CA209C1FFEDFFFEB8FAFC8F3AFFFC75" box="[661,743,894,918]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="9" pageNumber="400">Fig. 4E</figureCitation>
|
||
), which is also present in the second male from the Cameron
|
||
<collectingRegion id="065DDBA6FFEDFFFEBB6DFC4C3968FC37" box="[258,368,957,980]" country="Czech Republic" name="Vysocina" pageId="9" pageNumber="400">Highlands</collectingRegion>
|
||
(
|
||
<figureCitation id="5CA209C1FFEDFFFEBBEEFC4C39CCFC35" box="[385,468,957,982]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="9" pageNumber="400">Fig. 4F</figureCitation>
|
||
). However, an indistinctly elevated para-embolic plate is present in one of the two males from the Fraser’s Hill, whereas all other conspecific males possess only a distinct edge at this place (
|
||
<figureCitation id="5CA209C1FFEDFFFEB8FCFBEC3AF1FBD6" box="[659,745,1053,1077]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="9" pageNumber="400">Fig. 4D</figureCitation>
|
||
). Thus the
|
||
<typeStatus id="1B22ABE6FFEDFFFEBA90FBCD3944FBB0" box="[255,348,1084,1107]" pageId="9" pageNumber="400" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="03996EC7FFEDFFFEBBEBFBCF3A32FBB6" authorityName="Haupt" authorityYear="1983" box="[388,554,1086,1109]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="400" phylum="Arthropoda" rank="subSpecies" species="malayanus" subSpecies="cameroni">
|
||
<emphasis id="F6EDC956FFEDFFFEBBEBFBCF3A32FBB6" box="[388,554,1086,1109]" italics="true" pageId="9" pageNumber="400">L. m. cameroni</emphasis>
|
||
</taxonomicName>
|
||
differs from other conspecific males examined in possessing the most extreme states of three quite variable characters, but it is not representative for the population in the Cameron
|
||
<collectingRegion id="065DDBA6FFEDFFFEBAF8FB4A3910FB31" box="[151,264,1211,1234]" country="Czech Republic" name="Vysocina" pageId="9" pageNumber="400">Highlands</collectingRegion>
|
||
. I consequently place
|
||
<taxonomicName id="03996EC7FFEDFFFEB864FB4A38E3FB17" authorityName="Haupt" authorityYear="1983" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="400" phylum="Arthropoda" rank="subSpecies" species="malayanus" subSpecies="cameroni">
|
||
<emphasis id="F6EDC956FFEDFFFEB864FB4A38E3FB17" italics="true" pageId="9" pageNumber="400">Liphistius malayanus cameroni</emphasis>
|
||
</taxonomicName>
|
||
in the synonymy of
|
||
<taxonomicName id="03996EC7FFEDFFFEB868FB2A3AECFB11" authorityName="Abraham" authorityYear="1923" box="[519,756,1243,1266]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="400" phylum="Arthropoda" rank="species" species="malayanus">
|
||
<emphasis id="F6EDC956FFEDFFFEB868FB2A3AECFB11" box="[519,756,1243,1266]" italics="true" pageId="9" pageNumber="400">Liphistius malayanus</emphasis>
|
||
</taxonomicName>
|
||
which makes the species monotypic again.
|
||
</paragraph>
|
||
<paragraph id="C4261544FFEDFFFEBAF8FADB3A3FF903" blockId="9.[151,757,1322,2077]" pageId="9" pageNumber="400">
|
||
<emphasis id="F6EDC956FFEDFFFEBAF8FADB3914FAA2" bold="true" box="[151,268,1322,1345]" pageId="9" pageNumber="400">Variation:</emphasis>
|
||
Carapace lengths in males (n=6; including the
|
||
<typeStatus id="1B22ABE6FFEDFFFEBAABFABB393AFA82" box="[196,290,1354,1377]" pageId="9" pageNumber="400" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="03996EC7FFEDFFFEBB3FFABA39E4FA81" authorityName="Haupt" authorityYear="1983" box="[336,508,1355,1378]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="400" phylum="Arthropoda" rank="subSpecies" species="malayanus" subSpecies="cameroni">
|
||
<emphasis id="F6EDC956FFEDFFFEBB3FFABA39E4FA81" box="[336,508,1355,1378]" italics="true" pageId="9" pageNumber="400">L. m. cameroni</emphasis>
|
||
</taxonomicName>
|
||
) 7.90-11.23, carapace widths 7.04-9.69; in the largest female from each locality (n=4; not including the
|
||
<typeStatus id="1B22ABE6FFEDFFFEB870FA7B3A64FA41" box="[543,636,1418,1442]" pageId="9" pageNumber="400" type="paratype">paratype</typeStatus>
|
||
of
|
||
<taxonomicName id="03996EC7FFEDFFFEB8DBFA7A38E4FA22" authorityName="Haupt" authorityYear="1983" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="400" phylum="Arthropoda" rank="subSpecies" species="malayanus" subSpecies="cameroni">
|
||
<emphasis id="F6EDC956FFEDFFFEB8DBFA7A38E4FA22" italics="true" pageId="9" pageNumber="400">L. m. cameroni</emphasis>
|
||
</taxonomicName>
|
||
) 10.86-16.67 and 10.00-14.07, respectively. The
|
||
<typeStatus id="1B22ABE6FFEDFFFEBAA2FA383933FA03" box="[205,299,1481,1504]" pageId="9" pageNumber="400" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="03996EC7FFEDFFFEBB36FA3B3A1EFA02" authorityName="Haupt" authorityYear="1983" box="[345,518,1482,1505]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="400" phylum="Arthropoda" rank="subSpecies" species="malayanus" subSpecies="cameroni">
|
||
<emphasis id="F6EDC956FFEDFFFEBB36FA3B3A1EFA02" box="[345,518,1482,1505]" italics="true" pageId="9" pageNumber="400">L. m. cameroni</emphasis>
|
||
</taxonomicName>
|
||
is the smallest male examined, with carapace length 7.90, width 7.04; in the second male from the Cameron
|
||
<collectingRegion id="065DDBA6FFEDFFFEB871F9F93A94F9FC" box="[542,652,1544,1567]" country="Czech Republic" name="Vysocina" pageId="9" pageNumber="400">Highlands</collectingRegion>
|
||
it is 9.63 and 8.40, respectively. The largest female is from the Templer Park, measuring 16.67 and 14.07; the second largest female is from the Fraser’s Hill, measuring 16.54 and 14.07. The female
|
||
<typeStatus id="1B22ABE6FFEDFFFEBBBDF9783A36F942" box="[466,558,1673,1697]" pageId="9" pageNumber="400" type="paratype">paratype</typeStatus>
|
||
of
|
||
<taxonomicName id="03996EC7FFEDFFFEB83DF9783AECF943" authorityName="Haupt" authorityYear="1983" box="[594,756,1673,1696]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="400" phylum="Arthropoda" rank="subSpecies" species="malayanus" subSpecies="cameroni">
|
||
<emphasis id="F6EDC956FFEDFFFEB83DF9783AECF943" box="[594,756,1673,1696]" italics="true" pageId="9" pageNumber="400">L. m. cameroni</emphasis>
|
||
</taxonomicName>
|
||
measures 9.26 and 7.53, the second female from the Cameron
|
||
<collectingRegion id="065DDBA6FFEDFFFEBB6FF9363976F93D" box="[256,366,1735,1758]" country="Czech Republic" name="Vysocina" pageId="9" pageNumber="400">Highlands</collectingRegion>
|
||
10.86 and 10.12.
|
||
</paragraph>
|
||
<paragraph id="C4261544FFEDFFFEBAF8F91639F6F7FE" blockId="9.[151,757,1322,2077]" pageId="9" pageNumber="400">
|
||
The apical megaspines on the tibial apophysis are quite variable in length, but not as thin as illustrated in
|
||
<bibRefCitation id="A00868B5FFEDFFFEB8F1F8F63A52F8DC" author="Murphy J. A. & Platnick N. I." journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="9" pageNumber="400" pagination="46 - 56" part="170" refString="Murphy J. A., Platnick N. I. 1981. On Liphistius desultor Schiodte (Araneae, Liphistiidae). Bulletin of the American Museum of Natural History 170: 46 - 56." title="On Liphistius desultor Schiodte (Araneae, Liphistiidae)" type="journal article" year="1981">Murphy & Platnick (1981: figs 8, 11, 14, 17, 20)</bibRefCitation>
|
||
and in
|
||
<bibRefCitation id="A00868B5FFEDFFFEB8F2F8D639FCF8BE" author="Platnick N. & Sedgwick W. C." journalOrPublisher="American Museum Novitates" pageId="9" pageNumber="400" pagination="1 - 31" part="2781" refString="Platnick N. I., Sedgwick W. C. 1984. A revision of the spider genus Liphistius (Araneae, Mesothelae). American Museum Novitates 2781: 1 - 31." title="A revision of the spider genus Liphistius (Araneae, Mesothelae)" type="journal article" year="1984">Platnick & Sedgwick (1984: figs 63-67)</bibRefCitation>
|
||
; the ventral-most of them is mostly bent distad and has a thin, long apex (
|
||
<figureCitation id="5CA209C1FFEDFFFEB8A6F89738BDF87C" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="9" pageNumber="400">Fig. 4</figureCitation>
|
||
A-B). The ventral megaspine is bent in the male from the Templer Park (
|
||
<figureCitation id="5CA209C1FFEDFFFEBB0CF85739A4F85D" box="[355,444,1958,1982]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="9" pageNumber="400">Fig. 4A</figureCitation>
|
||
), whereas in the
|
||
<typeStatus id="1B22ABE6FFEDFFFEB817F8573ACDF85E" box="[632,725,1958,1981]" pageId="9" pageNumber="400" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="03996EC7FFEDFFFEBAF8F8363958F83D" authorityName="Haupt" authorityYear="1983" box="[151,320,1991,2014]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="400" phylum="Arthropoda" rank="subSpecies" species="malayanus" subSpecies="cameroni">
|
||
<emphasis id="F6EDC956FFEDFFFEBAF8F8363958F83D" box="[151,320,1991,2014]" italics="true" pageId="9" pageNumber="400">L. m. cameroni</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="5CA209C1FFEDFFFEBB3DF83439B1F83D" box="[338,425,1989,2014]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="9" pageNumber="400">Fig. 4B</figureCitation>
|
||
), in a male from the Genting
|
||
<collectingRegion id="065DDBA6FFEDFFFEBAF8F814391FF81F" box="[151,263,2021,2044]" country="Czech Republic" name="Vysocina" pageId="9" pageNumber="400">Highlands</collectingRegion>
|
||
, and also in a second male from the Cameron
|
||
<collectingRegion id="065DDBA6FFEDFFFEBAF8F7F7391DF7FE" box="[151,261,2054,2077]" country="Czech Republic" name="Vysocina" pageId="9" pageNumber="400">Highlands</collectingRegion>
|
||
it is slightly sigmoid.
|
||
</paragraph>
|
||
<paragraph id="C4261544FFEDFFFEB950FEF33D59FE58" blockId="9.[831,1437,258,1836]" pageId="9" pageNumber="400">
|
||
The distal edge of the contrategulum carries one or two slightly enlarged dorsal denticles at some distance from the projecting dorsal apex (
|
||
<figureCitation id="5CA209C1FFEDFFFEBE02FEB23CDDFEB8" box="[1133,1221,323,347]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="9" pageNumber="400">Fig. 4G</figureCitation>
|
||
, I-L); in two males there is a sharp edge at the same place (
|
||
<figureCitation id="5CA209C1FFEDFFFEBEB1FE923D2AFE9F" box="[1246,1330,355,380]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="9" pageNumber="400">Fig. 4H</figureCitation>
|
||
), but only on one palp. These larger denticles are followed by a variable number of smaller proventral denticles.
|
||
</paragraph>
|
||
<paragraph id="C4261544FFEDFFFEB950FE353C72FBE1" blockId="9.[831,1437,258,1836]" pageId="9" pageNumber="400">
|
||
The para-embolic plate is always low, most visibly elevated (but not with a sharp edge) in the
|
||
<typeStatus id="1B22ABE6FFEDFFFEBF50FE143D84FE1F" box="[1343,1436,485,508]" pageId="9" pageNumber="400" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="03996EC7FFEDFFFEB90CFDF73C08FDFE" authorityName="Haupt" authorityYear="1983" box="[867,1040,518,541]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="400" phylum="Arthropoda" rank="subSpecies" species="malayanus" subSpecies="cameroni">
|
||
<emphasis id="F6EDC956FFEDFFFEB90CFDF73C08FDFE" box="[867,1040,518,541]" italics="true" pageId="9" pageNumber="400">L. m. cameroni</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="5CA209C1FFEDFFFEBE4BFDF43C65FDFD" box="[1060,1149,517,542]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="9" pageNumber="400">Fig. 4E</figureCitation>
|
||
), in a second male from the Cameron
|
||
<collectingRegion id="065DDBA6FFEDFFFEB9B7FDD43C5EFDDF" box="[984,1094,549,572]" country="Czech Republic" name="Vysocina" pageId="9" pageNumber="400">Highlands</collectingRegion>
|
||
(
|
||
<figureCitation id="5CA209C1FFEDFFFEBE36FDD43CB5FDDD" box="[1113,1197,549,574]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="9" pageNumber="400">Fig. 4F</figureCitation>
|
||
) and in a male from the Fraser’s Hill. In the second male from the Fraser’s Hill it is not elevated; the remaining males examined are intermediate (
|
||
<figureCitation id="5CA209C1FFEDFFFEBE6BFD773C47FD7C" box="[1028,1119,646,671]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="9" pageNumber="400">Fig. 4D</figureCitation>
|
||
). An indistinct second edge, situated below the indistinct para-embolic plate, is found in the only available male from the Genting
|
||
<collectingRegion id="065DDBA6FFEDFFFEBF40FD363D85FD3D" box="[1327,1437,711,734]" country="Czech Republic" name="Vysocina" pageId="9" pageNumber="400">Highlands</collectingRegion>
|
||
and in the only available male from the Templer Park (
|
||
<figureCitation id="5CA209C1FFEDFFFEB928FCF93B85FCC2" box="[839,925,776,801]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="9" pageNumber="400">Fig. 4D</figureCitation>
|
||
), but not in other males examined (
|
||
<figureCitation id="5CA209C1FFEDFFFEBF79FCF93D4DFCC2" box="[1302,1365,776,801]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="9" pageNumber="400">Fig. 4</figureCitation>
|
||
E-F). The size of the embolus proper varies from relatively thick in the male from the Templer Park (
|
||
<figureCitation id="5CA209C1FFEDFFFEBF72FCB83D6DFC82" box="[1309,1397,841,865]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="9" pageNumber="400">Fig. 4D</figureCitation>
|
||
) to distinctly more slender in the
|
||
<typeStatus id="1B22ABE6FFEDFFFEBE13FC983CC1FC63" box="[1148,1241,873,896]" pageId="9" pageNumber="400" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="03996EC7FFEDFFFEBE93FC9B3D84FC62" authorityName="Haupt" authorityYear="1983" box="[1276,1436,874,897]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="400" phylum="Arthropoda" rank="subSpecies" species="malayanus" subSpecies="cameroni">
|
||
<emphasis id="F6EDC956FFEDFFFEBE93FC9B3D84FC62" box="[1276,1436,874,897]" italics="true" pageId="9" pageNumber="400">L. m. cameroni</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="5CA209C1FFEDFFFEB928FC783B83FC41" box="[839,923,905,930]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="9" pageNumber="400">Fig. 4E</figureCitation>
|
||
); the other males are intermediate (
|
||
<figureCitation id="5CA209C1FFEDFFFEBF7FFC783D79FC41" box="[1296,1377,905,930]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="9" pageNumber="400">Fig. 4F</figureCitation>
|
||
). The longitudinal ribs on the sclerotised part of the embolus proper vary from indistinct to distinct (
|
||
<figureCitation id="5CA209C1FFEDFFFEBE8FFC3B3D97FC00" box="[1248,1423,970,995]" captionStart="Fig. 3" captionStartId="5.[151,186,1940,1961]" captionTargetBox="[261,1329,261,1882]" captionTargetId="figure@5.[255,1334,257,1886]" captionTargetPageId="5" captionText="Fig. 3. SEM-micrographs of embolus proper of left palpal organ of two males: Liphistius desultor from Penang Hill (A, C, E, G) and Liphistius malayanus from Fraser’s Hill (B, D, F, H). (A) Retroventral-distal view. (B) Dorsal-distal view. (C) Distal view (dorsal side to the right). (D) Distal view (prolateral side to the right). (E) Retrodorsal view. (F) Prodorsal view. (G) Proventral view. (H) Prolateral view. Abbreviations: f - major embolic fold; m - membranous part of embolus proper; s - sclerotised part of embolus proper." httpUri="https://zenodo.org/record/1021943/files/figure.png" pageId="9" pageNumber="400">Fig. 3B, D, F, H</figureCitation>
|
||
), so do the denticles on them.
|
||
</paragraph>
|
||
<paragraph id="C4261544FFEDFFFEB950FBFA3D85F8C8" blockId="9.[831,1437,258,1836]" pageId="9" pageNumber="400">
|
||
There is considerable variation in the extent of the tarsal scopulae in males. The scopula on tarsus I is always thin and undivided, covering two-thirds of the ventral side in most males, only one half in the male from the Genting
|
||
<collectingRegion id="065DDBA6FFEDFFFEB9F0FB7D3C08FB40" box="[927,1040,1164,1187]" country="Czech Republic" name="Vysocina" pageId="9" pageNumber="400">Highlands</collectingRegion>
|
||
. The scopula on tarsus II is slightly denser than on tarsus I, undivided and covering twothirds of the ventral side in all males. Tarsal scopula III is like tarsal scopula II in most males, in the non-type male from the Cameron
|
||
<collectingRegion id="065DDBA6FFEDFFFEBE0FFAFF3CD6FAC6" box="[1120,1230,1294,1317]" country="Czech Republic" name="Vysocina" pageId="9" pageNumber="400">Highlands</collectingRegion>
|
||
it is light. Tarsal scopula IV is dense in all males except for the non-type male from the Cameron
|
||
<collectingRegion id="065DDBA6FFEDFFFEBE23FABE3CA2FA85" box="[1100,1210,1359,1382]" country="Czech Republic" name="Vysocina" pageId="9" pageNumber="400">Highlands</collectingRegion>
|
||
(light), undivided in all males except for the same male from the Cameron
|
||
<collectingRegion id="065DDBA6FFEDFFFEB950FA7E3BB5FA45" box="[831,941,1423,1446]" country="Czech Republic" name="Vysocina" pageId="9" pageNumber="400">Highlands</collectingRegion>
|
||
(distinctly divided) and for the male from the Genting
|
||
<collectingRegion id="065DDBA6FFEDFFFEB9A1FA413C24FA24" box="[974,1084,1456,1479]" country="Czech Republic" name="Vysocina" pageId="9" pageNumber="400">Highlands</collectingRegion>
|
||
(indistinctly divided), covering two-thirds of the ventral surface in most males except for a male from the Cameron
|
||
<collectingRegion id="065DDBA6FFEDFFFEBEE7FA013CEEF9E4" box="[1160,1270,1520,1543]" country="Czech Republic" name="Vysocina" pageId="9" pageNumber="400">Highlands</collectingRegion>
|
||
(covering only half of the surface) and the male from the Templer Park (covering three-quarters). The male
|
||
<typeStatus id="1B22ABE6FFEDFFFEBF74F9C03D60F9AB" box="[1307,1400,1585,1608]" pageId="9" pageNumber="400" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="03996EC7FFEDFFFEB950F9A23C5FF989" authorityName="Haupt" authorityYear="1983" box="[831,1095,1618,1642]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="400" phylum="Arthropoda" rank="subSpecies" species="malayanus" subSpecies="cameroni">
|
||
<emphasis id="F6EDC956FFEDFFFEB950F9A23C5FF989" box="[831,1095,1618,1642]" italics="true" pageId="9" pageNumber="400">L. malayanus cameroni</emphasis>
|
||
</taxonomicName>
|
||
had already been returned to its depository when the tarsal scopulae were examined. Even if both males from the Cameron
|
||
<collectingRegion id="065DDBA6FFEDFFFEBEBAF9633D5BF94A" box="[1237,1347,1682,1705]" country="Czech Republic" name="Vysocina" pageId="9" pageNumber="400">Highlands</collectingRegion>
|
||
differed from those of other localities in shape and extent of their tarsal scopulae (especially on tarsus IV), this would not warrant a subspecific distinction. The tarsal scopula in this species is too variable to be of high taxonomic value.
|
||
</paragraph>
|
||
<paragraph id="C4261544FFEDFFFDB950F8B238E5FD9D" blockId="9.[831,1437,1859,2078]" lastBlockId="10.[151,757,258,638]" lastPageId="10" lastPageNumber="401" pageId="9" pageNumber="400">
|
||
The vulval plates of large females have more lateral hairs in the genital atrium than those of small females (
|
||
<figureCitation id="5CA209C1FFEDFFFEB927F8753B8DF878" box="[840,917,1924,1947]" captionStart="Fig. 5" captionStartId="8.[151,186,1809,1830]" captionTargetBox="[154,1439,260,1762]" captionTargetId="figure@8.[148,1439,254,1779]" captionTargetPageId="8" captionText="Fig. 5. Liphistius malayanus, vulval plate of eight females: paratype of L. m. cameroni from Berinchang, Cameron Highlands (A- B); exuvia of female from Ringlet, Cameron Highlands (C); female (SMF 64093) from Fraser’s Hill (D); female (moulted 24. VI. 2002) from Fraser’s Hill; poreplate slightly deformed by brief immersion in cold KOH (E); female (SMF 56206) from Fraser’s Hill (F); female (SMF 60037) from Genting Highlands (G); female (moulted 8. XII. 2005) from Genting Highlands (H); exuvia of female from Templer Park (I-J). (A, C-I) Dorsal view. (B, J) Ventral view. Scale lines: 1.0 mm (A-B; C, F; D; E, G; H; I-J)." httpUri="https://zenodo.org/record/1021947/files/figure.png" pageId="9" pageNumber="400">
|
||
Fig.
|
||
<date id="B0273384FFEDFFFEB910F8753B8DF878" box="[895,917,1924,1947]" pageId="9" pageNumber="400">5I</date>
|
||
</figureCitation>
|
||
cf.
|
||
<figureCitation id="5CA209C1FFEDFFFEB9A6F8753C3CF878" box="[969,1060,1924,1947]" captionStart="Fig. 5" captionStartId="8.[151,186,1809,1830]" captionTargetBox="[154,1439,260,1762]" captionTargetId="figure@8.[148,1439,254,1779]" captionTargetPageId="8" captionText="Fig. 5. Liphistius malayanus, vulval plate of eight females: paratype of L. m. cameroni from Berinchang, Cameron Highlands (A- B); exuvia of female from Ringlet, Cameron Highlands (C); female (SMF 64093) from Fraser’s Hill (D); female (moulted 24. VI. 2002) from Fraser’s Hill; poreplate slightly deformed by brief immersion in cold KOH (E); female (SMF 56206) from Fraser’s Hill (F); female (SMF 60037) from Genting Highlands (G); female (moulted 8. XII. 2005) from Genting Highlands (H); exuvia of female from Templer Park (I-J). (A, C-I) Dorsal view. (B, J) Ventral view. Scale lines: 1.0 mm (A-B; C, F; D; E, G; H; I-J)." httpUri="https://zenodo.org/record/1021947/files/figure.png" pageId="9" pageNumber="400">Fig. 5H</figureCitation>
|
||
). In the largest female (from the Templer Park) hairs are also present posteromedially in the genital atrium (
|
||
<figureCitation id="5CA209C1FFEDFFFEBE7BF8343C7AF83F" box="[1044,1122,1989,2012]" captionStart="Fig. 5" captionStartId="8.[151,186,1809,1830]" captionTargetBox="[154,1439,260,1762]" captionTargetId="figure@8.[148,1439,254,1779]" captionTargetPageId="8" captionText="Fig. 5. Liphistius malayanus, vulval plate of eight females: paratype of L. m. cameroni from Berinchang, Cameron Highlands (A- B); exuvia of female from Ringlet, Cameron Highlands (C); female (SMF 64093) from Fraser’s Hill (D); female (moulted 24. VI. 2002) from Fraser’s Hill; poreplate slightly deformed by brief immersion in cold KOH (E); female (SMF 56206) from Fraser’s Hill (F); female (SMF 60037) from Genting Highlands (G); female (moulted 8. XII. 2005) from Genting Highlands (H); exuvia of female from Templer Park (I-J). (A, C-I) Dorsal view. (B, J) Ventral view. Scale lines: 1.0 mm (A-B; C, F; D; E, G; H; I-J)." httpUri="https://zenodo.org/record/1021947/files/figure.png" pageId="9" pageNumber="400">
|
||
Fig.
|
||
<date id="B0273384FFEDFFFEBE26F8343C7AF83F" box="[1097,1122,1989,2012]" pageId="9" pageNumber="400">5I</date>
|
||
</figureCitation>
|
||
). The shape of the posterior stalk is quite variable but mostly trapezium-shaped (
|
||
<figureCitation id="5CA209C1FFEDFFFEB927F7F73B89F7FE" box="[840,913,2054,2077]" captionStart="Fig. 5" captionStartId="8.[151,186,1809,1830]" captionTargetBox="[154,1439,260,1762]" captionTargetId="figure@8.[148,1439,254,1779]" captionTargetPageId="8" captionText="Fig. 5. Liphistius malayanus, vulval plate of eight females: paratype of L. m. cameroni from Berinchang, Cameron Highlands (A- B); exuvia of female from Ringlet, Cameron Highlands (C); female (SMF 64093) from Fraser’s Hill (D); female (moulted 24. VI. 2002) from Fraser’s Hill; poreplate slightly deformed by brief immersion in cold KOH (E); female (SMF 56206) from Fraser’s Hill (F); female (SMF 60037) from Genting Highlands (G); female (moulted 8. XII. 2005) from Genting Highlands (H); exuvia of female from Templer Park (I-J). (A, C-I) Dorsal view. (B, J) Ventral view. Scale lines: 1.0 mm (A-B; C, F; D; E, G; H; I-J)." httpUri="https://zenodo.org/record/1021947/files/figure.png" pageId="9" pageNumber="400">Fig. 5</figureCitation>
|
||
). Even more variable is the size and shape of the CDO of the poreplate, ranging from (mostly) longer than wide to (rarely) wider than long, from quadrangular to pentangular and from near-triangular to near-circular (
|
||
<figureCitation id="5CA209C1FFEEFFFDBB21FE9239BDFE99" box="[334,421,355,378]" captionStart="Fig. 5" captionStartId="8.[151,186,1809,1830]" captionTargetBox="[154,1439,260,1762]" captionTargetId="figure@8.[148,1439,254,1779]" captionTargetPageId="8" captionText="Fig. 5. Liphistius malayanus, vulval plate of eight females: paratype of L. m. cameroni from Berinchang, Cameron Highlands (A- B); exuvia of female from Ringlet, Cameron Highlands (C); female (SMF 64093) from Fraser’s Hill (D); female (moulted 24. VI. 2002) from Fraser’s Hill; poreplate slightly deformed by brief immersion in cold KOH (E); female (SMF 56206) from Fraser’s Hill (F); female (SMF 60037) from Genting Highlands (G); female (moulted 8. XII. 2005) from Genting Highlands (H); exuvia of female from Templer Park (I-J). (A, C-I) Dorsal view. (B, J) Ventral view. Scale lines: 1.0 mm (A-B; C, F; D; E, G; H; I-J)." httpUri="https://zenodo.org/record/1021947/files/figure.png" pageId="10" pageNumber="401">Fig. 5A</figureCitation>
|
||
, C-I). The receptacular cluster is always large, undivided and has a complex structure, covering most of the ventral side of the poreplate (
|
||
<figureCitation id="5CA209C1FFEEFFFDB8A6FE5538CBFE3D" captionStart="Fig. 5" captionStartId="8.[151,186,1809,1830]" captionTargetBox="[154,1439,260,1762]" captionTargetId="figure@8.[148,1439,254,1779]" captionTargetPageId="8" captionText="Fig. 5. Liphistius malayanus, vulval plate of eight females: paratype of L. m. cameroni from Berinchang, Cameron Highlands (A- B); exuvia of female from Ringlet, Cameron Highlands (C); female (SMF 64093) from Fraser’s Hill (D); female (moulted 24. VI. 2002) from Fraser’s Hill; poreplate slightly deformed by brief immersion in cold KOH (E); female (SMF 56206) from Fraser’s Hill (F); female (SMF 60037) from Genting Highlands (G); female (moulted 8. XII. 2005) from Genting Highlands (H); exuvia of female from Templer Park (I-J). (A, C-I) Dorsal view. (B, J) Ventral view. Scale lines: 1.0 mm (A-B; C, F; D; E, G; H; I-J)." httpUri="https://zenodo.org/record/1021947/files/figure.png" pageId="10" pageNumber="401">Fig. 5B, J</figureCitation>
|
||
); only in one female examined (from the Genting
|
||
<collectingRegion id="065DDBA6FFEEFFFDBAF8FE143910FE1F" box="[151,264,485,508]" country="Czech Republic" name="Vysocina" pageId="10" pageNumber="401">Highlands</collectingRegion>
|
||
) does it not protrude beyond the anterior margin of the poreplate (
|
||
<figureCitation id="5CA209C1FFEEFFFDBBF0FDF739ECFDFE" box="[415,500,518,541]" captionStart="Fig. 5" captionStartId="8.[151,186,1809,1830]" captionTargetBox="[154,1439,260,1762]" captionTargetId="figure@8.[148,1439,254,1779]" captionTargetPageId="8" captionText="Fig. 5. Liphistius malayanus, vulval plate of eight females: paratype of L. m. cameroni from Berinchang, Cameron Highlands (A- B); exuvia of female from Ringlet, Cameron Highlands (C); female (SMF 64093) from Fraser’s Hill (D); female (moulted 24. VI. 2002) from Fraser’s Hill; poreplate slightly deformed by brief immersion in cold KOH (E); female (SMF 56206) from Fraser’s Hill (F); female (SMF 60037) from Genting Highlands (G); female (moulted 8. XII. 2005) from Genting Highlands (H); exuvia of female from Templer Park (I-J). (A, C-I) Dorsal view. (B, J) Ventral view. Scale lines: 1.0 mm (A-B; C, F; D; E, G; H; I-J)." httpUri="https://zenodo.org/record/1021947/files/figure.png" pageId="10" pageNumber="401">Fig. 5G</figureCitation>
|
||
). The anterior poreplate margin varies from more or less strongly invaginated (in most cases;
|
||
<figureCitation id="5CA209C1FFEEFFFDBB73FDB63943FDBD" box="[284,347,583,606]" captionStart="Fig. 5" captionStartId="8.[151,186,1809,1830]" captionTargetBox="[154,1439,260,1762]" captionTargetId="figure@8.[148,1439,254,1779]" captionTargetPageId="8" captionText="Fig. 5. Liphistius malayanus, vulval plate of eight females: paratype of L. m. cameroni from Berinchang, Cameron Highlands (A- B); exuvia of female from Ringlet, Cameron Highlands (C); female (SMF 64093) from Fraser’s Hill (D); female (moulted 24. VI. 2002) from Fraser’s Hill; poreplate slightly deformed by brief immersion in cold KOH (E); female (SMF 56206) from Fraser’s Hill (F); female (SMF 60037) from Genting Highlands (G); female (moulted 8. XII. 2005) from Genting Highlands (H); exuvia of female from Templer Park (I-J). (A, C-I) Dorsal view. (B, J) Ventral view. Scale lines: 1.0 mm (A-B; C, F; D; E, G; H; I-J)." httpUri="https://zenodo.org/record/1021947/files/figure.png" pageId="10" pageNumber="401">Fig. 5</figureCitation>
|
||
A-E, G-J) to slightly arched medially (
|
||
<figureCitation id="5CA209C1FFEEFFFDBAF0FD9638E9FD9D" box="[159,241,615,638]" captionStart="Fig. 5" captionStartId="8.[151,186,1809,1830]" captionTargetBox="[154,1439,260,1762]" captionTargetId="figure@8.[148,1439,254,1779]" captionTargetPageId="8" captionText="Fig. 5. Liphistius malayanus, vulval plate of eight females: paratype of L. m. cameroni from Berinchang, Cameron Highlands (A- B); exuvia of female from Ringlet, Cameron Highlands (C); female (SMF 64093) from Fraser’s Hill (D); female (moulted 24. VI. 2002) from Fraser’s Hill; poreplate slightly deformed by brief immersion in cold KOH (E); female (SMF 56206) from Fraser’s Hill (F); female (SMF 60037) from Genting Highlands (G); female (moulted 8. XII. 2005) from Genting Highlands (H); exuvia of female from Templer Park (I-J). (A, C-I) Dorsal view. (B, J) Ventral view. Scale lines: 1.0 mm (A-B; C, F; D; E, G; H; I-J)." httpUri="https://zenodo.org/record/1021947/files/figure.png" pageId="10" pageNumber="401">Fig. 5F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8C8346CFFFEEFFFDBAF8FD663AADFBF0" pageId="10" pageNumber="401" type="distribution">
|
||
<paragraph id="C4261544FFEEFFFDBAF8FD663AADFBF0" blockId="10.[151,756,663,1045]" pageId="10" pageNumber="401">
|
||
<emphasis id="F6EDC956FFEEFFFDBAF8FD663936FD4D" bold="true" box="[151,302,663,686]" pageId="10" pageNumber="401">Distribution:</emphasis>
|
||
<taxonomicName id="03996EC7FFEEFFFDBB50FD663A28FD4D" authorityName="Abraham" authorityYear="1923" box="[319,560,663,686]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="401" phylum="Arthropoda" rank="species" species="malayanus">
|
||
<emphasis id="F6EDC956FFEEFFFDBB50FD663A28FD4D" box="[319,560,663,686]" italics="true" pageId="10" pageNumber="401">Liphistius malayanus</emphasis>
|
||
</taxonomicName>
|
||
is known from several lowland and upland localities in the western part of central
|
||
<collectingCountry id="BC8E55D4FFEEFFFDBB38FD2939A2FD0C" box="[343,442,728,751]" name="Malaysia" pageId="10" pageNumber="401">Malaysia</collectingCountry>
|
||
(
|
||
<collectingRegion id="065DDBA6FFEEFFFDBBBCFD293A0DFD0C" box="[467,533,728,751]" country="Malaysia" name="Perak" pageId="10" pageNumber="401">Perak</collectingRegion>
|
||
,
|
||
<collectingRegion id="065DDBA6FFEEFFFDB844FD293A66FD0C" box="[555,638,728,751]" country="Malaysia" name="Pahang" pageId="10" pageNumber="401">Pahang</collectingRegion>
|
||
,
|
||
<collectingRegion id="065DDBA6FFEEFFFDB8FBFD293AECFD0C" box="[660,756,728,751]" country="Malaysia" name="Selangor" pageId="10" pageNumber="401">Selangor</collectingRegion>
|
||
and
|
||
<collectingRegion id="065DDBA6FFEEFFFDBAA6FD09399FFCEC" box="[201,391,760,783]" country="Malaysia" name="Negeri Sembilan" pageId="10" pageNumber="401">Negeri Sembilan</collectingRegion>
|
||
;
|
||
<figureCitation id="5CA209C1FFEEFFFDBBF9FD0939C2FCF2" box="[406,474,760,785]" captionStart="Fig. 1" captionStartId="1.[151,186,1392,1413]" captionTargetBox="[153,758,625,1349]" captionTargetId="figure@1.[149,499,895,1170]" captionTargetPageId="1" captionText="Fig. 1. Localities of Liphistius species of the malayanus - group, tioman - group, linang - group and batuenis - group in peninsular Malaysia and southern Thailand (coast of Sumatra omitted): 1 - Gunung Angsi (type locality of L. malayanus); 2 - Templer Park and Gua Anak Takun (L. malayanus, L. batuensis); 3 - Genting Highlands (L. malayanus); 4 - Fraser’s Hill (L. malayanus); 5 - Cameron Highlands (type locality of L. malayanus cameroni); 6 - Sungai Jasin in Endau Rompin National Park (type locality of L. endau); 7 - Gunung Belumut (L. endau); 8 - Gunung Muntahak (L. endau; type locality of L. gracilis sp. nov.); 9 - Sungai Rengit (type locality of L. johore); 10 - Gunung Kajang on Tioman Island (type locality of L. tioman); 11 - Gua Charas in Bukit Charas (type locality of L. panching); 12 - Nusa Camp in Taman Negara (type locality of L. negara sp. nov.); 13 - Jeram Linang Waterfall (type locality of L. linang sp. nov.); 14 - Sankalakhierie Mountains (type locality of L. indra); 15 - Batu Caves (type locality of L. batuensis); 16 - Gua Tempurung (type locality of L. tempurung); 17 - Gua Cicak (L. tempurung); 18 - Gua Keris (type locality of L. priceae sp. nov). Localities with conspecific populations are encircled. Colours distinguish species groups." httpUri="https://zenodo.org/record/1021937/files/figure.png" pageId="10" pageNumber="401">Fig. 1</figureCitation>
|
||
, localities 1-5). See also
|
||
<bibRefCitation id="A00868B5FFEEFFFDBAF8FCE83A00FCD1" author="Platnick N. & Sedgwick W. C." box="[151,536,793,818]" journalOrPublisher="American Museum Novitates" pageId="10" pageNumber="401" pagination="1 - 31" part="2781" refString="Platnick N. I., Sedgwick W. C. 1984. A revision of the spider genus Liphistius (Araneae, Mesothelae). American Museum Novitates 2781: 1 - 31." title="A revision of the spider genus Liphistius (Araneae, Mesothelae)" type="journal article" year="1984">Platnick & Sedgwick (1984: 24)</bibRefCitation>
|
||
.
|
||
<taxonomicName id="03996EC7FFEEFFFDB844FCE83A8EFCD3" authorityName="Schiodte" authorityYear="1849" box="[555,662,793,816]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="401" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F6EDC956FFEEFFFDB844FCE83A8EFCD3" box="[555,662,793,816]" italics="true" pageId="10" pageNumber="401">Liphistius</emphasis>
|
||
</taxonomicName>
|
||
records from other localties near
|
||
<collectingRegion id="065DDBA6FFEEFFFDBBADFCC83A7FFCB1" box="[450,615,825,850]" country="Malaysia" name="Kuala Lumpur" pageId="10" pageNumber="401">Kuala Lumpur</collectingRegion>
|
||
(e.g. Klang Gates;
|
||
<bibRefCitation id="A00868B5FFEEFFFDBA8BFCAB39E5FC90" author="Murphy F. & Murphy J." bookContentInfo="625 pp." box="[228,509,858,883]" journalOrPublisher="Malaysian Nature Society, Kuala Lumpur" pageId="10" pageNumber="401" refString="Murphy F., Murphy J. 2000. An introduction to the spiders of South East Asia with notes on all the genera. Malaysian Nature Society, Kuala Lumpur, 625 pp., 32 colour plates." title="An introduction to the spiders of South East Asia with notes on all the genera" type="book" year="2000">Murphy & Murphy, 2000</bibRefCitation>
|
||
: plate 2.5) can also be attributed to
|
||
<taxonomicName id="03996EC7FFEEFFFDBB4CFC8D39ACFC72" authorityName="Abraham" authorityYear="1923" box="[291,436,890,915]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="401" phylum="Arthropoda" rank="species" species="malayanus">
|
||
<emphasis id="F6EDC956FFEEFFFDBB4CFC8D39ACFC72" box="[291,436,890,915]" italics="true" pageId="10" pageNumber="401">L. malayanus</emphasis>
|
||
</taxonomicName>
|
||
. This species has a relatively wide geographical range (the northernmost localitiy, in the Cameron
|
||
<collectingRegion id="065DDBA6FFEEFFFDBB5DFC4A39BBFC31" box="[306,419,955,978]" country="Czech Republic" name="Vysocina" pageId="10" pageNumber="401">Highlands</collectingRegion>
|
||
, is about
|
||
<quantity id="0361B8A1FFEEFFFDB876FC4D3A77FC31" box="[537,623,955,980]" metricMagnitude="5" metricUnit="m" metricValue="2.8" pageId="10" pageNumber="401" unit="km" value="280.0">280 km</quantity>
|
||
away from the southernmost locality, Gunung Angsi) and a large vertical distribution (from
|
||
<quantity id="0361B8A1FFEEFFFDBBDCFC0C3A56FBF6" box="[435,590,1021,1045]" metricMagnitude="3" metricUnit="m" metricValue="1.015" metricValueMax="1.8" metricValueMin="0.22999999999999998" pageId="10" pageNumber="401" unit="m" value="1015.0" valueMax="1800.0" valueMin="230.0">230 to 1800 m</quantity>
|
||
altitude).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8C8346CFFFEEFFFDBAF8FBDD3979F872" pageId="10" pageNumber="401" type="biology_ecology">
|
||
<paragraph id="C4261544FFEEFFFDBAF8FBDD38E0F9E9" blockId="10.[151,757,1068,1938]" pageId="10" pageNumber="401">
|
||
<emphasis id="F6EDC956FFEEFFFDBAF8FBDD38EFFBA0" bold="true" box="[151,247,1068,1091]" pageId="10" pageNumber="401">Biology:</emphasis>
|
||
Spiders were collected from quite different habitats: soil on exposed sides of roads and trails inside and outside forests (but never very far from a forest), sides of erosion gullies, sloping forest floor, and in decomposing wood of logs lying on the forest floor. Trapdoors of females were up to
|
||
<quantity id="0361B8A1FFEEFFFDBB96FB3E3A5AFB04" box="[505,578,1231,1255]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="10" pageNumber="401" unit="cm" value="3.8">3.8 cm</quantity>
|
||
long and
|
||
<quantity id="0361B8A1FFEEFFFDB8C4FB3E3AECFB04" box="[683,756,1231,1255]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="10" pageNumber="401" unit="cm" value="6.5">6.5 cm</quantity>
|
||
wide. Signal lines (not more than eight) were usually about
|
||
<quantity id="0361B8A1FFEEFFFDBAB2FAE1393CFACB" box="[221,292,1296,1320]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="401" unit="cm" value="10.0">10 cm</quantity>
|
||
long, in one immature male even
|
||
<quantity id="0361B8A1FFEEFFFDB8C1FAE13AECFACB" box="[686,756,1296,1320]" metricMagnitude="-1" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="401" unit="cm" value="24.0">24 cm</quantity>
|
||
long. The longest burrow measured was
|
||
<quantity id="0361B8A1FFEEFFFDB805FAC03AAAFAAB" box="[618,690,1329,1353]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="401" unit="cm" value="20.0">20 cm</quantity>
|
||
long. Two penultimate males had trapdoors with a length of
|
||
<quantity id="0361B8A1FFEEFFFDBAF8FA833913FA6A" box="[151,267,1394,1418]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="10" pageNumber="401" unit="cm" value="2.05" valueMax="2.2" valueMin="1.9">1.9-2.2 cm</quantity>
|
||
and a width of 3.0-
|
||
<quantity id="0361B8A1FFEEFFFDBBB3FA833A31FA6A" box="[476,553,1394,1418]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="10" pageNumber="401" unit="cm" value="3.4">3.4 cm</quantity>
|
||
, which is probably normal. A juvenile male (moulted again but died before reaching maturity) from near Ringlet in the Cameron
|
||
<collectingRegion id="065DDBA6FFEEFFFDBAF8FA22391DFA09" box="[151,261,1491,1514]" country="Czech Republic" name="Vysocina" pageId="10" pageNumber="401">Highlands</collectingRegion>
|
||
had a surprising
|
||
<quantity id="0361B8A1FFEEFFFDBBD2FA223A1EFA08" box="[445,518,1491,1515]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="10" pageNumber="401" unit="cm" value="2.9">2.9 cm</quantity>
|
||
long and 5.0 cm wide trapdoor.
|
||
</paragraph>
|
||
<paragraph id="C4261544FFEEFFFDBAF8F9E53998F90D" blockId="10.[151,757,1068,1938]" pageId="10" pageNumber="401">
|
||
The female
|
||
<typeStatus id="1B22ABE6FFEEFFFDBB70F9E43963F9CE" box="[287,379,1557,1581]" pageId="10" pageNumber="401" type="paratype">paratype</typeStatus>
|
||
of
|
||
<taxonomicName id="03996EC7FFEEFFFDBBC9F9E43A57F9CF" authorityName="Haupt" authorityYear="1983" box="[422,591,1557,1580]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="401" phylum="Arthropoda" rank="subSpecies" species="malayanus" subSpecies="cameroni">
|
||
<emphasis id="F6EDC956FFEEFFFDBBC9F9E43A57F9CF" box="[422,591,1557,1580]" italics="true" pageId="10" pageNumber="401">L. m. cameroni</emphasis>
|
||
</taxonomicName>
|
||
and two other females (from Fraser’s Hill and Templer Park) show bite marks on their carapaces and chelicerae. These were probably caused by specimens of
|
||
<taxonomicName id="03996EC7FFEEFFFDB848F9843AF5F96F" box="[551,749,1653,1676]" class="Arachnida" family="Laelapidae" genus="Ljunghia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="401" phylum="Arthropoda" rank="species" species="bristowi">
|
||
<emphasis id="F6EDC956FFEEFFFDB848F9843AF5F96F" box="[551,749,1653,1676]" italics="true" pageId="10" pageNumber="401">Ljunghia bristowi</emphasis>
|
||
</taxonomicName>
|
||
, a species of ectoparasitic laelapid mites originally described from
|
||
<taxonomicName id="03996EC7FFEEFFFDBB27F94639C5F92E" authorityName="Abraham" authorityYear="1923" box="[328,477,1718,1742]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="401" phylum="Arthropoda" rank="species" species="malayanus">
|
||
<emphasis id="F6EDC956FFEEFFFDBB27F94639C5F92E" box="[328,477,1718,1742]" italics="true" pageId="10" pageNumber="401">L. malayanus</emphasis>
|
||
</taxonomicName>
|
||
(see Halliday & Juvara- Bals, 2016: 837-845).
|
||
</paragraph>
|
||
<paragraph id="C4261544FFEEFFFDBAF8F9093979F872" blockId="10.[151,757,1068,1938]" pageId="10" pageNumber="401">
|
||
Mature males were collected in the field in October and December; in captivity males matured between July and March. The mating period in
|
||
<taxonomicName id="03996EC7FFEEFFFDBBBEF8C83A79F8AC" authorityName="Abraham" authorityYear="1923" box="[465,609,1848,1872]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="401" phylum="Arthropoda" rank="species" species="malayanus">
|
||
<emphasis id="F6EDC956FFEEFFFDBBBEF8C83A79F8AC" box="[465,609,1848,1872]" italics="true" pageId="10" pageNumber="401">L. malayanus</emphasis>
|
||
</taxonomicName>
|
||
appears to be much longer than in other congeneric species. I did not find any egg cases.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |