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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.981.53583" ID-GBIF-Dataset="8e14a5b5-aa20-4050-b2b4-f393764fc6c7" ID-PMC="PMC7644702" ID-Pensoft-Pub="1313-2970-981-1" ID-Pensoft-UUID="7C44C797C4125A71BAE032A55E6FA5DC" ID-PubMed="33199958" ID-ZooBank="5194AAC86B8A473F8A41470A60182A0B" ModsDocID="1313-2970-981-1" checkinTime="1604011718171" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Sajan, Sheikh, Tripathy, Basudev, Meng, Kaibaryer, Asami, Takahiro &amp; Ablett, Jonathan D." docDate="2020" docId="DFC7B62D7CFE5A408CA7486682E2034B" docLanguage="en" docName="ZooKeys 981: 1-220" docOrigin="ZooKeys 981" docSource="http://dx.doi.org/10.3897/zookeys.981.53583" docTitle="Dicharax elevatus" docType="treatment" docVersion="6" id="7C44C797C4125A71BAE032A55E6FA5DC" lastPageNumber="1" masterDocId="7C44C797C4125A71BAE032A55E6FA5DC" masterDocTitle="Genus-level revision of the Alycaeidae (Gastropoda, Cyclophoroidea), with an annotated species catalogue" masterLastPageNumber="220" masterPageNumber="1" pageNumber="1" updateTime="1668169683351" updateUser="ExternalLinkService">
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<mods:title>Genus-level revision of the Alycaeidae (Gastropoda, Cyclophoroidea), with an annotated species catalogue</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pall-Gergely, Barna</mods:namePart>
<mods:affiliation>Plant Protection Institute, Centre for Agricultural Research, Herman Otto ut 15, Budapest, H- 1022, Hungary</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sajan, Sheikh</mods:namePart>
<mods:affiliation>Zoological Survey of India, Prani Vigyan Bhawan, M Block, New Alipore, Kolkata 700053, West Bengal, India &amp; Wildlife Institute of India, Chandrabani, Dehradun 248 002, Uttarakhand, India</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Tripathy, Basudev</mods:namePart>
<mods:affiliation>Zoological Survey of India, Prani Vigyan Bhawan, M Block, New Alipore, Kolkata 700053, West Bengal, India</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Meng, Kaibaryer</mods:namePart>
<mods:affiliation>National Zoological Museum of China, Institute of Zoology, Chinese Academy of Sciences, Beijing, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Asami, Takahiro</mods:namePart>
<mods:affiliation>Department of Biology, Shinshu University, Matsumoto 390 - 8621, Japan</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-5706-0272</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ablett, Jonathan D.</mods:namePart>
<mods:affiliation>Mollusca Section, Invertebrates Division, Department of Life Sciences, The Natural History Museums, London SW 7 5 BD, United Kingdom</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>981</mods:number>
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<mods:end>220</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.981.53583</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.981.53583</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-981-1</mods:identifier>
<mods:identifier type="ZooBank">5194AAC86B8A473F8A41470A60182A0B</mods:identifier>
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<treatment ID-GBIF-Taxon="168715019" LSID="urn:lsid:plazi:treatment:DFC7B62D7CFE5A408CA7486682E2034B" httpUri="http://treatment.plazi.org/id/DFC7B62D7CFE5A408CA7486682E2034B" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName LSID="DFC7B62D-7CFE-5A40-8CA7-486682E2034B" authority="(Heude, 1886)" baseAuthorityName="Heude" baseAuthorityYear="1886" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dicharax elevatus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="elevatus">Dicharax elevatus (Heude, 1886)</taxonomicName>
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Dicharax (?) elevatus (Heude, 1886), syntype (HMT- 218 a). Photographs: Kaibaryer Meng." figureDoi="10.3897/zookeys.981.53583.figure16" httpUri="https://binary.pensoft.net/fig/468661" pageId="0" pageNumber="1">Fig. 16</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Heude" authorityYear="1886" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus elevatus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="elevatus">Alycaeus elevatus</taxonomicName>
Heude, 1886: 210.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Heude" authorityYear="1886" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus elevatus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="elevatus">Alycaeus elevatus</taxonomicName>
-
<bibRefCitation author="Heude, RPM" journalOrPublisher="Mission Catholique, Chang-Hai" pageId="0" pageNumber="1" pagination="125 - 188" refId="B84" refString="Heude, RPM, 1890. Memoires concernant l'histoire naturelle de l'empire chinois par des peres de la Compagnie de JESUS. Notes sur les Mollusques terrestres de la vallee du Fleuve Bleu. Mission Catholique, Chang-Hai 4: 125 - 188" title="Memoires concernant l'histoire naturelle de l'empire chinois par des peres de la Compagnie de JESUS. Notes sur les Mollusques terrestres de la vallee du Fleuve Bleu." volume="4" year="1890">Heude 1890</bibRefCitation>
: 129, pl. 36, fig. 19.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Heude" authorityYear="1886" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Chamalycaeus) elevatus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="elevatus" subGenus="Chamalycaeus">Alycaeus (Chamalycaeus) elevatus</taxonomicName>
-
<bibRefCitation author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/28228303" refId="B98" refString="Kobelt, W, Moellendorff, O von, 1897. Catalog der gegenvaertig lebend bekannten Pneumonopomen. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 29: 73-88, 105-120, 137-152. https://www.biodiversitylibrary.org/page/28228303" title="Catalog der gegenvaertig lebend bekannten Pneumonopomen. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 29: 73 - 88, 105 - 120, 137 - 152." url="https://www.biodiversitylibrary.org/page/28228303" year="1897">
Kobelt and
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
1897
</bibRefCitation>
: 148;
<bibRefCitation author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/1000211" refId="B97" refString="Kobelt, W, 1902. Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. In Verbindung mit der Deutschen Zoologischen Gesellschaft herausgegeben von der Koeniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R. Friedlaender und Sohn, Berlin, https://biodiversitylibrary.org/page/1000211" title="Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. In Verbindung mit der Deutschen Zoologischen Gesellschaft herausgegeben von der Koeniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R." url="https://biodiversitylibrary.org/page/1000211" year="1902">Kobelt 1902</bibRefCitation>
: 355.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Heude" baseAuthorityYear="1886" class="Gastropoda" family="Cyclophoridae" genus="Metalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metalycaeus elevatus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="elevatus">Metalycaeus (?) elevatus</taxonomicName>
-
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4331.1.1" author="Pall-Gergely, B" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 124" refId="B162" refString="Pall-Gergely, B, Hunyadi, A, Đỗ, ĐS, Naggs, F, Asami, T, 2017. Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus. Zootaxa 4331 (1): 1 - 124, DOI: https://doi.org/10.11646/zootaxa.4331.1.1" title="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus." url="https://doi.org/10.11646/zootaxa.4331.1.1" volume="4331" year="2017">
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2017
</bibRefCitation>
: 104-105, fig. 69I.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
&quot;Tchen
<normalizedToken originalValue="Keou”">K'eou&quot;</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
Cheng-Kou County, Chong-qing, China, HMT-218a, deposited in IZCAS (syntype: labelled as lectotype, but probably there was no valid lectotype designation). No type specimens deposited in American museums were reported by
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.141074" author="Johnson, RJ" journalOrPublisher="Cambridge, Massachusetts (Special Occasional Publication, Department of Mollusks, Museum of Comparative Zoology, Harvard University)" pageId="0" pageNumber="1" refId="B92" refString="Johnson, RJ, 1973. Heude's Molluscan Types: or, Asian land and fresh water mollusks, mostly from the People's Republic of China, described by P. M. Heude. Cambridge, Massachusetts (Special Occasional Publication, Department of Mollusks, Museum of Comparative Zoology, Harvard University), DOI: https://doi.org/10.5962/bhl.title.141074" title="Heude's Molluscan Types: or, Asian land and fresh water mollusks, mostly from the People's Republic of China, described by P. M. Heude." url="https://doi.org/10.5962/bhl.title.141074" year="1973">Johnson (1973)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species could be examined for the first time, since it was not examined in our previous paper (
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4331.1.1" author="Pall-Gergely, B" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 124" refId="B162" refString="Pall-Gergely, B, Hunyadi, A, Đỗ, ĐS, Naggs, F, Asami, T, 2017. Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus. Zootaxa 4331 (1): 1 - 124, DOI: https://doi.org/10.11646/zootaxa.4331.1.1" title="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus." url="https://doi.org/10.11646/zootaxa.4331.1.1" volume="4331" year="2017">
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2017
</bibRefCitation>
). Protoconch low, smooth; R1 without spiral striation, its beginning is densely, finely ribbed, which gradually changes to a more widely-spaced, strongly ribbed surface. R2 and R3 are of comparable length, R2 ribs dense, low, blunt, not elevated. The most similar species is
<taxonomicName lsidName="D. fargesianus" pageId="0" pageNumber="1" rank="species" species="fargesianus">
<emphasis italics="true" pageId="0" pageNumber="1">D. fargesianus</emphasis>
</taxonomicName>
, which has denser ribs on R1 and R3, and more marked swelling on R3.
</paragraph>
<paragraph pageId="0" pageNumber="1">
The examined specimen has a shorter R3 than the one illustrated by
<bibRefCitation author="Heude, RPM" journalOrPublisher="Mission Catholique, Chang-Hai" pageId="0" pageNumber="1" pagination="125 - 188" refId="B84" refString="Heude, RPM, 1890. Memoires concernant l'histoire naturelle de l'empire chinois par des peres de la Compagnie de JESUS. Notes sur les Mollusques terrestres de la vallee du Fleuve Bleu. Mission Catholique, Chang-Hai 4: 125 - 188" title="Memoires concernant l'histoire naturelle de l'empire chinois par des peres de la Compagnie de JESUS. Notes sur les Mollusques terrestres de la vallee du Fleuve Bleu." volume="4" year="1890">Heude (1890)</bibRefCitation>
(see also
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4331.1.1" author="Pall-Gergely, B" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 124" refId="B162" refString="Pall-Gergely, B, Hunyadi, A, Đỗ, ĐS, Naggs, F, Asami, T, 2017. Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus. Zootaxa 4331 (1): 1 - 124, DOI: https://doi.org/10.11646/zootaxa.4331.1.1" title="Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus." url="https://doi.org/10.11646/zootaxa.4331.1.1" volume="4331" year="2017">
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2017
</bibRefCitation>
: fig. 69I). This raises some doubts about the identity of this type.
</paragraph>
<caption doi="10.3897/zookeys.981.53583.figure16" httpUri="https://binary.pensoft.net/fig/468661" pageId="0" pageNumber="1" start="Figure 16" startId="F16">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 16.</emphasis>
<taxonomicName baseAuthorityName="Heude" baseAuthorityYear="1886" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dicharax elevatus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="elevatus">Dicharax (?) elevatus</taxonomicName>
(Heude, 1886), syntype (HMT-218a). Photographs: Kaibaryer Meng.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>