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<document ID-DOI="http://doi.org/10.5281/zenodo.5522467" ID-GBIF-Dataset="3a7a0350-3c58-40e6-8648-fe6e019bbe54" ID-GBIF-Taxon="157389655" ID-Zenodo-Dep="5522467" checkinTime="1559571790758" checkinUser="jeremy" docAuthor="Kormondy, E. J." docDate="1959" docId="03FD7535FFC4EF3CEBDBEE76F9B3FB0A" docLanguage="en" docName="Kormondy - 1959 - iLestes tikalusi, n. sp ABBYY.pdf.imf" docOrigin="The Ohio Journal of Science 59" docStyle="DocumentStyle{}" docTitle="Hetaerina pilula Calvert 1901" docType="treatment" docVersion="7" lastPageId="3" lastPageNumber="307" masterDocId="FFC40D4DFFC6EF3FEB21E867FFD2FFD0" masterDocTitle="Lestes tikalus, N. Sp. and other Odonata from Guatemala" masterLastPageNumber="312" masterPageNumber="305" pageId="2" pageNumber="306" updateTime="1632328732606" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Lestes tikalus, N. Sp. and other Odonata from Guatemala</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kormondy, E. J.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>The Ohio Journal of Science</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>1959</mods:date>
<mods:detail type="volume">
<mods:number>59</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>305</mods:start>
<mods:end>312</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5281/zenodo.3237708</mods:identifier>
<mods:identifier type="GBIF-Dataset">3a7a0350-3c58-40e6-8648-fe6e019bbe54</mods:identifier>
<mods:identifier type="Zenodo-Dep">3237708</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5522467" ID-GBIF-Taxon="157389655" ID-Zenodo-Dep="5522467" LSID="urn:lsid:plazi:treatment:03FD7535FFC4EF3CEBDBEE76F9B3FB0A" httpUri="http://treatment.plazi.org/id/03FD7535FFC4EF3CEBDBEE76F9B3FB0A" lastPageId="3" lastPageNumber="307" pageId="2" pageNumber="306">
<subSubSection pageId="2" pageNumber="306" type="nomenclature">
<paragraph blockId="2.[250,1753,1162,2633]" pageId="2" pageNumber="306">
<taxonomicName authority="Calvert" authorityName="Calvert" authorityYear="1901" box="[250,644,1553,1587]" class="Insecta" family="Calopterygidae" genus="Hetaerina" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="306" phylum="Arthropoda" rank="species" species="pilula">
<emphasis box="[250,500,1553,1587]" italics="true" pageId="2" pageNumber="306">Hetaerina pilula</emphasis>
Calvert.
</taxonomicName>
<materialsCitation ID-GBIF-Occurrence="2248043285" collectingDate="1956-04-16" collectionCode="University of Michigan Museum of Zoology" collectorName="Theodore H. Hubbell &amp; Irving J. Cantrall" country="Guatemala" location="Poptun in small clearing in gallery forest along large pool in Rio San Pedro" pageId="2" pageNumber="306" specimenCount="2" specimenCount-male="2">
<collectingRegion box="[683,796,1553,1587]" pageId="2" pageNumber="306">Poptun</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:03FD7535FFC4EF3CEBDBEE76F9B3FB0A:8E8B92F8FFC4EF3DE814EE76FE10F98B" country="Guatemala" name="in small clearing in gallery forest along large pool in Rio San Pedro" pageId="2" pageNumber="306" stateProvince="Poptun">in small clearing in gallery forest along large pool in Rio San Pedro</location>
, 8:
<emphasis bold="true" box="[495,536,1593,1627]" pageId="2" pageNumber="306">30</emphasis>
to 10
<emphasis box="[650,693,1598,1625]" italics="true" pageId="2" pageNumber="306">pm</emphasis>
,
<collectingDate box="[717,848,1593,1627]" pageId="2" pageNumber="306" value="2016-04">
<date box="[717,848,1593,1627]" pageId="2" pageNumber="306" value="2016-04">April 16</date>
</collectingDate>
,
<collectorName box="[872,1045,1593,1627]" pageId="2" pageNumber="306">THH+IJC</collectorName>
(178),
<specimenCount box="[1154,1208,1593,1627]" pageId="2" pageNumber="306" type="male">
<emphasis bold="true" box="[1154,1174,1593,1627]" pageId="2" pageNumber="306">2</emphasis>
<emphasis box="[1173,1208,1593,1627]" italics="true" pageId="2" pageNumber="306">♂</emphasis>
</specimenCount>
. According to the collectors, the river is a succession of deep (to
<quantity box="[781,847,1634,1668]" metricMagnitude="0" metricUnit="m" metricValue="3.048" pageId="2" pageNumber="306" unit="ft" value="10.0">10 ft</quantity>
) and clear pools with narrower and swifter reaches, some with rapids of small size over limestone rocks
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[352,1596,1718,1752]" pageId="2" pageNumber="306" type="discussion">
<paragraph blockId="2.[250,1753,1162,2633]" box="[352,1596,1718,1752]" pageId="2" pageNumber="306">This is apparently the first record of this species from the Yucatan Peninsula.</paragraph>
</subSubSection>
<paragraph blockId="2.[250,1753,1162,2633]" pageId="2" pageNumber="306">
<taxonomicName authority="Calvert" authorityName="Calvert" authorityYear="1901" box="[251,738,1759,1793]" class="Insecta" family="Calopterygidae" genus="Hetaerina" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="306" phylum="Arthropoda" rank="species" species="pilula">
<emphasis box="[251,513,1759,1793]" italics="true" pageId="2" pageNumber="306">Hetaerina pilula,</emphasis>
Plesiallotype
</taxonomicName>
Female (fig. 3). A female, collected with the two males of
<emphasis box="[285,381,1800,1834]" italics="true" pageId="2" pageNumber="306">
<taxonomicName box="[285,375,1800,1834]" class="Insecta" family="Calopterygidae" genus="Hetaerina" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="306" phylum="Arthropoda" rank="species" species="pilula">pilula</taxonomicName>
,
</emphasis>
was initially identified as
<taxonomicName authorityName="Selys" authorityYear="1853" box="[806,1001,1800,1834]" class="Insecta" family="Calopterygidae" genus="Hetaerina" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="306" phylum="Arthropoda" rank="species" species="macropus">
<emphasis box="[806,1001,1800,1834]" italics="true" pageId="2" pageNumber="306">H. macropus</emphasis>
</taxonomicName>
according to
<bibRefCitation author="Calvert, P. P." box="[1220,1469,1800,1834]" pageId="2" pageNumber="306" pagination="17 - 420" refId="ref6219" refString="Calvert, P. P. 1901 - 1908 a. Odonata, in Biologia Centrali-Americana. xxx + 17 - 420 pp." title="Odonata" type="book chapter" volumeTitle="Biologia Centrali-Americana" year="1901">Calvert’s (1901)</bibRefCitation>
key. It differed from
<taxonomicName authorityName="Selys" authorityYear="1853" box="[370,512,1842,1876]" class="Insecta" family="Calopterygidae" genus="Hetaerina" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="306" phylum="Arthropoda" rank="species" species="macropus">
<emphasis box="[370,512,1842,1876]" italics="true" pageId="2" pageNumber="306">macropus</emphasis>
</taxonomicName>
in several characters, notably in the presence of two spines on the dorsoapical margin of abdominal segment 10, one on each side. These spines are shorter and more slender than the spine which continues the middorsal carina. Comparison of the female with the description of the male of
<taxonomicName box="[601,850,1967,2001]" class="Insecta" family="Calopterygidae" genus="Hetaerina" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="306" phylum="Arthropoda" rank="species" species="pilula">
<emphasis box="[601,850,1967,2001]" italics="true" pageId="2" pageNumber="306">Hetaerina pilula</emphasis>
</taxonomicName>
indicated that this was the undescribed female of that species. The specimen was sent to Calvert for an opinion; his reply (
<date box="[1370,1596,2009,2043]" pageId="2" pageNumber="306" value="1957-08-08">August 8, 1957</date>
) contains the following: “Your Guatemala female does not differ more... from the description of
<taxonomicName box="[284,372,2090,2124]" class="Insecta" family="Calopterygidae" genus="Hetaerina" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="306" phylum="Arthropoda" rank="species" species="pilula">
<emphasis box="[284,372,2090,2124]" italics="true" pageId="2" pageNumber="306">pilula</emphasis>
</taxonomicName>
male than do females of
<taxonomicName authorityName="Selys" authorityYear="1853" box="[784,926,2090,2124]" class="Insecta" family="Calopterygidae" genus="Hetaerina" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="306" phylum="Arthropoda" rank="species" species="macropus">
<emphasis box="[784,926,2090,2124]" italics="true" pageId="2" pageNumber="306">macropus</emphasis>
</taxonomicName>
from
<taxonomicName authorityName="Selys" authorityYear="1853" box="[1027,1169,2090,2124]" class="Insecta" family="Calopterygidae" genus="Hetaerina" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="306" phylum="Arthropoda" rank="species" species="macropus">
<emphasis box="[1027,1169,2090,2124]" italics="true" pageId="2" pageNumber="306">macropus</emphasis>
</taxonomicName>
males; bearing this in mind I think it would be safe to call it
<emphasis box="[696,813,2132,2166]" italics="true" pageId="2" pageNumber="306">
<taxonomicName box="[696,795,2132,2166]" class="Insecta" family="Calopterygidae" genus="Hetaerina" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="306" phylum="Arthropoda" rank="species" species="pilula">pilula</taxonomicName>
.&quot;
</emphasis>
</paragraph>
<paragraph blockId="2.[250,1753,1162,2633]" box="[350,1747,2173,2207]" pageId="2" pageNumber="306">
The female differs from the description of the male (
<bibRefCitation author="Calvert, P. P." box="[1186,1398,2173,2207]" pageId="2" pageNumber="306" pagination="17 - 420" refId="ref6219" refString="Calvert, P. P. 1901 - 1908 a. Odonata, in Biologia Centrali-Americana. xxx + 17 - 420 pp." title="Odonata" type="book chapter" volumeTitle="Biologia Centrali-Americana" year="1901">Calvert, 1901</bibRefCitation>
, pp. 33-34) as follows:
</paragraph>
<paragraph blockId="2.[250,1753,1162,2633]" pageId="2" pageNumber="306">Head with nasus with a cupreous metallic reflection, vertex metallic dark green with a coppery-yellow reflection.</paragraph>
<paragraph blockId="2.[250,1753,1162,2633]" pageId="2" pageNumber="306">Prothorax and thoracic dorsum without coppery-red reflection; a narrow metallic green band along the thoracic carina, widened posteriorly to the humeral suture; a metallic green band on each of the mesepimeron, metepisternum, and metepimeron, that of mesepimeron interrupted posteriorly at three-fourths its length; yellow humeral stripe four to five times wider than the dark brown of the same side of the thoracic dorsum.</paragraph>
<paragraph blockId="2.[250,1753,1162,2633]" lastBlockId="3.[179,1648,297,1242]" lastPageId="3" lastPageNumber="307" pageId="2" pageNumber="306">
Abdomen with a small dorsolateral metallic green spot on segment 1; an elongate dorsolateral green spot extending one-half the length of segment
<emphasis bold="true" box="[1268,1287,2556,2590]" pageId="2" pageNumber="306">2</emphasis>
; segment 10 with middorsal carina extended and elevated as a slender black spine; two dorsoapical spines on segment 10, one on either side of the middorsal spine, one-half as long and more slender than middorsal spine; ventrolateral apical spines (2 on left, 3 on right) and ventroapical spines, one on each side, as long as the two dorsoapical spines.
</paragraph>
<paragraph blockId="3.[179,1648,297,1242]" pageId="3" pageNumber="307">Appendages nearly as long as segment 10, conical, acute; genital valvules extend to ventral apex of tergum of segment 10, apical half of inferior margin denticulated.</paragraph>
<paragraph blockId="3.[179,1648,297,1242]" pageId="3" pageNumber="307">
Stigma obscure in color, surmounting less than one-half cell on all wings; tip of each wing unicolorous with rest of wing, not brown; coppery-red metallic reflection on longitudinal veins from Ri to lower sector of arculus and outward to beyond nodus, otherwise no coloring of the wing to represent the coloring of the base of the wing in the male; median cross veins,
<quantity box="[179,235,669,703]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="3" pageNumber="307" unit="in" value="6.0">6 in</quantity>
forewing,
<quantity box="[402,457,669,703]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="3" pageNumber="307" unit="in" value="7.0">7 in</quantity>
hindwing (vs.
<quantity box="[693,749,669,703]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="3" pageNumber="307" unit="in" value="9.0">9 in</quantity>
male); quadrilateral crossveins,
<quantity box="[1276,1331,669,703]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="3" pageNumber="307" unit="in" value="6.0">6 in</quantity>
forewing,
<quantity box="[1498,1554,669,703]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="3" pageNumber="307" unit="in" value="5.0">5 in</quantity>
hindwing (vs.
<quantity box="[340,431,711,745]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" metricValueMax="2.286" metricValueMin="1.778" pageId="3" pageNumber="307" unit="in" value="8.0" valueMax="9.0" valueMin="7.0">7-9 in</quantity>
male); antenodal crossveins,
<quantity box="[915,990,711,745]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" pageId="3" pageNumber="307" unit="in" value="23.0">23 in</quantity>
left forewing,
<quantity box="[1230,1305,711,745]" metricMagnitude="-1" metricUnit="m" metricValue="5.334" pageId="3" pageNumber="307" unit="in" value="21.0">21 in</quantity>
right forewing,
<quantity box="[1570,1645,711,745]" metricMagnitude="-1" metricUnit="m" metricValue="5.588" pageId="3" pageNumber="307" unit="in" value="22.0">22 in</quantity>
left hindwing,
<quantity box="[410,482,753,787]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" pageId="3" pageNumber="307" unit="in" value="23.0">23 in</quantity>
right hindwing (vs.
<quantity box="[812,885,753,787]" metricMagnitude="-1" metricUnit="m" metricValue="5.588" pageId="3" pageNumber="307" unit="in" value="22.0">22 in</quantity>
male).
</paragraph>
<paragraph blockId="3.[179,1648,297,1242]" box="[246,812,794,828]" pageId="3" pageNumber="307">
Abdomen,
<quantity box="[417,525,794,828]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="3" pageNumber="307" unit="mm" value="34.0">34 mm</quantity>
; hindwing,
<quantity box="[711,812,794,828]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="307" unit="mm" value="30.0">30 mm</quantity>
</paragraph>
<paragraph blockId="3.[179,1648,297,1242]" pageId="3" pageNumber="307">
<materialsCitation ID-GBIF-Occurrence="2248043327" collectingDate="1956-04-16" collectionCode="University of Michigan Museum of Zoology" collectorName="I. J. Cantrall &amp; T. H. Hubbell" country="Guatemala" location="Poptun" pageId="3" pageNumber="307" specimenCount="1" specimenCount-female="1" stateProvince="Peten" typeStatus="plesiallotype">
The plesiallotype
<specimenCount box="[541,648,836,870]" pageId="3" pageNumber="307" type="female">female</specimenCount>
, labeled, “
<collectingCountry box="[837,1015,836,870]" name="Guatemala" pageId="3" pageNumber="307" value="Guatemala">Guatemala</collectingCountry>
:
<collectingRegion box="[1040,1338,836,870]" pageId="3" pageNumber="307">Department Peten</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:03FD7535FFC4EF3CEBDBEE76F9B3FB0A:8E8B92F8FFC5EF3CEE77EB23FA15FCB6" box="[1366,1479,836,870]" country="Guatemala" name="Poptun" pageId="3" pageNumber="307" stateProvince="Department Peten">Poptün</location>
,
<collectingDate pageId="3" pageNumber="307" value="1956-04-16">
<date pageId="3" pageNumber="307" value="1956-04-16">April 16, 1956</date>
</collectingDate>
,
<collectorName box="[271,479,877,911]" pageId="3" pageNumber="307">I. J. Cantrall</collectorName>
and
<collectorName box="[560,790,877,911]" pageId="3" pageNumber="307">T. H. Hubbell</collectorName>
, #178,” is deposited in the type collection of the
<collectionCode pageId="3" pageNumber="307">University of Michigan Museum of Zoology</collectionCode>
</materialsCitation>
.
</paragraph>
<subSection pageId="3" pageNumber="307" type="key">
<subSubSection pageId="3" pageNumber="307" type="key">
<paragraph blockId="3.[179,1648,297,1242]" pageId="3" pageNumber="307">
The following modification in Calvert’s key (1901, p. 21) will permit separation of
<taxonomicName authorityName="Selys" authorityYear="1853" box="[179,320,999,1033]" class="Insecta" family="Calopterygidae" genus="Hetaerina" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="307" phylum="Arthropoda" rank="species" species="macropus">
<emphasis box="[179,320,999,1033]" italics="true" pageId="3" pageNumber="307">macropus</emphasis>
</taxonomicName>
and
<taxonomicName box="[403,491,999,1033]" class="Insecta" family="Calopterygidae" genus="Hetaerina" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="307" phylum="Arthropoda" rank="species" species="pilula">
<emphasis box="[403,491,999,1033]" italics="true" pageId="3" pageNumber="307">pilula</emphasis>
</taxonomicName>
females.
</paragraph>
<paragraph blockId="3.[179,1648,297,1242]" box="[244,966,1041,1075]" pageId="3" pageNumber="307">f. Pterostigma, if present, obscure in colour.</paragraph>
<paragraph blockId="3.[179,1648,297,1242]" pageId="3" pageNumber="307">
fi. A dorsolateral apical spine on each side of spine which continues the middorsal carina................................................................................................................ 9.
<taxonomicName box="[1541,1633,1124,1158]" class="Insecta" family="Calopterygidae" genus="Hetaerina" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="307" phylum="Arthropoda" rank="species" species="pilula">
<emphasis box="[1541,1633,1124,1158]" italics="true" pageId="3" pageNumber="307">pilula</emphasis>
</taxonomicName>
</paragraph>
<paragraph blockId="3.[179,1648,297,1242]" pageId="3" pageNumber="307">
f-2. No dorsolateral apical spine on each side of spine which continues the middorsal carina.........................................................................................................10.
<taxonomicName authorityName="Selys" authorityYear="1853" box="[1490,1633,1208,1242]" class="Insecta" family="Calopterygidae" genus="Hetaerina" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="307" phylum="Arthropoda" rank="species" species="macropus">
<emphasis box="[1490,1633,1208,1242]" italics="true" pageId="3" pageNumber="307">macropus</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
</subSection>
</treatment>
</document>
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