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<document id="9228F455842E983AB3E3C4EB70D57298" ID-DOI="10.13133/2284-4880/545" ID-ISSN="2284-4880" ID-Zenodo-Dep="12764154" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721242503917" checkinUser="felipe" docAuthor="Ballerio, Alberto" docDate="2021" docId="0F7475501111FFAEFF16AC85184E96A8" docLanguage="en" docName="FragEntomol.53.2.105-282.pdf" docOrigin="Fragmenta entomologica 53 (2)" docSource="http://dx.doi.org/10.13133/2284-4880/545" docStyle="DocumentStyle:8499FF4308F512386C0E43DC290A16BD.1:FragEntomol.2021-.journal_article.0cover" docStyleId="8499FF4308F512386C0E43DC290A16BD" docStyleName="FragEntomol.2021-.journal_article.0cover" docStyleVersion="1" docTitle="Goudotostes laevis Ballerio, 2021, sp. nov." docType="treatment" docUuid="7C75918E-B152-423C-9050-22F8EEEA2427" docUuidSource="ZooBank" docVersion="2" lastPageNumber="181" masterDocId="F34D0D28115AFFE2FF9EAF31192A9224" masterDocTitle="The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)" masterLastPageNumber="282" masterPageNumber="105" pageNumber="180" updateTime="1721247602959" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
<mods:mods id="BEA4559D5D203F51B642477E9DEB70C4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4EFC32DCC3C938BF3A80A7846EAE8DFC">
<mods:title id="DE363D7D8CCF0D354EC75F63CA18D901">The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)</mods:title>
</mods:titleInfo>
<mods:name id="B188A99CC5BE0110E1930629BCEB0D58" type="personal">
<mods:role id="0DE7A950C80DC3F97E9968B11C770BA6">
<mods:roleTerm id="72D4924451EA48352684FC1ACB7BCE4B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3738E5DBB18115756A9BEB6286E7608C">Ballerio, Alberto</mods:namePart>
</mods:name>
<mods:typeOfResource id="FF75816333A55A325381A33848A5020C">text</mods:typeOfResource>
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<mods:titleInfo id="3ACEBD7ADA0EF5465BC9F68C0CCA1C60">
<mods:title id="4D4CD1A1A2EB629433B5FD8B908BB645">Fragmenta entomologica</mods:title>
</mods:titleInfo>
<mods:part id="D4B6C2A5EB66D7DE8C8B9696A556AA82">
<mods:date id="E57A5A31F711BCA9FE0AB15C7909EDF3">2021</mods:date>
<mods:detail id="927F3FD89460F44283728C7AD80BF700" type="pubDate">
<mods:number id="087283D6B2BACAEC385FC4D478EC0690">2021-11-30</mods:number>
</mods:detail>
<mods:detail id="2C671AAB9C7F1EEA1BD668FDB7809F91" type="volume">
<mods:number id="69144E46F43BF9A8990967065FE31C0A">53</mods:number>
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<mods:number id="893751DD4DDEAC6C6AF3683B05CA144C">2</mods:number>
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<mods:start id="0E7670B952203701CDC149662BB547C9">105</mods:start>
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<mods:location id="16D353F5FC8052A4902044E2FDC9B589">
<mods:url id="A09EECF7C6AA5E1C653B782209244C84">http://dx.doi.org/10.13133/2284-4880/545</mods:url>
</mods:location>
<mods:classification id="4E4BE1F21FCC846875324D8AEECE87A2">journal article</mods:classification>
<mods:identifier id="AFB0FA50F0CD7C325EC25175FE264466" type="DOI">10.13133/2284-4880/545</mods:identifier>
<mods:identifier id="C64A1AB242F029CFF6274125E689ED29" type="ISSN">2284-4880</mods:identifier>
<mods:identifier id="4E8FA0D5A24DE6762F969BB57576E482" type="Zenodo-Dep">12764154</mods:identifier>
<mods:identifier id="CFFF074C9550209ED0F1CB27FBE08E93" type="ZooBank">B3F75A2C-7603-4027-B8E1-F4A8B5A0CDD9</mods:identifier>
</mods:mods>
<treatment id="0F7475501111FFAEFF16AC85184E96A8" LSID="urn:lsid:zoobank.org:act:7C75918E-B152-423C-9050-22F8EEEA2427" httpUri="http://treatment.plazi.org/id/0F7475501111FFAEFF16AC85184E96A8" lastPageId="76" lastPageNumber="181" pageId="75" pageNumber="180">
<subSubSection id="CFC797CD1111FFA9FF16AC85188491E8" box="[136,430,948,972]" pageId="75" pageNumber="180" type="nomenclature">
<paragraph id="8762C4461111FFA9FF16AC85188491E8" blockId="75.[136,430,948,972]" box="[136,430,948,972]" pageId="75" pageNumber="180">
<heading id="DC2A732A1111FFA9FF16AC85188491E8" box="[136,430,948,972]" fontSize="10" level="2" pageId="75" pageNumber="180" reason="5">
<taxonomicName id="40DDBFC51111FFA9FF16AC85187891E8" ID-CoL="8SSN2" authorityName="Ballerio" authorityYear="2021" box="[136,338,948,972]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="180" phylum="Arthropoda" rank="species" species="laevis" status="sp. nov.">
<emphasis id="B5A918541111FFA9FF16AC85187891E8" box="[136,338,948,972]" italics="true" pageId="75" pageNumber="180">Goudotostes laevis</emphasis>
</taxonomicName>
<emphasis id="B5A918541111FFA9FEC7AC85188491E8" bold="true" box="[345,430,948,972]" pageId="75" pageNumber="180">
<taxonomicNameLabel id="AE9AA52F1111FFA9FEC7AC85188491E8" box="[345,430,948,972]" pageId="75" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CFC797CD1111FFA9FF16ACE518D69608" pageId="75" pageNumber="180" type="description">
<paragraph id="8762C4461111FFA9FF16ACE518D69608" blockId="75.[136,756,980,1068]" pageId="75" pageNumber="180">
(
<figureCitation id="1FE6D8C31111FFA9FF0EACE519D891C8" box="[144,242,980,1004]" captionStart="Fig" captionStartId="14.[151,182,1885,1904]" captionTargetBox="[153,1410,191,1847]" captionTargetId="figure-13@14.[148,1416,163,1855]" captionTargetPageId="14" captionText="Fig. 10 – Wings. a, Synarmostes tibialis; b, Synarmostes occidentalis sp. nov.; c, Synarmostes sp. C; d, Philharmostes metallicus sp. nov.; e, Synarmostes makirovanae sp. nov.; f, Goudotostes laevis sp. nov.." figureDoi="http://doi.org/10.5281/zenodo.12764184" httpUri="https://zenodo.org/record/12764184/files/figure.png" pageId="75" pageNumber="180">Figs 10 f</figureCitation>
,
<figureCitation id="1FE6D8C31111FFA9FF63ACE5183291C8" box="[253,280,980,1004]" captionStart="Fig" captionStartId="129.[136,167,1858,1877]" captionTargetBox="[148,1397,170,1835]" captionTargetId="figure-4@129.[134,1414,156,1835]" captionTargetPageId="129" captionText="Fig. 64 – Goudotostes laevis sp. nov., a, female paratype, dorsal view of distended specimen; b, paratype, ventral view of rolled up specimen; c, paratype, dorsal view of rolled up specimen; d, paratype, ventral view of rolled up specimen; e, paratype, lateral view of rolled up specimen; f, holotype, spiculum gastrale; g–j, holotype, four views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764309" httpUri="https://zenodo.org/record/12764309/files/figure.png" pageId="75" pageNumber="180">64</figureCitation>
a-j, 77) http://zoobank.org/urn:lsid:zoobank.org:act:7C75918EB152-423C-9050-22F8EEEA2427
</paragraph>
</subSubSection>
<subSubSection id="CFC797CD1111FFA9FF30AB65183D94C8" pageId="75" pageNumber="180" type="materials_examined">
<paragraph id="8762C4461111FFA9FF30AB65183D94C8" blockId="75.[135,758,1108,1932]" pageId="75" pageNumber="180">
<emphasis id="B5A918541111FFA9FF30AB65181A9648" bold="true" box="[174,304,1108,1132]" pageId="75" pageNumber="180">Type series.</emphasis>
<materialsCitation id="37B5CE1B1111FFA9FEABAB6518BA9768" collectingDate="2011-04-28" collectingDateMax="2011-04-29" collectingDateMin="2011-04-28" collectorName="San Francisco, CA &amp; B. L. Fisher" country="Madagascar" county="Academy of Sciences" determinerName="A. Ballerio" elevation="415" latitude="-14.170556" location="California Acad. of Sciences" longLatPrecision="21" longitude="49.954166" municipality="Makirovana Forest" pageId="75" pageNumber="180" specimenCount="1" specimenCount-male="1" stateProvince="Antsiranana" typeStatus="holotype">
<typeStatus id="58667AE41111FFA9FEABAB6518B19648" box="[309,411,1108,1132]" pageId="75" pageNumber="180" type="holotype">Holotype</typeStatus>
,
<specimenCount id="91DB0FCF1111FFA9FE3AAB6418909648" box="[420,442,1109,1132]" count="1" pageId="75" pageNumber="180" type="male">♂</specimenCount>
, deposited in the collection of the California
<collectingCounty id="6E03BCCA1111FFA9FEBBAB451B3996A8" box="[293,531,1140,1164]" pageId="75" pageNumber="180">Academy of Sciences</collectingCounty>
,
<collectorName id="2A28A1901111FFA9FDBEAB451BDA96A8" box="[544,752,1140,1164]" pageId="75" pageNumber="180">San Francisco, CA</collectorName>
, USA, labelled as follows: / CASTYPE 20268
<emphasis id="B5A918541111FFA9FDE2ABA51BA99688" bold="true" box="[636,643,1172,1196]" pageId="75" pageNumber="180">/</emphasis>
<collectingCountry id="FFCA84D61111FFA9FD15ABA5198796E8" name="Madagascar" pageId="75" pageNumber="180">Madagascar</collectingCountry>
:
<collectingRegion id="45190AA41111FFA9FF28AB85181D96E8" box="[182,311,1204,1228]" country="Madagascar" name="Antsiranana" pageId="75" pageNumber="180">Antsiranana</collectingRegion>
<collectingMunicipality id="67065E3C1111FFA9FEA0AB851B2396E8" box="[318,521,1204,1228]" pageId="75" pageNumber="180">Makirovana Forest</collectingMunicipality>
<date id="F363E2861111FFA9FD8EAB851BED96E8" box="[528,711,1204,1228]" pageId="75" pageNumber="180" value="2011-04-28" valueMax="2011-04-29" valueMin="2011-04-28">
<collectingDate id="E3271B6E1111FFA9FD8EAB851BED96E8" box="[528,711,1204,1228]" pageId="75" pageNumber="180" value="2011-04-28" valueMax="2011-04-29" valueMin="2011-04-28">28-29 April 2011</collectingDate>
</date>
<geoCoordinate id="E2E9A2811111FFA9FD51AB8519C196C8" degrees="14" direction="south" minutes="10" orientation="latitude" pageId="75" pageNumber="180" precision="15" seconds="14" value="-14.170556">14° 10’ 14” S</geoCoordinate>
<geoCoordinate id="E2E9A2811111FFA9FF6EABE518A196C8" box="[240,395,1236,1260]" degrees="049" direction="east" minutes="57" orientation="longitude" pageId="75" pageNumber="180" precision="15" seconds="15" value="49.954166">049° 57’ 15” E</geoCoordinate>
<location id="8202929D1111FFA9FE0FABE51BEB96C8" LSID="urn:lsid:plazi:treatment:0F7475501111FFAEFF16AC85184E96A8:8202929D1111FFA9FE0FABE51BEB96C8" box="[401,705,1236,1260]" country="Madagascar" county="Academy of Sciences" latitude="-14.170556" longLatPrecision="21" longitude="49.954166" municipality="Makirovana Forest" name="California Acad. of Sciences" pageId="75" pageNumber="180" stateProvince="Antsiranana">California Acad. of Sciences</location>
, collectors:
<collectorName id="2A28A1901111FFA9FF41ABC418709728" box="[223,346,1268,1292]" pageId="75" pageNumber="180">B.L. Fisher</collectorName>
et al. sifted leaf litter, rainforest elev.
<quantity id="402569A31111FFA9FF16AA2519E79708" box="[136,205,1300,1324]" metricMagnitude="2" metricUnit="m" metricValue="4.15" pageId="75" pageNumber="180" unit="m" value="415.0">
<elevation id="0CF023751111FFA9FF16AA2519E79708" box="[136,205,1300,1324]" metricMagnitude="2" metricUnit="m" metricValue="4.15" pageId="75" pageNumber="180" unit="m" value="415.0">415 m</elevation>
</quantity>
, collection code: BLF 26523 /
<taxonomicName id="40DDBFC51111FFA9FD8EAA251BF39708" authorityName="Ballerio" authorityYear="2021" box="[528,729,1300,1324]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="180" phylum="Arthropoda" rank="species" species="laevis" status="sp. nov.">
<emphasis id="B5A918541111FFA9FD8EAA251BF39708" box="[528,729,1300,1324]" italics="true" pageId="75" pageNumber="180">Goudotostes laevis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AE9AA52F1111FFA9FD7EAA24198C9768" pageId="75" pageNumber="180" rank="species">n. sp.</taxonomicNameLabel>
det.
<determinerName id="72BE28821111FFA9FF45AA0518799768" box="[219,339,1332,1356]" pageId="75" pageNumber="180">A. Ballerio</determinerName>
2020
</materialsCitation>
<typeStatus id="58667AE41111FFA9FE06AA051B2D9768" box="[408,519,1332,1356]" pageId="75" pageNumber="180" type="holotype">Holotypus</typeStatus>
<specimenCount id="91DB0FCF1111FFA9FD91AA041B099768" box="[527,547,1333,1356]" count="1" pageId="75" pageNumber="180" type="male">♂</specimenCount>
/ [distended, glued on card, dissected, male genitalia glued in DMHF resin on a separate card, same pin].
<materialsCitation id="37B5CE1B1111FFA9FE41AA44182994C8" collectingDate="2011-05-01" collectingDateMax="2011-05-02" collectingDateMin="2011-05-01" collectionCode="CASC, ABCB" collectorName="B. L. Fisher" country="Madagascar" county="Makirovana Forest" determinerName="A. Ballerio" elevation="550" latitude="-14.166667" location="Makirovana Forest" longLatPrecision="21" longitude="49.95" municipality="California Acad. of Sciences" pageId="75" pageNumber="180" specimenCount="3" specimenCount-male="3" stateProvince="Antsiranana" typeStatus="paratype">
<typeStatus id="58667AE41111FFA9FE41AA441B6D97A9" box="[479,583,1397,1421]" pageId="75" pageNumber="180" type="paratype">Paratypes</typeStatus>
[
<specimenCount id="91DB0FCF1111FFA9FDC6AA451BB297A8" box="[600,664,1396,1420]" count="3" pageId="75" pageNumber="180" type="male">3 ♂♂</specimenCount>
dissect- ed]: 8 exx. (
<quantity id="402569A31111FFA9FE92AAA5181D9788" box="[268,311,1428,1452]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="75" pageNumber="180" unit="in" value="6.0">6 in</quantity>
<collectionCode id="E1CC5C831111FFA9FEA1AAA518AF9788" box="[319,389,1428,1452]" pageId="75" pageNumber="180">CASC</collectionCode>
and
<quantity id="402569A31111FFA9FE25AAA518CC9788" box="[443,486,1428,1452]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="75" pageNumber="180" unit="in" value="2.0">2 in</quantity>
<collectionCode id="E1CC5C831111FFA9FE72AAA51B139788" box="[492,569,1428,1452]" pageId="75" pageNumber="180">ABCB</collectionCode>
), same collecting data as holotype. 6 exx. (
<quantity id="402569A31111FFA9FE3FAA8518E497E8" box="[417,462,1460,1484]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="75" pageNumber="180" unit="in" value="4.0">4 in</quantity>
<collectionCode id="E1CC5C831111FFA9FE49AA851B3797E8" box="[471,541,1460,1484]" pageId="75" pageNumber="180">CASC</collectionCode>
and
<quantity id="402569A31111FFA9FDC8AA851BA897E8" box="[598,642,1460,1484]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="75" pageNumber="180" unit="in" value="2.0">2 in</quantity>
<collectionCode id="E1CC5C831111FFA9FD14AA851BF297E8" box="[650,728,1460,1484]" pageId="75" pageNumber="180">ABCB</collectionCode>
): /
<collectingCountry id="FFCA84D61111FFA9FF16AAE5182797C8" box="[136,269,1492,1516]" name="Madagascar" pageId="75" pageNumber="180">Madagascar</collectingCountry>
:
<collectingRegion id="45190AA41111FFA9FE89AAE518B297C8" box="[279,408,1492,1516]" country="Madagascar" name="Antsiranana" pageId="75" pageNumber="180">Antsiranana</collectingRegion>
<collectingCounty id="6E03BCCA1111FFA9FE3FAAE51B4797C8" box="[417,621,1492,1516]" pageId="75" pageNumber="180">Makirovana Forest</collectingCounty>
elev.
<quantity id="402569A31111FFA9FD31AAE51BDE97C8" box="[687,756,1492,1516]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="75" pageNumber="180" unit="m" value="550.0">
<elevation id="0CF023751111FFA9FD31AAE51BDE97C8" box="[687,756,1492,1516]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="75" pageNumber="180" unit="m" value="550.0">550 m</elevation>
</quantity>
<date id="F363E2861111FFA9FF14AAC5180D9428" box="[138,295,1524,1549]" pageId="75" pageNumber="180" value="2011-05-01" valueMax="2011-05-02" valueMin="2011-05-01">
<collectingDate id="E3271B6E1111FFA9FF14AAC5180D9428" box="[138,295,1524,1549]" pageId="75" pageNumber="180" value="2011-05-01" valueMax="2011-05-02" valueMin="2011-05-01">1-2 May 2011</collectingDate>
</date>
<geoCoordinate id="E2E9A2811111FFA9FEAFAAC518E69428" box="[305,460,1524,1548]" degrees="14" direction="south" minutes="09" orientation="latitude" pageId="75" pageNumber="180" precision="15" seconds="38" value="-14.160555">14° 09’ 38” S</geoCoordinate>
<geoCoordinate id="E2E9A2811111FFA9FE48AAC51B559428" box="[470,639,1524,1548]" degrees="049" direction="east" minutes="57" orientation="longitude" pageId="75" pageNumber="180" precision="15" seconds="08" value="49.952225">049° 57’ 08” E</geoCoordinate>
<collectingMunicipality id="67065E3C1111FFA9FD14AAC518619408" pageId="75" pageNumber="180">California Acad. of Sciences</collectingMunicipality>
, collectors:
<collectorName id="2A28A1901111FFA9FE52A9241B6F9408" box="[460,581,1556,1580]" pageId="75" pageNumber="180">B.L. Fisher</collectorName>
et al. rainforest, sifted leaf litter collection code: BLF 26937 /. 4 exx. (
<quantity id="402569A31111FFA9FD55A9051BDE9468" box="[715,756,1588,1612]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="75" pageNumber="180" unit="in" value="3.0">3 in</quantity>
<collectionCode id="E1CC5C831111FFA9FF16A96519E49448" box="[136,206,1620,1644]" pageId="75" pageNumber="180">CASC</collectionCode>
and
<quantity id="402569A31111FFA9FE99A96518189448" box="[263,306,1620,1644]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="75" pageNumber="180" unit="in" value="1.0">1 in</quantity>
<collectionCode id="E1CC5C831111FFA9FEA7A96518A99448" box="[313,387,1620,1644]" pageId="75" pageNumber="180">ABCB</collectionCode>
):
<emphasis id="B5A918541111FFA9FE03A965188E9448" bold="true" box="[413,420,1620,1644]" pageId="75" pageNumber="180">/</emphasis>
<collectingCountry id="FFCA84D61111FFA9FE30A9651B199448" box="[430,563,1620,1644]" name="Madagascar" pageId="75" pageNumber="180">Madagascar</collectingCountry>
:
<collectingRegion id="45190AA41111FFA9FDA3A9651B949448" box="[573,702,1620,1644]" country="Madagascar" name="Antsiranana" pageId="75" pageNumber="180">Antsiranana</collectingRegion>
<location id="8202929D1111FFA9FD59A964180594A8" LSID="urn:lsid:plazi:treatment:0F7475501111FFAEFF16AC85184E96A8:8202929D1111FFA9FD59A964180594A8" country="Madagascar" county="Makirovana Forest" latitude="-14.166667" longLatPrecision="21" longitude="49.95" municipality="California Acad. of Sciences" name="Makirovana Forest" pageId="75" pageNumber="180" stateProvince="Antsiranana">Makirovana Forest</location>
<date id="F363E2861111FFA9FEA8A94518E694A8" box="[310,460,1652,1677]" pageId="75" pageNumber="180" value="2011-05-01" valueMax="2011-05-02" valueMin="2011-05-01">
<collectingDate id="E3271B6E1111FFA9FEA8A94518E694A8" box="[310,460,1652,1677]" pageId="75" pageNumber="180" value="2011-05-01" valueMax="2011-05-02" valueMin="2011-05-01">1-2 May 2011</collectingDate>
</date>
<geoCoordinate id="E2E9A2811111FFA9FE4DA9451B4F94A8" box="[467,613,1652,1676]" degrees="14" direction="south" minutes="10" orientation="latitude" pageId="75" pageNumber="180" precision="15" seconds="00" value="-14.166667">14° 10’ 00” S</geoCoordinate>
<geoCoordinate id="E2E9A2811111FFA9FDF5A94519B29488" degrees="049" direction="east" minutes="57" orientation="longitude" pageId="75" pageNumber="180" precision="15" seconds="00" value="49.95">049° 57’ 00” E</geoCoordinate>
<location id="8202929D1111FFA9FF3FA9A518F09488" LSID="urn:lsid:plazi:treatment:0F7475501111FFAEFF16AC85184E96A8:8202929D1111FFA9FF3FA9A518F09488" box="[161,474,1684,1708]" country="Madagascar" county="Makirovana Forest" latitude="-14.166667" longLatPrecision="21" longitude="49.95" municipality="California Acad. of Sciences" name="California Acad. of Sciences" pageId="75" pageNumber="180" stateProvince="Antsiranana">California Acad. of Sciences</location>
, collectors:
<collectorName id="2A28A1901111FFA9FDC0A9A41BF39488" box="[606,729,1684,1708]" pageId="75" pageNumber="180">B.L. Fisher</collectorName>
et al. sifted leaf litter, rainforest elev.
<quantity id="402569A31111FFA9FE65A9851B6A94E8" box="[507,576,1716,1740]" metricMagnitude="2" metricUnit="m" metricValue="7.15" pageId="75" pageNumber="180" unit="m" value="715.0">
<elevation id="0CF023751111FFA9FE65A9851B6A94E8" box="[507,576,1716,1740]" metricMagnitude="2" metricUnit="m" metricValue="7.15" pageId="75" pageNumber="180" unit="m" value="715.0">715 m</elevation>
</quantity>
, collection code: BLF 26827
</materialsCitation>
/.
</paragraph>
</subSubSection>
<subSubSection id="CFC797CD1111FFA9FF30A9C51DE991A8" pageId="75" pageNumber="180" type="diagnosis">
<paragraph id="8762C4461111FFA9FF30A9C51DE991A8" blockId="75.[135,758,1108,1932]" lastBlockId="75.[816,1438,180,1932]" pageId="75" pageNumber="180">
<emphasis id="B5A918541111FFA9FF30A9C518099528" bold="true" box="[174,291,1780,1804]" pageId="75" pageNumber="180">Diagnosis.</emphasis>
<taxonomicName id="40DDBFC51111FFA9FEB5A9C518DD9528" authorityName="Ballerio" authorityYear="2021" box="[299,503,1780,1804]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="180" phylum="Arthropoda" rank="species" species="laevis" status="sp. nov.">
<emphasis id="B5A918541111FFA9FEB5A9C518DD9528" box="[299,503,1780,1804]" italics="true" pageId="75" pageNumber="180">Goudotostes laevis</emphasis>
</taxonomicName>
<emphasis id="B5A918541111FFA9FD9EA9C51B729528" bold="true" box="[512,600,1780,1804]" pageId="75" pageNumber="180">
<taxonomicNameLabel id="AE9AA52F1111FFA9FD9EA9C51B729528" box="[512,600,1780,1804]" pageId="75" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be differentiated from all other species of
<taxonomicName id="40DDBFC51111FFA9FE6BA8251B509508" authorityName=", sensu Ballerio" authorityYear="2008" box="[501,634,1812,1836]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="180" phylum="Arthropoda" rank="genus">
<emphasis id="B5A918541111FFA9FE6BA8251B509508" box="[501,634,1812,1836]" italics="true" pageId="75" pageNumber="180">Goudotostes</emphasis>
</taxonomicName>
by the following combination of characters: a) small size, b) dorsal ocular area absent, c) clypeal apical projection pointed, d) clypeal punctation made of large impressed simple punctures, e) fronto-clypeal tubercle present but weak, f) pronotal anterior carina bilobate, g) pronotal basal carina bilobate, h) pronotal disc covered by sparse large simple impressed punctures, i) elytral dorsal area covered by longitudinal irregular blunt carinae, sometimes interrupt- ed multiple times, with regular profile in lateral view, j) elytral first interstria (area between sutural stria and first longitudinal carina) with sparse punctation (interpunctural distance superior than punctural diameter), k) elytral peridiscal area with sparse punctation (interpunctural distance superior than punctural diameter), l) left paramere with ventral margin invaginated (a character which places this species in the
<taxonomicName id="40DDBFC51111FFA9FC50AD051D269068" authorityName="Ballerio" authorityYear="2021" box="[974,1036,564,588]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="180" phylum="Arthropoda" rank="species" species="laevis">
<emphasis id="B5A918541111FFA9FC50AD051D269068" box="[974,1036,564,588]" italics="true" pageId="75" pageNumber="180">laevis</emphasis>
</taxonomicName>
group of species), m) the distinctive shape of parameres (as in
<figureCitation id="1FE6D8C31111FFA9FBD1AD651D8B9048" box="[1103,1185,596,620]" captionStart="Fig" captionStartId="129.[136,167,1858,1877]" captionTargetBox="[148,1397,170,1835]" captionTargetId="figure-4@129.[134,1414,156,1835]" captionTargetPageId="129" captionText="Fig. 64 – Goudotostes laevis sp. nov., a, female paratype, dorsal view of distended specimen; b, paratype, ventral view of rolled up specimen; c, paratype, dorsal view of rolled up specimen; d, paratype, ventral view of rolled up specimen; e, paratype, lateral view of rolled up specimen; f, holotype, spiculum gastrale; g–j, holotype, four views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764309" httpUri="https://zenodo.org/record/12764309/files/figure.png" pageId="75" pageNumber="180">Figs 64</figureCitation>
g-j). The species more similar to it are
<taxonomicName id="40DDBFC51111FFA9FC77AD451C1290A8" authorityName="Ballerio" authorityYear="2021" box="[1001,1336,628,652]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="180" phylum="Arthropoda" rank="species" species="rafanomezantsoai" status="sp. nov.">
<emphasis id="B5A918541111FFA9FC77AD451C1290A8" box="[1001,1336,628,652]" italics="true" pageId="75" pageNumber="180">Goudotostes rafanomezantsoai</emphasis>
</taxonomicName>
<emphasis id="B5A918541111FFA9FADDAD451CB790A8" bold="true" box="[1347,1437,628,652]" pageId="75" pageNumber="180">
<taxonomicNameLabel id="AE9AA52F1111FFA9FADDAD451CB790A8" box="[1347,1437,628,652]" pageId="75" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="40DDBFC51111FFA9FCC3ADA51D469088" authorityName="Ballerio" authorityYear="2021" box="[861,1132,660,684]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="180" phylum="Arthropoda" rank="species" species="rajemisonae" status="sp. nov.">
<emphasis id="B5A918541111FFA9FCC3ADA51D469088" box="[861,1132,660,684]" italics="true" pageId="75" pageNumber="180">Goudotostes rajemisonae</emphasis>
</taxonomicName>
<emphasis id="B5A918541111FFA9FBECADA51DED9088" bold="true" box="[1138,1223,660,684]" pageId="75" pageNumber="180">
<taxonomicNameLabel id="AE9AA52F1111FFA9FBECADA51DED9088" box="[1138,1223,660,684]" pageId="75" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
.
<taxonomicName id="40DDBFC51111FFA9FB4DADA51CB69088" authorityName="Ballerio" authorityYear="2021" box="[1235,1436,660,684]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="180" phylum="Arthropoda" rank="species" species="laevis" status="sp. nov.">
<emphasis id="B5A918541111FFA9FB4DADA51CB69088" box="[1235,1436,660,684]" italics="true" pageId="75" pageNumber="180">Goudotostes laevis</emphasis>
</taxonomicName>
<emphasis id="B5A918541111FFA9FCAEAD851AAF90E8" bold="true" box="[816,901,692,716]" pageId="75" pageNumber="180">
<taxonomicNameLabel id="AE9AA52F1111FFA9FCAEAD851AAF90E8" box="[816,901,692,716]" pageId="75" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from them because of the elytral first interstria with sparse punctation (interpunctural distance superior than punctural diameter), elytral peridiscal area with sparse punctation (interpunctural distance superior than punctural diameter), elytral carinae longer, left paramere longer and more strongly curved outwards and spiculum of spiculum gastrale distinctly longer.
</paragraph>
</subSubSection>
<subSubSection id="CFC797CD1111FFAEFCC8ACA51B289188" lastPageId="76" lastPageNumber="181" pageId="75" pageNumber="180" type="description">
<paragraph id="8762C4461111FFAEFCC8ACA51B289188" blockId="75.[816,1438,180,1932]" lastBlockId="76.[151,772,180,1164]" lastPageId="76" lastPageNumber="181" pageId="75" pageNumber="180">
<emphasis id="B5A918541111FFA9FCC8ACA51AC99188" bold="true" box="[854,995,916,940]" pageId="75" pageNumber="180">Description.</emphasis>
Size: HL =
<quantity id="402569A31111FFA9FBF5ACA51DE89188" box="[1131,1218,916,940]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="75" pageNumber="180" unit="mm" value="0.8">0.8 mm</quantity>
; HW =
<quantity id="402569A31111FFA9FA84ACA51C5B9188" box="[1306,1393,916,940]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="75" pageNumber="180" unit="mm" value="1.2">1.2 mm</quantity>
; PL =
<quantity id="402569A31111FFA9FCD6AC851AB591E8" box="[840,927,948,972]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="75" pageNumber="180" unit="mm" value="1.2">1.2 mm</quantity>
; PW = 2.0 mm; EL = 2.0 mm; EW = 2.0 mm. Overall morphology as in generic diagnosis. Small-sized
<taxonomicName id="40DDBFC51111FFA9FCAFACC51A929628" authorityName=", sensu Ballerio" authorityYear="2008" box="[817,952,1012,1036]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="180" phylum="Arthropoda" rank="genus">
<emphasis id="B5A918541111FFA9FCAFACC51A929628" box="[817,952,1012,1036]" italics="true" pageId="75" pageNumber="180">Goudotostes</emphasis>
</taxonomicName>
. Body strongly convex. Black, shiny, dorsum with extremely short fine setation (50×), underside, tarsi and antennae dark reddish-brown, setation yellowish-brown.
<emphasis id="B5A918541111FFA9FC33AB651ACD9648" box="[941,999,1108,1132]" italics="true" pageId="75" pageNumber="180">Head</emphasis>
: W/L ratio = 1.49; clypeal apex obtuse, apical projection strongly developed and sexually dimorphic, sides almost rectilinear with slightly unaligned genae in correspondence with the genal suture; dorsal ocular area absent, genal canthus indistinct, ventral ocular area small-sized, genae acute, distinctly protruding outwards, genal suture indistinct; head dorsal surface uniformly covered by dense large impressed simple punctures dens- er on clypeal disc, fore margin marked by few irregular transverse lines. Interpunctural distance of punctures inferior than punctural diameter on clypeal disc, about as long as punctural diameter at sides, and on frons and vertex. Fronto-clypeal tubercle present and weak.
<emphasis id="B5A918541111FFA9FAB4AAE41CBF97C8" box="[1322,1429,1493,1516]" italics="true" pageId="75" pageNumber="180">Pronotum</emphasis>
: subrectangular (dorsal view) (W/L ratio = 1.88), convex, fore angles gently subtruncate; pronotal surface covered by dense large deep simple punctures becoming sparser smaller and comma-shaped at sides, area near fore angles with few irregular lines, interpunctural distance mostly as long as punctural diameter; pronotal bead complete although not visible from above along base; pronotal sculpturing made of large and gently raised tubercles and carinae as follows: bilobate posterior carina weakly raised, sides of disc with one gently raised transverse short carina and another gently raised transverse carina reaching outer margin, anterior bilobate carina weakly raised.
<taxonomicName id="40DDBFC51111FFA9FAB5A8651CBF9548" box="[1323,1429,1876,1900]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="75" pageNumber="180" phylum="Arthropoda" rank="genus">
<emphasis id="B5A918541111FFA9FAB5A8651CBF9548" box="[1323,1429,1876,1900]" italics="true" pageId="75" pageNumber="180">Scutellum</emphasis>
</taxonomicName>
: basal area and sides smooth, remaining surface covered by impressed transverse large comma-shaped punctures.
<emphasis id="B5A918541116FFAEFF09AFE519F092C8" box="[151,218,212,236]" italics="true" pageId="76" pageNumber="181">Elytra</emphasis>
: longer than wide (W/L ratio = 0.96), subovoidal (dorsal view), strongly convex; humeral callus marked by a tubercle; elytral surface covered by large deep sparse comma-shaped punctures with opening directed backwards, inwards or outwards, interpunctural distance spanning from almost equal to up to three times punctural diameter. Striated articular area hardly visible in lateral view, relatively narrow and short; inferior sutural stria visible and complete; marginal area large-sized; elytral articular process well developed, smooth and shiny, isolated from remaining elytral surface by a deep furrow. Elytral sculpturing as follows: base with one raised large tubercle below articular process with apex directed backwards (parasutural tubercle), next to it a humeral oblique larger raised tubercle with apex directed backwards (humeral tubercle), dorsal area of elytra with five (including elytral suture) irregular and asymmetrical rows of weakly raised smooth carinae, sometimes interrupted multiple times, sides and apical third with some more raised tubercles and longitudinal short carinae.
<emphasis id="B5A918541116FFAEFDD3AC041BA59168" box="[589,655,821,844]" italics="true" pageId="76" pageNumber="181">Wings</emphasis>
: flightless (macropterous) (
<figureCitation id="1FE6D8C31116FFAEFED5AC6518819148" box="[331,427,852,876]" captionStart="Fig" captionStartId="14.[151,182,1885,1904]" captionTargetBox="[153,1410,191,1847]" captionTargetId="figure-13@14.[148,1416,163,1855]" captionTargetPageId="14" captionText="Fig. 10 – Wings. a, Synarmostes tibialis; b, Synarmostes occidentalis sp. nov.; c, Synarmostes sp. C; d, Philharmostes metallicus sp. nov.; e, Synarmostes makirovanae sp. nov.; f, Goudotostes laevis sp. nov.." figureDoi="http://doi.org/10.5281/zenodo.12764184" httpUri="https://zenodo.org/record/12764184/files/figure.png" pageId="76" pageNumber="181">Fig. 10 f</figureCitation>
).
<emphasis id="B5A918541116FFAEFE21AC651B779148" box="[447,605,852,876]" italics="true" pageId="76" pageNumber="181">Male genitalia</emphasis>
: aedeagus (L = 1.0 mm) with parameres as in
<figureCitation id="1FE6D8C31116FFAEFE78AC451B1D91A8" box="[486,567,884,908]" captionStart="Fig" captionStartId="129.[136,167,1858,1877]" captionTargetBox="[148,1397,170,1835]" captionTargetId="figure-4@129.[134,1414,156,1835]" captionTargetPageId="129" captionText="Fig. 64 – Goudotostes laevis sp. nov., a, female paratype, dorsal view of distended specimen; b, paratype, ventral view of rolled up specimen; c, paratype, dorsal view of rolled up specimen; d, paratype, ventral view of rolled up specimen; e, paratype, lateral view of rolled up specimen; f, holotype, spiculum gastrale; g–j, holotype, four views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764309" httpUri="https://zenodo.org/record/12764309/files/figure.png" pageId="76" pageNumber="181">Figs 64</figureCitation>
g-j, spiculum gastrale (L =
<quantity id="402569A31116FFAEFE9DACA5187D9188" box="[259,343,916,940]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="76" pageNumber="181" unit="mm" value="1.2">1.2 mm</quantity>
) as in
<figureCitation id="1FE6D8C31116FFAEFE00ACA518D49188" box="[414,510,916,940]" captionStart="Fig" captionStartId="129.[136,167,1858,1877]" captionTargetBox="[148,1397,170,1835]" captionTargetId="figure-4@129.[134,1414,156,1835]" captionTargetPageId="129" captionText="Fig. 64 – Goudotostes laevis sp. nov., a, female paratype, dorsal view of distended specimen; b, paratype, ventral view of rolled up specimen; c, paratype, dorsal view of rolled up specimen; d, paratype, ventral view of rolled up specimen; e, paratype, lateral view of rolled up specimen; f, holotype, spiculum gastrale; g–j, holotype, four views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764309" httpUri="https://zenodo.org/record/12764309/files/figure.png" pageId="76" pageNumber="181">
Fig.
<specimenCount id="91DB0FCF1116FFAEFE4EACA518D49188" box="[464,510,916,940]" count="64" pageId="76" pageNumber="181" type="generic">64 f</specimenCount>
</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CFC797CD1116FFAEFF23AC8518F09628" pageId="76" pageNumber="181" type="etymology">
<paragraph id="8762C4461116FFAEFF23AC8518F09628" blockId="76.[151,772,180,1164]" pageId="76" pageNumber="181">
<emphasis id="B5A918541116FFAEFF23AC85181491E8" bold="true" box="[189,318,948,972]" pageId="76" pageNumber="181">Etymology.</emphasis>
Latin adjective in the nominative singular meaning “polished”, due to the overall polished appearance of the dorsal sculpturing.
</paragraph>
</subSubSection>
<subSubSection id="CFC797CD1116FFAEFF23AB25184E96A8" pageId="76" pageNumber="181" type="distribution">
<paragraph id="8762C4461116FFAEFF23AB25184E96A8" blockId="76.[151,772,180,1164]" pageId="76" pageNumber="181">
<emphasis id="B5A918541116FFAEFF23AB2518F69608" bold="true" box="[189,476,1044,1068]" pageId="76" pageNumber="181">Distribution and habitat.</emphasis>
Known only from the
<typeStatus id="58667AE41116FFAEFD4BAB241A299609" box="[725,771,1045,1069]" pageId="76" pageNumber="181">type</typeStatus>
locality. Collected by sifting litter in rainforest. For details on the collecting locality see under
<taxonomicName id="40DDBFC51116FFAEFDD2AB64182B96A8" authorityName="Ballerio" authorityYear="2021" class="Insecta" family="Hybosoridae" genus="Synarmostes" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="181" phylum="Arthropoda" rank="species" species="makirovanae" status="sp. nov.">
<emphasis id="B5A918541116FFAEFDD2AB64182B96A8" italics="true" pageId="76" pageNumber="181">Synarmostes makirovanae</emphasis>
</taxonomicName>
<emphasis id="B5A918541116FFAEFE99AB45187796A8" bold="true" box="[263,349,1140,1164]" pageId="76" pageNumber="181">
<taxonomicNameLabel id="AE9AA52F1116FFAEFE99AB45187796A8" box="[263,349,1140,1164]" pageId="76" pageNumber="181" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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