184 lines
16 KiB
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184 lines
16 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.981.53583" ID-GBIF-Dataset="8e14a5b5-aa20-4050-b2b4-f393764fc6c7" ID-Pensoft-Pub="1313-2970-981-1" ID-Pensoft-UUID="7C44C797C4125A71BAE032A55E6FA5DC" ID-ZooBank="5194AAC86B8A473F8A41470A60182A0B" ModsDocID="1313-2970-981-1" checkinTime="1604011718171" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Sajan, Sheikh, Tripathy, Basudev, Meng, Kaibaryer, Asami, Takahiro & Ablett, Jonathan D." docDate="2020" docId="D4363F5432255752818F836036CA4B3D" docLanguage="en" docName="ZooKeys 981: 1-220" docOrigin="ZooKeys 981" docSource="http://dx.doi.org/10.3897/zookeys.981.53583" docTitle="Dioryx Benson 1859" docType="treatment" docVersion="5" id="7C44C797C4125A71BAE032A55E6FA5DC" lastPageNumber="1" masterDocId="7C44C797C4125A71BAE032A55E6FA5DC" masterDocTitle="Genus-level revision of the Alycaeidae (Gastropoda, Cyclophoroidea), with an annotated species catalogue" masterLastPageNumber="220" masterPageNumber="1" pageNumber="1" updateTime="1634714035540" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Genus-level revision of the Alycaeidae (Gastropoda, Cyclophoroidea), with an annotated species catalogue</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pall-Gergely, Barna</mods:namePart>
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<mods:affiliation>Plant Protection Institute, Centre for Agricultural Research, Herman Otto ut 15, Budapest, H- 1022, Hungary</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
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<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sajan, Sheikh</mods:namePart>
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<mods:affiliation>Zoological Survey of India, Prani Vigyan Bhawan, M Block, New Alipore, Kolkata 700053, West Bengal, India & Wildlife Institute of India, Chandrabani, Dehradun 248 002, Uttarakhand, India</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-2785-6824</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tripathy, Basudev</mods:namePart>
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<mods:affiliation>Zoological Survey of India, Prani Vigyan Bhawan, M Block, New Alipore, Kolkata 700053, West Bengal, India</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Meng, Kaibaryer</mods:namePart>
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<mods:affiliation>National Zoological Museum of China, Institute of Zoology, Chinese Academy of Sciences, Beijing, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Asami, Takahiro</mods:namePart>
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<mods:affiliation>Department of Biology, Shinshu University, Matsumoto 390 - 8621, Japan</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-5706-0272</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ablett, Jonathan D.</mods:namePart>
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<mods:affiliation>Mollusca Section, Invertebrates Division, Department of Life Sciences, The Natural History Museums, London SW 7 5 BD, United Kingdom</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>981</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>220</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.981.53583</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.981.53583</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-981-1</mods:identifier>
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<mods:identifier type="ZooBank">5194AAC86B8A473F8A41470A60182A0B</mods:identifier>
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<mods:identifier type="Pensoft-UUID">7C44C797C4125A71BAE032A55E6FA5DC</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="168714974" LSID="urn:lsid:plazi:treatment:D4363F5432255752818F836036CA4B3D" httpUri="http://treatment.plazi.org/id/D4363F5432255752818F836036CA4B3D" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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Genus
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<taxonomicName LSID="D4363F54-3225-5752-818F-836036CA4B3D" authority="Benson, 1859" authorityName="Benson" authorityYear="1859" class="Gastropoda" family="Cyclophoridae" genus="Dioryx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dioryx" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Dioryx Benson, 1859</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Benson" authorityYear="1859" class="Gastropoda" family="Cyclophoridae" genus="Dioryx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dioryx" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Dioryx</taxonomicName>
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Benson, 1859: 177. [section (subgenus) of
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<taxonomicName authorityName="Gray" authorityYear="1850" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Alycaeus</taxonomicName>
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]
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Benson" authorityYear="1859" class="Gastropoda" family="Cyclophoridae" genus="Dioryx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dioryx" order="Gastropoda" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Dioryx</taxonomicName>
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-
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<bibRefCitation author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/28228303" refId="B98" refString="Kobelt, W, Moellendorff, O von, 1897. Catalog der gegenvaertig lebend bekannten Pneumonopomen. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 29: 73-88, 105-120, 137-152. https://www.biodiversitylibrary.org/page/28228303" title="Catalog der gegenvaertig lebend bekannten Pneumonopomen. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 29: 73 - 88, 105 - 120, 137 - 152." url="https://www.biodiversitylibrary.org/page/28228303" year="1897">
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Kobelt and
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<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
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1897
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</bibRefCitation>
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: 146;
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.12890" author="Gude, GK" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B74" refString="Gude, GK, 1921. The Fauna of British India including Ceylon and Burma. Mollusca. III. Land operculates (Cyclophoridae, Truncatellidae, Assimineidae, Helicinidae). Taylor and Francis, London, DOI: https://doi.org/10.5962/bhl.title.12890" title="The Fauna of British India including Ceylon and Burma. Mollusca. III. Land operculates (Cyclophoridae, Truncatellidae, Assimineidae, Helicinidae)." url="https://doi.org/10.5962/bhl.title.12890" year="1921">Gude 1921</bibRefCitation>
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: 198;
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<bibRefCitation author="Thiele, J" journalOrPublisher="Verlag von Gustav Fischer, Jena" pageId="0" pageNumber="1" refId="B213" refString="Thiele, J, 1929. Handbuch der systematischen Weichtierkunde, Teil 1. Verlag von Gustav Fischer, Jena" title="Handbuch der systematischen Weichtierkunde, Teil 1." year="1929">Thiele 1929</bibRefCitation>
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: 107; Wenz 1938: 477;
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<bibRefCitation author="Egorov, RV" journalOrPublisher="Egorov, Moscow" pageId="0" pageNumber="1" refId="B45" refString="Egorov, RV, 2013. Treasure of Russian Shells. Supplement 3. A review of the genera of the terrestrial pectinibranch molluscs (synopsis mainly based on published data). Part III. Littoriniformes: Liareidae, Pupinidae, Diplommatinidae, Alycaeidae, Cochlostomidae. R. Egorov, Moscow" title="Treasure of Russian Shells. Supplement 3. A review of the genera of the terrestrial pectinibranch molluscs (synopsis mainly based on published data). Part III. Littoriniformes: Liareidae, Pupinidae, Diplommatinidae, Alycaeidae, Cochlostomidae. R." year="2013">Egorov 2013</bibRefCitation>
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: 34.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type species">
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<paragraph pageId="0" pageNumber="1">Type species.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Benson" authorityYear="1856" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus amphora" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="Amphora">
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<emphasis italics="true" pageId="0" pageNumber="1">Alycaeus amphora</emphasis>
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</taxonomicName>
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Benson, 1856 (Fig.
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<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Dioryx amphora (Benson, 1856), probable syntype (UMZC I. 102850; type species of Dioryx). Photographs: Harold Taylor (NHM)." figureDoi="10.3897/zookeys.981.53583.figure26" httpUri="https://binary.pensoft.net/fig/468671" pageId="0" pageNumber="1">26</figureCitation>
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), SD
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.12890" author="Gude, GK" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B74" refString="Gude, GK, 1921. The Fauna of British India including Ceylon and Burma. Mollusca. III. Land operculates (Cyclophoridae, Truncatellidae, Assimineidae, Helicinidae). Taylor and Francis, London, DOI: https://doi.org/10.5962/bhl.title.12890" title="The Fauna of British India including Ceylon and Burma. Mollusca. III. Land operculates (Cyclophoridae, Truncatellidae, Assimineidae, Helicinidae)." url="https://doi.org/10.5962/bhl.title.12890" year="1921">Gude (1921</bibRefCitation>
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: 198).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Shell small to very large (3.5-9 mm), globose, sometimes the body whorl is angled or keeled; protoconch smooth, not spirally striated; R1 usually very finely reticulated due to fine radial ribs and fine spiral striation, or smooth; R2 short to long, usually without ribs (superficially smooth, with alternating lighter and darker stripes); R3 practically absent (the inner opening of the sutural tube is situated close to the aperture). Operculum thin, usually proteinaceous (
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<normalizedToken originalValue="“horny”">"horny"</normalizedToken>
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), in some species can be thicker, calcified. Central tooth with 5-7 cusps, broad, central cusp blunt.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
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<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The combination of the globular shell, the reduced sculpture (only very fine spiral and radial lines are present), and the absence of the R2 region characteristic for the vast majority of alycaeid species make
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<taxonomicName authorityName="Benson" authorityYear="1859" class="Gastropoda" family="Alycaeidae" genus="Dioryx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dioryx" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Dioryx</emphasis>
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</taxonomicName>
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species distinguishable from other alycaeid genera.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Benson" authorityYear="1859" class="Gastropoda" family="Alycaeidae" genus="Dioryx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dioryx" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Dioryx</emphasis>
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</taxonomicName>
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is distributed from the southeastern Himalayan region to Taiwan in the east, and down to the northern part of the Malay Peninsula and southern Vietnam in the south (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution of Alycaeus Gray, 1850 (dark shaded area) and Dioryx Benson, 1859 (light shaded area)." figureDoi="10.3897/zookeys.981.53583.figure7" httpUri="https://binary.pensoft.net/fig/468652" pageId="0" pageNumber="1">7</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Benson" authorityYear="1859" class="Gastropoda" family="Alycaeidae" genus="Dioryx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dioryx" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Dioryx</emphasis>
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</taxonomicName>
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is primarily classified based on the reduced R3. This character state is found in other genera of the
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<taxonomicName authorityName="Blanford" authorityYear="1864" family="Alycaeidae" lsidName="" pageId="0" pageNumber="1" rank="family">Alycaeidae</taxonomicName>
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as well, such as in
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<taxonomicName authorityName="Fulton" authorityYear="1902" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus conformis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="Conformis">
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<emphasis italics="true" pageId="0" pageNumber="1">Alycaeus conformis</emphasis>
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</taxonomicName>
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,
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<taxonomicName baseAuthorityName="Moellendorff" baseAuthorityYear="1887" class="Gastropoda" family="Alycaeidae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus microconus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="Microconus">
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<emphasis italics="true" pageId="0" pageNumber="1">Chamalycaeus microconus</emphasis>
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</taxonomicName>
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, and in
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<taxonomicName authorityName="Kuroda" authorityYear="1951" class="Gastropoda" family="Alycaeidae" genus="Awalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Awalycaeus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Awalycaeus</emphasis>
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</taxonomicName>
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(treated as a synonym of
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<taxonomicName authorityName="Kobelt & Moellendorff" authorityYear="1900" class="Gastropoda" family="Alycaeidae" genus="Dicharax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dicharax" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Dicharax</emphasis>
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</taxonomicName>
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). Even presuming that similar morphology has appeared multiple times in the
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<taxonomicName authorityName="Blanford" authorityYear="1864" family="Alycaeidae" lsidName="" pageId="0" pageNumber="1" rank="family">Alycaeidae</taxonomicName>
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, we can reasonably assume the monophyly of
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<taxonomicName authorityName="Benson" authorityYear="1859" class="Gastropoda" family="Alycaeidae" genus="Dioryx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dioryx" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Dioryx</emphasis>
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</taxonomicName>
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due to the uniform, simple sculpture across all species, and the generally globose shell.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |