375 lines
48 KiB
XML
375 lines
48 KiB
XML
<document id="B8A211B15C7EE10C1A834B1445A4F991" ID-CLB-Dataset="55715" ID-DOI="10.5852/ejt.2022.817.1765" ID-GBIF-Dataset="509d55a8-15d2-4c36-8083-b3ec7e9a8ecf" ID-ISSN="2118-9773" ID-Zenodo-Dep="6518305" ID-ZooBank="D55FA216-086F-4226-8DDD-E64CC980AE19" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1651674223624" checkinUser="felipe" docAuthor="Satkunanathan, Abira & Benjamin, Suresh P." docDate="2022" docId="686687F0A706D66E7B4D6E25DBA240B7" docLanguage="en" docName="ejt-817_78-110.pdf" docOrigin="European Journal of Taxonomy 817 (1)" docSource="http://dx.doi.org/10.5852/ejt.2022.817.1765" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Carrhotus atratus Satkunanathan & Benjamin 2022, sp. nov." docType="treatment" docUuid="D26852D1-B21E-41CE-8291-78FBAD0DE9A7" docUuidSource="ZooBank" docVersion="9" lastPageNumber="92" masterDocId="945FFF88A70CD66079306F2DDF304232" masterDocTitle="Phylogenetic placement of Carrhotus Thorell, 1891 with three new species from Sri Lanka (Araneae: Salticidae)" masterLastPageNumber="110" masterPageNumber="78" pageNumber="88" updateTime="1699329587011" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
|
||
<mods:mods id="23C906BAADA3831A34D73944E25E3ED1" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="BAADF12AAECE12F329AD768E2AAE41AD">
|
||
<mods:title id="AED1416D3065CC3A514CE20B06186BB4">Phylogenetic placement of Carrhotus Thorell, 1891 with three new species from Sri Lanka (Araneae: Salticidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="E197CB338FD2EFB73F75B47B056F1AD5" type="personal">
|
||
<mods:role id="AD1DDD69A8D3D88BAC4D06461B0BF809">
|
||
<mods:roleTerm id="41253A0E7A8134FB10511B5174D8FBA2">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="28A5F5307D5BEACB0E0857B0B89EFBD0">Satkunanathan, Abira</mods:namePart>
|
||
<mods:nameIdentifier id="DAA913B33FEB8241B63D655DBF52E926" type="LSID">65DE6669-0A51-4145-BBED-A57D17C49402</mods:nameIdentifier>
|
||
<mods:affiliation id="0F9B7BC23FCA6E4DADFD20801044CC2A">National Institute of Fundamental Studies, Hantana Road, Kandy, Sri Lanka.</mods:affiliation>
|
||
<mods:nameIdentifier id="F22569F61D2E2B6F3FA854EFC96C2E32" type="email">abirasatkuna11@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="293B27943739FA536ABD34F8564CD721" type="personal">
|
||
<mods:role id="27062F8877AEEE411848B6935307B0CE">
|
||
<mods:roleTerm id="1EBA48D994D222DAB40DB18F3DB43016">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="818F5ED258942FB58CD1AE652410DAEB">Benjamin, Suresh P.</mods:namePart>
|
||
<mods:nameIdentifier id="E96608228C475BE28870D2AE500461EB" type="LSID">986CED51-9425-4CB6-84A4-A9464BB9207E</mods:nameIdentifier>
|
||
<mods:affiliation id="E5E1BF980089A665EA8E3D7A11B84B42">National Institute of Fundamental Studies, Hantana Road, Kandy, Sri Lanka.</mods:affiliation>
|
||
<mods:nameIdentifier id="36DDCA50C9197421271672FD2A225BE4" type="email">suresh.benjamin@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource id="9DAAA8752CC296DBFBE32C530533C36D">text</mods:typeOfResource>
|
||
<mods:relatedItem id="946DB6DDF99B6C33572DFD48E7398E2E" type="host">
|
||
<mods:titleInfo id="E9775932E64106D6E2AFC7706CF8082D">
|
||
<mods:title id="72999D7368C1AF356593DEF8429A8B2C">European Journal of Taxonomy</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="CAA9CAE5F4688C8391931AF2C185A8F0">
|
||
<mods:date id="51C3B7DEDFA9F1D939F0C81D39A1C26C">2022</mods:date>
|
||
<mods:detail id="770F12ED4BC59EAA651B9A4E06AEE37B" type="pubDate">
|
||
<mods:number id="A8F6054F47F33CF12436461036A62A52">2022-05-02</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="2727B5BC122B8ACE9DD16118726418D6" type="volume">
|
||
<mods:number id="57D53BEFC7D5E866437F85A30DE8ED7A">817</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="2FFC867C11047646BC2D79E0A20B8861" type="issue">
|
||
<mods:number id="B4B18DD050F48BA8A75449EE558070CD">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="99B0F07FAEF60F43FF6ED8260750462A" unit="page">
|
||
<mods:start id="87C1BE25DB0728958C5364A6E15D4745">78</mods:start>
|
||
<mods:end id="A3A1C78BC9BAD2CF8853F25E757EA168">110</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="847D43DAC33315CC55413D106C3DE872">
|
||
<mods:url id="E73DFBF8094B0CBDE0BD290BF8607035">http://dx.doi.org/10.5852/ejt.2022.817.1765</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="1220767709396D7821763285B6AC3E9F">journal article</mods:classification>
|
||
<mods:identifier id="0A6FCF9B8545F68EE92E67EA5DA0F577" type="CLB-Dataset">55715</mods:identifier>
|
||
<mods:identifier id="7448025F635C059521B6E784A50861BF" type="DOI">10.5852/ejt.2022.817.1765</mods:identifier>
|
||
<mods:identifier id="32B96891D5989A4F6D35AE3A19AE55FF" type="GBIF-Dataset">509d55a8-15d2-4c36-8083-b3ec7e9a8ecf</mods:identifier>
|
||
<mods:identifier id="A4C3215B620F369D0C67C26825891404" type="ISSN">2118-9773</mods:identifier>
|
||
<mods:identifier id="A9EBDED6E7755E5FDFB26B08712AD1DD" type="Zenodo-Dep">6518305</mods:identifier>
|
||
<mods:identifier id="6F78547F1C6BF5B25230309EE74DD526" type="ZooBank">D55FA216-086F-4226-8DDD-E64CC980AE19</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="686687F0A706D66E7B4D6E25DBA240B7" ID-DOI="http://doi.org/10.5281/zenodo.6517924" ID-GBIF-Taxon="195191971" ID-Zenodo-Dep="6517924" LSID="urn:lsid:zoobank.org:act:D26852D1-B21E-41CE-8291-78FBAD0DE9A7" httpUri="http://treatment.plazi.org/id/686687F0A706D66E7B4D6E25DBA240B7" lastPageId="14" lastPageNumber="92" pageId="10" pageNumber="88">
|
||
<subSubSection id="A8D5656DA706D66A7B4D6E25DC874311" box="[637,951,264,291]" pageId="10" pageNumber="88" type="nomenclature">
|
||
<paragraph id="E07036E6A706D66A7B4D6E25DC874311" blockId="10.[360,1227,264,362]" box="[637,951,264,291]" pageId="10" pageNumber="88">
|
||
<heading id="BB38818AA706D66A7B4D6E25DC874311" box="[637,951,264,291]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="88" reason="2">
|
||
<taxonomicName id="27CF4D65A706D66A7B4D6E25DC654311" ID-CoL="992H3" authority="Satkunanathan & Benjamin, 2022" authorityName="Satkunanathan & Benjamin" authorityYear="2022" box="[637,853,264,291]" class="Arachnida" family="Salticidae" genus="Carrhotus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="88" phylum="Arthropoda" rank="species" species="atratus" status="sp. nov.">
|
||
<emphasis id="D2BBEAF4A706D66A7B4D6E25DC654311" bold="true" box="[637,853,264,291]" italics="true" pageId="10" pageNumber="88">Carrhotus atratus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C988578FA706D66A7A6C6E24DC874311" box="[860,951,265,291]" pageId="10" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A8D5656DA706D66A78586E01DCF1435B" pageId="10" pageNumber="88" type="description">
|
||
<paragraph id="E07036E6A706D66A78586E01DBFB4375" blockId="10.[360,1227,264,362]" box="[360,1227,300,327]" pageId="10" pageNumber="88">
|
||
<uri id="945E3AE4A706D66A78586E01DBFB4375" box="[360,1227,300,327]" pageId="10" pageNumber="88">
|
||
urn:lsid:zoobank.org:act:
|
||
<uuid id="94690C33A706D66A7BB96E01DBFB4375" box="[649,1227,300,327]" pageId="10" pageNumber="88">D26852D1-B21E-41CE-8291-78FBAD0DE9A7</uuid>
|
||
</uri>
|
||
</paragraph>
|
||
<paragraph id="E07036E6A706D66A7B426E62DCF1435B" blockId="10.[360,1227,264,362]" box="[626,961,335,362]" pageId="10" pageNumber="88">
|
||
<figureCitation id="78F42A63A706D66A7B426E62DDDB4358" box="[626,747,335,362]" captionStart="Fig 5" captionStartId="10.[189,230,1962,1988]" captionTargetBox="[189,1398,684,1918]" captionTargetId="figure-104@10.[189,1398,684,1918]" captionTargetPageId="10" captionText="Fig 5. Photographs of live Carrhotus atratus sp. nov. A–B, E–F, I. Males. C–D, G–H. Females. A–H from Mandaitivu, I from Hiyare." figureDoi="http://doi.org/10.5281/zenodo.6518317" httpUri="https://zenodo.org/record/6518317/files/figure.png" pageId="10" pageNumber="88">Figs 5A–I</figureCitation>
|
||
,
|
||
<figureCitation id="78F42A63A706D66A7BC76E62DC72435B" box="[759,834,335,361]" captionStart="Fig 6" captionStartId="12.[189,230,1807,1833]" captionTargetBox="[189,1398,269,1754]" captionTargetId="figure-11@12.[189,1398,269,1754]" captionTargetPageId="12" captionText="Fig 6. Carrhotus atratus sp. nov. A–B. Male habitus. A. Dorsal view. B. Ventral view. C–D. Female habitus.C. Dorsal view. D. Ventral view. E–F. Palp. E. Ventral view. F. Retrolateral view. G. Epigynum. Abbreviations: E = embolus; PLP = posterior lateral protrusion; RTA = retrolateral tibial apophysis. Scale bars: A–D = 2 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6518321" httpUri="https://zenodo.org/record/6518321/files/figure.png" pageId="10" pageNumber="88">6A–G</figureCitation>
|
||
,
|
||
<figureCitation id="78F42A63A706D66A7A7D6E7DDCA84358" box="[845,920,336,362]" captionStart="Fig 7" captionStartId="13.[189,230,1829,1855]" captionTargetBox="[207,1392,281,1766]" captionTargetId="figure-15@13.[189,1398,266,1776]" captionTargetPageId="13" captionText="Fig 7. Carrhotus atratus sp. nov. A–B. Palp. A. Ventral view. B. Retrolateral view. C. Epigynum ventral view. D. Vulva dorsal view. Abbreviations: AG = accessory gland; CO = copulatory opening; E = embolus; FD = fertilization duct; PLP = posterior lateral protrusion; RTA = retrolateral tibial apophysis; S = spermatheca; TE = tegulum. Scale bars: A–B = 0.2 mm; C–D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6518323" httpUri="https://zenodo.org/record/6518323/files/figure.png" pageId="10" pageNumber="88">7A–D</figureCitation>
|
||
,
|
||
<figureCitation id="78F42A63A706D66A7A946E62DCF1435B" box="[932,961,335,361]" captionStart="Fig 17" captionStartId="28.[189,230,1982,2008]" captionTargetBox="[189,1382,719,1923]" captionTargetId="figure-115@28.[189,1397,718,1944]" captionTargetPageId="28" captionText="Fig 17. Distribution of Carrhotus spp. of Sri Lanka." figureDoi="http://doi.org/10.5281/zenodo.6518352" httpUri="https://zenodo.org/record/6518352/files/figure.png" pageId="10" pageNumber="88">17</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A8D5656DA706D66A798D6EBDDA234054" pageId="10" pageNumber="88" type="diagnosis">
|
||
<paragraph id="E07036E6A706D66A798D6EBDDE064398" blockId="10.[189,310,400,426]" box="[189,310,400,426]" pageId="10" pageNumber="88">
|
||
<heading id="BB38818AA706D66A798D6EBDDE064398" bold="true" box="[189,310,400,426]" fontSize="11" level="3" pageId="10" pageNumber="88" reason="3">
|
||
<emphasis id="D2BBEAF4A706D66A798D6EBDDE064398" bold="true" box="[189,310,400,426]" pageId="10" pageNumber="88">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="E07036E6A706D66A798D6E92DA234054" blockId="10.[188,1399,447,615]" pageId="10" pageNumber="88">
|
||
Males of
|
||
<taxonomicName id="27CF4D65A706D66A781E6EEDDE9443EB" authority="Satkunanathan & Benjamin, 2022" authorityName="Satkunanathan & Benjamin" authorityYear="2022" box="[302,420,448,474]" class="Arachnida" family="Salticidae" genus="Carrhotus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="88" phylum="Arthropoda" rank="species" species="atratus" status="sp. nov.">
|
||
<emphasis id="D2BBEAF4A706D66A781E6EEDDE9443EB" box="[302,420,448,474]" italics="true" pageId="10" pageNumber="88">C. atratus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C988578FA706D66A789E6EEDDD3A43E8" box="[430,522,448,474]" pageId="10" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be separated from those of other congeners by their triangular-shaped PLP and cone-shaped anticlockwise directed embolus tip (
|
||
<figureCitation id="78F42A63A706D66A7A4D6ECFDCCC43CE" box="[893,1020,482,508]" captionStart="Fig 6" captionStartId="12.[189,230,1807,1833]" captionTargetBox="[189,1398,269,1754]" captionTargetId="figure-11@12.[189,1398,269,1754]" captionTargetPageId="12" captionText="Fig 6. Carrhotus atratus sp. nov. A–B. Male habitus. A. Dorsal view. B. Ventral view. C–D. Female habitus.C. Dorsal view. D. Ventral view. E–F. Palp. E. Ventral view. F. Retrolateral view. G. Epigynum. Abbreviations: E = embolus; PLP = posterior lateral protrusion; RTA = retrolateral tibial apophysis. Scale bars: A–D = 2 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6518321" httpUri="https://zenodo.org/record/6518321/files/figure.png" pageId="10" pageNumber="88">Figs 6E–F</figureCitation>
|
||
,
|
||
<figureCitation id="78F42A63A706D66A7D3A6ECEDB6543CF" box="[1034,1109,483,509]" captionStart="Fig 7" captionStartId="13.[189,230,1829,1855]" captionTargetBox="[207,1392,281,1766]" captionTargetId="figure-15@13.[189,1398,266,1776]" captionTargetPageId="13" captionText="Fig 7. Carrhotus atratus sp. nov. A–B. Palp. A. Ventral view. B. Retrolateral view. C. Epigynum ventral view. D. Vulva dorsal view. Abbreviations: AG = accessory gland; CO = copulatory opening; E = embolus; FD = fertilization duct; PLP = posterior lateral protrusion; RTA = retrolateral tibial apophysis; S = spermatheca; TE = tegulum. Scale bars: A–B = 0.2 mm; C–D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6518323" httpUri="https://zenodo.org/record/6518323/files/figure.png" pageId="10" pageNumber="88">7A–B</figureCitation>
|
||
). The palp of
|
||
<taxonomicName id="27CF4D65A706D66A7DCE6ECEDA4643CE" authorityName="Satkunanathan & Benjamin" authorityYear="2022" box="[1278,1398,483,509]" class="Arachnida" family="Salticidae" genus="Carrhotus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="88" phylum="Arthropoda" rank="species" species="atratus">
|
||
<emphasis id="D2BBEAF4A706D66A7DCE6ECEDA4643CE" box="[1278,1398,483,509]" italics="true" pageId="10" pageNumber="88">C. atratus</emphasis>
|
||
</taxonomicName>
|
||
closely resembles that of
|
||
<taxonomicName id="27CF4D65A706D66A78C16D2BDC2B4012" authority="Jastrzebski 1999" authorityName="Jastrzebski" authorityYear="1999" box="[497,795,518,544]" class="Arachnida" family="Salticidae" genus="Carrhotus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="88" phylum="Arthropoda" rank="species" species="erus">
|
||
<emphasis id="D2BBEAF4A706D66A78C16D2BDD794012" box="[497,585,518,544]" italics="true" pageId="10" pageNumber="88">C. erus</emphasis>
|
||
<bibRefCitation id="845E4B17A706D66A7B636D2BDC2B4012" author="Jastrzebski P." box="[595,795,518,544]" pageId="10" pageNumber="88" pagination="1 - 9" refId="ref13287" refString="Jastrzebski P. 1999. Salticidae from the Himalaya: the genus Carrhotus Thorell 1891 (Araneae, Salticidae). Senckenbergiana Biologica 9: 1 - 9." type="journal article" year="1999">Jastrzebski 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
; however, its PLP is relatively short, rounded and RTA is not pointed. RTA is smaller and tapered (
|
||
<figureCitation id="78F42A63A706D66A7BD96D04DC704076" box="[745,832,553,580]" captionStart="Fig 6" captionStartId="12.[189,230,1807,1833]" captionTargetBox="[189,1398,269,1754]" captionTargetId="figure-11@12.[189,1398,269,1754]" captionTargetPageId="12" captionText="Fig 6. Carrhotus atratus sp. nov. A–B. Male habitus. A. Dorsal view. B. Ventral view. C–D. Female habitus.C. Dorsal view. D. Ventral view. E–F. Palp. E. Ventral view. F. Retrolateral view. G. Epigynum. Abbreviations: E = embolus; PLP = posterior lateral protrusion; RTA = retrolateral tibial apophysis. Scale bars: A–D = 2 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6518321" httpUri="https://zenodo.org/record/6518321/files/figure.png" pageId="10" pageNumber="88">Fig. 6F</figureCitation>
|
||
) than in other
|
||
<taxonomicName id="27CF4D65A706D66A7AD46D04DB894071" box="[996,1209,553,579]" pageId="10" pageNumber="88">
|
||
<emphasis id="D2BBEAF4A706D66A7AD46D04DB6C4071" box="[996,1116,553,579]" italics="true" pageId="10" pageNumber="88">Carrhotus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="98D87676A706D66A7DCE6D04DA434071" box="[1278,1395,553,579]" name="Sri Lanka" pageId="10" pageNumber="88">Sri Lanka</collectingCountry>
|
||
. Females are distinguished by the prominent hood at the CO and hook-shaped CD (
|
||
<figureCitation id="78F42A63A706D66A7DB56D61DA374054" box="[1157,1287,588,614]" captionStart="Fig 7" captionStartId="13.[189,230,1829,1855]" captionTargetBox="[207,1392,281,1766]" captionTargetId="figure-15@13.[189,1398,266,1776]" captionTargetPageId="13" captionText="Fig 7. Carrhotus atratus sp. nov. A–B. Palp. A. Ventral view. B. Retrolateral view. C. Epigynum ventral view. D. Vulva dorsal view. Abbreviations: AG = accessory gland; CO = copulatory opening; E = embolus; FD = fertilization duct; PLP = posterior lateral protrusion; RTA = retrolateral tibial apophysis; S = spermatheca; TE = tegulum. Scale bars: A–B = 0.2 mm; C–D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6518323" httpUri="https://zenodo.org/record/6518323/files/figure.png" pageId="10" pageNumber="88">Fig. 7C–D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="B4B0666EA706D66A798D6887DD0C45DA" ID-DOI="http://doi.org/10.5281/zenodo.6518317" ID-Zenodo-Dep="6518317" httpUri="https://zenodo.org/record/6518317/files/figure.png" pageId="10" pageNumber="88" startId="10.[189,230,1962,1988]" targetBox="[189,1398,684,1918]" targetPageId="10">
|
||
<paragraph id="E07036E6A706D66A798D6887DD0C45DA" blockId="10.[188,1399,1962,2024]" pageId="10" pageNumber="88">
|
||
<emphasis id="D2BBEAF4A706D66A798D6887DFCD45F7" bold="true" box="[189,253,1962,1989]" pageId="10" pageNumber="88">Fig 5</emphasis>
|
||
. Photographs of live
|
||
<taxonomicName id="27CF4D65A706D66A78CD6886DDE145F6" authority="Satkunanathan & Benjamin, 2022" authorityName="Satkunanathan & Benjamin" authorityYear="2022" box="[509,721,1963,1989]" class="Arachnida" family="Salticidae" genus="Carrhotus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="88" phylum="Arthropoda" rank="species" species="atratus" status="sp. nov.">
|
||
<emphasis id="D2BBEAF4A706D66A78CD6886DDE145F6" box="[509,721,1963,1989]" italics="true" pageId="10" pageNumber="88">Carrhotus atratus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C988578FA706D66A7BE86886DC0445F7" box="[728,820,1963,1989]" pageId="10" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="D2BBEAF4A706D66A7A0D6887DC4345F6" bold="true" box="[829,883,1962,1988]" pageId="10" pageNumber="88">A–B</emphasis>
|
||
,
|
||
<emphasis id="D2BBEAF4A706D66A7AB36886DC8645F7" bold="true" box="[899,950,1963,1989]" pageId="10" pageNumber="88">E–F</emphasis>
|
||
,
|
||
<emphasis id="D2BBEAF4A706D66A7AF76886DCE245F7" bold="true" box="[967,978,1963,1989]" pageId="10" pageNumber="88">I</emphasis>
|
||
. Males.
|
||
<emphasis id="D2BBEAF4A706D66A7D0A6887DB4245F6" bold="true" box="[1082,1138,1962,1988]" pageId="10" pageNumber="88">C–D</emphasis>
|
||
,
|
||
<emphasis id="D2BBEAF4A706D66A7DB26887DB8E45F6" bold="true" box="[1154,1214,1962,1988]" pageId="10" pageNumber="88">G–H</emphasis>
|
||
. Females. A–H from Mandaitivu, I from Hiyare.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="A8D5656DA707D66B798D6E24DE884346" pageId="11" pageNumber="89" type="etymology">
|
||
<paragraph id="E07036E6A707D66B798D6E24DE744311" blockId="11.[189,324,265,291]" box="[189,324,265,291]" pageId="11" pageNumber="89">
|
||
<heading id="BB38818AA707D66B798D6E24DE744311" bold="true" box="[189,324,265,291]" fontSize="11" level="3" pageId="11" pageNumber="89" reason="3">
|
||
<emphasis id="D2BBEAF4A707D66B798D6E24DE744311" bold="true" box="[189,324,265,291]" pageId="11" pageNumber="89">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="E07036E6A707D66B798D6E1ADE884346" blockId="11.[188,1399,311,372]" pageId="11" pageNumber="89">
|
||
The specific epithet is derived from the Latin word ‘
|
||
<taxonomicName id="27CF4D65A707D66B7A2B6E15DC5E4363" authorityName="Satkunanathan & Benjamin" authorityYear="2022" box="[795,878,312,337]" class="Arachnida" family="Salticidae" genus="Carrhotus" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="89" phylum="Arthropoda" rank="species" species="atratus">
|
||
<emphasis id="D2BBEAF4A707D66B7A2B6E15DC5E4363" box="[795,878,312,337]" italics="true" pageId="11" pageNumber="89">atratus</emphasis>
|
||
</taxonomicName>
|
||
’ meaning ‘darkened’, loosely referring to its blackish body colour.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A8D5656DA707D66B798D6EB6DECA416C" pageId="11" pageNumber="89" type="materials_examined">
|
||
<paragraph id="E07036E6A707D66B798D6EB6DE9A4387" blockId="11.[189,426,411,437]" box="[189,426,411,437]" pageId="11" pageNumber="89">
|
||
<heading id="BB38818AA707D66B798D6EB6DE9A4387" bold="true" box="[189,426,411,437]" fontSize="11" level="3" pageId="11" pageNumber="89" reason="3">
|
||
<emphasis id="D2BBEAF4A707D66B798D6EB6DE9A4387" bold="true" box="[189,426,411,437]" pageId="11" pageNumber="89">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="E07036E6A707D66B79DA6EE7DCF84019" blockId="11.[189,1398,458,555]" pageId="11" pageNumber="89">
|
||
<materialsCitation id="50A73CBBA707D66B79DA6EE7DCF84019" ID-GBIF-Occurrence="3762961323" collectingDate="2020-01-12" collectingMethod="beating" collectionCode="NIFS" collectorName="S. P. Benjamin" country="Sri Lanka" elevation="12" location="Mandaitivu" municipality="Jaffna District" pageId="11" pageNumber="89" specimenCode="NIFS_SAL_1280" specimenCount="1" specimenCount-male="1" stateProvince="Northern Province" typeStatus="holotype">
|
||
<emphasis id="D2BBEAF4A707D66B79DA6EE7DE6C43D6" bold="true" box="[234,348,458,484]" pageId="11" pageNumber="89">
|
||
<typeStatus id="3F748844A707D66B79DA6EE7DE6C43D6" box="[234,348,458,484]" pageId="11" pageNumber="89" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="98D87676A707D66B798D6EC3DE66403A" box="[189,342,494,520]" name="Sri Lanka" pageId="11" pageNumber="89">SRI LANKA</collectingCountry>
|
||
•
|
||
<specimenCount id="F6C9FD6FA707D66B785C6EC3DEB5403A" box="[364,389,494,520]" pageId="11" pageNumber="89" type="male">♂</specimenCount>
|
||
;
|
||
<collectingRegion id="220BF804A707D66B78A06EC0DD5B4035" box="[400,619,493,519]" country="Sri Lanka" name="Northern" pageId="11" pageNumber="89">Northern Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="0014AC9CA707D66B7B456EC0DC2D4035" box="[629,797,493,519]" pageId="11" pageNumber="89">Jaffna District</collectingMunicipality>
|
||
,
|
||
<location id="E510603DA707D66B7A176EC0DC824035" LSID="urn:lsid:plazi:treatment:686687F0A706D66E7B4D6E25DBA240B7:E510603DA707D66B7A176EC0DC824035" box="[807,946,493,519]" country="Sri Lanka" municipality="Jaffna District" name="Mandaitivu" pageId="11" pageNumber="89" stateProvince="Northern Province">Mandaitivu</location>
|
||
; 09°61′67″ N, 79°99′22″ E;
|
||
<elevation id="6BE2D1D5A707D66B7DC86EC3DA404035" box="[1272,1392,493,520]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="89" unit="m" value="12.0">
|
||
<quantity id="27379B03A707D66B7DC86EC3DA02403A" box="[1272,1330,494,520]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="89" unit="m" value="12.0">12 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<date id="94711026A707D66B798D6D3CDE654019" box="[189,341,529,555]" pageId="11" pageNumber="89" value="2020-01-12">
|
||
<collectingDate id="8435E9CEA707D66B798D6D3CDE654019" box="[189,341,529,555]" pageId="11" pageNumber="89" value="2020-01-12">12 Jan. 2020</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="4D3A5330A707D66B78526D3CDD374019" box="[354,519,529,555]" pageId="11" pageNumber="89">S.P. Benjamin</collectorName>
|
||
<emphasis id="D2BBEAF4A707D66B7B3E6D3CDD794019" box="[526,585,529,555]" italics="true" pageId="11" pageNumber="89">et al.</emphasis>
|
||
leg.;
|
||
<collectingMethod id="398E4EF1A707D66B7BBB6D3CDDD54019" box="[651,741,529,555]" pageId="11" pageNumber="89">beating</collectingMethod>
|
||
;
|
||
<specimenCode id="B0699E9DA707D66B7BC16D3CDCF44019" box="[753,964,529,555]" pageId="11" pageNumber="89">NIFS_SAL_1280</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="E07036E6A707D66B79DA6D7FDB0940BD" blockId="11.[189,1081,594,656]" pageId="11" pageNumber="89">
|
||
<materialsCitation id="50A73CBBA707D66B79DA6D7FDB0440BD" ID-GBIF-Occurrence="3762961332" collectingDate="2020-01-12" collectingMethod="beating" collectionCode="NIFS" collectorName="S. P. Benjamin" country="Sri Lanka" elevation="12" location="SRI LANKA" municipality="Jaffna District" pageId="11" pageNumber="89" specimenCode="NIFS_SAL_1281" specimenCount="1" specimenCount-female="1" stateProvince="Northern" typeStatus="paratype">
|
||
<emphasis id="D2BBEAF4A707D66B79DA6D7FDE6C405E" bold="true" box="[234,348,594,620]" pageId="11" pageNumber="89">
|
||
<typeStatus id="3F748844A707D66B79DA6D7FDE6C405E" box="[234,348,594,620]" pageId="11" pageNumber="89" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="98D87676A707D66B798D6D58DE6740BD" box="[189,343,629,655]" name="Sri Lanka" pageId="11" pageNumber="89">SRI LANKA</collectingCountry>
|
||
•
|
||
<specimenCount id="F6C9FD6FA707D66B78406D58DEAD40A2" box="[368,413,629,656]" pageId="11" pageNumber="89" type="female">1 ♀</specimenCount>
|
||
; same collection data as for holotype;
|
||
<specimenCode id="B0699E9DA707D66B7A526D58DB0440BD" box="[866,1076,629,655]" pageId="11" pageNumber="89">NIFS_SAL_1281</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="E07036E6A707D66B79DA6D9BDE9640E2" blockId="11.[188,1399,694,862]" box="[234,422,694,720]" pageId="11" pageNumber="89">
|
||
<emphasis id="D2BBEAF4A707D66B79DA6D9BDE9640E2" bold="true" box="[234,422,694,720]" pageId="11" pageNumber="89">Other material</emphasis>
|
||
</paragraph>
|
||
<paragraph id="E07036E6A707D66B798D6DF7DECA416C" blockId="11.[188,1399,694,862]" pageId="11" pageNumber="89">
|
||
<materialsCitation id="50A73CBBA707D66B798D6DF7DE7D4125" ID-GBIF-Occurrence="3762961306" collectingDate="2016-10-20" collectingDateMax="2016-10-22" collectingDateMin="2016-10-20" collectingMethod="beating" collectionCode="NIFS" collectorName="S. P. Benjamin" country="Sri Lanka" elevation="12" location="SRI LANKA" municipality="Jaffna District" pageId="11" pageNumber="89" specimenCode="NIFS_SAL_932 to SAL_938" specimenCount="7" specimenCount-female="5" specimenCount-male="2" stateProvince="Northern">
|
||
<collectingCountry id="98D87676A707D66B798D6DF7DE6840C6" box="[189,344,730,756]" name="Sri Lanka" pageId="11" pageNumber="89">SRI LANKA</collectingCountry>
|
||
•
|
||
<specimenCount id="F6C9FD6FA707D66B785E6DF7DE8140C6" box="[366,433,730,756]" pageId="11" pageNumber="89" type="male">2 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="F6C9FD6FA707D66B788D6DF7DECF40C6" box="[445,511,730,757]" pageId="11" pageNumber="89" type="female">5 ♀♀</specimenCount>
|
||
; same collection data as for holotype but
|
||
<date id="94711026A707D66B7AD46DF7DB9B40C6" box="[996,1195,730,756]" pageId="11" pageNumber="89" value="2016-10-20" valueMax="2016-10-22" valueMin="2016-10-20">
|
||
<collectingDate id="8435E9CEA707D66B7AD46DF7DB9B40C6" box="[996,1195,730,756]" pageId="11" pageNumber="89" value="2016-10-20" valueMax="2016-10-22" valueMin="2016-10-20">20–22 Oct. 2016</collectingDate>
|
||
</date>
|
||
;
|
||
<specimenCode id="B0699E9DA707D66B7D866DF7DE7D4125" pageId="11" pageNumber="89">NIFS_SAL_932 to SAL_938</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="50A73CBBA707D66B78576DD0DA174125" ID-GBIF-Occurrence="3762961317" box="[359,1319,765,792]" collectingDate="2020-01-12" collectingMethod="beating" collectionCode="NIFS" collectorName="S. P. Benjamin" country="Sri Lanka" elevation="12" location="Mandaitivu" municipality="Jaffna District" pageId="11" pageNumber="89" specimenCode="NIFS_SAL_1282 to SAL_1284" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Northern">
|
||
<specimenCount id="F6C9FD6FA707D66B78576DD0DEA7412A" box="[359,407,765,792]" pageId="11" pageNumber="89" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="F6C9FD6FA707D66B78946DD0DED8412A" box="[420,488,765,792]" pageId="11" pageNumber="89" type="female">2 ♀♀</specimenCount>
|
||
; same collection data as for holotype;
|
||
<specimenCode id="B0699E9DA707D66B7A816DD0DA174125" box="[945,1319,765,792]" pageId="11" pageNumber="89">NIFS_SAL_1282 to SAL_1284</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="50A73CBBA707D66B7C726DD0DEC5416C" ID-GBIF-Occurrence="3762961307" collectingDate="2021-01-26" collectingMethod="beating" collectionCode="NIFS" collectorName="S. P. Benjamin" country="Sri Lanka" elevation="10" latitude="9.620556" location="Jaffna District" longLatPrecision="21" longitude="79.99777" municipality="Vantharavathai" pageId="11" pageNumber="89" specimenCode="NIFS_SAL_1441" specimenCount="1" specimenCount-female="1">
|
||
<specimenCount id="F6C9FD6FA707D66B7C726DD0DA5F412A" box="[1346,1391,765,792]" pageId="11" pageNumber="89" type="female">1 ♀</specimenCount>
|
||
;
|
||
<location id="E510603DA707D66B798D6C0DDE554108" LSID="urn:lsid:plazi:treatment:686687F0A706D66E7B4D6E25DBA240B7:E510603DA707D66B798D6C0DDE554108" box="[189,357,800,826]" country="Sri Lanka" latitude="9.620556" longLatPrecision="21" longitude="79.99777" municipality="Vantharavathai" name="Jaffna District" pageId="11" pageNumber="89">Jaffna District</location>
|
||
,
|
||
<collectingMunicipality id="0014AC9CA707D66B785F6C0DDD134108" box="[367,547,800,826]" pageId="11" pageNumber="89">Vantharavathai</collectingMunicipality>
|
||
;
|
||
<geoCoordinate id="85FB5021A707D66B7B1E6C32DDF24109" box="[558,706,799,827]" degrees="09" direction="north" minutes="37" orientation="latitude" pageId="11" pageNumber="89" precision="15" seconds="14" value="9.620556">09°37′14″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="85FB5021A707D66B7BFD6C32DC6F4109" box="[717,863,799,827]" degrees="79" direction="east" minutes="59" orientation="longitude" pageId="11" pageNumber="89" precision="15" seconds="52" value="79.99777">79°59′52″ E</geoCoordinate>
|
||
;
|
||
<elevation id="6BE2D1D5A707D66B7A5A6C0DDCDB4108" box="[874,1003,800,827]" metricMagnitude="0" metricUnit="m" metricValue="9.5" pageId="11" pageNumber="89" unit="m" value="9.5">
|
||
<quantity id="27379B03A707D66B7A5A6C0DDC9C4109" box="[874,940,800,827]" metricMagnitude="0" metricUnit="m" metricValue="9.5" pageId="11" pageNumber="89" unit="m" value="9.5">9.5 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<date id="94711026A707D66B7AC76C0DDBBD4108" box="[1015,1165,800,827]" pageId="11" pageNumber="89" value="2021-01-26">
|
||
<collectingDate id="8435E9CEA707D66B7AC76C0DDBBD4108" box="[1015,1165,800,827]" pageId="11" pageNumber="89" value="2021-01-26">26 Jan. 2021</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="4D3A5330A707D66B7DA86C0DDA0D4108" box="[1176,1341,800,826]" pageId="11" pageNumber="89">S.P. Benjamin</collectorName>
|
||
leg.;
|
||
<collectingMethod id="398E4EF1A707D66B798D6C6EDE27416F" box="[189,279,835,861]" pageId="11" pageNumber="89">beating</collectingMethod>
|
||
;
|
||
<specimenCode id="B0699E9DA707D66B78136C69DEC5416C" box="[291,501,836,862]" pageId="11" pageNumber="89">NIFS_SAL_1441</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A8D5656DA707D66E798D6CA8DB8D43C3" lastPageId="14" lastPageNumber="92" pageId="11" pageNumber="89" type="description">
|
||
<paragraph id="E07036E6A707D66B798D6CA8DE7E41AD" blockId="11.[189,334,901,927]" box="[189,334,901,927]" pageId="11" pageNumber="89">
|
||
<heading id="BB38818AA707D66B798D6CA8DE7E41AD" bold="true" box="[189,334,901,927]" fontSize="11" level="3" pageId="11" pageNumber="89" reason="3">
|
||
<emphasis id="D2BBEAF4A707D66B798D6CA8DE7E41AD" bold="true" box="[189,334,901,927]" pageId="11" pageNumber="89">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="E07036E6A707D66B79DA6C99DE1A41FC" blockId="11.[189,1399,948,1115]" box="[234,298,948,974]" pageId="11" pageNumber="89">
|
||
<emphasis id="D2BBEAF4A707D66B79DA6C99DE1A41FC" bold="true" box="[234,298,948,974]" pageId="11" pageNumber="89">Male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="E07036E6A707D66B798D6CFADA214668" blockId="11.[189,1399,948,1115]" pageId="11" pageNumber="89">MEASUREMENTS. TL 4.5, PL 1.92, PW at PLEs 1.5, AL 1.83, AW 1.32. Eye field: diameter of AME 0.4; ALE 0.26; PME 0.06; PLE 0.12; PME–PME 1.34; PLE–PLE 1.3; ALE–PME 0.22; ALE–PLE 0.54. Leg I: Tr 1.02, Fm 0.46, Pt 1, Tb 0.64, Mt 0.5; Leg II: Tr 0.9, Fm 0.46, Pt 0.7, Tb 0.6, Mt 0.52; Leg III: Tr 1.08, Fm 0.46, Pt 0.88, Tb 0.82, Mt 0.44; Leg IV: Tr 0.9, Fm 0.84, Pt 0.4, Tb 0.72, Mt 0.42.</paragraph>
|
||
<paragraph id="E07036E6A707D66B798D6BAFDDE047EA" blockId="11.[188,1399,1154,1496]" pageId="11" pageNumber="89">
|
||
COLOR AND BODY (live spiders). Carapace black, sparsely covered with black hairs; lateral margins sparsely covered with white hairs; posterior region black (
|
||
<figureCitation id="78F42A63A707D66B7A5A6B88DB70468D" box="[874,1088,1189,1216]" captionStart="Fig 5" captionStartId="10.[189,230,1962,1988]" captionTargetBox="[189,1398,684,1918]" captionTargetId="figure-104@10.[189,1398,684,1918]" captionTargetPageId="10" captionText="Fig 5. Photographs of live Carrhotus atratus sp. nov. A–B, E–F, I. Males. C–D, G–H. Females. A–H from Mandaitivu, I from Hiyare." figureDoi="http://doi.org/10.5281/zenodo.6518317" httpUri="https://zenodo.org/record/6518317/files/figure.png" pageId="11" pageNumber="89">Fig. 5A–B, E–F, I</figureCitation>
|
||
). Clypeal region blackish; eyes surrounded by blackish orbital setae. Chelicerae black, curved outer margins, excavated inner margins; two promarginal and one retromarginal teeth. Sternum oval, blackish; labium and maxillae blackish brown, paler outer margins. Abdomen ovoid, sparsely covered with black hairs; anterior margin sparsely covered by white hairs; no prominent pattern present on posterior region, beige pairs of transverse lines covered with white hairs on mid dorsum. Venter blackish with a very broad dark black median region covered with two thin greyish longitudinal lines. Spinnerets blackish, covered with a patch of black hairs dorsally (
|
||
<figureCitation id="78F42A63A707D66B7B2D6AB6DDAF4784" box="[541,671,1435,1462]" captionStart="Fig 6" captionStartId="12.[189,230,1807,1833]" captionTargetBox="[189,1398,269,1754]" captionTargetId="figure-11@12.[189,1398,269,1754]" captionTargetPageId="12" captionText="Fig 6. Carrhotus atratus sp. nov. A–B. Male habitus. A. Dorsal view. B. Ventral view. C–D. Female habitus.C. Dorsal view. D. Ventral view. E–F. Palp. E. Ventral view. F. Retrolateral view. G. Epigynum. Abbreviations: E = embolus; PLP = posterior lateral protrusion; RTA = retrolateral tibial apophysis. Scale bars: A–D = 2 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6518321" httpUri="https://zenodo.org/record/6518321/files/figure.png" pageId="11" pageNumber="89">Fig. 6A–B</figureCitation>
|
||
). Leg I robust; femora I-IV black dorsally; patellae and tibiae black; tarsi and metatarsi of all legs blackish.
|
||
</paragraph>
|
||
<paragraph id="E07036E6A707D66B798D692DDDF2440F" blockId="11.[189,1399,1536,1597]" pageId="11" pageNumber="89">
|
||
PALP. Dark black colour and embolus long, bow-shaped and tip directed anticlockwise, its base visibly separated from tegulum (
|
||
<figureCitation id="78F42A63A707D66B78D4690EDD50440F" box="[484,608,1571,1597]" captionStart="Fig 6" captionStartId="12.[189,230,1807,1833]" captionTargetBox="[189,1398,269,1754]" captionTargetId="figure-11@12.[189,1398,269,1754]" captionTargetPageId="12" captionText="Fig 6. Carrhotus atratus sp. nov. A–B. Male habitus. A. Dorsal view. B. Ventral view. C–D. Female habitus.C. Dorsal view. D. Ventral view. E–F. Palp. E. Ventral view. F. Retrolateral view. G. Epigynum. Abbreviations: E = embolus; PLP = posterior lateral protrusion; RTA = retrolateral tibial apophysis. Scale bars: A–D = 2 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6518321" httpUri="https://zenodo.org/record/6518321/files/figure.png" pageId="11" pageNumber="89">Figs 6E–F</figureCitation>
|
||
,
|
||
<figureCitation id="78F42A63A707D66B7B5B690EDD86440F" box="[619,694,1571,1597]" captionStart="Fig 7" captionStartId="13.[189,230,1829,1855]" captionTargetBox="[207,1392,281,1766]" captionTargetId="figure-15@13.[189,1398,266,1776]" captionTargetPageId="13" captionText="Fig 7. Carrhotus atratus sp. nov. A–B. Palp. A. Ventral view. B. Retrolateral view. C. Epigynum ventral view. D. Vulva dorsal view. Abbreviations: AG = accessory gland; CO = copulatory opening; E = embolus; FD = fertilization duct; PLP = posterior lateral protrusion; RTA = retrolateral tibial apophysis; S = spermatheca; TE = tegulum. Scale bars: A–B = 0.2 mm; C–D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6518323" httpUri="https://zenodo.org/record/6518323/files/figure.png" pageId="11" pageNumber="89">7A–B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E07036E6A707D66B79DA6948DE75444D" blockId="11.[189,1399,1637,1804]" box="[234,325,1637,1663]" pageId="11" pageNumber="89">
|
||
<emphasis id="D2BBEAF4A707D66B79DA6948DE75444D" bold="true" box="[234,325,1637,1663]" pageId="11" pageNumber="89">Female</emphasis>
|
||
</paragraph>
|
||
<paragraph id="E07036E6A707D66B798D69A5DBED453E" blockId="11.[189,1399,1637,1804]" pageId="11" pageNumber="89">MEASUREMENTS. TL 5.75, PL 2.04, PW at PLEs 1.74, AL 2.7, AW 2.22. Eye field: diameter of AME 0.6; ALE 0.2; PME 0.1; PLE 0.2; PME–PME 1.4; PLE–PLE 1.54; ALE–PME 0.32; ALE–PLE 0.72. Leg I: Tr 0.8, Fm 0.46, Pt 0.7, Tb 0.56, Mt 0.44; Leg II: Tr 1.06, Fm 0.34, Pt 0.46, Tb 0.62, Mt 0.52; Leg III: Tr 0.72, Fm 0.46, Pt 0.9, Tb 0.46, Mt 0.54; Leg IV: Tr 0.7, Fm 0.4, Pt 0.8, Tb 0.4, Mt 0.5.</paragraph>
|
||
<paragraph id="E07036E6A707D66E798D681EDD664358" blockId="11.[188,1399,1843,2010]" lastBlockId="14.[189,1400,265,362]" lastPageId="14" lastPageNumber="92" pageId="11" pageNumber="89">
|
||
COLOR AND BODY (live spiders). Same as in male except for the following. Clypeal region covered with iridescent hairs providing a metallic sheen; lateral margins of carapace lined by a thin line of white hairs; posterior region black (
|
||
<figureCitation id="78F42A63A707D66B78F96854DDA345A1" box="[457,659,1913,1940]" captionStart="Fig 5" captionStartId="10.[189,230,1962,1988]" captionTargetBox="[189,1398,684,1918]" captionTargetId="figure-104@10.[189,1398,684,1918]" captionTargetPageId="10" captionText="Fig 5. Photographs of live Carrhotus atratus sp. nov. A–B, E–F, I. Males. C–D, G–H. Females. A–H from Mandaitivu, I from Hiyare." figureDoi="http://doi.org/10.5281/zenodo.6518317" httpUri="https://zenodo.org/record/6518317/files/figure.png" pageId="11" pageNumber="89">Fig. 5C–D, G–H</figureCitation>
|
||
). ALEs and PLEs surrounded by blackish orbital setae. Sternum oval, brownish; labium and maxillae reddish-brown, with paler outer margins. Abdomen ovoid, densely covered with rusty brown hairs; anterior margin covered with a thin line of white hairs; pattern present on black background with pair of two white spots anteriorly and pairs of transverse stripes following posteriorly. Mid-dorsum covered with scales of metallic sheen; venter yellowish, with a broad dark brown median region (
|
||
<figureCitation id="78F42A63A702D66E78F86E62DD7A4358" box="[456,586,335,362]" captionStart="Fig 6" captionStartId="12.[189,230,1807,1833]" captionTargetBox="[189,1398,269,1754]" captionTargetId="figure-11@12.[189,1398,269,1754]" captionTargetPageId="12" captionText="Fig 6. Carrhotus atratus sp. nov. A–B. Male habitus. A. Dorsal view. B. Ventral view. C–D. Female habitus.C. Dorsal view. D. Ventral view. E–F. Palp. E. Ventral view. F. Retrolateral view. G. Epigynum. Abbreviations: E = embolus; PLP = posterior lateral protrusion; RTA = retrolateral tibial apophysis. Scale bars: A–D = 2 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6518321" httpUri="https://zenodo.org/record/6518321/files/figure.png" pageId="14" pageNumber="92">Fig. 6C–D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="B4B0666EA700D66C798D6822DC0D45A1" ID-DOI="http://doi.org/10.5281/zenodo.6518321" ID-Zenodo-Dep="6518321" httpUri="https://zenodo.org/record/6518321/files/figure.png" pageId="12" pageNumber="90" startId="12.[189,230,1807,1833]" targetBox="[189,1398,269,1754]" targetPageId="12">
|
||
<paragraph id="E07036E6A700D66C798D6822DC0D45A1" blockId="12.[188,1399,1807,1939]" pageId="12" pageNumber="90">
|
||
<emphasis id="D2BBEAF4A700D66C798D6822DE35451B" bold="true" box="[189,261,1807,1833]" pageId="12" pageNumber="90">Fig 6.</emphasis>
|
||
<taxonomicName id="27CF4D65A700D66C783F6822DED3451B" authority="Satkunanathan & Benjamin, 2022" authorityName="Satkunanathan & Benjamin" authorityYear="2022" box="[271,483,1807,1833]" class="Arachnida" family="Salticidae" genus="Carrhotus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="90" phylum="Arthropoda" rank="species" species="atratus" status="sp. nov.">
|
||
<emphasis id="D2BBEAF4A700D66C783F6822DED3451B" box="[271,483,1807,1833]" italics="true" pageId="12" pageNumber="90">Carrhotus atratus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C988578FA700D66C78DF683DDD7C4518" box="[495,588,1808,1834]" pageId="12" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="D2BBEAF4A700D66C7B656822DDBA451B" bold="true" box="[597,650,1807,1833]" pageId="12" pageNumber="90">A–B</emphasis>
|
||
. Male habitus.
|
||
<emphasis id="D2BBEAF4A700D66C7A796822DC6C451B" bold="true" box="[841,860,1807,1833]" pageId="12" pageNumber="90">A</emphasis>
|
||
. Dorsal view.
|
||
<emphasis id="D2BBEAF4A700D66C7D3E6822DB2F451B" bold="true" box="[1038,1055,1807,1833]" pageId="12" pageNumber="90">B</emphasis>
|
||
. Ventral view.
|
||
<emphasis id="D2BBEAF4A700D66C7DE86822DA20451B" bold="true" box="[1240,1296,1807,1833]" pageId="12" pageNumber="90">C–D</emphasis>
|
||
. Female habitus.
|
||
<emphasis id="D2BBEAF4A700D66C782F681FDE04457E" bold="true" box="[287,308,1842,1868]" pageId="12" pageNumber="90">C</emphasis>
|
||
. Dorsal view.
|
||
<emphasis id="D2BBEAF4A700D66C78E8681FDEDD457E" bold="true" box="[472,493,1842,1868]" pageId="12" pageNumber="90">D</emphasis>
|
||
. Ventral view.
|
||
<emphasis id="D2BBEAF4A700D66C7BA9681FDDFD457E" bold="true" box="[665,717,1842,1868]" pageId="12" pageNumber="90">E–F</emphasis>
|
||
. Palp.
|
||
<emphasis id="D2BBEAF4A700D66C7A2C681FDC00457E" bold="true" box="[796,816,1842,1868]" pageId="12" pageNumber="90">E</emphasis>
|
||
. Ventral view.
|
||
<emphasis id="D2BBEAF4A700D66C7AEB681FDCDD457E" bold="true" box="[987,1005,1842,1868]" pageId="12" pageNumber="90">F</emphasis>
|
||
. Retrolateral view.
|
||
<emphasis id="D2BBEAF4A700D66C7DFF681FDBD6457E" bold="true" box="[1231,1254,1842,1868]" pageId="12" pageNumber="90">G</emphasis>
|
||
. Epigynum. Abbreviations: E = embolus; PLP = posterior lateral protrusion; RTA = retrolateral tibial apophysis. Scale bars: A–D = 2 mm; E–F = 0.2 mm; G = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B4B0666EA701D66D798D6808DB0D459B" ID-DOI="http://doi.org/10.5281/zenodo.6518323" ID-Zenodo-Dep="6518323" httpUri="https://zenodo.org/record/6518323/files/figure.png" pageId="13" pageNumber="91" startId="13.[189,230,1829,1855]" targetBox="[207,1392,281,1766]" targetPageId="13">
|
||
<paragraph id="E07036E6A701D66D798D6808DB0D459B" blockId="13.[188,1399,1829,1961]" pageId="13" pageNumber="91">
|
||
<emphasis id="D2BBEAF4A701D66D798D6808DE32450D" bold="true" box="[189,258,1829,1855]" pageId="13" pageNumber="91">Fig 7.</emphasis>
|
||
<taxonomicName id="27CF4D65A701D66D78396808DEEA450D" authority="Satkunanathan & Benjamin, 2022" authorityName="Satkunanathan & Benjamin" authorityYear="2022" box="[265,474,1829,1855]" class="Arachnida" family="Salticidae" genus="Carrhotus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="91" phylum="Arthropoda" rank="species" species="atratus" status="sp. nov.">
|
||
<emphasis id="D2BBEAF4A701D66D78396808DEEA450D" box="[265,474,1829,1855]" italics="true" pageId="13" pageNumber="91">Carrhotus atratus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C988578FA701D66D78D1680BDD0B4572" box="[481,571,1830,1856]" pageId="13" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="D2BBEAF4A701D66D7B706808DD47450D" bold="true" box="[576,631,1829,1855]" pageId="13" pageNumber="91">A–B</emphasis>
|
||
. Palp.
|
||
<emphasis id="D2BBEAF4A701D66D7BF76808DDEC450D" bold="true" box="[711,732,1829,1855]" pageId="13" pageNumber="91">A</emphasis>
|
||
. Ventral view.
|
||
<emphasis id="D2BBEAF4A701D66D7AB96808DCAD450D" bold="true" box="[905,925,1829,1855]" pageId="13" pageNumber="91">B</emphasis>
|
||
. Retrolateral view.
|
||
<emphasis id="D2BBEAF4A701D66D7DB06808DBA5450D" bold="true" box="[1152,1173,1829,1855]" pageId="13" pageNumber="91">C</emphasis>
|
||
. Epigynum ventral view.
|
||
<emphasis id="D2BBEAF4A701D66D78376865DE2C4550" bold="true" box="[263,284,1864,1890]" pageId="13" pageNumber="91">D</emphasis>
|
||
. Vulva dorsal view. Abbreviations: AG = accessory gland; CO = copulatory opening; E = embolus; FD = fertilization duct; PLP = posterior lateral protrusion; RTA = retrolateral tibial apophysis; S = spermatheca; TE = tegulum. Scale bars: A–B = 0.2 mm; C–D = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E07036E6A702D66E798D6EBCDB8D43C3" blockId="14.[188,1399,401,498]" pageId="14" pageNumber="92">
|
||
EPIGYNUM. Epigyne with a pair of copulatory ducts with a loop or flap-like structure arising from elongated pear-shaped spermathecae, whose latter part is narrowing. Accessory glands open at 3 o’clock position at narrowing anterior region of lateral walls of spermathecae (
|
||
<figureCitation id="78F42A63A702D66E7AC86EFADB6943C3" box="[1016,1113,471,497]" captionStart="Fig 6" captionStartId="12.[189,230,1807,1833]" captionTargetBox="[189,1398,269,1754]" captionTargetId="figure-11@12.[189,1398,269,1754]" captionTargetPageId="12" captionText="Fig 6. Carrhotus atratus sp. nov. A–B. Male habitus. A. Dorsal view. B. Ventral view. C–D. Female habitus.C. Dorsal view. D. Ventral view. E–F. Palp. E. Ventral view. F. Retrolateral view. G. Epigynum. Abbreviations: E = embolus; PLP = posterior lateral protrusion; RTA = retrolateral tibial apophysis. Scale bars: A–D = 2 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6518321" httpUri="https://zenodo.org/record/6518321/files/figure.png" pageId="14" pageNumber="92">Figs 6G</figureCitation>
|
||
,
|
||
<figureCitation id="78F42A63A702D66E7D566EFADB8143C3" box="[1126,1201,471,497]" captionStart="Fig 7" captionStartId="13.[189,230,1829,1855]" captionTargetBox="[207,1392,281,1766]" captionTargetId="figure-15@13.[189,1398,266,1776]" captionTargetPageId="13" captionText="Fig 7. Carrhotus atratus sp. nov. A–B. Palp. A. Ventral view. B. Retrolateral view. C. Epigynum ventral view. D. Vulva dorsal view. Abbreviations: AG = accessory gland; CO = copulatory opening; E = embolus; FD = fertilization duct; PLP = posterior lateral protrusion; RTA = retrolateral tibial apophysis; S = spermatheca; TE = tegulum. Scale bars: A–B = 0.2 mm; C–D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6518323" httpUri="https://zenodo.org/record/6518323/files/figure.png" pageId="14" pageNumber="92">7C–D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A8D5656DA702D66E798D6D35DBA240B7" pageId="14" pageNumber="92" type="distribution">
|
||
<paragraph id="E07036E6A702D66E798D6D35DEDE4000" blockId="14.[189,494,536,563]" box="[189,494,536,563]" pageId="14" pageNumber="92">
|
||
<heading id="BB38818AA702D66E798D6D35DEDE4000" bold="true" box="[189,494,536,563]" fontSize="11" level="3" pageId="14" pageNumber="92" reason="3">
|
||
<emphasis id="D2BBEAF4A702D66E798D6D35DEDE4000" bold="true" box="[189,494,536,563]" pageId="14" pageNumber="92">Distribution and habitat</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="E07036E6A702D66E798D6D6ADBA240B7" blockId="14.[189,1398,583,645]" pageId="14" pageNumber="92">
|
||
This species occurs in the mangrove forest, in the arid zone and lowland secondary rainforest of
|
||
<collectingCountry id="98D87676A702D66E7C646D6ADE3640B6" name="Sri Lanka" pageId="14" pageNumber="92">Sri Lanka</collectingCountry>
|
||
(
|
||
<figureCitation id="78F42A63A702D66E78266D47DE5D40B6" box="[278,365,618,644]" captionStart="Fig 17" captionStartId="28.[189,230,1982,2008]" captionTargetBox="[189,1382,719,1923]" captionTargetId="figure-115@28.[189,1397,718,1944]" captionTargetPageId="28" captionText="Fig 17. Distribution of Carrhotus spp. of Sri Lanka." figureDoi="http://doi.org/10.5281/zenodo.6518352" httpUri="https://zenodo.org/record/6518352/files/figure.png" pageId="14" pageNumber="92">Fig. 17</figureCitation>
|
||
). Specimens were collected by beating foliage up to a height of
|
||
<specimenCount id="F6C9FD6FA702D66E7D6E6D47DBA240B7" box="[1118,1170,618,645]" pageId="14" pageNumber="92" type="generic">
|
||
<quantity id="27379B03A702D66E7D6E6D47DBBE40B7" box="[1118,1166,618,645]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="92" unit="m" value="2.0">2 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |