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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.872.35361" ID-GBIF-Dataset="ed1a1322-b118-4434-b9ff-503a0fb64792" ID-PMC="PMC6711934" ID-Pensoft-Pub="1313-2970-872-13" ID-Pensoft-UUID="A9BE568D9BF65A179464AB8C7CB8BEB2" ID-PubMed="31496884" ID-ZooBank="DCDEFCB1E33D476F90D7178BC08C311A" ModsDocID="1313-2970-872-13" checkinTime="1566364211746" checkinUser="pensoft" docAuthor="Astafurova, Yulia V., Proshchalykin, Maxim Yu. &amp; Schwarz, Maximilian" docDate="2019" docId="D12FCFAA24895241B1E7149122E79AA7" docLanguage="en" docName="ZooKeys 872: 13-40" docOrigin="ZooKeys 872" docSource="http://dx.doi.org/10.3897/zookeys.872.35361" docTitle="Sphecodes villosulus Schwarz 2010" docType="treatment" docVersion="4" id="A9BE568D9BF65A179464AB8C7CB8BEB2" lastPageNumber="33" masterDocId="A9BE568D9BF65A179464AB8C7CB8BEB2" masterDocTitle="The distribution of the genus Sphecodes Latreille (Hymenoptera, Halictidae) of the Arabian Peninsula and surrounding countries with description of hitherto unknown female of S. atlanticus Warncke, 1992 and male of S. dathei Schwarz, 2010" masterLastPageNumber="40" masterPageNumber="13" pageNumber="33" updateTime="1668167741671" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>The distribution of the genus Sphecodes Latreille (Hymenoptera, Halictidae) of the Arabian Peninsula and surrounding countries with description of hitherto unknown female of S. atlanticus Warncke, 1992 and male of S. dathei Schwarz, 2010</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Astafurova, Yulia V.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Proshchalykin, Maxim Yu.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schwarz, Maximilian</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>872</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>13</mods:start>
<mods:end>40</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.872.35361</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.872.35361</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-872-13</mods:identifier>
<mods:identifier type="ZooBank">DCDEFCB1E33D476F90D7178BC08C311A</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="159217126" LSID="urn:lsid:plazi:treatment:D12FCFAA24895241B1E7149122E79AA7" httpUri="http://treatment.plazi.org/id/D12FCFAA24895241B1E7149122E79AA7" lastPageNumber="33" pageId="20" pageNumber="33">
<subSubSection pageId="20" pageNumber="33" type="nomenclature">
<paragraph pageId="20" pageNumber="33">
<taxonomicName LSID="d12fcfaa-2489-5241-b1e7-149122e79aa7" authority="Schwarz, 2010" authorityName="Schwarz" authorityYear="2010" class="Insecta" family="Halictidae" genus="Sphecodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphecodes villosulus" order="Hymenoptera" pageId="20" pageNumber="33" phylum="Arthropoda" rank="species" species="villosulus">Sphecodes villosulus Schwarz, 2010</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="33" type="reference_group">
<paragraph pageId="20" pageNumber="33">
<taxonomicName class="Insecta" family="Halictidae" genus="Sphecodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphecodes villosulus" order="Hymenoptera" pageId="20" pageNumber="33" phylum="Arthropoda" rank="species" species="villosulus">Sphecodes villosulus</taxonomicName>
Schwarz, 2010: 486-491, ♀, ♂ (holotype: ♀, United Arab Emirates, Dubai, Nakhalai, 28-30.IV.1984, in Malaise trap, E. Sugden leg.; BLCU).
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="33" type="diagnosis">
<paragraph pageId="20" pageNumber="33">Diagnosis.</paragraph>
<paragraph pageId="20" pageNumber="33">
This species differs from other small Palaearctic species with 5-6 hamuli in the hind wing by having a unique combination of simple mandibles and the male gonocoxite dorsally with an impression. The female is closest to
<taxonomicName lsidName="S. armeniacus" pageId="20" pageNumber="33" rank="species" species="armeniacus">
<emphasis italics="true" pageId="20" pageNumber="33">S. armeniacus</emphasis>
</taxonomicName>
owing to dense appressed snow-white pubescence obscuring the integument on face, a transverse head and sparsely punctate mesoscutum, but differs from this species by sparser and finer punctate ocello-ocular area (3-5
<normalizedToken originalValue="μm">μm</normalizedToken>
/ 2-3 versus 5-10
<normalizedToken originalValue="μm">μm</normalizedToken>
/ 1-2) and strongly transverse F3 (almost square in
<taxonomicName lsidName="S. armeniacus" pageId="20" pageNumber="33" rank="species" species="armeniacus">
<emphasis italics="true" pageId="20" pageNumber="33">S. armeniacus</emphasis>
</taxonomicName>
). The male of
<taxonomicName lsidName="S. villosulus" pageId="20" pageNumber="33" rank="species" species="villosulus">
<emphasis italics="true" pageId="20" pageNumber="33">S. villosulus</emphasis>
</taxonomicName>
recalls
<taxonomicName lsidName="S. miniatus" pageId="20" pageNumber="33" rank="species" species="miniatus">
<emphasis italics="true" pageId="20" pageNumber="33">S. miniatus</emphasis>
</taxonomicName>
in the rectangular gonostylar shape, but clearly differs from this species by the less developed tyloids on the flagellomeres extending to approximately a half of ventral flagellar surfaces (versus those across 4/5).
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="33" type="materials_examined">
<paragraph pageId="20" pageNumber="33">Material examined.</paragraph>
<paragraph pageId="20" pageNumber="33">
UNITED ARAB EMIRATES: 1 ♀, 1♂, Abu Dhabi, 30.I.1987, I. Hamer (NHMUK 013380419, 013380423); 1 ♀, 2♂♂, Abu Dhabi, 31.III.1987 (NHMUK 013380418, 013380421, 013380420); 1 ♀, idem, 10.IV.1987, I. Hamer (NHMUK 013380422); 1 ♀, Hatta, 20.XII.1990 (NHMUK 013380424); 2 ♂♂, idem, 23.VIII.1991 (NHMUK 013380426, 013380427); 1 ♀, idem, 5.III.1993, I. Hamer (NHMUK 013380425); 3 ♀♀, North Ras, Al Khaimah, 17.II.2018 (M. Mokrousov) (ZISP); SAUDI ARABIA: 1 ♀, Riyadh, El
<normalizedToken originalValue="Hair">Ha'ir</normalizedToken>
, 19.III.1980, K.M. Guichard (NHMUK 013380448); OMAN: 1 ♀, Rostaq, 350 m, 21-31.III.1976, K.M.Guichard (NHMUK 013380444).
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="33" type="published records">
<paragraph pageId="20" pageNumber="33">Published records.</paragraph>
<paragraph pageId="20" pageNumber="33">
<bibRefCitation author="Schwarz, M" journalOrPublisher="Arthropod fauna of the UAE" pageId="23" pageNumber="36" pagination="483 - 491" refId="B45" refString="Schwarz, M, 2010. Order Hymenoptera, family Halictidae. Supplementary records of the genus Sphecodes Latreille. . Arthropod fauna of the UAE 3: 483 - 491" title="Order Hymenoptera, family Halictidae. Supplementary records of the genus Sphecodes Latreille." volume="3" year="2010">Schwarz 2010</bibRefCitation>
: 486 (United Arab Emirates);
<bibRefCitation author="Ascher, JS" journalOrPublisher="Zoologica Scripta" pageId="21" pageNumber="34" publicationUrl="http://www.discoverlife.org/mp/20q?guide=Apoidea_species" refId="B3" refString="Ascher, JS, Pickering, J, 2019. . http://www.discoverlife.org/mp/20q?guide=Apoidea_species" url="http://www.discoverlife.org/mp/20q?guide=Apoidea_species" year="2019">Ascher and Pickering 2019</bibRefCitation>
(United Arab Emirates).
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="33" type="distribution">
<paragraph pageId="20" pageNumber="33">Distribution.</paragraph>
<paragraph pageId="20" pageNumber="33">United Arab Emirates, *Oman, *Saudi Arabia.</paragraph>
</subSubSection>
</treatment>
</document>