372 lines
47 KiB
XML
372 lines
47 KiB
XML
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<mods:title id="283009003F2B938F28925D28590050DA">Taxonomic review of the genus Calyptopsis of Turkey and adjacent areas (Coleoptera: Tenebrionidae: Tentyriini)</mods:title>
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<mods:namePart id="F32BAA40DA647B2C4A0E5516305938E6">Chigray, Svetlana</mods:namePart>
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<mods:namePart id="BA1B0F61E67512F3A6B0C08D1D2FE721">Nabozhenko, Maxim</mods:namePart>
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<mods:affiliation id="E3993C50890AFB090152BDE70A71160C">St. Petersburg 199034 Russia; e-mail: s. voloboeva 95 @ mail. ru & Dagestan State University, M. Gadzhiev str., 43 a, Makhachkala, Republic of Dagestan 367000 Russia</mods:affiliation>
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<heading id="4550FC60FF97891971C8F90AFB52F8F8" box="[995,1269,1779,1803]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="336" reason="3">
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<taxonomicName id="D9A7308FFF97891971C8F90AFB3EF8F8" authority="Chigray & Nabozhenko & Keskin & Abdurakhmanov, 2018" authorityName="Chigray & Nabozhenko & Keskin & Abdurakhmanov" authorityYear="2018" box="[995,1177,1779,1803]" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="ottoi" status="sp. nov.">
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<emphasis id="2CD3971EFF97891971C8F90AFB3EF8F8" bold="true" box="[995,1177,1779,1803]" italics="true" pageId="5" pageNumber="336">Calyptopsis ottoi</emphasis>
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(
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C–G, 9)
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locality.
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,
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, Görentaş,
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(
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: ♁, ‘
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/
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/ leg.
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, B.
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. Pektaş’ (
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<collectionCode id="78B6D3C9FF97891976ACF85FFB17F84A" box="[1159,1200,1958,1977]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="336" type="Museum">ZIN</collectionCode>
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)
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.
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:
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:
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, same data as
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(
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);
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, ‘
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<collectingCountry id="66B00B9CFF9789197688F838FB43F827" box="[1187,1252,1985,2004]" name="Turkey" pageId="5" pageNumber="336">Turkey</collectingCountry>
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/
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/ Berçelan Yayla
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,
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/ leg. M.V. and S.V. Nabozhenko, B. Keskin, A. Pektaş’ (
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|
||
<specimenCount id="08A18085FF978919770FF80FFAE4F7F9" box="[1316,1347,2035,2058]" pageId="5" pageNumber="336" type="female">1 ♀</specimenCount>
|
||
, ‘
|
||
<collectingCountry id="66B00B9CFF978919777BF80FFA36F7FA" box="[1360,1425,2038,2057]" name="Turkey" pageId="5" pageNumber="336">Turkey</collectingCountry>
|
||
/
|
||
<collectingRegion id="DC6385EEFF9789197117F7E8FC08F7D7" box="[828,943,2065,2084]" country="Turkey" name="Van" pageId="5" pageNumber="336">Van Province</collectingRegion>
|
||
/ Erekdağ
|
||
<quantity id="D95FE6E9FF9789197625F7E8FB22F7D7" box="[1038,1157,2065,2084]" metricMagnitude="3" metricUnit="m" metricValue="2.3385" metricValueMax="2.6" metricValueMin="2.077" pageId="5" pageNumber="336" unit="m" value="2338.5" valueMax="2600.0" valueMin="2077.0">2077–2600 m</quantity>
|
||
/
|
||
<geoCoordinate id="7B932DCBFF97891976BDF7E8FAB7F7D7" box="[1174,1296,2065,2084]" degrees="38" direction="north" minutes="25" orientation="latitude" pageId="5" pageNumber="336" precision="1" seconds="53.1" value="38.43142">38°25’53.1”N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="7B932DCBFF9789197731F7E8FA37F7D7" box="[1306,1424,2065,2084]" degrees="43" direction="east" minutes="29" orientation="longitude" pageId="5" pageNumber="336" precision="1" seconds="03.6" value="43.484333">43°29’03.6”E</geoCoordinate>
|
||
/
|
||
<date id="6A196DCCFF94891A72BCF8C6FF41F8A1" box="[151,230,1855,1874]" pageId="6" pageNumber="337" value="2013-06-01">1.vi.2013</date>
|
||
/ leg.M.V.and S.V. Nabozhenko, B. Keskin,A.Pektaş’ (
|
||
<collectionCode id="78B6D3C9FF94891A7098F8C6FD57F8A1" box="[691,752,1855,1874]" country="Turkey" httpUri="http://grbio.org/cool/s077-eqrn" name="Zoology Department, Ege University" pageId="6" pageNumber="337">ZDEU</collectionCode>
|
||
).
|
||
<emphasis id="2CD3971EFF94891A72BCF8A0FF71F89F" bold="true" box="[151,214,1881,1900]" pageId="6" pageNumber="337">
|
||
<collectingCountry id="66B00B9CFF94891A72BCF8A0FF75F89F" box="[151,210,1881,1900]" name="Iraq" pageId="6" pageNumber="337">IRAQ</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="08A18085FF94891A72F6F8A0FF58F89E" box="[221,255,1878,1901]" pageId="6" pageNumber="337" type="female">1 ♀</specimenCount>
|
||
, ‘
|
||
<collectingCountry id="66B00B9CFF94891A7324F8A0FE90F89F" box="[271,311,1881,1900]" name="Iraq" pageId="6" pageNumber="337">Iraq</collectingCountry>
|
||
, Hasarost /
|
||
<date id="6A196DCCFF94891A738AF8A0FDA5F89F" box="[417,514,1881,1900]" pageId="6" pageNumber="337" value="1971-07-13">13.vii.1971</date>
|
||
/ M.S. Abdulxassa [unclear name of collector]’ (
|
||
<collectionCode id="78B6D3C9FF94891A736FF88DFE2DF874" box="[324,394,1908,1927]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="6" pageNumber="337">HNHM</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="4AD81B84FF94891A72BCF942FB87F8F0" ID-DOI="http://doi.org/10.5281/zenodo.4504778" ID-Zenodo-Dep="4504778" httpUri="https://zenodo.org/record/4504778/files/figure.png" pageId="6" pageNumber="337" startId="6.[151,182,1723,1742]" targetBox="[265,1329,189,1703]" targetPageId="6">
|
||
<paragraph id="1E184B0CFF94891A72BCF942FB87F8F0" blockId="6.[151,1437,1719,1795]" pageId="6" pageNumber="337">
|
||
Fig. 3.
|
||
<taxonomicName id="D9A7308FFF94891A72FDF942FE9CF93D" authorityName="Solier" authorityYear="1835" box="[214,315,1723,1742]" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="337" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2CD3971EFF94891A72FDF942FE9CF93D" box="[214,315,1723,1742]" italics="true" pageId="6" pageNumber="337">Calyptopsis</emphasis>
|
||
</taxonomicName>
|
||
spp., habitus and details of structure. A, B, C –
|
||
<taxonomicName id="D9A7308FFF94891A70CBF942FC6CF93D" authority="Reitter, 1900" authorityName="Reitter" authorityYear="1900" box="[736,971,1723,1742]" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="337" phylum="Arthropoda" rank="species" species="escherichi">
|
||
<emphasis id="2CD3971EFF94891A70CBF942FCF5F93D" box="[736,850,1723,1742]" italics="true" pageId="6" pageNumber="337">C. escherichi</emphasis>
|
||
<bibRefCitation id="7A3636FDFF94891A7172F942FC6CF93D" author="REITTER E." box="[857,971,1723,1742]" pageId="6" pageNumber="337" pagination="82 - 197" refId="ref11458" refString="REITTER E. 1900: Bestimmungs-Tabelle der Tenebrioniden-Abtheilungen: Tentyrini und Adelostomini aus Europa und den angrenzenden Landern. Verhandlungen des Naturforschendes Vereines in Brunn 39: 82 - 197." type="journal article" year="1900">Reitter, 1900</bibRefCitation>
|
||
</taxonomicName>
|
||
, ♀ and labels (Eskişehir, HNHM); D –
|
||
<taxonomicName id="D9A7308FFF94891A770EF942FA30F93D" authorityName="Reitter" authorityYear="1900" box="[1317,1431,1723,1742]" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="337" phylum="Arthropoda" rank="species" species="escherichi">
|
||
<emphasis id="2CD3971EFF94891A770EF942FA30F93D" box="[1317,1431,1723,1742]" italics="true" pageId="6" pageNumber="337">C. escherichi</emphasis>
|
||
</taxonomicName>
|
||
, mesoventrite; E –
|
||
<taxonomicName id="D9A7308FFF94891A731DF92CFE0FF91B" authorityName="Reitter" authorityYear="1900" box="[310,424,1749,1768]" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="337" phylum="Arthropoda" rank="species" species="escherichi">
|
||
<emphasis id="2CD3971EFF94891A731DF92CFE0FF91B" box="[310,424,1749,1768]" italics="true" pageId="6" pageNumber="337">C. escherichi</emphasis>
|
||
</taxonomicName>
|
||
, ♁ (Niğde: Maden köy, ZIN); F – the same, ♀; G –
|
||
<taxonomicName id="D9A7308FFF94891A715FF92CFBC4F91B" authority="Reitter, 1897" authorityName="Reitter" authorityYear="1897" box="[884,1123,1749,1768]" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="337" phylum="Arthropoda" rank="species" species="lineimargo">
|
||
<emphasis id="2CD3971EFF94891A715FF92CFC4CF91B" box="[884,1003,1749,1768]" italics="true" pageId="6" pageNumber="337">C. lineimargo</emphasis>
|
||
<bibRefCitation id="7A3636FDFF94891A71DAF92CFBC4F91B" author="REITTER E." box="[1009,1123,1749,1768]" pageId="6" pageNumber="337" pagination="305 - 311" refId="ref11436" refString="REITTER E. 1897: Uebersicht der Arten der Coleopteren-Gattung Calyptopsis Sol. Deutsche Entomologische Zeitschrift 1896: 305 - 311." type="journal article" year="1897">Reitter, 1897</bibRefCitation>
|
||
</taxonomicName>
|
||
, lectotype and labels; H –
|
||
<taxonomicName id="D9A7308FFF94891A7767F92CFEF9F8F0" authority="Reiche & Saulcy, 1857" authorityName="Reiche & Saulcy" authorityYear="1857" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="337" phylum="Arthropoda" rank="species" species="solieri">
|
||
<emphasis id="2CD3971EFF94891A7767F92CFA3AF91B" box="[1356,1437,1749,1768]" italics="true" pageId="6" pageNumber="337">C. solieri</emphasis>
|
||
<bibRefCitation id="7A3636FDFF94891A72BCF909FEF9F8F0" author="REICHE L. J. & SAULCY F." box="[151,350,1776,1795]" pageId="6" pageNumber="337" pagination="169 - 276" refId="ref11344" refString="REICHE L. J. & SAULCY F. 1857: Especes nouvelles ou peu connues de coleopteres, recueillis par M. F. de Saulcy, membre de l'lnstitut, dans son voyage en Orient. Annales de la Societe Entomologique de France, Troisieme Serie 5: 169 - 276 + pl. 5." type="journal article" year="1857">Reiche & Saulcy, 1857</bibRefCitation>
|
||
</taxonomicName>
|
||
, ♁ (Şanlıurfa Province, Birecik); I –
|
||
<taxonomicName id="D9A7308FFF94891A70B1F909FD4DF8F0" authorityName="Reiche & Saulcy" authorityYear="1857" box="[666,746,1776,1795]" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="337" phylum="Arthropoda" rank="species" species="solieri">
|
||
<emphasis id="2CD3971EFF94891A70B1F909FD4DF8F0" box="[666,746,1776,1795]" italics="true" pageId="6" pageNumber="337">C. solieri</emphasis>
|
||
</taxonomicName>
|
||
, ♁ (Gaziantep Province, Şahinbey).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="56BD1887FF94891B72BCF859FBA2F832" lastPageId="7" lastPageNumber="338" pageId="6" pageNumber="337" type="description">
|
||
<paragraph id="1E184B0CFF94891B72BCF859FD05FA31" blockId="6.[151,759,1951,2071]" lastBlockId="7.[151,760,1067,2074]" lastPageId="7" lastPageNumber="338" pageId="6" pageNumber="337">
|
||
<emphasis id="2CD3971EFF94891A72BCF859FED1F844" bold="true" box="[151,374,1951,1975]" pageId="6" pageNumber="337">
|
||
Description.
|
||
<emphasis id="2CD3971EFF94891A731EF866FED1F844" bold="true" box="[309,374,1951,1975]" italics="true" pageId="6" pageNumber="337">Male.</emphasis>
|
||
</emphasis>
|
||
Body slender, with weak greasy shine. Apex of frontoclypeus (frontoclypeal tooth) acute, vertically projected. Anterior margin of frontoclypeus separated from the other surface of head by wide transverse protuberance, which is separated from the other surface of head by deep transverse depression. Lateral margin of head irregularly weakly rounded, sinuated. Genae weakly emarginate. Supraocular keels extended almost from level of basal margin of eye. Head surface with wide transverse depression on each side between frontoclypeus and anterior margin of supraocular keels. Head widest at eye level. Ratio of head width at eye level to distance between eyes 1.5. Antennae short, with apical antennomere reaching basal third of pronotum. Ratio of length (width) of antennomeres II–XI: 0.6(0.5): 0.6(0.4): 0.8(0.5): 0.7(0.4): 0.6(0.4): 0.6(0.4): 0.7(0.5): 0.5(0.5): 0.5(0.5): 0.3(0.4). Punctation of head moderately coarse and sparse, puncture diameter 1.5–2.0× as wide as distance among punctures. Vertex with deep transverse depression, shiny, with coarse and dense rasp-shaped punctation (puncture diameter 1.5–2.0× as wide as distance among punctures). Mentum weakly shiny, moderately coarsely and densely punctate by rasp-shaped punctures (puncture diameter 1.5–2.0× as wide as distance among punctures).
|
||
</paragraph>
|
||
<caption id="4AD81B84FF95891B72BCFC5BFE2EFC23" ID-DOI="http://doi.org/10.5281/zenodo.4504780" ID-Zenodo-Dep="4504780" httpUri="https://zenodo.org/record/4504780/files/figure.png" pageId="7" pageNumber="338" startId="7.[151,182,930,949]" targetBox="[189,1399,193,912]" targetPageId="7">
|
||
<paragraph id="1E184B0CFF95891B72BCFC5BFE2EFC23" blockId="7.[151,1437,924,976]" pageId="7" pageNumber="338">
|
||
Fig. 4.
|
||
<taxonomicName id="D9A7308FFF95891B72FFFC5BFE9EFC46" authorityName="Solier" authorityYear="1835" box="[212,313,930,949]" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="338" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2CD3971EFF95891B72FFFC5BFE9EFC46" box="[212,313,930,949]" italics="true" pageId="7" pageNumber="338">Calyptopsis</emphasis>
|
||
</taxonomicName>
|
||
spp., habitus, pronotum. A –
|
||
<taxonomicName id="D9A7308FFF95891B701EFC5BFCE8FC46" authority="(Faldermann, 1837)" baseAuthorityName="Faldermann" baseAuthorityYear="1837" box="[565,847,930,949]" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="338" phylum="Arthropoda" rank="species" species="pulchella">
|
||
<emphasis id="2CD3971EFF95891B701EFC5BFD3AFC46" box="[565,669,930,949]" italics="true" pageId="7" pageNumber="338">C. pulchella</emphasis>
|
||
(
|
||
<bibRefCitation id="7A3636FDFF95891B7082FC5BFCEFFC46" author="FALDERMANN F." box="[681,840,930,949]" pageId="7" pageNumber="338" refId="ref10602" refString="FALDERMANN F. 1837: Fauna Entomologica Trans-Caucasica. Coleoptera. Pars II. Auguste Semen, Moscou, 433 pp., 15 pls." type="book" year="1837">Faldermann, 1837</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, ♁; B –
|
||
<taxonomicName id="D9A7308FFF95891B71BCFC5BFC58FC46" baseAuthorityName="Faldermann" baseAuthorityYear="1837" box="[919,1023,930,949]" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="338" phylum="Arthropoda" rank="species" species="pulchella">
|
||
<emphasis id="2CD3971EFF95891B71BCFC5BFC58FC46" box="[919,1023,930,949]" italics="true" pageId="7" pageNumber="338">C. pulchella</emphasis>
|
||
</taxonomicName>
|
||
, ♀; C –
|
||
<taxonomicName id="D9A7308FFF95891B7662FC5BFA9CFC46" authority="Reitter, 1889" authorityName=": Reitter" authorityYear="1889" box="[1097,1339,930,949]" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="338" phylum="Arthropoda" rank="species" species="emarginata">
|
||
<emphasis id="2CD3971EFF95891B7662FC5BFB62FC46" box="[1097,1221,930,949]" italics="true" pageId="7" pageNumber="338">C. emarginata</emphasis>
|
||
<bibRefCitation id="7A3636FDFF95891B76E1FC5BFA9CFC46" author="REITTER E." box="[1226,1339,930,949]" pageId="7" pageNumber="338" pagination="17 - 44" refId="ref11402" refString="REITTER E. 1889: Neue Coleopteren aus Europa, den angrenzenden Landern und Sibirien, mit Bemerkungen uber bekannte Arten. Sechster Theil. Deutsche Entomologische Zeitschrift 1889: 17 - 44." type="journal article" year="1889">Reitter, 1889</bibRefCitation>
|
||
</taxonomicName>
|
||
, pronotum; D –
|
||
<taxonomicName id="D9A7308FFF95891B7297FC44FE83FC23" baseAuthorityName="Faldermann" baseAuthorityYear="1837" box="[188,292,957,976]" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="338" phylum="Arthropoda" rank="species" species="pulchella">
|
||
<emphasis id="2CD3971EFF95891B7297FC44FE83FC23" box="[188,292,957,976]" italics="true" pageId="7" pageNumber="338">C. pulchella</emphasis>
|
||
</taxonomicName>
|
||
, pronotum.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="1E184B0CFF95891B729EFA30FD44F869" blockId="7.[151,760,1067,2074]" pageId="7" pageNumber="338">Pronotum transverse (1.2× as wide as long), widest little before middle, 1.2× as wide as head. Ratio of pronotal width at anterior margin, in middle and at base 5.5: 7.0: 6.5. Anterior margin of pronotum widely emarginate, base bisinuate, weakly rounded in middle. Lateral margins of pronotum weakly rounded, slightly sinuated at base. Anterior angles of pronotum acute, narrowly rounded at apex, basal angles obtuse. All margins of pronotum beaded. Disc of pronotum moderately convex, without depressions in basal part. Disc of pronotum with fine, moderately dense punctation. Prothoracic hypomera completely punctate with moderately coarse, dense rasp-shaped punctures (puncture diameter 1.5–2.0× as wide as distance among punctures). Prosternal process laminar, strongly protruded, laterally acute, completely beaded, with longitudinal depression and very fine, moderately dense punctation.</paragraph>
|
||
<paragraph id="1E184B0CFF95891B729EF85AFC49FB12" blockId="7.[151,760,1067,2074]" lastBlockId="7.[828,1437,1066,2081]" pageId="7" pageNumber="338">Elytra smooth, with only fine microwrinkles, moderately elongate (1.6× as long as wide), slightly convex, 2.1× longer and 1.6× wider than pronotum, 2.0× wider than head. Elytra without striae, with fine, moderately dense punctation. Epipleura with poorly visible transverse wrinkles. Mesoventrite with mainly bisinuate bracket- shaped deep foveae. Metaventrite with poorly visible wrinkles and fine, moderately dense weakly rasp-shaped punctures along mesocoxae, remaining surface of metaventrite smooth.</paragraph>
|
||
<paragraph id="1E184B0CFF95891B7171FB13FA3AFA32" blockId="7.[828,1437,1066,2081]" pageId="7" pageNumber="338">First abdominal ventrite moderately coarsely and densely punctate by large round punctures along basal margin of metacoxae (puncture diameter 1.5–2.0× as wide as distance among punctures), remaining ventrites with smooth punctures on sides and very fine, sparse punctures middle. Only abdominal ventrite 1 completely laterally beaded, ventrite 2 beaded on less than half its surface, ventrites 3–5 not beaded.</paragraph>
|
||
<paragraph id="1E184B0CFF95891B7171FA33FB1EF992" blockId="7.[828,1437,1066,2081]" pageId="7" pageNumber="338">Legs short. Anterior tibiae not depressed along dorsal surface of outer margin. Pro- and mesotibiae straight, metatibiae slightly concavely curved. Ratio of femur/tibia/ tarsus 3.0: 3.5: 3.0 in fore legs, 5.0: 4.0: 3.0 in middle legs, and 6.0: 5.0: 4.0 in hind legs.</paragraph>
|
||
<paragraph id="1E184B0CFF95891B7171F993FB4BF972" blockId="7.[828,1437,1066,2081]" box="[858,1260,1642,1665]" pageId="7" pageNumber="338">
|
||
Body length 12.0 mm, width
|
||
<quantity id="D95FE6E9FF95891B76B8F993FB4FF972" box="[1171,1256,1642,1665]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="7" pageNumber="338" unit="mm" value="4.5">4.5 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="1E184B0CFF95891B7171F970FC28F932" blockId="7.[828,1437,1066,2081]" pageId="7" pageNumber="338">
|
||
<emphasis id="2CD3971EFF95891B7171F970FC14F952" bold="true" box="[858,947,1673,1697]" italics="true" pageId="7" pageNumber="338">Female.</emphasis>
|
||
Body more robust. Elytra 1.3–1.45× as wide as long.
|
||
</paragraph>
|
||
<paragraph id="1E184B0CFF95891B7117F933FBA2F832" blockId="7.[828,1437,1066,2081]" pageId="7" pageNumber="338">
|
||
<emphasis id="2CD3971EFF95891B7117F933FC1DF912" bold="true" box="[828,954,1738,1761]" pageId="7" pageNumber="338">Variability.</emphasis>
|
||
Specimen from Berçelan Yayla differs in finer and sparser punctation of pronotum (puncture diameter 1.5–2.0× as long as distance among punctures; prosternal process moderately protruded (not laminar), rounded, laterally rectangular; elytral punctation very fine and sparse (puncture diameter 3.0–4.0× shorter than distance among punctures). Ovipositor of females in all populations has the same structure.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="56BD1887FF95891B7117F833FBA7F7D2" pageId="7" pageNumber="338" type="etymology">
|
||
<paragraph id="1E184B0CFF95891B7117F833FBA7F7D2" blockId="7.[828,1437,1066,2081]" pageId="7" pageNumber="338">
|
||
<emphasis id="2CD3971EFF95891B7117F833FC1BF812" bold="true" box="[828,956,1994,2017]" pageId="7" pageNumber="338">Etymology.</emphasis>
|
||
The species is named in honour of Ottó Merkl (HNHM), who continuously and greatly helps us with the provided material.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="4AD81B84FF9A891472BCF93FFB20F907" ID-DOI="http://doi.org/10.5281/zenodo.4504782" ID-Zenodo-Dep="4504782" httpUri="https://zenodo.org/record/4504782/files/figure.png" pageId="8" pageNumber="339" startId="8.[151,182,1734,1753]" targetBox="[257,1330,189,1716]" targetPageId="8">
|
||
<paragraph id="1E184B0CFF9A891472BCF93FFB20F907" blockId="8.[151,1436,1727,1780]" pageId="8" pageNumber="339">
|
||
Fig. 5.
|
||
<taxonomicName id="D9A7308FFF9A891472FAF93FFE91F92A" authorityName="Solier" authorityYear="1835" box="[209,310,1734,1753]" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="339" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2CD3971EFF9A891472FAF93FFE91F92A" box="[209,310,1734,1753]" italics="true" pageId="8" pageNumber="339">Calyptopsis</emphasis>
|
||
</taxonomicName>
|
||
spp., habitus and details of structure.A –
|
||
<taxonomicName id="D9A7308FFF9A891470A5F93FFC12F92A" authority="Baudi di Selve, 1874" authorityName="Baudi di Selve" authorityYear="1874" box="[654,949,1734,1753]" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="339" phylum="Arthropoda" rank="species" species="armeniaca">
|
||
<emphasis id="2CD3971EFF9A891470A5F93FFCA7F92A" box="[654,768,1734,1753]" italics="true" pageId="8" pageNumber="339">C.armeniaca</emphasis>
|
||
<bibRefCitation id="7A3636FDFF9A8914712FF93FFC12F92A" author="BAUDI DI SELVE F." box="[772,949,1734,1753]" pageId="8" pageNumber="339" pagination="89 - 115" refId="ref10423" refString="BAUDI DI SELVE F. 1874: Catalogo dei tenebrioniti della fauna europea e circummediterranea del Museo Civico di Genova. Annali del Museo Civico di Storia Naturale di Genova 6: 89 - 115." type="journal article" year="1874">Baudi di Selve, 1874</bibRefCitation>
|
||
</taxonomicName>
|
||
, syntype,♁; B –
|
||
<taxonomicName id="D9A7308FFF9A89147669F93FFB12F92A" authorityName="Baudi di Selve" authorityYear="1874" box="[1090,1205,1734,1753]" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="339" phylum="Arthropoda" rank="species" species="armeniaca">
|
||
<emphasis id="2CD3971EFF9A89147669F93FFB12F92A" box="[1090,1205,1734,1753]" italics="true" pageId="8" pageNumber="339">C. armeniaca</emphasis>
|
||
</taxonomicName>
|
||
, syntype,♀; C–G –
|
||
<taxonomicName id="D9A7308FFF9A89147775F93FFA3BF92A" authority="Chigray & Nabozhenko & Keskin & Abdurakhmanov, 2018" authorityName="Chigray & Nabozhenko & Keskin & Abdurakhmanov" authorityYear="2018" box="[1374,1436,1734,1753]" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="339" phylum="Arthropoda" rank="species" species="ottoi" status="sp. nov.">
|
||
<emphasis id="2CD3971EFF9A89147775F93FFA3BF92A" box="[1374,1436,1734,1753]" italics="true" pageId="8" pageNumber="339">C. ottoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="37E02A65FF9A891472BCF918FF7AF907" box="[151,221,1761,1780]" pageId="8" pageNumber="339" rank="species">sp. nov.</taxonomicNameLabel>
|
||
: C – ♀ (Van, Erek Dağ); D – head; E – ♁ (Van, Görentaş); F – ♀ (Van, Görentaş); G – ♀ (Hakkari, Berçelan).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="56BD1887FF9A891572BCF8B5FECDF7E8" lastPageId="9" lastPageNumber="340" pageId="8" pageNumber="339" type="distribution">
|
||
<paragraph id="1E184B0CFF9A891472BCF8B5FD4AF890" blockId="8.[151,760,1868,2083]" box="[151,749,1868,1891]" pageId="8" pageNumber="339">
|
||
<emphasis id="2CD3971EFF9A891472BCF8B5FE82F890" bold="true" box="[151,293,1868,1891]" pageId="8" pageNumber="339">Distribution.</emphasis>
|
||
South-Eastern
|
||
<collectingCountry id="66B00B9CFF9A891473EDF8B5FDB5F890" box="[454,530,1868,1891]" name="Turkey" pageId="8" pageNumber="339">Turkey</collectingCountry>
|
||
, North-Western
|
||
<collectingCountry id="66B00B9CFF9A89147097F8B5FD4EF890" box="[700,745,1868,1891]" name="Iraq" pageId="8" pageNumber="339">Iraq</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="1E184B0CFF9A891572BCF895FECDF7E8" blockId="8.[151,760,1868,2083]" lastBlockId="9.[151,760,1796,2075]" lastPageId="9" lastPageNumber="340" pageId="8" pageNumber="339">
|
||
<emphasis id="2CD3971EFF9A891472BCF895FE03F870" bold="true" box="[151,420,1900,1923]" pageId="8" pageNumber="339">Comparative diagnosis.</emphasis>
|
||
The new species belongs to the species-group with beaded anterior margin, head separated from the other surface by transverse convexity. Externally
|
||
<taxonomicName id="D9A7308FFF9A891472BCF835FF40F810" authority="Chigray & Nabozhenko & Keskin & Abdurakhmanov, 2018" authorityName="Chigray & Nabozhenko & Keskin & Abdurakhmanov" authorityYear="2018" box="[151,231,1996,2019]" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="339" phylum="Arthropoda" rank="species" species="ottoi" status="sp. nov.">
|
||
<emphasis id="2CD3971EFF9A891472BCF835FF40F810" box="[151,231,1996,2019]" italics="true" pageId="8" pageNumber="339">C. ottoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="37E02A65FF9A891472C5F835FEE6F810" box="[238,321,1996,2019]" pageId="8" pageNumber="339" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is similar to
|
||
<taxonomicName id="D9A7308FFF9A891473E7F835FD1AF810" authority="Kaszab, 1962" authorityName="Kaszab" authorityYear="1962" box="[460,701,1996,2019]" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="339" phylum="Arthropoda" rank="species" species="gigas">
|
||
<emphasis id="2CD3971EFF9A891473E7F835FD82F810" box="[460,549,1996,2019]" italics="true" pageId="8" pageNumber="339">C. gigas</emphasis>
|
||
<bibRefCitation id="7A3636FDFF9A89147007F835FD1AF810" author="KASZAB Z." box="[556,701,1996,2019]" pageId="8" pageNumber="339" pagination="299 - 317" refId="ref10908" refString="KASZAB Z. 1962: Beitrage zur Kenntnis einiger asiatischen Tenebrioniden-Gattungen und Arten (Coleoptera). Annales Historico-Naturales Musei Nationalis Hungarici 54: 299 - 317." type="journal article" year="1962">Kaszab, 1962</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="DC6385EEFF9A891472BCF815FF53F7F0" box="[151,244,2028,2051]" country="Iran" name="Lorestan" pageId="8" pageNumber="339">Lorestan</collectingRegion>
|
||
(
|
||
<collectingCountry id="66B00B9CFF9A89147329F815FE97F7F0" box="[258,304,2028,2051]" name="Iran" pageId="8" pageNumber="339">Iran</collectingCountry>
|
||
) in form of its body, punctation of the pronotum and prothoracic hypomera but clearly differs from it in the head sculpture.
|
||
<taxonomicName id="D9A7308FFF9A89147637F8B4FB69F897" authority="Chigray & Nabozhenko & Keskin & Abdurakhmanov, 2018" authorityName="Chigray & Nabozhenko & Keskin & Abdurakhmanov" authorityYear="2018" box="[1052,1230,1869,1892]" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="339" phylum="Arthropoda" rank="species" species="ottoi" status="sp. nov.">
|
||
<emphasis id="2CD3971EFF9A89147637F8B4FB69F897" box="[1052,1230,1869,1892]" italics="true" pageId="8" pageNumber="339">Calyptopsis ottoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="37E02A65FF9A891476F8F8B4FA84F897" box="[1235,1315,1869,1892]" pageId="8" pageNumber="339" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has sharply beaded and elevated anterior margin of the head, which is separated from the other surface by deep depression, while
|
||
<taxonomicName id="D9A7308FFF9A89147117F854FC36F837" authorityName="Kaszab" authorityYear="1962" box="[828,913,1965,1988]" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="339" phylum="Arthropoda" rank="species" species="gigas">
|
||
<emphasis id="2CD3971EFF9A89147117F854FC36F837" box="[828,913,1965,1988]" italics="true" pageId="8" pageNumber="339">C. gigas</emphasis>
|
||
</taxonomicName>
|
||
possesses smooth beaded and not elevated anterior margin of the head, which is not separated from the other surface by transverse depression. The sharply beaded and separated anterior margin of head can be found also in
|
||
<taxonomicName id="D9A7308FFF9B891572BCF8FDFE7FF8E8" authority="Kaszab, 1962" authorityName="Kaszab" authorityYear="1962" box="[151,472,1796,1819]" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="theodoridesi">
|
||
<emphasis id="2CD3971EFF9B891572BCF8FDFE9AF8E8" box="[151,317,1796,1819]" italics="true" pageId="9" pageNumber="340">C. theodoridesi</emphasis>
|
||
<bibRefCitation id="7A3636FDFF9B8915736DF8FDFE7FF8E8" author="KASZAB Z." box="[326,472,1796,1819]" pageId="9" pageNumber="340" pagination="299 - 317" refId="ref10908" refString="KASZAB Z. 1962: Beitrage zur Kenntnis einiger asiatischen Tenebrioniden-Gattungen und Arten (Coleoptera). Annales Historico-Naturales Musei Nationalis Hungarici 54: 299 - 317." type="journal article" year="1962">Kaszab, 1962</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="D9A7308FFF9B89157024F8FDFE91F8C8" authority="Baudi di Selve, 1874" authorityName="Baudi di Selve" authorityYear="1874" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="harpaloides">
|
||
<emphasis id="2CD3971EFF9B89157024F8FDFD08F8E8" box="[527,687,1796,1819]" italics="true" pageId="9" pageNumber="340">C. harpaloides</emphasis>
|
||
<bibRefCitation id="7A3636FDFF9B89157093F8FDFE91F8C8" author="BAUDI DI SELVE F." pageId="9" pageNumber="340" pagination="89 - 115" refId="ref10423" refString="BAUDI DI SELVE F. 1874: Catalogo dei tenebrioniti della fauna europea e circummediterranea del Museo Civico di Genova. Annali del Museo Civico di Storia Naturale di Genova 6: 89 - 115." type="journal article" year="1874">Baudi di Selve, 1874</bibRefCitation>
|
||
</taxonomicName>
|
||
. The first species differs from
|
||
<taxonomicName id="D9A7308FFF9B891570ABF8DDFD75F8C8" authority="Chigray & Nabozhenko & Keskin & Abdurakhmanov, 2018" authorityName="Chigray & Nabozhenko & Keskin & Abdurakhmanov" authorityYear="2018" box="[640,722,1828,1851]" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="ottoi" status="sp. nov.">
|
||
<emphasis id="2CD3971EFF9B891570ABF8DDFD75F8C8" box="[640,722,1828,1851]" italics="true" pageId="9" pageNumber="340">C. ottoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="37E02A65FF9B891570F2F8DDFF62F8A8" pageId="9" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in robust body with very large pronotum 1.8× shorter than elytra, the second one differs in flattened and robust wide body, and emarginate near base lateral margins of pronotum. The new species has pronotum 2.20–2.35× shorter than elytra. From both species
|
||
<taxonomicName id="D9A7308FFF9B8915700EF83DFDD4F828" authority="Chigray & Nabozhenko & Keskin & Abdurakhmanov, 2018" authorityName="Chigray & Nabozhenko & Keskin & Abdurakhmanov" authorityYear="2018" box="[549,627,1988,2011]" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="ottoi" status="sp. nov.">
|
||
<emphasis id="2CD3971EFF9B8915700EF83DFDD4F828" box="[549,627,1988,2011]" italics="true" pageId="9" pageNumber="340">C. ottoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="37E02A65FF9B89157053F83DFD6EF828" box="[632,713,1988,2011]" pageId="9" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
|
||
also differs in the presence of distinct round impression in the middle of the frons.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |