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<document id="CEDDB49B79D20E9F9ADBFF7026B93A82" ID-DOI="10.5281/zenodo.8074943" ID-ISSN="2754-9844" ID-Zenodo-Dep="8074943" ID-ZooBank="BCAD06E8-0AFE-46ED-B7FA-930983CD44C4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="admin" checkinTime="1699388683108" checkinUser="felipe" docAuthor="Noyes, John Stuart" docDate="2023" docId="03BA87A7FF7FFF0BFDFDB9E0A4D3FE45" docLanguage="en" docName="TaxMonogNeotropHymenopt.2.1-921.pdf" docOrigin="Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (1)" docSource="http://dx.doi.org/10.5281/zenodo.8074943" docStyle="DocumentStyle:DAA2ADC4E839520FEEF1E0A10B782A41.1:TaxMonogNeotropHymenopt.2022-.journal_article" docStyleId="DAA2ADC4E839520FEEF1E0A10B782A41" docStyleName="TaxMonogNeotropHymenopt.2022-.journal_article" docStyleVersion="1" docTitle="Prionias nodutus Noyes 2023, sp.nov." docType="treatment" docUuid="00F36969-4673-4D16-A07C-78B93F6BDC31" docUuidSource="ZooBank" docVersion="2" lastPageNumber="171" masterDocId="FF83FFDFFFD5FFA7FFE2BC24A610FFC5" masterDocTitle="ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)" masterLastPageNumber="921" masterPageNumber="1" pageNumber="169" updateTime="1699637169064" updateUser="admin" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="BEB2492EA76E6A7EF064EB80BA02F73F">ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)</mods:title>
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<heading id="D0E481DDFF7FFF0DFDFDB9E0A533FA19" bold="true" box="[543,803,1476,1500]" centered="true" fontSize="10" level="2" pageId="170" pageNumber="169" reason="2">
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<taxonomicName id="4C134D32FF7FFF0DFDFDB9E0A4C2FA19" authority="Noyes, 2023" authorityName="Noyes" authorityYear="2023" box="[543,722,1476,1500]" class="Liliopsida" family="Poaceae" genus="Prionias" kingdom="Plantae" order="Poales" pageId="170" pageNumber="169" phylum="Tracheophyta" rank="species" species="nodutus" status="sp.nov.">
<emphasis id="B967EAA3FF7FFF0DFDFDB9E0A4C2FA19" bold="true" box="[543,722,1476,1500]" italics="true" pageId="170" pageNumber="169">Prionias nodutus</emphasis>
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<paragraph id="8BAC36B1FF7FFF0DFDFBB9FBA538FA32" blockId="170.[537,808,1476,1527]" box="[537,808,1503,1527]" pageId="170" pageNumber="169">
(
<figureCitation id="13282A34FF7FFF0DFDC2B9FBA4BCFA32" box="[544,684,1503,1527]" captionStart="Figs 426-438" captionStartId="767.[181,226,1526,1550]" captionTargetBox="[181,1163,227,1505]" captionTargetId="figure-10@767.[181,1163,223,1508]" captionTargetPageId="767" captionText="Figs 426-438. Georynus dionysus. HT (426) fore wing; (427) ovipositor. Prionias nodutus (428) apex of fore wing venation, HT; (429) fore wing, HT; (430) mesosoma, dorsal view, HT; (431) ovipositor, HT; (432) antenna (a, outer aspect of clava; b, inner aspect of clava), HT; (433) head, facial aspect, HT; (434) hypopygium. Strigenia laras, HT (435) antenna; (436) head, facial view; (437) apex of fore wing venation; (438) fore wing." pageId="170" pageNumber="169">Figs 428-434</figureCitation>
; Hab. E 69)
</paragraph>
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Female (
<typeStatus id="54A88813FF7FFF0DFEECBA30A77DF9E9" box="[270,365,1556,1580]" pageId="170" pageNumber="169">holotype</typeStatus>
): length, including ovipositor,
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; excluding ovipositor,
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. (
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)
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<paragraph id="8BAC36B1FF7FFF0DFF39BA0BA29CF952" blockId="170.[181,1165,1556,1687]" pageId="170" pageNumber="169">Head mainly black, occipital margin with a metallic dark blue sheen, area between occipital margin and posterior ocelli weakly purplish, frontovertex otherwise with a very weak, dark blue sheen, scrobal area with a distinct coppery purple sheen, temple and posterior gena with a weak brassy sheen, lower part of interantennal prominence weakly metallic dark green; piliferous</paragraph>
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<pageTitle id="CB8CEED6FF7EFF0CFEB5BCA4A5FBFF5D" box="[343,1003,128,152]" pageId="171" pageNumber="170">
Taxonomic Monographs on Neotropical
<taxonomicName id="4C134D32FF7EFF0CFD11BCA4A5C7FF5D" authority=", Volume" authorityName="Volume" box="[755,983,128,152]" class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="171" pageNumber="170" phylum="Arthropoda" rank="order">Hymenoptera, Volume</taxonomicName>
2
</pageTitle>
</paragraph>
<paragraph id="8BAC36B1FF7EFF0CFF57BCFBA539FD6F" blockId="171.[181,1165,222,1579]" pageId="171" pageNumber="170">punctures on frontovertex coppery purple; frontovertex, gena and interantennal prominence with short, moderately conspicuous, dark brown setae; radicle orange-brown margined dark brown, scape pale orange, pedicel ventrally pale orange and at apex, otherwise slightly dusky pale orange; funicle slightly dusky pale orange; clava brown; maxillary palpus off-white; pronotum black with a coppery purple sheen; prosternum dark brown; mesoscutum black with a slight, but distinct, metallic dark blue sheen, in some lights purple, especially posteriorly; tegula dark brown with a slight brassy sheen; scutellum metallic dark green, with some purple reflections, especially in anterior one-third; axilla dark purple-brown; dorsum of thorax clothed with dark brown setae; metanotum dark purple-brown with weak brassy sheen; mesopleuron dark purple-brown with a weak brassy sheen; mesosternum dark brown; fore coxa pale yellow, mid and hind coxae pale orange; legs generally pale yellow to pale orange, mid tibia with a small, dark brown subbasal spot, pretarsi brown; wings hyaline, fore wing venation pale brown; propodeum mainly dark purple-brown with a weak brassy sheen, area below spiracle with distinct metallic green sheen; setae surrounding spiracle white, translucent and silvery; gaster generally dark brown with a coppery purple sheen, Gt1 with a deep blue-green metallic sheen, side of Gt1 with a metallic dark green sheen, otherwise side and venter mainly with a mixed blue, brassy, purple and coppery sheen; gonostylus dark brown with extreme apex pale orange.</paragraph>
<paragraph id="8BAC36B1FF7EFF0CFF39BE89A5F2FB87" blockId="171.[181,1165,222,1579]" pageId="171" pageNumber="170">
Head (
<figureCitation id="13282A34FF7EFF0CFEC3BE89A76AFD00" box="[289,378,685,709]" captionStart="Figs 426-438" captionStartId="767.[181,226,1526,1550]" captionTargetBox="[181,1163,227,1505]" captionTargetId="figure-10@767.[181,1163,223,1508]" captionTargetPageId="767" captionText="Figs 426-438. Georynus dionysus. HT (426) fore wing; (427) ovipositor. Prionias nodutus (428) apex of fore wing venation, HT; (429) fore wing, HT; (430) mesosoma, dorsal view, HT; (431) ovipositor, HT; (432) antenna (a, outer aspect of clava; b, inner aspect of clava), HT; (433) head, facial aspect, HT; (434) hypopygium. Strigenia laras, HT (435) antenna; (436) head, facial view; (437) apex of fore wing venation; (438) fore wing." pageId="171" pageNumber="170">Fig. 433</figureCitation>
) about 4.4X as wide as frontovertex; ocelli forming an angle of about 50°; frontovertex with narrowest point about level with anterior ocellus, sculpture of mesh size much less than diameter eye facet; frontovertex with piliferous punctures shallow, 6 setae in front of anterior ocellus, a line of sparse setae along inner eye margin continued along lower eye margin; temple and gena about as shiny as frontovertex, with slightly shallower, longitudinally elongate, strigose to striate-reticulate sculpture, that on temple and posterior part of gena of conspicuously larger mesh; eye separated from scrobal margin by slightly less than diameter of ocellus, area between eye and scrobe with similar sculpture to that on frontovertex; scrobes meeting dorsally, with shallow sculpture; interantennal prominence, well separated from frontovertex, dorsally rounded, sculpture on interantennal prominence polygonally reticulate, about as deep that on frontovertex; antenna as in
<figureCitation id="13282A34FF7EFF0CFE24BF99A40CFC10" box="[454,540,957,981]" captionStart="Figs 426-438" captionStartId="767.[181,226,1526,1550]" captionTargetBox="[181,1163,227,1505]" captionTargetId="figure-10@767.[181,1163,223,1508]" captionTargetPageId="767" captionText="Figs 426-438. Georynus dionysus. HT (426) fore wing; (427) ovipositor. Prionias nodutus (428) apex of fore wing venation, HT; (429) fore wing, HT; (430) mesosoma, dorsal view, HT; (431) ovipositor, HT; (432) antenna (a, outer aspect of clava; b, inner aspect of clava), HT; (433) head, facial aspect, HT; (434) hypopygium. Strigenia laras, HT (435) antenna; (436) head, facial view; (437) apex of fore wing venation; (438) fore wing." pageId="171" pageNumber="170">Fig. 432</figureCitation>
; scape about 5.3X as long as broad; funicle with linear sensilla present on F2-F6; clava hardly broadened, about 1.2X as wide as F6, nearly as long as F3-F6 combined, clava with sensory area extending not quite one-third along its ventral surface; clypeal margin weakly concave, slightly protuberant medially. Relative measurements: HW 70,
<collectingRegion id="49D7F853FF7EFF0CFBA2B82BA275FBE3" box="[1088,1125,1039,1062]" country="Swaziland" name="Hhohho" pageId="171" pageNumber="170">HH</collectingRegion>
66, FV 16, POL 7, OOL 0, OCL 5.5, AOL 8, EL 49, EW 40, MS 22, SL 31.5, SW 6.
</paragraph>
<paragraph id="8BAC36B1FF7EFF0CFF39B861A786FADE" blockId="171.[181,1165,222,1579]" pageId="171" pageNumber="170">
Thorax (
<figureCitation id="13282A34FF7EFF0CFED6B861A79DFB98" box="[308,397,1093,1117]" captionStart="Figs 426-438" captionStartId="767.[181,226,1526,1550]" captionTargetBox="[181,1163,227,1505]" captionTargetId="figure-10@767.[181,1163,223,1508]" captionTargetPageId="767" captionText="Figs 426-438. Georynus dionysus. HT (426) fore wing; (427) ovipositor. Prionias nodutus (428) apex of fore wing venation, HT; (429) fore wing, HT; (430) mesosoma, dorsal view, HT; (431) ovipositor, HT; (432) antenna (a, outer aspect of clava; b, inner aspect of clava), HT; (433) head, facial aspect, HT; (434) hypopygium. Strigenia laras, HT (435) antenna; (436) head, facial view; (437) apex of fore wing venation; (438) fore wing." pageId="171" pageNumber="170">Fig. 430</figureCitation>
) with imbricate-reticulate to polygonally reticulate sculpture on pronotum and mesoscutum that is similar in depth to that on frontovertex below anterior ocellus; axilla with polygonally reticulate sculpture that is a little deeper and coarser than that on mesoscutum; posterior margin of pronotum moderately concave; visible part of mesoscutum about 1.4X as broad as long; scutellum about 0.9X as long as broad; fore wing with venation and distribution of setae as in
<figureCitation id="13282A34FF7EFF0CFEABB8E9A7CBFB20" box="[329,475,1229,1253]" captionStart="Figs 426-438" captionStartId="767.[181,226,1526,1550]" captionTargetBox="[181,1163,227,1505]" captionTargetId="figure-10@767.[181,1163,223,1508]" captionTargetPageId="767" captionText="Figs 426-438. Georynus dionysus. HT (426) fore wing; (427) ovipositor. Prionias nodutus (428) apex of fore wing venation, HT; (429) fore wing, HT; (430) mesosoma, dorsal view, HT; (431) ovipositor, HT; (432) antenna (a, outer aspect of clava; b, inner aspect of clava), HT; (433) head, facial aspect, HT; (434) hypopygium. Strigenia laras, HT (435) antenna; (436) head, facial view; (437) apex of fore wing venation; (438) fore wing." pageId="171" pageNumber="170">Figs 428, 429</figureCitation>
; propodeum (
<figureCitation id="13282A34FF7EFF0CFD89B8E9A4D5FB20" box="[619,709,1229,1253]" captionStart="Figs 426-438" captionStartId="767.[181,226,1526,1550]" captionTargetBox="[181,1163,227,1505]" captionTargetId="figure-10@767.[181,1163,223,1508]" captionTargetPageId="767" captionText="Figs 426-438. Georynus dionysus. HT (426) fore wing; (427) ovipositor. Prionias nodutus (428) apex of fore wing venation, HT; (429) fore wing, HT; (430) mesosoma, dorsal view, HT; (431) ovipositor, HT; (432) antenna (a, outer aspect of clava; b, inner aspect of clava), HT; (433) head, facial aspect, HT; (434) hypopygium. Strigenia laras, HT (435) antenna; (436) head, facial view; (437) apex of fore wing venation; (438) fore wing." pageId="171" pageNumber="170">Fig. 430</figureCitation>
) about 0.14X as long as scutellum, with an indistinct, single, longitudinal, median carina. Relative measurements: FWL 72, FWW 27.5; HWL 53, HWW 12.5.
</paragraph>
<paragraph id="8BAC36B1FF7EFF0CFF39B93BA702FA61" blockId="171.[181,1165,222,1579]" pageId="171" pageNumber="170">
Gaster a little longer than thorax, with hypopygium (
<figureCitation id="13282A34FF7EFF0CFD1FB93BA544FAF2" box="[765,852,1311,1335]" captionStart="Figs 426-438" captionStartId="767.[181,226,1526,1550]" captionTargetBox="[181,1163,227,1505]" captionTargetId="figure-10@767.[181,1163,223,1508]" captionTargetPageId="767" captionText="Figs 426-438. Georynus dionysus. HT (426) fore wing; (427) ovipositor. Prionias nodutus (428) apex of fore wing venation, HT; (429) fore wing, HT; (430) mesosoma, dorsal view, HT; (431) ovipositor, HT; (432) antenna (a, outer aspect of clava; b, inner aspect of clava), HT; (433) head, facial aspect, HT; (434) hypopygium. Strigenia laras, HT (435) antenna; (436) head, facial view; (437) apex of fore wing venation; (438) fore wing." pageId="171" pageNumber="170">Fig. 434</figureCitation>
) reaching about 0.8X towards apex; syntergum about 0.9X as long as mid tibia, with tongue-like extension a little more than a quarter as long as remainder of syntergum; ovipositor (
<figureCitation id="13282A34FF7EFF0CFD00B971A528FAA8" box="[738,824,1365,1389]" captionStart="Figs 426-438" captionStartId="767.[181,226,1526,1550]" captionTargetBox="[181,1163,227,1505]" captionTargetId="figure-10@767.[181,1163,223,1508]" captionTargetPageId="767" captionText="Figs 426-438. Georynus dionysus. HT (426) fore wing; (427) ovipositor. Prionias nodutus (428) apex of fore wing venation, HT; (429) fore wing, HT; (430) mesosoma, dorsal view, HT; (431) ovipositor, HT; (432) antenna (a, outer aspect of clava; b, inner aspect of clava), HT; (433) head, facial aspect, HT; (434) hypopygium. Strigenia laras, HT (435) antenna; (436) head, facial view; (437) apex of fore wing venation; (438) fore wing." pageId="171" pageNumber="170">Fig. 431</figureCitation>
) exserted, the exserted part about 0.2X as long as gaster or nearly as long as mid tibial spur. Relative measurements: OL 75; GS 22.5 [MT 38].
</paragraph>
<paragraph id="8BAC36B1FF7EFF0CFF39B983A773F9EE" blockId="171.[181,1165,222,1579]" pageId="171" pageNumber="170">
Variation. Females vary in overall length from
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, the
<typeStatus id="54A88813FF7EFF0CFC94B983A5DDFA7A" box="[886,973,1447,1471]" pageId="171" pageNumber="170">paratype</typeStatus>
is distinctly larger than the
<typeStatus id="54A88813FF7EFF0CFEE8B9E6A773FA1F" box="[266,355,1474,1498]" pageId="171" pageNumber="170">holotype</typeStatus>
and has paler legs that lack the small, dark brown, subbasal spot on the mid tibia, the fore wing has a slight yellow hue, the head is about 5X as wide as the frontovertex, the flagellar segments are a little more slender, F1 has linear sensilla and the propodeum has several irregular carinae medially.
</paragraph>
<paragraph id="8BAC36B1FF7EFF0CFF57BA6EA771F9A7" blockId="171.[181,353,1610,1634]" box="[181,353,1610,1634]" pageId="171" pageNumber="170">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C309653AFF7EFF0BFF57BAA4A29CFF5A" box="[181,1164,130,1688]" lastPageId="172" lastPageNumber="171" pageId="171" pageNumber="170" type="distribution">
<paragraph id="8BAC36B1FF7EFF0CFF57BAA4A7F9F95D" blockId="171.[181,489,1664,1688]" box="[181,489,1664,1688]" pageId="171" pageNumber="170">
<pageTitle id="CB8CEED6FF7EFF0CFF57BAA4A7F9F95D" box="[181,489,1664,1688]" pageId="171" pageNumber="170">
DISTRIBUTION.
<collectingCountry id="F3047621FF7EFF0CFE91BAA4A7F5F95D" box="[371,485,1664,1688]" name="Costa Rica" pageId="171" pageNumber="170">Costa Rica</collectingCountry>
.
</pageTitle>
</paragraph>
<paragraph id="8BAC36B1FF79FF0BFBBCBCA6A29CFF5A" blockId="172.[1118,1164,130,159]" box="[1118,1164,130,159]" pageId="172" pageNumber="171">171</paragraph>
</subSubSection>
<subSubSection id="C309653AFF79FF0BFF57BCFBA76EFF33" box="[181,382,222,247]" pageId="172" pageNumber="171" type="biology_ecology">
<paragraph id="8BAC36B1FF79FF0BFF57BCFBA76EFF33" blockId="172.[181,382,222,247]" box="[181,382,222,247]" pageId="172" pageNumber="171">HOSTS. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C309653AFF79FF0BFF57BD3CA4D3FE45" pageId="172" pageNumber="171" type="materials_examined">
<paragraph id="8BAC36B1FF79FF0BFF57BD3CA7BDFEE8" blockId="172.[181,1165,280,385]" box="[181,429,280,301]" pageId="172" pageNumber="171">MATERIAL EXAMINED.</paragraph>
<paragraph id="8BAC36B1FF79FF0BFF39BD10A4D3FE45" blockId="172.[181,1165,280,385]" pageId="172" pageNumber="171">
Type material.
<materialsCitation id="3B7B3CECFF79FF0BFE89BD17A77FFEA0" collectingDate="1996-01-26" collectingDateMax="1996-02-24" collectingDateMin="1996-01-26" collectionCode="E, MT, YPT, J, S" country="Costa Rica" elevation="1100" location="Guanacaste" pageId="172" pageNumber="171" specimenCount="1" stateProvince="Guanacaste" typeStatus="holotype">
<typeStatus id="54A88813FF79FF0BFE89BD17A7AFFE8C" box="[363,447,307,329]" pageId="172" pageNumber="171">Holotype</typeStatus>
<collectionCode id="ED02AE74FF79FF0BFE2BBD11A7CBFE8D" box="[457,475,309,328]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="172" pageNumber="171" type="Herbarium">E</collectionCode>
:
<emphasis id="B967EAA3FF79FF0BFE08BD17A469FE8C" bold="true" box="[490,633,307,329]" pageId="172" pageNumber="171">
<collectingCountry id="F3047621FF79FF0BFE08BD17A469FE8C" box="[490,633,307,329]" name="Costa Rica" pageId="172" pageNumber="171">COSTA RICA</collectingCountry>
</emphasis>
,
<emphasis id="B967EAA3FF79FF0BFD6ABD17A4E9FE8C" bold="true" box="[648,761,307,329]" pageId="172" pageNumber="171">
<collectingRegion id="49D7F853FF79FF0BFD6ABD17A4E9FE8C" box="[648,761,307,329]" country="Costa Rica" name="Guanacaste" pageId="172" pageNumber="171">Guanacaste</collectingRegion>
</emphasis>
,
<taxonomicName id="4C134D32FF79FF0BFCEABD10A59BFE8F" authority="(ACG)" baseAuthorityName="ACG" box="[776,907,308,330]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="172" pageNumber="171" phylum="Tracheophyta" rank="species" species="cacao">Cacao (ACG)</taxonomicName>
,
<quantity id="4CEB9B54FF79FF0BFC7ABD10A5C9FE8C" box="[920,985,308,329]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="172" pageNumber="171" unit="m" value="1100.0">
<elevation id="003ED182FF79FF0BFC7ABD10A5C9FE8C" box="[920,985,308,329]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="172" pageNumber="171" unit="m" value="1100.0">1100m</elevation>
</quantity>
,
<collectionCode id="ED02AE74FF79FF0BFC07BD10A217FE8C" box="[997,1031,308,329]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="172" pageNumber="171">MT</collectionCode>
/
<collectionCode id="ED02AE74FF79FF0BFBECBD10A229FE8C" box="[1038,1081,308,329]" pageId="172" pageNumber="171">YPT</collectionCode>
,
<date id="FFAD1071FF79FF0BFBA4BD17A6E5FEA0" pageId="172" pageNumber="171" value="1996-01-26" valueMax="1996-02-24" valueMin="1996-01-26">
<collectingDate id="EFE9E999FF79FF0BFBA4BD17A6E5FEA0" pageId="172" pageNumber="171" value="1996-01-26" valueMax="1996-02-24" valueMin="1996-01-26">26.i-24. ii.1996</collectingDate>
</date>
(
<collectionCode id="ED02AE74FF79FF0BFEE6BD6BA71FFEA1" box="[260,271,335,356]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="172" pageNumber="171" type="Herbarium">J</collectionCode>
.
<collectionCode id="ED02AE74FF79FF0BFEF1BD6BA731FEA1" box="[275,289,335,356]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="172" pageNumber="171" type="Herbarium">S</collectionCode>
. Noyes)
</materialsCitation>
.
<materialsCitation id="3B7B3CECFF79FF0BFE9BBD74A58DFEA0" box="[377,925,335,357]" collectionCode="E, ACG" country="Costa Rica" location="Pitilla" pageId="172" pageNumber="171" specimenCount="1" stateProvince="Guanacaste" typeStatus="paratype">
<typeStatus id="54A88813FF79FF0BFE9BBD74A7C5FEA0" box="[377,469,336,357]" pageId="172" pageNumber="171">Paratypes</typeStatus>
:
<emphasis id="B967EAA3FF79FF0BFE3DBD6BA47BFEA0" bold="true" box="[479,619,335,357]" pageId="172" pageNumber="171">
<collectingCountry id="F3047621FF79FF0BFE3DBD6BA47BFEA0" box="[479,619,335,357]" name="Costa Rica" pageId="172" pageNumber="171">COSTA RICA</collectingCountry>
</emphasis>
, 1
<collectionCode id="ED02AE74FF79FF0BFD61BD75A485FEA1" box="[643,661,337,356]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="172" pageNumber="171" type="Herbarium">E</collectionCode>
,
<emphasis id="B967EAA3FF79FF0BFD43BD6BA502FEA0" bold="true" box="[673,786,335,357]" pageId="172" pageNumber="171">
<collectingRegion id="49D7F853FF79FF0BFD43BD6BA502FEA0" box="[673,786,335,357]" country="Costa Rica" name="Guanacaste" pageId="172" pageNumber="171">Guanacaste</collectingRegion>
</emphasis>
,
<location id="8ECC606AFF79FF0BFCFDBD6BA545FEA0" LSID="urn:lsid:plazi:treatment:03BA87A7FF7FFF0BFDFDB9E0A4D3FE45:8ECC606AFF79FF0BFCFDBD6BA545FEA0" box="[799,853,335,357]" country="Costa Rica" name="Pitilla" pageId="172" pageNumber="171" stateProvince="Guanacaste">Pitilla</location>
(
<collectionCode id="ED02AE74FF79FF0BFC80BD6BA586FEA0" box="[866,918,335,357]" pageId="172" pageNumber="171">ACG</collectionCode>
)
</materialsCitation>
,
<quantity id="4CEB9B54FF79FF0BFC45BD6BA5CDFEA1" box="[935,989,335,356]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="172" pageNumber="171" unit="m" value="700.0">700m</quantity>
,
<materialsCitation id="3B7B3CECFF79FF0BFC0ABD6BA6E5FE44" collectingDate="1996-01-24" collectingDateMax="1996-02-27" collectingDateMin="1996-01-24" collectionCode="MT, YPT" country="Costa Rica" pageId="172" pageNumber="171" specimenCount="1" stateProvince="Guanacaste" typeStatus="paratype">
<collectionCode id="ED02AE74FF79FF0BFC0ABD6BA21AFEA1" box="[1000,1034,335,356]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="172" pageNumber="171">MT</collectionCode>
/
<collectionCode id="ED02AE74FF79FF0BFBF2BD6BA22BFEA1" box="[1040,1083,335,356]" pageId="172" pageNumber="171">YPT</collectionCode>
,
<date id="FFAD1071FF79FF0BFBA4BD6BA6E5FE44" pageId="172" pageNumber="171" value="1996-01-24" valueMax="1996-02-27" valueMin="1996-01-24">
<collectingDate id="EFE9E999FF79FF0BFBA4BD6BA6E5FE44" pageId="172" pageNumber="171" value="1996-01-24" valueMax="1996-02-27" valueMin="1996-01-24">24.i-27. ii.1996</collectingDate>
</date>
</materialsCitation>
(
<collectionCode id="ED02AE74FF79FF0BFEE1BD4FA71EFE45" box="[259,270,363,384]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="172" pageNumber="171" type="Herbarium">J</collectionCode>
.
<collectionCode id="ED02AE74FF79FF0BFEF3BD4FA70FFE45" box="[273,287,363,384]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="172" pageNumber="171" type="Herbarium">S</collectionCode>
. Noyes).
<materialsCitation id="3B7B3CECFF79FF0BFE99BD4FA4D0FE45" box="[379,704,363,385]" collectionCode="NHMUK" country="Costa Rica" pageId="172" pageNumber="171" specimenCount="2" stateProvince="Guanacaste" typeStatus="holotype">
<typeStatus id="54A88813FF79FF0BFE99BD4FA7DFFE44" box="[379,463,363,385]" pageId="172" pageNumber="171">Holotype</typeStatus>
and
<typeStatus id="54A88813FF79FF0BFE1EBD4FA45AFE45" box="[508,586,363,384]" pageId="172" pageNumber="171">paratype</typeStatus>
in
<collectionCode id="ED02AE74FF79FF0BFD85BD4FA4D0FE45" box="[615,704,363,384]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="172" pageNumber="171" type="Museum">NHMUK</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>