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<document id="97B10B0BC2A8D859D59853B7C52F2135" ID-CLB-Dataset="32713" ID-DOI="10.11646/zootaxa.4287.1.1" ID-GBIF-Dataset="91e8f5fd-1e15-4c4f-872f-3d4c50d0c8a7" ID-ISSN="1175-5326" ID-Zenodo-Dep="829051" ID-ZooBank="13C4F702-EF00-4F04-B38E-3F0AA6CAF718" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500110702684" checkinUser="plazi" docAuthor="ROBERTO CALDARA, CHARLES W. OBRIEN &amp; MASSIMO MEREGALLI" docDate="2017" docId="D17787854F561F210DD76E61164BFBBB" docLanguage="en" docName="zootaxa.4287.1.1.pdf.imf" docOrigin="Zootaxa 4287 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Bagous argillaceus" docType="treatment" docVersion="8" lastPageNumber="44" masterDocId="2D4EFFFD4F7D1F0A0D406F35171FFFD9" masterDocTitle="A phylogenetic analysis of the aquatic weevil tribe Bagoini (Coleoptera: Curculionidae) based on morphological characters of adults" masterLastPageNumber="63" masterPageNumber="1" pageNumber="44" updateTime="1698465391549" updateUser="ExternalLinkService">
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<mods:title id="889258B07D21D9135B221FAFA5073443">A phylogenetic analysis of the aquatic weevil tribe Bagoini (Coleoptera: Curculionidae) based on morphological characters of adults</mods:title>
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<mods:namePart id="EEF2144EB308B071DA6BCE443879B2B9">ROBERTO CALDARA</mods:namePart>
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<mods:namePart id="E73AEAEE13AFD8DA29425028FC677EAD">CHARLES W. OBRIEN</mods:namePart>
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<treatment id="D17787854F561F210DD76E61164BFBBB" ID-DOI="http://doi.org/10.5281/zenodo.4386690" ID-GBIF-Taxon="132549348" ID-Zenodo-Dep="4386690" LSID="urn:lsid:plazi:treatment:D17787854F561F210DD76E61164BFBBB" httpUri="http://treatment.plazi.org/id/D17787854F561F210DD76E61164BFBBB" lastPageNumber="44" pageId="43" pageNumber="44">
<subSubSection id="11C465184F561F210DD76E6116E2FEB7" box="[151,509,340,366]" pageId="43" pageNumber="44" type="nomenclature">
<paragraph id="596136934F561F210DD76E6116E2FEB7" blockId="43.[151,509,340,366]" box="[151,509,340,366]" pageId="43" pageNumber="44">
<heading id="022981FF4F561F210DD76E6116E2FEB7" bold="true" box="[151,509,340,366]" fontSize="11" level="1" pageId="43" pageNumber="44" reason="1">
<emphasis id="6BAAEA814F561F210DD76E6116E2FEB7" bold="true" box="[151,509,340,366]" pageId="43" pageNumber="44">
12.
<taxonomicName id="9EDE4D104F561F210D836E6116B5FEB7" ID-CoL="KDMD" box="[195,426,340,366]" class="Insecta" family="Curculionidae" genus="Bagous" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="argillaceus">
<emphasis id="6BAAEA814F561F210D836E6116B5FEB7" bold="true" box="[195,426,340,366]" italics="true" pageId="43" pageNumber="44">Bagous argillaceus</emphasis>
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group
</emphasis>
</heading>
</paragraph>
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<subSubSection id="11C465184F561F210DD76EAE1561FD55" pageId="43" pageNumber="44" type="diagnosis">
<paragraph id="596136934F561F210DD76EAE1561FD55" blockId="43.[151,1437,411,1122]" pageId="43" pageNumber="44">
<emphasis id="6BAAEA814F561F210DD76EAE160EFE6D" bold="true" box="[151,273,411,436]" pageId="43" pageNumber="44">Diagnosis.</emphasis>
Scales smooth and shiny, with at most a very fine pit (char. 3.1). Supraocular setae very slender, long, erect and curved. Pronotum with complete median longitudinal sulcus or impression. Prosternum with scarcely evident canal, lateral margin more or less parallel and weakly raised (char. 31.1). Sclerites complex within tube of penis body with two posteriorly joined pairs of subapical sclerites, one submedian and the other sublateral (char. 84.4). Penis body with pseudo-orifice large to very large, apparent orifice strongly extended proximally, ventrally sinuate in area beneath orifice, lateral surface ventral to flange concave. Apodemes of spiculum ventrale strongly divergent, contiguous only at extreme base.
</paragraph>
</subSubSection>
<subSubSection id="11C465184F561F210D876DA3164BFBBB" pageId="43" pageNumber="44" type="discussion">
<paragraph id="596136934F561F210D876DA3127DFC2D" blockId="43.[151,1437,411,1122]" pageId="43" pageNumber="44">
<emphasis id="6BAAEA814F561F210D876DA31624FD76" bold="true" box="[199,315,662,687]" pageId="43" pageNumber="44">Remarks.</emphasis>
This group is characterized by one synapomorphy from external morphology, the vestiture composed of almost smooth and distinctly shiny scales, and two synapomorphies from the genitalia. It was supported in all of the phylogenetic analyses. Prevously,
<taxonomicName id="9EDE4D104F561F210E646DD51493FD2E" box="[804,908,735,759]" class="Insecta" family="Curculionidae" genus="Bagous" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="6BAAEA814F561F210E646DD51493FD2E" box="[804,908,735,759]" italics="true" pageId="43" pageNumber="44">B. affinis</emphasis>
</taxonomicName>
was placed in a separate monotypic group by OBrien &amp; Askevold (1995) due to two autapomorphies (scales on pronotal disc sublunate and distinctly emarginate and penis apodemes extremely short; char. 28.1 and 110.2, respectively) and the lack of four synapomorphies of the genitalia possessed by the other species of the
<taxonomicName id="9EDE4D104F561F210E986C791366FCBD" box="[984,1145,843,868]" class="Insecta" family="Curculionidae" genus="Bagous" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="argillaceus">
<emphasis id="6BAAEA814F561F210E986C791366FCBD" box="[984,1145,843,868]" italics="true" pageId="43" pageNumber="44">B. argillaceus</emphasis>
</taxonomicName>
group—laminar orificial lateral sclerites present although sometimes only distinguishable at the apical portion (char. 74.1); orificial lateral sclerites obliquely uniformly raised (char. 75.1) and with basal margin distinct; orifice with lateral, transverse vertical sclerite (char. 83.1); and presence of a bursa with sclerites more or less bilobed and convex (char. 116.1). It is represented by four Palaearctic species, three of which with an eastern distribution, and two Indian species.
</paragraph>
<paragraph id="596136934F561F210D876CCB164BFBBB" blockId="43.[151,1437,411,1122]" pageId="43" pageNumber="44">
Species included.
<emphasis id="6BAAEA814F561F210CD66CCB16C2FBCE" bold="true" box="[406,477,1022,1047]" pageId="43" pageNumber="44">ORR:</emphasis>
<taxonomicName id="9EDE4D104F561F210CBC6B351443FBCE" authority="Hustache, 1926" authorityName="Hustache" authorityYear="1926" box="[508,860,1022,1047]" class="Insecta" family="Curculionidae" genus="Bagous" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="6BAAEA814F561F210CBC6B351580FBCE" box="[508,671,1023,1047]" italics="true" pageId="43" pageNumber="44">Bagous affinis</emphasis>
Hustache, 1926
</taxonomicName>
; ¹
<taxonomicName id="9EDE4D104F561F210E326B351250FBCE" authority="O'Brien &amp; Pajni, 1989" authorityName="O'Brien &amp; Pajni" authorityYear="1989" box="[882,1359,1022,1047]" class="Insecta" family="Curculionidae" genus="Bagous" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="laevigatus">
<emphasis id="6BAAEA814F561F210E326B35135DFBCE" box="[882,1090,1022,1047]" italics="true" pageId="43" pageNumber="44">Bagous laevigatus</emphasis>
<bibRefCitation id="3D4F4B624F561F21090B6CCB1250FBCE" author="O'Brien" box="[1099,1359,1022,1047]" pageId="43" pageNumber="44" refString="O'Brien, C. W. &amp; Pajni, R. H. (1989) Two Indian Bagous weevils (Coleoptera, Curculionidae), tuber feeders of Hydrilla verticillata (Hydrocharitaceae), one a potential biocontrol agent in Florida. Entomologist, 72 (3), 462 - 468. https: // doi. org / 10.2307 / 3495184" type="journal article" year="1989">O'Brien &amp; Pajni, 1989</bibRefCitation>
</taxonomicName>
;
<emphasis id="6BAAEA814F561F21081C6CCB1285FBCE" bold="true" box="[1372,1434,1022,1047]" pageId="43" pageNumber="44">PAL:</emphasis>
*§¹
<taxonomicName id="9EDE4D104F561F210DF56B121506FBE7" authority="Gyllenhal, 1836" authorityName="Gyllenhal" authorityYear="1836" box="[181,537,1061,1086]" class="Insecta" family="Curculionidae" genus="Bagous" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="argillaceus">
<emphasis id="6BAAEA814F561F210DF56B12164BFBE7" box="[181,340,1061,1086]" italics="true" pageId="43" pageNumber="44">B. argillaceus</emphasis>
Gyllenhal, 1836
</taxonomicName>
, ¹
<taxonomicName id="9EDE4D104F561F210F6F6B12146AFBE7" authority="Hartmann, 1899" authorityName="Hartmann" authorityYear="1899" box="[559,885,1062,1086]" class="Insecta" family="Curculionidae" genus="Bagous" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="foersteri">
<emphasis id="6BAAEA814F561F210F6F6B1215AFFBE7" box="[559,688,1062,1086]" italics="true" pageId="43" pageNumber="44">B. foersteri</emphasis>
Hartmann, 1899
</taxonomicName>
, *²
<taxonomicName id="9EDE4D104F561F210EDB6B1213EDFBE7" authority="Dieckmann, 1975" authorityName="Dieckmann" authorityYear="1975" box="[923,1266,1061,1086]" class="Insecta" family="Curculionidae" genus="Bagous" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="fremuthi">
<emphasis id="6BAAEA814F561F210EDB6B121304FBE7" box="[923,1051,1061,1086]" italics="true" pageId="43" pageNumber="44">B. fremuthi</emphasis>
Dieckmann, 1975
</taxonomicName>
, ¹
<taxonomicName id="9EDE4D104F561F2108486B12164FFBBB" authority="Hartmann, 1899" authorityName="Hartmann" authorityYear="1899" class="Insecta" family="Curculionidae" genus="Bagous" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="6BAAEA814F561F2108486B121283FBE7" box="[1288,1436,1061,1086]" italics="true" pageId="43" pageNumber="44">B. sulcicollis</emphasis>
Hartmann, 1899
</taxonomicName>
.
</paragraph>
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