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<document ID-DOI="http://dx.doi.org/10.3897/afrinvertebr.64.91446" ID-Pensoft-Pub="2305-2562-2-139" ID-Pensoft-UUID="CC5D8054AA77554CA516D6FAF06E05DB" ID-ZooBank="C41321E7060F460EA89461759A1176F7" ModsDocID="2305-2562-64-2-139" checkinTime="1683304276267" checkinUser="pensoft" docAuthor="Sinclair, Bradley J." docDate="2023" docId="DA4228F4CC76561E88075830E535BC0A" docLanguage="en" docName="AfrInvert 64(2): 139-147" docOrigin="African Invertebrates 64 (2)" docPubDate="2023-05-05" docSource="http://dx.doi.org/10.3897/afrinvertebr.64.91446" docTitle="Wiedemannia londti Sinclair 2023, sp. nov." docType="treatment" docUuid="23A887FB-D28F-49D1-BA14-78C77CD16AC2" docUuidSource="ZooBank" docVersion="1" id="CC5D8054AA77554CA516D6FAF06E05DB" lastPageNumber="139" masterDocId="CC5D8054AA77554CA516D6FAF06E05DB" masterDocTitle="New Afrotropical species of Wiedemannia Zetterstedt (Diptera, Empididae, Clinocerinae)" masterLastPageNumber="147" masterPageNumber="139" pageNumber="139" updateTime="1683304276267" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New Afrotropical species of Wiedemannia Zetterstedt (Diptera, Empididae, Clinocerinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sinclair, Bradley J.</mods:namePart>
<mods:affiliation>Canadian National Collection of Insects, K. W. Neatby Bldg., C. E. F., 960 Carling Ave., ON, K 1 A 0 C 6, Ottawa, Canada &amp; Canadian Food Inspection Agency, OPL Entomology, K. W. Neatby Bldg., C. E. F., 960 Carling Ave., ON, K 1 A 0 C 6, Ottawa, Canada</mods:affiliation>
<mods:nameIdentifier type="email">bradley.sinclair@inspection.gc.ca</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-05-05</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>64</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>139</mods:start>
<mods:end>147</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/afrinvertebr.64.91446</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/afrinvertebr.64.91446</mods:identifier>
<mods:identifier type="Pensoft-Pub">2305-2562-2-139</mods:identifier>
<mods:identifier type="ZooBank">C41321E7060F460EA89461759A1176F7</mods:identifier>
<mods:identifier type="Pensoft-UUID">CC5D8054AA77554CA516D6FAF06E05DB</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:23A887FB-D28F-49D1-BA14-78C77CD16AC2" httpUri="http://treatment.plazi.org/id/DA4228F4CC76561E88075830E535BC0A" lastPageNumber="139" pageId="0" pageNumber="139">
<subSubSection pageId="0" pageNumber="139" type="nomenclature">
<paragraph pageId="0" pageNumber="139">
<taxonomicName LSID="https://zoobank.org/23A887FB-D28F-49D1-BA14-78C77CD16AC2" authority="Sinclair, 2023" authorityName="Sinclair" authorityYear="2023" class="Insecta" family="Empididae" genus="Wiedemannia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wiedemannia londti" order="Diptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="species" species="londti" status="sp. nov.">Wiedemannia londti</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="139">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="recognition">
<paragraph pageId="0" pageNumber="139">Recognition.</paragraph>
<paragraph pageId="0" pageNumber="139">This species is distinguished by its fully developed wings, fore coxae without erect, outstanding setae, clasping cercus with narrow base and Y-shaped apex, blade-like subepandrial sclerite and oddly shaped phallus, with reduced distiphallus.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="etymology">
<paragraph pageId="0" pageNumber="139">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="139">It is my pleasure to name this species in honour of Jason Londt. Jason was very welcoming to me as a young visitor to the Natal Museum in 1994 and I always look forward to meeting up with him at meetings of the International Congress of Dipterology.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="description">
<paragraph pageId="0" pageNumber="139">Description.</paragraph>
<paragraph pageId="0" pageNumber="139">
<emphasis bold="true" pageId="0" pageNumber="139">
Male.
<emphasis italics="true" pageId="0" pageNumber="139">Head</emphasis>
</emphasis>
: face, gena and occiput with blue pruinescence (Fig.
<figureCitation captionStart="Figures 46" captionStartId="F2" captionText="Figures 4 - 6. Afrotropical Wiedemannia, holotypes 4 W. kilimanjaro sp. nov., habitus, lateral view 5 W. kilimanjaro sp. nov., labels 6 W. londti sp. nov., habitus, lateral view 7 W. londti sp. nov., labels. Scale bar: 1.0 mm (4); 0.75 mm (6)." figureDoi="10.3897/afrinvertebr.64.91446.figures4-6" httpUri="https://binary.pensoft.net/fig/846417" pageId="0" pageNumber="139">6</figureCitation>
); vertex brown. Face with faint brownish median stripe; apex of facial cleft velvety brown with bright oval blue pruinescent patches on either side of cleft in frontal view. Ocellar triangle thinly pruinescent; ocellar setae long, slender, longer than postpronotal seta; several pairs of short posterior setulae; 4 upper postocular setae strong, spine-like, not overlapping eye; lower postocular setae slender, longer. Genal width one-third height of eye. Antenna brown; scape slightly longer than pedicel; postpedicel pointed ovate; arista-like stylus short, tip blunt, about twice as long as postpedicel.
</paragraph>
<caption doi="10.3897/afrinvertebr.64.91446.figures1-3" httpUri="https://binary.pensoft.net/fig/846416" pageId="0" pageNumber="139" start="Figures 13" startId="F1">
<paragraph pageId="0" pageNumber="139">
<emphasis bold="true" pageId="0" pageNumber="139">Figures 1-3.</emphasis>
Male terminalia of Afrotropical
<taxonomicName class="Insecta" family="Empididae" genus="Wiedemannia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wiedemannia" order="Diptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="139">Wiedemannia</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="139">1</emphasis>
<taxonomicName lsidName="W. kilimanjaro" pageId="0" pageNumber="139" rank="species" species="kilimanjaro">
<emphasis italics="true" pageId="0" pageNumber="139">W. kilimanjaro</emphasis>
</taxonomicName>
sp. nov.. lateral view
<emphasis bold="true" pageId="0" pageNumber="139">2</emphasis>
<taxonomicName lsidName="W. kilimanjaro" pageId="0" pageNumber="139" rank="species" species="kilimanjaro">
<emphasis italics="true" pageId="0" pageNumber="139">W. kilimanjaro</emphasis>
</taxonomicName>
sp. nov., clasping cercus, anterior view
<emphasis bold="true" pageId="0" pageNumber="139">3</emphasis>
<taxonomicName lsidName="W. londti" pageId="0" pageNumber="139" rank="species" species="londti">
<emphasis italics="true" pageId="0" pageNumber="139">W. londti</emphasis>
</taxonomicName>
sp. nov., lateral view. Scale bars: 0.1 mm. Abbreviations: cerc pl - cercal plate; cl cerc - clasping cercus; epand - epandrium; hypd - hypandrium; ph - phallus; sbepand scl - subepandrial sclerite; sur - surstylus.
</paragraph>
</caption>
<caption doi="10.3897/afrinvertebr.64.91446.figures4-6" httpUri="https://binary.pensoft.net/fig/846417" pageId="0" pageNumber="139" start="Figures 46" startId="F2">
<paragraph pageId="0" pageNumber="139">
<emphasis bold="true" pageId="0" pageNumber="139">Figures 4-6.</emphasis>
Afrotropical
<taxonomicName class="Insecta" family="Empididae" genus="Wiedemannia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wiedemannia" order="Diptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="139">Wiedemannia</emphasis>
</taxonomicName>
, holotypes
<emphasis bold="true" pageId="0" pageNumber="139">4</emphasis>
<taxonomicName lsidName="W. kilimanjaro" pageId="0" pageNumber="139" rank="species" species="kilimanjaro">
<emphasis italics="true" pageId="0" pageNumber="139">W. kilimanjaro</emphasis>
</taxonomicName>
sp. nov., habitus, lateral view
<emphasis bold="true" pageId="0" pageNumber="139">5</emphasis>
<taxonomicName lsidName="W. kilimanjaro" pageId="0" pageNumber="139" rank="species" species="kilimanjaro">
<emphasis italics="true" pageId="0" pageNumber="139">W. kilimanjaro</emphasis>
</taxonomicName>
sp. nov., labels
<emphasis bold="true" pageId="0" pageNumber="139">6</emphasis>
<taxonomicName lsidName="W. londti" pageId="0" pageNumber="139" rank="species" species="londti">
<emphasis italics="true" pageId="0" pageNumber="139">W. londti</emphasis>
</taxonomicName>
sp. nov., habitus, lateral view
<emphasis bold="true" pageId="0" pageNumber="139">7</emphasis>
<taxonomicName lsidName="W. londti" pageId="0" pageNumber="139" rank="species" species="londti">
<emphasis italics="true" pageId="0" pageNumber="139">W. londti</emphasis>
</taxonomicName>
sp. nov., labels. Scale bar: 1.0 mm (
<emphasis bold="true" pageId="0" pageNumber="139">4</emphasis>
); 0.75 mm (
<emphasis bold="true" pageId="0" pageNumber="139">6</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="139">
<emphasis bold="true" italics="true" pageId="0" pageNumber="139">Thorax</emphasis>
: pleura and postpronotal lobe with blue pruinescence; scutum brown with faint pair of black vittae. Antepronotum with strong, long lateral seta, nearly as long as postpronotal seta. Proepisternum with several long, pale setae. Mesonotum with 5 long dorsocentral setae, with several short setulae intermixed; several short acrostichal setulae, anterior to second dorsocentral seta; 1 postpronotal seta, with several short, dark setulae; 2 notopleural setae, lower seta weaker, shorter than upper, with numerous black setulae; 1 presutural supra-alar seta, with numerous black setulae; 1 postsutural supra-alar seta, with several setulae; 1 postalar seta; 1 pair scutellar setae, longer than dorsocentral setae, with dark marginal and discal setae and setulae. Laterotergite with cluster of pale setae.
</paragraph>
<paragraph pageId="0" pageNumber="139">
<emphasis bold="true" italics="true" pageId="0" pageNumber="139">Wing</emphasis>
: length 4.1 mm; membrane infuscate, veins darker; basal costa seta not extending to humeral crossvein. Origin of veins M1 and M2 widely separated at end of cell dm, subequal to length of crossvein r-m. Vein CuA+CuP absent. Pterostigma not visible. Costal margin with short, strong erect setae beyond apex of Sc. Halter with dark knob; base of shaft pale brown.
</paragraph>
<paragraph pageId="0" pageNumber="139">
<emphasis bold="true" italics="true" pageId="0" pageNumber="139">Legs</emphasis>
: coxae with blue pruinescence, concolourous with pleura; remaining leg segments dark brown. Fore femur with posteroventral row of fine setae, shorter than width of femur; fore tibia with erect ventral setae, shorter than width of tibia; otherwise uniformly covered with rows of small dark setulae; fore femur without strong preapical setae; fore coxa without erect setae; hind tibia with 2-3 pairs of strong ventral setae.
</paragraph>
<paragraph pageId="0" pageNumber="139">
<emphasis bold="true" italics="true" pageId="0" pageNumber="139">Abdomen</emphasis>
: blue pruinescent, concolourous with thorax, covered in short setae. Tergite 8 narrow, band-like.
</paragraph>
<paragraph pageId="0" pageNumber="139">
<emphasis bold="true" pageId="0" pageNumber="139">
<emphasis italics="true" pageId="0" pageNumber="139">Terminalia</emphasis>
(Fig.
<figureCitation captionStart="Figures 13" captionStartId="F1" captionText="Figures 1 - 3. Male terminalia of Afrotropical Wiedemannia 1 W. kilimanjaro sp. nov .. lateral view 2 W. kilimanjaro sp. nov., clasping cercus, anterior view 3 W. londti sp. nov., lateral view. Scale bars: 0.1 mm. Abbreviations: cerc pl - cercal plate; cl cerc - clasping cercus; epand - epandrium; hypd - hypandrium; ph - phallus; sbepand scl - subepandrial sclerite; sur - surstylus." figureDoi="10.3897/afrinvertebr.64.91446.figures1-3" httpUri="https://binary.pensoft.net/fig/846416" pageId="0" pageNumber="139">3</figureCitation>
)
</emphasis>
: hypandrium robust, subequal to length of epandrium. Epandrium subrectangular, produced dorsally; surstylus rounded, knob-like, with several marginal setulae. Clasping cercus arched, Y-shaped in lateral view; apical lobes, broadly rounded; inner face without peg-like setae; shorter than height of epandrium; without inner basal lobes. Subepandrial sclerite robust, blade-like, with pointed apex, projecting free beyond bases of clasping cercus. Cercal plate with 4 long setae confined to upper sclerite. Phallus straight; apex of shaft without membranous sacs, broadly rounded; apex of shaft prolonged anteroapically as slender process; distiphallus greatly reduced.
</paragraph>
<paragraph pageId="0" pageNumber="139">
<emphasis bold="true" pageId="0" pageNumber="139">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="distribution">
<paragraph pageId="0" pageNumber="139">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="139">
This species is known only from the type locality in Malawi (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F3" captionText="Figure 8. Distribution of Wiedemannia in the Afrotropical Region." figureDoi="10.3897/afrinvertebr.64.91446.figure8" httpUri="https://binary.pensoft.net/fig/846418" pageId="0" pageNumber="139">8</figureCitation>
).
</paragraph>
<caption doi="10.3897/afrinvertebr.64.91446.figure8" httpUri="https://binary.pensoft.net/fig/846418" pageId="0" pageNumber="139" start="Figure 8" startId="F3">
<paragraph pageId="0" pageNumber="139">
<emphasis bold="true" pageId="0" pageNumber="139">Figure 8.</emphasis>
Distribution of
<taxonomicName class="Insecta" family="Empididae" genus="Wiedemannia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wiedemannia" order="Diptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="139">Wiedemannia</emphasis>
</taxonomicName>
in the Afrotropical Region.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="type material">
<paragraph pageId="0" pageNumber="139">Type material.</paragraph>
<paragraph pageId="0" pageNumber="139">
<emphasis bold="true" italics="true" pageId="0" pageNumber="139">Holotype</emphasis>
♂, labelled (Fig. 7): &quot;MALAWI Mulanje Mnt./ Likabula river valley [-15.938°, 35.519°]/ 28-30.xi.1980 1535Dc/ 1000m Stuckenberg &amp;/ Londt Riverine/ Brachystegia woodland&quot;; &quot;HOLOTYPE/
<taxonomicName class="Insecta" family="Empididae" genus="Wiedemannia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wiedemannia" order="Diptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">Wiedemannia</taxonomicName>
/
<taxonomicName authorityName="Sinclair" authorityYear="2023" class="Insecta" family="Empididae" genus="Wiedemannia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wiedemannia londti" order="Diptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="species" species="londti">Wiedemannia londti</taxonomicName>
/ Sinclair&quot; [dissected] (NMSA).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="remarks">
<paragraph pageId="0" pageNumber="139">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="139">
This is the first species of
<taxonomicName class="Insecta" family="Empididae" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="family">Empididae</taxonomicName>
to be described from Malawi.
<taxonomicName authorityName="Sinclair" authorityYear="2023" class="Insecta" family="Empididae" genus="Wiedemannia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wiedemannia londti" order="Diptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="species" species="londti">
<emphasis italics="true" pageId="0" pageNumber="139">Wiedemannia londti</emphasis>
</taxonomicName>
is most similar to the odd South African species,
<taxonomicName lsidName="W. hughesi" pageId="0" pageNumber="139" rank="species" species="hughesi">
<emphasis italics="true" pageId="0" pageNumber="139">W. hughesi</emphasis>
</taxonomicName>
with absence of erect setae on the fore coxae, enlarged hypandrium and reduction of the distiphallus.
</paragraph>
</subSubSection>
</treatment>
</document>