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<document id="8B59DF07521492DC0837D1E7087178EB" ID-CLB-Dataset="46084" ID-DOI="10.5281/zenodo.201284" ID-GBIF-Dataset="57b8e376-da3c-4eef-8fe8-30630518f583" ID-ISSN="1175-5326" ID-Zenodo-Dep="201284" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460404852598" checkinUser="plazi" docAuthor="Hausmann, Axel &amp; Huemer, Peter" docDate="2011" docId="D1232C65FF996046FF45F9E2FEE2C8D7" docLanguage="en" docName="zt03070p068.pdf" docOrigin="Zootaxa 3070" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Acasis appensata subsp. callaina Hausmann &amp; Huemer, ssp. nov." docType="treatment" docVersion="10" lastPageNumber="66" masterDocId="2D1A541DFF9B6040FFD2FF9DFFCFCD25" masterDocTitle="Taxonomic decision as a compromise: Acasis appensata (Eversmann, 1832) in Central Italy — a case of conflicting evidence between DNA barcode and morphology (Lepidoptera: Geometridae)" masterLastPageNumber="68" masterPageNumber="60" pageNumber="62" updateTime="1698299646792" updateUser="ExternalLinkService">
<mods:mods id="AEF3412172DC3C0041A3CA6698E6EF07" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D335D154361B5F0D963C63D0DE4534D1">
<mods:title id="10B957C832F0BB074E4DD35800D243BE">Taxonomic decision as a compromise: Acasis appensata (Eversmann, 1832) in Central Italy — a case of conflicting evidence between DNA barcode and morphology (Lepidoptera: Geometridae)</mods:title>
</mods:titleInfo>
<mods:name id="A92B87EC2571BAE463B2CD49381BF78A" type="personal">
<mods:role id="6A2779B8B8396B145B1E96080554E3A7">
<mods:roleTerm id="26B1A6656649D7D5C5F240AA9C1EA304">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="669C9DE1D535E6FDE7B91398AE40FCEE">Hausmann, Axel</mods:namePart>
</mods:name>
<mods:name id="3C7742A25ABEB21C9F65E7E016241019" type="personal">
<mods:role id="EBE0B99DDC546010C1D67C37CB1C12FF">
<mods:roleTerm id="AB56647FDA0D7F74886E20D821B51F26">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="55DC2C68421AED909393D5AC99ABEB0C">Huemer, Peter</mods:namePart>
</mods:name>
<mods:typeOfResource id="52141BD19CCE67A1466EF07662444CB9">text</mods:typeOfResource>
<mods:relatedItem id="4DA1C6583CABFEA0EE5EA4EE4D1ADD35" type="host">
<mods:titleInfo id="ACF2459C611DC7EF3DCD55051EA01B6E">
<mods:title id="B8C8A25EA5CE4F861F7163BA0AEBFB4C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A41B5998ABB78288CC256137B491E3EA">
<mods:date id="7E294D45A464035C5AE1CB2E3292BE4A">2011</mods:date>
<mods:detail id="05DED8F54B0A1AB4B77C489B79B4DBF6" type="volume">
<mods:number id="1CDBF27EE258C31E0A443C6904C22B5A">3070</mods:number>
</mods:detail>
<mods:extent id="49BF3760013C9EFF1F91C6A5BE8465CC" unit="page">
<mods:start id="529387A553F603244BE4D031FF77A197">60</mods:start>
<mods:end id="795D5756FA26A23D9E3A8931E894F878">68</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="4E532FD4D5C7CFF4EF91D69683A8C15E">journal article</mods:classification>
<mods:identifier id="5C29F2ADF23CC47483E1878AD9083265" type="CLB-Dataset">46084</mods:identifier>
<mods:identifier id="2C6B4FB225D3C3867DFBCFA099B71E51" type="DOI">10.5281/zenodo.201284</mods:identifier>
<mods:identifier id="42F3886E33748CB05D086EFE1436E710" type="GBIF-Dataset">57b8e376-da3c-4eef-8fe8-30630518f583</mods:identifier>
<mods:identifier id="B8D10B4BFD5D03E217A925FFE993ACF9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="104DA8EECD1794B02DEA8DE95F8238DB" type="Zenodo-Dep">201284</mods:identifier>
</mods:mods>
<treatment id="D1232C65FF996046FF45F9E2FEE2C8D7" ID-DOI="http://doi.org/10.5281/zenodo.5659766" ID-GBIF-Taxon="119421039" ID-Zenodo-Dep="5659766" LSID="urn:lsid:plazi:treatment:D1232C65FF996046FF45F9E2FEE2C8D7" httpUri="http://treatment.plazi.org/id/D1232C65FF996046FF45F9E2FEE2C8D7" lastPageId="6" lastPageNumber="66" pageId="2" pageNumber="62">
<subSubSection id="1190CEF8FF996042FF45F9E2FCF1CBBD" box="[151,830,1663,1688]" pageId="2" pageNumber="62" type="nomenclature">
<paragraph id="59359D73FF996042FF45F9E2FCF1CBBD" blockId="2.[151,830,1663,1688]" box="[151,830,1663,1688]" pageId="2" pageNumber="62">
<heading id="027D2A1FFF996042FF45F9E2FCF1CBBD" bold="true" box="[151,830,1663,1688]" fontSize="10" level="2" pageId="2" pageNumber="62" reason="2">
<emphasis id="6BFE4161FF996042FF45F9E2FCF1CBBD" bold="true" box="[151,830,1663,1688]" pageId="2" pageNumber="62">
<taxonomicName id="9E8AE6F0FF996042FF45F9E2FD1DCBBD" authority="Hausmann &amp; Huemer" authorityName="Hausmann &amp; Huemer" box="[151,722,1663,1688]" class="Insecta" family="Geometridae" genus="Acasis" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="62" phylum="Arthropoda" rank="subSpecies" species="appensata" status="subsp. nov." subSpecies="callaina">
<emphasis id="6BFE4161FF996042FF45F9E2FE0ECBBD" bold="true" box="[151,449,1663,1688]" italics="true" pageId="2" pageNumber="62">Acasis appensata callaina</emphasis>
Hausmann &amp; Huemer
</taxonomicName>
<taxonomicNameLabel id="70CDFC1AFF996042FD0BF91DFCF1CBBD" box="[729,830,1664,1688]" pageId="2" pageNumber="62" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1190CEF8FF996042FF45F955FDAECA9D" pageId="2" pageNumber="62" type="materials_examined">
<paragraph id="59359D73FF996042FF45F955FDAECA9D" blockId="2.[151,1436,1735,1976]" pageId="2" pageNumber="62">
<emphasis id="6BFE4161FF996042FF45F955FE89CBC5" bold="true" box="[151,326,1735,1760]" pageId="2" pageNumber="62">
<typeStatus id="863123D1FF996042FF45F955FF1DCBC5" box="[151,210,1736,1760]" pageId="2" pageNumber="62">Type</typeStatus>
material.
</emphasis>
<typeStatus id="863123D1FF996042FE9CF955FE79CBC5" box="[334,438,1736,1760]" pageId="2" pageNumber="62" type="holotype">Holotype</typeStatus>
Ƥ, ‘
<collectingCountry id="219DDDE3FF996042FE35F95AFD85CBFA" box="[487,586,1735,1759]" name="Italy" pageId="2" pageNumber="62">ITALIA</collectingCountry>
, Abruzzen, Monte Terminillo,
<quantity id="9E723096FF996042FC78F95AFBCDCBFA" box="[938,1026,1735,1759]" metricMagnitude="3" metricUnit="m" metricValue="1.73" pageId="2" pageNumber="62" unit="m" value="1730.0">1730 m</quantity>
,
<geoCoordinate id="3CBEFBB4FF996042FBDDF95AFB4BCBFA" box="[1039,1156,1735,1759]" direction="east" orientation="longitude" pageId="2" pageNumber="62" precision="92" value="13.01">13°00,6´E</geoCoordinate>
,
<geoCoordinate id="3CBEFBB4FF996042FB43F95AFAC5CBFA" box="[1169,1290,1735,1759]" direction="north" orientation="latitude" pageId="2" pageNumber="62" precision="92" value="42.816666">42°49,0´N</geoCoordinate>
,
<date id="2D34BBB3FF996042FAC5F95AFA58CBFA" box="[1303,1431,1735,1759]" pageId="2" pageNumber="62" value="2010-07-16">16.07.2010</date>
, Huemer leg.
<collectionCode id="3F9B05B6FF996042FEF5F970FEBDCA21" box="[295,370,1773,1796]" httpUri="http://grbio.org/cool/7159-6y1v" name="Tiroler Landesmuseum Ferdinandeum" pageId="2" pageNumber="62">TLMF</collectionCode>
2010-015’ (
<collectionCode id="3F9B05B6FF996042FE2EF971FD82CA21" box="[508,589,1772,1796]" httpUri="http://grbio.org/cool/7159-6y1v" name="Tiroler Landesmuseum Ferdinandeum" pageId="2" pageNumber="62">TLMF</collectionCode>
).
<typeStatus id="863123D1FF996042FD8CF970FD00CA20" box="[606,719,1773,1797]" pageId="2" pageNumber="62" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="219DDDE3FF996042FD0BF971FCDECA21" box="[729,785,1772,1796]" name="Italy" pageId="2" pageNumber="62">Italy</collectingCountry>
:
<date id="2D34BBB3FF996042FCC9F971FCA1CA21" box="[795,878,1772,1796]" pageId="2" pageNumber="62" value="2010-01-03">1 3, 10</date>
Ƥ, same data as
<typeStatus id="863123D1FF996042FBF8F971FB43CA21" box="[1066,1164,1772,1796]" pageId="2" pageNumber="62" type="holotype">holotype</typeStatus>
(genitalia slides G 1456 Ƥ, G 1458 Ƥ, G 1461 3 (
<collectionCode id="3F9B05B6FF996042FE12F892FDDECA02" box="[448,529,1807,1831]" httpUri="http://grbio.org/cool/7159-6y1v" name="Tiroler Landesmuseum Ferdinandeum" pageId="2" pageNumber="62">TLMF</collectionCode>
); 4 Ƥ, ditto, but
<date id="2D34BBB3FF996042FD02F892FC8FCA02" box="[720,832,1807,1831]" pageId="2" pageNumber="62" value="2010-12-07">12.7.2010</date>
(
<collectionCode id="3F9B05B6FF996042FC83F892FC6DCA02" box="[849,930,1807,1831]" httpUri="http://grbio.org/cool/7159-6y1v" name="Tiroler Landesmuseum Ferdinandeum" pageId="2" pageNumber="62">TLMF</collectionCode>
); 1 3, ditto, but
<date id="2D34BBB3FF996042FBB2F892FB00CA02" box="[1120,1231,1807,1831]" pageId="2" pageNumber="62" value="2010-11-07">11.7.2010</date>
(genitalia slide G 1464 3 (
<collectionCode id="3F9B05B6FF996042FF2BF8A9FE85CA69" box="[249,330,1844,1868]" httpUri="http://grbio.org/cool/7159-6y1v" name="Tiroler Landesmuseum Ferdinandeum" pageId="2" pageNumber="62">TLMF</collectionCode>
); 6 3, 6 Ƥ, ditto, but
<date id="2D34BBB3FF996042FDEAF8A9FD65CA69" box="[568,682,1844,1868]" pageId="2" pageNumber="62" value="2010-11-07">11.7.2010</date>
, leg. Mayr (
<collectionCode id="3F9B05B6FF996042FCE2F8A9FC4ACA69" box="[816,901,1844,1868]" pageId="2" pageNumber="62">RCTM</collectionCode>
; coll. N. Pöll);
<date id="2D34BBB3FF996042FBFCF8A9FB5FCA69" box="[1070,1168,1844,1868]" pageId="2" pageNumber="62" value="1916-10-03">10 3, 16</date>
Ƥ, ditto, but
<date id="2D34BBB3FF996042FAF7F8A9FA58CA69" box="[1317,1431,1844,1868]" pageId="2" pageNumber="62" value="2010-12-07">12.7.2010</date>
, leg. Mayr (
<collectionCode id="3F9B05B6FF996042FEC6F8CAFEA5CA4A" box="[276,362,1879,1903]" pageId="2" pageNumber="62">RCTM</collectionCode>
);
<date id="2D34BBB3FF996042FEAFF8CAFE1DCA4A" box="[381,466,1879,1904]" pageId="2" pageNumber="62" value="1912-02-03">2 3, 12</date>
Ƥ, ditto, but
<date id="2D34BBB3FF996042FDB9F8CAFD12CA4A" box="[619,733,1879,1903]" pageId="2" pageNumber="62" value="2010-07-16">16.7.2010</date>
, leg. Mayr (
<collectionCode id="3F9B05B6FF996042FCB4F8CAFC73CA4A" box="[870,956,1879,1903]" pageId="2" pageNumber="62">RCTM</collectionCode>
); 1 3, 6 Ƥ, ditto, but
<date id="2D34BBB3FF996042FB62F8CAFAEDCA4A" box="[1200,1314,1879,1903]" pageId="2" pageNumber="62" value="2010-07-17">17.7.2010</date>
, leg. Mayr (
<collectionCode id="3F9B05B6FF996042FF4DF8E1FF3ACAB1" box="[159,245,1916,1940]" pageId="2" pageNumber="62">RCTM</collectionCode>
). 1 3, 1 Ƥ, ditto, but
<quantity id="9E723096FF996042FE22F8E1FD87CAB1" box="[496,584,1916,1940]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="62" unit="m" value="1800.0">1800 m</quantity>
,
<date id="2D34BBB3FF996042FD84F8E1FD06CAB1" box="[598,713,1916,1940]" pageId="2" pageNumber="62" value="2004-07-15">15.7.2004</date>
, leg. H. Rietz (
<collectionCode id="3F9B05B6FF996042FCAAF8E1FC78CAB1" box="[888,951,1916,1940]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="2" pageNumber="62">ZSM</collectionCode>
); 1 3, ditto, but
<date id="2D34BBB3FF996042FBAAF8E1FB24CAB1" box="[1144,1259,1916,1940]" pageId="2" pageNumber="62" value="2007-07-15">15.7.2007</date>
, leg. et coll. H. Rietz; 1 3, 2 Ƥ, ditto, but 17.-
<date id="2D34BBB3FF996042FE30F83DFD92CA9D" box="[482,605,1952,1976]" pageId="2" pageNumber="62" value="2004-07-20">20.7.2004</date>
.
</paragraph>
</subSubSection>
<caption id="0DF5CDFBFF986043FF45F90FFE80CBE2" httpUri="https://zenodo.org/record/201285/files/figure.png" pageId="3" pageNumber="63" targetBox="[161,1428,183,1651]" targetPageId="3">
<paragraph id="59359D73FF986043FF45F90FFE80CBE2" blockId="3.[151,1436,1682,1735]" pageId="3" pageNumber="63">
<emphasis id="6BFE4161FF986043FF45F90FFEF3CB8D" bold="true" box="[151,316,1682,1705]" pageId="3" pageNumber="63">FIGURES 1–6.</emphasis>
Adults of
<taxonomicName id="9E8AE6F0FF986043FE7CF90EFE3FCB8D" box="[430,496,1683,1704]" class="Insecta" family="Geometridae" genus="Acasis" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="6BFE4161FF986043FE7CF90EFE3FCB8D" box="[430,496,1683,1704]" italics="true" pageId="3" pageNumber="63">Acasis</emphasis>
</taxonomicName>
. 1
<emphasis id="6BFE4161FF986043FDDEF90EFDD7CB8D" bold="true" box="[524,536,1683,1704]" pageId="3" pageNumber="63">–</emphasis>
2,
<taxonomicName id="9E8AE6F0FF986043FDE6F90EFC76CB8C" authority="Eversmann" authorityName="Eversmann" box="[564,953,1683,1705]" class="Insecta" family="Geometridae" genus="Acasis" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="63" phylum="Arthropoda" rank="subSpecies" species="appensata" subSpecies="appensata">
<emphasis id="6BFE4161FF986043FDE6F90EFCE4CB8C" box="[564,811,1683,1705]" italics="true" pageId="3" pageNumber="63">A. appensata appensata</emphasis>
(Eversmann)
</taxonomicName>
; 3–4,
<taxonomicName id="9E8AE6F0FF986043FC28F90EFB13CB8C" box="[1018,1244,1683,1705]" class="Insecta" family="Geometridae" genus="Acasis" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="63" phylum="Arthropoda" rank="subSpecies" species="appensata" status="subsp. nov." subSpecies="callaina">
<emphasis id="6BFE4161FF986043FC28F90EFB13CB8C" box="[1018,1244,1683,1705]" italics="true" pageId="3" pageNumber="63">A. appensata callaina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70CDFC1AFF986043FB37F90EFA8ECB8D" box="[1253,1345,1683,1704]" pageId="3" pageNumber="63" rank="subSpecies">
<emphasis id="6BFE4161FF986043FB37F90EFA8ECB8D" bold="true" box="[1253,1345,1683,1704]" pageId="3" pageNumber="63">ssp. nov.</emphasis>
</taxonomicNameLabel>
; 5
<emphasis id="6BFE4161FF986043FA8CF90EFAA4CB8D" bold="true" box="[1374,1387,1683,1704]" pageId="3" pageNumber="63">–</emphasis>
6,
<taxonomicName id="9E8AE6F0FF986043FA54F90EFE84CBE2" authority="Hubner" authorityName="Hubner" class="Insecta" family="Geometridae" genus="Acasis" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="63" phylum="Arthropoda" rank="species" species="viretata">
<emphasis id="6BFE4161FF986043FA54F90EFF2ACBE2" italics="true" pageId="3" pageNumber="63">A. viretata</emphasis>
(Hübner)
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="1190CEF8FF986046FF17F969FE95C8A3" lastPageId="6" lastPageNumber="66" pageId="3" pageNumber="63" type="description">
<paragraph id="59359D73FF986046FF17F969FE58C9D3" blockId="3.[151,1437,1780,2021]" lastBlockId="6.[151,1436,1138,1522]" lastPageId="6" lastPageNumber="66" pageId="3" pageNumber="63">
<emphasis id="6BFE4161FF986043FF17F969FE98CA28" bold="true" box="[197,343,1780,1805]" pageId="3" pageNumber="63">Description.</emphasis>
Adult (
<figureCitation id="C1B181F6FF986043FE64F969FDE6CA29" box="[438,553,1780,1804]" captionStart="FIGURES 1 – 6" captionStartId="3.[151,263,1682,1705]" captionTargetBox="[161,1428,183,1651]" captionTargetId="figure@3.[151,1436,165,1661]" captionTargetPageId="3" captionText="FIGURES 1 – 6. Adults of Acasis. 1 – 2, A. appensata appensata (Eversmann); 3 – 4, A. appensata callaina ssp. nov.; 5 – 6, A. viretata (Hübner)." httpUri="https://zenodo.org/record/201285/files/figure.png" pageId="3" pageNumber="63">Figs. 3–4</figureCitation>
). Head, thorax and tegula mixed grayish-brown, labial palpus dark brown. Wingspan
<date id="2D34BBB3FF986043FEC2F885FEBCCA15" box="[272,371,1816,1840]" pageId="3" pageNumber="63" value="1920-03-21">
3
<quantity id="9E723096FF986043FEFFF885FEBCCA15" box="[301,371,1816,1840]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.1" metricValueMin="2.0" pageId="3" pageNumber="63" unit="mm" value="20.5" valueMax="21.0" valueMin="20.0">20–21</quantity>
</date>
mm; Ƥ
<quantity id="9E723096FF986043FE07F885FD9DCA15" box="[469,594,1816,1840]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="3" pageNumber="63" unit="mm" value="21.5" valueMax="23.0" valueMin="20.0">20–23 mm</quantity>
; forewing with strongly convex outer margin; ground colour light greyish covered with pale green, darker grey suffusion particularly along undulated antemedian and postmedian fascia; postmedian fascia light greyish with indistinct darker line; veins with short dark brown lines along both sides of postmedian fascia; dark brown discal spot and dark brown patches along fringe line well developed; fringes chequered greyish-brown, outer part lighter. Hindwing grey-brown with slightly darker basal half; small discal spot; darker grey terminal line and weakly checkered fringes; base of anal area in male with short lobe. Male genitalia (
<figureCitation id="C1B181F6FF9F6044FF4DFF0AFED3CD8A" box="[159,284,151,175]" captionStart="FIGURES 7 – 14" captionStartId="4.[151,263,1955,1978]" captionTargetBox="[152,1362,269,1954]" captionTargetId="figure@4.[151,1436,269,1968]" captionTargetPageId="4" captionText="FIGURES 7 – 14. Male genitalia of Acasis. 7 – 8, A. appensata appensata (Eversmann), slide G 1460; 9 – 10, A. appensata callaina ssp. nov., paratype, slide G 1464; 11 – 12, ditto, paratype, slide G 1461; 13 – 14, A. viretata (Hübner), slide G 1462." httpUri="https://zenodo.org/record/201286/files/figure.png" pageId="4" pageNumber="64">Figs. 9–12</figureCitation>
). As described under the nominotypical subspecies, but sacculus slightly more tapered towards apex. Female genitalia (
<figureCitation id="C1B181F6FF9F6044FEB2FF21FE26CDF1" box="[352,489,188,212]" captionStart="FIGURES 15 – 18" captionStartId="5.[151,263,1966,1989]" captionTargetBox="[292,1308,236,1882]" captionTargetId="figure@5.[224,1358,193,1931]" captionTargetPageId="5" captionText="FIGURES 15 – 18. Female genitalia of Acasis. 15, A. appensata appensata (Eversmann), slide G 1459; 16, A. appensata callaina ssp. nov., paratype, slide G 1456; 17, ditto, paratype, slide G 1458; 18, A. viretata (Hübner), slide G 1463." httpUri="https://zenodo.org/record/201287/files/figure.png" pageId="4" pageNumber="64">Figs. 16–17</figureCitation>
). As described under the nominotypical subspecies, but antrum (colliculum) shorter, length approx.
<quantity id="9E723096FF9F6044FEEDFF42FE59CDD2" box="[319,406,223,247]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="64" unit="mm" value="0.3">0.3 mm</quantity>
, posteriorly more clearly funnel-shaped (with deeper posterior excavation), posterior part of
<emphasis id="6BFE4161FF9D6046FF17FBEFFEF0C9AE" bold="true" box="[197,319,1138,1163]" pageId="6" pageNumber="66">Diagnosis.</emphasis>
This new subspecies is characterized by the pale green suffusion of the forewings which is greyishbrown in the nominotypical subspecies (
<figureCitation id="C1B181F6FF9D6046FDB2FB0BFD04C98B" box="[608,715,1174,1198]" captionStart="FIGURES 1 – 6" captionStartId="3.[151,263,1682,1705]" captionTargetBox="[161,1428,183,1651]" captionTargetId="figure@3.[151,1436,165,1661]" captionTargetPageId="3" captionText="FIGURES 1 – 6. Adults of Acasis. 1 – 2, A. appensata appensata (Eversmann); 3 – 4, A. appensata callaina ssp. nov.; 5 – 6, A. viretata (Hübner)." httpUri="https://zenodo.org/record/201285/files/figure.png" pageId="6" pageNumber="66">Figs. 1–2</figureCitation>
) and yellowish-green in
<taxonomicName id="9E8AE6F0FF9D6046FC34FB0BFB92C98B" box="[998,1117,1174,1198]" class="Insecta" family="Geometridae" genus="Acasis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="66" phylum="Arthropoda" rank="species" species="viretata">
<emphasis id="6BFE4161FF9D6046FC34FB0BFB92C98B" box="[998,1117,1174,1198]" italics="true" pageId="6" pageNumber="66">A. viretata</emphasis>
</taxonomicName>
(
<figureCitation id="C1B181F6FF9D6046FBBFFB0BFB1BC98B" box="[1133,1236,1174,1198]" captionStart="FIGURES 1 – 6" captionStartId="3.[151,263,1682,1705]" captionTargetBox="[161,1428,183,1651]" captionTargetId="figure@3.[151,1436,165,1661]" captionTargetPageId="3" captionText="FIGURES 1 – 6. Adults of Acasis. 1 – 2, A. appensata appensata (Eversmann); 3 – 4, A. appensata callaina ssp. nov.; 5 – 6, A. viretata (Hübner)." httpUri="https://zenodo.org/record/201285/files/figure.png" pageId="6" pageNumber="66">Figs 5–6</figureCitation>
). The genitalia of both sexes are very similar to those of
<taxonomicName id="9E8AE6F0FF9D6046FD8AFB27FCA7C9F7" box="[600,872,1210,1234]" class="Insecta" family="Geometridae" genus="Acasis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="66" phylum="Arthropoda" rank="subSpecies" species="appensata" subSpecies="appensata">
<emphasis id="6BFE4161FF9D6046FD8AFB27FCA7C9F7" box="[600,872,1210,1234]" italics="true" pageId="6" pageNumber="66">A. appensata appensata</emphasis>
</taxonomicName>
but strongly differ from those of
<taxonomicName id="9E8AE6F0FF9D6046FB3CFB27FAA9C9F7" box="[1262,1382,1210,1234]" class="Insecta" family="Geometridae" genus="Acasis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="66" phylum="Arthropoda" rank="species" species="viretata">
<emphasis id="6BFE4161FF9D6046FB3CFB27FAA9C9F7" box="[1262,1382,1210,1234]" italics="true" pageId="6" pageNumber="66">A. viretata</emphasis>
</taxonomicName>
(see above and
<figureCitation id="C1B181F6FF9D6046FEC2FB43FE45C9D3" box="[272,394,1246,1270]" captionStart-0="FIGURES 7 – 14" captionStart-1="FIGURES 15 – 18" captionStartId-0="4.[151,263,1955,1978]" captionStartId-1="5.[151,263,1966,1989]" captionTargetBox-0="[152,1362,269,1954]" captionTargetBox-1="[292,1308,236,1882]" captionTargetId-0="figure@4.[151,1436,269,1968]" captionTargetId-1="figure@5.[224,1358,193,1931]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 7 – 14. Male genitalia of Acasis. 7 – 8, A. appensata appensata (Eversmann), slide G 1460; 9 – 10, A. appensata callaina ssp. nov., paratype, slide G 1464; 11 – 12, ditto, paratype, slide G 1461; 13 – 14, A. viretata (Hübner), slide G 1462." captionText-1="FIGURES 15 – 18. Female genitalia of Acasis. 15, A. appensata appensata (Eversmann), slide G 1459; 16, A. appensata callaina ssp. nov., paratype, slide G 1456; 17, ditto, paratype, slide G 1458; 18, A. viretata (Hübner), slide G 1463." httpUri-0="https://zenodo.org/record/201286/files/figure.png" httpUri-1="https://zenodo.org/record/201287/files/figure.png" pageId="6" pageNumber="66">Figs. 7–18</figureCitation>
).
</paragraph>
<caption id="0DF5CDFBFF9F6044FF45F83EFA9DCAFD" httpUri="https://zenodo.org/record/201286/files/figure.png" pageId="4" pageNumber="64" targetBox="[152,1362,269,1954]" targetPageId="4">
<paragraph id="59359D73FF9F6044FF45F83EFA9DCAFD" blockId="4.[151,1435,1955,2008]" pageId="4" pageNumber="64">
<emphasis id="6BFE4161FF9F6044FF45F83EFE88CA9C" bold="true" box="[151,327,1955,1978]" pageId="4" pageNumber="64">FIGURES 7–14.</emphasis>
Male genitalia of
<taxonomicName id="9E8AE6F0FF9F6044FDD7F839FD88CA9C" box="[517,583,1956,1977]" class="Insecta" family="Geometridae" genus="Acasis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="6BFE4161FF9F6044FDD7F839FD88CA9C" box="[517,583,1956,1977]" italics="true" pageId="4" pageNumber="64">Acasis</emphasis>
</taxonomicName>
. 7
<emphasis id="6BFE4161FF9F6044FDB0F839FDA0CA9C" bold="true" box="[610,623,1956,1977]" pageId="4" pageNumber="64">–</emphasis>
8,
<taxonomicName id="9E8AE6F0FF9F6044FD5BF839FBC5CA9F" authority="Eversmann" authorityName="Eversmann" box="[649,1034,1956,1978]" class="Insecta" family="Geometridae" genus="Acasis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="64" phylum="Arthropoda" rank="subSpecies" species="appensata" subSpecies="appensata">
<emphasis id="6BFE4161FF9F6044FD5BF839FCB2CA9F" box="[649,893,1956,1978]" italics="true" pageId="4" pageNumber="64">A. appensata appensata</emphasis>
(Eversmann)
</taxonomicName>
, slide G 1460; 9
<emphasis id="6BFE4161FF9F6044FB66F839FB0FCA9C" bold="true" box="[1204,1216,1956,1977]" pageId="4" pageNumber="64">–</emphasis>
10,
<taxonomicName id="9E8AE6F0FF9F6044FB3AF839FF04CAFD" class="Insecta" family="Geometridae" genus="Acasis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="64" phylum="Arthropoda" rank="subSpecies" species="appensata" status="subsp. nov." subSpecies="callaina">
<emphasis id="6BFE4161FF9F6044FB3AF839FF04CAFD" italics="true" pageId="4" pageNumber="64">A. appensata callaina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70CDFC1AFF9F6044FF03F85EFEE4CAFD" box="[209,299,1987,2008]" pageId="4" pageNumber="64" rank="subSpecies">
<emphasis id="6BFE4161FF9F6044FF03F85EFEE4CAFD" bold="true" box="[209,299,1987,2008]" pageId="4" pageNumber="64">ssp. nov.</emphasis>
</taxonomicNameLabel>
, paratype, slide G 1464; 11
<emphasis id="6BFE4161FF9F6044FD96F85EFD9FCAFD" bold="true" box="[580,592,1987,2008]" pageId="4" pageNumber="64">–</emphasis>
12,
<emphasis id="6BFE4161FF9F6044FDA4F85FFD64CAFD" box="[630,683,1986,2008]" italics="true" pageId="4" pageNumber="64">ditto,</emphasis>
paratype, slide G 1461; 13
<emphasis id="6BFE4161FF9F6044FC6FF85EFC05CAFD" bold="true" box="[957,970,1987,2008]" pageId="4" pageNumber="64">–</emphasis>
14,
<taxonomicName id="9E8AE6F0FF9F6044FC22F85EFB70CAFD" authority="Hubner" authorityName="Hubner" box="[1008,1215,1986,2008]" class="Insecta" family="Geometridae" genus="Acasis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="64" phylum="Arthropoda" rank="species" species="viretata">
<emphasis id="6BFE4161FF9F6044FC22F85EFB95CAFD" box="[1008,1114,1987,2008]" italics="true" pageId="4" pageNumber="64">A. viretata</emphasis>
(Hübner)
</taxonomicName>
, slide G 1462.
</paragraph>
</caption>
<caption id="0DF5CDFBFF9E6045FF45F833FAC8CAC6" httpUri="https://zenodo.org/record/201287/files/figure.png" pageId="5" pageNumber="65" targetBox="[292,1308,236,1882]" targetPageId="5">
<paragraph id="59359D73FF9E6045FF45F833FAC8CAC6" blockId="5.[151,1436,1966,2019]" pageId="5" pageNumber="65">
<emphasis id="6BFE4161FF9E6045FF45F833FE9ACAE1" bold="true" box="[151,341,1966,1989]" pageId="5" pageNumber="65">FIGURES 15–18.</emphasis>
Female genitalia of
<taxonomicName id="9E8AE6F0FF9E6045FDFBF832FDA4CAE1" box="[553,619,1967,1988]" class="Insecta" family="Geometridae" genus="Acasis" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="6BFE4161FF9E6045FDFBF832FDA4CAE1" box="[553,619,1967,1988]" italics="true" pageId="5" pageNumber="65">Acasis</emphasis>
</taxonomicName>
. 15,
<taxonomicName id="9E8AE6F0FF9E6045FD72F832FBEDCAE0" authority="Eversmann" authorityName="Eversmann" box="[672,1058,1967,1989]" class="Insecta" family="Geometridae" genus="Acasis" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="65" phylum="Arthropoda" rank="subSpecies" species="appensata" subSpecies="appensata">
<emphasis id="6BFE4161FF9E6045FD72F832FC5ACAE0" box="[672,917,1967,1989]" italics="true" pageId="5" pageNumber="65">A. appensata appensata</emphasis>
(Eversmann)
</taxonomicName>
, slide G 1459; 16,
<taxonomicName id="9E8AE6F0FF9E6045FB35F832FF04CAC6" class="Insecta" family="Geometridae" genus="Acasis" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="65" phylum="Arthropoda" rank="subSpecies" species="appensata" status="subsp. nov." subSpecies="callaina">
<emphasis id="6BFE4161FF9E6045FB35F832FF04CAC6" italics="true" pageId="5" pageNumber="65">A. appensata callaina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70CDFC1AFF9E6045FF03F850FEE4CAC7" box="[209,299,1997,2018]" pageId="5" pageNumber="65" rank="subSpecies">
<emphasis id="6BFE4161FF9E6045FF03F850FEE4CAC7" bold="true" box="[209,299,1997,2018]" pageId="5" pageNumber="65">ssp. nov.</emphasis>
</taxonomicNameLabel>
, paratype, slide G 1456; 17,
<emphasis id="6BFE4161FF9E6045FD83F850FD49CAC6" box="[593,646,1997,2019]" italics="true" pageId="5" pageNumber="65">ditto,</emphasis>
paratype, slide G 1458; 18,
<taxonomicName id="9E8AE6F0FF9E6045FC76F853FBBBCAC6" authority="Hubner" authorityName="Hubner" box="[932,1140,1997,2019]" class="Insecta" family="Geometridae" genus="Acasis" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="65" phylum="Arthropoda" rank="species" species="viretata">
<emphasis id="6BFE4161FF9E6045FC76F853FBC0CAC6" box="[932,1039,1998,2019]" italics="true" pageId="5" pageNumber="65">A. viretata</emphasis>
(Hübner)
</taxonomicName>
, slide G 1463.
</paragraph>
</caption>
<caption id="0DF5CDFBFF9D6046FF45FBB2FBB8C960" box="[151,1143,1071,1094]" httpUri="https://zenodo.org/record/201288/files/figure.png" pageId="6" pageNumber="66" targetBox="[151,1436,193,1049]" targetPageId="6">
<paragraph id="59359D73FF9D6046FF45FBB2FBB8C960" blockId="6.[151,1143,1071,1094]" box="[151,1143,1071,1094]" pageId="6" pageNumber="66">
<emphasis id="6BFE4161FF9D6046FF45FBB2FED0C960" bold="true" box="[151,287,1071,1094]" pageId="6" pageNumber="66">FIGURE 19.</emphasis>
Type-locality of
<taxonomicName id="9E8AE6F0FF9D6046FE1FFBADFD64C960" box="[461,683,1071,1094]" class="Insecta" family="Geometridae" genus="Acasis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="66" phylum="Arthropoda" rank="subSpecies" species="appensata" status="subsp. nov." subSpecies="callaina">
<emphasis id="6BFE4161FF9D6046FE1FFBADFD64C960" box="[461,683,1071,1094]" italics="true" pageId="6" pageNumber="66">A. appensata callaina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70CDFC1AFF9D6046FD63FBADFCC4C960" box="[689,779,1072,1093]" pageId="6" pageNumber="66" rank="subSpecies">
<emphasis id="6BFE4161FF9D6046FD63FBADFCC4C960" bold="true" box="[689,779,1072,1093]" pageId="6" pageNumber="66">ssp. nov.</emphasis>
</taxonomicNameLabel>
(Monte Terminillo, Abruzzi, Italy).
</paragraph>
</caption>
<paragraph id="59359D73FF9D6046FF17FA9FFE95C8A3" blockId="6.[151,1436,1138,1522]" pageId="6" pageNumber="66">
<emphasis id="6BFE4161FF9D6046FF17FA9FFE87C83E" bold="true" box="[197,328,1282,1307]" pageId="6" pageNumber="66">Bionomics.</emphasis>
Host-plant and early stages are unknown but it is very likely that the new subspecies also feeds on
<taxonomicName id="9E8AE6F0FF9D6046FF45FABBFEF2C81B" box="[151,317,1318,1342]" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="66" phylum="Arthropoda" rank="species" species="spicata">
<emphasis id="6BFE4161FF9D6046FF45FABBFEF2C81B" box="[151,317,1318,1342]" italics="true" pageId="6" pageNumber="66">Actaea spicata</emphasis>
</taxonomicName>
which occurs at the type-locality. The adults have been collected in mid-July nearby the upper edge of a beech forest on limestone at high altitudes of about
<quantity id="9E723096FF9D6046FCD5FAD7FCADC847" box="[775,866,1354,1378]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="66" unit="m" value="1700.0">1700 m</quantity>
.s.l. (
<figureCitation id="C1B181F6FF9D6046FC43FAD7FC2BC847" box="[913,996,1354,1378]" captionStart="FIGURE 19" captionStartId="6.[151,249,1071,1094]" captionTargetBox="[151,1436,193,1049]" captionTargetId="figure@6.[151,1436,193,1050]" captionTargetPageId="6" captionText="FIGURE 19. Type-locality of A. appensata callaina ssp. nov. (Monte Terminillo, Abruzzi, Italy)." httpUri="https://zenodo.org/record/201288/files/figure.png" pageId="6" pageNumber="66">Fig. 19</figureCitation>
). They were attracted to artificial lights in large numbers.
</paragraph>
</subSubSection>
<subSubSection id="1190CEF8FF9D6046FF17FA0FFA83C88F" box="[197,1356,1426,1451]" pageId="6" pageNumber="66" type="distribution">
<paragraph id="59359D73FF9D6046FF17FA0FFA83C88F" blockId="6.[151,1436,1138,1522]" box="[197,1356,1426,1451]" pageId="6" pageNumber="66">
<emphasis id="6BFE4161FF9D6046FF17FA0FFE91C88E" bold="true" box="[197,350,1426,1451]" pageId="6" pageNumber="66">Distribution.</emphasis>
Only known from the northern slopes of Monte Terminillo in the Central Italian Abruzzi.
</paragraph>
</subSubSection>
<subSubSection id="1190CEF8FF9D6046FF17FA2BFEE2C8D7" pageId="6" pageNumber="66" type="etymology">
<paragraph id="59359D73FF9D6046FF17FA2BFEE2C8D7" blockId="6.[151,1436,1138,1522]" pageId="6" pageNumber="66">
<emphasis id="6BFE4161FF9D6046FF17FA2BFE84C8EB" bold="true" box="[197,331,1462,1486]" pageId="6" pageNumber="66">Etymology.</emphasis>
The specific name, an adjective derived from the Latin
<emphasis id="6BFE4161FF9D6046FC6EFA2BFBECC8EB" box="[956,1059,1462,1486]" italics="true" pageId="6" pageNumber="66">callainus</emphasis>
, refers to the pale green colour of the forewing.
</paragraph>
</subSubSection>
</treatment>
</document>
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