1639 lines
195 KiB
XML
1639 lines
195 KiB
XML
<document id="197C642DE6DDD9469C1D48A8B82834BC" ID-CLB-Dataset="270520" ID-DOI="10.11646/zootaxa.5351.3.2" ID-GBIF-Dataset="5b1114e9-6e56-4af5-8f2d-3671cf5d4c0d" ID-ISSN="1175-5326" ID-Zenodo-Dep="8391692" ID-ZooBank="40E9413B-90BC-41C5-944C-F3271B5FD5FC" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="jonas" IM.treatments_approvedBy="julia" checkinTime="1695990468755" checkinUser="plazi" docAuthor="Allsopp, Peter G. & Hutchinson, Paul M." docDate="2023" docId="039ADD0BFFA40A1CFF4AC6BEFA0A08EE" docLanguage="en" docName="zootaxa.5351.3.2.pdf" docOrigin="Zootaxa 5351 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5351.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dynastinae" docType="key" docVersion="5" lastPageNumber="339" masterDocId="FFA3A573FFA80A0DFFDDC445FF960B64" masterDocTitle="Philcarneum new genus and Constricticollis new genus, two distinctive rhinoceros beetles (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) from opposite sides of Australia with a revised key to the Australian dynastine genera" masterLastPageNumber="340" masterPageNumber="322" pageNumber="334" updateTime="1729606666396" updateUser="jonas" zenodo-license-document="CLOSED">
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<mods:title id="592DF195D73D666F4C583A9524D089D7">Philcarneum new genus and Constricticollis new genus, two distinctive rhinoceros beetles (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) from opposite sides of Australia with a revised key to the Australian dynastine genera</mods:title>
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<mods:name id="19E44A39569368FE62F7FB31E4B3A353" type="personal">
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<mods:namePart id="3ED5708E1CF62B8E2BE96998C1EC10F5">Allsopp, Peter G.</mods:namePart>
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<mods:namePart id="CF48D240C3199986C15D55245F924149">Hutchinson, Paul M.</mods:namePart>
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<mods:affiliation id="1507044C155B913DBFC17F57B0EEC7D5">Department of Primary Industries and Regional Development (Quarantine WA), Locked Bag 69, Welshpool DC, Western Australia 6986, Australia</mods:affiliation>
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<mods:title id="B62F89715BDF6034F77A21E1B71A8FAB">Zootaxa</mods:title>
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<mods:date id="E9BCE6D0AC8316AA63FB9E8107D1AA05">2023</mods:date>
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<mods:number id="3BB85918DF45F2DBFED4AED8CDAB735F">2023-09-27</mods:number>
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<treatment id="039ADD0BFFA40A1CFF4AC6BEFA0A08EE" ID-DOI="http://doi.org/10.5281/zenodo.8400467" ID-GBIF-Taxon="213635936" ID-Zenodo-Dep="8400467" LSID="urn:lsid:plazi:treatment:039ADD0BFFA40A1CFF4AC6BEFA0A08EE" httpUri="http://treatment.plazi.org/id/039ADD0BFFA40A1CFF4AC6BEFA0A08EE" lastPageId="17" lastPageNumber="339" pageId="12" pageNumber="334">
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<subSubSection id="C3293F96FFA40A01FF4AC6BEFD120870" box="[151,644,762,789]" pageId="12" pageNumber="334" type="nomenclature">
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<paragraph id="8B8C6C1DFFA40A01FF4AC6BEFD120870" blockId="12.[151,644,762,789]" box="[151,644,762,789]" pageId="12" pageNumber="334">
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<emphasis id="B947B00FFFA40A01FF4AC6BEFD120870" bold="true" box="[151,644,762,789]" pageId="12" pageNumber="334">
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Key to genera of Australian
|
||
<taxonomicName id="4C33179EFFA40A01FE25C6BFFD120870" ID-CoL="7NLQQ" box="[504,644,762,788]" class="Insecta" family="Scarabaeidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="339" phylum="Arthropoda" rank="subFamily" subFamily="Dynastinae">Dynastinae</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3293F96FFA40A01FF4AC706FE7A08AD" pageId="12" pageNumber="334" type="multiple">
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<paragraph id="8B8C6C1DFFA40A01FF4AC706FE7A08AD" blockId="12.[151,1437,835,969]" pageId="12" pageNumber="334">
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This key is modified from
|
||
<bibRefCitation id="EFA211ECFFA40A01FE60C706FD140839" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[445,642,835,861]" pageId="12" pageNumber="334" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA40A01FE25C706FDB90839" box="[504,559,835,861]" italics="true" pageId="12" pageNumber="334">et al.</emphasis>
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(2019)
|
||
</bibRefCitation>
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||
, and incorporates
|
||
<taxonomicName id="4C33179EFFA40A01FC96C706FAD60839" authority="(Hutchinson & Allsopp 2021)" baseAuthorityName="Hutchinson & Allsopp" baseAuthorityYear="2021" box="[843,1344,835,861]" class="Insecta" family="Scarabaeidae" genus="Erbmahcedius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="334" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA40A01FC96C706FC7B0839" box="[843,1005,835,861]" italics="true" pageId="12" pageNumber="334">Erbmahcedius</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA211ECFFA40A01FC27C706FAAE0839" author="Hutchinson, P. M. & Allsopp, P. G." box="[1018,1336,835,861]" pageId="12" pageNumber="334" pagination="439 - 462" refId="ref19528" refString="Hutchinson, P. M. & Allsopp, P. G. (2021) Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa, 5072 (5), 439 - 462. https: // doi. org / 10.11646 / zootaxa. 5072.5.2" type="journal article" year="2021">Hutchinson & Allsopp 2021</bibRefCitation>
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||
)
|
||
</taxonomicName>
|
||
, the two genera described here and
|
||
<taxonomicName id="4C33179EFFA40A01FE1CC722FC3208E5" authority="Reid & Tees, 2023" authorityName="Reid & Tees" authorityYear="2023" box="[449,932,871,897]" class="Insecta" family="Scarabaeidae" genus="Semanopterus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="334" phylum="Arthropoda" rank="species" species="kingstoni">
|
||
<emphasis id="B947B00FFFA40A01FE1CC722FD5B08E5" box="[449,717,871,897]" italics="true" pageId="12" pageNumber="334">Semanopterus kingstoni</emphasis>
|
||
<bibRefCitation id="EFA211ECFFA40A01FD0EC722FC3208E5" author="Reid, C. A. M. & Tees, N. A." box="[723,932,871,897]" pageId="12" pageNumber="334" pagination="563 - 570" refId="ref19857" refString="Reid, C. A. M. & Tees, N. A. (2023) A new, but possibly extinct, species of Semanopterus Hope, 1847 from Lord Howe Island, in the southwestern Pacific Ocean (Coleoptera: Scarabaeidae: Dynastinae). Zootaxa, 5306 (5), 563 - 570. https: // doi. org / 10.11646 / zootaxa. 5306.5.4" type="journal article" year="2023">Reid & Tees, 2023</bibRefCitation>
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||
</taxonomicName>
|
||
from Lord Howe Island (
|
||
<bibRefCitation id="EFA211ECFFA40A01FB1CC722FA0708E5" author="Reid, C. A. M. & Tees, N. A." box="[1217,1425,871,897]" pageId="12" pageNumber="334" pagination="563 - 570" refId="ref19857" refString="Reid, C. A. M. & Tees, N. A. (2023) A new, but possibly extinct, species of Semanopterus Hope, 1847 from Lord Howe Island, in the southwestern Pacific Ocean (Coleoptera: Scarabaeidae: Dynastinae). Zootaxa, 5306 (5), 563 - 570. https: // doi. org / 10.11646 / zootaxa. 5306.5.4" type="journal article" year="2023">Reid & Tees 2023</bibRefCitation>
|
||
). The presence of elytral horns in
|
||
<taxonomicName id="4C33179EFFA40A01FE20C7CEFCEF08C1" authority="Sharp, 1873" authorityName="Sharp" authorityYear="1873" box="[509,889,907,933]" class="Insecta" family="Scarabaeidae" genus="Nephrodopus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="334" phylum="Arthropoda" rank="species" species="enigma">
|
||
<emphasis id="B947B00FFFA40A01FE20C7CEFD7D08C0" box="[509,747,907,933]" italics="true" pageId="12" pageNumber="334">Nephrodopus enigma</emphasis>
|
||
Sharp, 1873
|
||
</taxonomicName>
|
||
was previously discounted as a generic character (
|
||
<bibRefCitation id="EFA211ECFFA40A01FF42C7EAFE7708AD" author="Hutchinson, P. M. & Allsopp, P. G." box="[159,481,943,969]" pageId="12" pageNumber="334" pagination="1 - 14" refId="ref19412" refString="Hutchinson, P. M. & Allsopp, P. G. (2019) Revisiting Nephrodopus Sharp, 1873, with the description of a new species and a female (Coleoptera: Scarabaeidae: Dynastinae). Australian Entomologist, 46, 1 - 14." type="journal article" year="2019">Hutchinson & Allsopp 2019</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3293F96FFA40A1CFF4AC7B1FA0A08EE" lastPageId="17" lastPageNumber="339" pageId="12" pageNumber="334" type="key">
|
||
<key id="FFA23B2CFFA40A1CFF4AC7B1FA0A08EE" lastPageId="17" lastPageNumber="339" pageId="12" pageNumber="334">
|
||
<keyStep id="30C774B8FFA40A01FF4AC7B1FA0A0F42" pageId="12" pageNumber="334">
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<paragraph id="8B8C6C1DFFA40A01FF4AC7B1FA0A0F6D" blockId="12.[151,1437,1012,2031]" box="[151,1436,1012,1034]" pageId="12" pageNumber="334">
|
||
<keyLead id="30C2CF28FFA40A01FF4AC7B1FA0A0F6D" box="[151,1436,1012,1034]" pageId="12" pageNumber="334">1 Mentum dilated, concealing at least bases of labial palps and usually most of maxillae and labium … Phileurini.......... 2</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA40A01FF4AC054FA0A0F42" blockId="12.[151,1437,1012,2031]" box="[151,1436,1041,1063]" pageId="12" pageNumber="334">
|
||
<keyLead id="30C2CF28FFA40A01FF4AC054FA0A0F42" box="[151,1436,1041,1063]" pageId="12" pageNumber="334">- Mentum normal, more-or-less narrow, not concealing bases of maxillary palps.................................... 3</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFA40A01FF4AC06BFA0A0E4A" pageId="12" pageNumber="334">
|
||
<paragraph id="8B8C6C1DFFA40A01FF4AC06BFA0A0FFF" blockId="12.[151,1437,1012,2031]" pageId="12" pageNumber="334">
|
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<keyLead id="30C2CF28FFA40A01FF4AC06BFA0A0FFF" pageId="12" pageNumber="334">
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2 Mandibles produced beyond clypeus, which is apically narrowed; mentum not concealing maxillae (
|
||
<bibRefCitation id="EFA211ECFFA40A01FB1AC06BFAE60F27" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[1223,1392,1070,1092]" pageId="12" pageNumber="334" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA40A01FADDC06AFAA20F27" box="[1280,1332,1070,1092]" italics="true" pageId="12" pageNumber="334">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.2) and not produced at suture with submentum; antennal scape not expanded and wedge-like, not concealing funicular antennomeres; black,
|
||
<quantity id="4CCBC1F8FFA40A01FE69C02CFD8D0F1A" box="[436,539,1129,1150]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="2.9" metricValueMin="2.0" pageId="12" pageNumber="334" unit="mm" value="24.5" valueMax="29.0" valueMin="20.0">20–29 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA40A01FD70C02CFCDA0F1A" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[685,844,1129,1151]" pageId="12" pageNumber="334" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA40A01FD3FC02FFC850F1A" box="[738,787,1129,1151]" italics="true" pageId="12" pageNumber="334">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 56C); northern Queensland,
|
||
<collectingCountry id="F3242C8DFFA40A01FB46C02CFAAB0F1B" box="[1179,1341,1129,1151]" name="Christmas Island" pageId="12" pageNumber="334">Christmas Island</collectingCountry>
|
||
............................................................................. Phileurina:
|
||
<taxonomicName id="4C33179EFFA40A01FB74C0C3FA0A0FFF" ID-CoL="4CHR" authority="Arrow, 1908" authorityName="Arrow" authorityYear="1908" box="[1193,1436,1158,1179]" class="Insecta" family="Scarabaeidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="334" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA40A01FB74C0C3FA880FFF" box="[1193,1310,1158,1179]" italics="true" pageId="12" pageNumber="334">Eophileurus</emphasis>
|
||
Arrow, 1908
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA40A01FF4AC0E1FA0A0E4A" blockId="12.[151,1437,1012,2031]" pageId="12" pageNumber="334">
|
||
<keyLead id="30C2CF28FFA40A01FF4AC0E1FA0A0E4A" pageId="12" pageNumber="334">
|
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- Mandibles not visible beyond clypeus, which is broadly rounded or truncate at apex; mentum greatly enlarged, concealing all mouthparts (
|
||
<bibRefCitation id="EFA211ECFFA40A01FE86C084FE690FB2" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[347,511,1217,1239]" pageId="12" pageNumber="334" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA40A01FE4FC087FE530FB2" box="[402,453,1217,1239]" italics="true" pageId="12" pageNumber="334">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.3), and produced and often notched at suture with submentum; antennal scape apically dilated and wedge-like, concealing all funicular segments; brown-black, red-brown, shiny dark brownish black or black,
|
||
<quantity id="4CCBC1F8FFA40A01FABCC09BFE910E75" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.7" metricValueMin="1.1" pageId="12" pageNumber="334" unit="mm" value="19.0" valueMax="27.0" valueMin="11.0">11–27 mm</quantity>
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long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA40A01FE4AC0B9FDA30E75" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[407,565,1276,1298]" pageId="12" pageNumber="334" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
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Weir
|
||
<emphasis id="B947B00FFFA40A01FE11C0B9FE6B0E75" box="[460,509,1276,1297]" italics="true" pageId="12" pageNumber="334">et al.</emphasis>
|
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2019
|
||
</bibRefCitation>
|
||
: plate 52K);
|
||
<collectingRegion id="49F7A2FFFFA40A01FD72C0B9FCCA0E76" box="[687,860,1276,1298]" country="Australia" name="New South Wales" pageId="12" pageNumber="334">New South Wales</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA40A01FCB9C0B9FB800E76" box="[868,1046,1276,1298]" country="Australia" name="Northern Territory" pageId="12" pageNumber="334">Northern Territory</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA40A01FBC2C0B9FB050E76" box="[1055,1171,1276,1298]" country="Australia" name="Queensland" pageId="12" pageNumber="334">Queensland</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA40A01FB46C0B9FAA40E76" box="[1179,1330,1276,1298]" country="Australia" name="South Australia" pageId="12" pageNumber="334">South Australia</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA40A01FAE6C0B9FA0F0E76" box="[1339,1433,1276,1298]" country="Australia" name="Tasmania" pageId="12" pageNumber="334">Tasmania</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA40A01FF3FC15CFEA40E4B" box="[226,306,1305,1327]" country="Australia" name="Victoria" pageId="12" pageNumber="334">Victoria</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA40A01FEE6C15CFE710E4B" box="[315,487,1305,1327]" country="Australia" name="Western Australia" pageId="12" pageNumber="334">Western Australia</collectingRegion>
|
||
, Lord Howe Island............................... Cryptodina:
|
||
<taxonomicName id="4C33179EFFA40A01FB40C15CFA0A0E4A" ID-CoL="3WL7" authority="MacLeay, 1819" authorityName="MacLeay" authorityYear="1819" box="[1181,1436,1305,1326]" class="Insecta" family="Scarabaeidae" genus="Cryptodus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="334" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA40A01FB40C15CFA970E4A" box="[1181,1281,1305,1326]" italics="true" pageId="12" pageNumber="334">Cryptodus</emphasis>
|
||
MacLeay, 1819
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFA40A01FF4AC173FA0A0EE3" pageId="12" pageNumber="334">
|
||
<paragraph id="8B8C6C1DFFA40A01FF4AC173FA0A0E2F" blockId="12.[151,1437,1012,2031]" box="[151,1436,1334,1356]" pageId="12" pageNumber="334">
|
||
<keyLead id="30C2CF28FFA40A01FF4AC173FA0A0E2F" box="[151,1436,1334,1356]" pageId="12" pageNumber="334">
|
||
3 Apex of metatibia scalloped (
|
||
<bibRefCitation id="EFA211ECFFA40A01FE21C173FD0A0E2F" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[508,668,1334,1356]" pageId="12" pageNumber="334" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA40A01FDEFC172FDF50E2F" box="[562,611,1334,1356]" italics="true" pageId="12" pageNumber="334">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.4) or with longer or shorter fixed teeth, never with cilia............ 4
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA40A01FF4AC111FA0A0EE3" blockId="12.[151,1437,1012,2031]" pageId="12" pageNumber="334">
|
||
<keyLead id="30C2CF28FFA40A01FF4AC111FA0A0EE3" pageId="12" pageNumber="334">
|
||
- Apex edge of metatibia truncate or very slightly produced, almost always with at least a few (Oryctoderini) or many cilia or cilia lacking (
|
||
<taxonomicName id="4C33179EFFA40A01FEBBC134FE740EE2" authorityName="Allsopp & Hutchinson" authorityYear="2023" box="[358,482,1393,1414]" class="Insecta" family="Scarabaeidae" genus="Philcarneum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="334" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA40A01FEBBC134FE740EE2" box="[358,482,1393,1414]" italics="true" pageId="12" pageNumber="334">Philcarneum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C33179EFFA40A01FE33C134FD130EE2" ID-CoL="7TPM" authorityName="Hope" authorityYear="1837" box="[494,645,1393,1414]" class="Insecta" family="Scarabaeidae" genus="Temnorhynchus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="334" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA40A01FE33C134FD130EE2" box="[494,645,1393,1414]" italics="true" pageId="12" pageNumber="334">Temnorhynchus</emphasis>
|
||
</taxonomicName>
|
||
)................................................................ 6
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFA40A01FF4AC1CBFA0A0D53" pageId="12" pageNumber="334">
|
||
<paragraph id="8B8C6C1DFFA40A01FF4AC1CBFA0A0E9F" blockId="12.[151,1437,1012,2031]" pageId="12" pageNumber="334">
|
||
<keyLead id="30C2CF28FFA40A01FF4AC1CBFA0A0E9F" pageId="12" pageNumber="334">
|
||
4 Propygidium enlarged and bearing transverse stridulating ridges (
|
||
<bibRefCitation id="EFA211ECFFA40A01FCB4C1CBFB870EC7" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[873,1041,1422,1444]" pageId="12" pageNumber="334" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA40A01FC7CC1CAFC430EC7" box="[929,981,1422,1444]" italics="true" pageId="12" pageNumber="334">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.5); apical pygidial ridge setose; metatarsomere 1 produced apically (
|
||
<bibRefCitation id="EFA211ECFFA40A01FDE0C1E9FD4A0EA5" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[573,732,1452,1474]" pageId="12" pageNumber="334" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA40A01FDAEC1E9FD350EA5" box="[627,675,1452,1473]" italics="true" pageId="12" pageNumber="334">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.6); sexual dimorphism of armature, slight, head with median horn in both sexes; black,
|
||
<quantity id="4CCBC1F8FFA40A01FE68C18CFD890EBA" box="[437,543,1481,1503]" metricMagnitude="-2" metricUnit="m" metricValue="4.8" metricValueMax="6.3" metricValueMin="3.3" pageId="12" pageNumber="334" unit="mm" value="48.0" valueMax="63.0" valueMin="33.0">33–63 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA40A01FD65C18CFC420EBA" author="Hutchinson, P. M. & Allsopp, P. G." box="[696,980,1481,1503]" pageId="12" pageNumber="334" pagination="1 - 14" refId="ref19412" refString="Hutchinson, P. M. & Allsopp, P. G. (2019) Revisiting Nephrodopus Sharp, 1873, with the description of a new species and a female (Coleoptera: Scarabaeidae: Dynastinae). Australian Entomologist, 46, 1 - 14." type="journal article" year="2019">Allsopp & Hutchinson 2019</bibRefCitation>
|
||
: figs. 14–15);
|
||
<collectingRegion id="49F7A2FFFFA40A01FBB9C18CFA8F0EBB" box="[1124,1305,1481,1503]" country="Australia" name="Northern Territory" pageId="12" pageNumber="334">Northern Territory</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA40A01FAF8C18CFA0F0EBB" box="[1317,1433,1481,1503]" country="Australia" name="Queensland" pageId="12" pageNumber="334">Queensland</collectingRegion>
|
||
, Cocos-Keeling Islands.......................................................... Oryctini:
|
||
<taxonomicName id="4C33179EFFA40A01FB0BC1A2FA0A0E9F" ID-CoL="63GZ3" authority="Illiger, 1798" authorityName="Illiger" authorityYear="1798" box="[1238,1436,1510,1532]" class="Insecta" family="Scarabaeidae" genus="Oryctes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="334" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA40A01FB0BC1A2FAB70E98" box="[1238,1313,1511,1532]" italics="true" pageId="12" pageNumber="334">Oryctes</emphasis>
|
||
Illiger, 1798
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA40A01FF4AC241FA0A0D53" blockId="12.[151,1437,1012,2031]" pageId="12" pageNumber="334">
|
||
<keyLead id="30C2CF28FFA40A01FF4AC241FA0A0D53" pageId="12" pageNumber="334">- Propygidium not enlarged, without stridulating ridges; apical pygidial ridge glabrous; metatarsomere 1 not produced apically; sexual dimorphism of armature great, head in female at most with tubercle … Dynastini............................. 5</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFA40A01FF4AC27BFA0A0C5A" pageId="12" pageNumber="334">
|
||
<paragraph id="8B8C6C1DFFA40A01FF4AC27BFA0A0DAD" blockId="12.[151,1437,1012,2031]" pageId="12" pageNumber="334">
|
||
<keyLead id="30C2CF28FFA40A01FF4AC27BFA0A0DAD" pageId="12" pageNumber="334">
|
||
5 Meso- and metatibiae with apices obtusely scalloped and intermediate and basal carinae non-ciliate (
|
||
<bibRefCitation id="EFA211ECFFA40A01FB47C27BFAA00D37" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[1178,1334,1598,1620]" pageId="12" pageNumber="334" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA40A01FB13C27AFB680D37" box="[1230,1278,1598,1620]" italics="true" pageId="12" pageNumber="334">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.7); clypeal apex bituberculate; mandibles strongly bidentate (
|
||
<bibRefCitation id="EFA211ECFFA40A01FCDAC219FC330D15" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[775,933,1628,1650]" pageId="12" pageNumber="334" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA40A01FCE0C219FCFB0D15" box="[829,877,1628,1649]" italics="true" pageId="12" pageNumber="334">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.8); male with bifid cephalic horn and median pronotal process; metatibial spurs extremely short; female with dorsal surfaces conspicuously punctate and head with traces of cephalic tubercles; black,
|
||
<quantity id="4CCBC1F8FFA40A01FE00C2D3FDD30DC8" box="[477,581,1686,1708]" metricMagnitude="-2" metricUnit="m" metricValue="4.75" metricValueMax="6.6" metricValueMin="2.9" pageId="12" pageNumber="334" unit="mm" value="47.5" valueMax="66.0" valueMin="29.0">29–66 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA40A01FD04C2D3FCED0DCF" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[729,891,1686,1708]" pageId="12" pageNumber="334" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA40A01FCCDC2D2FCD70DCF" box="[784,833,1686,1708]" italics="true" pageId="12" pageNumber="334">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plates 53H, 56G;
|
||
<bibRefCitation id="EFA211ECFFA40A01FBF2C2D3FAD30DCF" author="Hutchinson, P. M. & Allsopp, P. G." box="[1071,1349,1686,1708]" pageId="12" pageNumber="334" pagination="1 - 14" refId="ref19412" refString="Hutchinson, P. M. & Allsopp, P. G. (2019) Revisiting Nephrodopus Sharp, 1873, with the description of a new species and a female (Coleoptera: Scarabaeidae: Dynastinae). Australian Entomologist, 46, 1 - 14." type="journal article" year="2019">Allsopp & Hutchinson 2019</bibRefCitation>
|
||
: fig. 19);
|
||
<collectingRegion id="49F7A2FFFFA40A01FF3FC2F1FE060DAE" box="[226,400,1716,1738]" country="Australia" name="New South Wales" pageId="12" pageNumber="334">New South Wales</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA40A01FE44C2F1FDDD0DAE" box="[409,587,1716,1738]" country="Australia" name="Northern Territory" pageId="12" pageNumber="334">Northern Territory</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA40A01FD89C2F1FD530DAE" box="[596,709,1716,1738]" country="Australia" name="Queensland" pageId="12" pageNumber="334">Queensland</collectingRegion>
|
||
..........................................
|
||
<taxonomicName id="4C33179EFFA40A01FB63C2F1FA0A0DAD" ID-CoL="89LR" authority="Hope, 1837" authorityName="Hope" authorityYear="1837" box="[1214,1436,1716,1737]" class="Insecta" family="Scarabaeidae" genus="Xylotrupes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="334" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA40A01FB63C2F1FAB00DAD" box="[1214,1318,1716,1737]" italics="true" pageId="12" pageNumber="334">Xylotrupes</emphasis>
|
||
Hope, 1837
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA40A01FF4AC294FA0A0C5A" blockId="12.[151,1437,1012,2031]" pageId="12" pageNumber="334">
|
||
<keyLead id="30C2CF28FFA40A01FF4AC294FA0A0C5A" pageId="12" pageNumber="334">
|
||
- Meso- and metatibiae with apices strongly bispinose and intermediate and basal carinae ciliate (
|
||
<bibRefCitation id="EFA211ECFFA40A01FB50C294FAA70D82" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[1165,1329,1745,1767]" pageId="12" pageNumber="334" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA40A01FB19C297FB600D82" box="[1220,1270,1745,1767]" italics="true" pageId="12" pageNumber="334">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.9); clypeal apex truncate or rounded; mandibles untoothed (
|
||
<bibRefCitation id="EFA211ECFFA40A01FCDFC2ABFC320C67" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[770,932,1774,1796]" pageId="12" pageNumber="334" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA40A01FCE5C2AAFCFC0C67" box="[824,874,1774,1796]" italics="true" pageId="12" pageNumber="334">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.10); metatibial spurs elongate in both sexes; deep brown to black,
|
||
<quantity id="4CCBC1F8FFA40A01FE66C34EFDB20C45" box="[443,548,1803,1825]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" metricValueMax="5.6" metricValueMin="2.6" pageId="12" pageNumber="334" unit="mm" value="41.0" valueMax="56.0" valueMin="26.0">26–56 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA40A01FD63C349FCF30C45" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[702,869,1804,1826]" pageId="12" pageNumber="334" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA40A01FD2BC349FCBF0C45" box="[758,809,1804,1825]" italics="true" pageId="12" pageNumber="334">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plates 53G, 55L);
|
||
<collectingRegion id="49F7A2FFFFA40A01FBFBC349FB4F0C46" box="[1062,1241,1804,1826]" country="Australia" name="New South Wales" pageId="12" pageNumber="334">New South Wales</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA40A01FB39C349FA0F0C46" box="[1252,1433,1804,1826]" country="Australia" name="Northern Territory" pageId="12" pageNumber="334">Northern Territory</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA40A01FF3FC36CFEC00C5B" box="[226,342,1833,1855]" country="Australia" name="Queensland" pageId="12" pageNumber="334">Queensland</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA40A01FE82C36CFD9F0C5B" box="[351,521,1833,1855]" country="Australia" name="Western Australia" pageId="12" pageNumber="334">Western Australia</collectingRegion>
|
||
.....................................................
|
||
<taxonomicName id="4C33179EFFA40A01FB54C36CFA0A0C5A" ID-CoL="4TK9" authority="Arrow, 1908" authorityName="Arrow" authorityYear="1908" box="[1161,1436,1833,1854]" class="Insecta" family="Scarabaeidae" genus="Haploscapanes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="334" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA40A01FB54C36CFA8B0C5A" box="[1161,1309,1833,1854]" italics="true" pageId="12" pageNumber="334">Haploscapanes</emphasis>
|
||
Arrow, 1908
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<paragraph id="8B8C6C1DFFA40A00FF4AC303FA0A0BC9" blockId="12.[151,1437,1012,2031]" lastBlockId="13.[151,1437,151,2021]" lastPageId="13" lastPageNumber="335" pageId="12" pageNumber="334">
|
||
<keyStep id="30C774B8FFA40A00FF4AC303FA0A0BC9" lastPageId="13" lastPageNumber="335" pageId="12" pageNumber="334">
|
||
<keyLead id="30C2CF28FFA40A00FF4AC303FA0A0BC9" lastPageId="13" lastPageNumber="335" pageId="12" pageNumber="334">
|
||
6 Head and pronotum unarmed in both sexes, the latter simply convex; ratio of head width across eyes to interocular space at eye canthus greater than 1.7:1 (
|
||
<bibRefCitation id="EFA211ECFFA40A01FE38C321FD130C1D" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[485,645,1892,1914]" pageId="12" pageNumber="334" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA40A01FDC6C321FDDA0C1D" box="[539,588,1892,1913]" italics="true" pageId="12" pageNumber="334">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.11); mandibles extending beyond clypeus, outer sides straight and apically upturned; male protarsi thickened, with large asymmetrical claws, protarsomere 5 as long as 1–4 combined; metatibia not greatly enlarged, with slender, sharp spurs in male and short blunt spurs in female; propygidium without stridulatory area; light brownish yellow, shiny, pronotum and elytra with symmetrical and sometimes vaguely indicated dark brown markings,
|
||
<quantity id="4CCBC1F8FFA40A01FABDC3FEFE910C8A" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="12" pageNumber="334" unit="mm" value="13.5" valueMax="15.0" valueMin="12.0">12–15 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA40A01FE46C39CFDA80C8A" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[411,574,2009,2031]" pageId="12" pageNumber="334" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA40A01FE0FC39FFD920C8A" box="[466,516,2009,2031]" italics="true" pageId="12" pageNumber="334">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 55K;
|
||
<bibRefCitation id="EFA211ECFFA40A01FD6FC39CFC5E0C8A" author="Hutchinson, P. M. & Allsopp, P. G." box="[690,968,2009,2031]" pageId="12" pageNumber="334" pagination="1 - 14" refId="ref19412" refString="Hutchinson, P. M. & Allsopp, P. G. (2019) Revisiting Nephrodopus Sharp, 1873, with the description of a new species and a female (Coleoptera: Scarabaeidae: Dynastinae). Australian Entomologist, 46, 1 - 14." type="journal article" year="2019">Allsopp & Hutchinson 2019</bibRefCitation>
|
||
: fig. 1);
|
||
<collectingRegion id="49F7A2FFFFA40A01FBC4C39CFAB00C8B" box="[1049,1318,2009,2031]" country="Australia" name="Australian Capital Territory" pageId="12" pageNumber="334">Australian Capital Territory</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA40A01FAEDC39CFA0A0C8B" box="[1328,1436,2009,2031]" country="Australia" name="New South Wales" pageId="12" pageNumber="334">New South</collectingRegion>
|
||
Wales,
|
||
<collectingRegion id="49F7A2FFFFA50A00FEF5C4D2FE0A0BC9" box="[296,412,151,173]" country="Australia" name="Queensland" pageId="13" pageNumber="335">Queensland</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA50A00FE78C4D2FDAC0BC9" box="[421,570,151,173]" country="Australia" name="South Australia" pageId="13" pageNumber="335">South Australia</collectingRegion>
|
||
..................................... Cyclocephalini:
|
||
<taxonomicName id="4C33179EFFA50A00FB4FC4D2FA0A0BC9" ID-CoL="3XVL" authority="Dejean, 1821" authorityName="Dejean" authorityYear="1821" box="[1170,1436,151,173]" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="335" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA50A00FB4FC4D2FA800BC8" box="[1170,1302,151,172]" italics="true" pageId="13" pageNumber="335">Cyclocephala</emphasis>
|
||
Dejean, 1821
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</keyStep>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA50A00FF4AC4F0FA0A0A1E" blockId="13.[151,1437,151,2021]" pageId="13" pageNumber="335">
|
||
<keyLead id="30C2CF28FFA50A00FF4AC4F0FA0A0A1E" pageId="13" pageNumber="335">
|
||
- Head and pronotum with more-or-less distinct frontoclypeal suture, tubercles, horns, impressions, or hollows; if unarmed or armature not obvious, then ratio of head width across eyes to interocular space at eye canthus less than 1.4:1; and either propygidium with stridulating area (
|
||
<taxonomicName id="4C33179EFFA50A00FD53C4AAFC840A60" ID-CoL="62V4V" authorityName="Burmeister" authorityYear="1847" box="[654,786,239,260]" class="Insecta" family="Scarabaeidae" genus="Heteronychus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="335" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA50A00FD53C4AAFC840A60" box="[654,786,239,260]" italics="true" pageId="13" pageNumber="335">Heteronychus</emphasis>
|
||
</taxonomicName>
|
||
) or elytral striae very obvious (
|
||
<emphasis id="B947B00FFFA50A00FB86C4AAFD8F0A46" italics="true" pageId="13" pageNumber="335">
|
||
<taxonomicName id="4C33179EFFA50A00FB86C4AAFA9A0A60" ID-CoL="5Z6B" authorityName="Carne" authorityYear="1985" box="[1115,1292,239,260]" class="Insecta" family="Scarabaeidae" genus="Neocorynophyllus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="335" phylum="Arthropoda" rank="genus">Neocorynophyllus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C33179EFFA50A00FAC4C4AAFA0F0A60" ID-CoL="PDB" authorityName="Blackburn" authorityYear="1889" box="[1305,1433,239,260]" class="Insecta" family="Scarabaeidae" genus="Adoryphorus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="335" phylum="Arthropoda" rank="genus">Adoryphorus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C33179EFFA50A00FF3FC548FE100A46" ID-CoL="WS6" authorityName="Arrow" authorityYear="1908" box="[226,390,269,290]" class="Insecta" family="Scarabaeidae" genus="Anomalomorpha" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="335" phylum="Arthropoda" rank="genus">Anomalomorpha</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C33179EFFA50A00FE53C548FD8F0A46" ID-CoL="8SSQQ" baseAuthorityName="Hutchinson & Allsopp" baseAuthorityYear="2021" box="[398,537,269,290]" class="Insecta" family="Scarabaeidae" genus="Erbmahcedius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="335" phylum="Arthropoda" rank="genus">Erbmahcedius</taxonomicName>
|
||
</emphasis>
|
||
) or sides of clypeus deeply emarginate before ocular canthi (
|
||
<taxonomicName id="4C33179EFFA50A00FB86C548FB450A46" ID-CoL="63GZ8" authorityName="Boisduval" authorityYear="1835" box="[1115,1235,269,290]" class="Insecta" family="Scarabaeidae" genus="Oryctoderus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="335" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA50A00FB86C548FB450A46" box="[1115,1235,269,290]" italics="true" pageId="13" pageNumber="335">Oryctoderus</emphasis>
|
||
</taxonomicName>
|
||
); mandibles various; male protarsi thickened with asymmetrical claws or not; propygidium with or without stridulatory area; colour more or less uniform, brown to black, without pronotal and elytral colour pattern, sometimes bicoloured; length often greater than
|
||
<quantity id="4CCBC1F8FFA50A00FA91C502FA190A39" box="[1356,1423,327,349]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="335" unit="mm" value="15.0">15 mm</quantity>
|
||
................................................................................................... 7
|
||
</keyLead>
|
||
</paragraph>
|
||
<keyStep id="30C774B8FFA50A00FF4AC5C6FA0A0901" pageId="13" pageNumber="335">
|
||
<paragraph id="8B8C6C1DFFA50A00FF4AC5C6FA0A0A8B" blockId="13.[151,1437,151,2021]" pageId="13" pageNumber="335">
|
||
<keyLead id="30C2CF28FFA50A00FF4AC5C6FA0A0A8B" pageId="13" pageNumber="335">
|
||
7 Meso- and metatibial apices either with slight median projection separating cilia from spurs (
|
||
<bibRefCitation id="EFA211ECFFA50A00FBB3C5C7FA870AF3" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[1134,1297,386,408]" pageId="13" pageNumber="335" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA50A00FB78C5C6FB410AF3" box="[1189,1239,386,408]" italics="true" pageId="13" pageNumber="335">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.12) or with lateral projection and few cilia (
|
||
<bibRefCitation id="EFA211ECFFA50A00FD9CC5DAFD490AD1" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[577,735,415,437]" pageId="13" pageNumber="335" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA50A00FDABC5E5FD300AD0" box="[630,678,415,437]" italics="true" pageId="13" pageNumber="335">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.13); head with 1 or 2 tubercles or if without, then sides of clypeus deeply emarginate in front of ocular canthi; pronotum simply convex or with small depression and tubercle; elytra impunctate or with random punctation; propygidium without stridulatory area … Oryctoderini................................. 8
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA50A00FF4AC5BDFA0A0901" blockId="13.[151,1437,151,2021]" pageId="13" pageNumber="335">
|
||
<keyLead id="30C2CF28FFA50A00FF4AC5BDFA0A0901" pageId="13" pageNumber="335">
|
||
- Meso- and metatibial apices truncate or only slightly produced, cilia continuous with spurs or rarely entirely absent (
|
||
<bibRefCitation id="EFA211ECFFA50A00FAEAC5B2FE80094E" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." pageId="13" pageNumber="335" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA50A00FAB0C5BDFA0B0968" box="[1389,1437,503,525]" italics="true" pageId="13" pageNumber="335">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.14); head and pronotum with more-or-less distinct frontoclypeal suture, tubercles, horns, impressions or hollows; if unarmed or armature not obvious then either propygidium with stridulating area or elytral striae very obvious; propygidium with or without stridulating area … Pentodontini........................................................... 10
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFA50A00FF4AC628FA0A0871" pageId="13" pageNumber="335">
|
||
<paragraph id="8B8C6C1DFFA50A00FF4AC628FA0A09D9" blockId="13.[151,1437,151,2021]" pageId="13" pageNumber="335">
|
||
<keyLead id="30C2CF28FFA50A00FF4AC628FA0A09D9" pageId="13" pageNumber="335">
|
||
8 Clypeus with 2 distinct teeth; head with 2 low tubercles; meso- and metatibial apices with a lateral projection and only a few cilia (
|
||
<bibRefCitation id="EFA211ECFFA50A00FECBC6CFFE2409FB" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[278,434,650,672]" pageId="13" pageNumber="335" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA50A00FE96C6CEFEED09FB" box="[331,379,650,672]" italics="true" pageId="13" pageNumber="335">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.13); elytra with random ocellate punctures; red-brown,
|
||
<quantity id="4CCBC1F8FFA50A00FBD4C6CFFBF909C4" box="[1033,1135,650,672]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="13" pageNumber="335" unit="mm" value="21.5" valueMax="23.0" valueMin="20.0">20–23 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA50A00FB20C6CFFA0C09FB" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[1277,1434,650,672]" pageId="13" pageNumber="335" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA50A00FAEFC6CEFAF409FB" box="[1330,1378,650,672]" italics="true" pageId="13" pageNumber="335">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plare 59C;
|
||
<bibRefCitation id="EFA211ECFFA50A00FE96C6E2FDC909D9" author="Hutchinson, P. M. & Allsopp, P. G." box="[331,607,679,701]" pageId="13" pageNumber="335" pagination="1 - 14" refId="ref19412" refString="Hutchinson, P. M. & Allsopp, P. G. (2019) Revisiting Nephrodopus Sharp, 1873, with the description of a new species and a female (Coleoptera: Scarabaeidae: Dynastinae). Australian Entomologist, 46, 1 - 14." type="journal article" year="2019">Allsopp & Hutchinson 2019</bibRefCitation>
|
||
: fig. 16);
|
||
<collectingRegion id="49F7A2FFFFA50A00FD66C6E2FCBA09D9" box="[699,812,679,701]" country="Australia" name="Queensland" pageId="13" pageNumber="335">Queensland</collectingRegion>
|
||
...............................
|
||
<taxonomicName id="4C33179EFFA50A00FB71C6E2FA0A09D9" ID-CoL="63FJW" authority="Arrow, 1914" authorityName="Arrow" authorityYear="1914" box="[1196,1436,679,701]" class="Insecta" family="Scarabaeidae" genus="Onychionyx" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="335" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA50A00FB71C6E2FA8909D8" box="[1196,1311,679,700]" italics="true" pageId="13" pageNumber="335">Onychionyx</emphasis>
|
||
Arrow, 1914
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA50A00FF4AC680FA0A0871" blockId="13.[151,1437,151,2021]" pageId="13" pageNumber="335">
|
||
<keyLead id="30C2CF28FFA50A00FF4AC680FA0A0871" pageId="13" pageNumber="335">
|
||
- Clypeus rounded and truncate, with or without slight emargination; head with single tubercle or without tubercle; meso- and metatibial apices with slight median projection separating cilia from spurs (
|
||
<bibRefCitation id="EFA211ECFFA50A00FC61C6A7FBC80993" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[956,1118,738,760]" pageId="13" pageNumber="335" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA50A00FC2FC6A6FBB20993" box="[1010,1060,738,760]" italics="true" pageId="13" pageNumber="335">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.12); elytra impunctate or with some horseshoe shaped punctures.................................................................... 9
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFA50A00FF4AC758FA0A089B" pageId="13" pageNumber="335">
|
||
<paragraph id="8B8C6C1DFFA50A00FF4AC758FA0A08EF" blockId="13.[151,1437,151,2021]" pageId="13" pageNumber="335">
|
||
<keyLead id="30C2CF28FFA50A00FF4AC758FA0A08EF" pageId="13" pageNumber="335">
|
||
9 Clypeus deeply emarginate in front of ocular canthi, that of male deeply impressed; frons of male without tubercle; that of female with frontal tubercle feebly developed or not; pronotum without pit or tubercle in either sex; male protarsi thickened; elytra impunctate, smooth; dark brown-red,
|
||
<quantity id="4CCBC1F8FFA50A00FD58C71DFD7A0809" box="[645,748,856,877]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" metricValueMax="4.0" metricValueMin="3.3" pageId="13" pageNumber="335" unit="mm" value="36.5" valueMax="40.0" valueMin="33.0">33–40 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA50A00FCA6C712FB8F0809" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[891,1049,855,877]" pageId="13" pageNumber="335" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA50A00FC6CC71DFC770808" box="[945,993,855,877]" italics="true" pageId="13" pageNumber="335">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plates 54L, 56A;
|
||
<bibRefCitation id="EFA211ECFFA50A00FB19C712FE8008EE" author="Hutchinson, P. M. & Allsopp, P. G." pageId="13" pageNumber="335" pagination="1 - 14" refId="ref19412" refString="Hutchinson, P. M. & Allsopp, P. G. (2019) Revisiting Nephrodopus Sharp, 1873, with the description of a new species and a female (Coleoptera: Scarabaeidae: Dynastinae). Australian Entomologist, 46, 1 - 14." type="journal article" year="2019">Allsopp & Hutchinson 2019</bibRefCitation>
|
||
: fig. 17);
|
||
<collectingRegion id="49F7A2FFFFA50A00FEAFC730FE7508EF" box="[370,483,885,907]" country="Australia" name="Queensland" pageId="13" pageNumber="335">Queensland</collectingRegion>
|
||
.......................................................
|
||
<taxonomicName id="4C33179EFFA50A00FB5CC730FA0A08EF" ID-CoL="63GZ8" authority="Boisduval, 1835" authorityName="Boisduval" authorityYear="1835" box="[1153,1436,885,907]" class="Insecta" family="Scarabaeidae" genus="Oryctoderus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="335" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA50A00FB5CC730FB6F08EE" box="[1153,1273,885,906]" italics="true" pageId="13" pageNumber="335">Oryctoderus</emphasis>
|
||
Boisduval, 1835
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA50A00FF4AC7D6FA0A089B" blockId="13.[151,1437,151,2021]" pageId="13" pageNumber="335">
|
||
<keyLead id="30C2CF28FFA50A00FF4AC7D6FA0A089B" pageId="13" pageNumber="335">
|
||
- Clypeus not emarginate in front of ocular canthi; frons in both sexes with tubercle; pronotum in both sexes with tubercle and in males with small but distinct impression; male protarsi not thickened; elytra somewhat randomly punctate, with some horseshoeshaped punctures; black or pitch brown, venter red,
|
||
<quantity id="4CCBC1F8FFA50A00FD09C788FCAD0886" box="[724,827,973,994]" metricMagnitude="-2" metricUnit="m" metricValue="3.05" metricValueMax="3.3" metricValueMin="2.8" pageId="13" pageNumber="335" unit="mm" value="30.5" valueMax="33.0" valueMin="28.0">28–33 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA50A00FC10C788FBFB0886" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[973,1133,973,995]" pageId="13" pageNumber="335" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA50A00FBDEC78BFBA20886" box="[1027,1076,973,995]" italics="true" pageId="13" pageNumber="335">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 54A);
|
||
<collectingRegion id="49F7A2FFFFA50A00FB35C788FACF0887" box="[1256,1369,973,995]" country="Australia" name="Queensland" pageId="13" pageNumber="335">Queensland</collectingRegion>
|
||
.................................................................................
|
||
<taxonomicName id="4C33179EFFA50A00FB5FC7AFFA0A089B" ID-CoL="63GZ9" authority="Endrodi, 1978" authorityName="Endrodi" authorityYear="1978" box="[1154,1436,1002,1024]" class="Insecta" family="Scarabaeidae" genus="Oryctoderinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="335" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA50A00FB5FC7AFFA9B089B" box="[1154,1293,1002,1023]" italics="true" pageId="13" pageNumber="335">Oryctoderinus</emphasis>
|
||
Endrődi, 1978
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFA50A00FF4AC04DFA0A0FA9" pageId="13" pageNumber="335">
|
||
<paragraph id="8B8C6C1DFFA50A00FF4AC04DFA0A0F11" blockId="13.[151,1437,151,2021]" pageId="13" pageNumber="335">
|
||
<keyLead id="30C2CF28FFA50A00FF4AC04DFA0A0F11" pageId="13" pageNumber="335">
|
||
10 Protibia bidentate, apical tooth elongate (
|
||
<bibRefCitation id="EFA211ECFFA50A00FDAAC042FC880F79" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[631,798,1031,1053]" pageId="13" pageNumber="335" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA50A00FD72C04DFD740F78" box="[687,738,1031,1053]" italics="true" pageId="13" pageNumber="335">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.15); antennae with 9 antennomeres, club scarcely longer in male than in female; mentum not compressed; pronotum and elytra conspicuously punctate; legs very short and stout; metafemora bisulcate (
|
||
<bibRefCitation id="EFA211ECFFA50A00FE1CC007FDFC0F33" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[449,618,1090,1112]" pageId="13" pageNumber="335" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA50A00FE27C006FDB80F33" box="[506,558,1090,1112]" italics="true" pageId="13" pageNumber="335">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.14); black,
|
||
<quantity id="4CCBC1F8FFA50A00FCF2C007FC1B0F3C" box="[815,909,1090,1112]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="13" pageNumber="335" unit="mm" value="10.0" valueMax="12.0" valueMin="8.0">8–12 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA50A00FBFAC007FB590F33" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[1063,1231,1090,1112]" pageId="13" pageNumber="335" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA50A00FB82C006FB050F33" box="[1119,1171,1090,1112]" italics="true" pageId="13" pageNumber="335">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 54P);
|
||
<collectingRegion id="49F7A2FFFFA50A00FA93C006FEAC0F11" country="Australia" name="Western Australia" pageId="13" pageNumber="335">Western Australia</collectingRegion>
|
||
............................................................. Pseudoryctina:
|
||
<taxonomicName id="4C33179EFFA50A00FB67C025FA0A0F11" ID-CoL="63DQX" authority="Sharp, 1878" authorityName="Sharp" authorityYear="1878" box="[1210,1436,1119,1141]" class="Insecta" family="Scarabaeidae" genus="Neocnecus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="335" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA50A00FB67C025FAB40F11" box="[1210,1314,1120,1141]" italics="true" pageId="13" pageNumber="335">Neocnecus</emphasis>
|
||
Sharp, 1878
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA50A00FF4AC038FA0A0FA9" blockId="13.[151,1437,151,2021]" pageId="13" pageNumber="335">
|
||
<keyLead id="30C2CF28FFA50A00FF4AC038FA0A0FA9" pageId="13" pageNumber="335">
|
||
- Protibia tridentate, or if occasionally bidentate (some
|
||
<taxonomicName id="4C33179EFFA50A00FD26C038FCEE0FF6" ID-CoL="PDB" authorityName="Blackburn" authorityYear="1889" box="[763,888,1149,1170]" class="Insecta" family="Scarabaeidae" genus="Adoryphorus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="335" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA50A00FD26C038FCEE0FF6" box="[763,888,1149,1170]" italics="true" pageId="13" pageNumber="335">Adoryphorus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C33179EFFA50A00FC5AC038FB940FF6" ID-CoL="3MKX" authorityName="Hope" authorityYear="1837" box="[903,1026,1149,1170]" class="Insecta" family="Scarabaeidae" genus="Cheiroplatys" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="335" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA50A00FC5AC038FB940FF6" box="[903,1026,1149,1170]" italics="true" pageId="13" pageNumber="335">Cheiroplatys</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C33179EFFA50A00FBCCC038FB1B0FF6" authorityName="Allsopp & Hutchinson" authorityYear="2023" box="[1041,1165,1149,1170]" class="Insecta" family="Scarabaeidae" genus="Philcarneum" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="335" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA50A00FBCCC038FB1B0FF6" box="[1041,1165,1149,1170]" italics="true" pageId="13" pageNumber="335">Philcarneum</emphasis>
|
||
</taxonomicName>
|
||
), then apical tooth broadly rounded and poorly separated from second tooth; antennae with 9 or 10 antennomeres, with club of various lengths; mentum compressed or not; other characters variable............................................................... 11
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFA50A00FF4AC090FA0A0ED3" pageId="13" pageNumber="335">
|
||
<paragraph id="8B8C6C1DFFA50A00FF4AC090FA0A0E19" blockId="13.[151,1437,151,2021]" pageId="13" pageNumber="335">
|
||
<keyLead id="30C2CF28FFA50A00FF4AC090FA0A0E19" pageId="13" pageNumber="335">
|
||
11 Mentum and ligula compressed to form thin vertical lamina divided by ligular suture; apical maxillary palpomere remarkably enlarged and specialised (
|
||
<bibRefCitation id="EFA211ECFFA50A00FE3CC0B7FD1C0E63" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[481,650,1266,1288]" pageId="13" pageNumber="335" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA50A00FDC7C0B6FDD80E63" box="[538,590,1266,1288]" italics="true" pageId="13" pageNumber="335">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.16); pronotum of males with anterior horn and pair of lateral horns or tubercles, with an excavation extending to base; pronotum of females evenly rounded, without horns or tubercles; scutellum very large; metathoracic legs stout, the metafemora unisulcate; yellow, reddish-brown or black species, often bicoloured,
|
||
<quantity id="4CCBC1F8FFA50A00FABDC168FE910E04" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="13" pageNumber="335" unit="mm" value="14.0" valueMax="16.0" valueMin="12.0">12–16 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA50A00FE44C10FFD3B0E3B" author="Hutchinson, P. M. & Allsopp, P. G." box="[409,685,1354,1376]" pageId="13" pageNumber="335" pagination="1 - 14" refId="ref19412" refString="Hutchinson, P. M. & Allsopp, P. G. (2019) Revisiting Nephrodopus Sharp, 1873, with the description of a new species and a female (Coleoptera: Scarabaeidae: Dynastinae). Australian Entomologist, 46, 1 - 14." type="journal article" year="2019">Hutchinson & Allsopp 2019</bibRefCitation>
|
||
: figs. 4, 9, 14, 20, 25, 31);
|
||
<collectingRegion id="49F7A2FFFFA50A00FC73C10FFBF60E04" box="[942,1120,1354,1376]" country="Australia" name="Northern Territory" pageId="13" pageNumber="335">Northern Territory</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA50A00FBB4C10FFB4B0E04" box="[1129,1245,1354,1376]" country="Australia" name="Queensland" pageId="13" pageNumber="335">Queensland</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA50A00FB3BC10EFA070E04" box="[1254,1425,1354,1376]" country="Australia" name="Western Australia" pageId="13" pageNumber="335">Western Australia</collectingRegion>
|
||
................................................................... Pseudoryctina:
|
||
<taxonomicName id="4C33179EFFA50A00FB7FC122FA0A0E19" ID-CoL="637X" authority="Sharp, 1873" authorityName="Sharp" authorityYear="1873" box="[1186,1436,1383,1405]" class="Insecta" family="Scarabaeidae" genus="Nephrodopus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="335" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA50A00FB7FC122FAB40E18" box="[1186,1314,1383,1404]" italics="true" pageId="13" pageNumber="335">Nephrodopus</emphasis>
|
||
Sharp, 1873
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA50A00FF4AC1C0FA0A0ED3" blockId="13.[151,1437,151,2021]" pageId="13" pageNumber="335">
|
||
<keyLead id="30C2CF28FFA50A00FF4AC1C0FA0A0ED3" pageId="13" pageNumber="335">- Mentum flat or strongly compressed but not produced to form vertical lamina; apical maxillary palpomere not enlarged; pronotum various.................................................................................... 12</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFA50A00FF4AC185FA0A0E96" pageId="13" pageNumber="335">
|
||
<paragraph id="8B8C6C1DFFA50A00FF4AC185FA0A0EB1" blockId="13.[151,1437,151,2021]" box="[151,1436,1471,1493]" pageId="13" pageNumber="335">
|
||
<keyLead id="30C2CF28FFA50A00FF4AC185FA0A0EB1" box="[151,1436,1471,1493]" pageId="13" pageNumber="335">12 Meso- and metatibial apices without cilia, glabrous......................................................... 13</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA50A00FF4AC198FA0A0E96" blockId="13.[151,1437,151,2021]" box="[151,1436,1501,1523]" pageId="13" pageNumber="335">
|
||
<keyLead id="30C2CF28FFA50A00FF4AC198FA0A0E96" box="[151,1436,1501,1523]" pageId="13" pageNumber="335">- Meso- and metatibial apices with numerous cilia, continuous with spurs......................................... 14</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFA50A00FF4AC1BFFA0B0DDB" pageId="13" pageNumber="335">
|
||
<paragraph id="8B8C6C1DFFA50A00FF4AC1BFFA0A0D03" blockId="13.[151,1437,151,2021]" pageId="13" pageNumber="335">
|
||
<keyLead id="30C2CF28FFA50A00FF4AC1BFFA0A0D03" pageId="13" pageNumber="335">
|
||
13 Head plate-like, flattened, more-or-less vertical and coarsely, somewhat rugosely punctate (
|
||
<bibRefCitation id="EFA211ECFFA50A00FB8FC1BFFB650D6B" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[1106,1267,1530,1552]" pageId="13" pageNumber="335" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA50A00FB55C1BEFB2F0D6B" box="[1160,1209,1530,1552]" italics="true" pageId="13" pageNumber="335">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.17); front of clypeus straight with a small tubercle at either side; mandibles visible in front of clypeus; red-brown,
|
||
<quantity id="4CCBC1F8FFA50A00FB25C25DFAF40D49" box="[1272,1378,1560,1581]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="13" pageNumber="335" unit="mm" value="15.0" valueMax="20.0" valueMin="10.0">10–20 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA50A00FEEBC270FE400D2E" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[310,470,1589,1611]" pageId="13" pageNumber="335" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA50A00FEB1C273FE0B0D2E" box="[364,413,1589,1611]" italics="true" pageId="13" pageNumber="335">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 54H;
|
||
<bibRefCitation id="EFA211ECFFA50A00FD95C270FCCD0D2E" author="Hutchinson, P. M. & Allsopp, P. G." box="[584,859,1589,1611]" pageId="13" pageNumber="335" pagination="1 - 14" refId="ref19412" refString="Hutchinson, P. M. & Allsopp, P. G. (2019) Revisiting Nephrodopus Sharp, 1873, with the description of a new species and a female (Coleoptera: Scarabaeidae: Dynastinae). Australian Entomologist, 46, 1 - 14." type="journal article" year="2019">Allsopp & Hutchinson 2019</bibRefCitation>
|
||
: fig. 12);
|
||
<collectingRegion id="49F7A2FFFFA50A00FC6AC270FBF30D2F" box="[951,1125,1589,1611]" country="Australia" name="New South Wales" pageId="13" pageNumber="335">New South Wales</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA50A00FBB3C270FA8E0D2F" box="[1134,1304,1589,1611]" country="Australia" name="Western Australia" pageId="13" pageNumber="335">Western Australia</collectingRegion>
|
||
............................................................................ Pentodontina:
|
||
<taxonomicName id="4C33179EFFA50A00FB4DC217FA0A0D03" ID-CoL="7TPM" authority="Hope, 1837" authorityName="Hope" authorityYear="1837" box="[1168,1436,1618,1640]" class="Insecta" family="Scarabaeidae" genus="Temnorhynchus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="335" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA50A00FB4DC217FAB10D03" box="[1168,1319,1618,1639]" italics="true" pageId="13" pageNumber="335">Temnorhynchus</emphasis>
|
||
Hope, 1837
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA50A00FF4AC235FA0B0DDB" blockId="13.[151,1437,151,2021]" pageId="13" pageNumber="335">
|
||
<keyLead id="30C2CF28FFA50A00FF4AC235FA0B0DDB" pageId="13" pageNumber="335">
|
||
- Head not plate-like, not vertical, with concentric sculpturing, impunctate (
|
||
<figureCitation id="13087098FFA50A00FC7AC22AFC770DE1" box="[935,993,1647,1669]" captionStart="FIGURES 1–5" captionStartId="5.[151,264,1301,1325]" captionTargetBox="[151,1435,309,1278]" captionTargetId="figure-89@5.[151,1435,309,1278]" captionTargetPageId="5" captionText="FIGURES 1–5. Philcarneum aenigma Allsopp & Hutchinson new genus, new species, holotype female: 1, dorsal view; 2, ventral view; 3, lateral view; 4, head, dorsal view; 5, labels." figureDoi="http://doi.org/10.5281/zenodo.8391694" httpUri="https://zenodo.org/record/8391694/files/figure.png" pageId="13" pageNumber="335">Fig. 4</figureCitation>
|
||
); front of clypeus rounded, indented medially, without tubercles; mandibles concealed by clypeus; dorsum (
|
||
<figureCitation id="13087098FFA50A00FCE8C2C8FCE60DC6" box="[821,880,1677,1699]" captionStart="FIGURES 1–5" captionStartId="5.[151,264,1301,1325]" captionTargetBox="[151,1435,309,1278]" captionTargetId="figure-89@5.[151,1435,309,1278]" captionTargetPageId="5" captionText="FIGURES 1–5. Philcarneum aenigma Allsopp & Hutchinson new genus, new species, holotype female: 1, dorsal view; 2, ventral view; 3, lateral view; 4, head, dorsal view; 5, labels." figureDoi="http://doi.org/10.5281/zenodo.8391694" httpUri="https://zenodo.org/record/8391694/files/figure.png" pageId="13" pageNumber="335">Fig. 1</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFA211ECFFA50A00FCA6C2C8FB040DC6" author="Hutchinson, P. M. & Allsopp, P. G." box="[891,1170,1677,1699]" pageId="13" pageNumber="335" pagination="1 - 14" refId="ref19412" refString="Hutchinson, P. M. & Allsopp, P. G. (2019) Revisiting Nephrodopus Sharp, 1873, with the description of a new species and a female (Coleoptera: Scarabaeidae: Dynastinae). Australian Entomologist, 46, 1 - 14." type="journal article" year="2019">Allsopp & Hutchinson 2019</bibRefCitation>
|
||
: fig. 13); brown with black markings,
|
||
<quantity id="4CCBC1F8FFA50A00FE9AC2EFFE380DA4" box="[327,430,1706,1728]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="13" pageNumber="335" unit="mm" value="13.5" valueMax="14.0" valueMin="13.0">13–14 mm</quantity>
|
||
long;
|
||
<collectingRegion id="49F7A2FFFFA50A00FE31C2EEFD010DA4" box="[492,663,1706,1728]" country="Australia" name="New South Wales" pageId="13" pageNumber="335">New South Wales</collectingRegion>
|
||
.............. Cheiroplatina:
|
||
<taxonomicName id="4C33179EFFA50A00FC0CC2EFFABB0DA4" authority="Allsopp & Hutchinson" authorityName="Allsopp & Hutchinson" authorityYear="2023" box="[977,1325,1706,1728]" class="Insecta" family="Scarabaeidae" genus="Philcarneum" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="335" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B947B00FFFA50A00FC0CC2EFFBDB0DDB" box="[977,1101,1706,1727]" italics="true" pageId="13" pageNumber="335">Philcarneum</emphasis>
|
||
Allsopp & Hutchinson
|
||
</taxonomicName>
|
||
<emphasis id="B947B00FFFA50A00FAEFC2EFFA0B0DDB" bold="true" box="[1330,1437,1706,1727]" pageId="13" pageNumber="335">
|
||
<taxonomicNameLabel id="A2740D74FFA50A00FAEFC2EFFA0B0DDB" box="[1330,1437,1706,1727]" pageId="13" pageNumber="335" rank="genus">new genus</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFA50A00FF4AC28DFA0A0C14" pageId="13" pageNumber="335">
|
||
<paragraph id="8B8C6C1DFFA50A00FF4AC28DFA0A0C51" blockId="13.[151,1437,151,2021]" pageId="13" pageNumber="335">
|
||
<keyLead id="30C2CF28FFA50A00FF4AC28DFA0A0C51" pageId="13" pageNumber="335">
|
||
14 Propygidium enlarged, its posterior edge convex and its disc covered with numerous fine transverse stridulating ridges (
|
||
<bibRefCitation id="EFA211ECFFA50A00FA8BC282FEA30D9E" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." pageId="13" pageNumber="335" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA50A00FA56C28DFF6D0D9E" italics="true" pageId="13" pageNumber="335">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.18); head plate-like, vertical and smooth, forming a horn in males (
|
||
<bibRefCitation id="EFA211ECFFA50A00FBD6C2A0FB3A0D9E" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[1035,1196,1765,1787]" pageId="13" pageNumber="335" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA50A00FB9CC2A3FBE40D9E" box="[1089,1138,1765,1787]" italics="true" pageId="13" pageNumber="335">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.19); apical labial palpomere globular; glossy reddish brown to reddish black,
|
||
<quantity id="4CCBC1F8FFA50A00FCFDC347FC110C7C" box="[800,903,1794,1816]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" metricValueMax="3.2" metricValueMin="1.2" pageId="13" pageNumber="335" unit="mm" value="22.0" valueMax="32.0" valueMin="12.0">12–32 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA50A00FBC5C347FB210C73" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[1048,1207,1794,1816]" pageId="13" pageNumber="335" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA50A00FB90C346FBE80C73" box="[1101,1150,1794,1816]" italics="true" pageId="13" pageNumber="335">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 55D);
|
||
<collectingRegion id="49F7A2FFFFA50A00FAECC346FE890C51" country="Australia" name="New South Wales" pageId="13" pageNumber="335">New South Wales</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA50A00FEF5C35AFE4C0C51" box="[296,474,1823,1845]" country="Australia" name="Northern Territory" pageId="13" pageNumber="335">Northern Territory</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA50A00FE3EC35AFDC10C51" box="[483,599,1823,1845]" country="Australia" name="Queensland" pageId="13" pageNumber="335">Queensland</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA50A00FDBDC365FC9D0C51" box="[608,779,1823,1845]" country="Australia" name="Western Australia" pageId="13" pageNumber="335">Western Australia</collectingRegion>
|
||
........................... Dipelicina:
|
||
<taxonomicName id="4C33179EFFA50A00FB17C35AFA0A0C51" ID-CoL="632SP" authority="Hope, 1845" authorityName="Hope" authorityYear="1845" box="[1226,1436,1823,1845]" class="Insecta" family="Scarabaeidae" genus="Dipelicus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="335" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA50A00FB17C35AFAB00C50" box="[1226,1318,1823,1844]" italics="true" pageId="13" pageNumber="335">Dipelicus</emphasis>
|
||
Hope, 1845
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA50A00FF4AC378FA0A0C14" blockId="13.[151,1437,151,2021]" pageId="13" pageNumber="335">
|
||
<keyLead id="30C2CF28FFA50A00FF4AC378FA0A0C14" pageId="13" pageNumber="335">- Propygidium not enlarged, with or without stridulating ridges; if ridges present, then these are either scattered or form a pair of paramedian bands; head without vertical plate; apical labial palpomere cylindrical................................. 15</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<paragraph id="8B8C6C1DFFA50A03FF4AC332FA0A0BAF" blockId="13.[151,1437,151,2021]" lastBlockId="14.[151,1437,151,2021]" lastPageId="14" lastPageNumber="336" pageId="13" pageNumber="335">
|
||
<keyStep id="30C774B8FFA50A03FF4AC332FA0A0BAF" lastPageId="14" lastPageNumber="336" pageId="13" pageNumber="335">
|
||
<keyLead id="30C2CF28FFA50A03FF4AC332FA0A0BAF" lastPageId="14" lastPageNumber="336" pageId="13" pageNumber="335">15 Galeae reduced to small conical or pyramidal pieces, not toothed; mandibles frequently incapable of apposition, usually exposed beyond clypeus; mentum moderately to highly compressed; clypeus often with a deep, apical, truncate face; males and females sexually dimorphic with regard to the antennal club; male antennal club enlarged and antennomeres leaf-like and flattened, longer than shaft; female antennal club (where known) shorter than shaft; male head and pronotum often with bizarre armature; female head and pronotum (where known) unarmed; females either represented in collections by few specimens or unknown; only males can be successfully keyed out in couplets 16–26 … Pseudoryctina (part)............................... 16</keyLead>
|
||
</keyStep>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA60A03FF4AC496FA0A0A1E" blockId="14.[151,1437,151,2021]" pageId="14" pageNumber="336">
|
||
<keyLead id="30C2CF28FFA60A03FF4AC496FA0A0A1E" pageId="14" pageNumber="336">
|
||
- Galeae usually strongly developed; mandibles functional, sometimes concealed beneath clypeus; mentum not appreciably compressed; clypeus without a deep, apical, truncate face (except in
|
||
<taxonomicName id="4C33179EFFA60A03FCB1C4AAFC7F0A60" ID-CoL="3ZZY" authorityName="MacLeay" authorityYear="1819" box="[876,1001,239,260]" class="Insecta" family="Scarabaeidae" genus="Dasygnathus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="336" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA60A03FCB1C4AAFC7F0A60" box="[876,1001,239,260]" italics="true" pageId="14" pageNumber="336">Dasygnathus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C33179EFFA60A03FBCBC4AAFB230A60" ID-CoL="5Z9T" authorityName="Carne" authorityYear="1957" box="[1046,1205,239,260]" class="Insecta" family="Scarabaeidae" genus="Neodasygnathus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="336" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA60A03FBCBC4AAFB230A60" box="[1046,1205,239,260]" italics="true" pageId="14" pageNumber="336">Neodasygnathus</emphasis>
|
||
</taxonomicName>
|
||
); males and females not sexually dimorphic with regard to the antennal club; antennal club subequal in both sexes, not enlarged as above, antennomeres not as flattened, shorter than or equal in length to shaft, usually shorter than funicle; male and female pronotum similar or different; females and males more-or-less equally represented in collections; combined male and female characters used from couplet 27.......................................................................................... 27
|
||
</keyLead>
|
||
</paragraph>
|
||
<keyStep id="30C774B8FFA60A03FF4AC5C7FA0A0923" pageId="14" pageNumber="336">
|
||
<paragraph id="8B8C6C1DFFA60A03FF4AC5C7FA0A0A94" blockId="14.[151,1437,151,2021]" pageId="14" pageNumber="336">
|
||
<keyLead id="30C2CF28FFA60A03FF4AC5C7FA0A0A94" pageId="14" pageNumber="336">
|
||
16 Frontoclypeal suture absent; clypeus only slightly narrowed anteriorly, with thickened bead all around; setae on mentum very long, dense and extending well past front edge of clypeus; antennal lamellae widest near apex; male protarsal claws asymmetrical; black,
|
||
<quantity id="4CCBC1F8FFA60A03FE6DC5F8FE650AB6" box="[432,499,445,466]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="14" pageNumber="336" unit="mm" value="14.0">14 mm</quantity>
|
||
long; female unknown; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA60A03FCEFC5F8FC440AB6" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[818,978,445,467]" pageId="14" pageNumber="336" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA60A03FCB5C5F8FC0F0AB6" box="[872,921,445,466]" italics="true" pageId="14" pageNumber="336">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 54G);
|
||
<collectingRegion id="49F7A2FFFFA60A03FB90C5F8FB280AB7" box="[1101,1214,445,467]" country="Australia" name="Queensland" pageId="14" pageNumber="336">Queensland</collectingRegion>
|
||
.............................................................................................
|
||
<taxonomicName id="4C33179EFFA60A03FBAFC59FFA0A0A94" ID-CoL="5Z6B" authority="Carne, 1985" authorityName="Carne" authorityYear="1985" box="[1138,1436,474,496]" class="Insecta" family="Scarabaeidae" genus="Neocorynophyllus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="336" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA60A03FBAFC59FFAB70A8B" box="[1138,1313,474,495]" italics="true" pageId="14" pageNumber="336">Neocorynophyllus</emphasis>
|
||
Carne, 1985
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA60A03FF4AC5BDFA0A0923" blockId="14.[151,1437,151,2021]" pageId="14" pageNumber="336">
|
||
<keyLead id="30C2CF28FFA60A03FF4AC5BDFA0A0923" pageId="14" pageNumber="336">- Frontoclypeal suture usually well developed, often giving rise to tubercles or horns of various sizes; if suture weak or not evident then clypeus narrowed to reflexed apex; setae on mentum shorter; antennal lamellae usually widest at middle; male claws symmetrical................................................................................... 17</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFA60A03FF4AC615FA0A0871" pageId="14" pageNumber="336">
|
||
<paragraph id="8B8C6C1DFFA60A03FF4AC615FA0A09BF" blockId="14.[151,1437,151,2021]" pageId="14" pageNumber="336">
|
||
<keyLead id="30C2CF28FFA60A03FF4AC615FA0A09BF" pageId="14" pageNumber="336">
|
||
17 Clypeus deflexed ventrad, either evenly curved or abruptly declivous, not from clypeofrontal suture, but from a ridge joining antennal angles, and narrowed distally, sides sinuate to very narrow anterior face that is distinctly upturned; frontoclypeal suture subobsolete; apical pygidial ridge glabrous; pronotum with basal ridge continuous and with anteromedian tubercle and impression; dorsal surface conspicuously punctate; black,
|
||
<quantity id="4CCBC1F8FFA60A03FCD3C6EDFCE309D9" box="[782,885,680,701]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="1.9" metricValueMin="1.3" pageId="14" pageNumber="336" unit="mm" value="16.0" valueMax="19.0" valueMin="13.0">13–19 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA60A03FBDBC6E2FA8E09D9" author="Hutchinson, P. M. & Allsopp, P. G." box="[1030,1304,679,701]" pageId="14" pageNumber="336" pagination="428 - 448" refId="ref19461" refString="Hutchinson, P. M. & Allsopp, P. G. (2020) Revision of Epironastes Carne, 1957 (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Pseudoryctina) with description of the female, behaviours, and habitats. Zootaxa, 4852 (4), 428 - 448. https: // doi. org / 10.11646 / zootaxa. 4852.4.2" type="journal article" year="2020">Hutchinson & Allsopp 2020</bibRefCitation>
|
||
: figs. 1, 3, 11, 19, 21, 30, 32, 41, 43, 53;
|
||
<bibRefCitation id="EFA211ECFFA60A03FE03C680FDEA09BE" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[478,636,709,731]" pageId="14" pageNumber="336" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA60A03FDCFC680FDD509BE" box="[530,579,709,730]" italics="true" pageId="14" pageNumber="336">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 54O);
|
||
<collectingRegion id="49F7A2FFFFA60A03FD2AC680FC3409BF" box="[759,930,709,731]" country="Australia" name="Western Australia" pageId="14" pageNumber="336">Western Australia</collectingRegion>
|
||
......................
|
||
<taxonomicName id="4C33179EFFA60A03FB72C680FA0A09BF" ID-CoL="4DCC" authority="Carne, 1957" authorityName="Carne" authorityYear="1957" box="[1199,1436,709,731]" class="Insecta" family="Scarabaeidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="336" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA60A03FB72C680FAB709BE" box="[1199,1313,709,730]" italics="true" pageId="14" pageNumber="336">Epironastes</emphasis>
|
||
<bibRefCitation id="EFA211ECFFA60A03FAFAC680FA0A09BF" author="Carne, P. B." box="[1319,1436,709,731]" pageId="14" pageNumber="336" refId="ref19138" refString="Carne, P. B. (1957) Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne, 1957</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA60A03FF4AC6A6FA0A0871" blockId="14.[151,1437,151,2021]" pageId="14" pageNumber="336">
|
||
<keyLead id="30C2CF28FFA60A03FF4AC6A6FA0A0871" pageId="14" pageNumber="336">- Clypeus subrectangular, rounded or slightly narrowed, only rarely apically reflexed; with carinate or raised frontoclypeal suture, median tubercle or horn; other characters variable.......................................................... 18</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFA60A03FF4AC758FA0A08EF" pageId="14" pageNumber="336">
|
||
<paragraph id="8B8C6C1DFFA60A03FF4AC758FA0A082B" blockId="14.[151,1437,151,2021]" pageId="14" pageNumber="336">
|
||
<keyLead id="30C2CF28FFA60A03FF4AC758FA0A082B" pageId="14" pageNumber="336">18 Head with raised or carinate frontoclypeal suture, median tubercle or small to medium horn; if with horn then pronotum with anteromedial tubercle and impression not extending to posterior edge and not limited by an elevated transverse ridge..... 19</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA60A03FF4AC71DFA0A08EF" blockId="14.[151,1437,151,2021]" pageId="14" pageNumber="336">
|
||
<keyLead id="30C2CF28FFA60A03FF4AC71DFA0A08EF" pageId="14" pageNumber="336">- Head armed with a conspicuous horn; pronotum with either a major excavation extending to posterior edge or an excavation limited by a sharp elevated transverse ridge or sharp elevations................................................ 25</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFA60A03FF4AC7D7FA0A08A1" pageId="14" pageNumber="336">
|
||
<paragraph id="8B8C6C1DFFA60A03FF4AC7D7FA0A08C3" blockId="14.[151,1437,151,2021]" box="[151,1436,914,936]" pageId="14" pageNumber="336">
|
||
<keyLead id="30C2CF28FFA60A03FF4AC7D7FA0A08C3" box="[151,1436,914,936]" pageId="14" pageNumber="336">19 Head with carinate frontoclypeal suture only............................................................... 20</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA60A03FF4AC7F5FA0A08A1" blockId="14.[151,1437,151,2021]" box="[151,1436,943,965]" pageId="14" pageNumber="336">
|
||
<keyLead id="30C2CF28FFA60A03FF4AC7F5FA0A08A1" box="[151,1436,943,965]" pageId="14" pageNumber="336">- Head with raised frontoclypeal suture, median tubercle or small to medium horn.................................. 26</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFA60A03FF4AC788FA0A0F79" pageId="14" pageNumber="336">
|
||
<paragraph id="8B8C6C1DFFA60A03FF4AC788FA0A089B" blockId="14.[151,1437,151,2021]" pageId="14" pageNumber="336">
|
||
<keyLead id="30C2CF28FFA60A03FF4AC788FA0A089B" pageId="14" pageNumber="336">20 Anterior edge of pronotum narrowly membranous, at least on either side median third (where this is occupied by a horn-like process), as well as bearing a fringe of setae............................................................... 21</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA60A03FF4AC04DFA0A0F79" blockId="14.[151,1437,151,2021]" box="[151,1436,1031,1053]" pageId="14" pageNumber="336">
|
||
<keyLead id="30C2CF28FFA60A03FF4AC04DFA0A0F79" box="[151,1436,1031,1053]" pageId="14" pageNumber="336">- Anterior edge of pronotum entirely sclerotised, only with fringe of setae......................................... 23</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFA60A03FF4AC060FA0A0FCB" pageId="14" pageNumber="336">
|
||
<paragraph id="8B8C6C1DFFA60A03FF4AC060FA0A0FF7" blockId="14.[151,1437,151,2021]" pageId="14" pageNumber="336">
|
||
<keyLead id="30C2CF28FFA60A03FF4AC060FA0A0FF7" pageId="14" pageNumber="336">
|
||
21 Mandibles concealed beneath clypeus, the latter with sides convex; antennal club not much longer than funicle; legs moderately stout; metatibia contracted subapically; frontoclypeal ridge transverse, elevated; dorsal surfaces shiny and impunctate; pronotum with small, anteromedian tubercle and shallow depression; sutural stria linear; black, ~
|
||
<quantity id="4CCBC1F8FFA60A03FBE4C025FBEB0F11" box="[1081,1149,1120,1141]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="14" pageNumber="336" unit="mm" value="14.0">14 mm</quantity>
|
||
long; female unknown; male head and pronotum, dorsum (
|
||
<bibRefCitation id="EFA211ECFFA60A03FE26C038FD300FF7" author="Dechambre, R. - P." box="[507,678,1149,1171]" pageId="14" pageNumber="336" refId="ref19268" refString="Dechambre, R. - P. (2005) Dynastidae Australiens et Oceaniens. Les Coleopteres du Monde 30. Hillside Books, Canterbury, 132 pp." type="book" year="2005">Dechambre 2005</bibRefCitation>
|
||
: p. 67);
|
||
<collectingRegion id="49F7A2FFFFA60A03FD2EC038FC1E0FF7" box="[755,904,1149,1171]" country="Australia" name="South Australia" pageId="14" pageNumber="336">South Australia</collectingRegion>
|
||
.......................
|
||
<taxonomicName id="4C33179EFFA60A03FB44C038FA0A0FF7" ID-CoL="63H4T" authority="Carne, 1957" authorityName="Carne" authorityYear="1957" box="[1177,1436,1149,1171]" class="Insecta" family="Scarabaeidae" genus="Orthocavonus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="336" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA60A03FB44C038FAB70FF6" box="[1177,1313,1149,1170]" italics="true" pageId="14" pageNumber="336">Orthocavonus</emphasis>
|
||
<bibRefCitation id="EFA211ECFFA60A03FAFAC038FA0A0FF7" author="Carne, P. B." box="[1319,1436,1149,1171]" pageId="14" pageNumber="336" refId="ref19138" refString="Carne, P. B. (1957) Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne, 1957</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA60A03FF4AC0DEFA0A0FCB" blockId="14.[151,1437,151,2021]" box="[151,1436,1178,1200]" pageId="14" pageNumber="336">
|
||
<keyLead id="30C2CF28FFA60A03FF4AC0DEFA0A0FCB" box="[151,1436,1178,1200]" pageId="14" pageNumber="336">- Mandibles exposed beyond clypeus; antennal club much longer than funicle; other characters variable................. 22</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFA60A03FF4AC0FDFA0A0EB1" pageId="14" pageNumber="336">
|
||
<paragraph id="8B8C6C1DFFA60A03FF4AC0FDFA0A0E27" blockId="14.[151,1437,151,2021]" pageId="14" pageNumber="336">
|
||
<keyLead id="30C2CF28FFA60A03FF4AC0FDFA0A0E27" pageId="14" pageNumber="336">
|
||
22 Clypeus with sides rounded or sinuately narrowed towards apex; if rounded then epipleural setae sparse, not extending on to apical third of elytra; antennal club not much longer than shaft, kidney shaped (
|
||
<bibRefCitation id="EFA211ECFFA60A03FC36C090FB180F8E" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[1003,1166,1237,1259]" pageId="14" pageNumber="336" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA60A03FBFFC090FBC20F8E" box="[1058,1108,1237,1258]" italics="true" pageId="14" pageNumber="336">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.20); mentum flat or compressed anteriorly, strongly reflexed (
|
||
<bibRefCitation id="EFA211ECFFA60A03FDA4C0B7FC890E63" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[633,799,1266,1288]" pageId="14" pageNumber="336" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA60A03FD6DC0B6FD750E63" box="[688,739,1266,1288]" italics="true" pageId="14" pageNumber="336">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.20); metatibia strongly contracted subapically; pronotum with anteromedian horn or tubercle and impression, sometimes bounded by lateral horns; black,
|
||
<quantity id="4CCBC1F8FFA60A03FBABC155FB4B0E41" box="[1142,1245,1296,1317]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="2.1" metricValueMin="1.2" pageId="14" pageNumber="336" unit="mm" value="16.5" valueMax="21.0" valueMin="12.0">12–21 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA60A03FAB1C14AFEDA0E26" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." pageId="14" pageNumber="336" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA60A03FF3FC168FE850E26" box="[226,275,1325,1346]" italics="true" pageId="14" pageNumber="336">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 55C);
|
||
<collectingRegion id="49F7A2FFFFA60A03FE1BC168FDE20E27" box="[454,628,1325,1347]" country="Australia" name="New South Wales" pageId="14" pageNumber="336">New South Wales</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA60A03FDA0C168FC850E27" box="[637,787,1325,1347]" country="Australia" name="South Australia" pageId="14" pageNumber="336">South Australia</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA60A03FCC0C168FCFB0E27" box="[797,877,1325,1347]" country="Australia" name="Victoria" pageId="14" pageNumber="336">Victoria</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA60A03FCABC168FBB60E27" box="[886,1056,1325,1347]" country="Australia" name="Western Australia" pageId="14" pageNumber="336">Western Australia</collectingRegion>
|
||
..............
|
||
<taxonomicName id="4C33179EFFA60A03FB13C168FA0A0E27" ID-CoL="3JTF" authority="Sharp, 1875" authorityName="Sharp" authorityYear="1875" box="[1230,1436,1325,1347]" class="Insecta" family="Scarabaeidae" genus="Cavonus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="336" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA60A03FB13C168FAB40E26" box="[1230,1314,1325,1346]" italics="true" pageId="14" pageNumber="336">Cavonus</emphasis>
|
||
Sharp, 1875
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA60A03FF4AC10EFA0A0EB1" blockId="14.[151,1437,151,2021]" pageId="14" pageNumber="336">
|
||
<keyLead id="30C2CF28FFA60A03FF4AC10EFA0A0EB1" pageId="14" pageNumber="336">
|
||
- Clypeus rounded; antennal club much longer than shaft, its sides more-or-less subparallel; mentum highly compressed (
|
||
<bibRefCitation id="EFA211ECFFA60A03FAB1C10FFEDA0E19" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." pageId="14" pageNumber="336" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA60A03FF3FC12DFE850E18" box="[226,275,1383,1405]" italics="true" pageId="14" pageNumber="336">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.21); metatibia evenly dilated from femoral articulation to distal edge; pronotum with small, rarely strongly elevated, anteromedian tubercle plus an impression, or with surface evenly convex; epipleural setae dense, extending or nearly extending to elytral apices; reddish-brown to black,
|
||
<quantity id="4CCBC1F8FFA60A03FD05C1E7FCA80EDC" box="[728,830,1442,1464]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.7" metricValueMin="1.1" pageId="14" pageNumber="336" unit="mm" value="14.0" valueMax="17.0" valueMin="11.0">11–17 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA60A03FC0CC1E7FBE40ED3" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[977,1138,1442,1464]" pageId="14" pageNumber="336" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA60A03FBD5C1E6FBAF0ED3" box="[1032,1081,1442,1464]" italics="true" pageId="14" pageNumber="336">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 54F);
|
||
<collectingRegion id="49F7A2FFFFA60A03FB36C1E6FA0F0EDC" box="[1259,1433,1442,1464]" country="Australia" name="New South Wales" pageId="14" pageNumber="336">New South Wales</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA60A03FF3FC1FAFEC00EB1" box="[226,342,1471,1493]" country="Australia" name="Queensland" pageId="14" pageNumber="336">Queensland</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA60A03FE82C1FAFE600EB1" box="[351,502,1471,1493]" country="Australia" name="South Australia" pageId="14" pageNumber="336">South Australia</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA60A03FE22C1FAFDD90EB1" box="[511,591,1471,1493]" country="Australia" name="Victoria" pageId="14" pageNumber="336">Victoria</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA60A03FD85C185FC950EB1" box="[600,771,1471,1493]" country="Australia" name="Western Australia" pageId="14" pageNumber="336">Western Australia</collectingRegion>
|
||
.............................
|
||
<taxonomicName id="4C33179EFFA60A03FBBEC185FADE0EB1" ID-CoL="7TCH" authority="Sharp, 1873" authorityName="Sharp" authorityYear="1873" box="[1123,1352,1471,1493]" class="Insecta" family="Scarabaeidae" genus="Teinogenys" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="336" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA60A03FBBEC185FB580EB1" box="[1123,1230,1472,1493]" italics="true" pageId="14" pageNumber="336">Teinogenys</emphasis>
|
||
Sharp, 1873
|
||
</taxonomicName>
|
||
(in part)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFA60A03FF4AC198FA0A0DC6" pageId="14" pageNumber="336">
|
||
<paragraph id="8B8C6C1DFFA60A03FF4AC198FA0A0D2E" blockId="14.[151,1437,151,2021]" pageId="14" pageNumber="336">
|
||
<keyLead id="30C2CF28FFA60A03FF4AC198FA0A0D2E" pageId="14" pageNumber="336">
|
||
23 Maxillary palps short and inconspicuous, not much longer than labial palps; mandibles and labrum concealed; lateral pronotal horns irregular and turret-like; metatibia unicarinate; spurs straight and subequal in length; metatarsomere 1 strongly dilated and twice the length of metatarsomere 2; reddish brown, bicolored or black,
|
||
<quantity id="4CCBC1F8FFA60A03FC1AC25DFBB90D49" box="[967,1071,1560,1581]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.4" metricValueMin="1.4" pageId="14" pageNumber="336" unit="mm" value="19.0" valueMax="24.0" valueMin="14.0">14–24 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA60A03FB1EC252FAF20D49" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[1219,1380,1559,1581]" pageId="14" pageNumber="336" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA60A03FB24C25DFABD0D48" box="[1273,1323,1559,1581]" italics="true" pageId="14" pageNumber="336">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 53F);
|
||
<collectingRegion id="49F7A2FFFFA60A03FEC1C270FC200D2F" box="[284,950,1589,1611]" isEnumeration="true" pageId="14" pageNumber="336">New South Wales, Northern Territory, Queensland, Western Australia</collectingRegion>
|
||
...................
|
||
<taxonomicName id="4C33179EFFA60A03FB7FC270FA0A0D2E" ID-CoL="63R44" authority="Sharp, 1873" authorityName="Sharp" authorityYear="1873" box="[1186,1436,1589,1611]" class="Insecta" family="Scarabaeidae" genus="Pseudoryctes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="336" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA60A03FB7FC270FAB40D2E" box="[1186,1314,1589,1610]" italics="true" pageId="14" pageNumber="336">Pseudoryctes</emphasis>
|
||
Sharp, 1873
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA60A03FF4AC216FA0A0DC6" blockId="14.[151,1437,151,2021]" pageId="14" pageNumber="336">
|
||
<keyLead id="30C2CF28FFA60A03FF4AC216FA0A0DC6" pageId="14" pageNumber="336">- Maxillary palps elongate, much longer than labial palps; mandibles and labrum more-or-less exposed; pronotum with simple, lateral horns or lateral elevated angles; metatibia bicarinate; metatarsomere 1 not dilated and equal in length to metatarsomere 2 ................................................................................................. 24</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFA60A03FF4AC2EFFA0A0C36" pageId="14" pageNumber="336">
|
||
<paragraph id="8B8C6C1DFFA60A03FF4AC2EFFA0A0D9E" blockId="14.[151,1437,151,2021]" pageId="14" pageNumber="336">
|
||
<keyLead id="30C2CF28FFA60A03FF4AC2EFFA0A0D9E" pageId="14" pageNumber="336">
|
||
24 Antennal club very large, sinuate, bitexturous (
|
||
<bibRefCitation id="EFA211ECFFA60A03FD78C2EFFCD30DDB" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[677,837,1706,1728]" pageId="14" pageNumber="336" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA60A03FD06C2EEFC9A0DDB" box="[731,780,1706,1728]" italics="true" pageId="14" pageNumber="336">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.22); pronotum with lateral elevated angles (
|
||
<bibRefCitation id="EFA211ECFFA60A03FAEBC2EFFE800DB9" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." pageId="14" pageNumber="336" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA60A03FAB1C2EEFA0B0DDB" box="[1388,1437,1706,1728]" italics="true" pageId="14" pageNumber="336">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.22) and median third of basal edge strongly produced over scutellum; dark reddish brown,
|
||
<quantity id="4CCBC1F8FFA60A03FB67C282FAB60DB9" box="[1210,1312,1735,1757]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="14" pageNumber="336" unit="mm" value="16.5" valueMax="17.0" valueMin="16.0">16–17 mm</quantity>
|
||
long; female unknown; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA60A03FE41C2A0FDAA0D9E" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[412,572,1765,1787]" pageId="14" pageNumber="336" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA60A03FE0FC2A0FD950D9E" box="[466,515,1765,1786]" italics="true" pageId="14" pageNumber="336">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 55H);
|
||
<collectingRegion id="49F7A2FFFFA60A03FD6AC2A0FCF70D9F" box="[695,865,1765,1787]" country="Australia" name="Western Australia" pageId="14" pageNumber="336">Western Australia</collectingRegion>
|
||
.....................
|
||
<taxonomicName id="4C33179EFFA60A03FBBCC2A0FA0A0D9E" ID-CoL="6ZGB" authority="Blackburn, 1890" authorityName="Blackburn" authorityYear="1890" box="[1121,1436,1765,1787]" class="Insecta" family="Scarabaeidae" genus="Pseudocavonus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="336" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA60A03FBBCC2A0FB600D9E" box="[1121,1270,1765,1786]" italics="true" pageId="14" pageNumber="336">Pseudocavonus</emphasis>
|
||
Blackburn, 1890
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA60A03FF4AC346FA0A0C36" blockId="14.[151,1437,151,2021]" pageId="14" pageNumber="336">
|
||
<keyLead id="30C2CF28FFA60A03FF4AC346FA0A0C36" pageId="14" pageNumber="336">
|
||
- Antennal club not strongly sinuate, unitexturous; pronotum with simple lateral horns and basal edge normally rounded; reddish brown, bicoloured or black,
|
||
<quantity id="4CCBC1F8FFA60A03FE20C365FDF10C51" box="[509,615,1824,1845]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.7" metricValueMin="1.2" pageId="14" pageNumber="336" unit="mm" value="19.5" valueMax="27.0" valueMin="12.0">12–27 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA60A03FCDCC35AFC3F0C51" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[769,937,1823,1845]" pageId="14" pageNumber="336" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA60A03FCE4C365FCFB0C50" box="[825,877,1823,1845]" italics="true" pageId="14" pageNumber="336">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 55J);
|
||
<collectingRegion id="49F7A2FFFFA60A03FBF8C365FC950C37" isEnumeration="true" pageId="14" pageNumber="336">New South Wales, Northern Territory, Queensland, South Australia, Victoria, Western Australia</collectingRegion>
|
||
...............................
|
||
<taxonomicName id="4C33179EFFA60A03FBA8C378FA0A0C36" ID-CoL="3HQG" authority="Ozdikmen, 2009" authorityName="Ozdikmen" authorityYear="2009" box="[1141,1436,1853,1876]" class="Insecta" family="Scarabaeidae" genus="Carneoryctes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="336" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA60A03FBA8C378FB600C36" box="[1141,1270,1853,1874]" italics="true" pageId="14" pageNumber="336">Carneoryctes</emphasis>
|
||
Özdikmen, 2009
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFA60A02FF4AC31FFA0A0BAE" lastPageId="15" lastPageNumber="337" pageId="14" pageNumber="336">
|
||
<paragraph id="8B8C6C1DFFA60A03FF4AC31FFA0A0CA3" blockId="14.[151,1437,151,2021]" pageId="14" pageNumber="336">
|
||
<keyLead id="30C2CF28FFA60A03FF4AC31FFA0A0CA3" pageId="14" pageNumber="336">
|
||
25 Head with horn extending back over more than half of pronotum (
|
||
<bibRefCitation id="EFA211ECFFA60A03FCA9C31FFB8D0C0B" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[884,1051,1882,1904]" pageId="14" pageNumber="336" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA60A03FC71C31EFC760C0B" box="[940,992,1882,1904]" italics="true" pageId="14" pageNumber="336">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.23); pronotum with excavation not extending to posterior edge and bounded posteriorly by a sharp elevated transverse ridge or sharp elevations; basal area of pygidium glabrous; reddish brown, black or bicolored,
|
||
<quantity id="4CCBC1F8FFA60A03FCC9C3D0FCEA0CCE" box="[788,892,1941,1962]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="2.0" metricValueMin="1.2" pageId="14" pageNumber="336" unit="mm" value="16.0" valueMax="20.0" valueMin="12.0">12–20 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA60A03FBCDC3D0FB240CCE" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[1040,1202,1941,1963]" pageId="14" pageNumber="336" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA60A03FB9AC3D0FBEE0CCE" box="[1095,1144,1941,1962]" italics="true" pageId="14" pageNumber="336">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 55G);
|
||
<collectingRegion id="49F7A2FFFFA60A03FAEDC3D0FE890CAC" country="Australia" name="New South Wales" pageId="14" pageNumber="336">New South Wales</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA60A03FEF5C3F7FE0A0CAC" box="[296,412,1970,1992]" country="Australia" name="Queensland" pageId="14" pageNumber="336">Queensland</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA60A03FE78C3F7FDAA0CAC" box="[421,572,1970,1992]" country="Australia" name="South Australia" pageId="14" pageNumber="336">South Australia</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA60A03FD98C3F7FD040CAC" box="[581,658,1970,1992]" country="Australia" name="Victoria" pageId="14" pageNumber="336">Victoria</collectingRegion>
|
||
....................................
|
||
<taxonomicName id="4C33179EFFA60A03FB98C3F7FADE0CAC" ID-CoL="62NWH" authority="Hope, 1845" authorityName="Hope" authorityYear="1845" box="[1093,1352,1970,1992]" class="Insecta" family="Scarabaeidae" genus="Corynophyllus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="336" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA60A03FB98C3F7FB440CA3" box="[1093,1234,1970,1991]" italics="true" pageId="14" pageNumber="336">Corynophyllus</emphasis>
|
||
Hope, 1845
|
||
</taxonomicName>
|
||
(in part)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA60A02FF4AC395FA0A0BAE" blockId="14.[151,1437,151,2021]" lastBlockId="15.[151,1437,151,2021]" lastPageId="15" lastPageNumber="337" pageId="14" pageNumber="336">
|
||
<keyLead id="30C2CF28FFA60A02FF4AC395FA0A0BAE" lastPageId="15" lastPageNumber="337" pageId="14" pageNumber="336">
|
||
- Head with horn not or only just extending back over pronotum; pronotum with excavation extending to posterior edge and bounded laterally by blunt lateral elevations; basal area of pygidium markedly setose; black,
|
||
<quantity id="4CCBC1F8FFA70A02FB80C4D2FB550BC9" box="[1117,1219,151,173]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="15" pageNumber="337" unit="mm" value="16.5" valueMax="18.0" valueMin="15.0">15–18 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA70A02FA88C4D2FEA30BAE" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." pageId="15" pageNumber="337" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA70A02FA56C4DDFF6D0BAE" italics="true" pageId="15" pageNumber="337">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plates 54N, 55A);
|
||
<collectingRegion id="49F7A2FFFFA70A02FE33C4F0FC630BAF" box="[494,1013,181,203]" isEnumeration="true" pageId="15" pageNumber="337">Northern Territory, South Australia, Western Australia</collectingRegion>
|
||
.........
|
||
<taxonomicName id="4C33179EFFA70A02FBB8C4F0FA0A0BAE" ID-CoL="6PJV" authority="Blackburn, 1890" authorityName="Blackburn" authorityYear="1890" box="[1125,1436,181,203]" class="Insecta" family="Scarabaeidae" genus="Phylliocephala" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="337" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA70A02FBB8C4F0FB600BAE" box="[1125,1270,181,202]" italics="true" pageId="15" pageNumber="337">Phylliocephala</emphasis>
|
||
Blackburn, 1890
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFA70A02FF4AC497FA0A0969" pageId="15" pageNumber="337">
|
||
<paragraph id="8B8C6C1DFFA70A02FF4AC497FA0A0A39" blockId="15.[151,1437,151,2021]" pageId="15" pageNumber="337">
|
||
<keyLead id="30C2CF28FFA70A02FF4AC497FA0A0A39" pageId="15" pageNumber="337">
|
||
26 Clypeus evenly rounded with moderately deep, truncate face; frontoclypeal ridge transverse or posteriorly angulate, usually giving rise to median tubercle, which is in line with or behind anterior edge of eye; pronotum strongly transverse, nontuberculate, evenly convex or with slight, submarginal impression in front; antennal club with sides more-or-less subparallel; apical pygidial ridge glabrous; black or reddish brown,
|
||
<quantity id="4CCBC1F8FFA70A02FCD4C56FFCE70A24" box="[777,881,298,320]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.7" metricValueMin="1.1" pageId="15" pageNumber="337" unit="mm" value="14.0" valueMax="17.0" valueMin="11.0">11–17 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA70A02FBD7C56FFB260A5B" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[1034,1200,298,320]" pageId="15" pageNumber="337" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA70A02FB9FC56EFBE30A5B" box="[1090,1141,298,320]" italics="true" pageId="15" pageNumber="337">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 54F);
|
||
<collectingRegion id="49F7A2FFFFA70A02FAF2C56EFCDF0A39" isEnumeration="true" pageId="15" pageNumber="337">New South Wales, Queensland, South Australia, Victoria, Western Australia</collectingRegion>
|
||
.......................
|
||
<taxonomicName id="4C33179EFFA70A02FBBEC50DFADE0A39" ID-CoL="7TCH" authority="Sharp, 1873" authorityName="Sharp" authorityYear="1873" box="[1123,1352,327,349]" class="Insecta" family="Scarabaeidae" genus="Teinogenys" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="337" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA70A02FBBEC50DFB580A39" box="[1123,1230,328,349]" italics="true" pageId="15" pageNumber="337">Teinogenys</emphasis>
|
||
Sharp, 1873
|
||
</taxonomicName>
|
||
(in part)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA70A02FF4AC520FA0A0969" blockId="15.[151,1437,151,2021]" pageId="15" pageNumber="337">
|
||
<keyLead id="30C2CF28FFA70A02FF4AC520FA0A0969" pageId="15" pageNumber="337">
|
||
- Clypeus elongate, narrowed anteriorly with truncate apex or if rarely evenly rounded then without truncate apical face; frontoclypeal ridge transverse, giving rise to median elevation, tubercle or distinct, sometimes bifurcate, horn all of which are in front of anterior edge of eye; antennal club greatly enlarged and of characteristic shape (
|
||
<bibRefCitation id="EFA211ECFFA70A02FBE5C5DAFB430AD1" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[1080,1237,415,437]" pageId="15" pageNumber="337" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA70A02FBB0C5E5FB0B0AD0" box="[1133,1181,415,437]" italics="true" pageId="15" pageNumber="337">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.24); pronotal margin with median tubercle or node; pronotum with anteromedian tubercle and impression; apical pygidial ridge setose; reddish brown, black or bicolored,
|
||
<quantity id="4CCBC1F8FFA70A02FDF0C59FFD040A94" box="[557,658,474,496]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="2.0" metricValueMin="1.2" pageId="15" pageNumber="337" unit="mm" value="16.0" valueMax="20.0" valueMin="12.0">12–20 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA70A02FCFCC59FFC2B0A8B" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[801,957,474,496]" pageId="15" pageNumber="337" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA70A02FC88C59EFC130A8B" box="[853,901,474,496]" italics="true" pageId="15" pageNumber="337">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 55G);
|
||
<collectingRegion id="49F7A2FFFFA70A02FBE8C59EFB760A94" box="[1077,1248,474,496]" country="Australia" name="New South Wales" pageId="15" pageNumber="337">New South Wales</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA70A02FB35C59FFACA0A94" box="[1256,1372,474,496]" country="Australia" name="Queensland" pageId="15" pageNumber="337">Queensland</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA70A02FAB9C59FFEAA0969" country="Australia" name="South Australia" pageId="15" pageNumber="337">South Australia</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA70A02FE98C5B2FE040969" box="[325,402,503,525]" country="Australia" name="Victoria" pageId="15" pageNumber="337">Victoria</collectingRegion>
|
||
..........................................................
|
||
<taxonomicName id="4C33179EFFA70A02FB98C5B2FADE0969" ID-CoL="62NWH" authority="Hope, 1845" authorityName="Hope" authorityYear="1845" box="[1093,1352,503,525]" class="Insecta" family="Scarabaeidae" genus="Corynophyllus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="337" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA70A02FB98C5B2FB440968" box="[1093,1234,503,524]" italics="true" pageId="15" pageNumber="337">Corynophyllus</emphasis>
|
||
Hope, 1845
|
||
</taxonomicName>
|
||
(in part)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFA70A02FF4AC650FA0A09D9" pageId="15" pageNumber="337">
|
||
<paragraph id="8B8C6C1DFFA70A02FF4AC650FA0A0901" blockId="15.[151,1437,151,2021]" pageId="15" pageNumber="337">
|
||
<keyLead id="30C2CF28FFA70A02FF4AC650FA0A0901" pageId="15" pageNumber="337">27 Mandibles either evenly rounded, slightly scalloped or straight sided, exposed beyond or beside clypeus or concealed; propygidium without a trace of stridulating ridges; last abdominal ventrite sometimes with well-defined transverse sulcus, especially in female … Cheiroplatina.................................................................... 27</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA70A02FF4AC628FA0A09D9" blockId="15.[151,1437,151,2021]" pageId="15" pageNumber="337">
|
||
<keyLead id="30C2CF28FFA70A02FF4AC628FA0A09D9" pageId="15" pageNumber="337">- Mandibles almost always conspicuously sinuate, bidentate or tridentate and exposed beyond or beside clypeus; propygidium with either short, scattered stridulating ridges or a pair of paramedian stridulatory bands; last abdominal ventrite without transverse sulcus … Pentodontina....................................................................... 37</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFA70A02FF4AC680FA0A0856" pageId="15" pageNumber="337">
|
||
<paragraph id="8B8C6C1DFFA70A02FF4AC680FA0A0993" blockId="15.[151,1437,151,2021]" pageId="15" pageNumber="337">
|
||
<keyLead id="30C2CF28FFA70A02FF4AC680FA0A0993" pageId="15" pageNumber="337">28 Clypeus (viewed dorsally) strongly contracted towards apex with sides concave; concave sides with characteristic vertical faces; mandibles somewhat straight-sided and exposed beyond or beside clypeus.................................. 29</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA70A02FF4AC745FA0A0856" blockId="15.[151,1437,151,2021]" pageId="15" pageNumber="337">
|
||
<keyLead id="30C2CF28FFA70A02FF4AC745FA0A0856" pageId="15" pageNumber="337">- Clypeus (viewed dorsally) broadly triangular, subrectangular or trapezoidal, with sides straight; mandibles rounded or slightly scalloped, exposed beyond or beside clypeus or concealed.................................................... 30</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFA70A02FF4AC77FFA0A0F11" pageId="15" pageNumber="337">
|
||
<paragraph id="8B8C6C1DFFA70A02FF4AC77FFA0A0887" blockId="15.[151,1437,151,2021]" pageId="15" pageNumber="337">
|
||
<keyLead id="30C2CF28FFA70A02FF4AC77FFA0A0887" pageId="15" pageNumber="337">
|
||
29 Frontoclypeal suture posteriorly angulate, giving rise to triangular elevation or large horn in male; pronotum somewhat pentagonal, especially in female, its sides contracted towards anterolateral angles; anterior edge of pronotum membranous only across median third; pronotum in male with large impression or excavation and without anteromedian tubercle; antennae with 10 antennomeres; base of pygidium setose; aedeagus with simple parameres; reddish brown to reddish black,
|
||
<quantity id="4CCBC1F8FFA70A02FAE8C7D7FA0A08CC" box="[1333,1436,914,936]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.5" metricValueMin="1.6" pageId="15" pageNumber="337" unit="mm" value="20.5" valueMax="25.0" valueMin="16.0">16–25 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA70A02FEB1C7EAFD9C08A1" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[364,522,943,965]" pageId="15" pageNumber="337" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA70A02FE7CC7F5FE4708A0" box="[417,465,943,965]" italics="true" pageId="15" pageNumber="337">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plates 55E, 55N);
|
||
<collectingRegion id="49F7A2FFFFA70A02FD63C7F5FE1B0887" isEnumeration="true" pageId="15" pageNumber="337">New South Wales, Northern Territory, Queensland, South Australia, Victoria, Western Australia</collectingRegion>
|
||
.....................................................................
|
||
<taxonomicName id="4C33179EFFA70A02FB13C788FA0A0887" ID-CoL="65NT" authority="Sharp, 1875" authorityName="Sharp" authorityYear="1875" box="[1230,1436,973,995]" class="Insecta" family="Scarabaeidae" genus="Novapus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="337" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA70A02FB13C788FAB40886" box="[1230,1314,973,994]" italics="true" pageId="15" pageNumber="337">Novapus</emphasis>
|
||
Sharp, 1875
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA70A02FF4AC7AEFA0A0F11" blockId="15.[151,1437,151,2021]" pageId="15" pageNumber="337">
|
||
<keyLead id="30C2CF28FFA70A02FF4AC7AEFA0A0F11" pageId="15" pageNumber="337">
|
||
- Frontoclypeal suture usually straight, rarely tuberculate; pronotum with sides more evenly curved towards anterolateral angles; anterior edge of pronotum wholly membranous; pronotum in male usually with anteromedian tubercle and small impression; antennae usually with 9 antennomeres (very rarely with 8 or 10); base of pygidium glabrous; aedeagus with complex parameres; dark reddish brown to black,
|
||
<quantity id="4CCBC1F8FFA70A02FDDDC007FDCA0F3C" box="[512,604,1090,1112]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.6" metricValueMin="0.9" pageId="15" pageNumber="337" unit="mm" value="12.5" valueMax="16.0" valueMin="9.0">9–16 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA70A02FD2CC007FC050F33" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[753,915,1090,1112]" pageId="15" pageNumber="337" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA70A02FCFAC006FCCF0F33" box="[807,857,1090,1112]" italics="true" pageId="15" pageNumber="337">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 54D);
|
||
<collectingRegion id="49F7A2FFFFA70A02FBCCC006FE1B0F11" isEnumeration="true" pageId="15" pageNumber="337">New South Wales, Queensland, Victoria, Western Australia</collectingRegion>
|
||
................................................................
|
||
<taxonomicName id="4C33179EFFA70A02FB4FC01AFA0A0F11" ID-CoL="82HY" authority="Burmeister, 1847" authorityName="Burmeister" authorityYear="1847" box="[1170,1436,1119,1141]" class="Insecta" family="Scarabaeidae" genus="Trissodon" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="337" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA70A02FB4FC01AFB670F10" box="[1170,1265,1119,1140]" italics="true" pageId="15" pageNumber="337">Trissodon</emphasis>
|
||
Burmeister, 1847
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFA70A02FF4AC038FA0A0E63" pageId="15" pageNumber="337">
|
||
<paragraph id="8B8C6C1DFFA70A02FF4AC038FA0A0FCB" blockId="15.[151,1437,151,2021]" pageId="15" pageNumber="337">
|
||
<keyLead id="30C2CF28FFA70A02FF4AC038FA0A0FCB" pageId="15" pageNumber="337">30 Clypeus broadly triangular, sides straight; mandibles exposed beyond or beside clypeus or concealed; galea strongly developed and conspicuously toothed; palps short................................................................... 31</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA70A02FF4AC0FDFA0A0E63" blockId="15.[151,1437,151,2021]" pageId="15" pageNumber="337">
|
||
<keyLead id="30C2CF28FFA70A02FF4AC0FDFA0A0E63" pageId="15" pageNumber="337">
|
||
- Clypeus broadly subrectangular or trapezoidal, sides straight or rounded; mandibles usually concealed beneath clypeus (exposed at sides in some
|
||
<taxonomicName id="4C33179EFFA70A02FE5DC090FD890F8E" ID-CoL="5Z9T" authorityName="Carne" authorityYear="1957" box="[384,543,1237,1258]" class="Insecta" family="Scarabaeidae" genus="Neodasygnathus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="337" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA70A02FE5DC090FD890F8E" box="[384,543,1237,1258]" italics="true" pageId="15" pageNumber="337">Neodasygnathus</emphasis>
|
||
</taxonomicName>
|
||
); galea reduced, either weakly toothed or having only a small, conical process terminating in a pencil of setae; palps elongate or not.................................................................... 32
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFA70A02FF4AC155FA0A0D2E" pageId="15" pageNumber="337">
|
||
<paragraph id="8B8C6C1DFFA70A02FF4AC155FA0A0EFE" blockId="15.[151,1437,151,2021]" pageId="15" pageNumber="337">
|
||
<keyLead id="30C2CF28FFA70A02FF4AC155FA0A0EFE" pageId="15" pageNumber="337">
|
||
31 Head completely unarmed; clypeus with oblique, sometimes serrate, anterior edge; galea strongly toothed; mandibles concealed or slightly exposed at sides of clypeus; pronotum with anterior tubercle and (usually) deep impression in male; armature reduced but still discernible in most females; protibia with 3 or 2 blunt teeth; metatarsomere 1 widened, flattened and with distinct longitudinal carina externally; deep reddish brown to black,
|
||
<quantity id="4CCBC1F8FFA70A02FCBAC122FC5B0E19" box="[871,973,1383,1405]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" metricValueMax="2.8" metricValueMin="1.5" pageId="15" pageNumber="337" unit="mm" value="21.5" valueMax="28.0" valueMin="15.0">15–28 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA70A02FB86C122FB610E19" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[1115,1271,1383,1405]" pageId="15" pageNumber="337" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA70A02FB4DC12DFB290E18" box="[1168,1215,1383,1405]" italics="true" pageId="15" pageNumber="337">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 55O);
|
||
<collectingRegion id="49F7A2FFFFA70A02FAB2C12DFCA10EFF" isEnumeration="true" pageId="15" pageNumber="337">New South Wales, Queensland, South Australia, Tasmania, Victoria</collectingRegion>
|
||
...............................
|
||
<taxonomicName id="4C33179EFFA70A02FB76C1C0FA0A0EFE" ID-CoL="3MKX" authority="Hope, 1837" authorityName="Hope" authorityYear="1837" box="[1195,1436,1413,1434]" class="Insecta" family="Scarabaeidae" genus="Cheiroplatys" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="337" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA70A02FB76C1C0FAB00EFE" box="[1195,1318,1413,1434]" italics="true" pageId="15" pageNumber="337">Cheiroplatys</emphasis>
|
||
Hope, 1837
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA70A02FF4AC1E6FA0A0D2E" blockId="15.[151,1437,151,2021]" pageId="15" pageNumber="337">
|
||
<keyLead id="30C2CF28FFA70A02FF4AC1E6FA0A0D2E" pageId="15" pageNumber="337">
|
||
- Head armed with low tubercle or small horn; clypeus with anterior edge slightly upturned, never oblique; galea weakly toothed; mandibles strongly exposed beyond clypeus; pronotum with anterior (usually) and posterior foveae (always) developed in both sexes, without anterior tubercle; protibia with 3 sharp teeth; metatarsomere 1 more rounded, not carinate externally; elytra sometimes costate; reddish black to black,
|
||
<quantity id="4CCBC1F8FFA70A02FD5DC1BFFD710D74" box="[640,743,1530,1552]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.6" metricValueMin="1.2" pageId="15" pageNumber="337" unit="mm" value="19.0" valueMax="26.0" valueMin="12.0">12–26 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA70A02FCA7C1BFFB8D0D6B" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[890,1051,1530,1552]" pageId="15" pageNumber="337" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA70A02FC6DC1BEFC740D6B" box="[944,994,1530,1552]" italics="true" pageId="15" pageNumber="337">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 55I;
|
||
<bibRefCitation id="EFA211ECFFA70A02FB5BC1BFFAAF0D6B" author="Reid, C. A. M. & Tees, N. A." box="[1158,1337,1530,1552]" pageId="15" pageNumber="337" pagination="563 - 570" refId="ref19857" refString="Reid, C. A. M. & Tees, N. A. (2023) A new, but possibly extinct, species of Semanopterus Hope, 1847 from Lord Howe Island, in the southwestern Pacific Ocean (Coleoptera: Scarabaeidae: Dynastinae). Zootaxa, 5306 (5), 563 - 570. https: // doi. org / 10.11646 / zootaxa. 5306.5.4" type="journal article" year="2023">Reid & Tees 2023</bibRefCitation>
|
||
: figs. 1–5, 7–10);
|
||
<collectingRegion id="49F7A2FFFFA70A02FEFAC25DFE420D49" box="[295,468,1559,1581]" country="Australia" name="New South Wales" pageId="15" pageNumber="337">New South Wales</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA70A02FE00C252FD060D49" box="[477,656,1559,1581]" country="Australia" name="Northern Territory" pageId="15" pageNumber="337">Northern Territory</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA70A02FD44C252FC9B0D49" box="[665,781,1559,1581]" country="Australia" name="Queensland" pageId="15" pageNumber="337">Queensland</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA70A02FCCBC252FC3B0D49" box="[790,941,1559,1581]" country="Australia" name="South Australia" pageId="15" pageNumber="337">South Australia</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA70A02FC6AC252FB910D49" box="[951,1031,1559,1581]" country="Australia" name="Victoria" pageId="15" pageNumber="337">Victoria</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA70A02FBD2C25DFB2A0D49" box="[1039,1212,1559,1581]" country="Australia" name="Western Australia" pageId="15" pageNumber="337">Western Australia</collectingRegion>
|
||
, Lord Howe Island.................................................................................
|
||
<taxonomicName id="4C33179EFFA70A02FB43C270FA0A0D2E" ID-CoL="7FC2" authority="Hope, 1847" authorityName="Hope" authorityYear="1847" box="[1182,1436,1589,1610]" class="Insecta" family="Scarabaeidae" genus="Semanopterus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="337" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA70A02FB43C270FAB00D2E" box="[1182,1318,1589,1610]" italics="true" pageId="15" pageNumber="337">Semanopterus</emphasis>
|
||
Hope, 1847
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFA70A02FF4AC217FA0A0C0B" pageId="15" pageNumber="337">
|
||
<paragraph id="8B8C6C1DFFA70A02FF4AC217FA0A0DDB" blockId="15.[151,1437,151,2021]" pageId="15" pageNumber="337">
|
||
<keyLead id="30C2CF28FFA70A02FF4AC217FA0A0DDB" pageId="15" pageNumber="337">
|
||
32 Length
|
||
<quantity id="4CCBC1F8FFA70A02FEF3C217FE0C0D0C" box="[302,410,1618,1640]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="3.2" metricValueMin="1.7" pageId="15" pageNumber="337" unit="mm" value="24.5" valueMax="32.0" valueMin="17.0">17–32 mm</quantity>
|
||
; head armed with tubercle or small horn on straight or recurved frontoclypeal suture; if tubercle hard to see or absent (some
|
||
<taxonomicName id="4C33179EFFA70A02FE76C22AFDBE0DE0" ID-CoL="3ZZY" authorityName="MacLeay" authorityYear="1819" box="[427,552,1647,1668]" class="Insecta" family="Scarabaeidae" genus="Dasygnathus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="337" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA70A02FE76C22AFDBE0DE0" box="[427,552,1647,1668]" italics="true" pageId="15" pageNumber="337">Dasygnathus</emphasis>
|
||
</taxonomicName>
|
||
), then base of mentum deeply hollowed or cleft; pronotum sometimes with a submarginal groove; elytra never conspicuously guttered; galea weakly toothed; pronotum of male with excavation of various development, sometimes with anteromedian impression and tubercle in both sexes............................................ 33
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFA70A02FF4AC28DFA0A0C0B" blockId="15.[151,1437,151,2021]" pageId="15" pageNumber="337">
|
||
<keyLead id="30C2CF28FFA70A02FF4AC28DFA0A0C0B" pageId="15" pageNumber="337">
|
||
- Length
|
||
<quantity id="4CCBC1F8FFA70A02FEF0C28DFE1B0DB9" box="[301,397,1736,1757]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.1" metricValueMin="0.9" pageId="15" pageNumber="337" unit="mm" value="15.0" valueMax="21.0" valueMin="9.0">9–21 mm</quantity>
|
||
; head either unarmed, or with small swelling on vertex (
|
||
<taxonomicName id="4C33179EFFA70A02FC64C282FBA00DB8" ID-CoL="PDB" authorityName="Blackburn" authorityYear="1889" box="[953,1078,1735,1756]" class="Insecta" family="Scarabaeidae" genus="Adoryphorus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="337" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA70A02FC64C282FBA00DB8" box="[953,1078,1735,1756]" italics="true" pageId="15" pageNumber="337">Adoryphorus</emphasis>
|
||
</taxonomicName>
|
||
), or with small tubercle on recurved frontoclypeal suture; if with small tubercle on recurved frontoclypeal suture then pronotum with a submarginal groove and elytra conspicuously guttered and epipleura horizontal in anterior half (
|
||
<taxonomicName id="4C33179EFFA70A02FC4FC347FBA50C73" ID-CoL="WS6" authorityName="Arrow" authorityYear="1908" box="[914,1075,1794,1815]" class="Insecta" family="Scarabaeidae" genus="Anomalomorpha" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="337" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA70A02FC4FC347FBA50C73" box="[914,1075,1794,1815]" italics="true" pageId="15" pageNumber="337">Anomalomorpha</emphasis>
|
||
</taxonomicName>
|
||
); base of mentum never deeply cleft; galea with only a small conical process terminating in a pencil of setae; pronotum either evenly convex with micropunctate surface (
|
||
<taxonomicName id="4C33179EFFA70A02FEE8C378FE240C36" ID-CoL="PDB" authorityName="Blackburn" authorityYear="1889" box="[309,434,1853,1874]" class="Insecta" family="Scarabaeidae" genus="Adoryphorus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="337" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA70A02FEE8C378FE240C36" box="[309,434,1853,1874]" italics="true" pageId="15" pageNumber="337">Adoryphorus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B947B00FFFA70A02FE63C378FD610C36" box="[446,759,1853,1874]" italics="true" pageId="15" pageNumber="337">
|
||
<taxonomicName id="4C33179EFFA70A02FE63C378FDF40C36" ID-CoL="WS6" authorityName="Arrow" authorityYear="1908" box="[446,610,1853,1874]" class="Insecta" family="Scarabaeidae" genus="Anomalomorpha" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="337" phylum="Arthropoda" rank="genus">Anomalomorpha</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C33179EFFA70A02FDB1C378FD610C36" ID-CoL="8SSQQ" baseAuthorityName="Hutchinson & Allsopp" baseAuthorityYear="2021" box="[620,759,1853,1874]" class="Insecta" family="Scarabaeidae" genus="Erbmahcedius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="337" phylum="Arthropoda" rank="genus">Erbmahcedius</taxonomicName>
|
||
</emphasis>
|
||
) or with small anterior tubercle and discernible punctures (
|
||
<taxonomicName id="4C33179EFFA70A02FAEDC378FA110C36" ID-CoL="4BZS" authorityName="Dechambre" authorityYear="1999" box="[1328,1415,1853,1874]" class="Insecta" family="Scarabaeidae" genus="Enracius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="337" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFA70A02FAEDC378FA110C36" box="[1328,1415,1853,1874]" italics="true" pageId="15" pageNumber="337">Enracius</emphasis>
|
||
</taxonomicName>
|
||
). .................................................................................................. 34
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFA70A1DFF4AC33DFA0A0A39" lastPageId="16" lastPageNumber="338" pageId="15" pageNumber="337">
|
||
<paragraph id="8B8C6C1DFFA70A1DFF4AC33DFA0A0BC9" blockId="15.[151,1437,151,2021]" lastBlockId="16.[151,1437,151,2021]" lastPageId="16" lastPageNumber="338" pageId="15" pageNumber="337">
|
||
<keyLead id="30C2CF28FFA70A1DFF4AC33DFA0A0BC9" lastPageId="16" lastPageNumber="338" pageId="15" pageNumber="337">
|
||
33 Head with preocular armature of tubercle or small horn, always anterior to a line joining the front of the eyes; frontoclypeal suture straight; mandibles exposed at side of clypeus or not; base of mentum not hollowed or cleft; pronotum with anteromedian tubercle in both sexes; elytral punctation heavily impressed; size and development of male armature not exhibiting marked variation; reddish brown to pitch black,
|
||
<quantity id="4CCBC1F8FFA70A02FDBFC395FD5F0C81" box="[610,713,2000,2021]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" metricValueMax="3.2" metricValueMin="2.0" pageId="15" pageNumber="337" unit="mm" value="26.0" valueMax="32.0" valueMin="20.0">20–32 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFA70A02FC86C38AFC6D0C81" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[859,1019,1999,2021]" pageId="15" pageNumber="337" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFA70A02FC4CC395FC540C80" box="[913,962,1999,2021]" italics="true" pageId="15" pageNumber="337">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 55M);
|
||
<collectingRegion id="49F7A2FFFFA70A02FBA7C395FABE0C81" box="[1146,1320,1999,2021]" country="Australia" name="New South Wales" pageId="15" pageNumber="337">New South Wales</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFA70A02FAECC38AFA1B0C81" box="[1329,1421,1999,2021]" country="Australia" name="Tasmania" pageId="15" pageNumber="337">Tasmania</collectingRegion>
|
||
..............................................................................
|
||
<taxonomicName id="4C33179EFFB80A1DFB5FC4D2FA0A0BC9" ID-CoL="5Z9T" authority="Carne, 1957" authorityName="Carne" authorityYear="1957" box="[1154,1436,151,173]" class="Insecta" family="Scarabaeidae" genus="Neodasygnathus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="338" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFB80A1DFB5FC4D2FAB70BC8" box="[1154,1313,151,172]" italics="true" pageId="16" pageNumber="338">Neodasygnathus</emphasis>
|
||
<bibRefCitation id="EFA211ECFFB80A1DFAFAC4DDFA0A0BC9" author="Carne, P. B." box="[1319,1436,151,173]" pageId="16" pageNumber="338" refId="ref19138" refString="Carne, P. B. (1957) Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne, 1957</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFB80A1DFF4AC4F0FA0A0A39" blockId="16.[151,1437,151,2021]" pageId="16" pageNumber="338">
|
||
<keyLead id="30C2CF28FFB80A1DFF4AC4F0FA0A0A39" pageId="16" pageNumber="338">
|
||
- Head with interocular armature; frontoclypeal suture recurved, giving rise to tubercle or small horn usually posterior to a line joining the front of the eyes; mandibles almost always concealed; base of mentum hollowed or deeply cleft (
|
||
<bibRefCitation id="EFA211ECFFB80A1DFB25C497FA0C0B83" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[1272,1434,210,232]" pageId="16" pageNumber="338" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFB80A1DFAF2C496FAF60B83" box="[1327,1376,210,232]" italics="true" pageId="16" pageNumber="338">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.25); pronotum never with anteromedian tubercle, variously excavated, often bounded posteriorly by sharp points; elytral punctation lightly impressed; extremely variable in size and development of male armature; reddish brown to reddish black,
|
||
<quantity id="4CCBC1F8FFB80A1DFF3FC56FFEDF0A24" box="[226,329,298,320]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="3.2" metricValueMin="1.7" pageId="16" pageNumber="338" unit="mm" value="24.5" valueMax="32.0" valueMin="17.0">17–32 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFB80A1DFE06C56FFDED0A5B" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[475,635,298,320]" pageId="16" pageNumber="338" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFB80A1DFDCCC56EFDD70A5B" box="[529,577,298,320]" italics="true" pageId="16" pageNumber="338">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 55B);
|
||
<collectingRegion id="49F7A2FFFFB80A1DFD28C56EFC340A24" box="[757,930,298,320]" country="Australia" name="New South Wales" pageId="16" pageNumber="338">New South Wales</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFB80A1DFC76C56FFB890A24" box="[939,1055,298,320]" country="Australia" name="Queensland" pageId="16" pageNumber="338">Queensland</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFB80A1DFBF4C56FFB290A24" box="[1065,1215,298,320]" country="Australia" name="South Australia" pageId="16" pageNumber="338">South Australia</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFB80A1DFB14C56FFA800A24" box="[1225,1302,298,320]" country="Australia" name="Victoria" pageId="16" pageNumber="338">Victoria</collectingRegion>
|
||
........................................................................................
|
||
<taxonomicName id="4C33179EFFB80A1DFB59C502FA0A0A39" ID-CoL="3ZZY" authority="MacLeay, 1819" authorityName="MacLeay" authorityYear="1819" box="[1156,1436,327,349]" class="Insecta" family="Scarabaeidae" genus="Dasygnathus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="338" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFB80A1DFB59C502FA970A38" box="[1156,1281,327,348]" italics="true" pageId="16" pageNumber="338">Dasygnathus</emphasis>
|
||
MacLeay, 1819
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFB80A1DFF4AC520FA0A094F" pageId="16" pageNumber="338">
|
||
<paragraph id="8B8C6C1DFFB80A1DFF4AC520FA0A0AB6" blockId="16.[151,1437,151,2021]" pageId="16" pageNumber="338">
|
||
<keyLead id="30C2CF28FFB80A1DFF4AC520FA0A0AB6" pageId="16" pageNumber="338">
|
||
34 Pronotum and elytra guttered with epipleura horizontal on anterior half; head with recurved frontoclypeal suture often terminating in a tubercle; males with all tarsi elongate, protarsomere 5 much longer than 1; claws enlarged, robust, slightly curved; dark brown or black with yellow legs,
|
||
<quantity id="4CCBC1F8FFB80A1DFD7DC5DAFCBC0AD1" box="[672,810,415,437]" metricMagnitude="-2" metricUnit="m" metricValue="1.6149999999999998" metricValueMax="2.08" metricValueMin="1.15" pageId="16" pageNumber="338" unit="mm" value="16.15" valueMax="20.8" valueMin="11.5">11.5–20.8 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFB80A1DFC66C5DAFBCC0AD1" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[955,1114,415,437]" pageId="16" pageNumber="338" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFB80A1DFC2CC5E5FBB70AD0" box="[1009,1057,415,437]" italics="true" pageId="16" pageNumber="338">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 54J;
|
||
<bibRefCitation id="EFA211ECFFB80A1DFB18C5DAFE800AB6" author="Hutchinson, P. M. & Allsopp, P. G." pageId="16" pageNumber="338" pagination="439 - 462" refId="ref19528" refString="Hutchinson, P. M. & Allsopp, P. G. (2021) Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa, 5072 (5), 439 - 462. https: // doi. org / 10.11646 / zootaxa. 5072.5.2" type="journal article" year="2021">Hutchinson & Allsopp 2021</bibRefCitation>
|
||
: figs. 15, 19, 23, 28, 35);
|
||
<collectingRegion id="49F7A2FFFFB80A1DFDD6C5F8FD2F0AB7" box="[523,697,445,467]" country="Australia" name="New South Wales" pageId="16" pageNumber="338">New South Wales</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFB80A1DFD1FC5F8FCA00AB7" box="[706,822,445,467]" country="Australia" name="Queensland" pageId="16" pageNumber="338">Queensland</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFB80A1DFCE2C5F8FC190AB7" box="[831,911,445,467]" country="Australia" name="Victoria" pageId="16" pageNumber="338">Victoria</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFB80A1DFC45C5F8FBD40AB7" box="[920,1090,445,467]" country="Australia" name="Western Australia" pageId="16" pageNumber="338">Western Australia</collectingRegion>
|
||
....
|
||
<taxonomicName id="4C33179EFFB80A1DFBA1C5F8FA0A0AB6" ID-CoL="WS6" authority="Arrow, 1908" authorityName="Arrow" authorityYear="1908" box="[1148,1436,445,466]" class="Insecta" family="Scarabaeidae" genus="Anomalomorpha" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="338" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFB80A1DFBA1C5F8FA8B0AB6" box="[1148,1309,445,466]" italics="true" pageId="16" pageNumber="338">Anomalomorpha</emphasis>
|
||
Arrow, 1908
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFB80A1DFF4AC59EFA0A094F" blockId="16.[151,1437,151,2021]" pageId="16" pageNumber="338">
|
||
<keyLead id="30C2CF28FFB80A1DFF4AC59EFA0A094F" pageId="16" pageNumber="338">
|
||
- Pronotum and elytra not guttered, epipleura vertical; head either unarmed or with small swelling on vertex; frontoclypeal suture straight; males with tarsi elongate or not, protarsomere 5 much longer than 1 or not; claws enlarged or not; southern
|
||
<collectingCountry id="F3242C8DFFB80A1DFAE3C5B2FA000969" box="[1342,1430,503,525]" name="Australia" pageId="16" pageNumber="338">Australia</collectingCountry>
|
||
.................................................................................................. 35
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFB80A1DFF4AC677FA0A099C" pageId="16" pageNumber="338">
|
||
<paragraph id="8B8C6C1DFFB80A1DFF4AC677FA0A09FB" blockId="16.[151,1437,151,2021]" pageId="16" pageNumber="338">
|
||
<keyLead id="30C2CF28FFB80A1DFF4AC677FA0A09FB" pageId="16" pageNumber="338">
|
||
35 Clypeus setose; pronotum with discernible punctures; male pronotum with small tubercle on anterior edge; head unarmed; frontoclypeal suture carinate; tarsi elongate in both male and female; male protarsomere 5 much longer than 1; claws enlarged and only slightly curved; brownish black, 13.5–21.0 mm long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFB80A1DFC55C628FBB209E6" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[904,1060,621,643]" pageId="16" pageNumber="338" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFB80A1DFC60C62BFC7A09E6" box="[957,1004,621,643]" italics="true" pageId="16" pageNumber="338">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 54I;
|
||
<bibRefCitation id="EFA211ECFFB80A1DFB56C628FA0C09E6" author="Hutchinson, P. M. & Allsopp, P. G." box="[1163,1434,621,643]" pageId="16" pageNumber="338" pagination="439 - 462" refId="ref19528" refString="Hutchinson, P. M. & Allsopp, P. G. (2021) Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa, 5072 (5), 439 - 462. https: // doi. org / 10.11646 / zootaxa. 5072.5.2" type="journal article" year="2021">Hutchinson & Allsopp 2021</bibRefCitation>
|
||
: figs. 1, 5, 8);
|
||
<collectingRegion id="49F7A2FFFFB80A1DFEBFC6CEFD9B09C4" box="[354,525,650,672]" country="Australia" name="Western Australia" pageId="16" pageNumber="338">Western Australia</collectingRegion>
|
||
.....................................................
|
||
<taxonomicName id="4C33179EFFB80A1DFB48C6CEFA0A09FB" ID-CoL="4BZS" authority="Dechambre, 1999" authorityName="Dechambre" authorityYear="1999" box="[1173,1436,650,672]" class="Insecta" family="Scarabaeidae" genus="Enracius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="338" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFB80A1DFB48C6CEFB7A09C4" box="[1173,1260,651,672]" italics="true" pageId="16" pageNumber="338">Enracius</emphasis>
|
||
Dechambre, 1999
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFB80A1DFF4AC6EDFA0A099C" blockId="16.[151,1437,151,2021]" pageId="16" pageNumber="338">
|
||
<keyLead id="30C2CF28FFB80A1DFF4AC6EDFA0A099C" pageId="16" pageNumber="338">
|
||
- Clypeus glabrous; pronotum evenly convex with surface micropunctate; frontoclypeal suture carinate or not; male tarsi elongate or not; protarsomere 5 much longer than 1 or not; claws enlarged or not; brown to black,
|
||
<quantity id="4CCBC1F8FFB80A1DFBE4C680FB0209BE" box="[1081,1172,709,730]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.7" metricValueMin="0.9" pageId="16" pageNumber="338" unit="mm" value="13.0" valueMax="17.0" valueMin="9.0">9–17 mm</quantity>
|
||
long; eastern
|
||
<collectingCountry id="F3242C8DFFB80A1DFAC7C680FAE409BF" box="[1306,1394,709,731]" name="Australia" pageId="16" pageNumber="338">Australia</collectingCountry>
|
||
... .................................................................................................. 36
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFB80A1DFF4AC6BAFA0A0F79" pageId="16" pageNumber="338">
|
||
<paragraph id="8B8C6C1DFFB80A1DFF4AC6BAFA0A08EE" blockId="16.[151,1437,151,2021]" pageId="16" pageNumber="338">
|
||
<keyLead id="30C2CF28FFB80A1DFF4AC6BAFA0A08EE" pageId="16" pageNumber="338">
|
||
36 Anterior edge of pronotum entirely, narrowly membranous; male with tarsi elongate, protarsomere 5 much longer than 1; claws enlarged, robust and only slightly curved; elytral striae vaguely impressed; female penultimate abdominal ventrite with row of setae removed from posterior edge; parameres with inner edges neither spinose not toothed; brown,
|
||
<quantity id="4CCBC1F8FFB80A1DFB70C77FFA830834" box="[1197,1301,826,848]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="16" pageNumber="338" unit="mm" value="13.0" valueMax="14.0" valueMin="12.0">12–14 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFB80A1DFF35C712FE6A0809" author="Hutchinson, P. M. & Allsopp, P. G." box="[232,508,855,877]" pageId="16" pageNumber="338" pagination="439 - 462" refId="ref19528" refString="Hutchinson, P. M. & Allsopp, P. G. (2021) Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa, 5072 (5), 439 - 462. https: // doi. org / 10.11646 / zootaxa. 5072.5.2" type="journal article" year="2021">Hutchinson & Allsopp 2021</bibRefCitation>
|
||
: figs. 52, 58);
|
||
<collectingRegion id="49F7A2FFFFB80A1DFD58C71DFCA50809" box="[645,819,855,877]" country="Australia" name="New South Wales" pageId="16" pageNumber="338">New South Wales</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFB80A1DFCE1C712FC0E0809" box="[828,920,855,877]" country="Australia" name="Tasmania" pageId="16" pageNumber="338">Tasmania</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFB80A1DFC74C712FC600809" box="[937,1014,855,877]" country="Australia" name="Victoria" pageId="16" pageNumber="338">Victoria</collectingRegion>
|
||
....................................................................................................
|
||
<taxonomicName id="4C33179EFFB80A1DFC25C730FA0A08EE" ID-CoL="8SSQQ" authority="Hutchinson & Allsopp, 2021" authorityName="Hutchinson & Allsopp" authorityYear="2021" box="[1016,1436,885,907]" class="Insecta" family="Scarabaeidae" genus="Erbmahcedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="338" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFB80A1DFC25C730FB1508EE" box="[1016,1155,885,906]" italics="true" pageId="16" pageNumber="338">Erbmahcedius</emphasis>
|
||
<bibRefCitation id="EFA211ECFFB80A1DFB55C730FA0A08EE" author="Hutchinson, P. M. & Allsopp, P. G." box="[1160,1436,885,907]" pageId="16" pageNumber="338" pagination="439 - 462" refId="ref19528" refString="Hutchinson, P. M. & Allsopp, P. G. (2021) Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa, 5072 (5), 439 - 462. https: // doi. org / 10.11646 / zootaxa. 5072.5.2" type="journal article" year="2021">Hutchinson & Allsopp, 2021</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFB80A1DFF4AC7D6FA0A0F79" blockId="16.[151,1437,151,2021]" pageId="16" pageNumber="338">
|
||
<keyLead id="30C2CF28FFB80A1DFF4AC7D6FA0A0F79" pageId="16" pageNumber="338">
|
||
- Anterior edge of pronotum with narrow membrane not extending to anterior angles; male tarsi not elongate, protarsomere 5 about as long as 1; claws not enlarged and more strongly curved; elytral striae more deeply impressed and distinctly punctate; female penultimate abdominal ventrite with row of setae close to posterior edge; parameres with inner edges spinose or toothed; brown to black,
|
||
<quantity id="4CCBC1F8FFB80A1DFE5FC7AFFE480F64" box="[386,478,1002,1024]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.7" metricValueMin="0.9" pageId="16" pageNumber="338" unit="mm" value="13.0" valueMax="17.0" valueMin="9.0">9–17 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFB80A1DFDAEC7AFFC81089B" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[627,791,1002,1024]" pageId="16" pageNumber="338" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFB80A1DFD76C7AEFD4B089B" box="[683,733,1002,1024]" italics="true" pageId="16" pageNumber="338">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 54B;
|
||
<bibRefCitation id="EFA211ECFFB80A1DFC50C7AFFB35089B" author="Hutchinson, P. M. & Allsopp, P. G." box="[909,1187,1002,1024]" pageId="16" pageNumber="338" pagination="439 - 462" refId="ref19528" refString="Hutchinson, P. M. & Allsopp, P. G. (2021) Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina). Zootaxa, 5072 (5), 439 - 462. https: // doi. org / 10.11646 / zootaxa. 5072.5.2" type="journal article" year="2021">Hutchinson & Allsopp 2021</bibRefCitation>
|
||
: figs. 64–65);
|
||
<collectingRegion id="49F7A2FFFFB80A1DFAEDC7AEFE890F79" country="Australia" name="New South Wales" pageId="16" pageNumber="338">New South Wales</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFB80A1DFEF5C042FE0A0F79" box="[296,412,1031,1053]" country="Australia" name="Queensland" pageId="16" pageNumber="338">Queensland</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFB80A1DFE78C042FDAA0F79" box="[421,572,1031,1053]" country="Australia" name="South Australia" pageId="16" pageNumber="338">South Australia</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFB80A1DFD98C042FD350F79" box="[581,675,1031,1053]" country="Australia" name="Tasmania" pageId="16" pageNumber="338">Tasmania</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFB80A1DFD70C042FD6C0F79" box="[685,762,1031,1053]" country="Australia" name="Victoria" pageId="16" pageNumber="338">Victoria</collectingRegion>
|
||
................................
|
||
<taxonomicName id="4C33179EFFB80A1DFBA4C042FA0A0F79" ID-CoL="PDB" authority="Blackburn, 1889" authorityName="Blackburn" authorityYear="1889" box="[1145,1436,1031,1053]" class="Insecta" family="Scarabaeidae" genus="Adoryphorus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="338" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFB80A1DFBA4C042FB600F78" box="[1145,1270,1031,1052]" italics="true" pageId="16" pageNumber="338">Adoryphorus</emphasis>
|
||
Blackburn, 1889
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFB80A1DFF4AC060FA0A0FF6" pageId="16" pageNumber="338">
|
||
<paragraph id="8B8C6C1DFFB80A1DFF4AC060FA0A0F33" blockId="16.[151,1437,151,2021]" pageId="16" pageNumber="338">
|
||
<keyLead id="30C2CF28FFB80A1DFF4AC060FA0A0F33" pageId="16" pageNumber="338">37 Protibiae with 3 main teeth and smaller basal tooth or with 3 main teeth and denticles between second and third tooth and behind third tooth; male protarsomere 5 enlarged with asymmetrical claws; black................................. 38</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFB80A1DFF4AC025FA0A0FF6" blockId="16.[151,1437,151,2021]" pageId="16" pageNumber="338">
|
||
<keyLead id="30C2CF28FFB80A1DFF4AC025FA0A0FF6" pageId="16" pageNumber="338">- Protibiae clearly with only 3 teeth externally; male protarsomere 5 enlarged or not and claws symmetrical or not; light brown, reddish brown, reddish black or black.................................................................... 40</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFB80A1DFF4AC0DFFA0A0E26" pageId="16" pageNumber="338">
|
||
<paragraph id="8B8C6C1DFFB80A1DFF4AC0DFFA0A0E63" blockId="16.[151,1437,151,2021]" pageId="16" pageNumber="338">
|
||
<keyLead id="30C2CF28FFB80A1DFF4AC0DFFA0A0E63" pageId="16" pageNumber="338">
|
||
38 Propygidium with 2 well defined fine stridulatory bands; head unarmed; pygidium impunctate and highly convex, more so in female; pronotum evenly convex; black,
|
||
<quantity id="4CCBC1F8FFB80A1DFDB5C0F2FD550FA9" box="[616,707,1207,1229]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.5" metricValueMin="0.9" pageId="16" pageNumber="338" unit="mm" value="12.0" valueMax="15.0" valueMin="9.0">9–15 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFB80A1DFC8EC0F2FC670FA9" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[851,1009,1207,1229]" pageId="16" pageNumber="338" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFB80A1DFC55C0FDFC2F0FA8" box="[904,953,1207,1229]" italics="true" pageId="16" pageNumber="338">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 53K;
|
||
<bibRefCitation id="EFA211ECFFB80A1DFBBFC0F2FAE50FA9" author="Hutchinson, P. M. & Allsopp, P. G." box="[1122,1395,1207,1229]" pageId="16" pageNumber="338" pagination="1 - 14" refId="ref19412" refString="Hutchinson, P. M. & Allsopp, P. G. (2019) Revisiting Nephrodopus Sharp, 1873, with the description of a new species and a female (Coleoptera: Scarabaeidae: Dynastinae). Australian Entomologist, 46, 1 - 14." type="journal article" year="2019">Allsopp & Hutchinson 2019</bibRefCitation>
|
||
: fig. 2); New South Wales, Queensland, South Australia, Victoria, Western Australia,
|
||
<collectingCountry id="F3242C8DFFB80A1DFC3BC090FBE30F8F" box="[998,1141,1237,1259]" name="Norfolk Island" pageId="16" pageNumber="338">Norfolk Island</collectingCountry>
|
||
, Lord Howe Island....................................................................................
|
||
<taxonomicName id="4C33179EFFB80A1DFBB1C0B7FA0A0E63" ID-CoL="62V4V" authority="Burmeister, 1847" authorityName="Burmeister" authorityYear="1847" box="[1132,1436,1266,1288]" class="Insecta" family="Scarabaeidae" genus="Heteronychus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="338" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFB80A1DFBB1C0B7FB660E63" box="[1132,1264,1266,1287]" italics="true" pageId="16" pageNumber="338">Heteronychus</emphasis>
|
||
Burmeister, 1847
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFB80A1DFF4AC155FA0A0E26" blockId="16.[151,1437,151,2021]" pageId="16" pageNumber="338">
|
||
<keyLead id="30C2CF28FFB80A1DFF4AC155FA0A0E26" pageId="16" pageNumber="338">- Propygidium with dispersed stridulatory ridges, not forming well-defined bands; head armed with either 1 or 2 tubercles or horns; pygidium variously punctuate and less convex; pronotum either evenly convex or with a depression anteriorly..... 39</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFB80A1DFF4AC10FFA0A0D6B" pageId="16" pageNumber="338">
|
||
<paragraph id="8B8C6C1DFFB80A1DFF4AC10FFA0A0EFE" blockId="16.[151,1437,151,2021]" pageId="16" pageNumber="338">
|
||
<keyLead id="30C2CF28FFB80A1DFF4AC10FFA0A0EFE" pageId="16" pageNumber="338">
|
||
39 Head armed with 2 tubercles or horns; pronotum evenly convex; clypeus narrowed and truncate anteriorly with 2 small, upturned teeth; black,
|
||
<quantity id="4CCBC1F8FFB80A1DFE80C12DFE550E19" box="[349,451,1384,1405]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="2.2" metricValueMin="1.2" pageId="16" pageNumber="338" unit="mm" value="17.0" valueMax="22.0" valueMin="12.0">12–22 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFB80A1DFD89C122FD650E19" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[596,755,1383,1405]" pageId="16" pageNumber="338" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFB80A1DFD57C12DFD2C0E18" box="[650,698,1383,1405]" italics="true" pageId="16" pageNumber="338">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plates 54K, 59I);
|
||
<collectingRegion id="49F7A2FFFFB80A1DFC7EC12DFBC60E19" box="[931,1104,1383,1405]" country="Australia" name="New South Wales" pageId="16" pageNumber="338">New South Wales</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFB80A1DFB84C122FB5B0E19" box="[1113,1229,1383,1405]" country="Australia" name="Queensland" pageId="16" pageNumber="338">Queensland</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFB80A1DFB0BC122FAB00E19" box="[1238,1318,1383,1405]" country="Australia" name="Victoria" pageId="16" pageNumber="338">Victoria</collectingRegion>
|
||
, Lord Howe Island....................................................................
|
||
<taxonomicName id="4C33179EFFB80A1DFB80C1C3FADE0EFE" ID-CoL="62ZVP" authority="Arrow, 1911" authorityName="Arrow" authorityYear="1911" box="[1117,1352,1413,1435]" class="Insecta" family="Scarabaeidae" genus="Metanastes" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="338" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFB80A1DFB80C1C3FB5C0EFF" box="[1117,1226,1414,1435]" italics="true" pageId="16" pageNumber="338">Metanastes</emphasis>
|
||
Arrow, 1911
|
||
</taxonomicName>
|
||
(in part)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFB80A1DFF4AC1E6FA0A0D6B" blockId="16.[151,1437,151,2021]" pageId="16" pageNumber="338">
|
||
<keyLead id="30C2CF28FFB80A1DFF4AC1E6FA0A0D6B" pageId="16" pageNumber="338">
|
||
- Head armed with a horn (male) or median tubercle (female); pronotum with anterior rugose depression, bounded posteriorly by a tubercle in both sexes; clypeus more broadly truncate anteriorly with both anterior angles bearing a sharp tooth; larger black,
|
||
<quantity id="4CCBC1F8FFB80A1DFF3FC198FEDF0E96" box="[226,329,1501,1522]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="2.8" metricValueMin="2.2" pageId="16" pageNumber="338" unit="mm" value="25.0" valueMax="28.0" valueMin="22.0">22–28 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFB80A1DFE06C198FDED0E96" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[475,635,1501,1523]" pageId="16" pageNumber="338" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFB80A1DFDCCC19BFDD70E96" box="[529,577,1501,1523]" italics="true" pageId="16" pageNumber="338">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 53I;
|
||
<bibRefCitation id="EFA211ECFFB80A1DFD3EC198FC610E96" author="Hutchinson, P. M. & Allsopp, P. G." box="[739,1015,1501,1523]" pageId="16" pageNumber="338" pagination="1 - 14" refId="ref19412" refString="Hutchinson, P. M. & Allsopp, P. G. (2019) Revisiting Nephrodopus Sharp, 1873, with the description of a new species and a female (Coleoptera: Scarabaeidae: Dynastinae). Australian Entomologist, 46, 1 - 14." type="journal article" year="2019">Allsopp & Hutchinson 2019</bibRefCitation>
|
||
: fig. 9);
|
||
<collectingRegion id="49F7A2FFFFB80A1DFB9AC198FB2E0E97" box="[1095,1208,1501,1523]" country="Australia" name="Queensland" pageId="16" pageNumber="338">Queensland</collectingRegion>
|
||
....................................................................................................
|
||
<taxonomicName id="4C33179EFFB80A1DFB14C1BEFA0A0D6B" ID-CoL="6DJH" authority="Arrow, 1911" authorityName="Arrow" authorityYear="1911" box="[1225,1436,1530,1552]" class="Insecta" family="Scarabaeidae" genus="Papuana" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="338" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFB80A1DFB14C1BEFAB60D74" box="[1225,1312,1531,1552]" italics="true" pageId="16" pageNumber="338">Papuana</emphasis>
|
||
Arrow, 1911
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<paragraph id="8B8C6C1DFFB80A1DFF4AC25DFA0A0DE1" blockId="16.[151,1437,151,2021]" pageId="16" pageNumber="338">
|
||
<keyLead id="30C2CF28FFB80A1DFF4AC25DFA0A0DE1" pageId="16" pageNumber="338">
|
||
<keyStep id="30C774B8FFB80A1DFF4AC25DFA0A0DE1" pageId="16" pageNumber="338">
|
||
<specimenCount id="9D35A794FFB80A1DFF4AC25DFEA40D49" box="[151,306,1559,1581]" pageId="16" pageNumber="338" type="female">40 Females</specimenCount>
|
||
with anterior and posterior margins of the pronotum conspicuously concave (
|
||
<figureCitation id="13087098FFB80A1DFBCAC252FBF00D49" box="[1047,1126,1559,1581]" captionStart="FIGURES 11–14" captionStartId="9.[151,264,1836,1860]" captionTargetBox="[170,1392,260,1812]" captionTargetId="figure-19@9.[151,1435,181,1812]" captionTargetPageId="9" captionText="FIGURES 11–14. Constricticollis clunis Hutchinson & Allsopp new genus, new species, holotype female: 11, dorsal view; 12, ventral view; 13, lateral view; 14, labels." figureDoi="http://doi.org/10.5281/zenodo.8391704" httpUri="https://zenodo.org/record/8391704/files/figure.png" pageId="16" pageNumber="338">Figs. 11</figureCitation>
|
||
,
|
||
<figureCitation id="13087098FFB80A1DFBB2C25DFB1C0D49" box="[1135,1162,1560,1581]" captionStart="FIGURES 15–20" captionStartId="10.[151,264,1560,1584]" captionTargetBox="[151,1433,181,1536]" captionTargetId="figure-17@10.[151,1435,181,1536]" captionTargetPageId="10" captionText="FIGURES 15–20. Constricticollis clunis Hutchinson &Allsopp new genus, new species, holotype female: 15, frontal view; 16, rear view; 17, head, dorsal view; 18, head, ventral view; 19, head, dorso-lateral view; 20, head and pronotum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8391708" httpUri="https://zenodo.org/record/8391708/files/figure.png" pageId="16" pageNumber="338">20</figureCitation>
|
||
); males unknown; head dark brown, pronotum brown, elytra brown with apical half dark brown,
|
||
<quantity id="4CCBC1F8FFB80A1DFCB0C270FC260D2E" box="[877,944,1589,1611]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="338" unit="mm" value="15.0">15 mm</quantity>
|
||
long; northern
|
||
<collectingRegion id="49F7A2FFFFB80A1DFB98C270FB790D2F" box="[1093,1263,1589,1611]" country="Australia" name="Western Australia" pageId="16" pageNumber="338">Western Australia</collectingRegion>
|
||
.........................................................................
|
||
<taxonomicName id="4C33179EFFB80A1DFC69C217FABE0D0C" authority="Hutchinson & Allsopp" authorityName="Hutchinson & Allsopp" authorityYear="2023" box="[948,1320,1618,1640]" class="Insecta" family="Scarabaeidae" genus="Constricticollis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="338" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B947B00FFFB80A1DFC69C217FBDE0D03" box="[948,1096,1618,1639]" italics="true" pageId="16" pageNumber="338">Constricticollis</emphasis>
|
||
Hutchinson & Allsopp
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B947B00FFFB80A1DFAEFC217FA0B0D03" bold="true" box="[1330,1437,1618,1639]" pageId="16" pageNumber="338">
|
||
<taxonomicNameLabel id="A2740D74FFB80A1DFAEFC217FA0B0D03" box="[1330,1437,1618,1639]" pageId="16" pageNumber="338" rank="genus">new genus</taxonomicNameLabel>
|
||
</emphasis>
|
||
Females with anterior and posterior margins convex or almost straight.......................................... 41
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<keyStep id="30C774B8FFB80A1DFF4AC2C8FA0A0C0B" pageId="16" pageNumber="338">
|
||
<paragraph id="8B8C6C1DFFB80A1DFF4AC2C8FA0A0C7C" blockId="16.[151,1437,151,2021]" pageId="16" pageNumber="338">
|
||
<keyLead id="30C2CF28FFB80A1DFF4AC2C8FA0A0C7C" pageId="16" pageNumber="338">
|
||
41 Centre of apical truncate faces of meso- and metatibiae densely ciliate (
|
||
<bibRefCitation id="EFA211ECFFB80A1DFC6DC2C8FBBE0DC7" author="Carne, P. B." box="[944,1064,1677,1699]" pageId="16" pageNumber="338" refId="ref19138" refString="Carne, P. B. (1957) Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne 1957</bibRefCitation>
|
||
: fig. 9); frontoclypeal suture strongly marked, posteriorly angulate, and giving rise to a median tubercle; clypeus narrowed to a slightly upturned apex, sides concave; mandibles conspicuously broadly bidentate; pronotum unarmed, evenly convex; male protarsal claws asymmetrical; reddish brown to reddish yellow with little or no elytral punctation,
|
||
<quantity id="4CCBC1F8FFB80A1DFCF6C2A0FC050D9E" box="[811,915,1765,1787]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="16" pageNumber="338" unit="mm" value="17.5" valueMax="20.0" valueMin="15.0">15–20 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFB80A1DFBF5C2A0FB5D0D9E" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[1064,1227,1765,1787]" pageId="16" pageNumber="338" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFB80A1DFB82C2A3FB070D9E" box="[1119,1169,1765,1787]" italics="true" pageId="16" pageNumber="338">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 55F);
|
||
<collectingRegion id="49F7A2FFFFB80A1DFA9BC2A0FEAE0C7C" country="Australia" name="Northern Territory" pageId="16" pageNumber="338">Northern Territory</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFB80A1DFE9CC347FE230C7C" box="[321,437,1794,1816]" country="Australia" name="Queensland" pageId="16" pageNumber="338">Queensland</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFB80A1DFE63C346FDFF0C7C" box="[446,617,1794,1816]" country="Australia" name="Western Australia" pageId="16" pageNumber="338">Western Australia</collectingRegion>
|
||
............................................
|
||
<taxonomicName id="4C33179EFFB80A1DFBA0C347FA0A0C7C" ID-CoL="52KF" authority="Blackburn, 1895" authorityName="Blackburn" authorityYear="1895" box="[1149,1436,1794,1816]" class="Insecta" family="Scarabaeidae" genus="Hyphoryctes" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="338" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFB80A1DFBA0C347FB600C73" box="[1149,1270,1794,1815]" italics="true" pageId="16" pageNumber="338">Hyphoryctes</emphasis>
|
||
Blackburn, 1895
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFB80A1DFF4AC365FA0A0C0B" blockId="16.[151,1437,151,2021]" pageId="16" pageNumber="338">
|
||
<keyLead id="30C2CF28FFB80A1DFF4AC365FA0A0C0B" pageId="16" pageNumber="338">
|
||
-
|
||
<collectingRegion id="49F7A2FFFFB80A1DFF3FC365FEB40C51" box="[226,290,1824,1845]" country="France" name="Centre" pageId="16" pageNumber="338">Centre</collectingRegion>
|
||
of apical truncate faces of meso- and metatibiae glabrous; frontoclypeal suture straight or obscurely recurved with 1 or 2 low tubercles or a horn; clypeus various; mandibles bidentate or tridentate; pronotum either unarmed or with a depression and/or tubercle anteriorly; male protarsal claws asymmetrical or not; reddish brown, reddish black or black............. 42
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFB80A1CFF4AC33DFA0A0A39" lastPageId="17" lastPageNumber="339" pageId="16" pageNumber="338">
|
||
<paragraph id="8B8C6C1DFFB80A1CFF4AC33DFA0A0B83" blockId="16.[151,1437,151,2021]" lastBlockId="17.[151,1437,151,907]" lastPageId="17" lastPageNumber="339" pageId="16" pageNumber="338">
|
||
<keyLead id="30C2CF28FFB80A1CFF4AC33DFA0A0B83" lastPageId="17" lastPageNumber="339" pageId="16" pageNumber="338">
|
||
42 Mandibles bidentate with inner apical tooth much narrower than outer one (
|
||
<bibRefCitation id="EFA211ECFFB80A1DFC08C332FBEB0CE9" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[981,1149,1911,1933]" pageId="16" pageNumber="338" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFB80A1DFBD3C33DFBD70CE8" box="[1038,1089,1911,1933]" italics="true" pageId="16" pageNumber="338">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.27); pronotum either completely unarmed or with a major excavation bounded posteriorly by a ridge (one species); frontoclypeal suture obscure, somewhat recurved, giving rise to an obtuse median swelling, tubercles or horn; clypeus more-or-less rounded, apex obliquely truncate with lower portion of truncate face slightly produced beyond upper sinuate edge; propygidium with dispersed stridulating ridges; reddish brown, highly convex, rather featureless,
|
||
<quantity id="4CCBC1F8FFB90A1CFC5BC4D2FC790BC9" box="[902,1007,151,173]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.6" metricValueMin="1.2" pageId="17" pageNumber="339" unit="mm" value="19.0" valueMax="26.0" valueMin="12.0">12–26 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFB90A1CFB5BC4D2FABD0BC9" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[1158,1323,151,173]" pageId="17" pageNumber="339" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFB90A1CFB63C4DDFB660BC8" box="[1214,1264,151,173]" italics="true" pageId="17" pageNumber="339">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 59J); New South Wales, Queensland, South Australia, Tasmania, Victoria, Western Australia,
|
||
<collectingCountry id="F3242C8DFFB90A1CFBF0C4F0FB2A0BAF" box="[1069,1212,181,203]" name="Norfolk Island" pageId="17" pageNumber="339">Norfolk Island</collectingCountry>
|
||
, Lord Howe Island..................................................................................
|
||
<taxonomicName id="4C33179EFFB90A1CFB7DC497FA0A0B83" ID-CoL="6QKN" authority="Erichson, 1842" authorityName="Erichson" authorityYear="1842" box="[1184,1436,210,232]" class="Insecta" family="Scarabaeidae" genus="Pimelopus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="339" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFB90A1CFB7DC497FA930B83" box="[1184,1285,210,231]" italics="true" pageId="17" pageNumber="339">Pimelopus</emphasis>
|
||
Erichson, 1842
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFB90A1CFF4AC4B5FA0A0A39" blockId="17.[151,1437,151,907]" pageId="17" pageNumber="339">
|
||
<keyLead id="30C2CF28FFB90A1CFF4AC4B5FA0A0A39" pageId="17" pageNumber="339">- Mandibles variously tridentate; pronotum either completely unarmed or with anteromedian tubercle and impression; frontoclypeal suture straight, giving rise to 2 tubercles or none; clypeus narrowed to apex, which is either somewhat rounded or bears 2 small tubercles or teeth; propygidium with either dispersed stridulating ridges or ridges organised into short, coarse paramedian bands.................................................................................... 43</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFB90A1CFF4AC520FA0A094E" pageId="17" pageNumber="339">
|
||
<paragraph id="8B8C6C1DFFB90A1CFF4AC520FA0A0AB7" blockId="17.[151,1437,151,907]" pageId="17" pageNumber="339">
|
||
<keyLead id="30C2CF28FFB90A1CFF4AC520FA0A0AB7" pageId="17" pageNumber="339">
|
||
43 Pygidium evenly and heavily punctate and glabrous; clypeus somewhat rounded and slightly upturned at apex; protibiae with distal tooth somewhat elongate, basal tooth small; mandibles with inner apical tooth markedly produced; propygidium with dispersed stridulating ridges; black,
|
||
<quantity id="4CCBC1F8FFB90A1CFDE0C5E5FD320AD1" box="[573,676,416,437]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="17" pageNumber="339" unit="mm" value="13.0" valueMax="14.0" valueMin="12.0">12–14 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFB90A1CFCEBC5DAFC400AD1" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[822,982,415,437]" pageId="17" pageNumber="339" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFB90A1CFCB1C5E5FC0B0AD0" box="[876,925,415,437]" italics="true" pageId="17" pageNumber="339">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 54C);
|
||
<collectingRegion id="49F7A2FFFFB90A1CFB8DC5DAFA940AD1" box="[1104,1282,415,437]" country="Australia" name="Northern Territory" pageId="17" pageNumber="339">Northern Territory</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFB90A1CFAD6C5DAFAEA0AD1" box="[1291,1404,415,437]" country="Australia" name="Queensland" pageId="17" pageNumber="339">Queensland</collectingRegion>
|
||
....................................................................................
|
||
<taxonomicName id="4C33179EFFB90A1CFB63C5F8FA0A0AB7" ID-CoL="623W" authority="Carne, 1957" authorityName="Carne" authorityYear="1957" box="[1214,1436,445,467]" class="Insecta" family="Scarabaeidae" genus="Neonastes" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="339" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFB90A1CFB63C5F8FAB70AB6" box="[1214,1313,445,466]" italics="true" pageId="17" pageNumber="339">Neonastes</emphasis>
|
||
<bibRefCitation id="EFA211ECFFB90A1CFAFAC5F8FA0A0AB7" author="Carne, P. B." box="[1319,1436,445,467]" pageId="17" pageNumber="339" refId="ref19138" refString="Carne, P. B. (1957) Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne, 1957</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFB90A1CFF4AC59EFA0A094E" blockId="17.[151,1437,151,907]" pageId="17" pageNumber="339">
|
||
<keyLead id="30C2CF28FFB90A1CFF4AC59EFA0A094E" pageId="17" pageNumber="339">- Pygidium either impunctate or punctate and densely setose; clypeus with apex bearing 2 small tubercles or teeth; protibiae with teeth more evenly sized; mandibles with inner apical tooth not so produced; propygidium with either dispersed stridulating ridges or ridges organised into short, coarse paramedian bands................................................ 44</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C774B8FFB90A1CFF4AC677FA0A08EE" pageId="17" pageNumber="339">
|
||
<paragraph id="8B8C6C1DFFB90A1CFF4AC677FA0A0993" blockId="17.[151,1437,151,907]" pageId="17" pageNumber="339">
|
||
<keyLead id="30C2CF28FFB90A1CFF4AC677FA0A0993" pageId="17" pageNumber="339">
|
||
44 Propygidium with more-or-less well-defined, short, coarse stridulating bands (except
|
||
<taxonomicName id="4C33179EFFB90A1CFBF1C677FB3B0923" box="[1068,1197,562,584]" class="Insecta" family="Scarabaeidae" genus="Constricticollis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="339" phylum="Arthropoda" rank="species" species="laevicollis">
|
||
<emphasis id="B947B00FFFB90A1CFBF1C677FBAA0923" box="[1068,1084,562,583]" italics="true" pageId="17" pageNumber="339">C</emphasis>
|
||
.
|
||
<emphasis id="B947B00FFFB90A1CFB97C677FB3B0923" box="[1098,1197,562,583]" italics="true" pageId="17" pageNumber="339">laevicollis</emphasis>
|
||
</taxonomicName>
|
||
); male pronotum armed with anteromedian tubercle and impression (except
|
||
<taxonomicName id="4C33179EFFB90A1CFD12C615FCB40900" box="[719,802,591,613]" class="Insecta" family="Scarabaeidae" genus="Constricticollis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="339" phylum="Arthropoda" rank="species" species="laevis">
|
||
<emphasis id="B947B00FFFB90A1CFD12C615FD490901" box="[719,735,592,613]" italics="true" pageId="17" pageNumber="339">C</emphasis>
|
||
.
|
||
<emphasis id="B947B00FFFB90A1CFD37C60AFCB40900" box="[746,802,591,612]" italics="true" pageId="17" pageNumber="339">laevis</emphasis>
|
||
</taxonomicName>
|
||
), female sometimes with slight tubercle or swelling; frontoclypeal suture carinate or bituberculate; clypeus apically with 2 small upturned tubercles or teeth; metatibial spurs more-or-less rounded apically, spatulate; prosternal process more-or-less rounded apically (
|
||
<bibRefCitation id="EFA211ECFFB90A1CFC5CC6CFFBB209FB" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[897,1060,650,672]" pageId="17" pageNumber="339" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFB90A1CFC65C6CEFC7F09FB" box="[952,1001,650,672]" italics="true" pageId="17" pageNumber="339">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.28); if male propretarsal claws asymmetrical then protarsomere 5 not enlarged; reddish brown to reddish black,
|
||
<quantity id="4CCBC1F8FFB90A1CFC2AC6EDFBF609D9" box="[1015,1120,680,701]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.7" metricValueMin="1.0" pageId="17" pageNumber="339" unit="mm" value="13.5" valueMax="17.0" valueMin="10.0">10–17 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFB90A1CFB28C6E2FA0C09D9" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[1269,1434,679,701]" pageId="17" pageNumber="339" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFB90A1CFAF0C6EDFAC909D8" box="[1325,1375,679,701]" italics="true" pageId="17" pageNumber="339">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plate 54E);
|
||
<collectingRegion id="49F7A2FFFFB90A1CFE8BC680FD9E09BF" box="[342,520,709,731]" country="Australia" name="New South Wales" pageId="17" pageNumber="339">New South Wales</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFB90A1CFDCEC680FD5109BF" box="[531,711,709,731]" country="Australia" name="Northern Territory" pageId="17" pageNumber="339">Northern Territory</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFB90A1CFD0EC680FCD109BF" box="[723,839,709,731]" country="Australia" name="Queensland" pageId="17" pageNumber="339">Queensland</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFB90A1CFC8FC680FC7D09BF" box="[850,1003,709,731]" country="Australia" name="South Australia" pageId="17" pageNumber="339">South Australia</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFB90A1CFC2AC680FBD109BF" box="[1015,1095,709,731]" country="Australia" name="Victoria" pageId="17" pageNumber="339">Victoria</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFB90A1CFB8CC680FA9609BF" box="[1105,1280,709,731]" country="Australia" name="Western Australia" pageId="17" pageNumber="339">Western Australia</collectingRegion>
|
||
, Cocos-Keeling Islands........................................................................
|
||
<taxonomicName id="4C33179EFFB90A1CFB57C6A7FA0A0993" ID-CoL="62KHY" authority="Ozdikmen, 2009" authorityName="Ozdikmen" authorityYear="2009" box="[1162,1436,738,762]" class="Insecta" family="Scarabaeidae" genus="Carneodon" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="339" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFB90A1CFB57C6A7FB600993" box="[1162,1270,738,759]" italics="true" pageId="17" pageNumber="339">Carneodon</emphasis>
|
||
Özdikmen, 2009
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8C6C1DFFB90A1CFF4AC745FA0A08EE" blockId="17.[151,1437,151,907]" pageId="17" pageNumber="339">
|
||
<keyLead id="30C2CF28FFB90A1CFF4AC745FA0A08EE" pageId="17" pageNumber="339">
|
||
- Propygidium with only dispersed stridulating ridges; pronotum with vague tubercle in male, evenly convex in female; frontoclypeal suture bituberculate; clypeus apically with 2 small forwardly projecting teeth; metatibial spurs sharp; prosternal process expanded and flattened apically (
|
||
<bibRefCitation id="EFA211ECFFB90A1CFDA6C77FFCB5082B" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[635,803,826,848]" pageId="17" pageNumber="339" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFB90A1CFD6EC77EFD71082B" box="[691,743,826,848]" italics="true" pageId="17" pageNumber="339">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: fig. 33.29); male propretarsal claws strongly asymmetrical and protarsomere 5 enlarged; black,
|
||
<quantity id="4CCBC1F8FFB90A1CFDCAC71DFDE80809" box="[535,638,856,877]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="17" pageNumber="339" unit="mm" value="18.5" valueMax="20.0" valueMin="17.0">17–20 mm</quantity>
|
||
long; dorsum (
|
||
<bibRefCitation id="EFA211ECFFB90A1CFCCDC712FC260809" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[784,944,855,877]" pageId="17" pageNumber="339" refId="ref20085" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae MacLeay, 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book" year="2019">
|
||
Weir
|
||
<emphasis id="B947B00FFFB90A1CFC9BC71DFCE10808" box="[838,887,855,877]" italics="true" pageId="17" pageNumber="339">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: plates 54K, 59I);
|
||
<collectingRegion id="49F7A2FFFFB90A1CFBBCC71DFA980809" box="[1121,1294,855,877]" country="Australia" name="New South Wales" pageId="17" pageNumber="339">New South Wales</collectingRegion>
|
||
,
|
||
<collectingRegion id="49F7A2FFFFB90A1CFAC5C712FA1F0809" box="[1304,1417,855,877]" country="Australia" name="Queensland" pageId="17" pageNumber="339">Queensland</collectingRegion>
|
||
..........................................................................
|
||
<taxonomicName id="4C33179EFFB90A1CFB83C730FADE08EE" ID-CoL="62ZVP" authority="Arrow, 1911" authorityName="Arrow" authorityYear="1911" box="[1118,1352,885,906]" class="Insecta" family="Scarabaeidae" genus="Metanastes" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="339" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947B00FFFB90A1CFB83C730FB5D08EE" box="[1118,1227,885,906]" italics="true" pageId="17" pageNumber="339">Metanastes</emphasis>
|
||
Arrow, 1911
|
||
</taxonomicName>
|
||
(in part)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |