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<document id="4103DB49A764B88A5CC593B8B374B99C" ID-CLB-Dataset="31806" ID-DOI="10.5281/zenodo.893555" ID-GBIF-Dataset="4a2efe36-c48b-456b-9223-7b5793025a72" ID-ISSN="0035-418" ID-Zenodo-Dep="893555" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1508361162557" checkinUser="plazi" docAuthor="Peter J. Schwendinger" docDate="2017" docId="4C30A452FFC3FFDDBA94FB783B8CF806" docLanguage="en" docName="rsz_124_2_11_Schwendinger.pdf" docOrigin="Revue suisse de Zoologie 124 (2)" docStyle="DocumentStyle:225056F0A23C51DDC06AC27EAEA15D5B.2:RevSuisseZool.2015-2019.journal_article" docStyleId="225056F0A23C51DDC06AC27EAEA15D5B" docStyleName="RevSuisseZool.2015-2019.journal_article" docStyleVersion="2" docTitle="Liphistius tempurung Platnick 1997" docType="treatment" docVersion="9" lastPageNumber="433" masterDocId="B009DC2AFFE4FFF7BA6FFFF13818FFE3" masterDocTitle="A revision of the trapdoor spider genus Liphistius (Mesothelae: Liphistiidae) in peninsular Malaysia; part 1" masterLastPageNumber="445" masterPageNumber="391" pageNumber="430" updateTime="1698469770161" updateUser="ExternalLinkService">
<mods:mods id="AC367E1F40835E0EB608CE3808F5F956" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="808C72B710B8521D4763E9A7A42432EE">
<mods:title id="DB63B232DA7F6C9010B256BC3F957735">A revision of the trapdoor spider genus Liphistius (Mesothelae: Liphistiidae) in peninsular Malaysia; part 1</mods:title>
</mods:titleInfo>
<mods:name id="ADD00AF106F1A07C3365C8AA0E793064" type="personal">
<mods:role id="F4A7E2D19C0C07F0BB3212C66BDFA6CE">
<mods:roleTerm id="4340E9E07AB50AF16F2A590141519AE7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="16632B078444F348C5D10F5EBEA985EA">Peter J. Schwendinger</mods:namePart>
</mods:name>
<mods:typeOfResource id="E1214F81BD8E17F31707DFC9974C3F8B">text</mods:typeOfResource>
<mods:relatedItem id="8972F92383C78C68EED250FF9B215C6E" type="host">
<mods:titleInfo id="16E2CEB66D9BE8E8F461F4682BF1966E">
<mods:title id="A9A835048165529D30D1F419C209F4D9">Revue suisse de Zoologie</mods:title>
</mods:titleInfo>
<mods:part id="6CF4B647FF6D6338A191155F6D413828">
<mods:date id="1B49C9151D71F14FEA6AA77E01D541E7">2017</mods:date>
<mods:detail id="D82AFF16B70C03B2AB8A61966AA0CECD" type="pubDate">
<mods:number id="9688A2F924AA6B6F781EE5CA6CAEDE37">2017-09-30</mods:number>
</mods:detail>
<mods:detail id="FF97748E21159947AA457B1E02328F84" type="volume">
<mods:number id="D20E80F5E3DA44BF598492B9555A1310">124</mods:number>
</mods:detail>
<mods:detail id="FDF60C3E5D07641B88EED8624D8D89B8" type="issue">
<mods:number id="55802102CD0BC5E83AB6E17B94055339">2</mods:number>
</mods:detail>
<mods:extent id="8794A1ACC9AB55E576D708CFEEDB58D0" unit="page">
<mods:start id="3D0AA69F1F65B86A7B08A8EDD3F356DE">391</mods:start>
<mods:end id="9DBC8B5CE4A770A93A2EB1ED1F5B0589">445</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="36EE4C5082B4375B09FD8AD4EB114592">journal article</mods:classification>
<mods:identifier id="38A9BE076E2F80BEDD7A49F4A04223DC" type="CLB-Dataset">31806</mods:identifier>
<mods:identifier id="4419B5CEC78A71F8B38A93057C6D9B5E" type="DOI">10.5281/zenodo.893555</mods:identifier>
<mods:identifier id="90FD731021E2AF33F0D6D4C9EF087A79" type="GBIF-Dataset">4a2efe36-c48b-456b-9223-7b5793025a72</mods:identifier>
<mods:identifier id="04BB77F83D0A7D18348F3380668FB5EA" type="ISSN">0035-418</mods:identifier>
<mods:identifier id="F1B6872195C1A97CD7F5287A5E14C0D8" type="Zenodo-Dep">893555</mods:identifier>
</mods:mods>
<treatment id="4C30A452FFC3FFDDBA94FB783B8CF806" ID-DOI="http://doi.org/10.5281/zenodo.6042384" ID-GBIF-Taxon="135450785" ID-Zenodo-Dep="6042384" LSID="urn:lsid:plazi:treatment:4C30A452FFC3FFDDBA94FB783B8CF806" httpUri="http://treatment.plazi.org/id/4C30A452FFC3FFDDBA94FB783B8CF806" lastPageId="42" lastPageNumber="433" pageId="39" pageNumber="430">
<subSubSection id="8C8346CFFFC3FFD0BA94FB7839E7FB21" pageId="39" pageNumber="430" type="nomenclature">
<paragraph id="C4261544FFC3FFD0BA94FB783A88FB43" blockId="39.[251,656,1161,1218]" box="[251,656,1161,1185]" pageId="39" pageNumber="430">
<heading id="9F6EA228FFC3FFD0BA94FB783A88FB43" bold="true" box="[251,656,1161,1185]" centered="true" fontSize="10" level="3" pageId="39" pageNumber="430" reason="4">
<taxonomicName id="03996EC7FFC3FFD0BA94FB783A88FB43" ID-CoL="72J6X" authority="Platnick, 1997" authorityName="Platnick" authorityYear="1997" box="[251,656,1161,1185]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="430" phylum="Arthropoda" rank="species" species="tempurung">
<emphasis id="F6EDC956FFC3FFD0BA94FB783A88FB43" bold="true" box="[251,656,1161,1185]" pageId="39" pageNumber="430">
<emphasis id="F6EDC956FFC3FFD0BA94FB7839F1FB42" bold="true" box="[251,489,1161,1185]" italics="true" pageId="39" pageNumber="430">Liphistius tempurung</emphasis>
<bibRefCitation id="A00868B5FFC3FFD0BB80FB783A88FB43" author="Platnick N. &amp; Schwendinger P. J. &amp; Steiner H." box="[495,656,1161,1184]" journalOrPublisher="American Museum Novitates" pageId="39" pageNumber="430" pagination="1 - 13" part="3209" refString="Platnick N. I., Schwendinger P. J., Steiner H. 1997. Three new species of the spider genus Liphistius (Araneae, Mesothelae) from Malaysia. American Museum Novitates 3209: 1 - 13." title="Three new species of the spider genus Liphistius (Araneae, Mesothelae) from Malaysia" type="journal article" year="1997">Platnick, 1997</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="C4261544FFC3FFD0BBE3FB5839E7FB21" blockId="39.[251,656,1161,1218]" box="[396,511,1193,1218]" pageId="39" pageNumber="430">
<figureCitation id="5CA209C1FFC3FFD0BBE3FB5839E7FB21" box="[396,511,1193,1218]" captionStart-0="Fig. 21" captionStart-1="Fig. 22" captionStartId-0="40.[151,186,1796,1817]" captionStartId-1="41.[147,182,1789,1810]" captionTargetBox-0="[157,1424,264,1742]" captionTargetBox-1="[153,1429,258,1732]" captionTargetId-0="figure@40.[148,1446,254,1764]" captionTargetId-1="figure@41.[148,1446,254,1752]" captionTargetPageId-0="40" captionTargetPageId-1="41" captionText-0="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." captionText-1="Fig. 22. Liphistius tempurung, details of copulatory organ and genital sternite of six females: Gua Cicak, specimen moulted 8. I. 2005 (A- B, N); Gua Tempurung, exuvia, moult 22. I. 2007 (C-D, M); Gua Tempurung, exuvia, moult 9. XII. 2004 (E-F); Gua Tempurung, exuvia, moult 21. X. 2004 (G-H); medium-sized female from Gua Cicak (I-J); small female from Gua Tempurung (K-L). (A, C, E, G, I, K) Vulval plate, dorsal view. (B, D, F, H) Same, ventral view. (J, L) Receptacular cluster, ventral view. (M-N) Posterior margin of genital sternite, ventral view. Scale lines: 1.0 mm (A-D, I-L; E-H; M-N)." httpUri-0="https://zenodo.org/record/1021981/files/figure.png" httpUri-1="https://zenodo.org/record/1021985/files/figure.png" pageId="39" pageNumber="430">Figs 21-22</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFC3FFD0BAF8FB263AFBFAEA" pageId="39" pageNumber="430" type="reference_group">
<paragraph id="C4261544FFC3FFD0BAF8FB263AFBFAEA" blockId="39.[151,756,1239,1290]" pageId="39" pageNumber="430">
<treatmentCitationGroup id="E489326AFFC3FFD0BAF8FB263AFBFAEA" pageId="39" pageNumber="430">
<taxonomicName id="03996EC7FFC3FFD0BAF8FB263964FAEA" ID-CoL="72J6X" authority="Platnick in Platnick, Schwendinger &amp; Steiner, 1997: 4" authorityName="Platnick in Platnick, Schwendinger &amp; Steiner" authorityPageNumber="4" authorityYear="1997" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="430" phylum="Arthropoda" rank="species" species="tempurung">
<emphasis id="F6EDC956FFC3FFD0BAF8FB263971FB0E" box="[151,361,1239,1261]" italics="true" pageId="39" pageNumber="430">Liphistius tempurung</emphasis>
Platnick in
<treatmentCitation id="45383355FFC3FFD0BB82FB263964FAEA" author="Platnick N. &amp; Schwendinger P. J. &amp; Steiner H." page="4" pageId="39" pageNumber="430" year="1997">
<bibRefCitation id="A00868B5FFC3FFD0BB82FB26397EFAE9" author="Platnick N. &amp; Schwendinger P. J. &amp; Steiner H." journalOrPublisher="American Museum Novitates" pageId="39" pageNumber="430" pagination="1 - 13" part="3209" refString="Platnick N. I., Schwendinger P. J., Steiner H. 1997. Three new species of the spider genus Liphistius (Araneae, Mesothelae) from Malaysia. American Museum Novitates 3209: 1 - 13." title="Three new species of the spider genus Liphistius (Araneae, Mesothelae) from Malaysia" type="journal article" year="1997">Platnick, Schwendinger &amp; Steiner, 1997</bibRefCitation>
: 4
</treatmentCitation>
</taxonomicName>
-6, figs 10-11 (description of female).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFC3FFD0BAF8FAD23AFEF8AF" pageId="39" pageNumber="430" type="materials_examined">
<paragraph id="C4261544FFC3FFD0BAF8FAD23911FA9A" blockId="39.[151,756,1314,1402]" pageId="39" pageNumber="430">
<materialsCitation id="74F11F19FFC3FFD0BAF8FAD23911FA9A" ID-GBIF-Occurrence="1675315686" collectingDate="1996-05-18" collectionCode="AMNH" collectorName="H. Steiner." country="Malaysia" location="Gua Tempurung" pageId="39" pageNumber="430" specimenCount="2" stateProvince="Perak" typeStatus="holotype">
<emphasis id="F6EDC956FFC3FFD0BAF8FAD238C1FAD8" bold="true" box="[151,217,1315,1339]" pageId="39" pageNumber="430">
<typeStatus id="1B22ABE6FFC3FFD0BAF8FAD238CCFAD8" box="[151,212,1315,1339]" pageId="39" pageNumber="430">Type</typeStatus>
:
</emphasis>
<collectionCode id="A2888D81FFC3FFD0BA81FAD2395CFAD8" box="[238,324,1315,1339]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="39" pageNumber="430">AMNH</collectionCode>
; female
<typeStatus id="1B22ABE6FFC3FFD0BBAFFAD33A38FADA" box="[448,544,1314,1337]" pageId="39" pageNumber="430" type="holotype">holotype</typeStatus>
(not examined);
<collectingCountry id="BC8E55D4FFC3FFD0BAF8FAB338E6FABA" box="[151,254,1346,1369]" name="Malaysia" pageId="39" pageNumber="430">Malaysia</collectingCountry>
,
<collectingRegion id="065DDBA6FFC3FFD0BB7BFAB3394DFABA" box="[276,341,1346,1369]" country="Malaysia" name="Perak" pageId="39" pageNumber="430">Perak</collectingRegion>
,
<location id="C146439FFFC3FFD0BB03FAB23A33FAB8" LSID="urn:lsid:plazi:treatment:4C30A452FFC3FFDDBA94FB783B8CF806:C146439FFFC3FFD0BB03FAB23A33FAB8" box="[364,555,1347,1371]" country="Malaysia" name="Gua Tempurung" pageId="39" pageNumber="430" stateProvince="Perak">Gua Tempurung</location>
;
<collectingDate id="A063CA6CFFC3FFD0B82DFAB33AADFABA" box="[578,693,1346,1369]" pageId="39" pageNumber="430" value="1996-05-18">
<date id="B0273384FFC3FFD0B82DFAB33AADFABA" box="[578,693,1346,1369]" pageId="39" pageNumber="430" value="1996-05-18">18.V.1996</date>
</collectingDate>
; leg.
<collectorName id="696C7092FFC3FFD0BAF8FA923911FA9A" box="[151,265,1378,1402]" pageId="39" pageNumber="430">H. Steiner.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="C4261544FFC3FFD0BAF8FA633AFEF8AF" blockId="39.[151,756,1426,1870]" pageId="39" pageNumber="430">
<emphasis id="F6EDC956FFC3FFD0BAF8FA633999FA4A" bold="true" box="[151,385,1426,1449]" pageId="39" pageNumber="430">Material examined:</emphasis>
MHNG, sample MAL-04/17; 3 males (matured
<date id="B0273384FFC3FFD0BB35FA4339BAFA2A" box="[346,418,1458,1481]" pageId="39" pageNumber="430" value="2004-09">25.IX.</date>
,
<date id="B0273384FFC3FFD0BBD8FA433A2DFA2A" box="[439,565,1458,1481]" pageId="39" pageNumber="430" value="2004-09-26">26.IX.2004</date>
,
<date id="B0273384FFC3FFD0B825FA433AD1FA2A" box="[586,713,1458,1481]" pageId="39" pageNumber="430" value="2005-11-13">13.XI.2005</date>
),
<materialsCitation id="74F11F19FFC3FFD0B888FA433A11F96F" ID-GBIF-Occurrence="1675315690" collectingDate="2004-10-21" collectingDateMax="2004-12-09" collectingDateMin="2004-10-21" elevation="90" latitude="4.416111" location="Gua Tempurung" longLatPrecision="21" longitude="101.1875" municipality="Ipoh" pageId="39" pageNumber="430" specimenCount="6" specimenCount-female="6" stateProvince="Perak">
<specimenCount id="D29FDECDFFC3FFD0B888FA4338F3FA0A" pageId="39" pageNumber="430" type="female">6 females</specimenCount>
(moulted
<collectingDate id="A063CA6CFFC3FFD0BB08FA2239C4FA08" box="[359,476,1491,1515]" pageId="39" pageNumber="430" value="2004-10-21">
<date id="B0273384FFC3FFD0BB08FA2239C4FA08" box="[359,476,1491,1515]" pageId="39" pageNumber="430" value="2004-10-21">21.X.2004</date>
</collectingDate>
;
<collectingDate id="A063CA6CFFC3FFD0BB81FA223A70FA08" box="[494,616,1491,1515]" pageId="39" pageNumber="430" value="2004-12-09">
<date id="B0273384FFC3FFD0BB81FA223A70FA08" box="[494,616,1491,1515]" pageId="39" pageNumber="430" value="2004-12-09">9.XII.2004</date>
</collectingDate>
;
<collectingDate id="A063CA6CFFC3FFD0B816FA22395EF9EA" isEnumeration="true" pageId="39" pageNumber="430">
<date id="B0273384FFC3FFD0B816FA223AECFA08" box="[633,756,1491,1515]" pageId="39" pageNumber="430" value="2004-11-19">19.XI.2004</date>
and
<date id="B0273384FFC3FFD0BAB8FA03395EF9EA" box="[215,326,1522,1545]" pageId="39" pageNumber="430" value="2005-04-03">3.IV.2005</date>
</collectingDate>
;
<collectingDate id="A063CA6CFFC3FFD0BB0CFA0239B1F9C8" isEnumeration="true" pageId="39" pageNumber="430">
<date id="B0273384FFC3FFD0BB0CFA0239F1F9E8" box="[355,489,1523,1547]" pageId="39" pageNumber="430" value="2004-12-30">30.XII.2004</date>
and
<date id="B0273384FFC3FFD0B82EFA033AACF9EA" box="[577,692,1522,1545]" pageId="39" pageNumber="430" value="2005-10-17">17.X.2005</date>
and
<date id="B0273384FFC3FFD0BAF8F9E2391CF9C9" box="[151,260,1555,1578]" pageId="39" pageNumber="430" value="2006-05-24">24.V.2006</date>
and
<date id="B0273384FFC3FFD0BB50F9E239B1F9C8" box="[319,425,1555,1579]" pageId="39" pageNumber="430" value="2007-01-22">22.I.2007</date>
</collectingDate>
;
<collectingDate id="A063CA6CFFC3FFD0BBD8F9E238E4F9A9" isEnumeration="true" pageId="39" pageNumber="430">
<date id="B0273384FFC3FFD0BBD8F9E23A17F9C8" box="[439,527,1555,1579]" pageId="39" pageNumber="430" value="2004-09">IX.2004</date>
and
<date id="B0273384FFC3FFD0B825F9E23ADCF9C9" box="[586,708,1555,1578]" pageId="39" pageNumber="430" value="2005-09-13">13.IX.2005</date>
and
<date id="B0273384FFC3FFD0BAF8F9C238E4F9A9" box="[151,252,1587,1610]" pageId="39" pageNumber="430" value="2006-02-08">8.II.2006</date>
</collectingDate>
;
<collectingDate id="A063CA6CFFC3FFD0BB67F9C53A16F9A9" box="[264,526,1587,1612]" isEnumeration="true" pageId="39" pageNumber="430">
<date id="B0273384FFC3FFD0BB67F9C53973F9AF" box="[264,363,1588,1612]" pageId="39" pageNumber="430" value="2004-10-08">8.X.2004</date>
and
<date id="B0273384FFC3FFD0BBCEF9C23A16F9A9" box="[417,526,1587,1610]" pageId="39" pageNumber="430" value="2005-03-01">1.III.2005</date>
</collectingDate>
);
<collectingRegion id="065DDBA6FFC3FFD0B84CF9C23A7BF9A9" box="[547,611,1587,1610]" country="Malaysia" name="Perak" pageId="39" pageNumber="430">Perak</collectingRegion>
,
<quantity id="0361B8A1FFC3FFD0B801F9C23AAAF9A9" box="[622,690,1587,1610]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="39" pageNumber="430" unit="km" value="25.0">25 km</quantity>
south of
<collectingMunicipality id="24428F3EFFC3FFD0BAD3F9A238E8F989" box="[188,240,1619,1642]" pageId="39" pageNumber="430">Ipoh</collectingMunicipality>
,
<location id="C146439FFFC3FFD0BB6DF9A539A3F98F" LSID="urn:lsid:plazi:treatment:4C30A452FFC3FFDDBA94FB783B8CF806:C146439FFFC3FFD0BB6DF9A539A3F98F" box="[258,443,1620,1644]" latitude="4.416111" longLatPrecision="21" longitude="101.1875" municipality="Ipoh" name="Gua Tempurung" pageId="39" pageNumber="430" stateProvince="Perak">Gua Tempurung</location>
,
<geoCoordinate id="A1AD7383FFC3FFD0BBA3F9A23A51F989" box="[460,585,1619,1642]" direction="north" orientation="latitude" pageId="39" pageNumber="430" precision="15" value="4.416111">4°24’58’’N</geoCoordinate>
,
<geoCoordinate id="A1AD7383FFC3FFD0B833F9A23AF7F989" box="[604,751,1619,1642]" direction="east" orientation="longitude" pageId="39" pageNumber="430" precision="15" value="101.1875">101°11’15’’E</geoCoordinate>
,
<elevation id="4FB4F277FFC3FFD0BAF8F98538C8F96F" box="[151,208,1652,1676]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="430" unit="m" value="90.0">
<quantity id="0361B8A1FFC3FFD0BAF8F98538C8F96F" box="[151,208,1652,1676]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="430" unit="m" value="90.0">90 m</quantity>
</elevation>
(limestone cave, dark zone)
</materialsCitation>
;
<materialsCitation id="74F11F19FFC3FFD0B879F9853A7BF92F" ID-GBIF-Occurrence="1675315677" collectingDate="2004-06-22" collectionCode="MHNG" collectorName="P. J. Schwendinger." elevation="80" location="Gua Cicak" municipality="Gunung Lanno" pageId="39" pageNumber="430" specimenCount="1" specimenCount-juvenile="1" stateProvince="Perak">
<collectingDate id="A063CA6CFFC3FFD0B879F9853A8BF96F" box="[534,659,1652,1676]" pageId="39" pageNumber="430" value="2004-06-22">
<date id="B0273384FFC3FFD0B879F9853A8BF96F" box="[534,659,1652,1676]" pageId="39" pageNumber="430" value="2004-06-22">22.VI.2004</date>
</collectingDate>
; leg.
<collectorName id="696C7092FFC3FFD0B8BFF984392FF948" pageId="39" pageNumber="430">P.J. Schwendinger.</collectorName>
–
<collectionCode id="A2888D81FFC3FFD0BB31F96539ABF94F" box="[350,435,1684,1708]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="39" pageNumber="430">MHNG</collectionCode>
, sample 200/01;
<specimenCount id="D29FDECDFFC3FFD0B814F9653AE8F948" box="[635,752,1684,1708]" pageId="39" pageNumber="430" type="juvenile">1 juvenile</specimenCount>
;
<collectingRegion id="065DDBA6FFC3FFD0BAF8F94538CFF928" box="[151,215,1716,1739]" country="Malaysia" name="Perak" pageId="39" pageNumber="430">Perak</collectingRegion>
,
<collectingMunicipality id="24428F3EFFC3FFD0BA89F9443996F92F" box="[230,398,1717,1741]" pageId="39" pageNumber="430">Gunung Lanno</collectingMunicipality>
,
<location id="C146439FFFC3FFD0BBF2F9443A0EF928" LSID="urn:lsid:plazi:treatment:4C30A452FFC3FFDDBA94FB783B8CF806:C146439FFFC3FFD0BBF2F9443A0EF928" box="[413,534,1716,1741]" municipality="Gunung Lanno" name="Gua Cicak" pageId="39" pageNumber="430" stateProvince="Perak">Gua Cicak</location>
,
<elevation id="4FB4F277FFC3FFD0B84AF9443A7BF92F" box="[549,611,1717,1741]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="39" pageNumber="430" unit="m" value="80.0">
<quantity id="0361B8A1FFC3FFD0B84AF9443A7BF92F" box="[549,611,1717,1741]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="39" pageNumber="430" unit="m" value="80.0">80 m</quantity>
</elevation>
</materialsCitation>
;
<date id="B0273384FFC3FFD0B81DF9453AE8F928" box="[626,752,1716,1739]" pageId="39" pageNumber="430" value="2001-11-16">16.XI.2001</date>
; leg. H. Steiner. – MHNG, sample MAL-04/16; 3 males (matured
<date id="B0273384FFC3FFD0BB25F90439D6F8EE" box="[330,462,1781,1805]" pageId="39" pageNumber="430" value="2004-12-31">31.XII.2004</date>
,
<date id="B0273384FFC3FFD0BBB2F9053A5DF8E8" box="[477,581,1780,1803]" pageId="39" pageNumber="430" value="2005-01-23">23.I.2005</date>
,
<date id="B0273384FFC3FFD0B83BF9053AC8F8E8" box="[596,720,1780,1803]" pageId="39" pageNumber="430" value="2005-06-17">17.VI.2005</date>
),
<materialsCitation id="74F11F19FFC3FFD0B888F9043946F8AF" ID-GBIF-Occurrence="1675315676" collectingDate="2005-01-08" collectingDateMax="2005-01-17" collectingDateMin="2005-01-08" location="Gua Cicak" municipality="Gunung Lanno" pageId="39" pageNumber="430" specimenCount="3" specimenCount-female="3" stateProvince="Perak">
<specimenCount id="D29FDECDFFC3FFD0B888F90438F2F8CF" pageId="39" pageNumber="430" type="female">3 females</specimenCount>
(moulted
<collectingDate id="A063CA6CFFC3FFD0BB0EF8E439A6F8CF" box="[353,446,1813,1836]" pageId="39" pageNumber="430" value="2005-01-08">
<date id="B0273384FFC3FFD0BB0EF8E439A6F8CF" box="[353,446,1813,1836]" pageId="39" pageNumber="430" value="2005-01-08">8.I.2005</date>
</collectingDate>
;
<collectingDate id="A063CA6CFFC3FFD0BBA2F8E43A2FF8CF" box="[461,567,1813,1836]" pageId="39" pageNumber="430" value="2005-01-17">
<date id="B0273384FFC3FFD0BBA2F8E43A2FF8CF" box="[461,567,1813,1836]" pageId="39" pageNumber="430" value="2005-01-17">17.I.2005</date>
</collectingDate>
);
<collectingRegion id="065DDBA6FFC3FFD0B820F8E43A97F8CF" box="[591,655,1813,1836]" country="Malaysia" name="Perak" pageId="39" pageNumber="430">Perak</collectingRegion>
,
<collectingMunicipality id="24428F3EFFC3FFD0B8F1F8E438C6F8AE" pageId="39" pageNumber="430">Gunung Lanno</collectingMunicipality>
,
<location id="C146439FFFC3FFD0BA86F8C73946F8AF" LSID="urn:lsid:plazi:treatment:4C30A452FFC3FFDDBA94FB783B8CF806:C146439FFFC3FFD0BA86F8C73946F8AF" box="[233,350,1845,1870]" municipality="Gunung Lanno" name="Gua Cicak" pageId="39" pageNumber="430" stateProvince="Perak">Gua Cicak</location>
</materialsCitation>
;
<date id="B0273384FFC3FFD0BB07F8C739FDF8AD" box="[360,485,1846,1870]" pageId="39" pageNumber="430" value="2004-06-21">21.VI.2004</date>
; leg. P.J. Schwendinger.
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFC3FFD0BAF8F8953B63FC61" pageId="39" pageNumber="430" type="diagnosis">
<paragraph id="C4261544FFC3FFD0BAF8F8953B63FC61" blockId="39.[151,756,1892,2078]" lastBlockId="39.[831,1437,258,898]" pageId="39" pageNumber="430">
<emphasis id="F6EDC956FFC3FFD0BAF8F895390AF898" bold="true" box="[151,274,1892,1915]" pageId="39" pageNumber="430">Diagnosis:</emphasis>
Medium-sized, light brown-coloured species in both sexes. Similar to
<emphasis id="F6EDC956FFC3FFD0B875F8773AB1F87F" box="[538,681,1925,1949]" italics="true" pageId="39" pageNumber="430">
<taxonomicName id="03996EC7FFC3FFD0B875F8773ABCF87F" authorityName="Abraham" authorityYear="1923" box="[538,676,1925,1949]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="430" phylum="Arthropoda" rank="species" species="batuensis">L. batuensis</taxonomicName>
,
</emphasis>
males distinguished by much shorter and stouter megaspines on tibial apophysis of palp (
<figureCitation id="5CA209C1FFC3FFD0BBBBF8343A3AF83D" box="[468,546,1989,2014]" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="39" pageNumber="430">Fig. 21</figureCitation>
E-G cf.
<figureCitation id="5CA209C1FFC3FFD0B8EEF8343AFFF83D" box="[641,743,1989,2014]" captionStart="Fig. 19" captionStartId="37.[151,186,1883,1904]" captionTargetBox="[158,1437,494,1820]" captionTargetId="figure@37.[148,1448,467,1845]" captionTargetPageId="37" captionText="Fig. 19. Liphistius batuensis, details of left palp of two males from type locality: specimen collected 24. VII. 1969 (A-B, F, I), specimen matured 8. II. 2002 (C-E, G-H). (A) Distal part of palp (palpal organ not indicated), ventral view. (B) Paracymbium, retrolateral view. (C) Distal part of cymbium, prolateral view. (D) Palpal organ, distal view (dorsal side up; arrow indicating proventral angle at base of para-embolic plate). (E) Palpal organ, proventral view (arrow indicating same structure as in D). (F) Embolus complex, proventral view. (G) Palpal organ, retroventral and slightly proximal view (arrow indicating same structure as in D-E). (H) Same, retrodorsal view. (I) Same, prolateral view. Abbreviation: T - tegulum. Scale lines: 1.0 mm (A, C; B) and 0.5 mm (D-I)." httpUri="https://zenodo.org/record/1021977/files/figure.png" pageId="39" pageNumber="430">Fig. 19A</figureCitation>
); proximal edge of tegulum without denticles (
<figureCitation id="5CA209C1FFC3FFD0B8FEF8143AECF81D" box="[657,756,2021,2046]" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="39" pageNumber="430">Fig. 21B</figureCitation>
cf.
<figureCitation id="5CA209C1FFC3FFD0BAABF7F7392AF7FD" box="[196,306,2054,2078]" captionStart="Fig. 19" captionStartId="37.[151,186,1883,1904]" captionTargetBox="[158,1437,494,1820]" captionTargetId="figure@37.[148,1448,467,1845]" captionTargetPageId="37" captionText="Fig. 19. Liphistius batuensis, details of left palp of two males from type locality: specimen collected 24. VII. 1969 (A-B, F, I), specimen matured 8. II. 2002 (C-E, G-H). (A) Distal part of palp (palpal organ not indicated), ventral view. (B) Paracymbium, retrolateral view. (C) Distal part of cymbium, prolateral view. (D) Palpal organ, distal view (dorsal side up; arrow indicating proventral angle at base of para-embolic plate). (E) Palpal organ, proventral view (arrow indicating same structure as in D). (F) Embolus complex, proventral view. (G) Palpal organ, retroventral and slightly proximal view (arrow indicating same structure as in D-E). (H) Same, retrodorsal view. (I) Same, prolateral view. Abbreviation: T - tegulum. Scale lines: 1.0 mm (A, C; B) and 0.5 mm (D-I)." httpUri="https://zenodo.org/record/1021977/files/figure.png" pageId="39" pageNumber="430">Fig. 19H</figureCitation>
); retrolateral bridge between tegulum and contrategulum unbroken (
<figureCitation id="5CA209C1FFC3FFD0BEFEFEF33CEDFEF8" box="[1169,1269,258,283]" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="39" pageNumber="430">Fig. 21B</figureCitation>
cf.
<figureCitation id="5CA209C1FFC3FFD0BF47FEF33D97FEF8" box="[1320,1423,258,283]" captionStart="Fig. 19" captionStartId="37.[151,186,1883,1904]" captionTargetBox="[158,1437,494,1820]" captionTargetId="figure@37.[148,1448,467,1845]" captionTargetPageId="37" captionText="Fig. 19. Liphistius batuensis, details of left palp of two males from type locality: specimen collected 24. VII. 1969 (A-B, F, I), specimen matured 8. II. 2002 (C-E, G-H). (A) Distal part of palp (palpal organ not indicated), ventral view. (B) Paracymbium, retrolateral view. (C) Distal part of cymbium, prolateral view. (D) Palpal organ, distal view (dorsal side up; arrow indicating proventral angle at base of para-embolic plate). (E) Palpal organ, proventral view (arrow indicating same structure as in D). (F) Embolus complex, proventral view. (G) Palpal organ, retroventral and slightly proximal view (arrow indicating same structure as in D-E). (H) Same, retrodorsal view. (I) Same, prolateral view. Abbreviation: T - tegulum. Scale lines: 1.0 mm (A, C; B) and 0.5 mm (D-I)." httpUri="https://zenodo.org/record/1021977/files/figure.png" pageId="39" pageNumber="430">Fig. 19H</figureCitation>
); contrategulum with more pronounced, rather conical ventral process; distal edge with several short parallel ridges situated near base of ventral process and directed towards embolus; dorsal apex of contrategulum wider, without ridges (
<figureCitation id="5CA209C1FFC3FFD0B991FE553C4CFE5E" box="[1022,1108,420,445]" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="39" pageNumber="430">Fig. 21</figureCitation>
I-L cf.
<figureCitation id="5CA209C1FFC3FFD0BEDAFE553D3BFE5E" box="[1205,1315,420,445]" captionStart="Fig. 19" captionStartId="37.[151,186,1883,1904]" captionTargetBox="[158,1437,494,1820]" captionTargetId="figure@37.[148,1448,467,1845]" captionTargetPageId="37" captionText="Fig. 19. Liphistius batuensis, details of left palp of two males from type locality: specimen collected 24. VII. 1969 (A-B, F, I), specimen matured 8. II. 2002 (C-E, G-H). (A) Distal part of palp (palpal organ not indicated), ventral view. (B) Paracymbium, retrolateral view. (C) Distal part of cymbium, prolateral view. (D) Palpal organ, distal view (dorsal side up; arrow indicating proventral angle at base of para-embolic plate). (E) Palpal organ, proventral view (arrow indicating same structure as in D). (F) Embolus complex, proventral view. (G) Palpal organ, retroventral and slightly proximal view (arrow indicating same structure as in D-E). (H) Same, retrodorsal view. (I) Same, prolateral view. Abbreviation: T - tegulum. Scale lines: 1.0 mm (A, C; B) and 0.5 mm (D-I)." httpUri="https://zenodo.org/record/1021977/files/figure.png" pageId="39" pageNumber="430">Fig. 19D</figureCitation>
); embolus proper more slender (
<figureCitation id="5CA209C1FFC3FFD0BE2FFE353C8CFE3E" box="[1088,1172,452,477]" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="39" pageNumber="430">Fig. 21</figureCitation>
A-B cf.
<figureCitation id="5CA209C1FFC3FFD0BE91FE353D97FE3E" box="[1278,1423,452,477]" captionStart="Fig. 19" captionStartId="37.[151,186,1883,1904]" captionTargetBox="[158,1437,494,1820]" captionTargetId="figure@37.[148,1448,467,1845]" captionTargetPageId="37" captionText="Fig. 19. Liphistius batuensis, details of left palp of two males from type locality: specimen collected 24. VII. 1969 (A-B, F, I), specimen matured 8. II. 2002 (C-E, G-H). (A) Distal part of palp (palpal organ not indicated), ventral view. (B) Paracymbium, retrolateral view. (C) Distal part of cymbium, prolateral view. (D) Palpal organ, distal view (dorsal side up; arrow indicating proventral angle at base of para-embolic plate). (E) Palpal organ, proventral view (arrow indicating same structure as in D). (F) Embolus complex, proventral view. (G) Palpal organ, retroventral and slightly proximal view (arrow indicating same structure as in D-E). (H) Same, retrodorsal view. (I) Same, prolateral view. Abbreviation: T - tegulum. Scale lines: 1.0 mm (A, C; B) and 0.5 mm (D-I)." httpUri="https://zenodo.org/record/1021977/files/figure.png" pageId="39" pageNumber="430">Fig. 19E, H</figureCitation>
), dorsal wall of its sclerotised part without sharply bent end overlapping membranous part (
<figureCitation id="5CA209C1FFC3FFD0BEB7FDF43D58FDFD" box="[1240,1344,517,542]" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="39" pageNumber="430">Fig. 21C</figureCitation>
, I-L cf.
<figureCitation id="5CA209C1FFC3FFD0B950FDD43B93FDDD" box="[831,907,549,574]" captionStart="Fig. 19" captionStartId="37.[151,186,1883,1904]" captionTargetBox="[158,1437,494,1820]" captionTargetId="figure@37.[148,1448,467,1845]" captionTargetPageId="37" captionText="Fig. 19. Liphistius batuensis, details of left palp of two males from type locality: specimen collected 24. VII. 1969 (A-B, F, I), specimen matured 8. II. 2002 (C-E, G-H). (A) Distal part of palp (palpal organ not indicated), ventral view. (B) Paracymbium, retrolateral view. (C) Distal part of cymbium, prolateral view. (D) Palpal organ, distal view (dorsal side up; arrow indicating proventral angle at base of para-embolic plate). (E) Palpal organ, proventral view (arrow indicating same structure as in D). (F) Embolus complex, proventral view. (G) Palpal organ, retroventral and slightly proximal view (arrow indicating same structure as in D-E). (H) Same, retrodorsal view. (I) Same, prolateral view. Abbreviation: T - tegulum. Scale lines: 1.0 mm (A, C; B) and 0.5 mm (D-I)." httpUri="https://zenodo.org/record/1021977/files/figure.png" pageId="39" pageNumber="430">Fig. 19</figureCitation>
D-F, I); base of embolus complex without sharp proventral angle (
<figureCitation id="5CA209C1FFC3FFD0BE69FDB73C4DFDBD" box="[1030,1109,582,606]" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="39" pageNumber="430">Fig. 21</figureCitation>
I-L cf.
<figureCitation id="5CA209C1FFC3FFD0BEC6FDB73CEFFDBD" box="[1193,1271,582,606]" captionStart="Fig. 19" captionStartId="37.[151,186,1883,1904]" captionTargetBox="[158,1437,494,1820]" captionTargetId="figure@37.[148,1448,467,1845]" captionTargetPageId="37" captionText="Fig. 19. Liphistius batuensis, details of left palp of two males from type locality: specimen collected 24. VII. 1969 (A-B, F, I), specimen matured 8. II. 2002 (C-E, G-H). (A) Distal part of palp (palpal organ not indicated), ventral view. (B) Paracymbium, retrolateral view. (C) Distal part of cymbium, prolateral view. (D) Palpal organ, distal view (dorsal side up; arrow indicating proventral angle at base of para-embolic plate). (E) Palpal organ, proventral view (arrow indicating same structure as in D). (F) Embolus complex, proventral view. (G) Palpal organ, retroventral and slightly proximal view (arrow indicating same structure as in D-E). (H) Same, retrodorsal view. (I) Same, prolateral view. Abbreviation: T - tegulum. Scale lines: 1.0 mm (A, C; B) and 0.5 mm (D-I)." httpUri="https://zenodo.org/record/1021977/files/figure.png" pageId="39" pageNumber="430">Fig. 19</figureCitation>
D-E, G), paraembolic plate less elevated (
<figureCitation id="5CA209C1FFC3FFD0BEE5FD973CE8FD9C" box="[1162,1264,614,639]" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="39" pageNumber="430">Fig. 21B</figureCitation>
cf.
<figureCitation id="5CA209C1FFC3FFD0BF49FD973D88FD9C" box="[1318,1424,614,639]" captionStart="Fig. 19" captionStartId="37.[151,186,1883,1904]" captionTargetBox="[158,1437,494,1820]" captionTargetId="figure@37.[148,1448,467,1845]" captionTargetPageId="37" captionText="Fig. 19. Liphistius batuensis, details of left palp of two males from type locality: specimen collected 24. VII. 1969 (A-B, F, I), specimen matured 8. II. 2002 (C-E, G-H). (A) Distal part of palp (palpal organ not indicated), ventral view. (B) Paracymbium, retrolateral view. (C) Distal part of cymbium, prolateral view. (D) Palpal organ, distal view (dorsal side up; arrow indicating proventral angle at base of para-embolic plate). (E) Palpal organ, proventral view (arrow indicating same structure as in D). (F) Embolus complex, proventral view. (G) Palpal organ, retroventral and slightly proximal view (arrow indicating same structure as in D-E). (H) Same, retrodorsal view. (I) Same, prolateral view. Abbreviation: T - tegulum. Scale lines: 1.0 mm (A, C; B) and 0.5 mm (D-I)." httpUri="https://zenodo.org/record/1021977/files/figure.png" pageId="39" pageNumber="430">Fig. 19H</figureCitation>
). Females distinguished from those of
<taxonomicName id="03996EC7FFC3FFD0BF62FD793D85FD7E" authorityName="Abraham" authorityYear="1923" box="[1293,1437,646,671]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="430" phylum="Arthropoda" rank="species" species="batuensis">
<emphasis id="F6EDC956FFC3FFD0BF62FD793D85FD7E" box="[1293,1437,646,671]" italics="true" pageId="39" pageNumber="430">L. batuensis</emphasis>
</taxonomicName>
by anterior and posterior margins of vulval plate distinctly invaginated (
<figureCitation id="5CA209C1FFC3FFD0BE2BFD363C8FFD03" box="[1092,1175,711,736]" captionStart="Fig. 22" captionStartId="41.[147,182,1789,1810]" captionTargetBox="[153,1429,258,1732]" captionTargetId="figure@41.[148,1446,254,1752]" captionTargetPageId="41" captionText="Fig. 22. Liphistius tempurung, details of copulatory organ and genital sternite of six females: Gua Cicak, specimen moulted 8. I. 2005 (A- B, N); Gua Tempurung, exuvia, moult 22. I. 2007 (C-D, M); Gua Tempurung, exuvia, moult 9. XII. 2004 (E-F); Gua Tempurung, exuvia, moult 21. X. 2004 (G-H); medium-sized female from Gua Cicak (I-J); small female from Gua Tempurung (K-L). (A, C, E, G, I, K) Vulval plate, dorsal view. (B, D, F, H) Same, ventral view. (J, L) Receptacular cluster, ventral view. (M-N) Posterior margin of genital sternite, ventral view. Scale lines: 1.0 mm (A-D, I-L; E-H; M-N)." httpUri="https://zenodo.org/record/1021985/files/figure.png" pageId="39" pageNumber="430">Fig. 22</figureCitation>
cf.
<figureCitation id="5CA209C1FFC3FFD0BEA2FD363D3BFD03" box="[1229,1315,711,736]" captionStart="Fig. 20" captionStartId="38.[151,186,1993,2014]" captionTargetBox="[155,1432,578,1928]" captionTargetId="figure@38.[148,1439,556,1950]" captionTargetPageId="38" captionText="Fig. 20. Liphistius batuensis, vulval plate of five females from the type locality: exuvia, moult 24. I. 2002 (A-B); specimen SMF 13907 (C); exuvia, moult 12. VII. 2001 (D-E); specimen leg. XI. 1976 (F); exuvia, moult 13. I. 2002 (G-H). (A, C-D, F-G) Dorsal view. (B, E, H) Ventral view. Scale lines: 1.0 mm (A-B, D-E, G-H; C, F)." httpUri="https://zenodo.org/record/1021979/files/figure.png" pageId="39" pageNumber="430">Fig. 20</figureCitation>
); posterior part of genital atrium more strongly curved ventrad, clearly visible on female specimen when viewed from posteroventrally (
<figureCitation id="5CA209C1FFC3FFD0BE69FCD93C44FCA2" box="[1030,1116,808,833]" captionStart="Fig. 22" captionStartId="41.[147,182,1789,1810]" captionTargetBox="[153,1429,258,1732]" captionTargetId="figure@41.[148,1446,254,1752]" captionTargetPageId="41" captionText="Fig. 22. Liphistius tempurung, details of copulatory organ and genital sternite of six females: Gua Cicak, specimen moulted 8. I. 2005 (A- B, N); Gua Tempurung, exuvia, moult 22. I. 2007 (C-D, M); Gua Tempurung, exuvia, moult 9. XII. 2004 (E-F); Gua Tempurung, exuvia, moult 21. X. 2004 (G-H); medium-sized female from Gua Cicak (I-J); small female from Gua Tempurung (K-L). (A, C, E, G, I, K) Vulval plate, dorsal view. (B, D, F, H) Same, ventral view. (J, L) Receptacular cluster, ventral view. (M-N) Posterior margin of genital sternite, ventral view. Scale lines: 1.0 mm (A-D, I-L; E-H; M-N)." httpUri="https://zenodo.org/record/1021985/files/figure.png" pageId="39" pageNumber="430">Fig. 22</figureCitation>
M-N); poreplate with much smaller CDO (
<figureCitation id="5CA209C1FFC3FFD0B9B3FCB83CC9FC82" box="[988,1233,841,865]" captionStart="Fig. 22" captionStartId="41.[147,182,1789,1810]" captionTargetBox="[153,1429,258,1732]" captionTargetId="figure@41.[148,1446,254,1752]" captionTargetPageId="41" captionText="Fig. 22. Liphistius tempurung, details of copulatory organ and genital sternite of six females: Gua Cicak, specimen moulted 8. I. 2005 (A- B, N); Gua Tempurung, exuvia, moult 22. I. 2007 (C-D, M); Gua Tempurung, exuvia, moult 9. XII. 2004 (E-F); Gua Tempurung, exuvia, moult 21. X. 2004 (G-H); medium-sized female from Gua Cicak (I-J); small female from Gua Tempurung (K-L). (A, C, E, G, I, K) Vulval plate, dorsal view. (B, D, F, H) Same, ventral view. (J, L) Receptacular cluster, ventral view. (M-N) Posterior margin of genital sternite, ventral view. Scale lines: 1.0 mm (A-D, I-L; E-H; M-N)." httpUri="https://zenodo.org/record/1021985/files/figure.png" pageId="39" pageNumber="430">Fig. 22A, C, E, G, I, K</figureCitation>
cf.
<figureCitation id="5CA209C1FFC3FFD0BE94FCB83D44FC82" box="[1275,1372,841,865]" captionStart="Fig. 20" captionStartId="38.[151,186,1993,2014]" captionTargetBox="[155,1432,578,1928]" captionTargetId="figure@38.[148,1439,556,1950]" captionTargetPageId="38" captionText="Fig. 20. Liphistius batuensis, vulval plate of five females from the type locality: exuvia, moult 24. I. 2002 (A-B); specimen SMF 13907 (C); exuvia, moult 12. VII. 2001 (D-E); specimen leg. XI. 1976 (F); exuvia, moult 13. I. 2002 (G-H). (A, C-D, F-G) Dorsal view. (B, E, H) Ventral view. Scale lines: 1.0 mm (A-B, D-E, G-H; C, F)." httpUri="https://zenodo.org/record/1021979/files/figure.png" pageId="39" pageNumber="430">Fig. 20A</figureCitation>
, C-D, F-G).
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFC3FFDDB950FC693C95FDFC" lastPageId="42" lastPageNumber="433" pageId="39" pageNumber="430" type="description">
<paragraph id="C4261544FFC3FFD0B950FC693CC8FBD2" blockId="39.[831,1437,920,2079]" pageId="39" pageNumber="430">
<emphasis id="F6EDC956FFC3FFD0B950FC693D32FC4C" bold="true" box="[831,1322,920,945]" pageId="39" pageNumber="430">
Description of male (matured
<date id="B0273384FFC3FFD0BEF1FC693D06FC4C" box="[1182,1310,920,943]" pageId="39" pageNumber="430" value="2004-09-26">26.IX.2004</date>
):
</emphasis>
<emphasis id="F6EDC956FFC3FFD0BF5CFC693B88FC33" italics="true" pageId="39" pageNumber="430">Colour in alcohol</emphasis>
(slightly darker in life): Sclerotised parts light brown, except for cream-coloured proximal portion of chelicerae, whitish membranes of prosoma and creamcoloured membranes of opisthosoma.
</paragraph>
<paragraph id="C4261544FFC3FFD0B950FBCB3CFFFB51" blockId="39.[831,1437,920,2079]" pageId="39" pageNumber="430">
<emphasis id="F6EDC956FFC3FFD0B950FBCB3C3FFBB0" box="[831,1063,1082,1107]" italics="true" pageId="39" pageNumber="430">Bristles on carapace</emphasis>
: Short bristles along all margins (longest and strongest in front of eye mound); few on coxal elevations; four short bristles anterior to fovea; thick (spinelike) bristles on eye mound.
</paragraph>
<paragraph id="C4261544FFC3FFD0B950FB4D3C72FAD6" blockId="39.[831,1437,920,2079]" pageId="39" pageNumber="430">
<emphasis id="F6EDC956FFC3FFD0B950FB4D3B8DFB30" box="[831,917,1212,1235]" italics="true" pageId="39" pageNumber="430">Scopula</emphasis>
: Thin (weakest and narrowest on tarsus IV) and only distally divided by median stripe on all leg tarsi; covering distal half of ventral side of tarsi I-III, only distal one-third of tarsus IV.
</paragraph>
<paragraph id="C4261544FFC3FFD0B950FACC3BE7FA74" blockId="39.[831,1437,920,2079]" pageId="39" pageNumber="430">
<emphasis id="F6EDC956FFC3FFD0B950FACC3BE8FAB7" box="[831,1008,1341,1364]" italics="true" pageId="39" pageNumber="430">Cheliceral teeth</emphasis>
: Eleven small teeth of different sizes on promargin of right cheliceral groove, twelve on left cheliceral groove.
</paragraph>
<paragraph id="C4261544FFC3FFDDB950FA6F3A03FE18" blockId="39.[831,1437,920,2079]" lastBlockId="42.[151,756,258,828]" lastPageId="42" lastPageNumber="433" pageId="39" pageNumber="430">
<emphasis id="F6EDC956FFC3FFD0B950FA6F3B69FA56" box="[831,881,1438,1461]" italics="true" pageId="39" pageNumber="430">Palp</emphasis>
(illustrations mostly of male matured
<date id="B0273384FFC3FFD0BF7CFA6F3D89FA56" box="[1299,1425,1438,1461]" pageId="39" pageNumber="430" value="2005-11-13">13.XI.2005</date>
): Tibial apophysis basally quite wide in ventral view, distinctly set back from distal margin of tibia (
<figureCitation id="5CA209C1FFC3FFD0BF5FFA2E3D88FA1B" box="[1328,1424,1503,1528]" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="39" pageNumber="430">Fig. 21E</figureCitation>
), carrying four medium-long apical megaspines, the second from ventral situated clearly lower (more proximal) than others (
<figureCitation id="5CA209C1FFC3FFD0B9FCF9B13BFAF9BA" box="[915,994,1600,1625]" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="39" pageNumber="430">Fig. 21</figureCitation>
E-F). Apical lobes of cymbium distinct, prodorsal one longer and narrower than proventral one (
<figureCitation id="5CA209C1FFC3FFD0B928F9703BA8F979" box="[839,944,1665,1690]" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="39" pageNumber="430">Fig. 21H</figureCitation>
). Paracymbium of average length but quite shallow (
<figureCitation id="5CA209C1FFC3FFD0B9C9F9503C2CF959" box="[934,1076,1697,1722]" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="39" pageNumber="430">Fig. 21A, D</figureCitation>
), carrying long thick bristles in a widely spaced group on non-elevated cumulus (
<figureCitation id="5CA209C1FFC3FFD0BF1EF9333B86F918" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="39" pageNumber="430">Fig. 21A, D</figureCitation>
). Subtegulum without apophysis. Tegulum with proximal edge indistinct and completely without denticles, distinguished from rest of tegulum only by a stronger pigmentation (
<figureCitation id="5CA209C1FFC3FFD0BE34F8B23CDAF8BF" box="[1115,1218,1859,1884]" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="39" pageNumber="430">Fig. 21B</figureCitation>
). Pigmented bridge between tegulum and contrategulum on retrolateral side of palpal organ unbroken (
<figureCitation id="5CA209C1FFC3FFD0BE16F8753CFAF87E" box="[1145,1250,1924,1949]" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="39" pageNumber="430">Fig. 21B</figureCitation>
). Contrategulum with distinct, somewhat conical (in distal view) ventral process, with several parallel ridges directed towards embolus and situated near base of ventral process; dorsal apex of contrategulum large and widely toungeshaped (
<figureCitation id="5CA209C1FFCEFFDDBA9CFEF33957FEF8" box="[243,335,258,283]" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="42" pageNumber="433">
Fig.
<date id="B0273384FFCEFFDDBB47FEF23957FEF8" box="[296,335,259,283]" pageId="42" pageNumber="433" value="1921-01">21I</date>
</figureCitation>
). Para-embolic plate short (
<figureCitation id="5CA209C1FFCEFFDDB8EDFEF33AFFFEF8" box="[642,743,258,283]" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="42" pageNumber="433">Fig. 21B</figureCitation>
); no sharp proventral angle on and no sharp edge below para-embolic plate. Embolus proper with moderately wide apex (
<figureCitation id="5CA209C1FFCEFFDDBB4FFE93396BFE98" box="[288,371,354,379]" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="42" pageNumber="433">Fig. 21</figureCitation>
A-B); dorsal and ventral walls of sclerotised part equally wide, without modification (
<figureCitation id="5CA209C1FFCEFFDDBAF0FE5338E0FE58" box="[159,248,418,443]" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="42" pageNumber="433">
Fig.
<date id="B0273384FFCEFFDDBABDFE5238E0FE58" box="[210,248,419,443]" pageId="42" pageNumber="433" value="1921-01">21I</date>
</figureCitation>
), retrolateral wall carrying a distinct distal keel (
<figureCitation id="5CA209C1FFCEFFDDBAF0FE333919FE38" box="[159,257,450,475]" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="42" pageNumber="433">Fig. 21B</figureCitation>
); membranous part of embolus proper narrow, unpigmented throughout (
<figureCitation id="5CA209C1FFCEFFDDBBC2FE123A17FE18" box="[429,527,483,507]" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="42" pageNumber="433">Fig. 21C</figureCitation>
).
</paragraph>
<caption id="90E645CCFFCCFFDFBAF8F8F53B30F824" httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="40" pageNumber="431" targetBox="[157,1424,264,1742]" targetPageId="40">
<paragraph id="C4261544FFCCFFDFBAF8F8F53B30F824" blockId="40.[151,1439,1796,1991]" pageId="40" pageNumber="431">
Fig. 21.
<taxonomicName id="03996EC7FFCCFFDFBA82F8F539A2F8F9" authorityName="Platnick in Platnick, Schwendinger &amp; Steiner" authorityYear="1997" box="[237,442,1796,1818]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="40" pageNumber="431" phylum="Arthropoda" rank="species" species="tempurung">
<emphasis id="F6EDC956FFCCFFDFBA82F8F539A2F8F9" box="[237,442,1796,1818]" italics="true" pageId="40" pageNumber="431">Liphistius tempurung</emphasis>
</taxonomicName>
, details of left palp of four males: specimen from type locality, matured 13.XI.2005 (A-F, I); from Gua Cicak, matured 23.I.2005 (G, J); from type locality, matured 26.IX.2004 (H, K); from type locality, matured 25.IX.2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L).
</paragraph>
</caption>
<caption id="90E645CCFFCDFFDEBAFCF90C3C10F865" httpUri="https://zenodo.org/record/1021985/files/figure.png" pageId="41" pageNumber="432" targetBox="[153,1429,258,1732]" targetPageId="41">
<paragraph id="C4261544FFCDFFDEBAFCF90C3C10F865" blockId="41.[147,1437,1789,1926]" pageId="41" pageNumber="432">
Fig. 22.
<taxonomicName id="03996EC7FFCDFFDEBA86F90C39ACF8F0" authorityName="Platnick in Platnick, Schwendinger &amp; Steiner" authorityYear="1997" box="[233,436,1789,1811]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="41" pageNumber="432" phylum="Arthropoda" rank="species" species="tempurung">
<emphasis id="F6EDC956FFCDFFDEBA86F90C39ACF8F0" box="[233,436,1789,1811]" italics="true" pageId="41" pageNumber="432">Liphistius tempurung</emphasis>
</taxonomicName>
, details of copulatory organ and genital sternite of six females: Gua Cicak, specimen moulted 8.I.2005 (A- B, N); Gua Tempurung, exuvia, moult 22.I.2007 (C-D, M); Gua Tempurung, exuvia, moult 9.XII.2004 (E-F); Gua Tempurung, exuvia, moult 21.X.2004 (G-H); medium-sized female from Gua Cicak (I-J); small female from Gua Tempurung (K-L). (A, C, E, G, I, K) Vulval plate, dorsal view. (B, D, F, H) Same, ventral view. (J, L) Receptacular cluster, ventral view. (M-N) Posterior margin of genital sternite, ventral view. Scale lines: 1.0 mm (A-D, I-L; E-H; M-N).
</paragraph>
</caption>
<paragraph id="C4261544FFCEFFDDBAF8FDF53939FCD8" blockId="42.[151,756,258,828]" pageId="42" pageNumber="433">
<emphasis id="F6EDC956FFCEFFDDBAF8FDF5392AFDF8" box="[151,306,516,539]" italics="true" pageId="42" pageNumber="433">Measurements</emphasis>
: Total length 14.55; carapace 5.69 long, 5.25 wide; opisthosoma 6.04 long, 5.15 wide; eye mound 0.67 long, 0.83 wide; palpal coxae 1.68 long, 1.19 wide; labium 0.45 long, 0.99 wide; sternum 2.52 long, 1.68 wide (on ventral surface 0.79); palp 9.80 long (2.97 + 1.73 + 3.47 + 1.63); leg I 16.78 long (4.85 + 2.13 + 3.61 +4.16 + 2.03); leg II 18.07 long (4.95 + 2.13 + 3.96 + 4.65 + 2.38); leg III 19.65 long (5.00 + 2.13 + 4.21 + 5.64 + 2.67); leg IV 25.45 long (6.04 + 2.38 + 5.25 + 8.12 + 3.66).
</paragraph>
<paragraph id="C4261544FFCEFFDDBAF8FCA23A5BFA4E" blockId="42.[151,756,851,1453]" pageId="42" pageNumber="433">
<emphasis id="F6EDC956FFCEFFDDBAF8FCA23A2BFC89" bold="true" box="[151,563,851,876]" pageId="42" pageNumber="433">Additions to description of female:</emphasis>
Posterior margin of genital sternite widely W-shaped (
<figureCitation id="5CA209C1FFCEFFDDB827FC823A85FC6F" box="[584,669,883,908]" captionStart="Fig. 22" captionStartId="41.[147,182,1789,1810]" captionTargetBox="[153,1429,258,1732]" captionTargetId="figure@41.[148,1446,254,1752]" captionTargetPageId="41" captionText="Fig. 22. Liphistius tempurung, details of copulatory organ and genital sternite of six females: Gua Cicak, specimen moulted 8. I. 2005 (A- B, N); Gua Tempurung, exuvia, moult 22. I. 2007 (C-D, M); Gua Tempurung, exuvia, moult 9. XII. 2004 (E-F); Gua Tempurung, exuvia, moult 21. X. 2004 (G-H); medium-sized female from Gua Cicak (I-J); small female from Gua Tempurung (K-L). (A, C, E, G, I, K) Vulval plate, dorsal view. (B, D, F, H) Same, ventral view. (J, L) Receptacular cluster, ventral view. (M-N) Posterior margin of genital sternite, ventral view. Scale lines: 1.0 mm (A-D, I-L; E-H; M-N)." httpUri="https://zenodo.org/record/1021985/files/figure.png" pageId="42" pageNumber="433">Fig. 22</figureCitation>
A-I), in slightly posteroventral view posterior edge of posterior stalk of vulval plate clearly visible between paramedian lobes (
<figureCitation id="5CA209C1FFCEFFDDBA85FC22395CFC0F" box="[234,324,979,1004]" captionStart="Fig. 22" captionStartId="41.[147,182,1789,1810]" captionTargetBox="[153,1429,258,1732]" captionTargetId="figure@41.[148,1446,254,1752]" captionTargetPageId="41" captionText="Fig. 22. Liphistius tempurung, details of copulatory organ and genital sternite of six females: Gua Cicak, specimen moulted 8. I. 2005 (A- B, N); Gua Tempurung, exuvia, moult 22. I. 2007 (C-D, M); Gua Tempurung, exuvia, moult 9. XII. 2004 (E-F); Gua Tempurung, exuvia, moult 21. X. 2004 (G-H); medium-sized female from Gua Cicak (I-J); small female from Gua Tempurung (K-L). (A, C, E, G, I, K) Vulval plate, dorsal view. (B, D, F, H) Same, ventral view. (J, L) Receptacular cluster, ventral view. (M-N) Posterior margin of genital sternite, ventral view. Scale lines: 1.0 mm (A-D, I-L; E-H; M-N)." httpUri="https://zenodo.org/record/1021985/files/figure.png" pageId="42" pageNumber="433">Fig. 22</figureCitation>
M-N). Vulval plate (
<figureCitation id="5CA209C1FFCEFFDDB82EFC223A82FC0F" box="[577,666,979,1004]" captionStart="Fig. 22" captionStartId="41.[147,182,1789,1810]" captionTargetBox="[153,1429,258,1732]" captionTargetId="figure@41.[148,1446,254,1752]" captionTargetPageId="41" captionText="Fig. 22. Liphistius tempurung, details of copulatory organ and genital sternite of six females: Gua Cicak, specimen moulted 8. I. 2005 (A- B, N); Gua Tempurung, exuvia, moult 22. I. 2007 (C-D, M); Gua Tempurung, exuvia, moult 9. XII. 2004 (E-F); Gua Tempurung, exuvia, moult 21. X. 2004 (G-H); medium-sized female from Gua Cicak (I-J); small female from Gua Tempurung (K-L). (A, C, E, G, I, K) Vulval plate, dorsal view. (B, D, F, H) Same, ventral view. (J, L) Receptacular cluster, ventral view. (M-N) Posterior margin of genital sternite, ventral view. Scale lines: 1.0 mm (A-D, I-L; E-H; M-N)." httpUri="https://zenodo.org/record/1021985/files/figure.png" pageId="42" pageNumber="433">Fig. 22</figureCitation>
A-I, K) moderately wide; large parts of poreplate unpigmented, particularly central area around CDO; CDO indistinct, composed of a small drop-shaped or longitudinally slit-shaped opening lying within a larger hollow with a distinct, sunken anterior margin and an indistinct (level with poreplate) posterior margin (
<figureCitation id="5CA209C1FFCEFFDDB868FB653AFEFB4E" box="[519,742,1172,1197]" captionStart="Fig. 22" captionStartId="41.[147,182,1789,1810]" captionTargetBox="[153,1429,258,1732]" captionTargetId="figure@41.[148,1446,254,1752]" captionTargetPageId="41" captionText="Fig. 22. Liphistius tempurung, details of copulatory organ and genital sternite of six females: Gua Cicak, specimen moulted 8. I. 2005 (A- B, N); Gua Tempurung, exuvia, moult 22. I. 2007 (C-D, M); Gua Tempurung, exuvia, moult 9. XII. 2004 (E-F); Gua Tempurung, exuvia, moult 21. X. 2004 (G-H); medium-sized female from Gua Cicak (I-J); small female from Gua Tempurung (K-L). (A, C, E, G, I, K) Vulval plate, dorsal view. (B, D, F, H) Same, ventral view. (J, L) Receptacular cluster, ventral view. (M-N) Posterior margin of genital sternite, ventral view. Scale lines: 1.0 mm (A-D, I-L; E-H; M-N)." httpUri="https://zenodo.org/record/1021985/files/figure.png" pageId="42" pageNumber="433">Fig. 22A, C, E, G, I</figureCitation>
); receptacular cluster large and complex (
<figureCitation id="5CA209C1FFCEFFDDB80DFB4538D6FB0E" captionStart="Fig. 22" captionStartId="41.[147,182,1789,1810]" captionTargetBox="[153,1429,258,1732]" captionTargetId="figure@41.[148,1446,254,1752]" captionTargetPageId="41" captionText="Fig. 22. Liphistius tempurung, details of copulatory organ and genital sternite of six females: Gua Cicak, specimen moulted 8. I. 2005 (A- B, N); Gua Tempurung, exuvia, moult 22. I. 2007 (C-D, M); Gua Tempurung, exuvia, moult 9. XII. 2004 (E-F); Gua Tempurung, exuvia, moult 21. X. 2004 (G-H); medium-sized female from Gua Cicak (I-J); small female from Gua Tempurung (K-L). (A, C, E, G, I, K) Vulval plate, dorsal view. (B, D, F, H) Same, ventral view. (J, L) Receptacular cluster, ventral view. (M-N) Posterior margin of genital sternite, ventral view. Scale lines: 1.0 mm (A-D, I-L; E-H; M-N)." httpUri="https://zenodo.org/record/1021985/files/figure.png" pageId="42" pageNumber="433">Fig. 22B, D, F, H</figureCitation>
), slightly protruding beyond widely invaginated anterior margin of poreplate; posterior stalk strongly pigmented, more or less completely separated from pigmented areas of poreplate, its posterior portion finely pitted and bent ventrad at right angles; genital atrium with few or without lateral hairs, most vulvae without posterior hairs (
<figureCitation id="5CA209C1FFCEFFDDBB50FA643A2FFA4E" box="[319,567,1429,1453]" captionStart="Fig. 22" captionStartId="41.[147,182,1789,1810]" captionTargetBox="[153,1429,258,1732]" captionTargetId="figure@41.[148,1446,254,1752]" captionTargetPageId="41" captionText="Fig. 22. Liphistius tempurung, details of copulatory organ and genital sternite of six females: Gua Cicak, specimen moulted 8. I. 2005 (A- B, N); Gua Tempurung, exuvia, moult 22. I. 2007 (C-D, M); Gua Tempurung, exuvia, moult 9. XII. 2004 (E-F); Gua Tempurung, exuvia, moult 21. X. 2004 (G-H); medium-sized female from Gua Cicak (I-J); small female from Gua Tempurung (K-L). (A, C, E, G, I, K) Vulval plate, dorsal view. (B, D, F, H) Same, ventral view. (J, L) Receptacular cluster, ventral view. (M-N) Posterior margin of genital sternite, ventral view. Scale lines: 1.0 mm (A-D, I-L; E-H; M-N)." httpUri="https://zenodo.org/record/1021985/files/figure.png" pageId="42" pageNumber="433">Fig. 22A, C, E, G, I, K</figureCitation>
).
</paragraph>
<paragraph id="C4261544FFCEFFDDBAF8FA3539B1F8BE" blockId="42.[151,756,1476,2078]" pageId="42" pageNumber="433">
<emphasis id="F6EDC956FFCEFFDDBAF8FA353914FA38" bold="true" box="[151,268,1476,1499]" pageId="42" pageNumber="433">Variation:</emphasis>
Carapace lengths in males (n=6) 5.51-6.46, carapace widths 4.92-5.87; in the largest females with a well-developed vulval plate (n=9) 5.94-7.87 and 4.80- 6.69, respectively. In two females from Gua Tempurung both AME are completely absent, in other specimens they are reduced (without cornea), tiny (smaller than the pits from which nearby bristles arise) or normally developed. There is only minor variation in the tarsal scopula of males: covering distal two-thirds of tarsus II in two males (both from Gua Cicak), only distal half in all other males; being very thin (in two males from Gua Cicak) or thin (in remaining males) on tarsus IV, always weaker than on tarsi I-III.
</paragraph>
<paragraph id="C4261544FFCEFFDDBAF8F8943C95FDFC" blockId="42.[151,756,1476,2078]" lastBlockId="42.[831,1437,258,543]" pageId="42" pageNumber="433">
Variation in the shape of the ventral process of the contrategulum is shown in
<figureCitation id="5CA209C1FFCEFFDDBBA7F8743A0FF87D" box="[456,535,1925,1950]" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="42" pageNumber="433">Fig. 21</figureCitation>
I-L, variation in the shape of vulval plates in
<figureCitation id="5CA209C1FFCEFFDDBBC4F85439E0F85D" box="[427,504,1957,1982]" captionStart="Fig. 22" captionStartId="41.[147,182,1789,1810]" captionTargetBox="[153,1429,258,1732]" captionTargetId="figure@41.[148,1446,254,1752]" captionTargetPageId="41" captionText="Fig. 22. Liphistius tempurung, details of copulatory organ and genital sternite of six females: Gua Cicak, specimen moulted 8. I. 2005 (A- B, N); Gua Tempurung, exuvia, moult 22. I. 2007 (C-D, M); Gua Tempurung, exuvia, moult 9. XII. 2004 (E-F); Gua Tempurung, exuvia, moult 21. X. 2004 (G-H); medium-sized female from Gua Cicak (I-J); small female from Gua Tempurung (K-L). (A, C, E, G, I, K) Vulval plate, dorsal view. (B, D, F, H) Same, ventral view. (J, L) Receptacular cluster, ventral view. (M-N) Posterior margin of genital sternite, ventral view. Scale lines: 1.0 mm (A-D, I-L; E-H; M-N)." httpUri="https://zenodo.org/record/1021985/files/figure.png" pageId="42" pageNumber="433">Fig. 22</figureCitation>
A-I, K. One male from the Gua Cicak has one of the four apical megaspines on its tibial apophysis weakly developed on both palps (
<figureCitation id="5CA209C1FFCEFFDDBAF0F7F73910F7FD" box="[159,264,2054,2078]" captionStart="Fig. 21" captionStartId="40.[151,186,1796,1817]" captionTargetBox="[157,1424,264,1742]" captionTargetId="figure@40.[148,1446,254,1764]" captionTargetPageId="40" captionText="Fig. 21. Liphistius tempurung, details of left palp of four males: specimen from type locality, matured 13. XI. 2005 (A-F, I); from Gua Cicak, matured 23. I. 2005 (G, J); from type locality, matured 26. IX. 2004 (H, K); from type locality, matured 25. IX. 2004 (L). (A) Cymbium and palpal organ, ventral view. (B) Palpal organ, retrodorsal and slightly distal view. (C) Distal margin of cymbium and palpal organ, prolateral view. (D) Paracymbium, retrolateral view. (E) Distal part of palpal tibia, ventral view. (F) Tibial apophysis, retrodorsal and slightly proximal view. (G) Same, retrolateral and slightly proximal view. (H) Distal part of cymbium, prolateral view. (I-L) Palpal organ, distal view (dorsal side up). Abbreviations: T - tegulum; vp - ventral process of contrategulum. Scale lines: 0.5 mm (A, D-G; B-C, H-L)." httpUri="https://zenodo.org/record/1021981/files/figure.png" pageId="42" pageNumber="433">Fig. 21G</figureCitation>
). The number of lateral hairs in the genital atrium of females ranges from a few to none (
<figureCitation id="5CA209C1FFCEFFDDBF5DFEF33D98FEF8" box="[1330,1408,258,283]" captionStart="Fig. 22" captionStartId="41.[147,182,1789,1810]" captionTargetBox="[153,1429,258,1732]" captionTargetId="figure@41.[148,1446,254,1752]" captionTargetPageId="41" captionText="Fig. 22. Liphistius tempurung, details of copulatory organ and genital sternite of six females: Gua Cicak, specimen moulted 8. I. 2005 (A- B, N); Gua Tempurung, exuvia, moult 22. I. 2007 (C-D, M); Gua Tempurung, exuvia, moult 9. XII. 2004 (E-F); Gua Tempurung, exuvia, moult 21. X. 2004 (G-H); medium-sized female from Gua Cicak (I-J); small female from Gua Tempurung (K-L). (A, C, E, G, I, K) Vulval plate, dorsal view. (B, D, F, H) Same, ventral view. (J, L) Receptacular cluster, ventral view. (M-N) Posterior margin of genital sternite, ventral view. Scale lines: 1.0 mm (A-D, I-L; E-H; M-N)." httpUri="https://zenodo.org/record/1021985/files/figure.png" pageId="42" pageNumber="433">Fig. 22</figureCitation>
A- I); one female (from Gua Cicak) has additionally two paramedian hairs in the posterior part of its genital atrium (
<figureCitation id="5CA209C1FFCEFFDDB928FE953BBAFE9F" box="[839,930,356,380]" captionStart="Fig. 22" captionStartId="41.[147,182,1789,1810]" captionTargetBox="[153,1429,258,1732]" captionTargetId="figure@41.[148,1446,254,1752]" captionTargetPageId="41" captionText="Fig. 22. Liphistius tempurung, details of copulatory organ and genital sternite of six females: Gua Cicak, specimen moulted 8. I. 2005 (A- B, N); Gua Tempurung, exuvia, moult 22. I. 2007 (C-D, M); Gua Tempurung, exuvia, moult 9. XII. 2004 (E-F); Gua Tempurung, exuvia, moult 21. X. 2004 (G-H); medium-sized female from Gua Cicak (I-J); small female from Gua Tempurung (K-L). (A, C, E, G, I, K) Vulval plate, dorsal view. (B, D, F, H) Same, ventral view. (J, L) Receptacular cluster, ventral view. (M-N) Posterior margin of genital sternite, ventral view. Scale lines: 1.0 mm (A-D, I-L; E-H; M-N)." httpUri="https://zenodo.org/record/1021985/files/figure.png" pageId="42" pageNumber="433">
Fig.
<date id="B0273384FFCEFFDDB914FE953BBAFE9F" box="[891,930,356,380]" pageId="42" pageNumber="433" value="1922-01">22I</date>
</figureCitation>
). Females from Gua Cicak have more hairs in the genital atrium than females from the
<typeStatus id="1B22ABE6FFCEFFDDBF60FE743D25FE7E" box="[1295,1341,389,413]" pageId="42" pageNumber="433">type</typeStatus>
locality. The CDO is relatively large and clearly discernible only in the two smallest females from each locality (
<figureCitation id="5CA209C1FFCEFFDDBF50FE343B4EFE1C" captionStart="Fig. 22" captionStartId="41.[147,182,1789,1810]" captionTargetBox="[153,1429,258,1732]" captionTargetId="figure@41.[148,1446,254,1752]" captionTargetPageId="41" captionText="Fig. 22. Liphistius tempurung, details of copulatory organ and genital sternite of six females: Gua Cicak, specimen moulted 8. I. 2005 (A- B, N); Gua Tempurung, exuvia, moult 22. I. 2007 (C-D, M); Gua Tempurung, exuvia, moult 9. XII. 2004 (E-F); Gua Tempurung, exuvia, moult 21. X. 2004 (G-H); medium-sized female from Gua Cicak (I-J); small female from Gua Tempurung (K-L). (A, C, E, G, I, K) Vulval plate, dorsal view. (B, D, F, H) Same, ventral view. (J, L) Receptacular cluster, ventral view. (M-N) Posterior margin of genital sternite, ventral view. Scale lines: 1.0 mm (A-D, I-L; E-H; M-N)." httpUri="https://zenodo.org/record/1021985/files/figure.png" pageId="42" pageNumber="433">
Fig.
<date id="B0273384FFCEFFDDBF1DFE373D80FE3D" box="[1394,1432,454,478]" pageId="42" pageNumber="433" value="1922-01">22I</date>
, K
</figureCitation>
); in all larger females examined it is small and lies in a depression (
<figureCitation id="5CA209C1FFCEFFDDB9AFFDF63C99FDFC" box="[960,1153,519,543]" captionStart="Fig. 22" captionStartId="41.[147,182,1789,1810]" captionTargetBox="[153,1429,258,1732]" captionTargetId="figure@41.[148,1446,254,1752]" captionTargetPageId="41" captionText="Fig. 22. Liphistius tempurung, details of copulatory organ and genital sternite of six females: Gua Cicak, specimen moulted 8. I. 2005 (A- B, N); Gua Tempurung, exuvia, moult 22. I. 2007 (C-D, M); Gua Tempurung, exuvia, moult 9. XII. 2004 (E-F); Gua Tempurung, exuvia, moult 21. X. 2004 (G-H); medium-sized female from Gua Cicak (I-J); small female from Gua Tempurung (K-L). (A, C, E, G, I, K) Vulval plate, dorsal view. (B, D, F, H) Same, ventral view. (J, L) Receptacular cluster, ventral view. (M-N) Posterior margin of genital sternite, ventral view. Scale lines: 1.0 mm (A-D, I-L; E-H; M-N)." httpUri="https://zenodo.org/record/1021985/files/figure.png" pageId="42" pageNumber="433">Fig. 22A, C, E, G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFCEFFDDB950FDC73C32FABE" pageId="42" pageNumber="433" type="biology_ecology">
<paragraph id="C4261544FFCEFFDDB950FDC73CA5FC16" blockId="42.[831,1437,566,1373]" pageId="42" pageNumber="433">
<emphasis id="F6EDC956FFCEFFDDB950FDC73BB9FDAE" bold="true" box="[831,929,566,589]" pageId="42" pageNumber="433">Biology:</emphasis>
All specimens examined were collected from burrows in sandy or loamy soil on the floor in the aphotic zone of two caves. Unlike in
<taxonomicName id="03996EC7FFCEFFDDBF7EFD883D8DFD6D" authorityName="Abraham" authorityYear="1923" box="[1297,1429,631,656]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="433" phylum="Arthropoda" rank="species" species="batuensis">
<emphasis id="F6EDC956FFCEFFDDBF7EFD883D8DFD6D" box="[1297,1429,631,656]" italics="true" pageId="42" pageNumber="433">L. batuensis</emphasis>
</taxonomicName>
, no nests were found on cave walls. Most burrows ran into the depth of the soil and were closed by a single trapdoor. Only one burrow ran under the soil surface and had a second door at the opposite end, about six centimetres away from the first door. From the main entrance up to eight quite long (as usual in cavedwelling
<taxonomicName id="03996EC7FFCEFFDDB9CAFCAD3C16FC90" authorityName="Schiodte" authorityYear="1849" box="[933,1038,860,883]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="433" phylum="Arthropoda" rank="genus">
<emphasis id="F6EDC956FFCEFFDDB9CAFCAD3C16FC90" box="[933,1038,860,883]" italics="true" pageId="42" pageNumber="433">Liphistius</emphasis>
</taxonomicName>
) signal lines (the longest
<quantity id="0361B8A1FFCEFFDDBF49FCAD3D73FC97" box="[1318,1387,860,884]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="42" pageNumber="433" unit="cm" value="20.0">20 cm</quantity>
) ran over the soil surface (in a few cases also over nearby rock). The trapdoor of the largest female was
<quantity id="0361B8A1FFCEFFDDBF20FC6F3D85FC56" box="[1359,1437,926,950]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="42" pageNumber="433" unit="cm" value="1.8">1.8 cm</quantity>
long and 3.0 cm wide, in penultimate males trapdoors were 1.2-1.8 long and 2.0-2.8 wide.
</paragraph>
<paragraph id="C4261544FFCEFFDDB950FC0E3B8AFB5B" blockId="42.[831,1437,566,1373]" pageId="42" pageNumber="433">Despite the fact that all spiders examined were found in the dark portions of these two caves, no noteworthy troglobiomorphic adaptations are visible, apart from a relatively light body colouration in all living spiders and a more or less distinct reduction of the AME in some spiders.</paragraph>
<paragraph id="C4261544FFCEFFDDB950FB323C32FABE" blockId="42.[831,1437,566,1373]" pageId="42" pageNumber="433">
An illustrated presentation of
<taxonomicName id="03996EC7FFCEFFDDBEF7FB323D37FB39" authorityName="Platnick in Platnick, Schwendinger &amp; Steiner" authorityYear="1997" box="[1176,1327,1219,1242]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="433" phylum="Arthropoda" rank="species" species="tempurung">
<emphasis id="F6EDC956FFCEFFDDBEF7FB323D37FB39" box="[1176,1327,1219,1242]" italics="true" pageId="42" pageNumber="433">L. tempurung</emphasis>
</taxonomicName>
(together with
<taxonomicName id="03996EC7FFCEFFDDB91BFB153BFDFB19" box="[884,997,1251,1275]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="433" phylum="Arthropoda" rank="species" species="kanthan">
<emphasis id="F6EDC956FFCEFFDDB91BFB153BFDFB19" box="[884,997,1251,1275]" italics="true" pageId="42" pageNumber="433">L. kanthan</emphasis>
</taxonomicName>
) and its habitat is given in
<bibRefCitation id="A00868B5FFCEFFDDBE91FB123D81FB18" author="Steiner H." box="[1278,1433,1251,1275]" journalOrPublisher="Munchen" pageId="42" pageNumber="433" pagination="148 - 150" part="44" refString="Steiner H. 1998. Lebende Fossilien aus den Hohlen Malaysias. Mitteilungen des Verbandes der deutschen Hohlen- und Karstforscher e. V. Munchen 44 (4): 148 - 150." title="Lebende Fossilien aus den Hohlen Malaysias. Mitteilungen des Verbandes der deutschen Hohlen- und Karstforscher e. V" type="journal article" year="1998">Steiner (1998)</bibRefCitation>
. An extensive bibliography on
<taxonomicName id="03996EC7FFCEFFDDBEE3FAF43CF0FAFF" authorityName="Schiødte" authorityYear="1849" box="[1164,1256,1285,1308]" class="Arachnida" family="Liphistiidae" genus="Lipistius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="433" phylum="Arthropoda" rank="genus">
<emphasis id="F6EDC956FFCEFFDDBEE3FAF43CF0FAFF" box="[1164,1256,1285,1308]" italics="true" pageId="42" pageNumber="433">Lipistius</emphasis>
</taxonomicName>
and many other animals in caves of
<collectingCountry id="BC8E55D4FFCEFFDDBE7DFAD53C6CFAD8" box="[1042,1140,1316,1339]" name="Malaysia" pageId="42" pageNumber="433">Malaysia</collectingCountry>
can be found on Liz Price’s website (
<bibRefCitation id="A00868B5FFCEFFDDB9F0FAB53C07FABE" author="Price L." box="[927,1055,1348,1373]" journalOrPublisher="Available at" pageId="42" pageNumber="433" pagination="2017" publicationUrl="http://www.cavesofmalaysia.com/newspage3.htm" refString="Price L. 2017. Malaysian biospeleology bibliography. Available at http: // www. cavesofmalaysia. com / newspage 3. htm (accessed in July 2017)." title="Malaysian biospeleology bibliography" type="book chapter" year="2017">Price, 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFCEFFDDB950FA853B87F9D3" pageId="42" pageNumber="433" type="distribution">
<paragraph id="C4261544FFCEFFDDB950FA853B87F9D3" blockId="42.[831,1437,1396,1584]" pageId="42" pageNumber="433">
<emphasis id="F6EDC956FFCEFFDDB950FA853BCDFA68" bold="true" box="[831,981,1396,1419]" pageId="42" pageNumber="433">Distribution:</emphasis>
Known only from inside two limestone caves (
<figureCitation id="5CA209C1FFCEFFDDB9FDFA643BC2FA4E" box="[914,986,1429,1453]" captionStart="Fig. 1" captionStartId="1.[151,186,1392,1413]" captionTargetBox="[153,758,625,1349]" captionTargetId="figure@1.[149,499,895,1170]" captionTargetPageId="1" captionText="Fig. 1. Localities of Liphistius species of the malayanus - group, tioman - group, linang - group and batuenis - group in peninsular Malaysia and southern Thailand (coast of Sumatra omitted): 1 - Gunung Angsi (type locality of L. malayanus); 2 - Templer Park and Gua Anak Takun (L. malayanus, L. batuensis); 3 - Genting Highlands (L. malayanus); 4 - Fraser’s Hill (L. malayanus); 5 - Cameron Highlands (type locality of L. malayanus cameroni); 6 - Sungai Jasin in Endau Rompin National Park (type locality of L. endau); 7 - Gunung Belumut (L. endau); 8 - Gunung Muntahak (L. endau; type locality of L. gracilis sp. nov.); 9 - Sungai Rengit (type locality of L. johore); 10 - Gunung Kajang on Tioman Island (type locality of L. tioman); 11 - Gua Charas in Bukit Charas (type locality of L. panching); 12 - Nusa Camp in Taman Negara (type locality of L. negara sp. nov.); 13 - Jeram Linang Waterfall (type locality of L. linang sp. nov.); 14 - Sankalakhierie Mountains (type locality of L. indra); 15 - Batu Caves (type locality of L. batuensis); 16 - Gua Tempurung (type locality of L. tempurung); 17 - Gua Cicak (L. tempurung); 18 - Gua Keris (type locality of L. priceae sp. nov). Localities with conspecific populations are encircled. Colours distinguish species groups." httpUri="https://zenodo.org/record/1021937/files/figure.png" pageId="42" pageNumber="433">Fig. 1</figureCitation>
, localities 16-17), about
<quantity id="0361B8A1FFCEFFDDBF69FA643D49FA4F" box="[1286,1361,1429,1453]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="42" pageNumber="433" unit="km" value="13.0">13 km</quantity>
apart, southeast of Ipoh in the
<collectingRegion id="065DDBA6FFCEFFDDBE28FA443C9CFA2F" box="[1095,1156,1461,1484]" country="Malaysia" name="Perak" pageId="42" pageNumber="433">Perak</collectingRegion>
State. A short description of Gua Tempurung and photographs of the cave and some spider nests are given in
<bibRefCitation id="A00868B5FFCEFFDDBEE5FA073D7FF9EC" author="Platnick N. &amp; Schwendinger P. J. &amp; Steiner H." box="[1162,1383,1526,1551]" journalOrPublisher="American Museum Novitates" pageId="42" pageNumber="433" pagination="1 - 13" part="3209" refString="Platnick N. I., Schwendinger P. J., Steiner H. 1997. Three new species of the spider genus Liphistius (Araneae, Mesothelae) from Malaysia. American Museum Novitates 3209: 1 - 13." title="Three new species of the spider genus Liphistius (Araneae, Mesothelae) from Malaysia" type="journal article" year="1997">
Platnick
<emphasis id="F6EDC956FFCEFFDDBE87FA093D00F9EE" box="[1256,1304,1526,1551]" italics="true" pageId="42" pageNumber="433">et al</emphasis>
. (1997
</bibRefCitation>
: 2-3, figs 1-3).
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFCEFFDDB950F9B93B8CF806" pageId="42" pageNumber="433" type="discussion">
<paragraph id="C4261544FFCEFFDDB950F9B93B8CF806" blockId="42.[831,1437,1607,2021]" pageId="42" pageNumber="433">
<emphasis id="F6EDC956FFCEFFDDB950F9B93BAAF983" bold="true" box="[831,946,1608,1632]" pageId="42" pageNumber="433">Remarks:</emphasis>
First reported from a “cave in Gunong Tempurong” by
<bibRefCitation id="A00868B5FFCEFFDDB980F9963CD1F963" author="Bullock J. A." box="[1007,1225,1639,1664]" journalOrPublisher="Malayan Nature Journal" pageId="42" pageNumber="433" pagination="135 - 141" part="25" refString="Bullock J. A. 1972. Cave biology in Malaysia. Malayan Nature Journal 25: 135 - 141." title="Cave biology in Malaysia" type="journal article" year="1972">Bullock (1972: 138)</bibRefCitation>
under
<taxonomicName id="03996EC7FFCEFFDDBF7BF9983D8EF99D" authorityName="Abraham" authorityYear="1923" box="[1300,1430,1639,1664]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="433" phylum="Arthropoda" rank="species" species="batuensis">
<emphasis id="F6EDC956FFCEFFDDBF7BF9983D8EF99D" box="[1300,1430,1639,1664]" italics="true" pageId="42" pageNumber="433">L. batuensis</emphasis>
</taxonomicName>
, this is a narrowly endemic species restricted to caves. Like the other cave-living
<taxonomicName id="03996EC7FFCEFFDDBEBBF9583D27F923" authorityName="Schiodte" authorityYear="1849" box="[1236,1343,1705,1728]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="433" phylum="Arthropoda" rank="genus">
<emphasis id="F6EDC956FFCEFFDDBEBBF9583D27F923" box="[1236,1343,1705,1728]" italics="true" pageId="42" pageNumber="433">Liphistius</emphasis>
</taxonomicName>
species in
<collectingCountry id="BC8E55D4FFCEFFDDB90CF9383BDEF903" box="[867,966,1737,1760]" name="Malaysia" pageId="42" pageNumber="433">Malaysia</collectingCountry>
[
<taxonomicName id="03996EC7FFCEFFDDB9B1F93A3C70F903" authorityName="Abraham" authorityYear="1923" box="[990,1128,1737,1762]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="433" phylum="Arthropoda" rank="species" species="batuensis">
<emphasis id="F6EDC956FFCEFFDDB9B1F93A3C70F903" box="[990,1128,1737,1762]" italics="true" pageId="42" pageNumber="433">L. batuensis</emphasis>
</taxonomicName>
,
<taxonomicName id="03996EC7FFCEFFDDBE11F93A3CE1F903" box="[1150,1273,1737,1762]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="433" phylum="Arthropoda" rank="species" species="kanthan">
<emphasis id="F6EDC956FFCEFFDDBE11F93A3CE1F903" box="[1150,1273,1737,1762]" italics="true" pageId="42" pageNumber="433">L. kanthan</emphasis>
</taxonomicName>
(currently on the IUCN list of critically endangered species; http:// www.iucnredlist.org/search?page=52) and
<taxonomicName id="03996EC7FFCEFFDDBF4BF8FD3D84F8C0" authority="Peter J. Schwendinger, 2017" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[1316,1436,1804,1827]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="433" phylum="Arthropoda" rank="species" species="priceae" status="sp. nov.">
<emphasis id="F6EDC956FFCEFFDDBF4BF8FD3D84F8C0" box="[1316,1436,1804,1827]" italics="true" pageId="42" pageNumber="433">L. priceae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDDE742DFFCEFFDDB950F8DD3B85F8A0" box="[831,925,1836,1859]" pageId="42" pageNumber="433" rank="species">sp. nov.</taxonomicNameLabel>
],
<taxonomicName id="03996EC7FFCEFFDDB9DAF8DD3C54F8A0" authorityName="Platnick in Platnick, Schwendinger &amp; Steiner" authorityYear="1997" box="[949,1100,1836,1859]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="433" phylum="Arthropoda" rank="species" species="tempurung">
<emphasis id="F6EDC956FFCEFFDDB9DAF8DD3C54F8A0" box="[949,1100,1836,1859]" italics="true" pageId="42" pageNumber="433">L. tempurung</emphasis>
</taxonomicName>
is at great risk of becoming extinct if overcollected, or if their habitats change or are destroyed. Quarrying of limestone hills for cement production is widespread in
<collectingCountry id="BC8E55D4FFCEFFDDBE16F87C3CC3F847" box="[1145,1243,1933,1956]" name="Malaysia" pageId="42" pageNumber="433">Malaysia</collectingCountry>
and elsewhere in Southeast Asia, and poses the greatest danger for these species.
</paragraph>
</subSubSection>
</treatment>
</document>
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