116 lines
15 KiB
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116 lines
15 KiB
XML
<document id="0FBA0F1F2B97A42CDE8C41A0290E82C5" ID-CLB-Dataset="32717" ID-DOI="10.11646/zootaxa.4290.2.10" ID-GBIF-Dataset="2b8d8e70-876c-4df9-b3f4-ccb54eb91ae8" ID-ISSN="1175-5326" ID-Zenodo-Dep="828995" ID-ZooBank="D70803D6-1D43-4876-B0B6-C2350BB15569" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500110389050" checkinUser="plazi" docAuthor="Lambkin, Kevin J." docDate="2017" docId="2671BD2D8621FFD4FF73FD59FC65F995" docLanguage="en" docName="zootaxa.4290.2.10.pdf" docOrigin="Zootaxa 4290 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Archebittacus Riek 1955" docType="treatment" docVersion="7" lastPageNumber="386" masterDocId="DA48C5558620FFD5FFE4FFE1FFF9FFB9" masterDocTitle="Archebittacus exilis Riek — the oldest hangingfly (Insecta: Mecoptera: Bittacidae)" masterLastPageNumber="389" masterPageNumber="385" pageNumber="386" updateTime="1698465363758" updateUser="ExternalLinkService">
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<mods:titleInfo id="FE716F9512FE3180615EF8633F75993C">
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<mods:title id="2C1668E9E3A704D4D2CED28EA9B57634">Archebittacus exilis Riek — the oldest hangingfly (Insecta: Mecoptera: Bittacidae)</mods:title>
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<mods:name id="526B9C179478607FB2AEEBBFEA98EC8E" type="personal">
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<mods:namePart id="B11F2FF356418218636D486F3BC1DDDA">Lambkin, Kevin J.</mods:namePart>
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<mods:typeOfResource id="C0ABC565CCB4F351E3AFF30F7C267BF9">text</mods:typeOfResource>
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<mods:title id="335FC855277705F3CCC94048EF9427FD">Zootaxa</mods:title>
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<mods:part id="D981B15DD4954BCE0BE6C359CD842CF5">
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<mods:date id="A0B7314CFD5DDCEC90DC3F74C9582213">2017</mods:date>
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<mods:number id="2C62A15879DEBAF7BAC525307C44AB5D">4290</mods:number>
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<mods:number id="FB4C7ADF36B919A0B9C0021102B6956F">2</mods:number>
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<mods:identifier id="2CB6DA41E707638289455CD63121329E" type="DOI">10.11646/zootaxa.4290.2.10</mods:identifier>
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<treatment id="2671BD2D8621FFD4FF73FD59FC65F995" ID-DOI="http://doi.org/10.5281/zenodo.6010617" ID-GBIF-Taxon="132549225" ID-Zenodo-Dep="6010617" LSID="urn:lsid:plazi:treatment:2671BD2D8621FFD4FF73FD59FC65F995" httpUri="http://treatment.plazi.org/id/2671BD2D8621FFD4FF73FD59FC65F995" lastPageNumber="386" pageId="1" pageNumber="386">
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<subSubSection id="E6C25FB08621FFD4FF73FD59FE2BFD6B" box="[151,466,696,722]" pageId="1" pageNumber="386" type="nomenclature">
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<paragraph id="AE670C3B8621FFD4FF73FD59FE2BFD6B" blockId="1.[151,1436,696,1580]" box="[151,466,696,722]" pageId="1" pageNumber="386">
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<heading id="F52FBB578621FFD4FF73FD59FE2BFD6B" bold="true" box="[151,466,696,722]" fontSize="11" level="1" pageId="1" pageNumber="386" reason="1">
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<taxonomicName id="69D877B88621FFD4FF73FD59FE2BFD6B" authority="Riek, 1955" authorityName="Riek" authorityYear="1955" box="[151,466,696,722]" class="Insecta" family="Bittacidae" genus="Archebittacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mecoptera" pageId="1" pageNumber="386" phylum="Arthropoda" rank="genus">
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<emphasis id="9CACD0298621FFD4FF73FD59FE2BFD6B" bold="true" box="[151,466,696,722]" pageId="1" pageNumber="386">
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Archebittacus
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<bibRefCitation id="CA4971CA8621FFD4FEAAFD59FE2BFD6B" author="Riek" box="[334,466,696,722]" pageId="1" pageNumber="386" refString="Riek, E. F. (1955) Fossil insects from the Triassic beds at Mt. Crosby, Queensland. Australian Journal of Zoology, 3 (4), 654 - 691. https: // doi. org / 10.1071 / ZO 9550654" type="journal article" year="1955">Riek, 1955</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="E6C25FB08621FFD4FF73FD3BFE3BFD56" box="[151,450,730,751]" pageId="1" pageNumber="386" type="reference_group">
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<paragraph id="AE670C3B8621FFD4FF73FD3BFE3BFD56" blockId="1.[151,1436,696,1580]" box="[151,450,730,751]" pageId="1" pageNumber="386">
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<treatmentCitationGroup id="8EC82B158621FFD4FF73FD3BFE3BFD56" box="[151,450,730,751]" pageId="1" pageNumber="386">
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<taxonomicName id="69D877B88621FFD4FF73FD3BFE47FD56" authority="Riek, 1955: 666" authorityName="Riek" authorityPageNumber="666" authorityYear="1955" box="[151,446,730,751]" class="Insecta" family="Bittacidae" genus="Archebittacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mecoptera" pageId="1" pageNumber="386" phylum="Arthropoda" rank="genus">
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<emphasis id="9CACD0298621FFD4FF73FD3BFEE2FD56" box="[151,283,730,751]" italics="true" pageId="1" pageNumber="386">Archebittacus</emphasis>
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<treatmentCitation id="2F792A2A8621FFD4FEC5FD3BFE47FD56" author="Riek" box="[289,446,730,751]" page="666" pageId="1" pageNumber="386" year="1955">
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<bibRefCitation id="CA4971CA8621FFD4FEC5FD3BFE77FD56" author="Riek" box="[289,398,730,751]" pageId="1" pageNumber="386" refString="Riek, E. F. (1955) Fossil insects from the Triassic beds at Mt. Crosby, Queensland. Australian Journal of Zoology, 3 (4), 654 - 691. https: // doi. org / 10.1071 / ZO 9550654" type="journal article" year="1955">Riek, 1955</bibRefCitation>
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: 666
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="E6C25FB08621FFD4FF73FD19FC65F995" pageId="1" pageNumber="386" type="materials_examined">
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<paragraph id="AE670C3B8621FFD4FF73FD19FC70FCB6" blockId="1.[151,1436,696,1580]" box="[151,905,759,783]" pageId="1" pageNumber="386">
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<emphasis id="9CACD0298621FFD4FF73FD19FED2FCB6" bold="true" box="[151,299,760,783]" pageId="1" pageNumber="386">Type species.</emphasis>
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<taxonomicName id="69D877B88621FFD4FED5FD16FD78FCB6" authority="Riek, 1955" authorityName="Riek" authorityYear="1955" box="[305,641,759,783]" class="Insecta" family="Bittacidae" genus="Archebittacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mecoptera" pageId="1" pageNumber="386" phylum="Arthropoda" rank="species" species="exilis">
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<emphasis id="9CACD0298621FFD4FED5FD16FDFAFCB6" box="[305,515,759,783]" italics="true" pageId="1" pageNumber="386">Archebittacus exilis</emphasis>
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<bibRefCitation id="CA4971CA8621FFD4FDEEFD19FD78FCB6" author="Riek" box="[522,641,760,783]" pageId="1" pageNumber="386" refString="Riek, E. F. (1955) Fossil insects from the Triassic beds at Mt. Crosby, Queensland. Australian Journal of Zoology, 3 (4), 654 - 691. https: // doi. org / 10.1071 / ZO 9550654" type="journal article" year="1955">Riek, 1955</bibRefCitation>
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</taxonomicName>
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, by original designation.
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</paragraph>
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<paragraph id="AE670C3B8621FFD4FF23FCFBFB05FB16" blockId="1.[151,1436,696,1580]" pageId="1" pageNumber="386">
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<emphasis id="9CACD0298621FFD4FF23FCFBFE3DFC88" bold="true" box="[199,452,794,817]" pageId="1" pageNumber="386">Diagnostic characters.</emphasis>
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Forewing sub-petiolate, base narrowed but not as much as in extant and most fossil genera; costal space without crossveins (humeral vein apparently not preserved); Sc simple, terminating just before level of R4+5 fork; one sc–r1 at level of primary fork of Rs; R1 with two short terminal branches, R1b evenly curved; pterostigma quite long, defined posteriorly by a fine, evenly curved groove; one pterostigmal crossvein, running to R2+3 not R2 (preserved only between pterostigmal groove and R2+3); one backwardly oblique r1–r2+3; primary fork of R distal to level of separation of M and CuA; R2+3 with two branches; stem of Rs much shorter than its branches; primary fork of M slightly proximal to that of Rs; M with five branches, M3+4 fork distinctly proximal to that of M1+2, M4 deeply forked; m–cu variable, running from M3+4 (
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<figureCitation id="36E310BE8621FFD4FD24FBECFD00FB9D" box="[704,761,1037,1060]" captionStart="FIGURES 4 – 5" captionStartId="3.[151,258,922,944]" captionTargetBox="[180,1413,208,877]" captionTargetId="figure@3.[170,1417,191,900]" captionTargetPageId="3" captionText="FIGURES 4 – 5. Archebittacus exilis Riek: 4, forewing; 5, hindwing; Pcv, pterostigmal crossvein, Pg, pterostigmal groove." httpUri="https://zenodo.org/record/828999/files/figure.png" pageId="1" pageNumber="386">Fig.4</figureCitation>
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) or M4 (
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<figureCitation id="36E310BE8621FFD4FCBCFBECFC6AFB9D" box="[856,915,1037,1060]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,258,1665,1687]" captionTargetBox="[200,1386,228,1584]" captionTargetId="figure@2.[200,1386,228,1585]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Archebittacus exilis Riek, forewing: 1, holotype, UQG C 2139 (mirrored); 2, paratype, UQG C 1495 (mirrored); 3, paratype, UQG C 1545." httpUri="https://zenodo.org/record/828997/files/figure.png" pageId="1" pageNumber="386">Fig.2</figureCitation>
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); 1r–m, m1+2–m3, and m–cua not aligned, quite discordantly oblique; apical fields of Rs and M with limited crossveins (although slight kinking detected in some longitudinal veins suggesting the presence of more than are preserved); stem of CuA oblique, fused with M for a very short distance; one, or possibly two, cua–cup; 1A long, extending to level of primary fork of M; 1A and 2A more widely spaced than in extant and most fossil genera; 2A extending beyond level of the separation of M and CuA.
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</paragraph>
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<paragraph id="AE670C3B8621FFD4FF23FB5BFD43FA18" blockId="1.[151,1436,696,1580]" pageId="1" pageNumber="386">Hindwing shape and costal space as in forewing; Sc simple, shorter than in forewing, terminating proximal to primary fork of Rs; one apical sc–r1; relative positions of forks of main veins and terminations of 1A and 2A as in forewing; apex of R1 and pterostigmal area indistinctly preserved; Rs as in forewing; M with four branches, M3+4 fork distinctly proximal to that of M1+2; r–m, im, and m–cua not aligned, the latter two less oblique than in forewing; apical fields of Rs and M indistinctly preserved, crossveins not detectable; one cua–cup; one cup–1a; CuP and anal veins of usual bittacid form (CuP and 1A fused basally, 1A after separation from CuP forming a Y-vein with 1a–2a), except 2A and 3A longer and more widely spaced; one 2a–3a.</paragraph>
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<paragraph id="AE670C3B8621FFD4FF23FA4CFC65F995" blockId="1.[151,1436,696,1580]" pageId="1" pageNumber="386">
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<emphasis id="9CACD0298621FFD4FF23FA4CFEF4FA7D" bold="true" box="[199,269,1453,1476]" pageId="1" pageNumber="386">Notes.</emphasis>
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As noted by
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<bibRefCitation id="CA4971CA8621FFD4FE78FA4CFDE6FA7D" author="Riek" box="[412,543,1453,1476]" pageId="1" pageNumber="386" refString="Riek, E. F. (1955) Fossil insects from the Triassic beds at Mt. Crosby, Queensland. Australian Journal of Zoology, 3 (4), 654 - 691. https: // doi. org / 10.1071 / ZO 9550654" type="journal article" year="1955">Riek (1955)</bibRefCitation>
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,
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<taxonomicName id="69D877B88621FFD4FDCFFA4DFD47FA7D" box="[555,702,1452,1476]" class="Insecta" family="Bittacidae" genus="Archebittacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mecoptera" pageId="1" pageNumber="386" phylum="Arthropoda" rank="genus">
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<emphasis id="9CACD0298621FFD4FDCFFA4DFD47FA7D" box="[555,702,1452,1476]" italics="true" pageId="1" pageNumber="386">Archebittacus</emphasis>
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</taxonomicName>
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is most similar to the Jurassic
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<taxonomicName id="69D877B88621FFD4FBECFA4DFACEFA7D" authority="Tillyard, 1933" authorityName="Tillyard" authorityYear="1933" box="[1032,1335,1452,1476]" class="Insecta" family="Bittacidae" genus="Protobittacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mecoptera" pageId="1" pageNumber="386" phylum="Arthropoda" rank="genus">
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<emphasis id="9CACD0298621FFD4FBECFA4DFB61FA7D" box="[1032,1176,1452,1476]" italics="true" pageId="1" pageNumber="386">Protobittacus</emphasis>
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<bibRefCitation id="CA4971CA8621FFD4FB7BFA4CFACEFA7D" author="Tillyard" box="[1183,1335,1453,1476]" pageId="1" pageNumber="386" refString="Tillyard, R. J. (1933) The panorpoid complex in the British Rhaetic and Lias. British Museum (Natural History) Fossil Insects, 3, 1 - 79." type="journal article" year="1933">Tillyard, 1933</bibRefCitation>
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</taxonomicName>
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(
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<bibRefCitation id="CA4971CA8621FFD4FAA1FA4CFED6FA5E" author="Tillyard" pageId="1" pageNumber="386" refString="Tillyard, R. J. (1933) The panorpoid complex in the British Rhaetic and Lias. British Museum (Natural History) Fossil Insects, 3, 1 - 79." type="journal article" year="1933">Tillyard 1933, Fig. 18</bibRefCitation>
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), but the forewing differs in the more proximal position of sc–r1, the presence of r1–r2+3 (absent in
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<taxonomicName id="69D877B88621FFD4FF73FA10FEDEF9B0" box="[151,295,1521,1545]" class="Insecta" family="Bittacidae" genus="Protobittacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mecoptera" pageId="1" pageNumber="386" phylum="Arthropoda" rank="genus">
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<emphasis id="9CACD0298621FFD4FF73FA10FEDEF9B0" box="[151,295,1521,1545]" italics="true" pageId="1" pageNumber="386">Protobittacus</emphasis>
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</taxonomicName>
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), the crossvein between M and CuA originating from M3+4 or stem of M4 (from posterior branch of M
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<quantity id="6920A1DE8621FFD4FA97FA13FA62F9B0" box="[1395,1435,1522,1545]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="1" pageNumber="386" unit="in" value="4.0">4 in</quantity>
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<taxonomicName id="69D877B88621FFD4FF73F9F5FEDEF995" box="[151,295,1556,1580]" class="Insecta" family="Bittacidae" genus="Protobittacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mecoptera" pageId="1" pageNumber="386" phylum="Arthropoda" rank="genus">
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<emphasis id="9CACD0298621FFD4FF73F9F5FEDEF995" box="[151,295,1556,1580]" italics="true" pageId="1" pageNumber="386">Protobittacus</emphasis>
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</taxonomicName>
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), and in the presence of cua–cup (absent in
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<taxonomicName id="69D877B88621FFD4FD18F9F5FC75F995" box="[764,908,1556,1580]" class="Insecta" family="Bittacidae" genus="Protobittacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mecoptera" pageId="1" pageNumber="386" phylum="Arthropoda" rank="genus">
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<emphasis id="9CACD0298621FFD4FD18F9F5FC75F995" box="[764,908,1556,1580]" italics="true" pageId="1" pageNumber="386">Protobittacus</emphasis>
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</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |