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<document id="8BFC01F0AE8F0B53C8AB6A3B06F13648" ID-DOI="10.11646/zootaxa.3955.4.9" ID-GBIF-Dataset="fcf372f9-cb4b-4d0b-bf83-a37d663d07c4" ID-ISSN="1175-5326" ID-Zenodo-Dep="239809" ID-ZooBank="F7CD1F43-D256-4EA7-96FF-159131EE0258" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461053270960" checkinUser="plazi" docAuthor="Fraaije, René H. B., Van Bakel, Barry W. M., Jagt, John W. M. &amp; Machalski, Marcin" docDate="2015" docId="03AB87AEFFDFFFD4FF2176962B229305" docLanguage="en" docName="zt03955p594.pdf" docOrigin="Zootaxa 3955 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paguristes liwinskii Fraaije, Bakel, Jagt &amp; Machalski, 2015, n. sp." docType="treatment" docVersion="7" lastPageNumber="592" masterDocId="FF92FFD6FFDEFFD0FFB675552F239260" masterDocTitle="A new hermit crab (Anomura, Paguroidea) from the upper Albian (Cretaceous) of Annopol, Poland" masterLastPageNumber="594" masterPageNumber="588" pageNumber="589" updateTime="1698580287021" updateUser="plazi">
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<mods:title id="2AE51B370050115C7B130A653C14BC0C">A new hermit crab (Anomura, Paguroidea) from the upper Albian (Cretaceous) of Annopol, Poland</mods:title>
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<mods:namePart id="0922AFFCA414A40EF2B17391E58F70A3">Fraaije, René H. B.</mods:namePart>
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<mods:namePart id="E8AF7A88783ACAFCE8BD2D88D1DCB08A">Van Bakel, Barry W. M.</mods:namePart>
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<mods:namePart id="2280ED74D15A6D824258D4376A7AF358">Jagt, John W. M.</mods:namePart>
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<mods:namePart id="6896CC1F5009F0E06A6B354437D6BEC1">Machalski, Marcin</mods:namePart>
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<paragraph id="8BBD36B8FFDFFFD1FF2176962EEE91BE" blockId="1.[151,461,963,1023]" box="[151,461,963,990]" pageId="1" pageNumber="589">
<heading id="D0F581D4FFDFFFD1FF2176962EEE91BE" bold="true" box="[151,461,963,990]" fontSize="11" level="1" pageId="1" pageNumber="589" reason="1">
<emphasis id="B976EAAAFFDFFFD1FF2176962EEE91BE" bold="true" box="[151,461,963,990]" pageId="1" pageNumber="589">
<taxonomicName id="4C024D3BFFDFFFD1FF2176962EA091BD" box="[151,387,963,989]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="589" phylum="Arthropoda" rank="species" species="liwinskii" status="sp. nov.">
<emphasis id="B976EAAAFFDFFFD1FF2176962EA091BD" bold="true" box="[151,387,963,989]" italics="true" pageId="1" pageNumber="589">Paguristes liwinskii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24557D1FFDFFFD1FE3D76912EEE91BE" box="[395,461,964,990]" pageId="1" pageNumber="589" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="8BBD36B8FFDFFFD1FF2176B32FDE919F" blockId="1.[151,461,963,1023]" box="[151,253,998,1023]" pageId="1" pageNumber="589">
(
<figureCitation id="13392A3DFFDFFFD1FF2976B32FFD919F" box="[159,222,998,1023]" captionStart="FIGURE 1. A" captionStartId="2.[151,250,1512,1534]" captionTargetBox="[158,1419,185,1479]" captionTargetId="figure@2.[150,1436,173,1491]" captionTargetPageId="2" captionText="FIGURE 1. A, Litho- and chronostratigraphic column of the Annopol sedimentary succession (modified from Machalski &amp; Kennedy 2013), level of Paguristes liwinskii sp. nov. is arrowed. B, Geological map of the Annopol area (inset: Poland). C, Paguristes liwinskii sp. nov., ZPAL Cr. 9 / 43 (holotype), from the lower part of unit 3, upper upper Albian, locality Kopiec; in outer (1), inner (2), ventral (3) and dorsal (4) views, respectively." httpUri="https://zenodo.org/record/239810/files/figure.png" pageId="1" pageNumber="589">Fig. 1</figureCitation>
C)
</paragraph>
</subSubSection>
<subSubSection id="C3186533FFDFFFD1FF21717B2E0396D3" pageId="1" pageNumber="589" type="diagnosis">
<paragraph id="8BBD36B8FFDFFFD1FF21717B2E0396D3" blockId="1.[151,1436,1070,2031]" pageId="1" pageNumber="589">
<emphasis id="B976EAAAFFDFFFD1FF21717B2E329627" bold="true" box="[151,273,1070,1095]" pageId="1" pageNumber="589">Diagnosis.</emphasis>
Cross section of right cheliped transversely oval; dorsal and ventral sides convex, with regular, dense cover of forwardly directed, fine tubercles, largest arranged in row along dorsal outer side and along central proximal inner side; fixed finger short, stout and triangular; dactylus short and triangular, about twice length of fixed finger.
</paragraph>
</subSubSection>
<subSubSection id="C3186533FFDFFFD1FF7171EB2D82977F" pageId="1" pageNumber="589" type="etymology">
<paragraph id="8BBD36B8FFDFFFD1FF7171EB2D82977F" blockId="1.[151,1436,1070,2031]" pageId="1" pageNumber="589">
<emphasis id="B976EAAAFFDFFFD1FF7171EB2E6D96B7" bold="true" box="[199,334,1214,1239]" pageId="1" pageNumber="589">Etymology.</emphasis>
Named in honour of the mayor of Annopol, Mr Wiesław Liwiński, who facilitated large-scale excavations conducted in this area in recent years by a team led by the last author (for details, see
<bibRefCitation id="EF934B49FFDFFFD1FABF71B72E18977F" author="Machalski" pageId="1" pageNumber="589" refString="Machalski, M. &amp; Kennedy, W. J. (2013) Oyster-bioimmured ammonites from the Upper Albian of Annopol, Poland: stratigraphic and palaeobiogeographic implications. Acta Geologica Polonica, 63, 545 - 554. http: // dx. doi. org / 10.2478 / agp- 2013 - 0024" type="journal article" year="2013">Machalski &amp; Kennedy 2013</bibRefCitation>
and
<bibRefCitation id="EF934B49FFDFFFD1FEC770522DB6977F" author="Popov" box="[369,661,1286,1311]" pageId="1" pageNumber="589" refString="Popov, E. V. &amp; Machalski, M. (2014) Late Albian chimeroid fishes (Chimaeroidei) from Annopol, Poland. Cretaceous Research, 47, 1 - 18. http: // dx. doi. org / 10.1016 / j. cretres. 2013.09.011" type="journal article" year="2014">Popov &amp; Machalski 2014</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3186533FFDFFFD1FF71707F2B8D94CA" pageId="1" pageNumber="589" type="materials_examined">
<paragraph id="8BBD36B8FFDFFFD1FF71707F2C8E9707" blockId="1.[151,1436,1070,2031]" pageId="1" pageNumber="589">
<emphasis id="B976EAAAFFDFFFD1FF71707F2E2B9723" bold="true" box="[199,264,1322,1347]" pageId="1" pageNumber="589">
<typeStatus id="54B9881AFFDFFFD1FF71707F2E279723" box="[199,260,1322,1347]" pageId="1" pageNumber="589">Type</typeStatus>
.
</emphasis>
The
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, and sole specimen known (
<collectionCode id="ED13AE7DFFDFFFD1FD68707E2C079723" box="[734,804,1323,1347]" httpUri="http://grbio.org/cool/yd4q-n37a" name="Zoological Institute of Paleobiology, Polish Academy of Sciences" pageId="1" pageNumber="589">ZPAL</collectionCode>
Cr.9/43), is a near-complete right palm with fixed finger and dactylus, measuring
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in length, and
<quantity id="4CFA9B5DFFDFFFD1FD15701A2DC79706" box="[675,740,1359,1382]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="589" unit="mm" value="7.0">7 mm</quantity>
in greatest width.
</paragraph>
<paragraph id="8BBD36B8FFDFFFD1FF7170272B8D94CA" blockId="1.[151,1436,1070,2031]" pageId="1" pageNumber="589">
<emphasis id="B976EAAAFFDFFFD1FF7170272D2397EB" bold="true" box="[199,512,1394,1419]" pageId="1" pageNumber="589">Locality and stratigraphy.</emphasis>
Kopiec near Annopol, lower part of unit 3 (
<figureCitation id="13392A3DFFDFFFD1FC4270272B6D97EB" box="[1012,1102,1394,1419]" captionStart="FIGURE 1. A" captionStartId="2.[151,250,1512,1534]" captionTargetBox="[158,1419,185,1479]" captionTargetId="figure@2.[150,1436,173,1491]" captionTargetPageId="2" captionText="FIGURE 1. A, Litho- and chronostratigraphic column of the Annopol sedimentary succession (modified from Machalski &amp; Kennedy 2013), level of Paguristes liwinskii sp. nov. is arrowed. B, Geological map of the Annopol area (inset: Poland). C, Paguristes liwinskii sp. nov., ZPAL Cr. 9 / 43 (holotype), from the lower part of unit 3, upper upper Albian, locality Kopiec; in outer (1), inner (2), ventral (3) and dorsal (4) views, respectively." httpUri="https://zenodo.org/record/239810/files/figure.png" pageId="1" pageNumber="589">Fig. 1A</figureCitation>
), upper upper Albian. Unit 3,
<emphasis id="B976EAAAFFDFFFD1FF2170C22F8097CE" box="[151,163,1431,1454]" italics="true" pageId="1" pageNumber="589">c</emphasis>
.
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thick, is composed of quartz sand with abundant glauconite, passing gradually upwards into quartzglauconitic marl. A distinct bed of phosphatic clasts and fossils occurs in the upper part of unit 3. Its upper boundary is sharp, whereas the lower boundary is diffuse due to bioturbation (
<bibRefCitation id="EF934B49FFDFFFD1FBBA708B2AB39797" author="Marcinowski" box="[1036,1424,1502,1527]" pageId="1" pageNumber="589" refString="Marcinowski, R. &amp; Walaszczyk, I. (1985) Middle Cretaceous deposits and biostratigraphy of the Annopol section, Central Polish Uplands. Osterreichische Akademie der Wissenschaften, Schriftenreiche der Erdwissenschaftlichen Komisssionen, 7, 27 - 41." type="journal article" year="1985">Marcinowski &amp; Walaszczyk 1985</bibRefCitation>
). The phosphatic clasts are
<emphasis id="B976EAAAFFDFFFD1FE7373512EF2947B" box="[453,465,1540,1563]" italics="true" pageId="1" pageNumber="589">c</emphasis>
.
<quantity id="4CFA9B5DFFDFFFD1FE5573562D15947A" box="[483,566,1539,1563]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="589" unit="mm" value="10.0">10 mm</quantity>
in diameter, but in the upper part of the layer they commonly form larger aggregates, up to
<quantity id="4CFA9B5DFFDFFFD1FEDE73722EEA945E" box="[360,457,1575,1599]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="589" unit="mm" value="120.0">120 mm</quantity>
in diameter. Unit 3 of the Annopol succession is dated on the basis of specifically indeterminate specimens of the ammonite
<taxonomicName id="4C024D3BFFDFFFD1FDC273192CD79403" authority="Subschloenbachia" authorityName="Subschloenbachia" box="[628,1012,1610,1635]" class="Cephalopoda" family="Brancoceratidae" genus="Mortoniceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonitina" pageId="1" pageNumber="589" phylum="Mollusca" rank="genus">
<emphasis id="B976EAAAFFDFFFD1FDC273192C2F9403" box="[628,780,1612,1635]" italics="true" pageId="1" pageNumber="589">Mortoniceras</emphasis>
(
<emphasis id="B976EAAAFFDFFFD1FCAB731F2CCA9403" box="[797,1001,1610,1635]" italics="true" pageId="1" pageNumber="589">Subschloenbachia</emphasis>
)
</taxonomicName>
, preserved as attachment scars on the left valves of some oysters (
<bibRefCitation id="EF934B49FFDFFFD1FE41733B2C4794E7" author="Machalski" box="[503,868,1646,1671]" pageId="1" pageNumber="589" refString="Machalski, M. &amp; Kennedy, W. J. (2013) Oyster-bioimmured ammonites from the Upper Albian of Annopol, Poland: stratigraphic and palaeobiogeographic implications. Acta Geologica Polonica, 63, 545 - 554. http: // dx. doi. org / 10.2478 / agp- 2013 - 0024" type="journal article" year="2013">Machalski and Kennedy 2013</bibRefCitation>
). These specimens are closest to
<taxonomicName id="4C024D3BFFDFFFD1FAB2733A2E5694CB" authority="Subschloenbachia" authorityName="Subschloenbachia" class="Cephalopoda" family="Brancoceratidae" genus="Mortoniceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonitina" pageId="1" pageNumber="589" phylum="Mollusca" rank="genus">
<emphasis id="B976EAAAFFDFFFD1FAB2733A2ABF94E6" box="[1284,1436,1647,1670]" italics="true" pageId="1" pageNumber="589">Mortoniceras</emphasis>
(
<emphasis id="B976EAAAFFDFFFD1FF1673C72E4F94CB" box="[160,364,1682,1707]" italics="true" pageId="1" pageNumber="589">Subschloenbachia</emphasis>
)
</taxonomicName>
that characterise the upper upper Albian
<taxonomicName id="4C024D3BFFDFFFD1FCF273C12B8D94CA" authority="Zone." authorityName="Zone." box="[836,1198,1682,1707]" class="Cephalopoda" family="Brancoceratidae" genus="Mortoniceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonitina" pageId="1" pageNumber="589" phylum="Mollusca" rank="species" species="perinflatum">
<emphasis id="B976EAAAFFDFFFD1FCF273C12B4594CB" box="[836,1126,1682,1707]" italics="true" pageId="1" pageNumber="589">Mortoniceras perinflatum</emphasis>
Zone.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3186533FFDFFFD1FF7173E32FF5958F" pageId="1" pageNumber="589" type="description">
<paragraph id="8BBD36B8FFDFFFD1FF7173E32FF5958F" blockId="1.[151,1436,1070,2031]" pageId="1" pageNumber="589">
<emphasis id="B976EAAAFFDFFFD1FF7173E32E7A94AF" bold="true" box="[199,345,1718,1743]" pageId="1" pageNumber="589">Description.</emphasis>
Globose right cheliped, cross section transversely oval; convex dorsal and ventral sides; all sides with regular, dense cover of forwardly directed, fine tubercles, largest in a row along dorsal outer and along central proximal inner sides. Fixed finger short, stout, curved, with a row of about four large setal pores on the inner and outer cutting edge and numerous irregular, longitudinal rows of finer pores on central and ventral parts; tip spatulate; cutting edge near-straight with concave central part. Dactylus short, triangular, curved, about twice the length of fixed finger; covered with irregular rows of setal pores smallest in dorsal half; tip pointed; strongly concave dorsal side here considered to be a preservational artefact, possibly resulting from unsuccessful predatory attack as some regeneration cuticle is visible on the proximal dorsal part of dactylus and the distal part of dorsal palm.
</paragraph>
</subSubSection>
<caption id="DF7D6630FFDCFFD2FF2170BD2C08943A" httpUri="https://zenodo.org/record/239810/files/figure.png" pageId="2" pageNumber="590" targetBox="[158,1419,185,1479]" targetPageId="2">
<paragraph id="8BBD36B8FFDCFFD2FF2170BD2C08943A" blockId="2.[151,1436,1512,1626]" pageId="2" pageNumber="590">
<emphasis id="B976EAAAFFDCFFD2FF2170BD2E36979D" bold="true" box="[151,277,1512,1534]" pageId="2" pageNumber="590">FIGURE 1.</emphasis>
A, Litho- and chronostratigraphic column of the Annopol sedimentary succession (modified from Machalski &amp; Kennedy 2013), level of
<taxonomicName id="4C024D3BFFDCFFD2FE2C735D2D47947D" box="[410,612,1543,1565]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="590" phylum="Arthropoda" rank="species" species="liwinskii" status="sp. nov.">
<emphasis id="B976EAAAFFDCFFD2FE2C735D2D47947D" box="[410,612,1543,1565]" italics="true" pageId="2" pageNumber="590">Paguristes liwinskii</emphasis>
</taxonomicName>
<emphasis id="B976EAAAFFDCFFD2FDDA73522D9D947C" bold="true" box="[620,702,1543,1564]" pageId="2" pageNumber="590">
<taxonomicNameLabel id="A24557D1FFDCFFD2FDDA73522D9D947C" box="[620,702,1543,1564]" pageId="2" pageNumber="590" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is arrowed. B, Geological map of the Annopol area (inset: Poland). C,
<taxonomicName id="4C024D3BFFDCFFD2FF2173722E43945B" box="[151,352,1573,1596]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="590" phylum="Arthropoda" rank="species" species="liwinskii" status="sp. nov.">
<emphasis id="B976EAAAFFDCFFD2FF2173722E43945B" box="[151,352,1573,1596]" italics="true" pageId="2" pageNumber="590">Paguristes liwinskii</emphasis>
</taxonomicName>
<emphasis id="B976EAAAFFDCFFD2FED173732E9A945B" bold="true" box="[359,441,1574,1595]" pageId="2" pageNumber="590">
<taxonomicNameLabel id="A24557D1FFDCFFD2FED173732E9A945B" box="[359,441,1574,1595]" pageId="2" pageNumber="590" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ZPAL Cr.9/43 (holotype), from the lower part of unit 3, upper upper Albian, locality Kopiec; in outer (1), inner (2), ventral (3) and dorsal (4) views, respectively.
</paragraph>
</caption>
<subSubSection id="C3186533FFDCFFD3FF7173D22B9490A9" lastPageId="3" lastPageNumber="591" pageId="2" pageNumber="590" type="discussion">
<paragraph id="8BBD36B8FFDCFFD2FF7173D22E8A9584" blockId="2.[151,1437,1671,2020]" pageId="2" pageNumber="590">
<emphasis id="B976EAAAFFDCFFD2FF7173D22E6F94C0" bold="true" box="[199,332,1671,1696]" pageId="2" pageNumber="590">Discussion.</emphasis>
The generic placement of fossil paguroid chelae is difficult in that claws and, occasionally, meri and carpi are all that is preserved. The specimen studied has been assigned to
<taxonomicName id="4C024D3BFFDCFFD2FB9C73F92B8194A3" box="[1066,1186,1708,1731]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="590" phylum="Arthropoda" rank="genus">
<emphasis id="B976EAAAFFDCFFD2FB9C73F92B8194A3" box="[1066,1186,1708,1731]" italics="true" pageId="2" pageNumber="590">Paguristes</emphasis>
</taxonomicName>
because the claws of
<taxonomicName id="4C024D3BFFDCFFD2FF2173852EEE9487" box="[151,461,1744,1767]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="590" phylum="Arthropoda" rank="species" species="santamartaensis">
<emphasis id="B976EAAAFFDCFFD2FF2173852EEE9487" box="[151,461,1744,1767]" italics="true" pageId="2" pageNumber="590">Paguristes santamartaensis</emphasis>
</taxonomicName>
are remarkably similar and members of
<taxonomicName id="4C024D3BFFDCFFD2FC3873852B259487" box="[910,1030,1744,1767]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="590" phylum="Arthropoda" rank="genus">
<emphasis id="B976EAAAFFDCFFD2FC3873852B259487" box="[910,1030,1744,1767]" italics="true" pageId="2" pageNumber="590">Paguristes</emphasis>
</taxonomicName>
tend to be isochelous, whereas most other diogenid genera comprise species that are predominantly heterochelous with the left chela considerably larger. We have to keep in mind that the lefthandedness of most diogenids probably is the result of gastropod shell inhabitation, the great majority of marine snails being dextral. Fraaije (2003) noted a shift in molluscan inhabitation from ammonites to gastropods during the Cretaceous Period; this affected the morphology of hermit claws that blocked the aperture. It is also highly probable that Cretaceous species of
<taxonomicName id="4C024D3BFFDCFFD2FBC272D12BCF95FB" box="[1140,1260,1924,1947]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="590" phylum="Arthropoda" rank="genus">
<emphasis id="B976EAAAFFDCFFD2FBC272D12BCF95FB" box="[1140,1260,1924,1947]" italics="true" pageId="2" pageNumber="590">Paguristes</emphasis>
</taxonomicName>
had isochelous chelae and that left heterochely mainly occurred subsequent to the Cretaceous/Paleogene boundary in relation to this shift in inhabitation.
</paragraph>
<paragraph id="8BBD36B8FFDDFFD3FF7175CD2EE993B0" blockId="3.[151,1436,151,464]" pageId="3" pageNumber="591">
To date, about 120 extant species of
<taxonomicName id="4C024D3BFFDDFFD3FDD975CC2DC492D0" box="[623,743,153,176]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="genus">
<emphasis id="B976EAAAFFDDFFD3FDD975CC2DC492D0" box="[623,743,153,176]" italics="true" pageId="3" pageNumber="591">Paguristes</emphasis>
</taxonomicName>
are on record (
<bibRefCitation id="EF934B49FFDDFFD3FC2F75C22B8D92D0" author="McLaughlin" box="[921,1198,151,176]" pageId="3" pageNumber="591" refString="McLaughlin, P. A., Komai, T., Lemaitre, R. &amp; Rahayu, D. L. (2010) Annotated checklist of anomuran decapod crustaceans of the world (exclusive of the Kiwaoidea and families Chirostylidae and Galatheidae of the Galatheoidea). Part I - Lithodoidea, Lomisoidea and Paguroidea. The Raffles Bulletin of Zoology, 23 (Supplement), 5 - 107." type="journal article" year="2010">
McLaughlin
<emphasis id="B976EAAAFFDDFFD3FB9875CC2B4B92D0" box="[1070,1128,151,176]" italics="true" pageId="3" pageNumber="591">et al.</emphasis>
2010
</bibRefCitation>
;
<bibRefCitation id="EF934B49FFDDFFD3FB0D75C22AB392D0" author="Komai" box="[1211,1424,151,176]" pageId="3" pageNumber="591" refString="Komai, T., Reshmi, R. &amp; Kumar, A. B. (2015) A new species of the hermit crab genus Paguristes Dana, 1851 (Crustacea: Decapoda: Anomura: Diogenidae) from southwestern India. Zootaxa, 3937 (3), 517 - 532. http: // dx. doi. org / 10.11646 / zootaxa. 3937.3.5" type="journal article" year="2015">
Komai
<emphasis id="B976EAAAFFDDFFD3FAA675CC2A6892D0" box="[1296,1355,151,176]" italics="true" pageId="3" pageNumber="591">et al.</emphasis>
2015
</bibRefCitation>
), illustrating that this genus is the most successful member of the
<taxonomicName id="4C024D3BFFDDFFD3FC1275E92B0692B5" box="[932,1061,188,213]" class="Malacostraca" family="Diogenidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="family">Diogenidae</taxonomicName>
by far. From the fossil record,
<bibRefCitation id="EF934B49FFDDFFD3FF21758A2E879298" box="[151,420,223,248]" pageId="3" pageNumber="591" refString="Schweitzer, C. E., Feldmann, R. M., Garassino, A., Karasawa, H. &amp; Schweigert, G. (2010) Systematic list of fossil decapod crustacean species. Crustaceana Monographs, 10, i - viii + 1 - 221. http: // dx. doi. org / 10.1163 / ej. 9789004178915. i- 222" type="journal article">
Schweitzer
<emphasis id="B976EAAAFFDDFFD3FEAC75B42E719298" box="[282,338,223,248]" italics="true" pageId="3" pageNumber="591">et al.</emphasis>
(2010)
</bibRefCitation>
recorded a total of 15 species. Here, we list 17 species (
<tableCitation id="C6800303FFDDFFD3FBA1758A2B7E9298" box="[1047,1117,223,248]" captionStart="TABLE 1" captionStartId="3.[151,239,512,535]" captionText="TABLE 1. Enumeration of extinct members of the genus Paguristes known to date, arranged in stratigraphical order. Paguristes substriatiformis Lõrenthey &amp; Beurlen, 1929, from the Eocene of Hungary, has subsequently been transferred to the genus Ciliopagurus Forest, 1995 (see Forest 1995; Van Bakel et al. 2003; Schweitzer et al. 2010). NOTE: In view of their poor preservation, P. chipolensis Rathbun, 1935, and P. subequalis Rathbun, 1926, both from the Miocene of the USA, are here considered to be nomina dubia." pageId="3" pageNumber="591">Tab. 1</tableCitation>
), inclusive of the new form, which extends the stratigraphic range of
<taxonomicName id="4C024D3BFFDDFFD3FDD074502DFD937C" box="[614,734,261,284]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="genus">
<emphasis id="B976EAAAFFDDFFD3FDD074502DFD937C" box="[614,734,261,284]" italics="true" pageId="3" pageNumber="591">Paguristes</emphasis>
</taxonomicName>
downwards into the late Albian by about 25 myr. Four of the listed species (
<taxonomicName id="4C024D3BFFDDFFD3FE88747C2D309320" box="[318,531,297,320]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="santamartaensis">
<emphasis id="B976EAAAFFDDFFD3FE88747C2D309320" box="[318,531,297,320]" italics="true" pageId="3" pageNumber="591">P. santamartaensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C024D3BFFDDFFD3FD94747C2DE89320" box="[546,715,295,320]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="ouachitensis">
<emphasis id="B976EAAAFFDDFFD3FD94747C2DE89320" box="[546,715,295,320]" italics="true" pageId="3" pageNumber="591">P. ouachitensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C024D3BFFDDFFD3FD6D747C2C729320" box="[731,849,295,320]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="whitteni">
<emphasis id="B976EAAAFFDDFFD3FD6D747C2C729320" box="[731,849,295,320]" italics="true" pageId="3" pageNumber="591">P. whitteni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C024D3BFFDDFFD3FC3D747C2CC89320" box="[907,1003,295,320]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="florae">
<emphasis id="B976EAAAFFDDFFD3FC3D747C2CC89320" box="[907,1003,295,320]" italics="true" pageId="3" pageNumber="591">P. florae</emphasis>
</taxonomicName>
) are of late Late Cretaceous, eight (
<taxonomicName id="4C024D3BFFDDFFD3FA31747C2E2C9305" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="baldoensis">
<emphasis id="B976EAAAFFDDFFD3FA31747C2E2C9305" italics="true" pageId="3" pageNumber="591">P. baldoensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C024D3BFFDDFFD3FEAA74182EB79304" box="[284,404,333,356]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="extentus">
<emphasis id="B976EAAAFFDDFFD3FEAA74182EB79304" box="[284,404,333,356]" italics="true" pageId="3" pageNumber="591">P. extentus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C024D3BFFDDFFD3FE1774182D109305" box="[417,563,332,357]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="prealpinus">
<emphasis id="B976EAAAFFDDFFD3FE1774182D109305" box="[417,563,332,357]" italics="true" pageId="3" pageNumber="591">P. prealpinus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C024D3BFFDDFFD3FDF774182D9E9305" box="[577,701,332,357]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="wheeleri">
<emphasis id="B976EAAAFFDDFFD3FDF774182D9E9305" box="[577,701,332,357]" italics="true" pageId="3" pageNumber="591">P. wheeleri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C024D3BFFDDFFD3FD7D74182C7F9304" box="[715,860,333,356]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="B976EAAAFFDDFFD3FD7D74182C7F9304" box="[715,860,333,356]" italics="true" pageId="3" pageNumber="591">P. mexicanus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C024D3BFFDDFFD3FCDD74182CC59305" box="[875,998,332,357]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="johnsoni">
<emphasis id="B976EAAAFFDDFFD3FCDD74182CC59305" box="[875,998,332,357]" italics="true" pageId="3" pageNumber="591">P. johnsoni</emphasis>
</taxonomicName>
,
<taxonomicName id="4C024D3BFFDDFFD3FC4274182B5D9305" box="[1012,1150,332,357]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="hokoensis">
<emphasis id="B976EAAAFFDDFFD3FC4274182B5D9305" box="[1012,1150,332,357]" italics="true" pageId="3" pageNumber="591">P. hokoensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C024D3BFFDDFFD3FB0274182AB09305" box="[1204,1427,332,357]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="lineatuberculatus">
<emphasis id="B976EAAAFFDDFFD3FB0274182AB09305" box="[1204,1427,332,357]" italics="true" pageId="3" pageNumber="591">P. lineatuberculatus</emphasis>
</taxonomicName>
) of Paleogene, two (
<taxonomicName id="4C024D3BFFDDFFD3FEC474242D6B93E8" box="[370,584,367,392]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="oligotuberculata">
<emphasis id="B976EAAAFFDDFFD3FEC474242D6B93E8" box="[370,584,367,392]" italics="true" pageId="3" pageNumber="591">P. oligotuberculata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C024D3BFFDDFFD3FD3674242C0593E8" box="[640,806,367,392]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="cserhatensis">
<emphasis id="B976EAAAFFDDFFD3FD3674242C0593E8" box="[640,806,367,392]" italics="true" pageId="3" pageNumber="591">P. cserhatensis</emphasis>
</taxonomicName>
) of Neogene and two (
<taxonomicName id="4C024D3BFFDDFFD3FB9A74242B9B93E8" box="[1068,1208,367,392]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="lymani">
<emphasis id="B976EAAAFFDDFFD3FB9A74242B1D93E8" box="[1068,1086,369,392]" italics="true" pageId="3" pageNumber="591">P.</emphasis>
cf.
<emphasis id="B976EAAAFFDDFFD3FBDA743A2B9B93E8" box="[1132,1208,367,392]" italics="true" pageId="3" pageNumber="591">lymani</emphasis>
</taxonomicName>
and
<taxonomicName id="4C024D3BFFDDFFD3FB4674242AB093E8" box="[1264,1427,367,392]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="syrtensis">
<emphasis id="B976EAAAFFDDFFD3FB4674242A2193E8" box="[1264,1282,369,392]" italics="true" pageId="3" pageNumber="591">P.</emphasis>
cf.
<emphasis id="B976EAAAFFDDFFD3FA8774242AB093E8" box="[1329,1427,369,392]" italics="true" pageId="3" pageNumber="591">syrtensis</emphasis>
</taxonomicName>
) of Quaternary age. Only one of these records, i.e., that of
<taxonomicName id="4C024D3BFFDDFFD3FC9D74C02CF193CC" box="[811,978,404,429]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="syrtensis">
<emphasis id="B976EAAAFFDDFFD3FC9D74C02C1D93CC" box="[811,830,405,428]" italics="true" pageId="3" pageNumber="591">P.</emphasis>
cf.
<emphasis id="B976EAAAFFDDFFD3FCDB74C02CF193CC" box="[877,978,405,428]" italics="true" pageId="3" pageNumber="591">syrtensis</emphasis>
</taxonomicName>
, is based on a fragmentary dorsal shield (see
<bibRefCitation id="EF934B49FFDDFFD3FF7D74E22E9D93B0" author="Garassino" box="[203,446,439,464]" pageId="3" pageNumber="591" refString="Garassino, A., Pasini, G., De Angeli, A. &amp; Hyzny, M. (2014) The decapod fauna (Axiidea, Anomura, Brachyura) from the Late Pleistocene of Trumbaca, Reggio Calabria (Calabria, southern Italy). Natural History Sciences, Atti della Societa italiana di Scienze naturali del Museo civico di Storia naturale di Milano, 1 (2), 119 - 130. http: // dx. doi. org / 10.4081 / nhs. 2014.60" type="journal article" year="2014">
Garassino
<emphasis id="B976EAAAFFDDFFD3FEF474EC2E5193B0" box="[322,370,439,464]" italics="true" pageId="3" pageNumber="591">et al</emphasis>
. 2014
</bibRefCitation>
).
</paragraph>
<caption id="DF7D6630FFDDFFD3FF2177552DA690F7" pageId="3" pageNumber="591">
<paragraph id="8BBD36B8FFDDFFD3FF2177552DA690F7" blockId="3.[151,1436,512,663]" pageId="3" pageNumber="591">
<emphasis id="B976EAAAFFDDFFD3FF2177552E2F9077" bold="true" box="[151,268,512,535]" pageId="3" pageNumber="591">TABLE 1.</emphasis>
Enumeration of extinct members of the genus
<taxonomicName id="4C024D3BFFDDFFD3FCA577552CA69077" box="[787,901,512,535]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="genus">
<emphasis id="B976EAAAFFDDFFD3FCA577552CA69077" box="[787,901,512,535]" italics="true" pageId="3" pageNumber="591">Paguristes</emphasis>
</taxonomicName>
known to date, arranged in stratigraphical order.
<taxonomicName id="4C024D3BFFDDFFD3FF2177752DC99057" authority="Lorenthey &amp; Beurlen, 1929" authorityName="Lorenthey &amp; Beurlen" authorityYear="1929" box="[151,746,543,567]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="substriatiformis">
<emphasis id="B976EAAAFFDDFFD3FF2177752E9A9057" box="[151,441,543,567]" italics="true" pageId="3" pageNumber="591">Paguristes substriatiformis</emphasis>
Lõrenthey &amp; Beurlen, 1929
</taxonomicName>
, from the Eocene of Hungary, has subsequently been transferred to the genus
<taxonomicName id="4C024D3BFFDDFFD3FEAB776A2D169037" authority="Forest, 1995" authorityName="Forest" authorityYear="1995" box="[285,565,575,599]" class="Malacostraca" family="Diogenidae" genus="Ciliopagurus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="genus">
<emphasis id="B976EAAAFFDDFFD3FEAB776A2E899037" box="[285,426,575,599]" italics="true" pageId="3" pageNumber="591">Ciliopagurus</emphasis>
Forest, 1995
</taxonomicName>
(see Forest 1995; Van Bakel
<emphasis id="B976EAAAFFDDFFD3FCD877152CBF9037" box="[878,924,575,599]" italics="true" pageId="3" pageNumber="591">et al</emphasis>
. 2003; Schweitzer
<emphasis id="B976EAAAFFDDFFD3FBDC77152BB49037" box="[1130,1175,575,599]" italics="true" pageId="3" pageNumber="591">et al</emphasis>
. 2010).
<emphasis id="B976EAAAFFDDFFD3FB4777152A669037" bold="true" box="[1265,1349,576,599]" pageId="3" pageNumber="591">NOTE:</emphasis>
In view of their poor preservation,
<taxonomicName id="4C024D3BFFDDFFD3FE0477352DCD9017" authority="Rathbun, 1935" authorityName="Rathbun" authorityYear="1935" box="[434,750,607,631]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="chipolensis">
<emphasis id="B976EAAAFFDDFFD3FE0477352D649017" box="[434,583,607,631]" italics="true" pageId="3" pageNumber="591">P. chipolensis</emphasis>
Rathbun, 1935
</taxonomicName>
, and
<taxonomicName id="4C024D3BFFDDFFD3FC9377352B799017" authority="Rathbun, 1926" authorityName="Rathbun" authorityYear="1926" box="[805,1114,607,631]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="subequalis">
<emphasis id="B976EAAAFFDDFFD3FC9377352C979017" box="[805,948,607,631]" italics="true" pageId="3" pageNumber="591">P. subequalis</emphasis>
Rathbun, 1926
</taxonomicName>
, both from the Miocene of the USA, are here considered to be nomina dubia.
</paragraph>
</caption>
<paragraph id="8BBD36B8FFDDFFD3FF2177E62B9490A9" blockId="3.[151,1207,691,714]" box="[151,1207,691,714]" pageId="3" pageNumber="591">
<emphasis id="B976EAAAFFDDFFD3FF2177E62B9490A9" bold="true" box="[151,1207,691,714]" pageId="3" pageNumber="591">Species Stratigraphic/geographic occurrence</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3186533FFDDFFD4FF2177BD2B229305" lastPageId="4" lastPageNumber="592" pageId="3" pageNumber="591" type="reference_group">
<paragraph id="8BBD36B8FFDDFFD3FF2177BD2B259138" blockId="3.[151,1302,742,1286]" pageId="3" pageNumber="591">
<taxonomicName id="4C024D3BFFDDFFD3FF2177BD2E25909C" box="[151,262,742,765]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="liwinskii" status="sp. nov.">
<emphasis id="B976EAAAFFDDFFD3FF2177BD2E25909C" box="[151,262,742,765]" italics="true" pageId="3" pageNumber="591">P. liwinskii</emphasis>
</taxonomicName>
<emphasis id="B976EAAAFFDDFFD3FEBA77B22E7F909C" bold="true" box="[268,348,743,764]" pageId="3" pageNumber="591">
<taxonomicNameLabel id="A24557D1FFDDFFD3FEBA77B22E7F909C" box="[268,348,743,764]" pageId="3" pageNumber="591" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
upper Albian,
<collectingCountry id="F3157628FFDDFFD3FC7B77B32B30909C" box="[973,1043,742,764]" name="Poland" pageId="3" pageNumber="591">Poland</collectingCountry>
<taxonomicName id="4C024D3BFFDDFFD3FF2176532DF4917B" authority="Feldmann, Tshudy &amp; Thomson, 1993" authorityName="Feldmann, Tshudy &amp; Thomson" authorityYear="1993" box="[151,727,773,795]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="santamartaensis">
<emphasis id="B976EAAAFFDDFFD3FF2176532E75917B" box="[151,342,774,795]" italics="true" pageId="3" pageNumber="591">P. santamartaensis</emphasis>
<bibRefCitation id="EF934B49FFDDFFD3FEEA76502DF4917B" author="Feldmann" box="[348,727,773,795]" pageId="3" pageNumber="591" refString="Feldmann, R. M., Tshudy, D. M. &amp; Thomson, M R. A. (1993) Late Cretaceous and Paleocene decapod crustaceans from James Ross Basin, Antarctic Peninsula. The Paleontological Society Memoir, 28, 1 - 41." type="journal article" year="1993">Feldmann, Tshudy &amp; Thomson, 1993</bibRefCitation>
</taxonomicName>
upper Santonian/lower Campanian,
<collectingCountry id="F3157628FFDDFFD3FB1176502A32917B" box="[1191,1297,773,795]" name="Antarctica" pageId="3" pageNumber="591">Antarctica</collectingCountry>
<taxonomicName id="4C024D3BFFDDFFD3FF2176702EEE915A" authority="Rathbun, 1935" authorityName="Rathbun" authorityYear="1935" box="[151,461,804,826]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="ouachitensis">
<emphasis id="B976EAAAFFDDFFD3FF2176702E11915A" box="[151,306,804,826]" italics="true" pageId="3" pageNumber="591">P. ouachitensis</emphasis>
<bibRefCitation id="EF934B49FFDDFFD3FE8E76712EEE915A" author="Rathbun" box="[312,461,804,826]" pageId="3" pageNumber="591" refString="Rathbun, M. J. (1935) Fossil Crustacea of the Atlantic and Gulf Coastal Plain. Geological Society of America, Special Paper, 2, 1 - 160. http: // dx. doi. org / 10.1130 / SPE 2 - p 1" type="journal article" year="1935">Rathbun, 1935</bibRefCitation>
</taxonomicName>
upper Campanian,
<collectingCountry id="F3157628FFDDFFD3FC4D76712B0D915A" box="[1019,1070,804,826]" name="United States of America" pageId="3" pageNumber="591">USA</collectingCountry>
<taxonomicName id="4C024D3BFFDDFFD3FF2176162CEA9138" authority="Bishop, 1983 Maastrichtian" authorityName="Bishop, 1983 Maastrichtian" box="[151,969,834,856]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="whitteni">
<emphasis id="B976EAAAFFDDFFD3FF2176162E269138" box="[151,261,834,856]" italics="true" pageId="3" pageNumber="591">P. whitteni</emphasis>
<bibRefCitation id="EF934B49FFDDFFD3FEBD76172EB19138" author="Bishop" box="[267,402,834,856]" pageId="3" pageNumber="591" refString="Bishop, G. (1983) Fossil decapod Crustacea from the Late Cretaceous Coon Creek Formation, Union County, Mississippi. Journal of Crustacean Biology, 3, 417 - 430. http: // dx. doi. org / 10.2307 / 1548142" type="journal article" year="1983">Bishop, 1983</bibRefCitation>
Maastrichtian
</taxonomicName>
,
<collectingCountry id="F3157628FFDDFFD3FC6576172B259138" box="[979,1030,834,856]" name="United States of America" pageId="3" pageNumber="591">USA</collectingCountry>
</paragraph>
<paragraph id="8BBD36B8FFDDFFD3FF2176372B2B91F6" blockId="3.[151,1302,742,1286]" pageId="3" pageNumber="591">
<taxonomicName id="4C024D3BFFDDFFD3FF2176372D369117" authority="Collins, Fraaye &amp; Jagt, 1995" authorityName="Collins, Fraaye &amp; Jagt" authorityYear="1995" box="[151,533,865,887]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="florae">
<emphasis id="B976EAAAFFDDFFD3FF2176372FCF9117" box="[151,236,865,887]" italics="true" pageId="3" pageNumber="591">P. florae</emphasis>
<bibRefCitation id="EF934B49FFDDFFD3FF4476342D369117" author="Collins" box="[242,533,865,887]" pageId="3" pageNumber="591" refString="Collins, J. S. H., Fraaye, R. H. B. &amp; Jagt, J. W. M. (1995) Late Cretaceous anomurans and brachyurans from the Maastrichtian type area. Acta Palaeontologica Polonica, 40, 165 - 210." type="journal article" year="1995">Collins, Fraaye &amp; Jagt, 1995</bibRefCitation>
</taxonomicName>
upper Maastrichtian,
<collectingCountry id="F3157628FFDDFFD3FBA576342B909117" box="[1043,1203,865,887]" name="Netherlands" pageId="3" pageNumber="591">the Netherlands</collectingCountry>
<collectingCountry id="F3157628FFDDFFD3FB7676342A359117" box="[1216,1302,865,887]" name="Belgium" pageId="3" pageNumber="591">Belgium</collectingCountry>
<taxonomicName id="4C024D3BFFDDFFD3FF2176D42DB891F6" authority="Garassino, De Angeli &amp; Pasini, 2009" authorityName="Garassino, De Angeli &amp; Pasini" authorityYear="2009" box="[151,667,896,918]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="baldoensis">
<emphasis id="B976EAAAFFDDFFD3FF2176D42E3F91F6" box="[151,284,896,918]" italics="true" pageId="3" pageNumber="591">P. baldoensis</emphasis>
<bibRefCitation id="EF934B49FFDDFFD3FE9476D52DB891F6" author="Garassino" box="[290,667,896,918]" pageId="3" pageNumber="591" refString="Garassino, A., De Angeli, A. &amp; Pasini, G. (2009) In situ hermit crab (Crustacea, Anomura, Paguroidea from the Early Eocene (Ypresian) of NE Italy. Atti della Societa italiana di Scienze naturali del Museo civico di Storia naturale di Milano, 150 (II), 229 - 238." type="journal article" year="2009">
Garassino,
<collectingCountry id="F3157628FFDDFFD3FE2276D42E9291F6" box="[404,433,897,918]" name="South Georgia and the South Sandwich Islands" pageId="3" pageNumber="591">De</collectingCountry>
Angeli &amp; Pasini, 2009
</bibRefCitation>
</taxonomicName>
Lower Eocene,
<collectingCountry id="F3157628FFDDFFD3FC6D76D52B2B91F6" box="[987,1032,896,918]" name="Italy" pageId="3" pageNumber="591">Italy</collectingCountry>
</paragraph>
<paragraph id="8BBD36B8FFDDFFD3FF2176F52B2B91D4" blockId="3.[151,1302,742,1286]" box="[151,1032,926,949]" pageId="3" pageNumber="591">
<taxonomicName id="4C024D3BFFDDFFD3FF2176F52DFA91D4" authority="Beschin, Busulini, De Angeli &amp; Tessier, 2007" authorityName="Beschin, Busulini, De Angeli &amp; Tessier" authorityYear="2007" box="[151,729,926,949]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="extentus">
<emphasis id="B976EAAAFFDDFFD3FF2176F52E2091D5" box="[151,259,928,949]" italics="true" pageId="3" pageNumber="591">P. extentus</emphasis>
<bibRefCitation id="EF934B49FFDDFFD3FEBF76CB2DFA91D4" author="Beschin" box="[265,729,926,948]" pageId="3" pageNumber="591" refString="Beschin, C., Busulini, A., De Angeli, A. &amp; Tessier G. (2007) I decapodi dell'Eocene inferiore di Contrada Gecchelina (Vicenza - Italia settentrionale) (Anomura e Brachiura). Museo di Archeologia e Scienze Naturali ' G. Zannato ', Montecchio Maggiore (Vicenza), 2007, 9 - 76." type="journal article" year="2007">
Beschin, Busulini,
<collectingCountry id="F3157628FFDDFFD3FE7C76CA2EC491D4" box="[458,487,927,948]" name="South Georgia and the South Sandwich Islands" pageId="3" pageNumber="591">De</collectingCountry>
Angeli &amp; Tessier, 2007
</bibRefCitation>
</taxonomicName>
Lower Eocene,
<collectingCountry id="F3157628FFDDFFD3FC6D76CB2B2B91D4" box="[987,1032,926,948]" name="Italy" pageId="3" pageNumber="591">Italy</collectingCountry>
</paragraph>
<paragraph id="8BBD36B8FFDDFFD3FF2176EB2B2A91B3" blockId="3.[151,1302,742,1286]" box="[151,1033,957,979]" pageId="3" pageNumber="591">
<taxonomicName id="4C024D3BFFDDFFD3FF2176EB2CF191B3" authority="Beschin, De Angeli, Checchi &amp; Zarantonello, 2005 Middle Eocene" authorityName="Beschin, De Angeli, Checchi &amp; Zarantonello, 2005 Middle Eocene" box="[151,978,957,979]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="prealpinus">
<emphasis id="B976EAAAFFDDFFD3FF2176EB2E3F91B3" box="[151,284,957,979]" italics="true" pageId="3" pageNumber="591">P. prealpinus</emphasis>
<bibRefCitation id="EF934B49FFDDFFD3FE9776E82C0A91B3" author="Beschin" box="[289,809,957,979]" pageId="3" pageNumber="591" refString="Beschin, C., De Angeli, A., Checchi, A. &amp; Zarantonello, G. (2005) Crostacei eocenici di Grola di Cornedo Vicentino presso Spagnago (Vicenza, Italia settentrionale). Studi e Ricerche - Associazione Amici Museo civico ' G. Zannato', Montecchio Maggiore (Vicenza), 12, 5 - 35." type="journal article" year="2005">
Beschin,
<collectingCountry id="F3157628FFDDFFD3FEC976EB2EBE91B3" box="[383,413,958,979]" name="South Georgia and the South Sandwich Islands" pageId="3" pageNumber="591">De</collectingCountry>
Angeli, Checchi &amp; Zarantonello, 2005
</bibRefCitation>
Middle Eocene
</taxonomicName>
,
<collectingCountry id="F3157628FFDDFFD3FC6A76E82B2A91B3" box="[988,1033,957,979]" name="Italy" pageId="3" pageNumber="591">Italy</collectingCountry>
</paragraph>
<paragraph id="8BBD36B8FFDDFFD3FF2176882DA39670" blockId="3.[151,1302,742,1286]" pageId="3" pageNumber="591">
<taxonomicName id="4C024D3BFFDDFFD3FF2176882CFB9192" authority="Blow &amp; Manning, 1996 Middle Eocene" authorityName="Blow &amp; Manning, 1996 Middle Eocene" box="[151,984,988,1010]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="wheeleri">
<emphasis id="B976EAAAFFDDFFD3FF2176882E299192" box="[151,266,988,1010]" italics="true" pageId="3" pageNumber="591">P. wheeleri</emphasis>
<bibRefCitation id="EF934B49FFDDFFD3FEA676892D209192" author="Blow" box="[272,515,988,1010]" pageId="3" pageNumber="591" refString="Blow, W. C. &amp; Manning, R. B. (1996) Preliminary descriptions of 25 new decapod crustaceans from the Middle Eocene of the Carolinas, U. S. A. Tulane Studies in Geology and Paleontology, 29, 1 - 26." type="journal article" year="1996">Blow &amp; Manning, 1996</bibRefCitation>
Middle Eocene
</taxonomicName>
,
<collectingCountry id="F3157628FFDDFFD3FC5476892B369192" box="[994,1045,988,1010]" name="United States of America" pageId="3" pageNumber="591">USA</collectingCountry>
<taxonomicName id="4C024D3BFFDDFFD3FF2176AE2D5F9670" authority="Vega, Cosma, Coutino, Feldmann" authorityName="Vega, Cosma, Coutino, Feldmann" box="[151,636,1018,1040]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="B976EAAAFFDDFFD3FF2176AE2E3D9670" box="[151,286,1019,1040]" italics="true" pageId="3" pageNumber="591">P. mexicanus</emphasis>
Vega, Cosma, Coutiño, Feldmann
</taxonomicName>
,
</paragraph>
<paragraph id="8BBD36B8FFDDFFD3FF21714C2CE5962E" blockId="3.[151,1302,742,1286]" pageId="3" pageNumber="591">
<bibRefCitation id="EF934B49FFDDFFD3FF21714C2D2B964F" author="Nyborg" box="[151,520,1049,1071]" pageId="3" pageNumber="591" refString="Vega, F. J., Cosma, T., Coutino, M. A., Feldmann, R. M., Nyborg, T. G., Schweitzer, C. E. &amp; Waugh, D. A. (2001) New Middle Eocene decapods (Crustacea) from Chiapas, Mexico. Journal of Paleontology, 75, 929 - 946. http: // dx. doi. org / 10.1666 / 0022 - 3360 (2001) 075 % 3 C 0929: NMEDCF % 3 E 2.0. CO; 2" type="journal article" year="2001">Nyborg, Schweitzer &amp; Waugh, 2001</bibRefCitation>
Middle Eocene,
<collectingCountry id="F3157628FFDDFFD3FC54714C2B0C964F" box="[994,1071,1049,1071]" name="Mexico" pageId="3" pageNumber="591">Mexico</collectingCountry>
<taxonomicName id="4C024D3BFFDDFFD3FF21716C2CAB962E" authority="Rathbun, 1935 Eocene" authorityName="Rathbun, 1935 Eocene" box="[151,904,1080,1102]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="johnsoni">
<emphasis id="B976EAAAFFDDFFD3FF21716C2E29962E" box="[151,266,1080,1102]" italics="true" pageId="3" pageNumber="591">P. johnsoni</emphasis>
<bibRefCitation id="EF934B49FFDDFFD3FEA6716D2E85962E" author="Rathbun" box="[272,422,1080,1102]" pageId="3" pageNumber="591" refString="Rathbun, M. J. (1935) Fossil Crustacea of the Atlantic and Gulf Coastal Plain. Geological Society of America, Special Paper, 2, 1 - 160. http: // dx. doi. org / 10.1130 / SPE 2 - p 1" type="journal article" year="1935">Rathbun, 1935</bibRefCitation>
Eocene
</taxonomicName>
,
<collectingCountry id="F3157628FFDDFFD3FC25716D2CE5962E" box="[915,966,1080,1102]" name="United States of America" pageId="3" pageNumber="591">USA</collectingCountry>
</paragraph>
<paragraph id="8BBD36B8FFDDFFD3FF21710D2B28960C" blockId="3.[151,1302,742,1286]" box="[151,1035,1110,1133]" pageId="3" pageNumber="591">
<taxonomicName id="4C024D3BFFDDFFD3FF21710D2D70960C" authority="Schweitzer &amp; Feldmann, 2001" authorityName="Schweitzer &amp; Feldmann" authorityYear="2001" box="[151,595,1110,1133]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="hokoensis">
<emphasis id="B976EAAAFFDDFFD3FF21710D2E34960C" box="[151,279,1110,1133]" italics="true" pageId="3" pageNumber="591">P. hokoensis</emphasis>
Schweitzer &amp; Feldmann, 2001
</taxonomicName>
Upper Eocene,
<collectingCountry id="F3157628FFDDFFD3FC6E71032B28960C" box="[984,1035,1110,1132]" name="United States of America" pageId="3" pageNumber="591">USA</collectingCountry>
</paragraph>
<paragraph id="8BBD36B8FFDDFFD3FF2171232CDA96A8" blockId="3.[151,1302,742,1286]" pageId="3" pageNumber="591">
<taxonomicName id="4C024D3BFFDDFFD3FF2171232D7A96EB" authority="Muller &amp; Collins, 1991" authorityName="Muller &amp; Collins" authorityYear="1991" box="[151,601,1141,1163]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="oligotuberculatus">
<emphasis id="B976EAAAFFDDFFD3FF2171232E4696EB" box="[151,357,1141,1163]" italics="true" pageId="3" pageNumber="591">P. oligotuberculatus</emphasis>
<bibRefCitation id="EF934B49FFDDFFD3FEDD71202D7A96EB" author="Muller" box="[363,601,1141,1163]" pageId="3" pageNumber="591" refString="Muller, P. &amp; Collins, J. S. H. (1991) Late Eocene coral-associated decapods (Crustacea) from Hungary. Contributions to Tertiary and Quaternary Geology, 28, 47 - 92." type="journal article" year="1991">Müller &amp; Collins, 1991</bibRefCitation>
</taxonomicName>
Upper Eocene,
<collectingCountry id="F3157628FFDDFFD3FC6E71232B1396EB" box="[984,1072,1142,1163]" name="Hungary" pageId="3" pageNumber="591">Hungary</collectingCountry>
<taxonomicName id="4C024D3BFFDDFFD3FF2171C02C1396CA" authority="Beschin, De Angeli, Checchi &amp; Mietto, 2006" authorityName="Beschin, De Angeli, Checchi &amp; Mietto" authorityYear="2006" box="[151,816,1172,1194]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="lineatuberculatus">
<emphasis id="B976EAAAFFDDFFD3FF2171C02E4396CA" box="[151,352,1172,1194]" italics="true" pageId="3" pageNumber="591">P. lineatuberculatus</emphasis>
<bibRefCitation id="EF934B49FFDDFFD3FED071C12C1396CA" author="Beschin" box="[358,816,1172,1194]" pageId="3" pageNumber="591" refString="Beschin, C., De Angeli, A., Checchi, A. &amp; Mietto, P. (2006) Crostacei del Priaboniano di Priabona (Vicenza - Italia settentrionale). Lavori - Societa Veneziana di Scienze Naturali, Venezia, 31, 95 - 112." type="journal article" year="2006">
Beschin,
<collectingCountry id="F3157628FFDDFFD3FE7271C02EC296CA" box="[452,481,1173,1194]" name="South Georgia and the South Sandwich Islands" pageId="3" pageNumber="591">De</collectingCountry>
Angeli, Checchi &amp; Mietto, 2006
</bibRefCitation>
</taxonomicName>
Upper EoceneOligocene,
<collectingCountry id="F3157628FFDDFFD3FBF871C12B5896CA" box="[1102,1147,1172,1194]" name="Italy" pageId="3" pageNumber="591">Italy</collectingCountry>
<taxonomicName id="4C024D3BFFDDFFD3FF2171E62CB596A8" authority="Muller, 1984 Miocene" authorityName="Muller, 1984 Miocene" box="[151,918,1202,1224]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="cserhatensis">
<emphasis id="B976EAAAFFDDFFD3FF2171E62E1396A8" box="[151,304,1202,1224]" italics="true" pageId="3" pageNumber="591">P. cserhatensis</emphasis>
<bibRefCitation id="EF934B49FFDDFFD3FE8071E72E9A96A8" author="Muller" box="[310,441,1202,1224]" pageId="3" pageNumber="591" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 1 - 317." type="journal article" year="1984">Müller, 1984</bibRefCitation>
Miocene
</taxonomicName>
,
<collectingCountry id="F3157628FFDDFFD3FC1771E62CDA96A8" box="[929,1017,1203,1224]" name="Hungary" pageId="3" pageNumber="591">Hungary</collectingCountry>
</paragraph>
<paragraph id="8BBD36B8FFDDFFD3FF2171872B2E9687" blockId="3.[151,1302,742,1286]" box="[151,1037,1233,1255]" pageId="3" pageNumber="591">
<taxonomicName id="4C024D3BFFDDFFD3FF2171872C929687" ID-CoL="f0b7dda1-0141-47bb-b42d-2a3ffe32c867" authority="A. Milne-Edwards &amp; Bouvier, 1893 Pleistocene" authorityName="A. Milne-Edwards &amp; Bouvier, 1893 Pleistocene" box="[151,945,1233,1255]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="lymani">
<emphasis id="B976EAAAFFDDFFD3FF2171872F8B9687" box="[151,168,1234,1255]" italics="true" pageId="3" pageNumber="591">P.</emphasis>
cf.
<emphasis id="B976EAAAFFDDFFD3FF6671842E369687" box="[208,277,1233,1255]" italics="true" pageId="3" pageNumber="591">lymani</emphasis>
<bibRefCitation id="EF934B49FFDDFFD3FEAD71872DAB9687" author="Milne-Edwards" box="[283,648,1233,1255]" pageId="3" pageNumber="591" refString="Milne-Edwards, A. &amp; Bouvier, E. L. (1893) Description des Crustaces de la famille des Paguriens recueillis pendant l'expedition. Reports of the results of dredging under the supervision of Alexander Agassiz in the Gulf of Mexico (1877 - 78) and along the Atlantic coast of the United States (1880) by the U. S. Survey steamer &quot; Blake, &quot; Lieut. - Com. S. D. Sigsbee and Commander J. R. Bartlett commanding. Memoirs of the Museum of Comparative Zoology, 14, 1 - 172. [Harvard]" type="journal article" year="1893">A. Milne-Edwards &amp; Bouvier, 1893</bibRefCitation>
Pleistocene
</taxonomicName>
,
<collectingCountry id="F3157628FFDDFFD3FC0D71842B2E9687" box="[955,1037,1233,1255]" name="Jamaica" pageId="3" pageNumber="591">Jamaica</collectingCountry>
</paragraph>
<paragraph id="8BBD36B8FFDDFFD3FF2171A42CCA9766" blockId="3.[151,1302,742,1286]" box="[151,1001,1264,1286]" pageId="3" pageNumber="591">
<taxonomicName id="4C024D3BFFDDFFD3FF2171A42E0A9766" ID-CoL="590da9b5-f9cd-4b59-8f56-a6de8b858f00" box="[151,297,1264,1286]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="syrtensis">
<emphasis id="B976EAAAFFDDFFD3FF2171A42F8B9766" box="[151,168,1265,1286]" italics="true" pageId="3" pageNumber="591">P.</emphasis>
cf.
<emphasis id="B976EAAAFFDDFFD3FF6671A42E0A9766" box="[208,297,1265,1286]" italics="true" pageId="3" pageNumber="591">syrtensis</emphasis>
</taxonomicName>
de
<bibRefCitation id="EF934B49FFDDFFD3FEFB71A52D359766" author="Saint" box="[333,534,1264,1286]" pageId="3" pageNumber="591" refString="Saint Laurent, M. de (1971) Paguristes syrtensis, espece nouvelle de cotes tunisiennes (Crustacea Decapoda Diogenidae). Bulletin du Museum national d'Histoire naturelle de Paris, Series 2, 42 (for 1970), 1009 - 1107." type="journal article" year="1971">Saint Laurent, 1971</bibRefCitation>
Pleistocene,
<collectingCountry id="F3157628FFDDFFD3FC0D71A52CCA9766" box="[955,1001,1264,1286]" name="Italy" pageId="3" pageNumber="591">Italy</collectingCountry>
</paragraph>
<paragraph id="8BBD36B8FFDDFFD3FF7170062EDC947F" blockId="3.[151,1437,1362,2035]" pageId="3" pageNumber="591">
In overall shape and dense uniform ornamentation,
<taxonomicName id="4C024D3BFFDDFFD3FC9670012B22970B" box="[800,1025,1362,1387]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="liwinskii" status="sp. nov.">
<emphasis id="B976EAAAFFDDFFD3FC9670012B22970B" box="[800,1025,1362,1387]" italics="true" pageId="3" pageNumber="591">Paguristes liwinskii</emphasis>
</taxonomicName>
<emphasis id="B976EAAAFFDDFFD3FBBD70072B4B970B" bold="true" box="[1035,1128,1362,1387]" pageId="3" pageNumber="591">
<taxonomicNameLabel id="A24557D1FFDDFFD3FBBD70072B4B970B" box="[1035,1128,1362,1387]" pageId="3" pageNumber="591" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
most closely resembles
<taxonomicName id="4C024D3BFFDDFFD3FA3170012E6C97EE" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="santamartaensis">
<emphasis id="B976EAAAFFDDFFD3FA3170012E6C97EE" italics="true" pageId="3" pageNumber="591">P. santamartaensis</emphasis>
</taxonomicName>
, although the latter differs in having a smooth inner surface, a flattened dorsal surface and coarser tuberculation. The new species can be differentiated from
<taxonomicName id="4C024D3BFFDDFFD3FC9570C92CFA97D3" box="[803,985,1434,1459]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="ouachitaensis">
<emphasis id="B976EAAAFFDDFFD3FC9570C92CFA97D3" box="[803,985,1434,1459]" italics="true" pageId="3" pageNumber="591">P. ouachitaensis</emphasis>
</taxonomicName>
in that the palm has a convex inner side and transverse rows of tubercles are lacking.
<taxonomicName id="4C024D3BFFDDFFD3FD0470EA2CB397B7" box="[690,912,1470,1495]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="whitteni">
<emphasis id="B976EAAAFFDDFFD3FD0470EA2CB397B7" box="[690,912,1470,1495]" italics="true" pageId="3" pageNumber="591">Paguristes whitteni</emphasis>
</taxonomicName>
has a much coarser ornament and a longer, strongly deflected fixed finger, while
<emphasis id="B976EAAAFFDDFFD3FDE970B12DE3979B" box="[607,704,1506,1531]" italics="true" pageId="3" pageNumber="591">P. f l or a e</emphasis>
is less globose, has fingers with a lesser curvature and the ornamentation is more subdued.
</paragraph>
<paragraph id="8BBD36B8FFDDFFD3FF7173792FCD9497" blockId="3.[151,1437,1362,2035]" pageId="3" pageNumber="591">
<taxonomicName id="4C024D3BFFDDFFD3FF7173792E9E9423" box="[199,445,1578,1603]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="baldoensis">
<emphasis id="B976EAAAFFDDFFD3FF7173792E9E9423" box="[199,445,1578,1603]" italics="true" pageId="3" pageNumber="591">Paguristes baldoensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C024D3BFFDDFFD3FE4473792D4F9423" box="[498,620,1580,1603]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="extensus">
<emphasis id="B976EAAAFFDDFFD3FE4473792D4F9423" box="[498,620,1580,1603]" italics="true" pageId="3" pageNumber="591">P. extensus</emphasis>
</taxonomicName>
are less convex, have straighter dorsal and ventral sides and much longer fingers, whereas
<taxonomicName id="4C024D3BFFDDFFD3FEE8731A2ED59407" box="[350,502,1614,1639]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="prealpinus">
<emphasis id="B976EAAAFFDDFFD3FEE8731A2ED59407" box="[350,502,1614,1639]" italics="true" pageId="3" pageNumber="591">P. prealpinus</emphasis>
</taxonomicName>
has a smoother inner surface and a much longer and downturned fixed finger.
<taxonomicName id="4C024D3BFFDDFFD3FF2173212EFC94EB" box="[151,479,1650,1675]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="lineatuberculatus">
<emphasis id="B976EAAAFFDDFFD3FF2173212EFC94EB" box="[151,479,1650,1675]" italics="true" pageId="3" pageNumber="591">Paguristes lineatuberculatus</emphasis>
</taxonomicName>
has a different ornamentation and the distal tip of the fixed finger is directed downwards, whereas
<taxonomicName id="4C024D3BFFDDFFD3FE3D73C22D2894CF" box="[395,523,1686,1711]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="wheeleri">
<emphasis id="B976EAAAFFDDFFD3FE3D73C22D2894CF" box="[395,523,1686,1711]" italics="true" pageId="3" pageNumber="591">P. wheeleri</emphasis>
</taxonomicName>
shows a strongly curved occlusal surface and much coarser tubercles and spines on both sides. The ornamentation in
<taxonomicName id="4C024D3BFFDDFFD3FD8973E92DF594B3" box="[575,726,1724,1747]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="B976EAAAFFDDFFD3FD8973E92DF594B3" box="[575,726,1724,1747]" italics="true" pageId="3" pageNumber="591">P. mexicanus</emphasis>
</taxonomicName>
is less uniform, with a reticulate outer surface near the upper margin.
</paragraph>
<paragraph id="8BBD36B8FFDDFFD3FF7172512CDF9502" blockId="3.[151,1437,1362,2035]" pageId="3" pageNumber="591">
<taxonomicName id="4C024D3BFFDDFFD3FF7172512E84957B" box="[199,423,1794,1819]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="johnsoni">
<emphasis id="B976EAAAFFDDFFD3FF7172512E84957B" box="[199,423,1794,1819]" italics="true" pageId="3" pageNumber="591">Paguristes johnsoni</emphasis>
</taxonomicName>
can be differentiated in having a less globose palm, a longer fixed finger, and less dense and coarser ornamentation, while
<taxonomicName id="4C024D3BFFDDFFD3FDAC72722D85955F" box="[538,678,1830,1855]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="hokoensis">
<emphasis id="B976EAAAFFDDFFD3FDAC72722D85955F" box="[538,678,1830,1855]" italics="true" pageId="3" pageNumber="591">P. hokoensis</emphasis>
</taxonomicName>
has a significantly longer, downturned fixed finger and fewer, yet larger tubercles, arranged in longitudinal rows on the outer side of the manus.
</paragraph>
<paragraph id="8BBD36B8FFDDFFD3FF71723A2ABE9593" blockId="3.[151,1437,1362,2035]" pageId="3" pageNumber="591">
<taxonomicName id="4C024D3BFFDDFFD3FF71723A2EEC95E7" box="[199,463,1902,1927]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="cserhatensis">
<emphasis id="B976EAAAFFDDFFD3FF71723A2EEC95E7" box="[199,463,1902,1927]" italics="true" pageId="3" pageNumber="591">Paguristes cserhatensis</emphasis>
</taxonomicName>
is less globose, and has fewer, yet much larger tubercles arranged in longitudinal rows; in addition, the fixed finger has a lesser curvature. The outer surface of the palm in
<taxonomicName id="4C024D3BFFDDFFD3FBE272C12A1595CB" box="[1108,1334,1938,1963]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="oligotuberculatus">
<emphasis id="B976EAAAFFDDFFD3FBE272C12A1595CB" box="[1108,1334,1938,1963]" italics="true" pageId="3" pageNumber="591">P. oligotuberculatus</emphasis>
</taxonomicName>
has only few rows of coarse tubercles and the dorsal margin is straight and spinose, while
<taxonomicName id="4C024D3BFFDDFFD3FB9E72E22B9795AF" ID-CoL="f0b7dda1-0141-47bb-b42d-2a3ffe32c867" box="[1064,1204,1974,1999]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="lymani">
<emphasis id="B976EAAAFFDDFFD3FB9E72E22B1895AE" box="[1064,1083,1975,1998]" italics="true" pageId="3" pageNumber="591">P.</emphasis>
cf.
<emphasis id="B976EAAAFFDDFFD3FBDE72E32B9795AF" box="[1128,1204,1974,1999]" italics="true" pageId="3" pageNumber="591">lymani</emphasis>
</taxonomicName>
differs in showing a smoother inner surface, straighter dorsal and ventral margins. Moreover, it does not have a uniform ornamentation.
</paragraph>
<paragraph id="8BBD36B8FFDAFFD4FF7175C22B229305" blockId="4.[151,1436,151,357]" pageId="4" pageNumber="592">
<bibRefCitation id="EF934B49FFDAFFD4FF7175C22E9B92D0" author="Rathbun" box="[199,440,151,176]" pageId="4" pageNumber="592" refString="Rathbun, M. J. (1926) The fossil stalk-eyed Crustacea of the Pacific Slope of North America. United States National Museum Bulletin, 138, i - vii + 1 - 155. http: // dx. doi. org / 10.5479 / si. 03629236.138. i" type="journal article" year="1926">Rathbun (1926: 101)</bibRefCitation>
described the preservation of the
<typeStatus id="54B9881AFFDAFFD4FCFC75CD2C5992D0" box="[842,890,152,176]" pageId="4" pageNumber="592">type</typeStatus>
and only known specimen of
<taxonomicName id="4C024D3BFFDAFFD4FB5575CC2A5992D0" box="[1251,1402,151,176]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="592" phylum="Arthropoda" rank="species" species="subequalis">
<emphasis id="B976EAAAFFDAFFD4FB5575CC2A5992D0" box="[1251,1402,151,176]" italics="true" pageId="4" pageNumber="592">P. subequalis</emphasis>
</taxonomicName>
as follows, [.....] so crumbly that no more can be uncovered without destroying it. Only the form of the fingers is seen, and in the case of the propodal finger, nothing but the impression.
<taxonomicName id="4C024D3BFFDAFFD4FC5E75B42BC49298" box="[1000,1255,223,248]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="592" phylum="Arthropoda" rank="species" species="chipolensis">
<emphasis id="B976EAAAFFDAFFD4FC5E75B42BC49298" box="[1000,1255,223,248]" italics="true" pageId="4" pageNumber="592">Paguristes chipolensis</emphasis>
</taxonomicName>
was erected by
<bibRefCitation id="EF934B49FFDAFFD4FF2174512E64937D" author="Rathbun" box="[151,327,260,285]" pageId="4" pageNumber="592" refString="Rathbun, M. J. (1935) Fossil Crustacea of the Atlantic and Gulf Coastal Plain. Geological Society of America, Special Paper, 2, 1 - 160. http: // dx. doi. org / 10.1130 / SPE 2 - p 1" type="journal article" year="1935">Rathbun (1935)</bibRefCitation>
on the basis of a single, incomplete dactylus and propodus. The preservation of both
<taxonomicName id="4C024D3BFFDAFFD4FABE74502ABF937D" box="[1288,1436,260,285]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="592" phylum="Arthropoda" rank="species" species="subequalis">
<emphasis id="B976EAAAFFDAFFD4FABE74502ABF937D" box="[1288,1436,260,285]" italics="true" pageId="4" pageNumber="592">P. subequalis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C024D3BFFDAFFD4FF7E747C2E409320" box="[200,355,295,320]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="592" phylum="Arthropoda" rank="species" species="chipolensis">
<emphasis id="B976EAAAFFDAFFD4FF7E747C2E409320" box="[200,355,295,320]" italics="true" pageId="4" pageNumber="592">P. chipolensis</emphasis>
</taxonomicName>
is so poor that a detailed comparison is impossible; we propose that they are best regarded as
<emphasis id="B976EAAAFFDAFFD4FF2174182E119305" box="[151,306,332,357]" italics="true" pageId="4" pageNumber="592">nomina dubia</emphasis>
and recommend removal from decapod crustacean species lists.
</paragraph>
</subSubSection>
</treatment>
</document>