546 lines
63 KiB
XML
546 lines
63 KiB
XML
<document id="1A58C9D7669F808838E7C5DDBDC27D63" ID-CLB-Dataset="46750" ID-DOI="10.5281/zenodo.277759" ID-GBIF-Dataset="1f445f56-2cd0-431e-83fe-8aa55c3da513" ID-ISSN="1175-5326" ID-Zenodo-Dep="277759" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460364359420" checkinUser="plazi" docAuthor="Pulido-B, Hannier W., Andrade-C, Gonzalo, Peña, Carlos & Lamas, Gerardo" docDate="2011" docId="03BA87A87A41EF66FF51FA0CC206F9F2" docLanguage="en" docName="zt02906p051.pdf" docOrigin="Zootaxa 2906" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Euptychia cesarense Pulido, Andrade, Pena & Lamas, new species" docType="treatment" docVersion="9" lastPageNumber="49" masterDocId="FF83FFD07A40EF60FFC6FFC5C627FFF7" masterDocTitle="Two new taxa of Euptychia Hübner, 1818 (Lepidoptera: Nymphalidae: Satyrinae) from the Andes of Colombia and Peru" masterLastPageNumber="51" masterPageNumber="43" pageNumber="44" updateTime="1698295003119" updateUser="ExternalLinkService">
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<mods:title id="933EAB1681121B10796B9051B6CEA1F3">Two new taxa of Euptychia Hübner, 1818 (Lepidoptera: Nymphalidae: Satyrinae) from the Andes of Colombia and Peru</mods:title>
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<mods:namePart id="10FE19EBF90FFC0ED4270D12436AE29A">Pulido-B, Hannier W.</mods:namePart>
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<mods:namePart id="6BFE946F2B97909DEB47A22FF4CBA648">Andrade-C, Gonzalo</mods:namePart>
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<mods:namePart id="1DB9514B71B8D138E25F6CD052841325">Peña, Carlos</mods:namePart>
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<mods:namePart id="D723D0E52525CCA01A01E5165CF4D1D1">Lamas, Gerardo</mods:namePart>
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<mods:date id="41DFB0B1075E9FD7EA511D8B70B350CC">2011</mods:date>
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<treatment id="03BA87A87A41EF66FF51FA0CC206F9F2" ID-DOI="http://doi.org/10.5281/zenodo.6187284" ID-GBIF-Taxon="119410404" ID-Zenodo-Dep="6187284" LSID="urn:lsid:plazi:treatment:03BA87A87A41EF66FF51FA0CC206F9F2" httpUri="http://treatment.plazi.org/id/03BA87A87A41EF66FF51FA0CC206F9F2" lastPageId="6" lastPageNumber="49" pageId="1" pageNumber="44">
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<subSubSection id="C30965357A41EF61FF51FA0CC7FDF9BE" pageId="1" pageNumber="44" type="nomenclature">
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<paragraph id="8BAC36BE7A41EF61FF51FA0CC5EBFA15" blockId="1.[151,972,1480,1540]" box="[151,972,1480,1507]" pageId="1" pageNumber="44">
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<heading id="D0E481D27A41EF61FF51FA0CC5EBFA15" bold="true" box="[151,972,1480,1507]" fontSize="11" level="1" pageId="1" pageNumber="44" reason="1">
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<emphasis id="B967EAAC7A41EF61FF51FA0CC5EBFA15" bold="true" box="[151,972,1480,1507]" pageId="1" pageNumber="44">
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<taxonomicName id="4C134D3D7A41EF61FF51FA0CC508FA15" authority="Pulido, Andrade, Pena & Lamas" authorityName="Pulido, Andrade, Pena & Lamas" box="[151,815,1480,1507]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="44" phylum="Arthropoda" rank="species" species="cesarense" status="sp. nov.">
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<emphasis id="B967EAAC7A41EF61FF51FA0CC7A8FA14" bold="true" box="[151,399,1481,1507]" italics="true" pageId="1" pageNumber="44">Euptychia cesarense</emphasis>
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Pulido, Andrade, Peña & Lamas
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25457D77A41EF61FCFAFA0DC5EBFA15" box="[828,972,1480,1506]" pageId="1" pageNumber="44" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BAC36BE7A41EF61FF51FA29C735F9F3" blockId="1.[151,972,1480,1540]" box="[151,274,1516,1540]" pageId="1" pageNumber="44">
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(
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<figureCitation id="13282A3B7A41EF61FF59FA29C72DF9F3" box="[159,266,1516,1540]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="3.[151,250,1975,1998]" captionStartId-1="4.[151,250,1779,1802]" captionStartId-2="5.[151,250,1077,1100]" captionStartId-3="6.[151,250,663,686]" captionTargetBox-0="[302,1273,191,1942]" captionTargetBox-1="[166,1415,1149,1741]" captionTargetBox-2="[176,1426,411,1061]" captionTargetBox-3="[164,1422,179,650]" captionTargetId-0="figure@3.[263,1320,169,1962]" captionTargetId-1="figure@4.[151,1436,1135,1758]" captionTargetId-2="figure@5.[151,1436,395,1071]" captionTargetId-3="figure@6.[151,1436,159,658]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 2. Male wing venation of E. cesarense showing line and ocellus pattern, forewing above and hindwing below. (a), discal line; (b), postdiscal line; (c), submarginal ocelli; (d), submarginal line; (e) marginal line." captionText-1="FIGURE 3. Male genitalia of E. cesarense. A, lateral view; B, dorsal view; C, aedeagus (lateral view); ae: aedeagus; un: uncus; te: tegumen; ft: fultura superior; sa: saccus; va: valva; su: superuncus." captionText-2="FIGURE 4. Female genitalia of E. cesarense; pap. a: papillae anales; du. bu: ductus bursae; crp. bu: corpus bursae; sig: signa." captionText-3="FIGURE 5. Adults of E. cesarense, A. Holotype from Río de Oro (Cesar, Colombia), B. Allotype from González (Cesar, Colombia). D: Dorsal view; V: Ventral view." httpUri-0="https://zenodo.org/record/277761/files/figure.png" httpUri-1="https://zenodo.org/record/277762/files/figure.png" httpUri-2="https://zenodo.org/record/277763/files/figure.png" httpUri-3="https://zenodo.org/record/277764/files/figure.png" pageId="1" pageNumber="44">Figs. 2–5</figureCitation>
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)
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</paragraph>
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<paragraph id="8BAC36BE7A41EF61FF51F9F1C7FDF9BE" blockId="1.[151,474,1587,1610]" box="[151,474,1587,1610]" pageId="1" pageNumber="44">
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<taxonomicName id="4C134D3D7A41EF61FF51F9F1C6D9F9BD" box="[151,254,1588,1610]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis id="B967EAAC7A41EF61FF51F9F1C6D9F9BD" box="[151,254,1588,1610]" italics="true" pageId="1" pageNumber="44">Euptychia</emphasis>
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</taxonomicName>
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[n. sp.]:
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<bibRefCitation id="EF824B4F7A41EF61FE9EF9F0C7FDF9BE" author="Lamas" box="[344,474,1588,1610]" pageId="1" pageNumber="44" refString="Lamas, G. (2004) Atlas of Neotropical Lepidoptera. Checklist part 4 A. Hesperioidea-Papilionoidea. Association for tropical Lepidoptera, Florida, 439 pp." type="book" year="2004">Lamas, 2004</bibRefCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30965357A41EF65FF51F9B3C248FB58" lastPageId="5" lastPageNumber="48" pageId="1" pageNumber="44" type="description">
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<paragraph id="8BAC36BE7A41EF61FF51F9B3C555F979" blockId="1.[151,1436,1654,2002]" box="[151,882,1654,1679]" pageId="1" pageNumber="44">
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<emphasis id="B967EAAC7A41EF61FF51F9B3C7A4F978" bold="true" box="[151,387,1654,1679]" pageId="1" pageNumber="44">Description. MALE</emphasis>
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, FW length:
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<quantity id="4CEB9B5B7A41EF61FDD1F9B3C45DF979" box="[535,634,1654,1678]" metricMagnitude="-2" metricUnit="m" metricValue="1.71" pageId="1" pageNumber="44" unit="mm" value="17.1">17.1 mm</quantity>
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(n=4) (
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<figureCitation id="13282A3B7A41EF61FD09F9B3C538F979" box="[719,799,1654,1678]" captionStart="FIGURE 2" captionStartId="3.[151,250,1975,1998]" captionTargetBox="[302,1273,191,1942]" captionTargetId="figure@3.[263,1320,169,1962]" captionTargetPageId="3" captionText="FIGURE 2. Male wing venation of E. cesarense showing line and ocellus pattern, forewing above and hindwing below. (a), discal line; (b), postdiscal line; (c), submarginal ocelli; (d), submarginal line; (e) marginal line." httpUri="https://zenodo.org/record/277761/files/figure.png" pageId="1" pageNumber="44">Figs. 2</figureCitation>
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,
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||
<figureCitation id="13282A3B7A41EF61FCEDF9B3C51DF979" box="[811,826,1654,1678]" captionStart="FIGURE 3" captionStartId="4.[151,250,1779,1802]" captionTargetBox="[166,1415,1149,1741]" captionTargetId="figure@4.[151,1436,1135,1758]" captionTargetPageId="4" captionText="FIGURE 3. Male genitalia of E. cesarense. A, lateral view; B, dorsal view; C, aedeagus (lateral view); ae: aedeagus; un: uncus; te: tegumen; ft: fultura superior; sa: saccus; va: valva; su: superuncus." httpUri="https://zenodo.org/record/277762/files/figure.png" pageId="1" pageNumber="44">3</figureCitation>
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||
,
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||
<figureCitation id="13282A3B7A41EF61FC80F9B3C574F979" box="[838,851,1654,1678]" captionStart="FIGURE 5" captionStartId="6.[151,250,663,686]" captionTargetBox="[164,1422,179,650]" captionTargetId="figure@6.[151,1436,159,658]" captionTargetPageId="6" captionText="FIGURE 5. Adults of E. cesarense, A. Holotype from Río de Oro (Cesar, Colombia), B. Allotype from González (Cesar, Colombia). D: Dorsal view; V: Ventral view." httpUri="https://zenodo.org/record/277764/files/figure.png" pageId="1" pageNumber="44">5</figureCitation>
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A)
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</paragraph>
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<paragraph id="8BAC36BE7A41EF61FF01F95EC4CFF921" blockId="1.[151,1436,1654,2002]" pageId="1" pageNumber="44">
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<emphasis id="B967EAAC7A41EF61FF01F95EC737F944" bold="true" box="[199,272,1691,1715]" pageId="1" pageNumber="44">Head:</emphasis>
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Labial palpi twice the eye diameter (
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<quantity id="4CEB9B5B7A41EF61FD69F95FC52DF945" box="[687,778,1690,1714]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="44" unit="mm" value="1.5">1.5 mm</quantity>
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), with long scales (
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<quantity id="4CEB9B5B7A41EF61FC27F95FC202F945" box="[993,1061,1690,1714]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="44" unit="mm" value="3.0">3 mm</quantity>
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), antennae yellowish, total length (
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<quantity id="4CEB9B5B7A41EF61FF59F97BC721F921" box="[159,262,1726,1750]" metricMagnitude="-3" metricUnit="m" metricValue="9.03" pageId="1" pageNumber="44" unit="mm" value="9.03">9.03 mm</quantity>
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) a little less than half of costal vein length.
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</paragraph>
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<paragraph id="8BAC36BE7A41EF61FF01F926C750F8E9" blockId="1.[151,1436,1654,2002]" pageId="1" pageNumber="44">
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<emphasis id="B967EAAC7A41EF61FF01F926C70EF90C" bold="true" box="[199,297,1763,1787]" pageId="1" pageNumber="44">Thorax:</emphasis>
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Dorsally blackish brown (R40, G42, B31), ventrally yellowish (R105, G92, B48), covered with dense light yellowish hair.
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</paragraph>
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<paragraph id="8BAC36BE7A41EF61FF01F8EFC2FAF8B5" blockId="1.[151,1436,1654,2002]" box="[199,1245,1834,1859]" pageId="1" pageNumber="44">
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<emphasis id="B967EAAC7A41EF61FF01F8EFC764F8B4" bold="true" box="[199,323,1834,1859]" pageId="1" pageNumber="44">Abdomen.</emphasis>
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Dorsally blackish brown (R40, G42, B31), ventrally yellowish (R105, G92, B48).
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</paragraph>
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<paragraph id="8BAC36BE7A41EF61FF01F88BC27CF825" blockId="1.[151,1436,1654,2002]" pageId="1" pageNumber="44">
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<emphasis id="B967EAAC7A41EF61FF01F88BC712F890" bold="true" box="[199,309,1870,1895]" pageId="1" pageNumber="44">Genitalia</emphasis>
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(
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<figureCitation id="13282A3B7A41EF61FE81F88BC7A9F891" box="[327,398,1870,1894]" captionStart="FIGURE 3" captionStartId="4.[151,250,1779,1802]" captionTargetBox="[166,1415,1149,1741]" captionTargetId="figure@4.[151,1436,1135,1758]" captionTargetPageId="4" captionText="FIGURE 3. Male genitalia of E. cesarense. A, lateral view; B, dorsal view; C, aedeagus (lateral view); ae: aedeagus; un: uncus; te: tegumen; ft: fultura superior; sa: saccus; va: valva; su: superuncus." httpUri="https://zenodo.org/record/277762/files/figure.png" pageId="1" pageNumber="44">Fig. 3</figureCitation>
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): Uncus has the same length (
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||
<quantity id="4CEB9B5B7A41EF61FD2AF88BC572F891" box="[748,853,1870,1894]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="1" pageNumber="44" unit="mm" value="0.48">0.48 mm</quantity>
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) of the tegumen; superuncus is long (
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||
<quantity id="4CEB9B5B7A41EF61FACDF88BC353F891" box="[1291,1396,1870,1894]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="1" pageNumber="44" unit="mm" value="0.28">0.28 mm</quantity>
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) in comparison with other species of the genus (
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<bibRefCitation id="EF824B4F7A41EF61FD66F8B7C51FF87D" author="Forster" box="[672,824,1906,1930]" pageId="1" pageNumber="44" refString="Forster, W. (1964) Beitrage zur Kenntnis der Insektenfauna Boliviens XIX. Lepidoptera III. Satyridae. Veroffentlichungen der zoologischen Staatssammlung Munchen 8, 51 - 188, pls. 27 - 35." type="journal article" year="1964">Forster 1964</bibRefCitation>
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); gnathos absent; fultura superior (ft) very elongated (
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<quantity id="4CEB9B5B7A41EF61FF58F853C72EF859" box="[158,265,1942,1966]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="1" pageNumber="44" unit="mm" value="0.36">0.36 mm</quantity>
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); valva elongated (
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||
<quantity id="4CEB9B5B7A41EF61FE1BF853C463F859" box="[477,580,1942,1966]" metricMagnitude="-3" metricUnit="m" metricValue="1.26" pageId="1" pageNumber="44" unit="mm" value="1.26">1.26 mm</quantity>
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) with pointed end and plain internal margin; saccus (
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||
<quantity id="4CEB9B5B7A41EF61FB5CF853C325F859" box="[1178,1282,1942,1966]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" pageId="1" pageNumber="44" unit="mm" value="0.39">0.39 mm</quantity>
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||
) rounded and short in ventral view; aedeagus (
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||
<quantity id="4CEB9B5B7A41EF61FDC5F87FC47CF825" box="[515,603,1978,2002]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="44" unit="mm" value="1.5">1.5 mm</quantity>
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||
) tubular and curved upwards without cornuti.
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</paragraph>
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<caption id="DF6C66367A42EF62FF51F85AC4DFF823" httpUri="https://zenodo.org/record/277760/files/figure.png" pageId="2" pageNumber="45" targetBox="[311,1335,212,1923]" targetPageId="2">
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||
<paragraph id="8BAC36BE7A42EF62FF51F85AC4DFF823" blockId="2.[151,1435,1951,2004]" pageId="2" pageNumber="45">
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||
<emphasis id="B967EAAC7A42EF62FF51F85AC735F843" bold="true" box="[151,274,1951,1974]" pageId="2" pageNumber="45">FIGURE 1.</emphasis>
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||
<taxonomicName id="4C134D3D7A42EF62FEDEF865C7B0F842" box="[280,407,1952,1973]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="cesarense">
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<emphasis id="B967EAAC7A42EF62FEDEF865C7B0F842" box="[280,407,1952,1973]" italics="true" pageId="2" pageNumber="45">E. cesarense</emphasis>
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</taxonomicName>
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distribution in the Andes. A, Río de Oro (Cesar, Colombia); B, Pozuzo (Pasco, Perú); C,
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<taxonomicName id="4C134D3D7A42EF62FADCF85AC3BCF842" authority="Pichita" authorityName="Pichita" box="[1306,1435,1951,1973]" class="Magnoliopsida" family="Convolvulaceae" genus="Mina" higherTaxonomySource="CoL" kingdom="Plantae" order="Solanales" pageId="2" pageNumber="45" phylum="Tracheophyta" rank="genus">Mina Pichita</taxonomicName>
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(Junín, Perú); D, Santa Isabel, Río Cosñipata (Cuzco, Perú).
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</paragraph>
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</caption>
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<caption id="DF6C66367A43EF63FF51F872C271F81B" httpUri="https://zenodo.org/record/277761/files/figure.png" pageId="3" pageNumber="46" targetBox="[302,1273,191,1942]" targetPageId="3">
|
||
<paragraph id="8BAC36BE7A43EF63FF51F872C271F81B" blockId="3.[151,1436,1975,2029]" pageId="3" pageNumber="46">
|
||
<emphasis id="B967EAAC7A43EF63FF51F872C733F83A" bold="true" box="[151,276,1975,1998]" pageId="3" pageNumber="46">FIGURE 2.</emphasis>
|
||
Male wing venation of
|
||
<taxonomicName id="4C134D3D7A43EF63FDC9F87CC4A8F839" box="[527,655,1977,1998]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="cesarense">
|
||
<emphasis id="B967EAAC7A43EF63FDC9F87CC4A8F839" box="[527,655,1977,1998]" italics="true" pageId="3" pageNumber="46">E. cesarense</emphasis>
|
||
</taxonomicName>
|
||
showing line and ocellus pattern, forewing above and hindwing below. (a), discal line; (b), postdiscal line; (c), submarginal ocelli; (d), submarginal line; (e) marginal line.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAC36BE7A44EF64FF01FF52C4C5FE08" blockId="4.[151,1436,151,1106]" pageId="4" pageNumber="47">
|
||
<emphasis id="B967EAAC7A44EF64FF01FF52C7ADFF47" bold="true" box="[199,394,151,176]" pageId="4" pageNumber="47">Forewing (FW).</emphasis>
|
||
Apex rounded; outer margin covered with long scales. Dorsal ground color dark brown (R130, G112, B62), darker (R71, G59, B35) and diffused discal, postdiscal marginal and submarginal lines are evident in DFW, the same as a submarginal dot in M1-M2, discal and postdiscal lines are much diffused and darker (R86, G62, B34) than the ground. VFW color light brown (R150, G129, B66); discal, postdiscal, submarginal and marginal lines reddish brown (R104, G67, B25), straight discal line extending from discal cell to 2A, the postdiscal line is branched at the top of the discal cell and it extends from R5 base to 2A, slightly undulated submarginal line from R3 to 2A, and a narrower marginal line from R4 to 2A. A single postdiscal black ocellus with a white pupil and encircled with a yellowish (R170, G140, B52) ring and an external reddish brown ring between R5 and M2, the center of the ocellus is in M2-M1, but its extent into neibouring cells may vary; two dark yellow patches between postdiscal and submarginal lines in M3-Cu1 and Cu1-Cu2.
|
||
</paragraph>
|
||
<paragraph id="8BAC36BE7A44EF64FF01FDC8C777FC68" blockId="4.[151,1436,151,1106]" pageId="4" pageNumber="47">
|
||
<emphasis id="B967EAAC7A44EF64FF01FDC8C7B7FDD1" bold="true" box="[199,400,525,550]" pageId="4" pageNumber="47">Hindwing (HW).</emphasis>
|
||
Anal apex rounded. Dorsal ground color the same as DFW, discal and postdiscal lines fade out from costa to anal margin, marginal and submarginal lines darker (R62, G56, B34) than in DFW, with a submarginal dark (R86, G62, B34) dot in M1-M2, a black ocellus with a white pupil and encircled by a yellowish (R167, R131, B43) ring in cell Cu1-Cu2. VHW ground color the same as VFW, discal, postdiscal submarginal and marginal lines the same brown color as VFW, straight discal line from costal margin to 2A-3A, postdiscal line undulated from costal margin to M2 and stays straight from M2 to 2A-3A, where it branches reaching submarginal ocellus in Cu2-2A, submarginal line with an irregular undulated pattern from costal margin to ocellus in Cu2-2A, slightly undulated marginal line, thinner than remaining lines, from Rs to 2A-3A, four black ocelli between postdiscal and submarginal lines, smallest in cell Rs-M1, largest in cells M1-M2 and Cu1-Cu2, and medium-sized in cell Cu2-2A, all of them with a white pupil and encircled by a yellowish (R170, G140, B52) ring and an external reddish brown ring.
|
||
</paragraph>
|
||
<paragraph id="8BAC36BE7A44EF64FF01FC6FC4A2FC10" blockId="4.[151,1436,151,1106]" pageId="4" pageNumber="47">
|
||
<emphasis id="B967EAAC7A44EF64FF01FC6FC718FC34" bold="true" box="[199,319,938,963]" pageId="4" pageNumber="47">FEMALE</emphasis>
|
||
: FW length:
|
||
<quantity id="4CEB9B5B7A44EF64FE13FC6FC41DFC35" box="[469,570,938,962]" metricMagnitude="-2" metricUnit="m" metricValue="1.73" pageId="4" pageNumber="47" unit="mm" value="17.3">17.3 mm</quantity>
|
||
(n=4) (
|
||
<figureCitation id="13282A3B7A44EF64FD49FC6FC4C7FC35" box="[655,736,938,962]" captionStart="FIGURE 2" captionStartId="3.[151,250,1975,1998]" captionTargetBox="[302,1273,191,1942]" captionTargetId="figure@3.[263,1320,169,1962]" captionTargetPageId="3" captionText="FIGURE 2. Male wing venation of E. cesarense showing line and ocellus pattern, forewing above and hindwing below. (a), discal line; (b), postdiscal line; (c), submarginal ocelli; (d), submarginal line; (e) marginal line." httpUri="https://zenodo.org/record/277761/files/figure.png" pageId="4" pageNumber="47">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation id="13282A3B7A44EF64FD2AFC6FC4DBFC35" box="[748,764,938,962]" captionStart="FIGURE 4" captionStartId="5.[151,250,1077,1100]" captionTargetBox="[176,1426,411,1061]" captionTargetId="figure@5.[151,1436,395,1071]" captionTargetPageId="5" captionText="FIGURE 4. Female genitalia of E. cesarense; pap. a: papillae anales; du. bu: ductus bursae; crp. bu: corpus bursae; sig: signa." httpUri="https://zenodo.org/record/277763/files/figure.png" pageId="4" pageNumber="47">4</figureCitation>
|
||
,
|
||
<figureCitation id="13282A3B7A44EF64FCCFFC6FC53FFC35" box="[777,792,938,962]" captionStart="FIGURE 5" captionStartId="6.[151,250,663,686]" captionTargetBox="[164,1422,179,650]" captionTargetId="figure@6.[151,1436,159,658]" captionTargetPageId="6" captionText="FIGURE 5. Adults of E. cesarense, A. Holotype from Río de Oro (Cesar, Colombia), B. Allotype from González (Cesar, Colombia). D: Dorsal view; V: Ventral view." httpUri="https://zenodo.org/record/277764/files/figure.png" pageId="4" pageNumber="47">5</figureCitation>
|
||
B): General color and pattern very similar to male, but in general paler and with more rounded wings.
|
||
</paragraph>
|
||
<paragraph id="8BAC36BE7A44EF64FF01FC37C75EFBA5" blockId="4.[151,1436,151,1106]" pageId="4" pageNumber="47">
|
||
<emphasis id="B967EAAC7A44EF64FF01FC37C712FBFC" bold="true" box="[199,309,1010,1035]" pageId="4" pageNumber="47">Genitalia</emphasis>
|
||
(
|
||
<figureCitation id="13282A3B7A44EF64FE82FC37C7AEFBFD" box="[324,393,1010,1034]" captionStart="FIGURE 4" captionStartId="5.[151,250,1077,1100]" captionTargetBox="[176,1426,411,1061]" captionTargetId="figure@5.[151,1436,395,1071]" captionTargetPageId="5" captionText="FIGURE 4. Female genitalia of E. cesarense; pap. a: papillae anales; du. bu: ductus bursae; crp. bu: corpus bursae; sig: signa." httpUri="https://zenodo.org/record/277763/files/figure.png" pageId="4" pageNumber="47">Fig. 4</figureCitation>
|
||
): Papillae anales long and hairy adorned with setae, ductus bursae (du.bu) not sclerotized and not spirally twisted, long and narrow distally, corpus bursae (crp.bu) rounded with two sclerotized lineal signa (sig) longer than broader.
|
||
</paragraph>
|
||
<caption id="DF6C66367A44EF64FF51F936C57DF8DE" httpUri="https://zenodo.org/record/277762/files/figure.png" pageId="4" pageNumber="47" targetBox="[166,1415,1149,1741]" targetPageId="4">
|
||
<paragraph id="8BAC36BE7A44EF64FF51F936C57DF8DE" blockId="4.[151,1436,1779,1833]" pageId="4" pageNumber="47">
|
||
<emphasis id="B967EAAC7A44EF64FF51F936C735F8FE" bold="true" box="[151,274,1779,1802]" pageId="4" pageNumber="47">FIGURE 3.</emphasis>
|
||
Male genitalia of
|
||
<taxonomicName id="4C134D3D7A44EF64FE0FF930C46FF8FD" box="[457,584,1781,1802]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="cesarense">
|
||
<emphasis id="B967EAAC7A44EF64FE0FF930C46FF8FD" box="[457,584,1781,1802]" italics="true" pageId="4" pageNumber="47">E. cesarense</emphasis>
|
||
</taxonomicName>
|
||
. A, lateral view; B, dorsal view; C, aedeagus (lateral view); ae: aedeagus; un: uncus; te: tegumen; ft: fultura superior; sa: saccus; va: valva; su: superuncus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAC36BE7A44EF65FF01F88AC53DFF20" blockId="4.[151,1436,1871,2007]" lastBlockId="5.[151,1436,151,372]" lastPageId="5" lastPageNumber="48" pageId="4" pageNumber="47">
|
||
<emphasis id="B967EAAC7A44EF64FF01F88AC7A0F89F" bold="true" box="[199,391,1871,1896]" pageId="4" pageNumber="47">Forewing (FW).</emphasis>
|
||
Dorsal ground color paler (R147, G115, B66) than of male, an ocellus between R5 and M2 with a white pupil and encircled by a yellowish ring. This ocellus and the discal, postdiscal, submarginal and marginal lines are more clearly defined than those of the male. VFW color pale yellowish brown (R179, G161, B99). In general, the lines are lighter colored and their relative position is the same as the male, the submarginal line with an undulated pattern from R3 to M3 that continues with an irregular pattern reaching 2A, with a continuous darker yellow (R155, G115, B47) patch from M3-Cu1 to Cu2-A2.
|
||
</paragraph>
|
||
<paragraph id="8BAC36BE7A45EF65FF01FF20C468FE98" blockId="5.[151,1436,151,372]" pageId="5" pageNumber="48">
|
||
<emphasis id="B967EAAC7A45EF65FF01FF20C7B5FF09" bold="true" box="[199,402,229,254]" pageId="5" pageNumber="48">Hindwing (HW).</emphasis>
|
||
Dorsal ground color the same as DFW, two postdiscal ocelli with a little white pupil and encircled by a yellow ring in cells M1-M2 and Cu1-Cu2, submarginal line a little wider than of male. VHW ground color the same as VFW, lines and ocellus pattern the same as the male, but in some specimens the ocellus in Rs-M1 is connected with the ocellus in M1-M2.
|
||
</paragraph>
|
||
<caption id="DF6C66367A45EF65FF51FBF0C3A5FBBC" box="[151,1410,1077,1100]" httpUri="https://zenodo.org/record/277763/files/figure.png" pageId="5" pageNumber="48" targetBox="[176,1426,411,1061]" targetPageId="5">
|
||
<paragraph id="8BAC36BE7A45EF65FF51FBF0C3A5FBBC" blockId="5.[151,1410,1077,1100]" box="[151,1410,1077,1100]" pageId="5" pageNumber="48">
|
||
<emphasis id="B967EAAC7A45EF65FF51FBF0C734FBBC" bold="true" box="[151,275,1077,1100]" pageId="5" pageNumber="48">FIGURE 4.</emphasis>
|
||
Female genitalia of
|
||
<taxonomicName id="4C134D3D7A45EF65FE25FBF2C445FBBB" box="[483,610,1079,1100]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="cesarense">
|
||
<emphasis id="B967EAAC7A45EF65FE25FBF2C445FBBB" box="[483,610,1079,1100]" italics="true" pageId="5" pageNumber="48">E. cesarense</emphasis>
|
||
</taxonomicName>
|
||
; pap.a: papillae anales; du.bu: ductus bursae; crp.bu: corpus bursae; sig: signa.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAC36BE7A45EF65FF01FBB6C248FB58" blockId="5.[151,1436,1139,2036]" pageId="5" pageNumber="48">
|
||
<emphasis id="B967EAAC7A45EF65FF01FBB6C40EFB7C" bold="true" box="[199,553,1139,1163]" pageId="5" pageNumber="48">Early stages and host plants.</emphasis>
|
||
Unknown for the new species, but
|
||
<bibRefCitation id="EF824B4F7A45EF65FC0BFBB6C28EFB7C" box="[973,1193,1139,1163]" pageId="5" pageNumber="48" refString="Singer, M. C., Ehrlich, P. R. & Gilbert, L. E. (1971) Butterfly feeding on Lycopsid. Science, 172, 1341 - 1342." type="journal article">
|
||
Singer
|
||
<emphasis id="B967EAAC7A45EF65FBE6FBB1C273FB7C" box="[1056,1108,1139,1163]" italics="true" pageId="5" pageNumber="48">et al</emphasis>
|
||
(1971)
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EF824B4F7A45EF65FB20FBB6C6C6FB58" pageId="5" pageNumber="48" refString="Beccaloni, G. W., Viloria, A. L., Hall, S. K., & Robison, G. S. (2008) Catologue of the hostplants of the Neotropical butterflies. Catalogo de las plantas huesped de las mariposas neotropicales. Sociedad Entomologica Aragonesa m 3 m: Monografias Tercer MilenioVol. 8. S. E. A., Zaragosa, 536 pp." type="book">
|
||
Beccaloni
|
||
<emphasis id="B967EAAC7A45EF65FAA7FBB1C3BBFB7C" box="[1377,1436,1139,1163]" italics="true" pageId="5" pageNumber="48">et al.</emphasis>
|
||
(2008)
|
||
</bibRefCitation>
|
||
report plants of the
|
||
<taxonomicName id="4C134D3D7A45EF65FE02FB52C455FB58" box="[452,626,1175,1199]" class="Lycopodiopsida" family="Selaginellaceae" kingdom="Plantae" order="Selaginellales" pageId="5" pageNumber="48" phylum="Tracheophyta" rank="family">Selaginellaceae</taxonomicName>
|
||
as host plants for some species of
|
||
<emphasis id="B967EAAC7A45EF65FC30FB52C248FB58" box="[1014,1135,1175,1199]" italics="true" pageId="5" pageNumber="48">
|
||
<taxonomicName id="4C134D3D7A45EF65FC30FB52C24CFB58" box="[1014,1131,1175,1199]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="genus">Euptychia</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30965357A45EF65FF01FB7EC57BFB24" box="[199,860,1211,1236]" pageId="5" pageNumber="48" type="etymology">
|
||
<paragraph id="8BAC36BE7A45EF65FF01FB7EC57BFB24" blockId="5.[151,1436,1139,2036]" box="[199,860,1211,1236]" pageId="5" pageNumber="48">
|
||
<emphasis id="B967EAAC7A45EF65FF01FB7EC769FB24" bold="true" box="[199,334,1211,1235]" pageId="5" pageNumber="48">Etymology.</emphasis>
|
||
The name
|
||
<taxonomicName id="4C134D3D7A45EF65FE0DFB79C41FFB24" box="[459,568,1212,1235]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="cesarense">
|
||
<emphasis id="B967EAAC7A45EF65FE0DFB79C41FFB24" box="[459,568,1212,1235]" italics="true" pageId="5" pageNumber="48">cesarense</emphasis>
|
||
</taxonomicName>
|
||
refers to the
|
||
<typeStatus id="54A8881C7A45EF65FD0DFB79C4DCFB23" box="[715,763,1212,1236]" pageId="5" pageNumber="48">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30965357A45EF65FF01FB1AC202F950" pageId="5" pageNumber="48" type="materials_examined">
|
||
<paragraph id="8BAC36BE7A45EF65FF01FB1AC362FB00" blockId="5.[151,1436,1139,2036]" box="[199,1349,1246,1271]" pageId="5" pageNumber="48">
|
||
<emphasis id="B967EAAC7A45EF65FF01FB1AC742FB00" bold="true" box="[199,357,1246,1271]" pageId="5" pageNumber="48">
|
||
<typeStatus id="54A8881C7A45EF65FF01FB1AC725FB00" box="[199,258,1247,1271]" pageId="5" pageNumber="48">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="F304762E7A45EF65FEAAFB1AC7FAFB00" box="[364,477,1247,1271]" name="Colombia" pageId="5" pageNumber="48">Colombia</collectingCountry>
|
||
, Cesar,
|
||
<collectingCountry id="F304762E7A45EF65FDF3FB1AC493FB00" box="[565,692,1247,1271]" name="Western Sahara" pageId="5" pageNumber="48">Río de Oro</collectingCountry>
|
||
, Vereda El Salobre,
|
||
<geoCoordinate id="EE2750797A45EF65FC5EFB1AC20CFB00" box="[920,1067,1247,1271]" direction="north" orientation="latitude" pageId="5" pageNumber="48" precision="1" value="8.328722">8º19’43.4’’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2750797A45EF65FBF1FB1AC2FAFB00" box="[1079,1245,1247,1271]" direction="west" orientation="longitude" pageId="5" pageNumber="48" precision="1" value="-73.405365">73º24’19.3’’W</geoCoordinate>
|
||
,
|
||
<quantity id="4CEB9B5B7A45EF65FB2FFB1AC367FB00" box="[1257,1344,1247,1271]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="48" unit="m" value="1300.0">1300 m</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAC36BE7A45EF65FF01FAC6C760FA94" blockId="5.[151,1436,1139,2036]" pageId="5" pageNumber="48">
|
||
<typeStatus id="54A8881C7A45EF65FF01FAC6C778FAEC" box="[199,351,1283,1307]" pageId="5" pageNumber="48" type="holotype">HOLOTYPE</typeStatus>
|
||
: Male, with the following labels:
|
||
<emphasis id="B967EAAC7A45EF65FCCDFAC7C58DFAEC" bold="true" box="[779,938,1282,1307]" pageId="5" pageNumber="48">
|
||
<collectingCountry id="F304762E7A45EF65FCCDFAC7C58DFAEC" box="[779,938,1282,1307]" name="Colombia" pageId="5" pageNumber="48">COLOMBIA</collectingCountry>
|
||
</emphasis>
|
||
, Cesar,
|
||
<collectingCountry id="F304762E7A45EF65FBD3FAC6C281FAEC" box="[1045,1190,1283,1307]" name="Western Sahara" pageId="5" pageNumber="48">Río de Oro</collectingCountry>
|
||
, Vereda El Salobre,
|
||
<geoCoordinate id="EE2750797A45EF65FF51FAE2C712FAC8" box="[151,309,1319,1343]" direction="north" orientation="latitude" pageId="5" pageNumber="48" precision="1" value="8.328722">8º19’43.4’’ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2750797A45EF65FE82FAE2C7D2FAC8" box="[324,501,1319,1343]" direction="west" orientation="longitude" pageId="5" pageNumber="48" precision="1" value="-73.405365">73º24’19.3’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CEB9B5B7A45EF65FDC5FAE2C47AFAC8" box="[515,605,1319,1343]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="48" unit="m" value="1300.0">1300 m</quantity>
|
||
,
|
||
<date id="FFAD107E7A45EF65FDAAFAE2C535FAC8" box="[620,786,1319,1343]" pageId="5" pageNumber="48" value="2007-03-06">6 March 2007</date>
|
||
, Hannier Pulido-B. leg., HP 1654, Gen No. 1461, ICN- MHN-L 20417.
|
||
</paragraph>
|
||
<paragraph id="8BAC36BE7A45EF65FF01FAAAC760FA5C" blockId="5.[151,1436,1139,2036]" pageId="5" pageNumber="48">
|
||
ALLOTYPE: Female, with the following labels:
|
||
<emphasis id="B967EAAC7A45EF65FCC2FAABC584FA70" bold="true" box="[772,931,1390,1415]" pageId="5" pageNumber="48">
|
||
<collectingCountry id="F304762E7A45EF65FCC2FAABC584FA70" box="[772,931,1390,1415]" name="Colombia" pageId="5" pageNumber="48">COLOMBIA</collectingCountry>
|
||
</emphasis>
|
||
, Cesar (same data as HT), HP 1653, ICN- MHN-L 20419.
|
||
</paragraph>
|
||
<paragraph id="8BAC36BE7A45EF65FF01FA72C202F950" blockId="5.[151,1436,1139,2036]" pageId="5" pageNumber="48">
|
||
<typeStatus id="54A8881C7A45EF65FF01FA72C778FA38" box="[199,351,1463,1487]" pageId="5" pageNumber="48" type="paratype">PARATYPES</typeStatus>
|
||
:
|
||
<emphasis id="B967EAAC7A45EF65FEB7FA73C428FA38" bold="true" box="[369,527,1462,1487]" pageId="5" pageNumber="48">
|
||
<collectingCountry id="F304762E7A45EF65FEB7FA73C428FA38" box="[369,527,1462,1487]" name="Colombia" pageId="5" pageNumber="48">COLOMBIA</collectingCountry>
|
||
</emphasis>
|
||
, Cesar, González, Vereda San Cayetano,
|
||
<geoCoordinate id="EE2750797A45EF65FC22FA72C24CFA38" box="[996,1131,1463,1487]" direction="north" orientation="latitude" pageId="5" pageNumber="48" precision="15" value="8.289167">8º17’21’’ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2750797A45EF65FBB1FA72C337FA38" box="[1143,1296,1463,1487]" direction="west" orientation="longitude" pageId="5" pageNumber="48" precision="15" value="-73.41667">73º24’60’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CEB9B5B7A45EF65FADAFA72C354FA38" box="[1308,1395,1463,1487]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="48" unit="m" value="1800.0">1800 m</quantity>
|
||
,
|
||
<date id="FFAD107E7A45EF65FA46FA72C72AFA04" pageId="5" pageNumber="48" value="2007-06-23">23 June 2007</date>
|
||
, H. Pulido-B. leg, HP 1979, Gen No. 1462, ICN-MHN-L 21429 (
|
||
<specimenCount id="9D15FD377A45EF65FC3AFA1EC278FA04" box="[1020,1119,1499,1523]" pageId="5" pageNumber="48" type="female">1 female</specimenCount>
|
||
) same data as HT,
|
||
<specimenCount id="9D15FD377A45EF65FAFCFA1EC3BBFA04" box="[1338,1436,1499,1523]" pageId="5" pageNumber="48" type="female">1 female</specimenCount>
|
||
(HP 1653, ICN-MHN-L 20419); Cesar,
|
||
<collectingCountry id="F304762E7A45EF65FDACFA3AC4C8F9E0" box="[618,751,1535,1559]" name="Western Sahara" pageId="5" pageNumber="48">Río de Oro</collectingCountry>
|
||
, Vereda Saninvilla,
|
||
<geoCoordinate id="EE2750797A45EF65FC1CFA3AC24CF9E0" box="[986,1131,1535,1559]" direction="north" orientation="latitude" pageId="5" pageNumber="48" precision="1" value="8.267278">8º16’2.2’’ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2750797A45EF65FBBCFA3AC331F9E0" box="[1146,1302,1535,1559]" direction="west" orientation="longitude" pageId="5" pageNumber="48" precision="15" value="-73.40639">73º24’23’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CEB9B5B7A45EF65FAE2FA3AC358F9E0" box="[1316,1407,1535,1559]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="5" pageNumber="48" unit="m" value="1550.0">1550 m</quantity>
|
||
,
|
||
<date id="FFAD107E7A45EF65FA48FA3AC705F9CC" pageId="5" pageNumber="48" value="2007-03-04">4 March 2007</date>
|
||
, H. Pulido-B. Leg., HP 1396, Gen No. 1457, ICN-MHNL 20420 (
|
||
<specimenCount id="9D15FD377A45EF65FBD4F9E6C247F9CC" box="[1042,1120,1571,1595]" pageId="5" pageNumber="48" type="male">1 male</specimenCount>
|
||
); Cesar,
|
||
<collectingCountry id="F304762E7A45EF65FB04F9E6C365F9CC" box="[1218,1346,1571,1595]" name="Western Sahara" pageId="5" pageNumber="48">Río de Oro</collectingCountry>
|
||
, Vereda El Gitano,
|
||
<geoCoordinate id="EE2750797A45EF65FED4F982C78AF9A8" box="[274,429,1607,1631]" direction="north" orientation="latitude" pageId="5" pageNumber="48" precision="1" value="8.309584">8º18’34.5’’ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2750797A45EF65FE7FF982C475F9A8" box="[441,594,1607,1631]" direction="west" orientation="longitude" pageId="5" pageNumber="48" precision="15" value="-73.412224">73º24’44’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CEB9B5B7A45EF65FD9BF982C493F9A8" box="[605,692,1607,1631]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="48" unit="m" value="1600.0">1600 m</quantity>
|
||
,
|
||
<date id="FFAD107E7A45EF65FD06F982C547F9A8" box="[704,864,1607,1631]" pageId="5" pageNumber="48" value="2007-03-05">5 March 2007</date>
|
||
, H. Pulido-B. Leg., HP 1528, Gen No. 1458, ICN- MHN-L 20421 (
|
||
<specimenCount id="9D15FD377A45EF65FE93F9AEC782F974" box="[341,421,1643,1667]" pageId="5" pageNumber="48" type="male">1 male</specimenCount>
|
||
); Cesar,
|
||
<collectingCountry id="F304762E7A45EF65FDCCF9AEC4ABF974" box="[522,652,1643,1667]" name="Western Sahara" pageId="5" pageNumber="48">Río de Oro</collectingCountry>
|
||
, Vereda El Gitano,
|
||
<geoCoordinate id="EE2750797A45EF65FCA8F9AEC22BF974" box="[878,1036,1643,1667]" direction="north" orientation="latitude" pageId="5" pageNumber="48" precision="1" value="8.324194">8º19’27.1’’ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2750797A45EF65FBDFF9AEC2EEF974" box="[1049,1225,1643,1667]" direction="west" orientation="longitude" pageId="5" pageNumber="48" precision="1" value="-73.409584">73º24’34.5’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CEB9B5B7A45EF65FB13F9AEC309F974" box="[1237,1326,1643,1667]" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" pageId="5" pageNumber="48" unit="m" value="1370.0">1370 m</quantity>
|
||
,
|
||
<date id="FFAD107E7A45EF65FAFAF9AEC6F5F950" pageId="5" pageNumber="48" value="2007-03-05">5 March 2007</date>
|
||
, H. Pulido-B. Leg., HP 1527, Gen. No. 1460, ICN-MHN-L 20276 (
|
||
<specimenCount id="9D15FD377A45EF65FC0CF94AC23FF950" box="[970,1048,1679,1703]" pageId="5" pageNumber="48" type="male">1 male</specimenCount>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30965357A45EF66FF01F977C352FBFA" lastPageId="6" lastPageNumber="49" pageId="5" pageNumber="48" type="diagnosis">
|
||
<paragraph id="8BAC36BE7A45EF65FF01F977C706F85F" blockId="5.[151,1436,1139,2036]" pageId="5" pageNumber="48">
|
||
<emphasis id="B967EAAC7A45EF65FF01F977C766F93C" bold="true" box="[199,321,1714,1739]" pageId="5" pageNumber="48">Diagnosis.</emphasis>
|
||
The most obvious unique feature of the species is having a tornal ocellus in 2A-Cu2. The genitalia of
|
||
<taxonomicName id="4C134D3D7A45EF65FF73F91EC766F905" box="[181,321,1755,1778]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="cesarense">
|
||
<emphasis id="B967EAAC7A45EF65FF73F91EC766F905" box="[181,321,1755,1778]" italics="true" pageId="5" pageNumber="48">E. cesarense</emphasis>
|
||
</taxonomicName>
|
||
have the fultura superior and the superuncus longer than
|
||
<emphasis id="B967EAAC7A45EF65FC00F91EC28FF905" box="[966,1192,1754,1778]" italics="true" pageId="5" pageNumber="48">
|
||
<taxonomicName id="4C134D3D7A45EF65FC00F91EC229F905" box="[966,1038,1755,1778]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="met">E. met</taxonomicName>
|
||
and,
|
||
<taxonomicName id="4C134D3D7A45EF65FB8BF91EC283F905" box="[1101,1188,1755,1778]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="enyo">E. enyo</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Forster’s (1964) genitalia drawings of
|
||
<taxonomicName id="4C134D3D7A45EF65FEA4F93BC7F3F8E1" box="[354,468,1790,1814]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B967EAAC7A45EF65FEA4F93BC7F3F8E1" box="[354,468,1790,1814]" italics="true" pageId="5" pageNumber="48">Euptychia</emphasis>
|
||
</taxonomicName>
|
||
show the internal margin of the valva is plain without any dorsal processes or teeth. This species can be readily distinguished from
|
||
<emphasis id="B967EAAC7A45EF65FD76F8E6C72DF8A9" italics="true" pageId="5" pageNumber="48">
|
||
<taxonomicName id="4C134D3D7A45EF65FD76F8E6C560F8CD" box="[688,839,1826,1850]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="rufocincta">E. rufocincta</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C134D3D7A45EF65FC92F8E6C58BF8CD" box="[852,940,1827,1850]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="enyo">E. enyo</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C134D3D7A45EF65FC7FF8E6C207F8CD" box="[953,1056,1826,1850]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="hilara">E. hilara</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C134D3D7A45EF65FBEBF8E6C28CF8CD" box="[1069,1195,1826,1850]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="insolata">E. insolata</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C134D3D7A45EF65FB7EF8E6C336F8CD" box="[1208,1297,1827,1850]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="jesia">E. jesia</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C134D3D7A45EF65FADBF8E6C350F8CD" box="[1309,1399,1827,1850]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="meta">E. meta</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C134D3D7A45EF65FA42F8E6C72DF8A9" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="rufocincta">E. rufocincta</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C134D3D7A45EF65FE87F882C7F1F8A9" box="[321,470,1862,1886]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="westwoodi">
|
||
<emphasis id="B967EAAC7A45EF65FE87F882C7F1F8A9" box="[321,470,1862,1886]" italics="true" pageId="5" pageNumber="48">E. westwoodi</emphasis>
|
||
</taxonomicName>
|
||
by having, on the VHW, four black ocelli with a white pupil encircled by a yellowish ring and an external reddish brown ring between the postdiscal and submarginal lines in Rs-M1, M1-M2, Cu1-Cu2 and 2A-Cu2.
|
||
</paragraph>
|
||
<paragraph id="8BAC36BE7A45EF65FF03F87DC32DF804" blockId="5.[151,1436,1139,2036]" pageId="5" pageNumber="48">
|
||
<taxonomicName id="4C134D3D7A45EF65FF03F87DC773F838" box="[197,340,1976,1999]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="cesarense">
|
||
<emphasis id="B967EAAC7A45EF65FF03F87DC773F838" box="[197,340,1976,1999]" italics="true" pageId="5" pageNumber="48">E. cesarense</emphasis>
|
||
</taxonomicName>
|
||
is perhaps most similar to
|
||
<taxonomicName id="4C134D3D7A45EF65FD53F87DC50DF838" box="[661,810,1975,1999]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="rufocincta">
|
||
<emphasis id="B967EAAC7A45EF65FD53F87DC50DF838" box="[661,810,1975,1999]" italics="true" pageId="5" pageNumber="48">E. rufocincta</emphasis>
|
||
</taxonomicName>
|
||
, but can be separated because
|
||
<taxonomicName id="4C134D3D7A45EF65FB53F87DC30DF838" box="[1173,1322,1975,1999]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="rufocincta">
|
||
<emphasis id="B967EAAC7A45EF65FB53F87DC30DF838" box="[1173,1322,1975,1999]" italics="true" pageId="5" pageNumber="48">E. rufocincta</emphasis>
|
||
</taxonomicName>
|
||
has three ocelli on the VHW and the postdiscal line on the VFW is not branched at the costa as in
|
||
<taxonomicName id="4C134D3D7A45EF65FBB0F818C325F803" box="[1142,1282,2013,2036]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="cesarense">
|
||
<emphasis id="B967EAAC7A45EF65FBB0F818C325F803" box="[1142,1282,2013,2036]" italics="true" pageId="5" pageNumber="48">E. cesarense</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF6C66367A46EF66FF51FD52C47BFD3B" httpUri="https://zenodo.org/record/277764/files/figure.png" pageId="6" pageNumber="49" targetBox="[164,1422,179,650]" targetPageId="6">
|
||
<paragraph id="8BAC36BE7A46EF66FF51FD52C47BFD3B" blockId="6.[151,1436,663,716]" pageId="6" pageNumber="49">
|
||
<emphasis id="B967EAAC7A46EF66FF51FD52C731FD5A" bold="true" box="[151,278,663,686]" pageId="6" pageNumber="49">FIGURE 5.</emphasis>
|
||
Adults of
|
||
<taxonomicName id="4C134D3D7A46EF66FE4DFD5CC42AFD59" box="[395,525,665,686]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="cesarense">
|
||
<emphasis id="B967EAAC7A46EF66FE4DFD5CC42AFD59" box="[395,525,665,686]" italics="true" pageId="6" pageNumber="49">E. cesarense</emphasis>
|
||
</taxonomicName>
|
||
, A. Holotype from Río de Oro (Cesar, Colombia), B. Allotype from González (Cesar, Colombia). D: Dorsal view; V: Ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAC36BE7A46EF66FF03FD31C352FBFA" blockId="6.[151,1436,755,1542]" pageId="6" pageNumber="49">
|
||
In general aspect, the wing pattern of
|
||
<taxonomicName id="4C134D3D7A46EF66FD44FD30C536FCFB" box="[642,785,757,780]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="cesarense">
|
||
<emphasis id="B967EAAC7A46EF66FD44FD30C536FCFB" box="[642,785,757,780]" italics="true" pageId="6" pageNumber="49">E. cesarense</emphasis>
|
||
</taxonomicName>
|
||
resembles
|
||
<taxonomicName id="4C134D3D7A46EF66FC50FD31C260FCFB" box="[918,1095,756,780]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="enyo">
|
||
<emphasis id="B967EAAC7A46EF66FC50FD31C260FCFB" box="[918,1095,756,780]" italics="true" pageId="6" pageNumber="49">Euptychia enyo</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B967EAAC7A46EF66FB42FD31C723FCC7" italics="true" pageId="6" pageNumber="49">
|
||
<taxonomicName id="4C134D3D7A46EF66FB42FD31C351FCFB" box="[1156,1398,756,780]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="rufocincta">Euptychia rufocincta</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C134D3D7A46EF66FA42FD30C723FCC7" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="cesarense">E. cesarense</taxonomicName>
|
||
</emphasis>
|
||
can be distinguished from
|
||
<taxonomicName id="4C134D3D7A46EF66FDFBFCDCC4B4FCC7" box="[573,659,793,816]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="enyo">
|
||
<emphasis id="B967EAAC7A46EF66FDFBFCDCC4B4FCC7" box="[573,659,793,816]" italics="true" pageId="6" pageNumber="49">E. enyo</emphasis>
|
||
</taxonomicName>
|
||
by lacking the two ocelli between M2-M3 and M3-Cu
|
||
<quantity id="4CEB9B5B7A46EF66FB32FCE2C33CFCD8" box="[1268,1307,791,821]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="6" pageNumber="49" unit="in" value="1.0">1 in</quantity>
|
||
VHW, and lacking the three non-pupilled ocelli in M2-M3, M3-Cu1 and Cu1-Cu
|
||
<quantity id="4CEB9B5B7A46EF66FC49FC8BC592FCA1" box="[911,949,830,860]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="6" pageNumber="49" unit="in" value="2.0">2 in</quantity>
|
||
VFW. The ground color in
|
||
<taxonomicName id="4C134D3D7A46EF66FB35FC85C3A6FCA0" box="[1267,1409,832,855]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="cesarense">
|
||
<emphasis id="B967EAAC7A46EF66FB35FC85C3A6FCA0" box="[1267,1409,832,855]" italics="true" pageId="6" pageNumber="49">E. cesarense</emphasis>
|
||
</taxonomicName>
|
||
is darker and the lines on the ventral side are narrower. The new species can be distinguished from
|
||
<taxonomicName id="4C134D3D7A46EF66FAC1FCA3C3BBFC8A" box="[1287,1436,869,893]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="rufocincta">
|
||
<emphasis id="B967EAAC7A46EF66FAC1FCA3C3BBFC8A" box="[1287,1436,869,893]" italics="true" pageId="6" pageNumber="49">E. rufocincta</emphasis>
|
||
</taxonomicName>
|
||
because in ventral side the lines are narrower and in VFW the postdiscal line is branched. The genitalia of
|
||
<taxonomicName id="4C134D3D7A46EF66FA84FC4FC6F4FC32" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="cesarense">
|
||
<emphasis id="B967EAAC7A46EF66FA84FC4FC6F4FC32" italics="true" pageId="6" pageNumber="49">E. cesarense</emphasis>
|
||
</taxonomicName>
|
||
is compared with Forster’s (1964) genitalia drawings of
|
||
<taxonomicName id="4C134D3D7A46EF66FCAEFC68C5FDFC32" box="[872,986,941,965]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B967EAAC7A46EF66FCAEFC68C5FDFC32" box="[872,986,941,965]" italics="true" pageId="6" pageNumber="49">Euptychia</emphasis>
|
||
</taxonomicName>
|
||
species and with
|
||
<taxonomicName id="4C134D3D7A46EF66FB6DFC6BC324FC32" box="[1195,1283,942,965]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="meta">
|
||
<emphasis id="B967EAAC7A46EF66FB6DFC6BC324FC32" box="[1195,1283,942,965]" italics="true" pageId="6" pageNumber="49">E. meta</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C134D3D7A46EF66FAF8FC6BC3B2FC32" box="[1342,1429,942,965]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="enyo">
|
||
<emphasis id="B967EAAC7A46EF66FAF8FC6BC3B2FC32" box="[1342,1429,942,965]" italics="true" pageId="6" pageNumber="49">E. enyo</emphasis>
|
||
</taxonomicName>
|
||
, showing notable differences: the superuncus is very long, being the half of uncus length, the fultura superior is longer and slightly ventrally curved and the valvae are elongated with a pointed end and a smooth internal margin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30965357A46EF66FF03FBDCC7FBFB8E" pageId="6" pageNumber="49" type="distribution">
|
||
<paragraph id="8BAC36BE7A46EF66FF03FBDCC7FBFB8E" blockId="6.[151,1436,755,1542]" pageId="6" pageNumber="49">
|
||
<emphasis id="B967EAAC7A46EF66FF03FBDCC7D3FBC5" bold="true" box="[197,500,1049,1074]" pageId="6" pageNumber="49">Distribution and ecology.</emphasis>
|
||
Only known from the
|
||
<typeStatus id="54A8881C7A46EF66FD3BFBDFC50AFBC5" box="[765,813,1050,1074]" pageId="6" pageNumber="49">type</typeStatus>
|
||
locality in the Serranía del Perijá. The species flies in secondary-growth Andean and subandean forest with species of the genus
|
||
<taxonomicName id="4C134D3D7A46EF66FC22FBF8C34BFBA2" authority="Hedyosmum, Weinmannia" authorityName="Hedyosmum, Weinmannia" box="[996,1388,1085,1109]" class="Magnoliopsida" family="Juglandaceae" genus="Alfaroa" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="49" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B967EAAC7A46EF66FC22FBF8C34BFBA2" box="[996,1388,1085,1109]" italics="true" pageId="6" pageNumber="49">Alfaroa, Hedyosmum, Weinmannia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C134D3D7A46EF66FF51FBA4C6C7FB8E" box="[151,224,1121,1145]" class="Insecta" family="Clusiidae" genus="Clusia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B967EAAC7A46EF66FF51FBA4C6C7FB8E" box="[151,224,1121,1145]" italics="true" pageId="6" pageNumber="49">Clusia</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<quantity id="4CEB9B5B7A46EF66FEE2FBA4C7FFFB8E" box="[292,472,1121,1145]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.8" metricValueMin="1.3" pageId="6" pageNumber="49" unit="m" value="1550.0" valueMax="1800.0" valueMin="1300.0">
|
||
<date id="FFAD107E7A46EF66FEE2FBA4C79FFB8E" box="[292,440,1121,1145]" pageId="6" pageNumber="49" value="1300" valueMax="1800">1300 to 1800</date>
|
||
m
|
||
</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30965357A46EF66FF01FB40C206F9F2" pageId="6" pageNumber="49" type="discussion">
|
||
<paragraph id="8BAC36BE7A46EF66FF01FB40C206F9F2" blockId="6.[151,1436,755,1542]" pageId="6" pageNumber="49">
|
||
<emphasis id="B967EAAC7A46EF66FF01FB40C76BFB69" bold="true" box="[199,332,1157,1182]" pageId="6" pageNumber="49">Discussion.</emphasis>
|
||
Although there is no consensus about the adult morphological characters that define
|
||
<taxonomicName id="4C134D3D7A46EF66FAE5FB40C3B2FB6A" box="[1315,1429,1157,1181]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B967EAAC7A46EF66FAE5FB40C3B2FB6A" box="[1315,1429,1157,1181]" italics="true" pageId="6" pageNumber="49">Euptychia</emphasis>
|
||
</taxonomicName>
|
||
, the new species is described in this genus because it shows several similarities in ventral markings with other species currently placed in this genus. The relationships among
|
||
<taxonomicName id="4C134D3D7A46EF66FC83FB08C590FB12" box="[837,951,1229,1253]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B967EAAC7A46EF66FC83FB08C590FB12" box="[837,951,1229,1253]" italics="true" pageId="6" pageNumber="49">Euptychia</emphasis>
|
||
</taxonomicName>
|
||
and other euptychiines are not clear (
|
||
<bibRefCitation id="EF824B4F7A46EF66FAA7FB08C7A0FAFE" author="Murray" pageId="6" pageNumber="49" refString="Murray, D. & Prowell, D. P. (2005) Molecular phylogenetics and evolutionary history of the Neotropical Satyrinae Subtribe Euptychiina (Nymphalidae: Satyrinae). Molecular Phylogenetics and Evolution, 34, 67 - 80." type="journal article" year="2005">Murray & Prowell 2005</bibRefCitation>
|
||
; Peña
|
||
<emphasis id="B967EAAC7A46EF66FE1FFB37C431FAFE" box="[473,534,1265,1289]" italics="true" pageId="6" pageNumber="49">et al.</emphasis>
|
||
2006), however it is possible to differentiate
|
||
<taxonomicName id="4C134D3D7A46EF66FBFCFB34C28BFAFE" box="[1082,1196,1265,1289]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B967EAAC7A46EF66FBFCFB34C28BFAFE" box="[1082,1196,1265,1289]" italics="true" pageId="6" pageNumber="49">Euptychia</emphasis>
|
||
</taxonomicName>
|
||
from related genera (
|
||
<bibRefCitation id="EF824B4F7A46EF66FF58FAD0C714FADA" author="Forster" box="[158,307,1301,1325]" pageId="6" pageNumber="49" refString="Forster, W. (1964) Beitrage zur Kenntnis der Insektenfauna Boliviens XIX. Lepidoptera III. Satyridae. Veroffentlichungen der zoologischen Staatssammlung Munchen 8, 51 - 188, pls. 27 - 35." type="journal article" year="1964">Forster 1964</bibRefCitation>
|
||
). The male genitalia in
|
||
<taxonomicName id="4C134D3D7A46EF66FDFAFAD0C489FADA" box="[572,686,1301,1325]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B967EAAC7A46EF66FDFAFAD0C489FADA" box="[572,686,1301,1325]" italics="true" pageId="6" pageNumber="49">Euptychia</emphasis>
|
||
</taxonomicName>
|
||
exhibit a smaller uncus, absent gnathos and very elongated fultura superior.
|
||
<taxonomicName id="4C134D3D7A46EF66FF38FAFCC757FAA6" box="[254,368,1337,1361]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B967EAAC7A46EF66FF38FAFCC757FAA6" box="[254,368,1337,1361]" italics="true" pageId="6" pageNumber="49">Euptychia</emphasis>
|
||
</taxonomicName>
|
||
can be separated from
|
||
<taxonomicName id="4C134D3D7A46EF66FDB4FAFCC583FAA6" authority="Forster, 1964" authorityName="Forster" authorityYear="1964" box="[626,932,1337,1361]" class="Insecta" family="Nymphalidae" genus="Euptychoides" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B967EAAC7A46EF66FDB4FAFCC52EFAA6" box="[626,777,1337,1361]" italics="true" pageId="6" pageNumber="49">Euptychoides</emphasis>
|
||
<bibRefCitation id="EF824B4F7A46EF66FCD6FAFFC583FAA6" author="Forster" box="[784,932,1337,1361]" pageId="6" pageNumber="49" refString="Forster, W. (1964) Beitrage zur Kenntnis der Insektenfauna Boliviens XIX. Lepidoptera III. Satyridae. Veroffentlichungen der zoologischen Staatssammlung Munchen 8, 51 - 188, pls. 27 - 35." type="journal article" year="1964">Forster, 1964</bibRefCitation>
|
||
</taxonomicName>
|
||
because male genitalia in
|
||
<taxonomicName id="4C134D3D7A46EF66FB0CFAFCC346FAA6" box="[1226,1377,1337,1361]" class="Insecta" family="Nymphalidae" genus="Euptychoides" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B967EAAC7A46EF66FB0CFAFCC346FAA6" box="[1226,1377,1337,1361]" italics="true" pageId="6" pageNumber="49">Euptychoides</emphasis>
|
||
</taxonomicName>
|
||
have a well developed gnathos and the aedeagus has cornuti (
|
||
<bibRefCitation id="EF824B4F7A46EF66FCE5FA98C59EFA82" author="Forster" box="[803,953,1373,1397]" pageId="6" pageNumber="49" refString="Forster, W. (1964) Beitrage zur Kenntnis der Insektenfauna Boliviens XIX. Lepidoptera III. Satyridae. Veroffentlichungen der zoologischen Staatssammlung Munchen 8, 51 - 188, pls. 27 - 35." type="journal article" year="1964">Forster 1964</bibRefCitation>
|
||
), while these structures are not present in
|
||
<taxonomicName id="4C134D3D7A46EF66FF51FA44C72EFA6E" box="[151,265,1409,1433]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B967EAAC7A46EF66FF51FA44C72EFA6E" box="[151,265,1409,1433]" italics="true" pageId="6" pageNumber="49">Euptychia</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
<taxonomicName id="4C134D3D7A46EF66FEB2FA47C42AFA6E" box="[372,525,1410,1433]" class="Insecta" family="Nymphalidae" genus="Forsterinaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B967EAAC7A46EF66FEB2FA47C42AFA6E" box="[372,525,1410,1433]" italics="true" pageId="6" pageNumber="49">Forsterinaria</emphasis>
|
||
</taxonomicName>
|
||
species present undulated dark brown discal, postdiscal, submarginal and marginal lines and submarginal ocelli in ventral view in both wings (
|
||
<bibRefCitation id="EF824B4F7A46EF66FCB4FA60C27AFA4A" author="Pena" box="[882,1117,1445,1469]" pageId="6" pageNumber="49" refString="Pena, C. & Lamas, G. (2005) Revision of the butterfly genus Forsterinaria Gray, 1973 (Lepidoptera: Nymphalidae, Satyrinae). Revista peruana de Biologia, 12, 5 - 48." type="journal article" year="2005">Peña & Lamas 2005</bibRefCitation>
|
||
);
|
||
<taxonomicName id="4C134D3D7A46EF66FBB4FA63C3BFFA4A" authority="Forster, 1964" authorityName="Forster" authorityYear="1964" box="[1138,1432,1445,1470]" class="Insecta" family="Nymphalidae" genus="Erichthodes" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B967EAAC7A46EF66FBB4FA63C2DEFA49" box="[1138,1273,1446,1470]" italics="true" pageId="6" pageNumber="49">Erichthodes</emphasis>
|
||
<bibRefCitation id="EF824B4F7A46EF66FAC6FA63C3BFFA4A" author="Forster" box="[1280,1432,1445,1469]" pageId="6" pageNumber="49" refString="Forster, W. (1964) Beitrage zur Kenntnis der Insektenfauna Boliviens XIX. Lepidoptera III. Satyridae. Veroffentlichungen der zoologischen Staatssammlung Munchen 8, 51 - 188, pls. 27 - 35." type="journal article" year="1964">Forster, 1964</bibRefCitation>
|
||
</taxonomicName>
|
||
, in ventral view, present ocelli with bigger white pupils, and some species present white lines instead of those brown lines in
|
||
<taxonomicName id="4C134D3D7A46EF66FF29FA2BC746F9F1" box="[239,353,1518,1542]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B967EAAC7A46EF66FF29FA2BC746F9F1" box="[239,353,1518,1542]" italics="true" pageId="6" pageNumber="49">Euptychia</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C134D3D7A46EF66FEB6FA2BC439F9F1" box="[368,542,1518,1542]" class="Insecta" family="Nymphalidae" genus="Magneuptychia" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B967EAAC7A46EF66FEB6FA2BC439F9F1" box="[368,542,1518,1542]" italics="true" pageId="6" pageNumber="49">Magneuptychia</emphasis>
|
||
</taxonomicName>
|
||
includes bigger sized adults and wider uncus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |