treatments-xml/data/4C/30/A4/4C30A452FFCFFFD9BB5AFEF03A8BF83D.xml

<document id="3CB23243BC5A3FBB6C85C4D7B374B4BC" ID-CLB-Dataset="31806" ID-DOI="10.5281/zenodo.893555" ID-GBIF-Dataset="4a2efe36-c48b-456b-9223-7b5793025a72" ID-ISSN="0035-418" ID-Zenodo-Dep="893555" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1508361162557" checkinUser="plazi" docAuthor="Peter J. Schwendinger" docDate="2017" docId="4C30A452FFCFFFD9BB5AFEF03A8BF83D" docLanguage="en" docName="rsz_124_2_11_Schwendinger.pdf" docOrigin="Revue suisse de Zoologie 124 (2)" docStyle="DocumentStyle:225056F0A23C51DDC06AC27EAEA15D5B.2:RevSuisseZool.2015-2019.journal_article" docStyleId="225056F0A23C51DDC06AC27EAEA15D5B" docStyleName="RevSuisseZool.2015-2019.journal_article" docStyleVersion="2" docTitle="Liphistius priceae Peter J. Schwendinger 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="437" masterDocId="B009DC2AFFE4FFF7BA6FFFF13818FFE3" masterDocTitle="A revision of the trapdoor spider genus Liphistius (Mesothelae: Liphistiidae) in peninsular Malaysia; part 1" masterLastPageNumber="445" masterPageNumber="391" pageNumber="434" updateTime="1698469770161" updateUser="ExternalLinkService">
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<mods:title id="CE4BF0129A3A56E2B33111AE8B65F2E9">A revision of the trapdoor spider genus Liphistius (Mesothelae: Liphistiidae) in peninsular Malaysia; part 1</mods:title>
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<treatment id="4C30A452FFCFFFD9BB5AFEF03A8BF83D" ID-DOI="http://doi.org/10.5281/zenodo.6042388" ID-GBIF-Taxon="135450782" ID-Zenodo-Dep="6042388" LSID="urn:lsid:plazi:treatment:4C30A452FFCFFFD9BB5AFEF03A8BF83D" httpUri="http://treatment.plazi.org/id/4C30A452FFCFFFD9BB5AFEF03A8BF83D" lastPageId="46" lastPageNumber="437" pageId="43" pageNumber="434">
<subSubSection id="8C8346CFFFCFFFDCBB5AFEF039E7FED9" pageId="43" pageNumber="434" type="nomenclature">
<paragraph id="C4261544FFCFFFDCBB5AFEF03A4EFEF9" blockId="43.[309,598,257,314]" box="[309,598,257,282]" pageId="43" pageNumber="434">
<heading id="9F6EA228FFCFFFDCBB5AFEF03A4EFEF9" bold="true" box="[309,598,257,282]" centered="true" fontSize="10" level="3" pageId="43" pageNumber="434" reason="4">
<emphasis id="F6EDC956FFCFFFDCBB5AFEF03A4EFEF9" bold="true" box="[309,598,257,282]" pageId="43" pageNumber="434">
<taxonomicName id="03996EC7FFCFFFDCBB5AFEF039E2FEFB" ID-CoL="72J6Z" authority="Peter J. Schwendinger, 2017" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[309,506,257,280]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="434" phylum="Arthropoda" rank="species" species="priceae" status="sp. nov.">
<emphasis id="F6EDC956FFCFFFDCBB5AFEF039E2FEFB" bold="true" box="[309,506,257,280]" italics="true" pageId="43" pageNumber="434">Liphistius priceae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDDE742DFFCFFFDCB86FFEF33A4EFEF9" box="[512,598,258,282]" pageId="43" pageNumber="434" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="C4261544FFCFFFDCBBE3FED339E7FED9" blockId="43.[309,598,257,314]" box="[396,511,290,314]" pageId="43" pageNumber="434">
<figureCitation id="5CA209C1FFCFFFDCBBE3FED339E7FED9" box="[396,511,290,314]" captionStart-0="Fig. 23" captionStart-1="Fig. 24" captionStartId-0="44.[147,182,1855,1876]" captionStartId-1="45.[149,184,1333,1354]" captionTargetBox-0="[157,1429,261,1790]" captionTargetBox-1="[156,1432,263,1274]" captionTargetId-0="figure@44.[148,1439,254,1807]" captionTargetId-1="figure@45.[148,1446,254,1296]" captionTargetPageId-0="44" captionTargetPageId-1="45" captionText-0="Fig. 23. Liphistius priceae sp. nov., details of palp of six males; holotype (A-D, H-K), male paratype collected mature (E), paratype matured 20. VII. 2004 (F), paratype matured 1. VIII. 2004 (G), paratype matured 18. IX. 2004 (L-M) and paratype matured 11. VIII. 2005 (N-O). (A) Left cymbium and palpal organ, ventral view. (B) Distal part of left tibia, ventral view. (C) Left paracymbium, retrolateral view. (D-E) Left palpal organ, distal view (dorsal side up). (F-G) Dorsal apex of contrategulum of right palpal organ, distal view (dorsal side to the left). (H) Distal part of left palpal organ, prolateral view. (I) Same, retrodorsal and slightly distal view. (J) Distal part of cymbium, prolateral view. (K-L, N) Left tibial apophysis, retrolateral and slightly proximal view. (M, O) Right tibial apophysis, same view. Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A-C, K-O), 0.5 mm (D-I; J)." captionText-1="Fig. 24. Liphistius priceae sp. nov., vulval plate of four female paratypes (all from exuviae): paratype, moult 1. I. 2006 (A-B), allotype, moult 24. XI. 2004 (C-D), paratype, moult 13. XI. 2005 (E), paratype, moult 2. IV. 2005 (F). (A, C, E-F) Dorsal view. (B) Lateral view. (D) Ventral view. Scale lines: 0.5 mm (A-F)." httpUri-0="https://zenodo.org/record/1021987/files/figure.png" httpUri-1="https://zenodo.org/record/1021989/files/figure.png" pageId="43" pageNumber="434">Figs 23-24</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFCFFFDCBAF8FEA33963FD4A" pageId="43" pageNumber="434" type="description">
<paragraph id="C4261544FFCFFFDCBAF8FEA33963FD4A" blockId="43.[151,756,337,683]" pageId="43" pageNumber="434">
<emphasis id="F6EDC956FFCFFFDCBAF8FEA338FBFE89" bold="true" box="[151,227,338,362]" pageId="43" pageNumber="434">Types:</emphasis>
MHNG (most types, including the holotype), SMF (1 male and 1 female paratypes), sample MAL- 04/10; male holotype (matured late 
<date id="B0273384FFCFFFDCB84AFE633A8BFE49" box="[549,659,402,426]" pageId="43" pageNumber="434" value="2004-08">VIII.2004</date>
), 
<materialsCitation id="74F11F19FFCFFFDCB8C6FE633AB7FD89" ID-GBIF-Occurrence="1675315689" collectingDate="2004-11-24" collectingDateMax="2005-08-11" collectingDateMin="2004-11-24" country="Malaysia" county="Gua Keris" elevation="130" location="Gua Pagar" municipality="Kris Cave" pageId="43" pageNumber="434" specimenCount="23" specimenCount-female="11" specimenCount-male="11" stateProvince="Kelantan" typeStatus="paratype">
<specimenCount id="D29FDECDFFCFFFDCB8C6FE633AECFE4B" box="[681,756,401,426]" pageId="43" pageNumber="434" type="male">8 male</specimenCount>
<typeStatus id="1B22ABE6FFCFFFDCBAF8FE4338E6FE29" box="[151,254,434,458]" pageId="43" pageNumber="434" type="paratype">paratypes</typeStatus>
(matured 
<collectingDate id="A063CA6CFFCFFFDCBB1CFE433AABFE29" box="[371,691,434,458]" isEnumeration="true" pageId="43" pageNumber="434">
<date id="B0273384FFCFFFDCBB1CFE4339DCFE29" box="[371,452,434,458]" pageId="43" pageNumber="434" value="2004-07">20.VII.</date>
, 
<date id="B0273384FFCFFFDCBBBDFE433A05FE29" box="[466,541,434,458]" pageId="43" pageNumber="434" value="2004-08">1.VIII.</date>
, 
<date id="B0273384FFCFFFDCB843FE433AABFE29" box="[556,691,434,458]" pageId="43" pageNumber="434" value="2004-09-18">18.IX. 2004</date>
</collectingDate>
, mid- VII.; 
<collectingDate id="A063CA6CFFCFFFDCBAB9FE20397CFE0B" box="[214,356,465,488]" pageId="43" pageNumber="434" value="2005-08-11">
<date id="B0273384FFCFFFDCBAB9FE20397CFE0B" box="[214,356,465,488]" pageId="43" pageNumber="434" value="2005-08-11">11.VIII.2005</date>
</collectingDate>
; 
<specimenCount id="D29FDECDFFCFFFDCBB1BFE2339D8FE0B" box="[372,448,465,490]" pageId="43" pageNumber="434" type="male">3 male</specimenCount>
<typeStatus id="1B22ABE6FFCFFFDCBBA4FE223A2AFE08" box="[459,562,467,491]" pageId="43" pageNumber="434" type="paratype">paratypes</typeStatus>
collected mature at the site) and 
<specimenCount id="D29FDECDFFCFFFDCBB2FFE0339B3FDEA" box="[320,427,498,522]" pageId="43" pageNumber="434" type="female">11 female</specimenCount>
<typeStatus id="1B22ABE6FFCFFFDCBBDCFE023A01FDE8" box="[435,537,499,523]" pageId="43" pageNumber="434" type="paratype">paratypes</typeStatus>
(including 
<typeStatus id="1B22ABE6FFCFFFDCB8F8FE033AE8FDEA" box="[663,752,498,521]" pageId="43" pageNumber="434" type="allotype">allotype</typeStatus>
, moulted 
<collectingDate id="A063CA6CFFCFFFDCBA96FDE33960FDC9" box="[249,376,530,554]" pageId="43" pageNumber="434" value="2004-11-24">
<date id="B0273384FFCFFFDCBA96FDE33960FDC9" box="[249,376,530,554]" pageId="43" pageNumber="434" value="2004-11-24">24.XI.2004</date>
</collectingDate>
); 
<collectingCountry id="BC8E55D4FFCFFFDCBBE1FDE339EBFDCA" box="[398,499,530,553]" name="Malaysia" pageId="43" pageNumber="434">Malaysia</collectingCountry>
, 
<collectingRegion id="065DDBA6FFCFFFDCB86DFDE33A7EFDCA" box="[514,614,530,553]" country="Malaysia" name="Kelantan" pageId="43" pageNumber="434">Kelantan</collectingRegion>
, about 
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<quantity id="0361B8A1FFCFFFDCB8D5FDE33AECFDCA" box="[698,756,530,553]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="43" pageNumber="434" unit="km" value="5.0">5 km</quantity>
S of Dabong
</locationDeviation>
, 
<collectingCounty id="2D476DC8FFCFFFDCBB5BFDC239BDFDAA" box="[308,421,562,587]" pageId="43" pageNumber="434">Gua Keris</collectingCounty>
(= 
<collectingMunicipality id="24428F3EFFCFFFDCBBBFFDC33A27FDA8" box="[464,575,562,587]" pageId="43" pageNumber="434">Kris Cave</collectingMunicipality>
) and 
<location id="C146439FFFCFFFDCB8EEFDC23AECFDA8" LSID="urn:lsid:plazi:treatment:4C30A452FFCFFFD9BB5AFEF03A8BF83D:C146439FFFCFFFDCB8EEFDC23AECFDA8" box="[641,756,563,587]" country="Malaysia" county="Gua Keris" municipality="Kris Cave" name="Gua Pagar" pageId="43" pageNumber="434" stateProvince="Kelantan">Gua Pagar</location>
(locally also called 
<location id="C146439FFFCFFFDCBBECFDA23A5CFD88" LSID="urn:lsid:plazi:treatment:4C30A452FFCFFFD9BB5AFEF03A8BF83D:C146439FFFCFFFDCBBECFDA23A5CFD88" box="[387,580,594,619]" country="Malaysia" county="Gua Keris" municipality="Kris Cave" name="Gua King Kong" pageId="43" pageNumber="434" stateProvince="Kelantan">Gua King Kong</location>
), 
<elevation id="4FB4F277FFCFFFDCB830FDA23AB7FD89" box="[607,687,595,619]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="43" pageNumber="434" unit="m" value="130.0">
<quantity id="0361B8A1FFCFFFDCB830FDA23AB7FD89" box="[607,687,595,619]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="43" pageNumber="434" unit="m" value="130.0">130 m</quantity>
</elevation>
</materialsCitation>
; 
<date id="B0273384FFCFFFDCB8ADFDA238ECFD68" pageId="43" pageNumber="434" value="2004-06-08" valueMax="2004-06-09" valueMin="2004-06-08">8.-9. VI.2004</date>
; leg. P.J. Schwendinger. The precise type locality is Gua Keris.
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFCFFFDCBAF8FD333A31FC7A" pageId="43" pageNumber="434" type="etymology">
<paragraph id="C4261544FFCFFFDCBAF8FD333A31FC7A" blockId="43.[151,756,706,921]" pageId="43" pageNumber="434">
<emphasis id="F6EDC956FFCFFFDCBAF8FD333904FD39" bold="true" box="[151,284,706,730]" pageId="43" pageNumber="434">Etymology:</emphasis>
The new species is named in honour of Liz Price (London, 
<collectingCountry id="BC8E55D4FFCFFFDCBB15FD1339BDFD19" box="[378,421,738,762]" name="United Kingdom" pageId="43" pageNumber="434">UK</collectingCountry>
), a former long-time resident in 
<collectingRegion id="065DDBA6FFCFFFDCBAD8FCF33943FCF9" box="[183,347,770,794]" country="Malaysia" name="Kuala Lumpur" pageId="43" pageNumber="434">Kuala Lumpur</collectingRegion>
, and a very active speleologist who over 30 years explored and published on caves and cave faunas all over Southeast Asia. She was also involved in conservation and trying to save some caves from destruction by quarrying in 
<collectingCountry id="BC8E55D4FFCFFFDCBBD0FC733A3CFC7A" box="[447,548,898,921]" name="Malaysia" pageId="43" pageNumber="434">Malaysia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFCFFFDCBAF8FC4338E6F92E" pageId="43" pageNumber="434" type="diagnosis">
<paragraph id="C4261544FFCFFFDCBAF8FC4338E6F92E" blockId="43.[151,757,946,1741]" pageId="43" pageNumber="434">
<emphasis id="F6EDC956FFCFFFDCBAF8FC433909FC2A" bold="true" box="[151,273,946,969]" pageId="43" pageNumber="434">Diagnosis:</emphasis>
Small to medium-sized, light-browncoloured species, similar and closely related to 
<taxonomicName id="03996EC7FFCFFFDCBAF8FC023936FBE9" authorityName="Platnick in Platnick, Schwendinger &amp; Steiner" authorityYear="1997" box="[151,302,1011,1034]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="434" phylum="Arthropoda" rank="species" species="tempurung">
<emphasis id="F6EDC956FFCFFFDCBAF8FC023936FBE9" box="[151,302,1011,1034]" italics="true" pageId="43" pageNumber="434">L. tempurung</emphasis>
</taxonomicName>
. Both sexes slightly smaller than those of 
<taxonomicName id="03996EC7FFCFFFDCBAD5FBE23949FBC9" authorityName="Platnick in Platnick, Schwendinger &amp; Steiner" authorityYear="1997" box="[186,337,1043,1066]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="434" phylum="Arthropoda" rank="species" species="tempurung">
<emphasis id="F6EDC956FFCFFFDCBAD5FBE23949FBC9" box="[186,337,1043,1066]" italics="true" pageId="43" pageNumber="434">L. tempurung</emphasis>
</taxonomicName>
(carapace length of males 4.42-5.23 versus 
<quantity id="0361B8A1FFCFFFDCBA8BFBC33970FBAA" box="[228,360,1074,1097]" metricMagnitude="-1" metricUnit="m" metricValue="1.52019" metricValueMax="1.64084" metricValueMin="1.39954" pageId="43" pageNumber="434" unit="in" value="5.984999999999999" valueMax="6.46" valueMin="5.51">5.51-6.46 in</quantity>
<taxonomicName id="03996EC7FFCFFFDCBB1EFBC53A1CFBA8" authorityName="Platnick in Platnick, Schwendinger &amp; Steiner" authorityYear="1997" box="[369,516,1076,1099]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="434" phylum="Arthropoda" rank="species" species="tempurung">
<emphasis id="F6EDC956FFCFFFDCBB1EFBC53A1CFBA8" box="[369,516,1076,1099]" italics="true" pageId="43" pageNumber="434">L. tempurung</emphasis>
</taxonomicName>
). Males distinguished by scopula on tarsus IV more extensive; palpal tibia with a longer retrolateral apophysis carrying much shorter megaspines (
<figureCitation id="5CA209C1FFCFFFDCBBECFB6239F3FB48" box="[387,491,1171,1195]" captionStart="Fig. 23" captionStartId="44.[147,182,1855,1876]" captionTargetBox="[157,1429,261,1790]" captionTargetId="figure@44.[148,1439,254,1807]" captionTargetPageId="44" captionText="Fig. 23. Liphistius priceae sp. nov., details of palp of six males; holotype (A-D, H-K), male paratype collected mature (E), paratype matured 20. VII. 2004 (F), paratype matured 1. VIII. 2004 (G), paratype matured 18. IX. 2004 (L-M) and paratype matured 11. VIII. 2005 (N-O). (A) Left cymbium and palpal organ, ventral view. (B) Distal part of left tibia, ventral view. (C) Left paracymbium, retrolateral view. (D-E) Left palpal organ, distal view (dorsal side up). (F-G) Dorsal apex of contrategulum of right palpal organ, distal view (dorsal side to the left). (H) Distal part of left palpal organ, prolateral view. (I) Same, retrodorsal and slightly distal view. (J) Distal part of cymbium, prolateral view. (K-L, N) Left tibial apophysis, retrolateral and slightly proximal view. (M, O) Right tibial apophysis, same view. Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A-C, K-O), 0.5 mm (D-I; J)." httpUri="https://zenodo.org/record/1021987/files/figure.png" pageId="43" pageNumber="434">Fig. 23B</figureCitation>
, K-O cf. 
<figureCitation id="5CA209C1FFCFFFDCB80CFB623AAEFB48" box="[611,694,1171,1195]" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="43" pageNumber="434">Fig. 21</figureCitation>
E-G); paracymbium larger, with a more prominent retrolateralproximal part (
<figureCitation id="5CA209C1FFCFFFDCBB2CFB2239B0FB0F" box="[323,424,1235,1260]" captionStart="Fig. 23" captionStartId="44.[147,182,1855,1876]" captionTargetBox="[157,1429,261,1790]" captionTargetId="figure@44.[148,1439,254,1807]" captionTargetPageId="44" captionText="Fig. 23. Liphistius priceae sp. nov., details of palp of six males; holotype (A-D, H-K), male paratype collected mature (E), paratype matured 20. VII. 2004 (F), paratype matured 1. VIII. 2004 (G), paratype matured 18. IX. 2004 (L-M) and paratype matured 11. VIII. 2005 (N-O). (A) Left cymbium and palpal organ, ventral view. (B) Distal part of left tibia, ventral view. (C) Left paracymbium, retrolateral view. (D-E) Left palpal organ, distal view (dorsal side up). (F-G) Dorsal apex of contrategulum of right palpal organ, distal view (dorsal side to the left). (H) Distal part of left palpal organ, prolateral view. (I) Same, retrodorsal and slightly distal view. (J) Distal part of cymbium, prolateral view. (K-L, N) Left tibial apophysis, retrolateral and slightly proximal view. (M, O) Right tibial apophysis, same view. Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A-C, K-O), 0.5 mm (D-I; J)." httpUri="https://zenodo.org/record/1021987/files/figure.png" pageId="43" pageNumber="434">Fig. 23A</figureCitation>
cf. 
<figureCitation id="5CA209C1FFCFFFDCBBB2FB223A5EFB0F" box="[477,582,1235,1260]" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="43" pageNumber="434">Fig. 21A</figureCitation>
); tegulum with more clearly outlined proximal edge (
<figureCitation id="5CA209C1FFCFFFDCB80AFB023ADFFAEF" box="[613,711,1267,1292]" captionStart="Fig. 23" captionStartId="44.[147,182,1855,1876]" captionTargetBox="[157,1429,261,1790]" captionTargetId="figure@44.[148,1439,254,1807]" captionTargetPageId="44" captionText="Fig. 23. Liphistius priceae sp. nov., details of palp of six males; holotype (A-D, H-K), male paratype collected mature (E), paratype matured 20. VII. 2004 (F), paratype matured 1. VIII. 2004 (G), paratype matured 18. IX. 2004 (L-M) and paratype matured 11. VIII. 2005 (N-O). (A) Left cymbium and palpal organ, ventral view. (B) Distal part of left tibia, ventral view. (C) Left paracymbium, retrolateral view. (D-E) Left palpal organ, distal view (dorsal side up). (F-G) Dorsal apex of contrategulum of right palpal organ, distal view (dorsal side to the left). (H) Distal part of left palpal organ, prolateral view. (I) Same, retrodorsal and slightly distal view. (J) Distal part of cymbium, prolateral view. (K-L, N) Left tibial apophysis, retrolateral and slightly proximal view. (M, O) Right tibial apophysis, same view. Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A-C, K-O), 0.5 mm (D-I; J)." httpUri="https://zenodo.org/record/1021987/files/figure.png" pageId="43" pageNumber="434">
Fig. 
<date id="B0273384FFCFFFDCB8CCFB053ADFFAEF" box="[675,711,1268,1292]" pageId="43" pageNumber="434" value="1923-01">23I</date>
</figureCitation>
cf. 
<figureCitation id="5CA209C1FFCFFFDCBAF8FAE238E2FACF" box="[151,250,1299,1324]" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="43" pageNumber="434">Fig. 21B</figureCitation>
); contrategulum with series of oblique ribs on distal edge extending further prolaterally, dorsal apex much smaller and narrower, with a more pointed tip (
<figureCitation id="5CA209C1FFCFFFDCBAF0FA8538E8FA6F" box="[159,240,1396,1420]" captionStart="Fig. 23" captionStartId="44.[147,182,1855,1876]" captionTargetBox="[157,1429,261,1790]" captionTargetId="figure@44.[148,1439,254,1807]" captionTargetPageId="44" captionText="Fig. 23. Liphistius priceae sp. nov., details of palp of six males; holotype (A-D, H-K), male paratype collected mature (E), paratype matured 20. VII. 2004 (F), paratype matured 1. VIII. 2004 (G), paratype matured 18. IX. 2004 (L-M) and paratype matured 11. VIII. 2005 (N-O). (A) Left cymbium and palpal organ, ventral view. (B) Distal part of left tibia, ventral view. (C) Left paracymbium, retrolateral view. (D-E) Left palpal organ, distal view (dorsal side up). (F-G) Dorsal apex of contrategulum of right palpal organ, distal view (dorsal side to the left). (H) Distal part of left palpal organ, prolateral view. (I) Same, retrodorsal and slightly distal view. (J) Distal part of cymbium, prolateral view. (K-L, N) Left tibial apophysis, retrolateral and slightly proximal view. (M, O) Right tibial apophysis, same view. Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A-C, K-O), 0.5 mm (D-I; J)." httpUri="https://zenodo.org/record/1021987/files/figure.png" pageId="43" pageNumber="434">Fig. 23</figureCitation>
D-G cf. 
<figureCitation id="5CA209C1FFCFFFDCBB38FA8539B1FA6F" box="[343,425,1396,1420]" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="43" pageNumber="434">Fig. 21</figureCitation>
I-L); para-embolic plate with more prominent, angular distal margin (
<figureCitation id="5CA209C1FFCFFFDCB83CFA653AA1FA4E" box="[595,697,1428,1453]" captionStart="Fig. 23" captionStartId="44.[147,182,1855,1876]" captionTargetBox="[157,1429,261,1790]" captionTargetId="figure@44.[148,1439,254,1807]" captionTargetPageId="44" captionText="Fig. 23. Liphistius priceae sp. nov., details of palp of six males; holotype (A-D, H-K), male paratype collected mature (E), paratype matured 20. VII. 2004 (F), paratype matured 1. VIII. 2004 (G), paratype matured 18. IX. 2004 (L-M) and paratype matured 11. VIII. 2005 (N-O). (A) Left cymbium and palpal organ, ventral view. (B) Distal part of left tibia, ventral view. (C) Left paracymbium, retrolateral view. (D-E) Left palpal organ, distal view (dorsal side up). (F-G) Dorsal apex of contrategulum of right palpal organ, distal view (dorsal side to the left). (H) Distal part of left palpal organ, prolateral view. (I) Same, retrodorsal and slightly distal view. (J) Distal part of cymbium, prolateral view. (K-L, N) Left tibial apophysis, retrolateral and slightly proximal view. (M, O) Right tibial apophysis, same view. Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A-C, K-O), 0.5 mm (D-I; J)." httpUri="https://zenodo.org/record/1021987/files/figure.png" pageId="43" pageNumber="434">Fig. 23A</figureCitation>
, D-E cf. 
<figureCitation id="5CA209C1FFCFFFDCBAD1FA45393DFA2E" box="[190,293,1460,1485]" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="43" pageNumber="434">Fig. 21A</figureCitation>
, I-L); sclerotised part of embolus proper with more pointed apex (
<figureCitation id="5CA209C1FFCFFFDCBBDAFA253A2CFA0F" box="[437,564,1492,1517]" captionStart="Fig. 23" captionStartId="44.[147,182,1855,1876]" captionTargetBox="[157,1429,261,1790]" captionTargetId="figure@44.[148,1439,254,1807]" captionTargetPageId="44" captionText="Fig. 23. Liphistius priceae sp. nov., details of palp of six males; holotype (A-D, H-K), male paratype collected mature (E), paratype matured 20. VII. 2004 (F), paratype matured 1. VIII. 2004 (G), paratype matured 18. IX. 2004 (L-M) and paratype matured 11. VIII. 2005 (N-O). (A) Left cymbium and palpal organ, ventral view. (B) Distal part of left tibia, ventral view. (C) Left paracymbium, retrolateral view. (D-E) Left palpal organ, distal view (dorsal side up). (F-G) Dorsal apex of contrategulum of right palpal organ, distal view (dorsal side to the left). (H) Distal part of left palpal organ, prolateral view. (I) Same, retrodorsal and slightly distal view. (J) Distal part of cymbium, prolateral view. (K-L, N) Left tibial apophysis, retrolateral and slightly proximal view. (M, O) Right tibial apophysis, same view. Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A-C, K-O), 0.5 mm (D-I; J)." httpUri="https://zenodo.org/record/1021987/files/figure.png" pageId="43" pageNumber="434">Fig. 23A, I</figureCitation>
cf. 
<figureCitation id="5CA209C1FFCFFFDCB809FA253AAEFA0E" box="[614,694,1492,1517]" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="43" pageNumber="434">Fig. 21</figureCitation>
A-B). Females distinguished from those of 
<taxonomicName id="03996EC7FFCFFFDCB856FA073AD6F9EE" authorityName="Platnick in Platnick, Schwendinger &amp; Steiner" authorityYear="1997" box="[569,718,1526,1549]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="434" phylum="Arthropoda" rank="species" species="tempurung">
<emphasis id="F6EDC956FFCFFFDCB856FA073AD6F9EE" box="[569,718,1526,1549]" italics="true" pageId="43" pageNumber="434">L. tempurung</emphasis>
</taxonomicName>
by larger and more prominent receptacular cluster (
<figureCitation id="5CA209C1FFCFFFDCB8CAF9E53AECF9CE" box="[677,756,1556,1581]" captionStart="Fig. 24" captionStartId="45.[149,184,1333,1354]" captionTargetBox="[156,1432,263,1274]" captionTargetId="figure@45.[148,1446,254,1296]" captionTargetPageId="45" captionText="Fig. 24. Liphistius priceae sp. nov., vulval plate of four female paratypes (all from exuviae): paratype, moult 1. I. 2006 (A-B), allotype, moult 24. XI. 2004 (C-D), paratype, moult 13. XI. 2005 (E), paratype, moult 2. IV. 2005 (F). (A, C, E-F) Dorsal view. (B) Lateral view. (D) Ventral view. Scale lines: 0.5 mm (A-F)." httpUri="https://zenodo.org/record/1021989/files/figure.png" pageId="43" pageNumber="434">Fig. 24</figureCitation>
cf. 
<figureCitation id="5CA209C1FFCFFFDCBAADF9C53902F9AE" box="[194,282,1588,1613]" captionStart="Fig. 22" captionStartId="41.[147,182,1789,1810]" captionTargetBox="[153,1429,258,1732]" captionTargetId="figure@41.[148,1446,254,1752]" captionTargetPageId="41" captionText="Fig. 22. Liphistius tempurung, details of copulatory organ and genital sternite of six females: Gua Cicak, specimen moulted 8. I. 2005 (A- B, N); Gua Tempurung, exuvia, moult 22. I. 2007 (C-D, M); Gua Tempurung, exuvia, moult 9. XII. 2004 (E-F); Gua Tempurung, exuvia, moult 21. X. 2004 (G-H); medium-sized female from Gua Cicak (I-J); small female from Gua Tempurung (K-L). (A, C, E, G, I, K) Vulval plate, dorsal view. (B, D, F, H) Same, ventral view. (J, L) Receptacular cluster, ventral view. (M-N) Posterior margin of genital sternite, ventral view. Scale lines: 1.0 mm (A-D, I-L; E-H; M-N)." httpUri="https://zenodo.org/record/1021985/files/figure.png" pageId="43" pageNumber="434">Fig. 22</figureCitation>
); CDO much wider and more distinctly outlined, not sitting in a hollow (
<figureCitation id="5CA209C1FFCFFFDCB862F9A43A8FF98E" box="[525,663,1621,1645]" captionStart="Fig. 24" captionStartId="45.[149,184,1333,1354]" captionTargetBox="[156,1432,263,1274]" captionTargetId="figure@45.[148,1446,254,1296]" captionTargetPageId="45" captionText="Fig. 24. Liphistius priceae sp. nov., vulval plate of four female paratypes (all from exuviae): paratype, moult 1. I. 2006 (A-B), allotype, moult 24. XI. 2004 (C-D), paratype, moult 13. XI. 2005 (E), paratype, moult 2. IV. 2005 (F). (A, C, E-F) Dorsal view. (B) Lateral view. (D) Ventral view. Scale lines: 0.5 mm (A-F)." httpUri="https://zenodo.org/record/1021989/files/figure.png" pageId="43" pageNumber="434">Fig. 24A, C</figureCitation>
, E-F cf. 
<figureCitation id="5CA209C1FFCFFFDCBAF8F9843975F96D" box="[151,365,1653,1678]" captionStart="Fig. 22" captionStartId="41.[147,182,1789,1810]" captionTargetBox="[153,1429,258,1732]" captionTargetId="figure@41.[148,1446,254,1752]" captionTargetPageId="41" captionText="Fig. 22. Liphistius tempurung, details of copulatory organ and genital sternite of six females: Gua Cicak, specimen moulted 8. I. 2005 (A- B, N); Gua Tempurung, exuvia, moult 22. I. 2007 (C-D, M); Gua Tempurung, exuvia, moult 9. XII. 2004 (E-F); Gua Tempurung, exuvia, moult 21. X. 2004 (G-H); medium-sized female from Gua Cicak (I-J); small female from Gua Tempurung (K-L). (A, C, E, G, I, K) Vulval plate, dorsal view. (B, D, F, H) Same, ventral view. (J, L) Receptacular cluster, ventral view. (M-N) Posterior margin of genital sternite, ventral view. Scale lines: 1.0 mm (A-D, I-L; E-H; M-N)." httpUri="https://zenodo.org/record/1021985/files/figure.png" pageId="43" pageNumber="434">Fig. 22A, C, E, G, I</figureCitation>
); anterior margin of poreplate not or only indistinctly invaginated; posterior stalk anteriorly narrower.
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFCFFFD9BAF8F9153AF8FBFE" lastPageId="46" lastPageNumber="437" pageId="43" pageNumber="434" type="description">
<paragraph id="C4261544FFCFFFDCBAF8F9153BF6FE99" blockId="43.[151,757,1764,2078]" lastBlockId="43.[831,1437,258,1901]" pageId="43" pageNumber="434">
<emphasis id="F6EDC956FFCFFFDCBAF8F9153A0DF918" bold="true" box="[151,533,1764,1789]" pageId="43" pageNumber="434">
Description of male (
<typeStatus id="1B22ABE6FFCFFFDCBBF0F9153A11F918" box="[415,521,1764,1787]" pageId="43" pageNumber="434" type="holotype">holotype</typeStatus>
):
</emphasis>
<emphasis id="F6EDC956FFCFFFDCB84BF9153AECF918" box="[548,756,1764,1789]" italics="true" pageId="43" pageNumber="434">Colour in alcohol</emphasis>
(slightly darker in life): Body mostly light brown, carapace with indistinct cream-coloured and flowershaped area around fovea. Chelicerae with proximal portion cream-coloured, distal portion light brown. Cheliceral fangs, palpal tarsi and sclerites of palpal organ reddish brown (darker than rest of body). Legs with indistinct annulations (light median rings) on all tibiae and metatarsi; legs and palps with a light distal zone on patellae. Opisthosomal tergite I uniformly light brown, following tergites with increasingly larger light areas in posterior part (except for bases of paired para-median spines); membranous part of opisthosoma cream-coloured.
</paragraph>
<paragraph id="C4261544FFCFFFDCB950FE723D3FFE1F" blockId="43.[831,1437,258,1901]" pageId="43" pageNumber="434">
<emphasis id="F6EDC956FFCFFFDCB950FE723C28FE7F" box="[831,1072,387,412]" italics="true" pageId="43" pageNumber="434">Bristles on carapace</emphasis>
: Stiff bristles on anterior and posterior margins, as well as on and behind eye mound; fewer weaker bristles on lateral margins and on posterior coxal elevations; no bristles anterior to fovea.
</paragraph>
<paragraph id="C4261544FFCFFFDCB950FDF43CA3FDDF" blockId="43.[831,1437,258,1901]" pageId="43" pageNumber="434">
<emphasis id="F6EDC956FFCFFFDCB950FDF43BE8FDFF" box="[831,1008,517,540]" italics="true" pageId="43" pageNumber="434">Cheliceral teeth</emphasis>
: Twelve small teeth on promargin of cheliceral groove of each chelicera.
</paragraph>
<paragraph id="C4261544FFCFFFDCB950FDB73D49FD3D" blockId="43.[831,1437,258,1901]" pageId="43" pageNumber="434">
<emphasis id="F6EDC956FFCFFFDCB950FDB73B8DFDBE" box="[831,917,582,605]" italics="true" pageId="43" pageNumber="434">Scopula</emphasis>
: All tarsi with indistinctly outlined scopula on roughly distal two-thirds of ventral side, only behind claw divided by a short median stripe; scopula on tarsus I very thin and its proximal limit difficult to identify; scopula on other tarsi denser and and more clearly delimited.
</paragraph>
<paragraph id="C4261544FFCFFFDCB950FD193CB0FCDC" blockId="43.[831,1437,258,1901]" pageId="43" pageNumber="434">
<emphasis id="F6EDC956FFCFFFDCB950FD193B9AFD1C" box="[831,898,744,767]" italics="true" pageId="43" pageNumber="434">Claws</emphasis>
: Paired tarsal claws with 3-4 denticles on anterior legs, 4-5 denticles on posterior legs; unpaired claws with one indistinct denticle or without.
</paragraph>
<paragraph id="C4261544FFCFFFDCB950FCB83CDDF9C9" blockId="43.[831,1437,258,1901]" pageId="43" pageNumber="434">
<emphasis id="F6EDC956FFCFFFDCB950FCB83B69FC83" box="[831,881,841,864]" italics="true" pageId="43" pageNumber="434">Palp</emphasis>
: Tibial apophysis well-developed and quite long, slightly set back from anterior margin of tibia, distinctly pointing away from axis of tibia (
<figureCitation id="5CA209C1FFCFFFDCBEF0FC783CE7FC41" box="[1183,1279,905,930]" captionStart="Fig. 23" captionStartId="44.[147,182,1855,1876]" captionTargetBox="[157,1429,261,1790]" captionTargetId="figure@44.[148,1439,254,1807]" captionTargetPageId="44" captionText="Fig. 23. Liphistius priceae sp. nov., details of palp of six males; holotype (A-D, H-K), male paratype collected mature (E), paratype matured 20. VII. 2004 (F), paratype matured 1. VIII. 2004 (G), paratype matured 18. IX. 2004 (L-M) and paratype matured 11. VIII. 2005 (N-O). (A) Left cymbium and palpal organ, ventral view. (B) Distal part of left tibia, ventral view. (C) Left paracymbium, retrolateral view. (D-E) Left palpal organ, distal view (dorsal side up). (F-G) Dorsal apex of contrategulum of right palpal organ, distal view (dorsal side to the left). (H) Distal part of left palpal organ, prolateral view. (I) Same, retrodorsal and slightly distal view. (J) Distal part of cymbium, prolateral view. (K-L, N) Left tibial apophysis, retrolateral and slightly proximal view. (M, O) Right tibial apophysis, same view. Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A-C, K-O), 0.5 mm (D-I; J)." httpUri="https://zenodo.org/record/1021987/files/figure.png" pageId="43" pageNumber="434">Fig. 23B</figureCitation>
), carrying four very short megaspines, the ventral one being longest, a median one set back from distal margin (
<figureCitation id="5CA209C1FFCFFFDCBF49FC3B3D89FC00" box="[1318,1425,970,995]" captionStart="Fig. 23" captionStartId="44.[147,182,1855,1876]" captionTargetBox="[157,1429,261,1790]" captionTargetId="figure@44.[148,1439,254,1807]" captionTargetPageId="44" captionText="Fig. 23. Liphistius priceae sp. nov., details of palp of six males; holotype (A-D, H-K), male paratype collected mature (E), paratype matured 20. VII. 2004 (F), paratype matured 1. VIII. 2004 (G), paratype matured 18. IX. 2004 (L-M) and paratype matured 11. VIII. 2005 (N-O). (A) Left cymbium and palpal organ, ventral view. (B) Distal part of left tibia, ventral view. (C) Left paracymbium, retrolateral view. (D-E) Left palpal organ, distal view (dorsal side up). (F-G) Dorsal apex of contrategulum of right palpal organ, distal view (dorsal side to the left). (H) Distal part of left palpal organ, prolateral view. (I) Same, retrodorsal and slightly distal view. (J) Distal part of cymbium, prolateral view. (K-L, N) Left tibial apophysis, retrolateral and slightly proximal view. (M, O) Right tibial apophysis, same view. Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A-C, K-O), 0.5 mm (D-I; J)." httpUri="https://zenodo.org/record/1021987/files/figure.png" pageId="43" pageNumber="434">Fig. 23K</figureCitation>
). Distal margin of cymbium with elongate prodorsal lobe (
<figureCitation id="5CA209C1FFCFFFDCB928FBFA3BB8FBC7" box="[839,928,1035,1060]" captionStart="Fig. 23" captionStartId="44.[147,182,1855,1876]" captionTargetBox="[157,1429,261,1790]" captionTargetId="figure@44.[148,1439,254,1807]" captionTargetPageId="44" captionText="Fig. 23. Liphistius priceae sp. nov., details of palp of six males; holotype (A-D, H-K), male paratype collected mature (E), paratype matured 20. VII. 2004 (F), paratype matured 1. VIII. 2004 (G), paratype matured 18. IX. 2004 (L-M) and paratype matured 11. VIII. 2005 (N-O). (A) Left cymbium and palpal organ, ventral view. (B) Distal part of left tibia, ventral view. (C) Left paracymbium, retrolateral view. (D-E) Left palpal organ, distal view (dorsal side up). (F-G) Dorsal apex of contrategulum of right palpal organ, distal view (dorsal side to the left). (H) Distal part of left palpal organ, prolateral view. (I) Same, retrodorsal and slightly distal view. (J) Distal part of cymbium, prolateral view. (K-L, N) Left tibial apophysis, retrolateral and slightly proximal view. (M, O) Right tibial apophysis, same view. Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A-C, K-O), 0.5 mm (D-I; J)." httpUri="https://zenodo.org/record/1021987/files/figure.png" pageId="43" pageNumber="434">Fig. 23J</figureCitation>
). Paracymbium quite large and moderately deep, with a distinct retrolateral-proximal bulge (
<figureCitation id="5CA209C1FFCFFFDCBF62FBDA3D97FBA7" box="[1293,1423,1067,1092]" captionStart="Fig. 23" captionStartId="44.[147,182,1855,1876]" captionTargetBox="[157,1429,261,1790]" captionTargetId="figure@44.[148,1439,254,1807]" captionTargetPageId="44" captionText="Fig. 23. Liphistius priceae sp. nov., details of palp of six males; holotype (A-D, H-K), male paratype collected mature (E), paratype matured 20. VII. 2004 (F), paratype matured 1. VIII. 2004 (G), paratype matured 18. IX. 2004 (L-M) and paratype matured 11. VIII. 2005 (N-O). (A) Left cymbium and palpal organ, ventral view. (B) Distal part of left tibia, ventral view. (C) Left paracymbium, retrolateral view. (D-E) Left palpal organ, distal view (dorsal side up). (F-G) Dorsal apex of contrategulum of right palpal organ, distal view (dorsal side to the left). (H) Distal part of left palpal organ, prolateral view. (I) Same, retrodorsal and slightly distal view. (J) Distal part of cymbium, prolateral view. (K-L, N) Left tibial apophysis, retrolateral and slightly proximal view. (M, O) Right tibial apophysis, same view. Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A-C, K-O), 0.5 mm (D-I; J)." httpUri="https://zenodo.org/record/1021987/files/figure.png" pageId="43" pageNumber="434">Fig. 23A, C</figureCitation>
); cumulus indistinctly elevated, carrying few long stiff bristles (
<figureCitation id="5CA209C1FFCFFFDCB9CFFB9D3C33FB66" box="[928,1067,1132,1157]" captionStart="Fig. 23" captionStartId="44.[147,182,1855,1876]" captionTargetBox="[157,1429,261,1790]" captionTargetId="figure@44.[148,1439,254,1807]" captionTargetPageId="44" captionText="Fig. 23. Liphistius priceae sp. nov., details of palp of six males; holotype (A-D, H-K), male paratype collected mature (E), paratype matured 20. VII. 2004 (F), paratype matured 1. VIII. 2004 (G), paratype matured 18. IX. 2004 (L-M) and paratype matured 11. VIII. 2005 (N-O). (A) Left cymbium and palpal organ, ventral view. (B) Distal part of left tibia, ventral view. (C) Left paracymbium, retrolateral view. (D-E) Left palpal organ, distal view (dorsal side up). (F-G) Dorsal apex of contrategulum of right palpal organ, distal view (dorsal side to the left). (H) Distal part of left palpal organ, prolateral view. (I) Same, retrodorsal and slightly distal view. (J) Distal part of cymbium, prolateral view. (K-L, N) Left tibial apophysis, retrolateral and slightly proximal view. (M, O) Right tibial apophysis, same view. Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A-C, K-O), 0.5 mm (D-I; J)." httpUri="https://zenodo.org/record/1021987/files/figure.png" pageId="43" pageNumber="434">Fig. 23A, C</figureCitation>
). Subtegulum without apophysis. Tegulum with wide, sharp, non-dentate proximal edge (
<figureCitation id="5CA209C1FFCFFFDCB928FB5C3BB8FB25" box="[839,928,1197,1222]" captionStart="Fig. 23" captionStartId="44.[147,182,1855,1876]" captionTargetBox="[157,1429,261,1790]" captionTargetId="figure@44.[148,1439,254,1807]" captionTargetPageId="44" captionText="Fig. 23. Liphistius priceae sp. nov., details of palp of six males; holotype (A-D, H-K), male paratype collected mature (E), paratype matured 20. VII. 2004 (F), paratype matured 1. VIII. 2004 (G), paratype matured 18. IX. 2004 (L-M) and paratype matured 11. VIII. 2005 (N-O). (A) Left cymbium and palpal organ, ventral view. (B) Distal part of left tibia, ventral view. (C) Left paracymbium, retrolateral view. (D-E) Left palpal organ, distal view (dorsal side up). (F-G) Dorsal apex of contrategulum of right palpal organ, distal view (dorsal side to the left). (H) Distal part of left palpal organ, prolateral view. (I) Same, retrodorsal and slightly distal view. (J) Distal part of cymbium, prolateral view. (K-L, N) Left tibial apophysis, retrolateral and slightly proximal view. (M, O) Right tibial apophysis, same view. Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A-C, K-O), 0.5 mm (D-I; J)." httpUri="https://zenodo.org/record/1021987/files/figure.png" pageId="43" pageNumber="434">
Fig. 
<date id="B0273384FFCFFFDCB916FB5F3BB8FB25" box="[889,928,1198,1222]" pageId="43" pageNumber="434" value="1923-01">23I</date>
</figureCitation>
). Contrategulum with somewhat widely conical (its apex narrowly rounded) ventral process; distal edge with long row of oblique parallel ridges pointing towards embolus; dorsal apex asymmetrical, quite small and narrow, with narrowly rounded tip (
<figureCitation id="5CA209C1FFCFFFDCBE8FFADF3D52FAA4" box="[1248,1354,1326,1351]" captionStart="Fig. 23" captionStartId="44.[147,182,1855,1876]" captionTargetBox="[157,1429,261,1790]" captionTargetId="figure@44.[148,1439,254,1807]" captionTargetPageId="44" captionText="Fig. 23. Liphistius priceae sp. nov., details of palp of six males; holotype (A-D, H-K), male paratype collected mature (E), paratype matured 20. VII. 2004 (F), paratype matured 1. VIII. 2004 (G), paratype matured 18. IX. 2004 (L-M) and paratype matured 11. VIII. 2005 (N-O). (A) Left cymbium and palpal organ, ventral view. (B) Distal part of left tibia, ventral view. (C) Left paracymbium, retrolateral view. (D-E) Left palpal organ, distal view (dorsal side up). (F-G) Dorsal apex of contrategulum of right palpal organ, distal view (dorsal side to the left). (H) Distal part of left palpal organ, prolateral view. (I) Same, retrodorsal and slightly distal view. (J) Distal part of cymbium, prolateral view. (K-L, N) Left tibial apophysis, retrolateral and slightly proximal view. (M, O) Right tibial apophysis, same view. Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A-C, K-O), 0.5 mm (D-I; J)." httpUri="https://zenodo.org/record/1021987/files/figure.png" pageId="43" pageNumber="434">Fig. 23D</figureCitation>
). Paraembolic plate low, its distal margin angular (
<figureCitation id="5CA209C1FFCFFFDCBF78FABE3B52FA6B" captionStart="Fig. 23" captionStartId="44.[147,182,1855,1876]" captionTargetBox="[157,1429,261,1790]" captionTargetId="figure@44.[148,1439,254,1807]" captionTargetPageId="44" captionText="Fig. 23. Liphistius priceae sp. nov., details of palp of six males; holotype (A-D, H-K), male paratype collected mature (E), paratype matured 20. VII. 2004 (F), paratype matured 1. VIII. 2004 (G), paratype matured 18. IX. 2004 (L-M) and paratype matured 11. VIII. 2005 (N-O). (A) Left cymbium and palpal organ, ventral view. (B) Distal part of left tibia, ventral view. (C) Left paracymbium, retrolateral view. (D-E) Left palpal organ, distal view (dorsal side up). (F-G) Dorsal apex of contrategulum of right palpal organ, distal view (dorsal side to the left). (H) Distal part of left palpal organ, prolateral view. (I) Same, retrodorsal and slightly distal view. (J) Distal part of cymbium, prolateral view. (K-L, N) Left tibial apophysis, retrolateral and slightly proximal view. (M, O) Right tibial apophysis, same view. Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A-C, K-O), 0.5 mm (D-I; J)." httpUri="https://zenodo.org/record/1021987/files/figure.png" pageId="43" pageNumber="434">Fig. 23A, D, I</figureCitation>
); embolus proper with slightly widened and obliquely truncate apex (
<figureCitation id="5CA209C1FFCFFFDCB98DFA7E3C47FA4B" box="[994,1119,1423,1448]" captionStart="Fig. 23" captionStartId="44.[147,182,1855,1876]" captionTargetBox="[157,1429,261,1790]" captionTargetId="figure@44.[148,1439,254,1807]" captionTargetPageId="44" captionText="Fig. 23. Liphistius priceae sp. nov., details of palp of six males; holotype (A-D, H-K), male paratype collected mature (E), paratype matured 20. VII. 2004 (F), paratype matured 1. VIII. 2004 (G), paratype matured 18. IX. 2004 (L-M) and paratype matured 11. VIII. 2005 (N-O). (A) Left cymbium and palpal organ, ventral view. (B) Distal part of left tibia, ventral view. (C) Left paracymbium, retrolateral view. (D-E) Left palpal organ, distal view (dorsal side up). (F-G) Dorsal apex of contrategulum of right palpal organ, distal view (dorsal side to the left). (H) Distal part of left palpal organ, prolateral view. (I) Same, retrodorsal and slightly distal view. (J) Distal part of cymbium, prolateral view. (K-L, N) Left tibial apophysis, retrolateral and slightly proximal view. (M, O) Right tibial apophysis, same view. Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A-C, K-O), 0.5 mm (D-I; J)." httpUri="https://zenodo.org/record/1021987/files/figure.png" pageId="43" pageNumber="434">Fig. 23A, I</figureCitation>
), dorsal and ventral walls of sclerotised part equally wide and lying close to each other (
<figureCitation id="5CA209C1FFCFFFDCB9E5FA213BEAFA0A" box="[906,1010,1488,1513]" captionStart="Fig. 23" captionStartId="44.[147,182,1855,1876]" captionTargetBox="[157,1429,261,1790]" captionTargetId="figure@44.[148,1439,254,1807]" captionTargetPageId="44" captionText="Fig. 23. Liphistius priceae sp. nov., details of palp of six males; holotype (A-D, H-K), male paratype collected mature (E), paratype matured 20. VII. 2004 (F), paratype matured 1. VIII. 2004 (G), paratype matured 18. IX. 2004 (L-M) and paratype matured 11. VIII. 2005 (N-O). (A) Left cymbium and palpal organ, ventral view. (B) Distal part of left tibia, ventral view. (C) Left paracymbium, retrolateral view. (D-E) Left palpal organ, distal view (dorsal side up). (F-G) Dorsal apex of contrategulum of right palpal organ, distal view (dorsal side to the left). (H) Distal part of left palpal organ, prolateral view. (I) Same, retrodorsal and slightly distal view. (J) Distal part of cymbium, prolateral view. (K-L, N) Left tibial apophysis, retrolateral and slightly proximal view. (M, O) Right tibial apophysis, same view. Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A-C, K-O), 0.5 mm (D-I; J)." httpUri="https://zenodo.org/record/1021987/files/figure.png" pageId="43" pageNumber="434">Fig. 23D</figureCitation>
), retrolateral side enforced by a long, distinct keel (
<figureCitation id="5CA209C1FFCFFFDCB9BDFA013C54F9EA" box="[978,1100,1520,1545]" captionStart="Fig. 23" captionStartId="44.[147,182,1855,1876]" captionTargetBox="[157,1429,261,1790]" captionTargetId="figure@44.[148,1439,254,1807]" captionTargetPageId="44" captionText="Fig. 23. Liphistius priceae sp. nov., details of palp of six males; holotype (A-D, H-K), male paratype collected mature (E), paratype matured 20. VII. 2004 (F), paratype matured 1. VIII. 2004 (G), paratype matured 18. IX. 2004 (L-M) and paratype matured 11. VIII. 2005 (N-O). (A) Left cymbium and palpal organ, ventral view. (B) Distal part of left tibia, ventral view. (C) Left paracymbium, retrolateral view. (D-E) Left palpal organ, distal view (dorsal side up). (F-G) Dorsal apex of contrategulum of right palpal organ, distal view (dorsal side to the left). (H) Distal part of left palpal organ, prolateral view. (I) Same, retrodorsal and slightly distal view. (J) Distal part of cymbium, prolateral view. (K-L, N) Left tibial apophysis, retrolateral and slightly proximal view. (M, O) Right tibial apophysis, same view. Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A-C, K-O), 0.5 mm (D-I; J)." httpUri="https://zenodo.org/record/1021987/files/figure.png" pageId="43" pageNumber="434">Fig. 23D, I</figureCitation>
); membranous part of embolus proper narrow, indistinct (
<figureCitation id="5CA209C1FFCFFFDCBE3AF9E03CA1F9C9" box="[1109,1209,1553,1578]" captionStart="Fig. 23" captionStartId="44.[147,182,1855,1876]" captionTargetBox="[157,1429,261,1790]" captionTargetId="figure@44.[148,1439,254,1807]" captionTargetPageId="44" captionText="Fig. 23. Liphistius priceae sp. nov., details of palp of six males; holotype (A-D, H-K), male paratype collected mature (E), paratype matured 20. VII. 2004 (F), paratype matured 1. VIII. 2004 (G), paratype matured 18. IX. 2004 (L-M) and paratype matured 11. VIII. 2005 (N-O). (A) Left cymbium and palpal organ, ventral view. (B) Distal part of left tibia, ventral view. (C) Left paracymbium, retrolateral view. (D-E) Left palpal organ, distal view (dorsal side up). (F-G) Dorsal apex of contrategulum of right palpal organ, distal view (dorsal side to the left). (H) Distal part of left palpal organ, prolateral view. (I) Same, retrodorsal and slightly distal view. (J) Distal part of cymbium, prolateral view. (K-L, N) Left tibial apophysis, retrolateral and slightly proximal view. (M, O) Right tibial apophysis, same view. Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A-C, K-O), 0.5 mm (D-I; J)." httpUri="https://zenodo.org/record/1021987/files/figure.png" pageId="43" pageNumber="434">Fig. 23H</figureCitation>
).
</paragraph>
<paragraph id="C4261544FFCFFFDCB950F9C23CABF88F" blockId="43.[831,1437,258,1901]" pageId="43" pageNumber="434">
<emphasis id="F6EDC956FFCFFFDCB950F9C23BC2F9A9" box="[831,986,1587,1610]" italics="true" pageId="43" pageNumber="434">Measurements</emphasis>
: Total length 9.96; carapace 4.51 long, 4.24 wide; opisthosoma 3.96 long, 2.61 wide; eye mound 0.64 long, 0.76 wide, AME well-developed; palpal coxae 0.99 long, 0.95 wide; labium 0.40 long, 0.87 wide; sternum 2.02 long, 1.47 wide (0.75 on ventral surface); palp 7.77 long (2.46 + 1.39 + 2.69 + 1.23); leg I 14.69 long (3.96 + 1.82 + 3.17 + 3.64 + 2.10); leg II 15.61 long (4.08 + 1.82 + 3.29 + 4.08 + 2.34); leg III 17.18 long (4.16 + 1.86 + 3.56 + 4.91 + 2.69); leg IV 21.62 long (5.07 + 1.90 + 4.51 + 6.69 + 3.45).
</paragraph>
<paragraph id="C4261544FFCFFFDCB950F8753D29F7FE" blockId="43.[831,1437,1924,2077]" pageId="43" pageNumber="434">
<emphasis id="F6EDC956FFCFFFDCB950F8753CDCF878" bold="true" box="[831,1220,1924,1947]" pageId="43" pageNumber="434">
Description of female (
<typeStatus id="1B22ABE6FFCFFFDCBE3AF8753CAFF878" box="[1109,1207,1924,1947]" pageId="43" pageNumber="434" type="allotype">allotype</typeStatus>
):
</emphasis>
<emphasis id="F6EDC956FFCFFFDCBEBEF8753D85F878" box="[1233,1437,1924,1948]" italics="true" pageId="43" pageNumber="434">Colour in alcohol</emphasis>
(slightly darker in life): Mostly as in male, annulations on tibiae and metatarsi of legs slightly more distinct; palpal tarsus reddish brown only at tip. Opisthosomal tergites generally darker, light patches smaller.
</paragraph>
<caption id="90E645CCFFC8FFDBBAFCF8CE3ADDF7FD" httpUri="https://zenodo.org/record/1021987/files/figure.png" pageId="44" pageNumber="435" targetBox="[157,1429,261,1790]" targetPageId="44">
<paragraph id="C4261544FFC8FFDBBAFCF8CE3ADDF7FD" blockId="44.[147,1436,1855,2079]" pageId="44" pageNumber="435">
Fig. 23. 
<taxonomicName id="03996EC7FFC8FFDBBA86F8CE398FF8B6" authority="Peter J. Schwendinger, 2017" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[233,407,1855,1877]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="435" phylum="Arthropoda" rank="species" species="priceae" status="sp. nov.">
<emphasis id="F6EDC956FFC8FFDBBA86F8CE398FF8B6" box="[233,407,1855,1877]" italics="true" pageId="44" pageNumber="435">Liphistius priceae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDDE742DFFC8FFDBBBF1F8B139F4F8B6" box="[414,492,1856,1877]" pageId="44" pageNumber="435" rank="species">sp. nov.</taxonomicNameLabel>
, details of palp of six males; holotype (A-D, H-K), male paratype collected mature (E), paratype matured 20.VII.2004 (F), paratype matured 1.VIII.2004 (G), paratype matured 18.IX.2004 (L-M) and paratype matured 11.VIII.2005 (N-O). (A) Left cymbium and palpal organ, ventral view. (B) Distal part of left tibia, ventral view. (C) Left paracymbium, retrolateral view. (D-E) Left palpal organ, distal view (dorsal side up). (F-G) Dorsal apex of contrategulum of right palpal organ, distal view (dorsal side to the left). (H) Distal part of left palpal organ, prolateral view. (I) Same, retrodorsal and slightly distal view. (J) Distal part of cymbium, prolateral view. (K-L, N) Left tibial apophysis, retrolateral and slightly proximal view. (M, O) Right tibial apophysis, same view. Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A-C, K-O), 0.5 mm (D-I; J).
</paragraph>
</caption>
<caption id="90E645CCFFC9FFDABAFAFAC43AC8FA66" httpUri="https://zenodo.org/record/1021989/files/figure.png" pageId="45" pageNumber="436" targetBox="[156,1432,263,1274]" targetPageId="45">
<paragraph id="C4261544FFC9FFDABAFAFAC43AC8FA66" blockId="45.[149,1437,1333,1413]" pageId="45" pageNumber="436">
Fig. 24. 
<taxonomicName id="03996EC7FFC9FFDABA84FAC4398FFAA8" authority="Peter J. Schwendinger, 2017" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[235,407,1333,1355]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="45" pageNumber="436" phylum="Arthropoda" rank="species" species="priceae" status="sp. nov.">
<emphasis id="F6EDC956FFC9FFDABA84FAC4398FFAA8" box="[235,407,1333,1355]" italics="true" pageId="45" pageNumber="436">Liphistius priceae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDDE742DFFC9FFDABBF2FAC739F1FAA8" box="[413,489,1334,1355]" pageId="45" pageNumber="436" rank="species">sp. nov.</taxonomicNameLabel>
, vulval plate of four female paratypes (all from exuviae): paratype, moult 1.I.2006 (A-B), allotype, moult 24.XI.2004 (C-D), paratype, moult 13.XI.2005 (E), paratype, moult 2.IV.2005 (F). (A, C, E-F) Dorsal view. (B) Lateral view. (D) Ventral view. Scale lines: 0.5 mm (A-F).
</paragraph>
</caption>
<paragraph id="C4261544FFC9FFDABAF8FA013A2FF988" blockId="45.[151,757,1520,2077]" pageId="45" pageNumber="436">
<emphasis id="F6EDC956FFC9FFDABAF8FA013967F9EA" box="[151,383,1520,1545]" italics="true" pageId="45" pageNumber="436">Bristles on carapace</emphasis>
: Mostly as in male, additionally with very few tiny bristles on anterior coxal elevations. 
<emphasis id="F6EDC956FFC9FFDABAF8F9C23959F9A9" box="[151,321,1587,1610]" italics="true" pageId="45" pageNumber="436">Cheliceral teeth</emphasis>
: Eleven strong teeth on promargin of left cheliceral groove, twelve on right side.
</paragraph>
<paragraph id="C4261544FFC9FFDABAF8F9873950F912" blockId="45.[151,757,1520,2077]" pageId="45" pageNumber="436">
<emphasis id="F6EDC956FFC9FFDABAF8F98738C2F96E" box="[151,218,1654,1677]" italics="true" pageId="45" pageNumber="436">Claws</emphasis>
: Each palpal claw with three worn denticles. Paired leg claws with 2-4 denticles; unpaired claws of legs I-III with 2-3 denticles, leg IV with 0-2. All tarsi without scopula.
</paragraph>
<paragraph id="C4261544FFC9FFDABAF8F90A3B66F82D" blockId="45.[151,757,1520,2077]" lastBlockId="45.[831,1437,1520,1998]" pageId="45" pageNumber="436">
<emphasis id="F6EDC956FFC9FFDABAF8F90A38CBF8F1" box="[151,211,1787,1810]" italics="true" pageId="45" pageNumber="436">Vulva</emphasis>
: Posterior margin of genital sternite widely W-shaped (
<figureCitation id="5CA209C1FFC9FFDABB7CF8ED397BF8D6" box="[275,355,1820,1845]" captionStart="Fig. 24" captionStartId="45.[149,184,1333,1354]" captionTargetBox="[156,1432,263,1274]" captionTargetId="figure@45.[148,1446,254,1296]" captionTargetPageId="45" captionText="Fig. 24. Liphistius priceae sp. nov., vulval plate of four female paratypes (all from exuviae): paratype, moult 1. I. 2006 (A-B), allotype, moult 24. XI. 2004 (C-D), paratype, moult 13. XI. 2005 (E), paratype, moult 2. IV. 2005 (F). (A, C, E-F) Dorsal view. (B) Lateral view. (D) Ventral view. Scale lines: 0.5 mm (A-F)." httpUri="https://zenodo.org/record/1021989/files/figure.png" pageId="45" pageNumber="436">Fig. 24</figureCitation>
C-D), in slightly posteroventral view posterior edge of posterior stalk clearly visible between paramedian lobes. Vulval plate (
<figureCitation id="5CA209C1FFC9FFDABB9AF8AE3A5BF89B" box="[501,579,1887,1912]" captionStart="Fig. 24" captionStartId="45.[149,184,1333,1354]" captionTargetBox="[156,1432,263,1274]" captionTargetId="figure@45.[148,1446,254,1296]" captionTargetPageId="45" captionText="Fig. 24. Liphistius priceae sp. nov., vulval plate of four female paratypes (all from exuviae): paratype, moult 1. I. 2006 (A-B), allotype, moult 24. XI. 2004 (C-D), paratype, moult 13. XI. 2005 (E), paratype, moult 2. IV. 2005 (F). (A, C, E-F) Dorsal view. (B) Lateral view. (D) Ventral view. Scale lines: 0.5 mm (A-F)." httpUri="https://zenodo.org/record/1021989/files/figure.png" pageId="45" pageNumber="436">Fig. 24</figureCitation>
C-D) wider than long; posterior part of poreplate partly unpigmented; CDO distinctly outlined, somewhat quadrangular, longer than wide (
<figureCitation id="5CA209C1FFC9FFDABB77F8323967F83F" box="[280,383,1987,2012]" captionStart="Fig. 24" captionStartId="45.[149,184,1333,1354]" captionTargetBox="[156,1432,263,1274]" captionTargetId="figure@45.[148,1446,254,1296]" captionTargetPageId="45" captionText="Fig. 24. Liphistius priceae sp. nov., vulval plate of four female paratypes (all from exuviae): paratype, moult 1. I. 2006 (A-B), allotype, moult 24. XI. 2004 (C-D), paratype, moult 13. XI. 2005 (E), paratype, moult 2. IV. 2005 (F). (A, C, E-F) Dorsal view. (B) Lateral view. (D) Ventral view. Scale lines: 0.5 mm (A-F)." httpUri="https://zenodo.org/record/1021989/files/figure.png" pageId="45" pageNumber="436">Fig. 24C</figureCitation>
), leading into large and complex receptacular cluster (
<figureCitation id="5CA209C1FFC9FFDABBE0F8153A18F81E" box="[399,512,2020,2045]" captionStart="Fig. 24" captionStartId="45.[149,184,1333,1354]" captionTargetBox="[156,1432,263,1274]" captionTargetId="figure@45.[148,1446,254,1296]" captionTargetPageId="45" captionText="Fig. 24. Liphistius priceae sp. nov., vulval plate of four female paratypes (all from exuviae): paratype, moult 1. I. 2006 (A-B), allotype, moult 24. XI. 2004 (C-D), paratype, moult 13. XI. 2005 (E), paratype, moult 2. IV. 2005 (F). (A, C, E-F) Dorsal view. (B) Lateral view. (D) Ventral view. Scale lines: 0.5 mm (A-F)." httpUri="https://zenodo.org/record/1021989/files/figure.png" pageId="45" pageNumber="436">Fig. 24D</figureCitation>
); the latter strongly protruding beyond slightly arched anterior margin of poreplate (
<figureCitation id="5CA209C1FFC9FFDAB9DDFA013C0CF9EA" box="[946,1044,1520,1545]" captionStart="Fig. 24" captionStartId="45.[149,184,1333,1354]" captionTargetBox="[156,1432,263,1274]" captionTargetId="figure@45.[148,1446,254,1296]" captionTargetPageId="45" captionText="Fig. 24. Liphistius priceae sp. nov., vulval plate of four female paratypes (all from exuviae): paratype, moult 1. I. 2006 (A-B), allotype, moult 24. XI. 2004 (C-D), paratype, moult 13. XI. 2005 (E), paratype, moult 2. IV. 2005 (F). (A, C, E-F) Dorsal view. (B) Lateral view. (D) Ventral view. Scale lines: 0.5 mm (A-F)." httpUri="https://zenodo.org/record/1021989/files/figure.png" pageId="45" pageNumber="436">Fig. 24C</figureCitation>
); posterior stalk strongly pigmented, trapezium-shaped, anteriorly much narrower than posteriorly, indistinctly separated from pigmented areas of poreplate, its posterior portion finely pitted and bent ventrad at right angles (
<figureCitation id="5CA209C1FFC9FFDABE53F9833C85F968" box="[1084,1181,1650,1675]" captionStart="Fig. 24" captionStartId="45.[149,184,1333,1354]" captionTargetBox="[156,1432,263,1274]" captionTargetId="figure@45.[148,1446,254,1296]" captionTargetPageId="45" captionText="Fig. 24. Liphistius priceae sp. nov., vulval plate of four female paratypes (all from exuviae): paratype, moult 1. I. 2006 (A-B), allotype, moult 24. XI. 2004 (C-D), paratype, moult 13. XI. 2005 (E), paratype, moult 2. IV. 2005 (F). (A, C, E-F) Dorsal view. (B) Lateral view. (D) Ventral view. Scale lines: 0.5 mm (A-F)." httpUri="https://zenodo.org/record/1021989/files/figure.png" pageId="45" pageNumber="436">Fig. 24B</figureCitation>
, showing 
<typeStatus id="1B22ABE6FFC9FFDABF66F9823D72F968" box="[1289,1386,1651,1675]" pageId="45" pageNumber="436" type="paratype">paratype</typeStatus>
). 
<emphasis id="F6EDC956FFC9FFDAB950F9653BC2F948" box="[831,986,1684,1707]" italics="true" pageId="45" pageNumber="436">Measurements</emphasis>
: Total length 15.38; carapace 6.18 long, 4.91 wide; opisthosoma 6.49 long, 4.83 wide; eye mound 0.69 long, 0.82 wide; palpal coxae 1.90 long, 1.43 wide; labium 0.63 long, 1.43 wide; sternum 2.85 long, 1.90 wide (1.11 on ventral surface); palp 10.26 long (3.33 + 1.82 + 2.65 + 2.46); leg I 12.91 long (4.12 + 2.06 + 2.81 + 2.61 + 1.31); leg II 13.42 long (4.12 + 2.10 + 2.85 + 2.81 + 1.54); leg III 14.30 long (4.20 + 2.14 + 2.93 + 3.33 + 1.70); leg IV 19.46 long (5.23 + 2.38 + 4.08 + 5.39 + 2.38).
</paragraph>
<paragraph id="C4261544FFC9FFD9B950F8143AF8FBFE" blockId="45.[831,1436,2021,2078]" lastBlockId="46.[151,757,258,1053]" lastPageId="46" lastPageNumber="437" pageId="45" pageNumber="436">
<emphasis id="F6EDC956FFC9FFDAB950F8143BAEF81F" bold="true" box="[831,950,2021,2044]" pageId="45" pageNumber="436">Variation:</emphasis>
Carapace lengths in males (n=9) 4.42- 5.23, carapace width 4.02-4.76; in females with welldeveloped copulatory organs (n=9) 5.54-5.85 and 4.61-5.04, respectively. In a male and in a female one of the two AME is absent, in all other specimens both are distinct. In two males (including the 
<typeStatus id="1B22ABE6FFCAFFD9B8E3FE933AF4FE9A" box="[652,748,354,377]" pageId="46" pageNumber="437" type="holotype">holotype</typeStatus>
) the W-shaped marking behind the eye mound is indiscernible, in the other males it is more or less well-developed. No noteworthy variation in size and density of the tarsal scopula of males was detected. The arrangement of distal megaspines on the tibial apophysis of the male palp is quite variable (
<figureCitation id="5CA209C1FFCAFFD9B8E6FDD23ACFFDDF" box="[649,727,547,572]" captionStart="Fig. 23" captionStartId="44.[147,182,1855,1876]" captionTargetBox="[157,1429,261,1790]" captionTargetId="figure@44.[148,1439,254,1807]" captionTargetPageId="44" captionText="Fig. 23. Liphistius priceae sp. nov., details of palp of six males; holotype (A-D, H-K), male paratype collected mature (E), paratype matured 20. VII. 2004 (F), paratype matured 1. VIII. 2004 (G), paratype matured 18. IX. 2004 (L-M) and paratype matured 11. VIII. 2005 (N-O). (A) Left cymbium and palpal organ, ventral view. (B) Distal part of left tibia, ventral view. (C) Left paracymbium, retrolateral view. (D-E) Left palpal organ, distal view (dorsal side up). (F-G) Dorsal apex of contrategulum of right palpal organ, distal view (dorsal side to the left). (H) Distal part of left palpal organ, prolateral view. (I) Same, retrodorsal and slightly distal view. (J) Distal part of cymbium, prolateral view. (K-L, N) Left tibial apophysis, retrolateral and slightly proximal view. (M, O) Right tibial apophysis, same view. Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A-C, K-O), 0.5 mm (D-I; J)." httpUri="https://zenodo.org/record/1021987/files/figure.png" pageId="46" pageNumber="437">Fig. 23</figureCitation>
K- O). A male 
<typeStatus id="1B22ABE6FFCAFFD9BB74FDB5396FFDBF" box="[283,375,580,604]" pageId="46" pageNumber="437" type="paratype">paratype</typeStatus>
has fewer bristles on the cumulus of both palps than the other males; one of the cumulus bristles of the illustrated 
<typeStatus id="1B22ABE6FFCAFFD9BBCDFD7239E7FD79" box="[418,511,643,666]" pageId="46" pageNumber="437" type="holotype">holotype</typeStatus>
palp runs to the dorsal side of the palpal organ (
<figureCitation id="5CA209C1FFCAFFD9BBC5FD523A29FD5F" box="[426,561,675,700]" captionStart="Fig. 23" captionStartId="44.[147,182,1855,1876]" captionTargetBox="[157,1429,261,1790]" captionTargetId="figure@44.[148,1439,254,1807]" captionTargetPageId="44" captionText="Fig. 23. Liphistius priceae sp. nov., details of palp of six males; holotype (A-D, H-K), male paratype collected mature (E), paratype matured 20. VII. 2004 (F), paratype matured 1. VIII. 2004 (G), paratype matured 18. IX. 2004 (L-M) and paratype matured 11. VIII. 2005 (N-O). (A) Left cymbium and palpal organ, ventral view. (B) Distal part of left tibia, ventral view. (C) Left paracymbium, retrolateral view. (D-E) Left palpal organ, distal view (dorsal side up). (F-G) Dorsal apex of contrategulum of right palpal organ, distal view (dorsal side to the left). (H) Distal part of left palpal organ, prolateral view. (I) Same, retrodorsal and slightly distal view. (J) Distal part of cymbium, prolateral view. (K-L, N) Left tibial apophysis, retrolateral and slightly proximal view. (M, O) Right tibial apophysis, same view. Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A-C, K-O), 0.5 mm (D-I; J)." httpUri="https://zenodo.org/record/1021987/files/figure.png" pageId="46" pageNumber="437">Fig. 23A, C</figureCitation>
). The dorsal apex of the contrategulum is narrowly rounded to pointed (
<figureCitation id="5CA209C1FFCAFFD9BAF0FD1238F5FD1F" box="[159,237,739,764]" captionStart="Fig. 23" captionStartId="44.[147,182,1855,1876]" captionTargetBox="[157,1429,261,1790]" captionTargetId="figure@44.[148,1439,254,1807]" captionTargetPageId="44" captionText="Fig. 23. Liphistius priceae sp. nov., details of palp of six males; holotype (A-D, H-K), male paratype collected mature (E), paratype matured 20. VII. 2004 (F), paratype matured 1. VIII. 2004 (G), paratype matured 18. IX. 2004 (L-M) and paratype matured 11. VIII. 2005 (N-O). (A) Left cymbium and palpal organ, ventral view. (B) Distal part of left tibia, ventral view. (C) Left paracymbium, retrolateral view. (D-E) Left palpal organ, distal view (dorsal side up). (F-G) Dorsal apex of contrategulum of right palpal organ, distal view (dorsal side to the left). (H) Distal part of left palpal organ, prolateral view. (I) Same, retrodorsal and slightly distal view. (J) Distal part of cymbium, prolateral view. (K-L, N) Left tibial apophysis, retrolateral and slightly proximal view. (M, O) Right tibial apophysis, same view. Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A-C, K-O), 0.5 mm (D-I; J)." httpUri="https://zenodo.org/record/1021987/files/figure.png" pageId="46" pageNumber="437">Fig. 23</figureCitation>
D-G). Variation in the shape of the vulval plates is shown in 
<figureCitation id="5CA209C1FFCAFFD9BB4EFCF2396BFCFF" box="[289,371,771,796]" captionStart="Fig. 24" captionStartId="45.[149,184,1333,1354]" captionTargetBox="[156,1432,263,1274]" captionTargetId="figure@45.[148,1446,254,1296]" captionTargetPageId="45" captionText="Fig. 24. Liphistius priceae sp. nov., vulval plate of four female paratypes (all from exuviae): paratype, moult 1. I. 2006 (A-B), allotype, moult 24. XI. 2004 (C-D), paratype, moult 13. XI. 2005 (E), paratype, moult 2. IV. 2005 (F). (A, C, E-F) Dorsal view. (B) Lateral view. (D) Ventral view. Scale lines: 0.5 mm (A-F)." httpUri="https://zenodo.org/record/1021989/files/figure.png" pageId="46" pageNumber="437">Fig. 24</figureCitation>
. The CDO is small to large, wider than long or longer than wide, circular, elliptical or quadrangular (
<figureCitation id="5CA209C1FFCAFFD9BB5AFCB539A4FCBF" box="[309,444,836,860]" captionStart="Fig. 24" captionStartId="45.[149,184,1333,1354]" captionTargetBox="[156,1432,263,1274]" captionTargetId="figure@45.[148,1446,254,1296]" captionTargetPageId="45" captionText="Fig. 24. Liphistius priceae sp. nov., vulval plate of four female paratypes (all from exuviae): paratype, moult 1. I. 2006 (A-B), allotype, moult 24. XI. 2004 (C-D), paratype, moult 13. XI. 2005 (E), paratype, moult 2. IV. 2005 (F). (A, C, E-F) Dorsal view. (B) Lateral view. (D) Ventral view. Scale lines: 0.5 mm (A-F)." httpUri="https://zenodo.org/record/1021989/files/figure.png" pageId="46" pageNumber="437">Fig. 24A, C</figureCitation>
, E-F). The posterior stalk is more or less distinctly separated from the pigmented area of the poreplate. The lateral walls of the genital atrium are developed as narrow folds (
<figureCitation id="5CA209C1FFCAFFD9B828FC553AB4FC5E" box="[583,684,932,957]" captionStart="Fig. 24" captionStartId="45.[149,184,1333,1354]" captionTargetBox="[156,1432,263,1274]" captionTargetId="figure@45.[148,1446,254,1296]" captionTargetPageId="45" captionText="Fig. 24. Liphistius priceae sp. nov., vulval plate of four female paratypes (all from exuviae): paratype, moult 1. I. 2006 (A-B), allotype, moult 24. XI. 2004 (C-D), paratype, moult 13. XI. 2005 (E), paratype, moult 2. IV. 2005 (F). (A, C, E-F) Dorsal view. (B) Lateral view. (D) Ventral view. Scale lines: 0.5 mm (A-F)." httpUri="https://zenodo.org/record/1021989/files/figure.png" pageId="46" pageNumber="437">Fig. 24D</figureCitation>
) or as simple trenches. Three of the females examined have 1-2 lateral hairs on only one side of the genital atrium (
<figureCitation id="5CA209C1FFCAFFD9BAF0FBF538F5FBFE" box="[159,237,1028,1053]" captionStart="Fig. 24" captionStartId="45.[149,184,1333,1354]" captionTargetBox="[156,1432,263,1274]" captionTargetId="figure@45.[148,1446,254,1296]" captionTargetPageId="45" captionText="Fig. 24. Liphistius priceae sp. nov., vulval plate of four female paratypes (all from exuviae): paratype, moult 1. I. 2006 (A-B), allotype, moult 24. XI. 2004 (C-D), paratype, moult 13. XI. 2005 (E), paratype, moult 2. IV. 2005 (F). (A, C, E-F) Dorsal view. (B) Lateral view. (D) Ventral view. Scale lines: 0.5 mm (A-F)." httpUri="https://zenodo.org/record/1021989/files/figure.png" pageId="46" pageNumber="437">Fig. 24</figureCitation>
A-B, E-F), the 
<typeStatus id="1B22ABE6FFCAFFD9BBE3FBF539FAFBF8" box="[396,482,1028,1051]" pageId="46" pageNumber="437" type="allotype">allotype</typeStatus>
has none (
<figureCitation id="5CA209C1FFCAFFD9B839FBF53ABBFBFE" box="[598,675,1028,1053]" captionStart="Fig. 24" captionStartId="45.[149,184,1333,1354]" captionTargetBox="[156,1432,263,1274]" captionTargetId="figure@45.[148,1446,254,1296]" captionTargetPageId="45" captionText="Fig. 24. Liphistius priceae sp. nov., vulval plate of four female paratypes (all from exuviae): paratype, moult 1. I. 2006 (A-B), allotype, moult 24. XI. 2004 (C-D), paratype, moult 13. XI. 2005 (E), paratype, moult 2. IV. 2005 (F). (A, C, E-F) Dorsal view. (B) Lateral view. (D) Ventral view. Scale lines: 0.5 mm (A-F)." httpUri="https://zenodo.org/record/1021989/files/figure.png" pageId="46" pageNumber="437">Fig. 24</figureCitation>
C-D).
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFCAFFD9BAF8FBC2396EFB68" pageId="46" pageNumber="437" type="distribution">
<paragraph id="C4261544FFCAFFD9BAF8FBC2396EFB68" blockId="46.[151,756,1075,1163]" pageId="46" pageNumber="437">
<emphasis id="F6EDC956FFCAFFD9BAF8FBC23936FBA9" bold="true" box="[151,302,1075,1098]" pageId="46" pageNumber="437">Distribution:</emphasis>
Known only from two caves in the same limestone hill (
<figureCitation id="5CA209C1FFCAFFD9BB12FBA539A6FB8F" box="[381,446,1108,1132]" captionStart="Fig. 1" captionStartId="1.[151,186,1392,1413]" captionTargetBox="[153,758,625,1349]" captionTargetId="figure@1.[149,499,895,1170]" captionTargetPageId="1" captionText="Fig. 1. Localities of Liphistius species of the malayanus - group, tioman - group, linang - group and batuenis - group in peninsular Malaysia and southern Thailand (coast of Sumatra omitted): 1 - Gunung Angsi (type locality of L. malayanus); 2 - Templer Park and Gua Anak Takun (L. malayanus, L. batuensis); 3 - Genting Highlands (L. malayanus); 4 - Fraser’s Hill (L. malayanus); 5 - Cameron Highlands (type locality of L. malayanus cameroni); 6 - Sungai Jasin in Endau Rompin National Park (type locality of L. endau); 7 - Gunung Belumut (L. endau); 8 - Gunung Muntahak (L. endau; type locality of L. gracilis sp. nov.); 9 - Sungai Rengit (type locality of L. johore); 10 - Gunung Kajang on Tioman Island (type locality of L. tioman); 11 - Gua Charas in Bukit Charas (type locality of L. panching); 12 - Nusa Camp in Taman Negara (type locality of L. negara sp. nov.); 13 - Jeram Linang Waterfall (type locality of L. linang sp. nov.); 14 - Sankalakhierie Mountains (type locality of L. indra); 15 - Batu Caves (type locality of L. batuensis); 16 - Gua Tempurung (type locality of L. tempurung); 17 - Gua Cicak (L. tempurung); 18 - Gua Keris (type locality of L. priceae sp. nov). Localities with conspecific populations are encircled. Colours distinguish species groups." httpUri="https://zenodo.org/record/1021937/files/figure.png" pageId="46" pageNumber="437">Fig. 1</figureCitation>
, locality 18) in the north of peninsular 
<collectingCountry id="BC8E55D4FFCAFFD9BB62FB85396AFB68" box="[269,370,1140,1163]" name="Malaysia" pageId="46" pageNumber="437">Malaysia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFCAFFD9BAF8FB523A8BF83D" pageId="46" pageNumber="437" type="biology_ecology">
<paragraph id="C4261544FFCAFFD9BAF8FB523907FA99" blockId="46.[151,757,1187,2015]" pageId="46" pageNumber="437">
<emphasis id="F6EDC956FFCAFFD9BAF8FB5238EFFB59" bold="true" box="[151,247,1187,1210]" pageId="46" pageNumber="437">Biology:</emphasis>
The spiders examined were all collected in different zones (from the euphotic entrance area to the aphotic interior) of both caves. No such spiders were found in the adjacent degraded rain forest. Despite apparently being confined to these caves, 
<taxonomicName id="03996EC7FFCAFFD9B8EFFAD43AECFADF" authority="Peter J. Schwendinger, 2017" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[640,756,1317,1340]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="437" phylum="Arthropoda" rank="species" species="priceae" status="sp. nov.">
<emphasis id="F6EDC956FFCAFFD9B8EFFAD43AECFADF" box="[640,756,1317,1340]" italics="true" pageId="46" pageNumber="437">L. priceae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDDE742DFFCAFFD9BAF8FAB438E8FABF" box="[151,240,1349,1372]" pageId="46" pageNumber="437" rank="species">sp. nov.</taxonomicNameLabel>
shows no noteworthy cave adaptations in its morphology.
</paragraph>
<paragraph id="C4261544FFCAFFD9BAF8FA753A8BF83D" blockId="46.[151,757,1187,2015]" pageId="46" pageNumber="437">
Quite short burrows with a single door and relatively long signal lines (no measurements were taken), as usual for cave-dwelling 
<taxonomicName id="03996EC7FFCAFFD9BB07FA3539C9FA38" authorityName="Schiodte" authorityYear="1849" box="[360,465,1476,1499]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="437" phylum="Arthropoda" rank="genus">
<emphasis id="F6EDC956FFCAFFD9BB07FA3539C9FA38" box="[360,465,1476,1499]" italics="true" pageId="46" pageNumber="437">Liphistius</emphasis>
</taxonomicName>
, were built in horizontal or sloping loam at or near the cave entrance, as well as in cracks and holes of the walls at the entrance and in deeper portions of the caves. No sac-like retreats on rock surface, as constructed by some other cave-dwelling 
<taxonomicName id="03996EC7FFCAFFD9BAF8F9953918F998" authorityName="Schiodte" authorityYear="1849" box="[151,256,1636,1659]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="437" phylum="Arthropoda" rank="genus">
<emphasis id="F6EDC956FFCAFFD9BAF8F9953918F998" box="[151,256,1636,1659]" italics="true" pageId="46" pageNumber="437">Liphistius</emphasis>
</taxonomicName>
, were seen. The trapdoor of the largest female was 
<quantity id="0361B8A1FFCAFFD9BAA4F974390FF97E" box="[203,279,1669,1693]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="46" pageNumber="437" unit="cm" value="1.4">1.4 cm</quantity>
long and 
<quantity id="0361B8A1FFCAFFD9BBE4F97439C0F97E" box="[395,472,1669,1693]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="46" pageNumber="437" unit="cm" value="2.4">2.4 cm</quantity>
wide, in penultimate and adult males they were 1.15-1.6 long and 1.55-2.0 wide. Three males were adult when collected in early 
<date id="B0273384FFCAFFD9B8ACF93738C8F91E" pageId="46" pageNumber="437" value="2004-06">June 2004</date>
; four others matured not long afterwards, between late July and late September; the remaining two males matured in mid-July and late August of the following year. Adult females usually moulted twice per year: between March and August and again between September and January. No egg cases were found in the field and no eggs were laid in captivity, which means that the females had not yet mated when collected in early June.
</paragraph>
</subSubSection>
</treatment>
</document>