689 lines
58 KiB
XML
689 lines
58 KiB
XML
<document id="F4EA1B81C155EE47E628C9A04FA15A73" ID-DOI="10.3897/zookeys.1227.138100" ID-ZooBank="F259B26F-4DA1-452E-ABEE-7D0957CFE261" ID-publisher-id="138100" URI-arpha="C4646FFE-9C91-508A-94DF-6C3BA18F2656" URI-zoobank="http://zoobank.org/F259B26F-4DA1-452E-ABEE-7D0957CFE261" article-type="research-article" checkinTime="1739298295086" checkinUser="pensoft" docAuthor="Kaltenbach, Thomas, Kluge, Nikita J. & Gattolliat, Jean-Luc" docDate="2025" docId="EA88C84262745D8682259B48D59E7A64" docLanguage="en" docName="ZooKeys 1227: 159-347" docOrigin="ZooKeys 1227" docSource="https://zookeys.pensoft.net/article/138100/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Papuanatula (Papuafiliola) tuberculata Kaltenbach, Kluge & Gattolliat, 2025, sp. nov." docType="treatment" docVersion="1" dtd-version="3.0" lastPageNumber="347" masterDocId="C4646FFE9C91508A94DF6C3BA18F2656" masterDocTitle="Redescription of Papuanatula Lugo-Ortiz & McCafferty (Ephemeroptera, Baetidae), with description of a new subgenus and 20 new species" masterLastPageNumber="347" masterPageNumber="159" pageNumber="159" updateTime="1739298573857" updateUser="pensoft">
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<mods:mods id="2F853DFCD14E9854E88CEF4F2D8F28F1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="60DA2609C2A71CCB7FB6EF9E13958CAC">
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<mods:title id="C2A32AD3D263473770C49C6250041DCD">Redescription of Papuanatula Lugo-Ortiz & McCafferty (Ephemeroptera, Baetidae), with description of a new subgenus and 20 new species</mods:title>
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</mods:titleInfo>
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<mods:name id="E34A9F229AB8CCD7229E65F08D9FDA96" type="personal">
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<mods:role id="2BC535EA9A218CCB4D626F71C9EE1CE9">
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<mods:roleTerm id="76048296192CFC8A82790142E0216C7F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="88F84636340C089368FC9CBE6552A516">Kaltenbach, Thomas</mods:namePart>
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<mods:nameIdentifier id="267C612A9656E1C895643F11B006DBF7" type="ORCID">0000-0001-8052-0388</mods:nameIdentifier>
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<mods:affiliation id="D6A7EEF7288427F21697B2FD6A3CAC8A">Département de zoologie, Muséum cantonal des Sciences Naturelles, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland & Department of Ecology and Evolution, University of Lausanne (UNIL), CH- 1015 Lausanne, Switzerland</mods:affiliation>
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</mods:name>
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<mods:name id="C35D4860880BA2452DD92BA7FA7A17F4" type="personal">
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<mods:role id="C61A65EAA42F3EFB2A25C4DF6599EFE2">
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<mods:roleTerm id="4149636342DB63510B06EF720838D8FD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7DB3B3248230AD7E61EE427BAC314DB3">Kluge, Nikita J.</mods:namePart>
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<mods:nameIdentifier id="0BC8E60DA48B2A6FE494A4C10B65AE50" type="ORCID">0000-0001-9741-7790</mods:nameIdentifier>
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<mods:affiliation id="17D7CB8D9633A96045F5439AC6B2578E">Department of Entomology, Biological Faculty, Saint-Petersburg State University, Universitetskaya nab., 7 / 9, Saint Petersburg, 199034, Russia</mods:affiliation>
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</mods:name>
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<mods:name id="B771131DA4F347A736E256E23507ED16" type="personal">
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<mods:role id="F591870AD34F8732E24B43F133C9819C">
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<mods:roleTerm id="CA436E860A2242A4D4B14BAF4F0D0A96">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="39E4A33A2CBB9A0C015B92312D5999AB">Gattolliat, Jean-Luc</mods:namePart>
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<mods:nameIdentifier id="8F346CEABCE38392834EF86F27EB2A68" type="ORCID">0000-0001-5873-5083</mods:nameIdentifier>
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<mods:affiliation id="02D443E44D5403AFCB22F3075DBD6ECB">Département de zoologie, Muséum cantonal des Sciences Naturelles, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland & Department of Ecology and Evolution, University of Lausanne (UNIL), CH- 1015 Lausanne, Switzerland</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="29F131397A8B30BD262243265CC6E9B8">text</mods:typeOfResource>
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<mods:titleInfo id="5FD72968E124F9FE3EC5F1494F3EB07F">
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<mods:title id="EFC1DA8D7AD3E649B8D9D228B6FF5C00">ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="CB1F9B5F12E4CAE0C67564CD53BF11CB">
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<mods:date id="FBEF0D96AB26D4EDC3328CFA2B7E981E">2025</mods:date>
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<mods:detail id="6D3772FB6DFF9C90A61008A465740404" type="pubDate">
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<mods:number id="CF1A3F81885E47D4163B14A3B92F8C2D">2025-02-11</mods:number>
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</mods:detail>
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<mods:detail id="D557E23D15C6E09151790D9DECC0FA37" type="volume">
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<mods:number id="F364AC80E1A24B42EDD2E693FCA0B792">1227</mods:number>
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</mods:detail>
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<mods:extent id="504AA4964A233D324B72589C6558276A" unit="page">
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<mods:start id="EBE97FE1025B487C7EDDC1CC4C1FFB61">159</mods:start>
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<mods:end id="D663F662CA5E94C796ECC34AF0E574EB">347</mods:end>
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</mods:extent>
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<mods:classification id="E9AE1FEB758723C16536D118E98D1195">journal article</mods:classification>
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<mods:identifier id="ACAF9178C04190AB5ACD10132344F4A7" type="DOI">10.3897/zookeys.1227.138100</mods:identifier>
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<mods:identifier id="E64B7F7487E412919E8611228C427CC4" type="ZooBank">F259B26F-4DA1-452E-ABEE-7D0957CFE261</mods:identifier>
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</mods:mods>
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<treatment id="EA88C84262745D8682259B48D59E7A64" ID-arpha="EA88C842-6274-5D86-8225-9B48D59E7A64" ID-zoobank="https://zoobank.org/2315FD83-BEAF-41B3-9164-42A700945CC8" LSID="urn:lsid:plazi:treatment:EA88C84262745D8682259B48D59E7A64" LSID-ZBK="urn:lsid:zoobank.org:act:2315FD83-BEAF-41B3-9164-42A700945CC8" httpUri="http://treatment.plazi.org/id/EA88C84262745D8682259B48D59E7A64">
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<subSubSection id="E33035D0159A944AA8C87B7CCC2D833A" type="nomenclature">
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||
<paragraph id="A817DEBB6ACA40CADDE2ADBE39E48082">
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||
<taxonomicName id="FA2C8CEBB5855DDB093FDA7D8EE45780" authorityName="Kaltenbach & Kluge & Gattolliat" authorityYear="2025" class="Insecta" family="Baetidae" genus="Papuanatula" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov." subGenus="Papuafiliola">
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||
<emphasis id="71976AE7DD4721808911E4FF8D209D16" italics="true">Papuanatula</emphasis>
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||
(
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<emphasis id="F3043D4F94B0B92403ACBE77C02A0F6D" italics="true">Papuafiliola</emphasis>
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||
)
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<emphasis id="790040532D595E5358B9CF14D5044E4C" italics="true">tuberculata</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="F0D922CBE5D290BD4BB86F3652C1727E" rank="species">sp. nov.</taxonomicNameLabel>
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="78D995BF60FED468DBD3A85283610523" type="description">
|
||
<paragraph id="6BCDB19A3B077D5D22341AFA86911893">
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||
<figureCitation id="F93A610681468750818EF0257A1052F2" captionStart="Figure 138" captionStartId="F138" captionText="Figure 138. a, b Papuanatula (Papuafiliola) stenophylla sp. nov., eggs (extracted from mature larva) c Papuanatula (Papuafiliola) tuberculata sp. nov., larva." figureDoi="10.3897/zookeys.1227.138100.figure138" httpUri="https://binary.pensoft.net/fig/1247552">Figs 138</figureCitation>
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||
,
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||
<figureCitation id="1E97C81A16D45AC7C65FECF5AAC7DC19" captionStart="Figure 139" captionStartId="F139" captionText="Figure 139. Papuanatula (Papuafiliola) tuberculata sp. nov., larval exuviae (holotype, in one magnification): a head b fore, middle, and hind legs (densely covered with diatom algae) c half of pronotum and mesonotum d abdomen." figureDoi="10.3897/zookeys.1227.138100.figure139" httpUri="https://binary.pensoft.net/fig/1247553">139</figureCitation>
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,
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<figureCitation id="727F5F20350516FD85D2439D5CF19606" captionStart="Figure 140" captionStartId="F140" captionText="Figure 140. Papuanatula (Papuafiliola) tuberculata sp. nov., larva (a – c, e, h, i holotype): a labrum b, c left and right mandibles d, e maxillae f, g hypopharynx h – j labium (i focus on dorsal side)." figureDoi="10.3897/zookeys.1227.138100.figure140" httpUri="https://binary.pensoft.net/fig/1247554">140</figureCitation>
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||
,
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<figureCitation id="B05ADCA65F7EB685CC7BA6CACB08DBF7" captionStart="Figure 141" captionStartId="F141" captionText="Figure 141. Papuanatula (Papuafiliola) tuberculata sp. nov., larva: a leg b tarsus of fore leg (holotype)." figureDoi="10.3897/zookeys.1227.138100.figure141" httpUri="https://binary.pensoft.net/fig/1247555">141</figureCitation>
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||
,
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<figureCitation id="C46882C9A63C6831644A8E085957FB8C" captionStart="Figure 142" captionStartId="F142" captionText="Figure 142. Papuanatula (Papuafiliola) tuberculata sp. nov., larva: a tergum III b paraprocts c antenna d tergalius IV e its enlarged anal rib (a – c holotype)." figureDoi="10.3897/zookeys.1227.138100.figure142" httpUri="https://binary.pensoft.net/fig/1247556">142</figureCitation>
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,
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<figureCitation id="D10585411729AA32DABA713238CD55CC" captionStart="Figure 143" captionStartId="F143" captionText="Figure 143. Papuanatula (Papuafiliola) tuberculata sp. nov., subimaginal exuviae (holotype): a half of mesonotum b mesopleuron c hind tarsus d fore tarsus." figureDoi="10.3897/zookeys.1227.138100.figure143" httpUri="https://binary.pensoft.net/fig/1247557">143</figureCitation>
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,
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<figureCitation id="0383A3317A0A311CD2F4562FE3E22677" captionStart="Figure 144" captionStartId="F144" captionText="Figure 144. Papuanatula (Papuafiliola) tuberculata sp. nov., female imago (holotype): a dorsal view b lateral view c – e fore, middle and hind legs f pterostigma." figureDoi="10.3897/zookeys.1227.138100.figure144" httpUri="https://binary.pensoft.net/fig/1247559">144</figureCitation>
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,
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<figureCitation id="68569DE840B83EBBC3F403E9627B13D9" captionStart="Figure 145" captionStartId="F145" captionText="Figure 145. a – e Papuanatula (Papuafiliola) tuberculata sp. nov., eggs (extracted from imago)." figureDoi="10.3897/zookeys.1227.138100.figure145" httpUri="https://binary.pensoft.net/fig/1247561">145</figureCitation>
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||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="SECID0EQRCM" type="etymology">
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||
<paragraph id="36C7D458F70C39051815844D9A2B5D72">
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||
<heading id="77F64BF300707C31F4B71F05F42DAACF" reason="title">Etymology.</heading>
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||
</paragraph>
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<paragraph id="29573AB79A14D3CBE8C81EA3C4F8AF4D">
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The species name
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<emphasis id="60D7F703A691A7389C8C6A586D0C3E9B" italics="true">tuberculata</emphasis>
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refers to the unpaired tubercles on abdominal terga of the larva (Fig.
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<figureCitation id="5C2D3D915D37B5E067B33B36009D48D5" captionStart="Figure 138" captionStartId="F138" captionText="Figure 138. a, b Papuanatula (Papuafiliola) stenophylla sp. nov., eggs (extracted from mature larva) c Papuanatula (Papuafiliola) tuberculata sp. nov., larva." figureDoi="10.3897/zookeys.1227.138100.figure138" httpUri="https://binary.pensoft.net/fig/1247552">138 c</figureCitation>
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||
).
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||
</paragraph>
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||
<caption id="2D6C5587111812DC3FB103DA9DEE47E4" ID-DOI="10.3897/zookeys.1227.138100.figure138" ID-arpha="C332D8F1-5102-5FEF-A515-37A02183768D" httpUri="https://binary.pensoft.net/fig/1247552" startId="F138">
|
||
<paragraph id="5730427973C20627023D07E38129C91A">
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||
<label id="9AA24A1FA3CBE8AB1E2EEB04A10E3F00">Figure 138.</label>
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||
</paragraph>
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||
<paragraph id="C754AA300990B91EE950B653DBF58E4B">
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||
<emphasis id="B4DB977665376CB8780334A1C6ABF132" bold="true">a, b</emphasis>
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||
<taxonomicName id="0FACAD4DE8FCC9A0D9F273029D4B7F7D" authorityName="Kaltenbach & Kluge & Gattolliat" authorityYear="2025" class="Insecta" family="Baetidae" genus="Papuanatula" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="stenophylla" status="sp. nov." subGenus="Papuafiliola">
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||
<emphasis id="066DD45BA982679908FD889E3F49015F" italics="true">Papuanatula</emphasis>
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||
(
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||
<emphasis id="2B466C747EA9F6F21540C94687B1A28D" italics="true">Papuafiliola</emphasis>
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)
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<emphasis id="0AC235A61C25BCDE4940780D670BB129" italics="true">stenophylla</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F5DE7FEF95B2E7A7F4D637DE01493EB0" rank="species">sp. nov.</taxonomicNameLabel>
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, eggs (extracted from mature larva)
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<emphasis id="8338F1FED6989A2AE0276CC06468837F" bold="true">c</emphasis>
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<taxonomicName id="D6D6388F89F7DF1476826A3FA84AA031" authorityName="Kaltenbach & Kluge & Gattolliat" authorityYear="2025" class="Insecta" family="Baetidae" genus="Papuanatula" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov." subGenus="Papuafiliola">
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<emphasis id="ECC011B121A93D4463BDD792F5B3067D" italics="true">Papuanatula</emphasis>
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(
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<emphasis id="E5D55F28094F1BE99A91BDABB3584D5D" italics="true">Papuafiliola</emphasis>
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)
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<emphasis id="8424CD16D2AECEE983810434123DAA2D" italics="true">tuberculata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="ED9E5A2F1977E55407B0289D348202E0" rank="species">sp. nov.</taxonomicNameLabel>
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, larva.
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</paragraph>
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</caption>
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</subSubSection>
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||
<subSubSection id="SECID0EJTCM" type="material">
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<paragraph id="A8720286DC042CAF3D1FD4CF4783646C">
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||
<heading id="158D566AF9A1A078B2B47C047F3F7FF8" reason="title">Material examined.</heading>
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||
</paragraph>
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<paragraph id="96C1B6AC4395888644B4EA75973C4338">
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||
<materialsCitation id="B636B8D59DB5FFEF9CAAAB735A65B4F8" collectingDate="2012-08-16" collectionCode="SPbU" collectorName="N. Kluge & L. Sheyko" country="Indonesia" county="Baliem valley" location="river Elagaima" municipality="Wamena" specimenCount="1" specimenCount-female="1" stateProvince="Papua Barat" typeStatus="holotype">
|
||
<emphasis id="98AC288DCC53DC2362F74883F871E5CC" bold="true">
|
||
<emphasis id="7BF10F5E6A21FC381AD3AE23EFC9D2EE" bold="true" italics="true">
|
||
<typeStatus id="07B1A7E98CD86502BF581C9B9CC5E51E" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
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||
.
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||
</emphasis>
|
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L-S-I
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<specimenCount id="081177E50092882DCE446B4943F371E3" count="1" type="female">♀</specimenCount>
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||
{specimen number [V] (2) 2012};
|
||
<collectingCountry id="59AE1BB729D72B67BC9BC703B8D41B7E" name="Indonesia">Indonesia</collectingCountry>
|
||
•
|
||
<collectingRegion id="FAB735A9EB248E43B135CC56A85D3404" country="Indonesia" name="Papua Barat">Papua</collectingRegion>
|
||
,
|
||
<collectingCounty id="A172042C0A07AC50BD9F319355F24189">Baliem valley</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="AF887745A6ADF27289D562EEE1C4A419">Wamena</collectingMunicipality>
|
||
,
|
||
<location id="0699241F8B83C65EF2DB416310A2C526" LSID="urn:lsid:plazi:treatment:EA88C84262745D8682259B48D59E7A64:0699241F8B83C65EF2DB416310A2C526" country="Indonesia" county="Baliem valley" municipality="Wamena" name="river Elagaima" stateProvince="Papua Barat">river Elagaima</location>
|
||
;
|
||
<date id="06FD8AABF02B4E02E5E8BED8226D1789" value="2012-08-16">
|
||
<collectingDate id="C5D27ADF1D7258A9A99971700758D078" value="2012-08-16">16. viii. 2012</collectingDate>
|
||
</date>
|
||
; coll.
|
||
<collectorName id="401FF93B5C9E1517BAB951178F161162">N. Kluge</collectorName>
|
||
&
|
||
<collectorName id="44E10298D515F620F75DF7716DC60D34">L. Sheyko</collectorName>
|
||
;
|
||
<abbrev id="ABBRID0EUTCM" xlink_title="Saint-Petersburg State University">
|
||
<collectionCode id="C6B251AC28C23D79FA28A8F2B1302709">SPbU</collectionCode>
|
||
</abbrev>
|
||
</materialsCitation>
|
||
.
|
||
<emphasis id="6F9EB2FBC1D56F38F92A470BE6BB4812" bold="true">
|
||
<emphasis id="F8813EBAF8A5A40D6992471079A6E95E" bold="true" italics="true">
|
||
<typeStatus id="9587B409059FB54C717BA16CE5F7E80A" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
.
|
||
</emphasis>
|
||
Same locality and collectors;
|
||
<date id="CCB8E937B13A63561609584336205391" value="2012-08-15" valueMax="2012-08-19" valueMin="2012-08-15">15–19. viii. 2012</date>
|
||
:
|
||
<specimenCount id="F64B513E6BC3EC6EA1C3EB22E82BE7B8" count="2" type="larva">2 larvae</specimenCount>
|
||
;
|
||
<abbrev id="ABBRID0E3TCM" xlink_title="Saint-Petersburg State University">
|
||
<collectionCode id="0ED7F7AF6CB57C2964D20DF26D181694">SPbU</collectionCode>
|
||
</abbrev>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EAUCM" type="diagnosis">
|
||
<paragraph id="E42BB4DEF7226C492AE62910D2699085">
|
||
<heading id="E9C6DE7B0F4C059403FF3FB9D3ACCAD7" reason="title">Diagnosis.</heading>
|
||
</paragraph>
|
||
<paragraph id="BEB6752D6E7FE3332D2884C68E46407C">
|
||
<emphasis id="2941B23AA261832C3083D3BCB9876072" bold="true">Larva.</emphasis>
|
||
The following combination of characters distinguishes
|
||
<taxonomicName id="881BBE3F58C5AC5C132EE31E69CF90E8" authorityName="Kaltenbach & Kluge & Gattolliat" authorityYear="2025" class="Insecta" family="Baetidae" genus="Papuanatula" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov.">
|
||
<emphasis id="5413C24D587CCBFE2B7381E34473919C" italics="true">P. tuberculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6EF4EDD2EE1CECFEE27A6E3AB0C47F8D" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from the other species of
|
||
<taxonomicName id="D7BC8350A7BAE9646CC02556D4B7BEF7" authorityName="Kaltenbach & Kluge & Gattolliat" authorityYear="2025" class="Insecta" family="Baetidae" genus="Papuanatula" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Papuafiliola">
|
||
<emphasis id="691FF82307BB823BE0D5ED16C0CC4A2E" italics="true">Papuafiliola</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DEF15DE42CC724D61F80FED037C6A21D" rank="subGenus">subgen. nov.</taxonomicNameLabel>
|
||
: absence of brown hypodermal maculae on femora and abdomen; presence of median protuberances on abdominal terga; tergalii rather wide.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EAVCM" type="description">
|
||
<paragraph id="A880CA99277AA454BDE3B4F7A2E17713">
|
||
<heading id="67332545FC96BF6A70F4186DFAA4301E" reason="title">Description.</heading>
|
||
</paragraph>
|
||
<paragraph id="7B2DC0D5B087226C4B9F5AD13D49622F">
|
||
<emphasis id="A207C69AD57FD35D46EC91E17AAE5561" bold="true">Larva</emphasis>
|
||
(Figs
|
||
<figureCitation id="AA872360878BF6D461C13BBF6A420B56" captionStart="Figure 138" captionStartId="F138" captionText="Figure 138. a, b Papuanatula (Papuafiliola) stenophylla sp. nov., eggs (extracted from mature larva) c Papuanatula (Papuafiliola) tuberculata sp. nov., larva." figureDoi="10.3897/zookeys.1227.138100.figure138" httpUri="https://binary.pensoft.net/fig/1247552">138</figureCitation>
|
||
–
|
||
<figureCitation id="F5E2D9785E64DEACFC732B2951AD2CB7" captionStart="Figure 142" captionStartId="F142" captionText="Figure 142. Papuanatula (Papuafiliola) tuberculata sp. nov., larva: a tergum III b paraprocts c antenna d tergalius IV e its enlarged anal rib (a – c holotype)." figureDoi="10.3897/zookeys.1227.138100.figure142" httpUri="https://binary.pensoft.net/fig/1247556">142</figureCitation>
|
||
).
|
||
<emphasis id="B46A5DEF6F5E80D32C1A6DA020ADD84B" bold="true">
|
||
<emphasis id="42FBA09BB97655750BFD64EA08026B6D" bold="true" italics="true">Cuticular coloration</emphasis>
|
||
.
|
||
</emphasis>
|
||
Head, pronotum and mesonotum mostly pale brownish, with some paler areas. Fore protopteron pale brownish, with narrow paler lines along concave veins (Fig.
|
||
<figureCitation id="BE5D8F56D41FF3287211BD41B4CED10E" captionStart="Figure 139" captionStartId="F139" captionText="Figure 139. Papuanatula (Papuafiliola) tuberculata sp. nov., larval exuviae (holotype, in one magnification): a head b fore, middle, and hind legs (densely covered with diatom algae) c half of pronotum and mesonotum d abdomen." figureDoi="10.3897/zookeys.1227.138100.figure139" httpUri="https://binary.pensoft.net/fig/1247553">139 a, c</figureCitation>
|
||
). Metanotum darker brownish medially, colorless laterally. Thoracic pleura pale brownish, sterna colorless. Cuticle of legs mostly colorless, with brownish outer margin and two brownish transverse bands on anterior surface: one close to base and another in apical part; cuticle of these brownish areas serrate (Figs
|
||
<figureCitation id="6563B6F6464E044E049DE162FEB04E36" captionStart="Figure 139" captionStartId="F139" captionText="Figure 139. Papuanatula (Papuafiliola) tuberculata sp. nov., larval exuviae (holotype, in one magnification): a head b fore, middle, and hind legs (densely covered with diatom algae) c half of pronotum and mesonotum d abdomen." figureDoi="10.3897/zookeys.1227.138100.figure139" httpUri="https://binary.pensoft.net/fig/1247553">139 b</figureCitation>
|
||
,
|
||
<figureCitation id="0FD8A665896A5269E847869407908D96" captionStart="Figure 141" captionStartId="F141" captionText="Figure 141. Papuanatula (Papuafiliola) tuberculata sp. nov., larva: a leg b tarsus of fore leg (holotype)." figureDoi="10.3897/zookeys.1227.138100.figure141" httpUri="https://binary.pensoft.net/fig/1247555">141 a, b</figureCitation>
|
||
). Abdominal terga mostly pale brownish, with lateral areas paler or colorless; tergum VI more or less paler than others; sterna colorless (Figs
|
||
<figureCitation id="04F82B4D000E7247E52007EF6D1EEEE5" captionStart="Figure 138" captionStartId="F138" captionText="Figure 138. a, b Papuanatula (Papuafiliola) stenophylla sp. nov., eggs (extracted from mature larva) c Papuanatula (Papuafiliola) tuberculata sp. nov., larva." figureDoi="10.3897/zookeys.1227.138100.figure138" httpUri="https://binary.pensoft.net/fig/1247552">138</figureCitation>
|
||
,
|
||
<figureCitation id="BCBAF28D6A41DCF7A08F5B85BEE7F729" captionStart="Figure 139" captionStartId="F139" captionText="Figure 139. Papuanatula (Papuafiliola) tuberculata sp. nov., larval exuviae (holotype, in one magnification): a head b fore, middle, and hind legs (densely covered with diatom algae) c half of pronotum and mesonotum d abdomen." figureDoi="10.3897/zookeys.1227.138100.figure139" httpUri="https://binary.pensoft.net/fig/1247553">139 d</figureCitation>
|
||
). Cerci uniformly pale brownish.
|
||
</paragraph>
|
||
<caption id="D5C22FDE2C137A325F7D6AB8B71FDE86" ID-DOI="10.3897/zookeys.1227.138100.figure139" ID-arpha="E11797EB-514A-576F-9ED5-12B53E90FE62" httpUri="https://binary.pensoft.net/fig/1247553" startId="F139">
|
||
<paragraph id="CEB0845BEA4C6AE556D36889F16306FA">
|
||
<label id="CAB7FA7DA7164FE5D10B8F0062704D71">Figure 139.</label>
|
||
</paragraph>
|
||
<paragraph id="364F4271A426B5964D2DE8C23F6AAF2B">
|
||
<taxonomicName id="D0D1064E51F965BB9898F281E9A0EC2E" authorityName="Kaltenbach & Kluge & Gattolliat" authorityYear="2025" class="Insecta" family="Baetidae" genus="Papuanatula" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov." subGenus="Papuafiliola">
|
||
<emphasis id="1FB2E0432137AD3FCB31BD8484763E4A" italics="true">Papuanatula</emphasis>
|
||
(
|
||
<emphasis id="1B560626D5C9952F1067479F050B0C6B" italics="true">Papuafiliola</emphasis>
|
||
)
|
||
<emphasis id="DB723C6FEE9553202A46A6FAAC7AE5AF" italics="true">tuberculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7A23E0502328482AB470F836F3979AF0" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, larval exuviae (holotype, in one magnification):
|
||
<emphasis id="124D2858D18ECE40F6AC85AF4485528D" bold="true">a</emphasis>
|
||
head
|
||
<emphasis id="56DCF61EA43FA48FBC26AF5E07382A03" bold="true">b</emphasis>
|
||
fore, middle, and hind legs (densely covered with diatom algae)
|
||
<emphasis id="E0AEE7B4A675674CBB2C96162B7E5C4C" bold="true">c</emphasis>
|
||
half of pronotum and mesonotum
|
||
<emphasis id="932C78FD3E9319259F55A905D32D2E2E" bold="true">d</emphasis>
|
||
abdomen.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5894EF1093F3A0E7B106B2C171CD5AB6">
|
||
<emphasis id="AEFCFB0C1C60CBEC4B31934A16980F63" bold="true">
|
||
<emphasis id="7581915EE268417BC3B766762701471E" bold="true" italics="true">Hypodermal coloration</emphasis>
|
||
.
|
||
</emphasis>
|
||
Not expressed (at least in penultimate larval instar). Tissues surrounding tracheae of tergalii (main trachea and its branches) with brown pigmentation (Fig.
|
||
<figureCitation id="92362CDDC71B199330067AF720603871" captionStart="Figure 142" captionStartId="F142" captionText="Figure 142. Papuanatula (Papuafiliola) tuberculata sp. nov., larva: a tergum III b paraprocts c antenna d tergalius IV e its enlarged anal rib (a – c holotype)." figureDoi="10.3897/zookeys.1227.138100.figure142" httpUri="https://binary.pensoft.net/fig/1247556">142 d, e</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8C4A2E97E1684C76FB4A868DC5FD74FB">
|
||
<emphasis id="D183629E863734F20FB9F8BEF401D267" bold="true">
|
||
<emphasis id="5EFFE7EA1C63BC5D262A0FE264CE8E65" bold="true" italics="true">Head</emphasis>
|
||
.
|
||
<emphasis id="9905273F53999F3B2ED0595F08FF49AB" bold="true" italics="true">Antenna</emphasis>
|
||
</emphasis>
|
||
(Fig.
|
||
<figureCitation id="1845E950A8A1191447993BC9AD691D83" captionStart="Figure 142" captionStartId="F142" captionText="Figure 142. Papuanatula (Papuafiliola) tuberculata sp. nov., larva: a tergum III b paraprocts c antenna d tergalius IV e its enlarged anal rib (a – c holotype)." figureDoi="10.3897/zookeys.1227.138100.figure142" httpUri="https://binary.pensoft.net/fig/1247556">142 c</figureCitation>
|
||
) with each flagellomere symmetric, cylindrical.
|
||
<emphasis id="325DA123BD4185C316F87C34A9B9752A" bold="true">
|
||
<emphasis id="DDF1604BE900431366CC97D709951987" bold="true" italics="true">Developing turbinate eyes in last instar male larva</emphasis>
|
||
.
|
||
</emphasis>
|
||
Unknown.
|
||
<emphasis id="3265B8A32BA7EBC6D046D2059CE24433" bold="true" italics="true">Labrum</emphasis>
|
||
(Fig.
|
||
<figureCitation id="B5D1834FE62735BE2CCCDE9BFB2CBABB" captionStart="Figure 140" captionStartId="F140" captionText="Figure 140. Papuanatula (Papuafiliola) tuberculata sp. nov., larva (a – c, e, h, i holotype): a labrum b, c left and right mandibles d, e maxillae f, g hypopharynx h – j labium (i focus on dorsal side)." figureDoi="10.3897/zookeys.1227.138100.figure140" httpUri="https://binary.pensoft.net/fig/1247554">140 a</figureCitation>
|
||
) short and widest at base; long setae on dorsal surface simple (not branched), constitute two pairs: submedian pair and one more pair laterad of them.
|
||
<emphasis id="00EEA1C8199C196E80E7D7910B768BC9" bold="true" italics="true">Right mandible</emphasis>
|
||
(Fig.
|
||
<figureCitation id="0209B3D15F61C2B7E179529503C237CD" captionStart="Figure 140" captionStartId="F140" captionText="Figure 140. Papuanatula (Papuafiliola) tuberculata sp. nov., larva (a – c, e, h, i holotype): a labrum b, c left and right mandibles d, e maxillae f, g hypopharynx h – j labium (i focus on dorsal side)." figureDoi="10.3897/zookeys.1227.138100.figure140" httpUri="https://binary.pensoft.net/fig/1247554">140 c</figureCitation>
|
||
) short, with incisor not elongated; incisor terminating with four denticles, among which most distal denticle shorter than others; other three denticles subequal; mola with seta on proximal corner.
|
||
<emphasis id="32DC66B287414F9E502D79DC37A85100" bold="true" italics="true">Left mandible</emphasis>
|
||
(Fig.
|
||
<figureCitation id="199702116CEBC347E7642826AF3C1E1A" captionStart="Figure 140" captionStartId="F140" captionText="Figure 140. Papuanatula (Papuafiliola) tuberculata sp. nov., larva (a – c, e, h, i holotype): a labrum b, c left and right mandibles d, e maxillae f, g hypopharynx h – j labium (i focus on dorsal side)." figureDoi="10.3897/zookeys.1227.138100.figure140" httpUri="https://binary.pensoft.net/fig/1247554">140 b</figureCitation>
|
||
) short, with incisor not elongated; incisor terminating with four denticles, among which most distal denticle shorter than others; other three denticles subequal; mola with seta on proximal corner.
|
||
<emphasis id="31433DE6DF0F1C647CC027AE7F4AD5F3" bold="true" italics="true">Hypopharynx</emphasis>
|
||
(Fig.
|
||
<figureCitation id="CDBEF37DAFD349835ABD3BEE22185DFA" captionStart="Figure 140" captionStartId="F140" captionText="Figure 140. Papuanatula (Papuafiliola) tuberculata sp. nov., larva (a – c, e, h, i holotype): a labrum b, c left and right mandibles d, e maxillae f, g hypopharynx h – j labium (i focus on dorsal side)." figureDoi="10.3897/zookeys.1227.138100.figure140" httpUri="https://binary.pensoft.net/fig/1247554">140 f, g</figureCitation>
|
||
) with bunch of long, straight, stout, setae-like spines situated on common projection.
|
||
<emphasis id="FB44FBE7212D917E57C642A46E03FDF0" bold="true" italics="true">Maxilla</emphasis>
|
||
(Fig.
|
||
<figureCitation id="6630931452AC1FA826AFA864C5214C1A" captionStart="Figure 140" captionStartId="F140" captionText="Figure 140. Papuanatula (Papuafiliola) tuberculata sp. nov., larva (a – c, e, h, i holotype): a labrum b, c left and right mandibles d, e maxillae f, g hypopharynx h – j labium (i focus on dorsal side)." figureDoi="10.3897/zookeys.1227.138100.figure140" httpUri="https://binary.pensoft.net/fig/1247554">140 d, e</figureCitation>
|
||
) short and wide; maxillary palp 2 - segmented, as long as galea-lacinia.
|
||
<emphasis id="136CF6E1424BC7688488EF7F4BF30534" bold="true" italics="true">Labium</emphasis>
|
||
(Fig.
|
||
<figureCitation id="607DE5797C2B4C576E9819D02CC29284" captionStart="Figure 140" captionStartId="F140" captionText="Figure 140. Papuanatula (Papuafiliola) tuberculata sp. nov., larva (a – c, e, h, i holotype): a labrum b, c left and right mandibles d, e maxillae f, g hypopharynx h – j labium (i focus on dorsal side)." figureDoi="10.3897/zookeys.1227.138100.figure140" httpUri="https://binary.pensoft.net/fig/1247554">140 h – j</figureCitation>
|
||
). Paraglossae widest at base, lateral side with slight concavity near base; three apical setal rows parallel to apical-lateral margin only (not bent parallel to median margin). Glossa longer than half of paraglossa, with finger-like (distal) portion much longer than triangular (proximal) portion. Ventral side of glossa with several irregularly arranged setae both on triangular and finger-like portions. Labial palp with long, narrow, arched distomedian projection on segment II; segment III with median margin shorter than lateral margin.
|
||
</paragraph>
|
||
<caption id="BA66120F5FC9C5F34D34E7DB417A2F82" ID-DOI="10.3897/zookeys.1227.138100.figure140" ID-arpha="249C9D16-B7EC-52DF-A82D-5300B64773E1" httpUri="https://binary.pensoft.net/fig/1247554" startId="F140">
|
||
<paragraph id="02748947AAAFDE38C37A46FBC47A624D">
|
||
<label id="860BE257D6E318C0F4ACADD07F7FCE8F">Figure 140.</label>
|
||
</paragraph>
|
||
<paragraph id="4B79F764E0D24E75A84A4ABED5DB94F5">
|
||
<taxonomicName id="69C829988D3387FFFE039F1CC5CB71C9" authorityName="Kaltenbach & Kluge & Gattolliat" authorityYear="2025" class="Insecta" family="Baetidae" genus="Papuanatula" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov." subGenus="Papuafiliola">
|
||
<emphasis id="82858004E0124E25F7C530706D833061" italics="true">Papuanatula</emphasis>
|
||
(
|
||
<emphasis id="43499F25D9EB99F7474BBC711442C11B" italics="true">Papuafiliola</emphasis>
|
||
)
|
||
<emphasis id="44C168336960F3189F735FFD8F16A0D5" italics="true">tuberculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CE6E95CD61B7409062F8F70886F50101" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, larva (
|
||
<emphasis id="4C5BB6ADE7A9512C0ADBD8CF2382FECB" bold="true">a – c, e, h, i</emphasis>
|
||
holotype):
|
||
<emphasis id="E0F62F9EDB3470A807E4D64BBAF58292" bold="true">a</emphasis>
|
||
labrum
|
||
<emphasis id="A2E6E52A631CED7D04EF8B6BFF272C56" bold="true">b</emphasis>
|
||
,
|
||
<emphasis id="7D23786DA24F1F1161D8BDC4C7F13418" bold="true">c</emphasis>
|
||
left and right mandibles
|
||
<emphasis id="E25EBE4C6AE8C3D800EA08B08FEFC3A6" bold="true">d, e</emphasis>
|
||
maxillae
|
||
<emphasis id="4F1CB49112D546A8B3CB9D22CC6D1BE8" bold="true">f, g</emphasis>
|
||
hypopharynx
|
||
<emphasis id="B252F630E4B683A895822C8FA3671BC3" bold="true">h – j</emphasis>
|
||
labium (
|
||
<emphasis id="6585477DE16CF2A0FD8C4E72B2720971" bold="true">i</emphasis>
|
||
focus on dorsal side).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="902FA25774E49FA352D17B5D88C5E1EE">
|
||
<emphasis id="970280549664B3B6120DD41975482F8E" bold="true">
|
||
<emphasis id="4E63E730E7DAFE54029BB6F86D0B28FE" bold="true" italics="true">Thorax</emphasis>
|
||
.
|
||
<emphasis id="2199F76476204664CF617620AB2BB6ED" bold="true" italics="true">Sterna</emphasis>
|
||
</emphasis>
|
||
without protuberances.
|
||
<emphasis id="A87D6E4EE8A7D7DD2C15119126D5B6B5" bold="true">
|
||
<emphasis id="D9D2A4577067B60A05CC6E2466FEE1A6" bold="true" italics="true">Terga</emphasis>
|
||
.
|
||
</emphasis>
|
||
Without long setae on midline; without protuberances. Metanotum without hind protoptera or their vestiges.
|
||
<emphasis id="92DD692E59835A3831E256DDFE4A0D5D" bold="true" italics="true">Legs</emphasis>
|
||
(Figs
|
||
<figureCitation id="CB0E5B340B7B1EE1B2365C22DDE71F60" captionStart="Figure 139" captionStartId="F139" captionText="Figure 139. Papuanatula (Papuafiliola) tuberculata sp. nov., larval exuviae (holotype, in one magnification): a head b fore, middle, and hind legs (densely covered with diatom algae) c half of pronotum and mesonotum d abdomen." figureDoi="10.3897/zookeys.1227.138100.figure139" httpUri="https://binary.pensoft.net/fig/1247553">139 b</figureCitation>
|
||
,
|
||
<figureCitation id="863A2B7A7DB45290C48A440D92B34432" captionStart="Figure 141" captionStartId="F141" captionText="Figure 141. Papuanatula (Papuafiliola) tuberculata sp. nov., larva: a leg b tarsus of fore leg (holotype)." figureDoi="10.3897/zookeys.1227.138100.figure141" httpUri="https://binary.pensoft.net/fig/1247555">141 a, b</figureCitation>
|
||
). Fore femur widened in proximal part; hind tibia shorter than others.
|
||
<emphasis id="233245944CE76D8FC2001140988A64EE" bold="true">
|
||
<emphasis id="87951DC8CDCFBFDB23DFCB76F449B267" bold="true" italics="true">Femur</emphasis>
|
||
.
|
||
</emphasis>
|
||
Outer side of each femur with single regular row of long, slender, flattened, parallel-sided setae with blunt apices. Cuticle of anterior surface of each femur with serrate areas corresponding with brownish pigmentation (see above).
|
||
<emphasis id="A6E0630A75EB0A498D9C26E699718374" bold="true">
|
||
<emphasis id="9193A367F7713815EC79EAF6DB8270EE" bold="true" italics="true">Tibia</emphasis>
|
||
.
|
||
</emphasis>
|
||
Patella-tibial suture present on all legs, terminated proximad of inner margin of tibia. Tibia-tarsal condylus turned to anterior side. Anterior side of each tibia with regular row of long, slender, flattened, parallel-sided setae with blunt apices, similar to that on femur, but shorter.
|
||
<emphasis id="CCD6FA2034673812BBA80DDD532986CC" bold="true">
|
||
<emphasis id="5227006C6947AE8F250A8A9FF6D6ABBC" bold="true" italics="true">Tarsus</emphasis>
|
||
.
|
||
</emphasis>
|
||
Anterior side of each tarsus with regular row of similar, but shorter setae. Posterior side of each tarsus with regular row of few small, stout, pointed setae, most distal of which not longer or slightly longer than others.
|
||
<emphasis id="A950D52DAE74FF6F25F5B5213A62B6F1" bold="true" italics="true">Claw</emphasis>
|
||
with row of 8–10 subequal denticles and with long, arched seta on posterior side.
|
||
</paragraph>
|
||
<caption id="576FA76561BD0DDB225AF2582A161C45" ID-DOI="10.3897/zookeys.1227.138100.figure141" ID-arpha="F2CF38BB-EE89-5E42-BFF4-E87CC2BF7A7C" httpUri="https://binary.pensoft.net/fig/1247555" startId="F141">
|
||
<paragraph id="41FE6EA68CAE18ABE0BF6DB9B081F706">
|
||
<label id="71953FD5D51F1997D4D01F6AD5B07157">Figure 141.</label>
|
||
</paragraph>
|
||
<paragraph id="37CF6AB27A46815994D6B3D60F3BAF3E">
|
||
<taxonomicName id="AC66EDD335159D639593AB0429A6B723" authorityName="Kaltenbach & Kluge & Gattolliat" authorityYear="2025" class="Insecta" family="Baetidae" genus="Papuanatula" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov." subGenus="Papuafiliola">
|
||
<emphasis id="EA23F455E5225780BDFA53692FDA51E1" italics="true">Papuanatula</emphasis>
|
||
(
|
||
<emphasis id="AFB3D18D36C8F33E30BB18399473E111" italics="true">Papuafiliola</emphasis>
|
||
)
|
||
<emphasis id="631478B013AD550C0429A956A1DF40AB" italics="true">tuberculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="161B2D4113B99D8F9C7C7060CDD9A747" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, larva:
|
||
<emphasis id="6619FD910BD9A20B1505EE37D5700BD1" bold="true">a</emphasis>
|
||
leg
|
||
<emphasis id="5DD7D9738BD68A3024C109A1C23C4A83" bold="true">b</emphasis>
|
||
tarsus of fore leg (holotype).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="38A782D76267B0C67B8E05E1D9B4209E">
|
||
<emphasis id="B58ED8C049279A037A300AADAAF19240" bold="true">
|
||
<emphasis id="B14F6982CDBE8918DDCA124000E586A9" bold="true" italics="true">Abdomen</emphasis>
|
||
.
|
||
<emphasis id="719B4484355F6B6F75F943E0A9172050" bold="true" italics="true">Terga</emphasis>
|
||
</emphasis>
|
||
(Figs
|
||
<figureCitation id="DF640121A413CE11139D781CD824AC2B" captionStart="Figure 138" captionStartId="F138" captionText="Figure 138. a, b Papuanatula (Papuafiliola) stenophylla sp. nov., eggs (extracted from mature larva) c Papuanatula (Papuafiliola) tuberculata sp. nov., larva." figureDoi="10.3897/zookeys.1227.138100.figure138" httpUri="https://binary.pensoft.net/fig/1247552">138</figureCitation>
|
||
,
|
||
<figureCitation id="A004157B89744F9545D4C96E9B83F12E" captionStart="Figure 139" captionStartId="F139" captionText="Figure 139. Papuanatula (Papuafiliola) tuberculata sp. nov., larval exuviae (holotype, in one magnification): a head b fore, middle, and hind legs (densely covered with diatom algae) c half of pronotum and mesonotum d abdomen." figureDoi="10.3897/zookeys.1227.138100.figure139" httpUri="https://binary.pensoft.net/fig/1247553">139 d</figureCitation>
|
||
,
|
||
<figureCitation id="81849539A857D9D82D2D13F1B03DECD5" captionStart="Figure 142" captionStartId="F142" captionText="Figure 142. Papuanatula (Papuafiliola) tuberculata sp. nov., larva: a tergum III b paraprocts c antenna d tergalius IV e its enlarged anal rib (a – c holotype)." figureDoi="10.3897/zookeys.1227.138100.figure142" httpUri="https://binary.pensoft.net/fig/1247556">142 a</figureCitation>
|
||
) without long setae on midline (e. g., in contrast to
|
||
<taxonomicName id="F238F9A0E596DEA9661FF9B7CD618E25" authorityName="Kaltenbach & Kluge & Gattolliat" authorityYear="2025" class="Insecta" family="Baetidae" genus="Papuanatula" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="obscura" status="sp. nov.">
|
||
<emphasis id="24908A8723FF4749BFF0BB9DB18EEEC0" italics="true">P. obscura</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D14433106A04E0D33D1A91C89771A42F" rank="species">sp. nov.</taxonomicNameLabel>
|
||
). Each abdominal tergum I – IX with slightly expressed median ridge elevating toward posterior margin. Surface of abdominal terga serrate, with numerous short sensilla; posterior margins of abdominal terga II – X with very small, blunt, dark brown denticles. Abdominal sterna without serrations and denticles.
|
||
<emphasis id="CA087300AAC116E7DB4FBE30BA6D56B0" bold="true" italics="true">Tergalii</emphasis>
|
||
(Fig.
|
||
<figureCitation id="5A80563E3FB5680FF2F9CB8C870C3A4B" captionStart="Figure 142" captionStartId="F142" captionText="Figure 142. Papuanatula (Papuafiliola) tuberculata sp. nov., larva: a tergum III b paraprocts c antenna d tergalius IV e its enlarged anal rib (a – c holotype)." figureDoi="10.3897/zookeys.1227.138100.figure142" httpUri="https://binary.pensoft.net/fig/1247556">142 d, e</figureCitation>
|
||
) of abdominal segment I absent; tergalii II – VII subequal, oval. Each tergalius with anal rib longer than costal rib; ribs with very small, irregularly situated denticles on dorsal side.
|
||
<emphasis id="9551AB30633C9B57E5D1A2F0CCCC7828" bold="true" italics="true">Paraproct</emphasis>
|
||
(Fig.
|
||
<figureCitation id="1CA1A1797B8A8AB900A8EE43F36408BF" captionStart="Figure 142" captionStartId="F142" captionText="Figure 142. Papuanatula (Papuafiliola) tuberculata sp. nov., larva: a tergum III b paraprocts c antenna d tergalius IV e its enlarged anal rib (a – c holotype)." figureDoi="10.3897/zookeys.1227.138100.figure142" httpUri="https://binary.pensoft.net/fig/1247556">142 b</figureCitation>
|
||
) with many small, equal denticles on median and posterior margins, without posterior prolongation.
|
||
<emphasis id="1D5DE515E5BE1AA474F9F4069368B0B4" bold="true" italics="true">Caudalii</emphasis>
|
||
(Fig.
|
||
<figureCitation id="02C3F6B8C2020E7456A08F4618A322BA" captionStart="Figure 139" captionStartId="F139" captionText="Figure 139. Papuanatula (Papuafiliola) tuberculata sp. nov., larval exuviae (holotype, in one magnification): a head b fore, middle, and hind legs (densely covered with diatom algae) c half of pronotum and mesonotum d abdomen." figureDoi="10.3897/zookeys.1227.138100.figure139" httpUri="https://binary.pensoft.net/fig/1247553">139 d</figureCitation>
|
||
). Cerci without swimming setae. Paracercus short, consisting of ~ 12 segments.
|
||
</paragraph>
|
||
<caption id="9A88E36A737FCB4AF833BC51472618B3" ID-DOI="10.3897/zookeys.1227.138100.figure142" ID-arpha="A82B7BDA-11EA-5E9A-82CD-BED53C7FAFFC" httpUri="https://binary.pensoft.net/fig/1247556" startId="F142">
|
||
<paragraph id="7846ACD6320F9CBE4F465D834B3FC72F">
|
||
<label id="421F1D7C59A8DEE9F1A254A73E72080B">Figure 142.</label>
|
||
</paragraph>
|
||
<paragraph id="4B801BB31FC1D3F5688C10A755845769">
|
||
<taxonomicName id="03549BA9A44081BBB869FA021FED5A72" authorityName="Kaltenbach & Kluge & Gattolliat" authorityYear="2025" class="Insecta" family="Baetidae" genus="Papuanatula" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov." subGenus="Papuafiliola">
|
||
<emphasis id="E087B2E51CCD78714836C45BEE211520" italics="true">Papuanatula</emphasis>
|
||
(
|
||
<emphasis id="613496DDDC8E1FE151C4ED3836BF1B09" italics="true">Papuafiliola</emphasis>
|
||
)
|
||
<emphasis id="09DFE8AE0731DEF4428F9B4B4C988C55" italics="true">tuberculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EA2FAF0071AA2529FCD11BBC67BBC9C1" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, larva:
|
||
<emphasis id="38D0A8470EC9FEBE58E1D33E4027C8B6" bold="true">a</emphasis>
|
||
tergum III
|
||
<emphasis id="751F926627F8FBA65A6D8B8A36CD2F1A" bold="true">b</emphasis>
|
||
paraprocts
|
||
<emphasis id="CF00ADD8F3DDA86561E09E9BA2BA183E" bold="true">c</emphasis>
|
||
antenna
|
||
<emphasis id="A9C58B4138661AB09E47092712296756" bold="true">d</emphasis>
|
||
tergalius IV
|
||
<emphasis id="1BCB98BF67ADB01D994557BD9B4E09D5" bold="true">e</emphasis>
|
||
its enlarged anal rib (
|
||
<emphasis id="ECDEDCB9AF1584C46BE221D0D9D9153B" bold="true">a – c</emphasis>
|
||
holotype).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E0FA0E1587E01992C198D15A10EA01A1">
|
||
<emphasis id="E6EE5B8EC5C2232E4BC81AC18B7B3666" bold="true">
|
||
<emphasis id="99F342BEDA21A1714CE1EE644311E3C9" bold="true" italics="true">Pose of subimaginal gonostyli under larval cuticle</emphasis>
|
||
.
|
||
</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<paragraph id="0B369CB5BB5FB8FB4F4267CD2194B240">
|
||
<emphasis id="7387330AB2DA2635E679499ADC21A93B" bold="true">
|
||
Subimago.
|
||
<emphasis id="6638E0CF677013FE851EA7C20066D8C8" bold="true" italics="true">Cuticular coloration</emphasis>
|
||
.
|
||
</emphasis>
|
||
Pronotum partly brownish. Mesonotum pale brown with medioparapsidal suture colorless, other sutures darker brown (Fig.
|
||
<figureCitation id="A451E5339BEF68660A090FEE6694F5F5" captionStart="Figure 143" captionStartId="F143" captionText="Figure 143. Papuanatula (Papuafiliola) tuberculata sp. nov., subimaginal exuviae (holotype): a half of mesonotum b mesopleuron c hind tarsus d fore tarsus." figureDoi="10.3897/zookeys.1227.138100.figure143" httpUri="https://binary.pensoft.net/fig/1247557">143 a</figureCitation>
|
||
). Meso- and metathoracic pleura and sterna with colorless, pale brownish and dark brown areas (Fig.
|
||
<figureCitation id="243555B28B6FC7D512B6B0FFCDC9F54E" captionStart="Figure 143" captionStartId="F143" captionText="Figure 143. Papuanatula (Papuafiliola) tuberculata sp. nov., subimaginal exuviae (holotype): a half of mesonotum b mesopleuron c hind tarsus d fore tarsus." figureDoi="10.3897/zookeys.1227.138100.figure143" httpUri="https://binary.pensoft.net/fig/1247557">143 b</figureCitation>
|
||
). Cuticle of wings colorless, with microtrichiae brownish. Legs nearly colorless, with pale brown bordering on femur and base of tibia (as in Fig.
|
||
<figureCitation id="DBFE1FB6DD14E8E0EEC800DB7A8A0D65" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Papuanatula (Papuanatula) cyclopomontana sp. nov. a head and thorax of male imago b, c fore and middle legs of male imago d subimaginal exuviae of fore leg e subimaginal exuviae of mesopleuron f subimaginal exuviae of mesonotum g abdomen of male imago h, i abdomen of female subimago, dorsally and ventrally (a – g holotype)." figureDoi="10.3897/zookeys.1227.138100.figure36" httpUri="https://binary.pensoft.net/fig/1247450">36 d</figureCitation>
|
||
). Abdomen very pale brownish. Cerci colorless with setae brownish.
|
||
</paragraph>
|
||
<caption id="715A7B1714DEE26DFE8C08C091B89F22" ID-DOI="10.3897/zookeys.1227.138100.figure143" ID-arpha="944638ED-6538-507F-94FA-858F20959AE3" httpUri="https://binary.pensoft.net/fig/1247557" startId="F143">
|
||
<paragraph id="A10B78F39F39343BE8B4E9C0978732DF">
|
||
<label id="70DF92DA055AEA424459B1951454233D">Figure 143.</label>
|
||
</paragraph>
|
||
<paragraph id="300CFA49D5E0673FB4B1092101EC0F24">
|
||
<taxonomicName id="8AF723F179566875D83688F9DB88D68B" authorityName="Kaltenbach & Kluge & Gattolliat" authorityYear="2025" class="Insecta" family="Baetidae" genus="Papuanatula" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov." subGenus="Papuafiliola">
|
||
<emphasis id="4A351193C6ADB77906BE366811832797" italics="true">Papuanatula</emphasis>
|
||
(
|
||
<emphasis id="F686570904590CA8349615B9575B78E9" italics="true">Papuafiliola</emphasis>
|
||
)
|
||
<emphasis id="4F41E70C74B4BC487843126D429602AF" italics="true">tuberculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="59FD438DF9E0C77CDE810A2436998904" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, subimaginal exuviae (holotype):
|
||
<emphasis id="C31D6E0A3BBC9E16CA3C967E477A57B5" bold="true">a</emphasis>
|
||
half of mesonotum
|
||
<emphasis id="BE6DECA69E620763F0A841E593E617AA" bold="true">b</emphasis>
|
||
mesopleuron
|
||
<emphasis id="927A57B00E008D508BCA48AA62E51B0C" bold="true">c</emphasis>
|
||
hind tarsus
|
||
<emphasis id="D7B4BFEB94B0BFB5B3D98B8288C99FB6" bold="true">d</emphasis>
|
||
fore tarsus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4052E9C65ABA36C79C42E58593608305">
|
||
<emphasis id="BAD448FCA90D182A99140CF4EF191949" bold="true">
|
||
<emphasis id="FC1A060FEC14E83D6EC1175FE35B0EFC" bold="true" italics="true">Hypodermal coloration</emphasis>
|
||
.
|
||
</emphasis>
|
||
As in imago.
|
||
</paragraph>
|
||
<paragraph id="C4401DCA9BD4A35067E3F2753D0853FC">
|
||
<emphasis id="A50F5748AA209B1A3117710D48A2BEF7" bold="true">
|
||
<emphasis id="F5D6D43EC8146A73F6041FF10D376B8E" bold="true" italics="true">Texture</emphasis>
|
||
.
|
||
</emphasis>
|
||
On all legs of female, terminal tarsomere covered with pointed microlepides only; other tarsomeres covered partly with pointed, partly with blunt microlepides (Fig.
|
||
<figureCitation id="F044D5B307247E30381C3153C2208288" captionStart="Figure 143" captionStartId="F143" captionText="Figure 143. Papuanatula (Papuafiliola) tuberculata sp. nov., subimaginal exuviae (holotype): a half of mesonotum b mesopleuron c hind tarsus d fore tarsus." figureDoi="10.3897/zookeys.1227.138100.figure143" httpUri="https://binary.pensoft.net/fig/1247557">143 c, d</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="38056093E7864F6BA18F79DF1271B2BD">
|
||
<emphasis id="155D8468A50B54185F9C8298261314FF" bold="true">
|
||
Imago.
|
||
<emphasis id="1550ED7C5F600A9392E283BBAA3B2346" bold="true" italics="true">Imago, male</emphasis>
|
||
.
|
||
</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<paragraph id="4870871DED898AF036A180710DB2E3F2">
|
||
<emphasis id="4A23E97F6AAC2B1F946A89BEDB083E2B" bold="true" italics="true">Imago, female</emphasis>
|
||
(Fig.
|
||
<figureCitation id="166B26A9D8AACC6CD2CC1AA95D0F954B" captionStart="Figure 144" captionStartId="F144" captionText="Figure 144. Papuanatula (Papuafiliola) tuberculata sp. nov., female imago (holotype): a dorsal view b lateral view c – e fore, middle and hind legs f pterostigma." figureDoi="10.3897/zookeys.1227.138100.figure144" httpUri="https://binary.pensoft.net/fig/1247559">144 a – f</figureCitation>
|
||
). Head ochre with brown. Antennae ochre, distal parts of scapus and pedicellum brown. Thorax brown with ochre, equally dark dorsally, laterally, and ventrally. Fore wing with membrane mostly colorless, veins pale ochre; base of wing, including costal brace contrastingly pigmented with brown and ochre. Pterostigma with three or four oblique, incomplete cross veins (Fig.
|
||
<figureCitation id="F44A534E639CD6B85FE86254BB2069A5" captionStart="Figure 144" captionStartId="F144" captionText="Figure 144. Papuanatula (Papuafiliola) tuberculata sp. nov., female imago (holotype): a dorsal view b lateral view c – e fore, middle and hind legs f pterostigma." figureDoi="10.3897/zookeys.1227.138100.figure144" httpUri="https://binary.pensoft.net/fig/1247559">144 f</figureCitation>
|
||
). Legs ochre; femur of each leg with wide reddish-brown band in distal part; femur of fore leg, besides this, with longitudinal reddish-brown macula occupying most part of proximal 1 / 2 (Fig.
|
||
<figureCitation id="049FB83B28678A3F58EF6C3DA7966298" captionStart="Figure 144" captionStartId="F144" captionText="Figure 144. Papuanatula (Papuafiliola) tuberculata sp. nov., female imago (holotype): a dorsal view b lateral view c – e fore, middle and hind legs f pterostigma." figureDoi="10.3897/zookeys.1227.138100.figure144" httpUri="https://binary.pensoft.net/fig/1247559">144 c – e</figureCitation>
|
||
). On each leg, tarsus with two apical spines: on 2
|
||
<superScript id="CC9FCAE846E512AA3E3D583EE4798442">nd</superScript>
|
||
and 3
|
||
<superScript id="DAECDDDCB9162D30AB014A3BF555525E">rd</superScript>
|
||
tarsomeres on fore leg, on 1
|
||
<superScript id="77A59AFC5F748143E9B11CA4C1C26711">st</superScript>
|
||
+ 2
|
||
<superScript id="2D646C4FADC6C1D3294164D86C68D0CA">nd</superScript>
|
||
and 3
|
||
<superScript id="67CBB470379F862EA26B4BCDCECEC1F2">rd</superScript>
|
||
tarsomeres on middle and hind legs. Each abdominal tergum I – IX brown with paler ochre areas; tergum X brownish ochre; sterna ochre (Fig.
|
||
<figureCitation id="0ABDAF21C6FE1F8B3E8FA0FD92932EDB" captionStart="Figure 144" captionStartId="F144" captionText="Figure 144. Papuanatula (Papuafiliola) tuberculata sp. nov., female imago (holotype): a dorsal view b lateral view c – e fore, middle and hind legs f pterostigma." figureDoi="10.3897/zookeys.1227.138100.figure144" httpUri="https://binary.pensoft.net/fig/1247559">144 a, b</figureCitation>
|
||
). Cerci pale ochre.
|
||
</paragraph>
|
||
<caption id="C48E8FDC538AA41916873369DFE42C94" ID-DOI="10.3897/zookeys.1227.138100.figure144" ID-arpha="DC72487A-8B66-5D27-8C3C-6804DE33C07B" httpUri="https://binary.pensoft.net/fig/1247559" startId="F144">
|
||
<paragraph id="3F25DF6D1C366AA8FD2DD15D0A283063">
|
||
<label id="68A46ABB8B0D1F8ACB50F55AB9411AAA">Figure 144.</label>
|
||
</paragraph>
|
||
<paragraph id="9633383705FC1036239E3C697A0AEB0B">
|
||
<taxonomicName id="393B4B9DA996EADEE30C3DF1918C6F9F" authorityName="Kaltenbach & Kluge & Gattolliat" authorityYear="2025" class="Insecta" family="Baetidae" genus="Papuanatula" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov." subGenus="Papuafiliola">
|
||
<emphasis id="4350C712C0AEFDE5FC9400962014E20E" italics="true">Papuanatula</emphasis>
|
||
(
|
||
<emphasis id="A8EBFFFE51EAF9CDBFFF20DCF7137F2B" italics="true">Papuafiliola</emphasis>
|
||
)
|
||
<emphasis id="98224DF4DB59A1035F2CA9439A947227" italics="true">tuberculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B34941014B96E6BCE0520F32170D1002" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, female imago (holotype):
|
||
<emphasis id="B4997A9A2719050F2E806DB8BD86FFE5" bold="true">a</emphasis>
|
||
dorsal view
|
||
<emphasis id="1E4EC08639337E83E6D02B4CEB7E0077" bold="true">b</emphasis>
|
||
lateral view
|
||
<emphasis id="92615FEB4E1EA2E8FCEEE8749CC447F6" bold="true">c – e</emphasis>
|
||
fore, middle and hind legs
|
||
<emphasis id="69E74FE6552CB34F2AE8BA95E705EF70" bold="true">f</emphasis>
|
||
pterostigma.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C657F15EAAD70DCEBC005717E6BA5B0C">
|
||
<emphasis id="24EB3F499510E3D29945F94FD02D3CCB" bold="true">Egg</emphasis>
|
||
(Fig.
|
||
<figureCitation id="7FD32D9669FD6C326DDD10CA9734C0A6" captionStart="Figure 145" captionStartId="F145" captionText="Figure 145. a – e Papuanatula (Papuafiliola) tuberculata sp. nov., eggs (extracted from imago)." figureDoi="10.3897/zookeys.1227.138100.figure145" httpUri="https://binary.pensoft.net/fig/1247561">145 a – e</figureCitation>
|
||
). Elongate oval. Chorion entirely and evenly covered with ridges forming a net-like relief.
|
||
</paragraph>
|
||
<caption id="BF40643CC73C0A9B8AF2B1A0DE47CB6C" ID-DOI="10.3897/zookeys.1227.138100.figure145" ID-arpha="1CC94E57-C956-58E2-8A16-C24089EECB5F" httpUri="https://binary.pensoft.net/fig/1247561" startId="F145">
|
||
<paragraph id="E59F27DE90A67470169613F6C7BCE5A5">
|
||
<label id="5BB29E37083F6533CCEFBCB980386724">Figure 145.</label>
|
||
</paragraph>
|
||
<paragraph id="568F520DCEA01F7DA40429A64CFC1A0D">
|
||
<emphasis id="531F27621134A85F16D49200321CDC7E" bold="true">a – e</emphasis>
|
||
<taxonomicName id="57D70EEAE5DDC7AA91016467AABEB099" authorityName="Kaltenbach & Kluge & Gattolliat" authorityYear="2025" class="Insecta" family="Baetidae" genus="Papuanatula" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov." subGenus="Papuafiliola">
|
||
<emphasis id="D6DA049C2FD9D518BC1CAFE8B5E356F8" italics="true">Papuanatula</emphasis>
|
||
(
|
||
<emphasis id="8E298E7AD14242E738A88B49A4C6BEAA" italics="true">Papuafiliola</emphasis>
|
||
)
|
||
<emphasis id="6DC91BEEAF413F95B0FF15653681FD39" italics="true">tuberculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="22EB15A35AEAEA9EE83C711543610CDC" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, eggs (extracted from imago).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="23C2BAFDC3E0E160DCCB914076F94205" ID-DOI="10.3897/zookeys.1227.138100.figure146" ID-arpha="2CD976F7-56A1-5B62-B14F-B71790B04BD7" httpUri="https://binary.pensoft.net/fig/1247563" startId="F146">
|
||
<paragraph id="902C87F4A521F06C94D3987F0CE6B9F6">
|
||
<label id="3351B52ACA53772B878D735DAAB55E96">Figure 146.</label>
|
||
</paragraph>
|
||
<paragraph id="A67B5D063F4AC2446ECFD909185499C2">
|
||
Distribution of
|
||
<taxonomicName id="82A3B26EF3CB7E22206EC11BE43570CB">
|
||
<emphasis id="C29196374B37CF70792724613AC6E650" italics="true">Papuanatula</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
(x: type locality).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="EA4E04569373D33316A7F62EB254BAB4" ID-DOI="10.3897/zookeys.1227.138100.figure147" ID-arpha="756D01AE-D72A-546D-81F2-EEAA34CB09DC" httpUri="https://binary.pensoft.net/fig/1247565" startId="F147">
|
||
<paragraph id="3D68EB5AE5E48E00494033A1498C811A">
|
||
<label id="E9ABBE23761919D15B5EFE68DCABBCC6">Figure 147.</label>
|
||
</paragraph>
|
||
<paragraph id="FB13DD353047B427C9FECF54392E11D8">
|
||
Distribution of
|
||
<taxonomicName id="B712B76BC8A8CEA3CA643F7EE8B83D6D">
|
||
<emphasis id="6E1B7C469FFD751E4094058EF6B64AAF" italics="true">Papuanatula</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
(x: type locality).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ELHDM" type="Dimension">
|
||
<paragraph id="5A601F8D698DD49F75E3DAB8368A88B7">
|
||
<heading id="06CC985E71046C7FC0022EB4710DFDA3" reason="title">Dimension.</heading>
|
||
</paragraph>
|
||
<paragraph id="4A79ECEAFC16802BE380A2094AC95ECE">
|
||
Fore wing length of female (and approximate body length)
|
||
<quantity id="269DE46A5AA2B66074EC520392D6A001" metricMagnitude="-2" metricUnit="m" metricValue="5.0" unit="mm" value="50.0">50 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EQHDM" type="comparisons">
|
||
<paragraph id="F173E6A34A2D6677F5AF5AAF516C2438">
|
||
<heading id="E95BFAC26DC93C8EEE1AD1F6EC2144B7" reason="title">Comparison.</heading>
|
||
</paragraph>
|
||
<paragraph id="376EFB947E94BFE2090BF93755F8FB36">
|
||
Larva of
|
||
<taxonomicName id="5C74F9ED3E64DEAB88BEB2083FDBF51B" authorityName="Kaltenbach & Kluge & Gattolliat" authorityYear="2025" class="Insecta" family="Baetidae" genus="Papuanatula" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov." subGenus="Papuafiliola">
|
||
<emphasis id="C33BC74632618A4890D784938D26AF77" italics="true">Papuanatula</emphasis>
|
||
(
|
||
<emphasis id="31436C91B4078A16C6B962EFBC519B99" italics="true">Papuafiliola</emphasis>
|
||
)
|
||
<emphasis id="CF50D669504484347B5515E019AC896D" italics="true">tuberculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="AAE83AFA88CEE7513C424F8CBC96C1C9" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="AB4228E48AC309EB23EA73EA9FB05E12" authorityName="Kaltenbach & Kluge & Gattolliat" authorityYear="2025" class="Insecta" family="Baetidae" genus="Papuanatula" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="stenophylla" status="sp. nov." subGenus="Papuafiliola">
|
||
<emphasis id="3B5640082915656687D1B651759A9163" italics="true">P.</emphasis>
|
||
(
|
||
<emphasis id="C046CB98C2E93821797C66A7F9DFA3E0" italics="true">Papuafiliola</emphasis>
|
||
)
|
||
<emphasis id="25E30F34B84A1B6090A3C7AEF18F2496" italics="true">stenophylla</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5DC828C54F7EA9FD3FEBAFB069F9228F" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by absence of brown hypodermal maculae on femora and abdomen, presence of median protuberances on abdominal terga and wider tergalii. Imago of
|
||
<taxonomicName id="388B5099A00210470AC5516668C0B15D" authorityName="Kaltenbach & Kluge & Gattolliat" authorityYear="2025" class="Insecta" family="Baetidae" genus="Papuanatula" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov." subGenus="Papuafiliola">
|
||
<emphasis id="E61D32C236721D5B52C4FFA27ED68F8D" italics="true">P.</emphasis>
|
||
(
|
||
<emphasis id="EE0411E00B35F8C2E8672FB183CF7C3A" italics="true">Papuafiliola</emphasis>
|
||
)
|
||
<emphasis id="AB8B8336406E414B9537B5A7790D3439" italics="true">tuberculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DB302491714F5A8BB0804AE2E7B58B1A" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="360B70A1310326B01F163753B5D6469B" authorityName="Kaltenbach & Kluge & Gattolliat" authorityYear="2025" class="Insecta" family="Baetidae" genus="Papuanatula" kingdom="Animalia" order="Ephemeroptera" phylum="Arthropoda" rank="species" species="stenophylla" status="sp. nov." subGenus="Papuafiliola">
|
||
<emphasis id="B960A9F0FC609CEEDFFA8ECD3C894532" italics="true">P.</emphasis>
|
||
(
|
||
<emphasis id="C98AA2A3179C6F2CCDFFA7E905F868A8" italics="true">Papuafiliola</emphasis>
|
||
)
|
||
<emphasis id="DCFD65844B5F1587657A76C2B7C268D2" italics="true">stenophylla</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BFA0D301488463B63B3DC84A3CCAEECA" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by absence of brown hypodermal maculae on proximal part of middle and hind femora.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EOJDM" type="distribution">
|
||
<paragraph id="70F744A3E3848FEC8E37026ED8BBBFE3">
|
||
<heading id="EEF7ED164BE7A3E7DBF94832E6E5F490" reason="title">Distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="5AC8B24B2A9387A9B92964AD8EBC126E">
|
||
New
|
||
<collectingCountry id="3B07C7AD3F03C73271579A2AEAE644B3" name="Guinea-Bissau">Guinea</collectingCountry>
|
||
(Fig.
|
||
<figureCitation id="E0F06D3EC5B3AF5ECCDC0C776722CAEC" captionStart="Figure 148" captionStartId="F148" captionText="Figure 148. Distribution of Papuanatula species (x: type locality)." figureDoi="10.3897/zookeys.1227.138100.figure148" httpUri="https://binary.pensoft.net/fig/1247567">148</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="61C5967A016A4C6AE54959807231E0A3" ID-DOI="10.3897/zookeys.1227.138100.figure148" ID-arpha="10E0E6B1-D09B-5460-BE50-F1018F718550" httpUri="https://binary.pensoft.net/fig/1247567" startId="F148">
|
||
<paragraph id="BE2BA12CD0B77E26B133BA164E7B8FEE">
|
||
<label id="6AD343E2A5D2D42C17AB3B820822347E">Figure 148.</label>
|
||
</paragraph>
|
||
<paragraph id="6C1AF4B8A3A3D3C00DBE2076F05EED12">
|
||
Distribution of
|
||
<taxonomicName id="729F4380F2D1CBA1B9B3F7EC9294A138">
|
||
<emphasis id="A542648D5B7BFF7B72D201CCD5484BEF" italics="true">Papuanatula</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
(x: type locality).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |