910 lines
108 KiB
XML
910 lines
108 KiB
XML
<document id="DF0C22D1ECF92B5AF638EFA818149443" ID-DOI="10.5281/zenodo.185821" ID-GBIF-Dataset="70965309-90ae-4c30-8ec3-615e2f1119d4" ID-ISSN="1175-5326" ID-Zenodo-Dep="185821" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460024783984" checkinUser="plazi" docAuthor="Wood, Perry L., Grismer, Jesse L., Grismer, Lee, Ahmad, Norhayati, Onn, Chan Kin & Bauer, Aaron M." docDate="2009" docId="039A879BFF9AFFCFD3F4F9E4CC636F9C" docLanguage="en" docName="zt02012p046.pdf" docOrigin="Zootaxa 2012" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Acanthosaura titiwangsaensis Wood, Grismer, Grismer, Ahmad, Onn & Bauer, 2009, sp. nov." docType="treatment" docVersion="9" lastPageNumber="43" masterDocId="FFA3FFE3FF92FFC0D363FFDDCD696D53" masterDocTitle="Two new montane species of Acanthosaura Gray, 1831 (Squamata: Agamidae) from Peninsular Malaysia" masterLastPageNumber="46" masterPageNumber="28" pageNumber="36" updateTime="1698233838824" updateUser="plazi">
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<mods:titleInfo id="C5F67055DAFCCBC8B6CD60527F1C841C">
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<mods:title id="5544B53822126513E74FDFCD63E333CE">Two new montane species of Acanthosaura Gray, 1831 (Squamata: Agamidae) from Peninsular Malaysia</mods:title>
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<mods:namePart id="11732A501AA5CEE93BBC6EB4796C4F2B">Wood, Perry L.</mods:namePart>
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<mods:roleTerm id="E785379053D66AF17FE617C8C48B1E6E">Author</mods:roleTerm>
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<mods:namePart id="35798E1562DB3EDB53716043B3439A06">Grismer, Jesse L.</mods:namePart>
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<mods:roleTerm id="9E6EF0EF0E8D2CA3ACC02A7134EA60E7">Author</mods:roleTerm>
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<mods:namePart id="B48AAAACDBFAC48CC68CC47F2B9357BF">Grismer, Lee</mods:namePart>
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</mods:name>
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<mods:name id="8D443B5E45C7B19317F8C4BCCD758E21" type="personal">
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<mods:roleTerm id="DA801F5C2D31AD3162FEFB6AE7BC8FED">Author</mods:roleTerm>
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<mods:namePart id="9979CE7DC3E6F1B3D8174F1AC8541868">Ahmad, Norhayati</mods:namePart>
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<mods:namePart id="B34FA5CA844ECB001240F81E931695C7">Onn, Chan Kin</mods:namePart>
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</mods:role>
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<mods:namePart id="96EA453F5ED1D1F78797DDAABDCA3958">Bauer, Aaron M.</mods:namePart>
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<mods:typeOfResource id="5BED8DE846AA02243D01FDF38AA126AC">text</mods:typeOfResource>
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<mods:title id="7591E5867629A7D93942E697D48DCB51">Zootaxa</mods:title>
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<mods:part id="CB29C6417A2803AF777FF1A4233A58FC">
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<mods:date id="67A21AEBF81A695CF32303C166DCA08C">2009</mods:date>
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<mods:detail id="E4291958107BBBC64FE2CEC49D70A8FF" type="volume">
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<mods:number id="4893CFD6B459C982223DA6379C71CAA0">2012</mods:number>
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<mods:classification id="D1E0FB870DD3E44C679000FD1F48EDF2">journal article</mods:classification>
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<mods:identifier id="91C78B6603457831C712B4E594C01627" type="DOI">10.5281/zenodo.185821</mods:identifier>
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<mods:identifier id="13DCF55202A8D9F6F92A0A20DED36BC4" type="ISSN">1175-5326</mods:identifier>
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<treatment id="039A879BFF9AFFCFD3F4F9E4CC636F9C" ID-DOI="http://doi.org/10.5281/zenodo.5684741" ID-GBIF-Taxon="119382002" ID-Zenodo-Dep="5684741" LSID="urn:lsid:plazi:treatment:039A879BFF9AFFCFD3F4F9E4CC636F9C" httpUri="http://treatment.plazi.org/id/039A879BFF9AFFCFD3F4F9E4CC636F9C" lastPageId="15" lastPageNumber="43" pageId="8" pageNumber="36">
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<subSubSection id="C3296506FF9AFFC8D3F4F9E4CC7F6B25" pageId="8" pageNumber="36" type="nomenclature">
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<paragraph id="8B8C368DFF9AFFC8D3F4F9E4CF0F6B01" blockId="8.[151,614,1592,1692]" box="[151,614,1592,1619]" pageId="8" pageNumber="36">
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<heading id="D0C481E1FF9AFFC8D3F4F9E4CF0F6B01" bold="true" box="[151,614,1592,1619]" fontSize="11" level="1" pageId="8" pageNumber="36" reason="1">
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<emphasis id="B947EA9FFF9AFFC8D3F4F9E4CF0F6B01" bold="true" box="[151,614,1592,1619]" pageId="8" pageNumber="36">
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<taxonomicName id="4C334D0EFF9AFFC8D3F4F9E4CF686B00" box="[151,513,1593,1619]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="36" phylum="Chordata" rank="species" species="titiwangsaensis" status="sp. nov.">
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<emphasis id="B947EA9FFF9AFFC8D3F4F9E4CF686B00" bold="true" box="[151,513,1593,1619]" italics="true" pageId="8" pageNumber="36">Acanthosaura titiwangsaensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27457E4FF9AFFC8D16BF9E5CF0F6B01" box="[520,614,1592,1618]" pageId="8" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8C368DFF9AFFC8D3F4F981CC7F6B25" blockId="8.[151,614,1592,1692]" box="[151,278,1628,1654]" pageId="8" pageNumber="36">
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<figureCitation id="13082A08FF9AFFC8D3F4F981CC7F6B25" box="[151,278,1628,1654]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="11.[151,258,1152,1176]" captionStartId-1="12.[151,255,1259,1283]" captionTargetBox-0="[151,1434,194,1128]" captionTargetBox-1="[151,1434,388,1235]" captionTargetId-0="figure@11.[151,1436,194,1128]" captionTargetId-1="figure@12.[151,1436,388,1235]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="FIGURE 5. Adult male holotype (ZRC 2.6805) of Acanthosaura titiwangsaensis from Bishop’s trail Fraser’s Hill, Pahang, Peninsular Malaysia. Upper left: in life. Lower left: lateral view of the scalation on the left side of the body. Upper right: lateral view of the head. Lower right: dorsal view." captionText-1="FIGURE 6. Type series of Acanthosaura titiwangsaensis. Left: male holotype (ZRC 2.6805). Middle: female paratype (ZRC 2.5994). Right: juvenile male paratype (LSUHC 9094)." httpUri-0="https://zenodo.org/record/185827/files/figure.png" httpUri-1="https://zenodo.org/record/185828/files/figure.png" pageId="8" pageNumber="36">Figure 5–6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3296506FF9AFFC8D3F4F95FCF556BCF" box="[151,572,1666,1692]" pageId="8" pageNumber="36" type="description">
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<paragraph id="8B8C368DFF9AFFC8D3F4F95FCF556BCF" blockId="8.[151,614,1592,1692]" box="[151,572,1666,1692]" pageId="8" pageNumber="36">Malayan Mountain Horned Agamid</paragraph>
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</subSubSection>
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<subSubSection id="C3296506FF9AFFC8D3F4F912CC886A81" pageId="8" pageNumber="36" type="materials_examined">
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<paragraph id="8B8C368DFF9AFFC8D3F4F912CCC16A64" blockId="8.[151,1437,1743,2002]" pageId="8" pageNumber="36">
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<emphasis id="B947EA9FFF9AFFC8D3F4F912CC706BBA" bold="true" box="[151,281,1743,1769]" pageId="8" pageNumber="36">
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<typeStatus id="5488882FFF9AFFC8D3F4F912CC7A6BBA" box="[151,275,1743,1769]" pageId="8" pageNumber="36" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Adult male (
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<collectionCode id="ED22AE48FF9AFFC8D2A3F90DCC976BB9" box="[448,510,1744,1770]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="36">ZRC</collectionCode>
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2.6805) from Bishop’s Trail, Fraser’s Hill, Pahang, Peninsular
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<collectingCountry id="F324761DFF9AFFC8D649F90DC8F46BB9" box="[1322,1437,1744,1770]" name="Malaysia" pageId="8" pageNumber="36">Malaysia</collectingCountry>
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(
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<geoCoordinate id="EE07504AFF9AFFC8D3FCF92BCC4E6A43" box="[159,295,1782,1808]" direction="north" orientation="latitude" pageId="8" pageNumber="36" precision="1" value="3.7205">3º43.230'N</geoCoordinate>
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;
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<geoCoordinate id="EE07504AFF9AFFC8D251F92BCCB86A43" box="[306,465,1782,1808]" direction="east" orientation="longitude" pageId="8" pageNumber="36" precision="1" value="101.748116">101º44.887'E</geoCoordinate>
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) collected on
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<date id="FF8D104DFF9AFFC8D110F92BCE666A43" box="[627,783,1782,1808]" pageId="8" pageNumber="36" value="2008-06-28">28 June 2008</date>
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by Jesse L. Grismer, L. Lee Grismer, Perry L. Wood Jr., at 2330 h
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<quantity id="4CCB9B68FF9AFFC8D268F8C0CC036A64" box="[267,362,1821,1847]" metricMagnitude="3" metricUnit="m" metricValue="1.112" pageId="8" pageNumber="36" unit="m" value="1112.0">1,112 m</quantity>
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||
a.s.l.
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||
</paragraph>
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||
<paragraph id="8B8C368DFF9AFFC8D3A5F89ECC886A81" blockId="8.[151,1437,1743,2002]" pageId="8" pageNumber="36">
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<emphasis id="B947EA9FFF9AFFC8D3A5F89ECC226A0E" bold="true" box="[198,331,1859,1885]" pageId="8" pageNumber="36">
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<typeStatus id="5488882FFF9AFFC8D3A5F89ECC2F6A0E" box="[198,326,1859,1885]" pageId="8" pageNumber="36" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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||
<collectionCode id="ED22AE48FF9AFFC8D23BF899CCF86A0D" box="[344,401,1860,1886]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="36">ZRC</collectionCode>
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2.5994; gravid female from Pahang,
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<collectingCountry id="F324761DFF9AFFC8D004F899CEBA6A0D" box="[871,979,1860,1886]" name="Cameroon" pageId="8" pageNumber="36">Cameron</collectingCountry>
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Highlands, Telom Valley, Peninsular
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<collectingCountry id="F324761DFF9AFFC8D3F4F8B7CC676AD7" box="[151,270,1898,1924]" name="Malaysia" pageId="8" pageNumber="36">Malaysia</collectingCountry>
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, collected during March, 1935.
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<collectionCode id="ED22AE48FF9AFFC8D1C0F8B7CE616AD7" box="[675,776,1898,1924]" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="8" pageNumber="36">LSUHC</collectionCode>
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9094; juvenile male from Fraser’s Hill collected on Bishop’s Trial, Pahang, Peninsular
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<collectingCountry id="F324761DFF9AFFC8D15AF84CCFCC6AF8" box="[569,677,1937,1963]" name="Malaysia" pageId="8" pageNumber="36">Malaysia</collectingCountry>
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(
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||
<geoCoordinate id="EE07504AFF9AFFC8D1D5F84CCE576AF8" box="[694,830,1937,1963]" direction="north" orientation="latitude" pageId="8" pageNumber="36" precision="1" value="3.7205667">3º43.234'N</geoCoordinate>
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;
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<geoCoordinate id="EE07504AFF9AFFC8D028F84CCE836AF8" box="[843,1002,1937,1963]" direction="east" orientation="longitude" pageId="8" pageNumber="36" precision="1" value="101.748184">101º44.891'E</geoCoordinate>
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) on
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<date id="FF8D104DFF9AFFC8D743F84CC9D66AF8" box="[1056,1215,1937,1963]" pageId="8" pageNumber="36" value="2008-06-27">27 June 2008</date>
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by Perry L. Wood Jr. at 2100h at
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<quantity id="4CCB9B68FF9AFFC8D221F865CCCA6A81" box="[322,419,1976,2002]" metricMagnitude="3" metricUnit="m" metricValue="1.13" pageId="8" pageNumber="36" unit="m" value="1130.0">1,130 m</quantity>
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a.s.l.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3296506FF9BFFC9D3A5FF4AC9B86F67" pageId="9" pageNumber="37" type="diagnosis">
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<paragraph id="8B8C368DFF9BFFC9D3A5FF4AC9B86F67" blockId="9.[151,1437,151,1957]" pageId="9" pageNumber="37">
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<emphasis id="B947EA9FFF9BFFC9D3A5FF4ACC2F6DE2" bold="true" box="[198,326,151,177]" pageId="9" pageNumber="37">Diagnosis.</emphasis>
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Maximum SVL of adult males
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<quantity id="4CCB9B68FF9BFFC9D187FF45CE0D6DE1" box="[740,868,152,178]" metricMagnitude="-1" metricUnit="m" metricValue="1.184" pageId="9" pageNumber="37" unit="mm" value="118.4">118.4 mm</quantity>
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, maximum SVL of females
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||
<quantity id="4CCB9B68FF9BFFC9D7B1FF45C8386DE1" box="[1234,1361,152,178]" metricMagnitude="-1" metricUnit="m" metricValue="1.127" pageId="9" pageNumber="37" unit="mm" value="112.7">112.7 mm</quantity>
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||
; head triangular, snout short (SL/HL 46–52%); eye moderate (EYE/HL 28–30%); no enlarged elliptical scales below orbit; scales on the canthus rostralis-supraorbital ridge 14 or 15; large rectangular rostral scale; a short cylindrical spine above posterior margin of eye surrounded by a rosette of four smaller keeled scales at base; spine on occiput between tympanum and nuchal crest present; tympanum naked, oblong, taller than wide; dewlap present in males and females; body triangular in cross section; scales of flanks intermixed with randomly arranged clusters of larger keeled scales; nuchal crest present; 11–13 scales in nape diastema; vertebral crest composed of enlarged, pointed scales beginning at shoulder region and tapering to a series of enlarged keeled scales on anterior portion of tail; crest scales largest at base of nape; tail 140–150% of SVL; black collar present; posterior 2/3 of tail black to dark brown; greenish-yellow to light orange spots encircled by gray on body, gular region orangish yellow; no yellow medial stripe on dewlap (
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<tableCitation id="C6B10336FF9BFFC9D704FDC7C9D66F67" box="[1127,1215,538,564]" captionStart="TABLE 1" captionStartId="3.[151,239,847,871]" captionText="TABLE 1. All measurements (in mm) for all currently recognized species of Acanthosaura. PS – postorbital spine length, measured from the base to the tip of the spine; NS – the maximum length of the largest spine measured from the base to the tip; NSL – longest nuchal scale, measured at the base; DS – maximum length of longest spine on the body measured from the base to the tip; DSL – longest dorsal scale, measured at the base below the dorsal crest; WNC – maximum width of the spines in the nuchal crest, measured at the base; WDS – maximum width of the largest dorsal scale below the dorsal crest, measured at the base; DIAS – length of the diastema, measured from the posterior end of the nuchal crest to the anterior end of the nuchal crest; FOREL – forelimb length, measured from axilla to the proximal edge of the palmar region; HINDL – hindlimb length, measured from groin to the proximal edge plantar region; RS – number of scales bordering the rostral scale; NCS – number of scales between the fifth canthals; NSCSL – number of scales from the fifth canthal to the fifth supralabial; NR – number of scales between the nasals and the rostrals; NSSLC – number of scales between the seventh supralabial and the sixth canthal; LKP – presence or absence of light knee patch; GP – size of gular pouch scored as absent, small, medium or large; OF – presence or absence of oblique fold anterior to the fore limb insertions. For additional character abbreviations see materials and methods. / Indicates missing data. All data were taken from specimens or from the literature." pageId="9" pageNumber="37">Table 1</tableCitation>
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||
).
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3296506FF9BFFCBD3A5FD9DCE046BFF" lastPageId="11" lastPageNumber="39" pageId="9" pageNumber="37" type="description">
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<paragraph id="8B8C368DFF9BFFC9D3A5FD9DC9686B1B" blockId="9.[151,1437,151,1957]" pageId="9" pageNumber="37">
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<emphasis id="B947EA9FFF9BFFC9D3A5FD9DCF486F09" bold="true" box="[198,545,576,602]" pageId="9" pageNumber="37">
|
||
Description of the
|
||
<typeStatus id="5488882FFF9BFFC9D2CCFD9DCF486F09" box="[431,545,576,602]" pageId="9" pageNumber="37" type="holotype">Holotype</typeStatus>
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||
</emphasis>
|
||
. (
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||
<figureCitation id="13082A08FF9BFFC9D15AFD9CCFE86F08" box="[569,641,577,603]" captionStart="FIGURE 5" captionStartId="11.[151,258,1152,1176]" captionTargetBox="[151,1434,194,1128]" captionTargetId="figure@11.[151,1436,194,1128]" captionTargetPageId="11" captionText="FIGURE 5. Adult male holotype (ZRC 2.6805) of Acanthosaura titiwangsaensis from Bishop’s trail Fraser’s Hill, Pahang, Peninsular Malaysia. Upper left: in life. Lower left: lateral view of the scalation on the left side of the body. Upper right: lateral view of the head. Lower right: dorsal view." httpUri="https://zenodo.org/record/185827/files/figure.png" pageId="9" pageNumber="37">Fig. 5</figureCitation>
|
||
). Adult male, SVL
|
||
<quantity id="4CCB9B68FF9BFFC9D009FD9CCE8D6F08" box="[874,996,577,603]" metricMagnitude="-1" metricUnit="m" metricValue="1.184" pageId="9" pageNumber="37" unit="mm" value="118.4">118.4 mm</quantity>
|
||
, TL 174.0 mm. Head short (HL/SVL 20%), head narrow (HW/SVL 20%), head tall (HD/HL 80%), head triangular in dorsal and lateral profile; moderate snout (SL/HL 50%); interorbital and frontal regions, and rostrum wide (RW/RH 250%); steeply sloping anteriorly; canthus rostralis prominent, forming a large projecting shelf extending posteriorly above eye, composed of 15 enlarged scales, shelf terminates with a notch anterior to the postorbital spine surrounded by four smaller keeled scales; rostral large, rectangular, bordered laterally by the first supralabials and posteriorly by nine smaller scales; nasals crater-like, surrounded by two prenasal and four postnasal scales; eight scales between nasal scales; supranasals somewhat elongate; enlarged scales above orbit weakly keeled; an enlarged row of keeled scales below orbit extending from anterior margin of eye to middle of eye; eye large (EYE/HL 30%); interorbital, frontal, and prefrontal scales keeled and smaller then enlarged row of keeled scales below the orbit; no hair organs on anterior end of keels in lateral frontal scales; three enlarged, keeled, azygous prefrontals forming a Y-shaped pattern; scales of occiput flat to weakly keeled; enlarged parietals, no pineal eyespot; short cylindrical epidermal spine above posterior margin of eye; occipital spine surrounded by four enlarged keeled scales on the left side and five on the right side; suborbital scales enlarged, keeled extending in a single row of six from below posterior margin of eye to anterior margin of tympanum, increasing in size posteriorly, terminal scale spine-like; elongate cylindrical epidermal spine on lateral margin of nape; tympanum exposed, oblong, taller than wide surrounded by small conical scales; supralabials 12, rectangular, not decreasing in size posteriorly; mental somewhat square anteriorly and becoming triangular posteriorly, much larger than adjacent infralabials; two postmentals similar in size, all five scales contacting mental; chinshields enlarged, extending posteriorly past angle of jaw, separated from infralabials by one scale row anteriorly and three at angle of jaw; infralabials 12, rectangular; gulars sharply keeled and spinose with no enlarged midventral row; extensible dewlap present; large gular pouch; nuchal crest with a diastema 10 scales wide at base of nape; nuchal crest composed of eight elongate, lancolate scales bordered on each side by two rows of enlarged, spinose scales; dorsal body crest present, extending from posterior margin of diastema onto base of tail, vertebral crest composed of enlarged epidermal, laterally compressed, spinose scales, bordered by a single row of slightly smaller paravertebral squarish, spinose scales; vertebral crest tapers slightly to base of tail then fades approximately one-third down tail.
|
||
</paragraph>
|
||
<paragraph id="8B8C368DFF9BFFC9D3A6F988CF786AF6" blockId="9.[151,1437,151,1957]" pageId="9" pageNumber="37">Body moderately developed, triangular in cross-section; dorsal scales small with medially projecting spines; larger randomly arranged scale clusters among dorsals; scales of pectoral region and abdomen larger than dorsals, keeled, spinose, arranged in semi-transverse rows; no enlarged, keeled scales anterior to vent; limbs relatively long; dorsal and ventral scales of forelimbs keeled, spinose, and nearly the same size; five digits on manus; subdigital scales keeled; scales of hind limbs keeled and spinose; postfemoral scales small, interspersed with larger spinose scales; five digits on appendages; subdigital scales keeled; subdigital lamellae on the fourth finger 21; subdigital lamellae on fourth toe 25; tail 146% of SVL, covered with keeled spinose scales; keels on subcaudals posteriorly directed, same size as keels of other caudal scales; base of tail in males nearly twice as wide as females.</paragraph>
|
||
<caption id="DF4C6605FF98FFCAD3F4F872CF0F6ABA" httpUri="https://zenodo.org/record/185826/files/figure.png" pageId="10" pageNumber="38" targetBox="[199,1385,194,1937]" targetPageId="10">
|
||
<paragraph id="8B8C368DFF98FFCAD3F4F872CF0F6ABA" blockId="10.[151,1436,1967,2025]" pageId="10" pageNumber="38">
|
||
<emphasis id="B947EA9FFF98FFCAD3F4F872CC496A94" bold="true" box="[151,288,1967,1991]" pageId="10" pageNumber="38">FIGURE 4.</emphasis>
|
||
Habitat of type locality of
|
||
<taxonomicName id="4C334D0EFF98FFCAD13BF872CEE46A95" box="[600,909,1967,1990]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="38" phylum="Chordata" rank="species" species="bintangensis">
|
||
<emphasis id="B947EA9FFF98FFCAD13BF872CEE46A95" box="[600,909,1967,1990]" italics="true" pageId="10" pageNumber="38">Acanthosaura bintangensis</emphasis>
|
||
</taxonomicName>
|
||
from Bukit Larut, Perak, Peninsular Malaysia (04º51.446'N; 100º48.455'E, 1351 m a.s.l.).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4C6605FF99FFCBD3F4FB5DCE5F698E" httpUri="https://zenodo.org/record/185827/files/figure.png" pageId="11" pageNumber="39" targetBox="[151,1434,194,1128]" targetPageId="11">
|
||
<paragraph id="8B8C368DFF99FFCBD3F4FB5DCE5F698E" blockId="11.[151,1437,1152,1245]" pageId="11" pageNumber="39">
|
||
<emphasis id="B947EA9FFF99FFCBD3F4FB5DCC7769CB" bold="true" box="[151,286,1152,1176]" pageId="11" pageNumber="39">FIGURE 5.</emphasis>
|
||
Adult male holotype (ZRC 2.6805) of
|
||
<taxonomicName id="4C334D0EFF99FFCBD1B5FB5CC94C69CB" box="[726,1061,1153,1176]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="39" phylum="Chordata" rank="species" species="titiwangsaensis">
|
||
<emphasis id="B947EA9FFF99FFCBD1B5FB5CC94C69CB" box="[726,1061,1153,1176]" italics="true" pageId="11" pageNumber="39">Acanthosaura titiwangsaensis</emphasis>
|
||
</taxonomicName>
|
||
from Bishop’s trail Fraser’s Hill, Pahang, Peninsular Malaysia. Upper left: in life. Lower left: lateral view of the scalation on the left side of the body. Upper right: lateral view of the head. Lower right: dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8C368DFF99FFCBD3A5FAD2CE6E6B42" blockId="11.[151,1437,1295,2017]" pageId="11" pageNumber="39">
|
||
<emphasis id="B947EA9FFF99FFCBD3A5FAD2CCF4687A" bold="true" box="[198,413,1295,1321]" pageId="11" pageNumber="39">Coloration in life</emphasis>
|
||
. (
|
||
<figureCitation id="13082A08FF99FFCBD2D6FAD2CC97687A" box="[437,510,1295,1321]" captionStart="FIGURE 5" captionStartId="11.[151,258,1152,1176]" captionTargetBox="[151,1434,194,1128]" captionTargetId="figure@11.[151,1436,194,1128]" captionTargetPageId="11" captionText="FIGURE 5. Adult male holotype (ZRC 2.6805) of Acanthosaura titiwangsaensis from Bishop’s trail Fraser’s Hill, Pahang, Peninsular Malaysia. Upper left: in life. Lower left: lateral view of the scalation on the left side of the body. Upper right: lateral view of the head. Lower right: dorsal view." httpUri="https://zenodo.org/record/185827/files/figure.png" pageId="11" pageNumber="39">Fig. 5</figureCitation>
|
||
). Dorsal ground color of the head, body, and limbs orangish-yellow; sides and the top of the head immaculate orangish-yellow; eye patch dark and restricted to orbit not extending into the postorbital region; iris brownish-orange; black collar present fading dorsally; gular region same orangishyellow; body covered with medium-sized light orange spots edged in a faded black color and extend onto base of tail; undersides of limbs pale orange; forelimbs nearly uniform in color; smaller spots similar to that of the body on hind limbs; no faint knee patches; posterior 2/3 of tail black remainder of tail similar to color of body; abdomen pale orange with irregular darker markings.
|
||
</paragraph>
|
||
<paragraph id="8B8C368DFF99FFCBD3A6F9C3CE046BFF" blockId="11.[151,1437,1295,2017]" pageId="11" pageNumber="39">
|
||
<emphasis id="B947EA9FFF99FFCBD3A6F9C3CC296B6B" bold="true" box="[197,320,1566,1592]" pageId="11" pageNumber="39">Variation.</emphasis>
|
||
(
|
||
<figureCitation id="13082A08FF99FFCBD231F9C3CCF36B6B" box="[338,410,1566,1592]" captionStart="FIGURE 6" captionStartId="12.[151,255,1259,1283]" captionTargetBox="[151,1434,388,1235]" captionTargetId="figure@12.[151,1436,388,1235]" captionTargetPageId="12" captionText="FIGURE 6. Type series of Acanthosaura titiwangsaensis. Left: male holotype (ZRC 2.6805). Middle: female paratype (ZRC 2.5994). Right: juvenile male paratype (LSUHC 9094)." httpUri="https://zenodo.org/record/185828/files/figure.png" pageId="11" pageNumber="39">Fig. 6</figureCitation>
|
||
). The
|
||
<typeStatus id="5488882FFF99FFCBD286F9C3CF3C6B6B" box="[485,597,1566,1592]" pageId="11" pageNumber="39" type="paratype">paratypes</typeStatus>
|
||
resemble the
|
||
<typeStatus id="5488882FFF99FFCBD199F9C3CE086B6B" box="[762,865,1566,1592]" pageId="11" pageNumber="39" type="holotype">holotype</typeStatus>
|
||
in all aspects of morphology except the juvenile male LSUHC 9094 has more but smaller spots on the dorsum. The head is light green and the dorsum is yellowish-green with yellow spots. Adult female ZRC 2.5994 has fewer but larger spots on the dorsum. Scalation and morphological variation is presented in
|
||
<tableCitation id="C6B10336FF99FFCBD06CF94FCE016BFF" box="[783,872,1682,1708]" captionStart="TABLE 3" captionStartId="13.[151,246,151,175]" captionText="TABLE 3. Selected measurements (in mm) for the type series of Acanthosaura titiwangsaensis. For character abbreviations see materials and methods." pageId="11" pageNumber="39">Table 3</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3296506FF99FFCCD3A6F965C9F06DAC" lastPageId="12" lastPageNumber="40" pageId="11" pageNumber="39" type="distribution">
|
||
<paragraph id="8B8C368DFF99FFCBD3A6F965CC5F6A3E" blockId="11.[151,1437,1295,2017]" pageId="11" pageNumber="39">
|
||
<emphasis id="B947EA9FFF99FFCBD3A6F965CC1F6B81" bold="true" box="[197,374,1720,1746]" pageId="11" pageNumber="39">Distribution.</emphasis>
|
||
<taxonomicName id="4C334D0EFF99FFCBD21DF964CE686B81" box="[382,769,1721,1746]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="39" phylum="Chordata" rank="species" species="titiwangsaensis">
|
||
<emphasis id="B947EA9FFF99FFCBD21DF964CE686B81" box="[382,769,1721,1746]" italics="true" pageId="11" pageNumber="39">Acanthosaura titiwangsaensis</emphasis>
|
||
</taxonomicName>
|
||
is only known from the montane areas of
|
||
<collectingCountry id="F324761DFF99FFCBD64AF964C8F46B80" box="[1321,1437,1721,1747]" name="Cameroon" pageId="11" pageNumber="39">Cameron</collectingCountry>
|
||
Highlands and Bukit Fraser in the Banjaran Titi Wangsa, Peninsular
|
||
<collectingCountry id="F324761DFF99FFCBD0A5F902C95F6BAA" box="[966,1078,1759,1785]" name="Malaysia" pageId="11" pageNumber="39">Malaysia</collectingCountry>
|
||
. Given that this range extends into southern
|
||
<collectingCountry id="F324761DFF99FFCBD25AF8DBCCC86A73" box="[313,417,1798,1824]" name="Thailand" pageId="11" pageNumber="39">Thailand</collectingCountry>
|
||
and other montane endemics restricted to this range extend across the Thai-Malaysian border (
|
||
<bibRefCitation id="EFA24B7CFF99FFCBD396F8F0CCD26A14" author="Boulenger" box="[245,443,1837,1863]" pageId="11" pageNumber="44" refString="Boulenger, G. A. (1903) Report on the batrachians and reptiles. P. 131 - 176 in Fasciculii Malayenses. Anthropological and zoological results of an Expedition to Perak and the Siamese Malay States 1901 - 1902, undertaken by Nelson Annandale and Herbert C. Robinson. Vol. 1. Zoology. University Press, Liverpool." type="book chapter" year="1903">Boulenger 1903</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA24B7CFF99FFCBD2ABF8F0CF0D6A14" author="Flower" box="[456,612,1837,1863]" pageId="11" pageNumber="44" refString="Flower, S. S. (1896) Notes on a collection of reptiles and batrachians made in the Malay Peninsula in 1895 - 96, with a list of species recorded from that region. Proceedings of the Zoological Society of London, 1896, 856 - 914." type="proceedings paper" year="1896">Flower 1896</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFA24B7CFF99FFCBD112F8F0CFD86A14" author="Flower" box="[625,689,1837,1863]" pageId="11" pageNumber="44" refString="Flower, S. S. (1899) Notes on a second collection of batrachians made in the Malay Peninsula and Siam in 1895 - 96, with a list of the species recorded from that region. Proceedings of the Zoological Society of London, 1899, 600 - 697." type="proceedings paper" year="1899">1899</bibRefCitation>
|
||
), it is possible that
|
||
<taxonomicName id="4C334D0EFF99FFCBD0FFF8F0C91F6A15" box="[924,1142,1837,1862]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="39" phylum="Chordata" rank="species" species="titiwangsaensis">
|
||
<emphasis id="B947EA9FFF99FFCBD0FFF8F0C91F6A15" box="[924,1142,1837,1862]" italics="true" pageId="11" pageNumber="39">A. titiwangsaensis</emphasis>
|
||
</taxonomicName>
|
||
may also extend further north as well.
|
||
</paragraph>
|
||
<paragraph id="8B8C368DFF99FFCCD3A5F8A7C9F06DAC" blockId="11.[151,1437,1295,2017]" lastBlockId="12.[151,1437,152,332]" lastPageId="12" lastPageNumber="40" pageId="11" pageNumber="39">
|
||
<emphasis id="B947EA9FFF99FFCBD3A5F8A7CCFC6AC7" bold="true" box="[198,405,1914,1940]" pageId="11" pageNumber="39">Natural history.</emphasis>
|
||
This species was found at night sleeping head-up on the trunks of small trees, approximately
|
||
<quantity id="4CCB9B68FF99FFCBD228F87CCCEE6AE8" box="[331,391,1953,1979]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="11" pageNumber="39" unit="cm" value="8.0">8 cm</quantity>
|
||
in diameter, approximately
|
||
<quantity id="4CCB9B68FF99FFCBD1B9F87CCE4F6AE8" box="[730,806,1953,1979]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="11" pageNumber="39" unit="m" value="2.5" valueMax="3.0" valueMin="2.0">2–3 m</quantity>
|
||
above the ground. We did not see lizards during the day but presume its diurnal habits resemble those of
|
||
<taxonomicName id="4C334D0EFF99FFCBD053F815C9ED6AB2" box="[816,1156,1992,2017]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="39" phylum="Chordata" rank="species" species="bintangensis">
|
||
<emphasis id="B947EA9FFF99FFCBD053F815C9ED6AB2" box="[816,1156,1992,2017]" italics="true" pageId="11" pageNumber="39">Acanthosaura bintangensis</emphasis>
|
||
</taxonomicName>
|
||
. Like other species of
|
||
<emphasis id="B947EA9FFF9EFFCCD3F4FF45CC2B6DE2" box="[151,322,152,177]" italics="true" pageId="12" pageNumber="40">
|
||
<taxonomicName id="4C334D0EFF9EFFCCD3F4FF45CC566DE2" box="[151,319,152,177]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="40" phylum="Chordata" rank="genus">Acanthosaura</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
we believe this species forages on the ground for food, digging through the soil (
|
||
<bibRefCitation id="EFA24B7CFF9EFFCCD661FF45C8FD6DE1" author="Taylor" box="[1282,1428,152,178]" pageId="12" pageNumber="45" refString="Taylor, E. H. (1963) The Lizards of Thailand. Kansas University Science Bulletin, 46 (5), 687 - 1077." type="journal article" year="1963">Taylor 1963</bibRefCitation>
|
||
) based on soil fragments found on the labial scales and within the external nares of specimens from Fraser’s Hill. A gravid female was found during March (ZRC 25994) from
|
||
<collectingCountry id="F324761DFF9EFFCCD0C6FF38C9786DAC" box="[933,1041,229,255]" name="Cameroon" pageId="12" pageNumber="40">Cameron</collectingCountry>
|
||
Highlands.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3296506FF9EFFCDD3A6FED6C90269FC" lastPageId="13" lastPageNumber="41" pageId="12" pageNumber="40" type="etymology">
|
||
<paragraph id="8B8C368DFF9EFFCCD3A6FED6CEA86C1F" blockId="12.[151,1437,152,332]" pageId="12" pageNumber="40">
|
||
<emphasis id="B947EA9FFF9EFFCCD3A6FED6CC346C76" bold="true" box="[197,349,267,293]" pageId="12" pageNumber="40">Etymology.</emphasis>
|
||
This specific epithet
|
||
<emphasis id="B947EA9FFF9EFFCCD115FED1CE5B6C76" box="[630,818,268,293]" italics="true" pageId="12" pageNumber="40">titwangsaensis</emphasis>
|
||
is in reference to the mountain range, Banjaran Titiwangsa, to which we believe this species is likely to be restricted.
|
||
</paragraph>
|
||
<caption id="DF4C6605FF9EFFCCD3F4FB36CE406876" httpUri="https://zenodo.org/record/185828/files/figure.png" pageId="12" pageNumber="40" targetBox="[151,1434,388,1235]" targetPageId="12">
|
||
<paragraph id="8B8C368DFF9EFFCCD3F4FB36CE406876" blockId="12.[151,1436,1259,1317]" pageId="12" pageNumber="40">
|
||
<emphasis id="B947EA9FFF9EFFCCD3F4FB36CC726850" bold="true" box="[151,283,1259,1283]" pageId="12" pageNumber="40">FIGURE 6.</emphasis>
|
||
Type series of
|
||
<taxonomicName id="4C334D0EFF9EFFCCD2A3FB36CE6A6851" box="[448,771,1259,1282]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="40" phylum="Chordata" rank="species" species="titiwangsaensis">
|
||
<emphasis id="B947EA9FFF9EFFCCD2A3FB36CE6A6851" box="[448,771,1259,1282]" italics="true" pageId="12" pageNumber="40">Acanthosaura titiwangsaensis</emphasis>
|
||
</taxonomicName>
|
||
. Left: male holotype (ZRC 2.6805). Middle: female paratype (ZRC 2.5994). Right: juvenile male paratype (LSUHC 9094).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4C6605FF9EFFCCD3F4F88ACF236AC2" httpUri="https://zenodo.org/record/185829/files/figure.png" pageId="12" pageNumber="40" targetBox="[151,1434,1371,1854]" targetPageId="12">
|
||
<paragraph id="8B8C368DFF9EFFCCD3F4F88ACF236AC2" blockId="12.[151,1435,1879,1937]" pageId="12" pageNumber="40">
|
||
<emphasis id="B947EA9FFF9EFFCCD3F4F88ACC706A3C" bold="true" box="[151,281,1879,1903]" pageId="12" pageNumber="40">FIGURE 7.</emphasis>
|
||
Illustration of the two enlarged, keeled, elliptical suborbital scales in
|
||
<taxonomicName id="4C334D0EFF9EFFCCD762F88AC84D6A3D" box="[1025,1316,1879,1902]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="40" phylum="Chordata" rank="species" species="bintangensis">
|
||
<emphasis id="B947EA9FFF9EFFCCD762F88AC84D6A3D" box="[1025,1316,1879,1902]" italics="true" pageId="12" pageNumber="40">Acanthosaura bintangensis</emphasis>
|
||
</taxonomicName>
|
||
(right) and their absence in
|
||
<taxonomicName id="4C334D0EFF9EFFCCD226F8A7CF6E6AC2" box="[325,519,1914,1937]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="40" phylum="Chordata" rank="species" species="titiwangsaensis">
|
||
<emphasis id="B947EA9FFF9EFFCCD226F8A7CF6E6AC2" box="[325,519,1914,1937]" italics="true" pageId="12" pageNumber="40">A. titiwangsaensis</emphasis>
|
||
</taxonomicName>
|
||
(left).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4C6605FF9FFFCDD3F4FF4ACF246D82" pageId="13" pageNumber="41">
|
||
<paragraph id="8B8C368DFF9FFFCDD3F4FF4ACF246D82" blockId="13.[151,1437,151,2009]" pageId="13" pageNumber="41">
|
||
<emphasis id="B947EA9FFF9FFFCDD3F4FF4ACC7C6DFC" bold="true" box="[151,277,151,175]" pageId="13" pageNumber="41">TABLE 3.</emphasis>
|
||
Selected measurements (in mm) for the type series of
|
||
<taxonomicName id="4C334D0EFF9FFFCDD0F6FF45C99B6DFC" box="[917,1266,152,175]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="41" phylum="Chordata" rank="species" species="titiwangsaensis">
|
||
<emphasis id="B947EA9FFF9FFFCDD0F6FF45C99B6DFC" box="[917,1266,152,175]" italics="true" pageId="13" pageNumber="41">Acanthosaura titiwangsaensis</emphasis>
|
||
</taxonomicName>
|
||
. For character abbreviations see materials and methods.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8C368DFF9FFFCDD3FCFF30C9346F0A" blockId="13.[151,1437,151,2009]" pageId="13" pageNumber="41">
|
||
Sex ZRC 2.6805
|
||
<typeStatus id="5488882FFF9FFFCDD285FF30CF206C57" box="[486,585,237,260]" pageId="13" pageNumber="41" type="holotype">Holotype</typeStatus>
|
||
Male LSUHC 9094
|
||
<typeStatus id="5488882FFF9FFFCDD013FF30CEA76C57" box="[880,974,237,260]" pageId="13" pageNumber="41" type="paratype">Paratype</typeStatus>
|
||
Juvenile ZRC 2.5994
|
||
<typeStatus id="5488882FFF9FFFCDD7B9FF30C85E6C57" box="[1242,1335,237,260]" pageId="13" pageNumber="41" type="paratype">Paratype</typeStatus>
|
||
Female SVL 118.4 91.8 112.7 TL 174 136 160 HL 24 20 24.3 HW 23.4 17.5 20.7 HD 20.2 15.7 18.6 SL 12.5 9.7 11.4 EYE 7.2 5.6 7.7 TD 3.7 2.7 4
|
||
</paragraph>
|
||
<paragraph id="8B8C368DFF9FFFCDD3FCFDB0C9346FD7" blockId="13.[151,1437,151,2009]" box="[159,1117,621,644]" pageId="13" pageNumber="41">TN 1 1 1</paragraph>
|
||
<paragraph id="8B8C368DFF9FFFCDD3FCFD45C9346E0A" blockId="13.[151,1437,151,2009]" pageId="13" pageNumber="41">PS 4.4 3.3 3.5 NS 4.4 3.8 2.7 NSL 2.4 1.6 1.9 DS 2.1 2 1.7 DSL 1.9 1.5 2</paragraph>
|
||
<paragraph id="8B8C368DFF9FFFCDD3FCFCB0C9036957" blockId="13.[151,1437,151,2009]" pageId="13" pageNumber="41">
|
||
WN 1.6 1.6 1.4 WDS 2.3 1.6 2.1 DIAS 5.9 5.1 7.6 DIASN
|
||
<date id="FF8D104DFF9FFFCDD23FFC30C9036957" box="[348,1130,1005,1028]" pageId="13" pageNumber="41" value="1910-11-13">10 13 11</date>
|
||
</paragraph>
|
||
<paragraph id="8B8C368DFF9FFFCDD3FCFBC5C90269FC" blockId="13.[151,1437,151,2009]" pageId="13" pageNumber="41">
|
||
DIAS/SVL 0.05 0.06 0.07 FOREL 47.6 38 51.7 HINDL 65.6 48.5 60.5 SUPRAL
|
||
<date id="FF8D104DFF9FFFCDD23FFB45C90269FC" box="[348,1131,1176,1199]" pageId="13" pageNumber="41" value="1912-12-13">12 13 12</date>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3296506FF9FFFCFD3FCFB1FCC636F9C" lastPageId="15" lastPageNumber="43" pageId="13" pageNumber="41" type="description">
|
||
<paragraph id="8B8C368DFF9FFFCDD3FCFB1FC903698A" blockId="13.[151,1437,151,2009]" box="[159,1130,1218,1241]" pageId="13" pageNumber="41">
|
||
INFRAL
|
||
<date id="FF8D104DFF9FFFCDD23FFB1FC903698A" box="[348,1130,1218,1241]" pageId="13" pageNumber="41" value="1911-11-12">12 11 11</date>
|
||
</paragraph>
|
||
<paragraph id="8B8C368DFF9FFFCDD3FCFB30C9026857" blockId="13.[151,1437,151,2009]" box="[159,1131,1261,1284]" pageId="13" pageNumber="41">VENT 57 54 47</paragraph>
|
||
<paragraph id="8B8C368DFF9FFFCDD3FCFAC5C902687C" blockId="13.[151,1437,151,2009]" box="[159,1131,1304,1327]" pageId="13" pageNumber="41">FI 21 21 20</paragraph>
|
||
<paragraph id="8B8C368DFF9FFFCDD3FCFA9FC902680A" blockId="13.[151,1437,151,2009]" box="[159,1131,1346,1369]" pageId="13" pageNumber="41">TO 25 27 23</paragraph>
|
||
<paragraph id="8B8C368DFF9FFFCDD3FCFAB0C934688A" blockId="13.[151,1437,151,2009]" pageId="13" pageNumber="41">TL/SVL 1.5 1.5 1.4 OS 2.3 1.8 2.2 NSSOS 4 or 5 5 5</paragraph>
|
||
<paragraph id="8B8C368DFF9FFFCDD3FCFA30C9026B57" blockId="13.[151,1437,151,2009]" box="[159,1131,1517,1540]" pageId="13" pageNumber="41">CS 15 15 14</paragraph>
|
||
<paragraph id="8B8C368DFF9FFFCDD3FCF9C5C9346B0A" blockId="13.[151,1437,151,2009]" pageId="13" pageNumber="41">RW 4.5 3.6 5.2 RH 1.8 1.4 2</paragraph>
|
||
<paragraph id="8B8C368DFF9FFFCDD3FCF9B0C9346BD7" blockId="13.[151,1437,151,2009]" box="[159,1117,1645,1668]" pageId="13" pageNumber="41">RS 9 9 9</paragraph>
|
||
<paragraph id="8B8C368DFF9FFFCDD3FCF945C9346BFC" blockId="13.[151,1437,151,2009]" box="[159,1117,1688,1711]" pageId="13" pageNumber="41">NS 8 8 8</paragraph>
|
||
<paragraph id="8B8C368DFF9FFFCDD3FCF91FC9036B8A" blockId="13.[151,1437,151,2009]" box="[159,1130,1730,1753]" pageId="13" pageNumber="41">
|
||
NCS
|
||
<date id="FF8D104DFF9FFFCDD23FF91FC9036B8A" box="[348,1130,1730,1753]" pageId="13" pageNumber="41" value="1911-12-11">11 12 11</date>
|
||
</paragraph>
|
||
<paragraph id="8B8C368DFF9FFFCDD3FCF930C9036A57" blockId="13.[151,1437,151,2009]" box="[159,1130,1773,1796]" pageId="13" pageNumber="41">
|
||
NSCSL
|
||
<date id="FF8D104DFF9FFFCDD23FF930C9036A57" box="[348,1130,1773,1796]" pageId="13" pageNumber="41" value="1911-09-07">7 9 11</date>
|
||
</paragraph>
|
||
<paragraph id="8B8C368DFF9FFFCDD3FCF8C5C9346A7C" blockId="13.[151,1437,151,2009]" box="[159,1117,1816,1839]" pageId="13" pageNumber="41">NR 2 1 1</paragraph>
|
||
<paragraph id="8B8C368DFF9FFFCDD3FCF89FC9026A0A" blockId="13.[151,1437,151,2009]" box="[159,1131,1858,1881]" pageId="13" pageNumber="41">
|
||
NSSLC
|
||
<date id="FF8D104DFF9FFFCDD23FF89FC9026A0A" box="[348,1131,1858,1881]" pageId="13" pageNumber="41" value="1914-12-11">11 12 14</date>
|
||
</paragraph>
|
||
<paragraph id="8B8C368DFF9FFFCDD3FCF8B0C9346AD7" blockId="13.[151,1437,151,2009]" box="[159,1117,1901,1924]" pageId="13" pageNumber="41">MW 1.8 1.4 2</paragraph>
|
||
<paragraph id="8B8C368DFF9FFFCDD3FCF845C9186AFC" blockId="13.[151,1437,151,2009]" box="[159,1137,1944,1967]" pageId="13" pageNumber="41">MH 1.8 1.4 2.4</paragraph>
|
||
<paragraph id="8B8C368DFF9FFFCDD78AF81FC8FC6A8A" blockId="13.[151,1437,151,2009]" box="[1257,1429,1986,2009]" pageId="13" pageNumber="41">to be continued.</paragraph>
|
||
<paragraph id="8B8C368DFF9CFFCED3FCFF42C9346F64" pageId="14" pageNumber="42">
|
||
<table id="F933C42DFF9C003FD3FCFF42C8E46F64" box="[159,1421,159,567]" colsContinueFrom="8.[159,1418,159,865]" gridcols="3" gridrows="10" pageId="14" pageNumber="42">
|
||
<tr id="350334CFFF9C003FD3FCFF42C8E46DE4" box="[159,1421,159,183]" gridrow="0" pageId="14" pageNumber="42" rowspan-1="1" rowspan-2="1">
|
||
<th id="76D25DB3FF9C003FD3FCFF42CFEE6DE4" box="[159,647,159,183]" gridcol="0" gridrow="0" pageId="14" pageNumber="42">
|
||
<emphasis id="B947EA9FFF9CFFCED3FCFF42CC7D6DE4" bold="true" box="[159,276,159,183]" pageId="14" pageNumber="42">TABLE 3.</emphasis>
|
||
(continued)
|
||
</th>
|
||
</tr>
|
||
<tr id="350334CFFF9C003FD3FCFF17C8E46DB2" box="[159,1421,202,225]" gridrow="1" pageId="14" pageNumber="42">
|
||
<th id="76D25DB3FF9C003FD3FCFF17CFEE6DB2" box="[159,647,202,225]" gridcol="0" gridrow="1" pageId="14" pageNumber="42">Sex ZRC 2.6805 Holotype Male</th>
|
||
<td id="76D25DB3FF9C003FD1B5FF17C9446DB2" box="[726,1069,202,225]" gridcol="1" gridrow="1" pageId="14" pageNumber="42">LSUHC 9094 Paratype Juvenile</td>
|
||
<td id="76D25DB3FF9C003FD733FF17C8E46DB2" box="[1104,1421,202,225]" gridcol="2" gridrow="1" pageId="14" pageNumber="42">ZRC 2.5994 Paratype Female</td>
|
||
</tr>
|
||
<tr id="350334CFFF9C003FD3FCFF28C8E46C5F" box="[159,1421,245,268]" gridrow="2" pageId="14" pageNumber="42">
|
||
<th id="76D25DB3FF9C003FD3FCFF28CFEE6C5F" box="[159,647,245,268]" gridcol="0" gridrow="2" pageId="14" pageNumber="42">PM 5</th>
|
||
<td id="76D25DB3FF9C003FD1B5FF28C9446C5F" box="[726,1069,245,268]" gridcol="1" gridrow="2" pageId="14" pageNumber="42">5</td>
|
||
<td id="76D25DB3FF9C003FD733FF28C8E46C5F" box="[1104,1421,245,268]" gridcol="2" gridrow="2" pageId="14" pageNumber="42">5</td>
|
||
</tr>
|
||
<tr id="350334CFFF9C003FD3FCFEFDC8E46C64" box="[159,1421,288,311]" gridrow="3" pageId="14" pageNumber="42">
|
||
<th id="76D25DB3FF9C003FD3FCFEFDCFEE6C64" box="[159,647,288,311]" gridcol="0" gridrow="3" pageId="14" pageNumber="42">YAS 1</th>
|
||
<td id="76D25DB3FF9C003FD1B5FEFDC9446C64" box="[726,1069,288,311]" gridcol="1" gridrow="3" pageId="14" pageNumber="42">1</td>
|
||
<td id="76D25DB3FF9C003FD733FEFDC8E46C64" box="[1104,1421,288,311]" gridcol="2" gridrow="3" pageId="14" pageNumber="42">1</td>
|
||
</tr>
|
||
<tr id="350334CFFF9C003FD3FCFE97C8E46C32" box="[159,1421,330,353]" gridrow="4" pageId="14" pageNumber="42">
|
||
<th id="76D25DB3FF9C003FD3FCFE97CFEE6C32" box="[159,647,330,353]" gridcol="0" gridrow="4" pageId="14" pageNumber="42">ND 1</th>
|
||
<td id="76D25DB3FF9C003FD1B5FE97C9446C32" box="[726,1069,330,353]" gridcol="1" gridrow="4" pageId="14" pageNumber="42">1</td>
|
||
<td id="76D25DB3FF9C003FD733FE97C8E46C32" box="[1104,1421,330,353]" gridcol="2" gridrow="4" pageId="14" pageNumber="42">1</td>
|
||
</tr>
|
||
<tr id="350334CFFF9C003FD3FCFEA8C8E46CDF" box="[159,1421,373,396]" gridrow="5" pageId="14" pageNumber="42">
|
||
<th id="76D25DB3FF9C003FD3FCFEA8CFEE6CDF" box="[159,647,373,396]" gridcol="0" gridrow="5" pageId="14" pageNumber="42">LKP 0</th>
|
||
<td id="76D25DB3FF9C003FD1B5FEA8C9446CDF" box="[726,1069,373,396]" gridcol="1" gridrow="5" pageId="14" pageNumber="42">0</td>
|
||
<td id="76D25DB3FF9C003FD733FEA8C8E46CDF" box="[1104,1421,373,396]" gridcol="2" gridrow="5" pageId="14" pageNumber="42">0</td>
|
||
</tr>
|
||
<tr id="350334CFFF9C003FD3FCFE7DC8E46CE4" box="[159,1421,416,439]" gridrow="6" pageId="14" pageNumber="42">
|
||
<th id="76D25DB3FF9C003FD3FCFE7DCFEE6CE4" box="[159,647,416,439]" gridcol="0" gridrow="6" pageId="14" pageNumber="42">BEP 1</th>
|
||
<td id="76D25DB3FF9C003FD1B5FE7DC9446CE4" box="[726,1069,416,439]" gridcol="1" gridrow="6" pageId="14" pageNumber="42">1</td>
|
||
<td id="76D25DB3FF9C003FD733FE7DC8E46CE4" box="[1104,1421,416,439]" gridcol="2" gridrow="6" pageId="14" pageNumber="42">1</td>
|
||
</tr>
|
||
<tr id="350334CFFF9C003FD3FCFE17C8E46CB2" box="[159,1421,458,481]" gridrow="7" pageId="14" pageNumber="42">
|
||
<th id="76D25DB3FF9C003FD3FCFE17CFEE6CB2" box="[159,647,458,481]" gridcol="0" gridrow="7" pageId="14" pageNumber="42">ESBO 0</th>
|
||
<td id="76D25DB3FF9C003FD1B5FE17C9446CB2" box="[726,1069,458,481]" gridcol="1" gridrow="7" pageId="14" pageNumber="42">0</td>
|
||
<td id="76D25DB3FF9C003FD733FE17C8E46CB2" box="[1104,1421,458,481]" gridcol="2" gridrow="7" pageId="14" pageNumber="42">0</td>
|
||
</tr>
|
||
<tr id="350334CFFF9C003FD3FCFE28C8E46F5F" box="[159,1421,501,524]" gridrow="8" pageId="14" pageNumber="42">
|
||
<th id="76D25DB3FF9C003FD3FCFE28CFEE6F5F" box="[159,647,501,524]" gridcol="0" gridrow="8" pageId="14" pageNumber="42">GP 4</th>
|
||
<td id="76D25DB3FF9C003FD1B5FE28C9446F5F" box="[726,1069,501,524]" gridcol="1" gridrow="8" pageId="14" pageNumber="42">3</td>
|
||
<td id="76D25DB3FF9C003FD733FE28C8E46F5F" box="[1104,1421,501,524]" gridcol="2" gridrow="8" pageId="14" pageNumber="42">2</td>
|
||
</tr>
|
||
<tr id="350334CFFF9C003FD3FCFDFDC8E46F64" box="[159,1421,544,567]" gridrow="9" pageId="14" pageNumber="42">
|
||
<th id="76D25DB3FF9C003FD3FCFDFDCFEE6F64" box="[159,647,544,567]" gridcol="0" gridrow="9" pageId="14" pageNumber="42">OF 1</th>
|
||
<td id="76D25DB3FF9C003FD1B5FDFDC9446F64" box="[726,1069,544,567]" gridcol="1" gridrow="9" pageId="14" pageNumber="42">1</td>
|
||
<td id="76D25DB3FF9C003FD733FDFDC8E46F64" box="[1104,1421,544,567]" gridcol="2" gridrow="9" pageId="14" pageNumber="42">1</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8B8C368DFF9CFFCED3A6FD7ECE656825" blockId="14.[151,1437,675,2016]" pageId="14" pageNumber="42">
|
||
<emphasis id="B947EA9FFF9CFFCED3A6FD7ECF506FEE" bold="true" box="[197,569,675,701]" pageId="14" pageNumber="42">Comparisons to other species.</emphasis>
|
||
<taxonomicName id="4C334D0EFF9CFFCED122FD79CEE96FEE" box="[577,896,676,701]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="bintangensis">
|
||
<emphasis id="B947EA9FFF9CFFCED122FD79CEE96FEE" box="[577,896,676,701]" italics="true" pageId="14" pageNumber="42">Acanthosaura bintangensis</emphasis>
|
||
</taxonomicName>
|
||
can be differentiated from all other species of
|
||
<taxonomicName id="4C334D0EFF9CFFCED3F4FD17CC286FB0" box="[151,321,714,739]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="B947EA9FFF9CFFCED3F4FD17CC286FB0" box="[151,321,714,739]" italics="true" pageId="14" pageNumber="42">Acanthosaura</emphasis>
|
||
</taxonomicName>
|
||
by having elliptical keeled scales below the orbit of the eye and by having more subdigital lamellae on the fourth finger (23 vs. 16–18 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED1A5FD2CCE546E59" box="[710,829,753,778]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="armata">
|
||
<emphasis id="B947EA9FFF9CFFCED1A5FD2CCE546E59" box="[710,829,753,778]" italics="true" pageId="14" pageNumber="42">A. armata</emphasis>
|
||
</taxonomicName>
|
||
, 16–17 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED0ABFD2CC9446E59" box="[968,1069,753,778]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="capra">
|
||
<emphasis id="B947EA9FFF9CFFCED0ABFD2CC9446E59" box="[968,1069,753,778]" italics="true" pageId="14" pageNumber="42">A. capra</emphasis>
|
||
</taxonomicName>
|
||
, 19–22 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED7DAFD2CC82A6E59" box="[1209,1347,753,778]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="coronata">
|
||
<emphasis id="B947EA9FFF9CFFCED7DAFD2CC82A6E59" box="[1209,1347,753,778]" italics="true" pageId="14" pageNumber="42">A. coronata</emphasis>
|
||
</taxonomicName>
|
||
, 14–18 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED3A1FCC5CC306E62" box="[194,345,792,817]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="crucigera">
|
||
<emphasis id="B947EA9FFF9CFFCED3A1FCC5CC306E62" box="[194,345,792,817]" italics="true" pageId="14" pageNumber="42">A. crucigera</emphasis>
|
||
</taxonomicName>
|
||
, 16–21 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED284FCC5CF026E62" box="[487,619,792,817]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="nataliae">
|
||
<emphasis id="B947EA9FFF9CFFCED284FCC5CF026E62" box="[487,619,792,817]" italics="true" pageId="14" pageNumber="42">A. nataliae</emphasis>
|
||
</taxonomicName>
|
||
, and 17–19 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED04EFCC5CE8C6E62" box="[813,997,792,817]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="lepidogaster">
|
||
<emphasis id="B947EA9FFF9CFFCED04EFCC5CE8C6E62" box="[813,997,792,817]" italics="true" pageId="14" pageNumber="42">A. lepidogaster</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C334D0EFF9CFFCED09DFCC5C82F6E62" box="[1022,1350,792,817]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="bintangensis">
|
||
<emphasis id="B947EA9FFF9CFFCED09DFCC5C82F6E62" box="[1022,1350,792,817]" italics="true" pageId="14" pageNumber="42">Acanthosaura bintangensis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C334D0EFF9CFFCED3B9FCE2CC2D6E0B" box="[218,324,831,856]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="capra">
|
||
<emphasis id="B947EA9FFF9CFFCED3B9FCE2CC2D6E0B" box="[218,324,831,856]" italics="true" pageId="14" pageNumber="42">A. capra</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C334D0EFF9CFFCED2E2FCE2CF616E0B" box="[385,520,831,856]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="nataliae">
|
||
<emphasis id="B947EA9FFF9CFFCED2E2FCE2CF616E0B" box="[385,520,831,856]" italics="true" pageId="14" pageNumber="42">A. nataliae</emphasis>
|
||
</taxonomicName>
|
||
by having a spine on the occiput between tympanum and nuchal crest (vs. lacking), from
|
||
<taxonomicName id="4C334D0EFF9CFFCED227FCB8CCA46E2D" box="[324,461,869,894]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="coronata">
|
||
<emphasis id="B947EA9FFF9CFFCED227FCB8CCA46E2D" box="[324,461,869,894]" italics="true" pageId="14" pageNumber="42">A. coronata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C334D0EFF9CFFCED2B8FCB8CF046E2D" box="[475,621,869,894]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="crucigera">
|
||
<emphasis id="B947EA9FFF9CFFCED2B8FCB8CF046E2D" box="[475,621,869,894]" italics="true" pageId="14" pageNumber="42">A. crucigera</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C334D0EFF9CFFCED1CFFCB8CE376E2D" box="[684,862,869,894]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="lepidogaster">
|
||
<emphasis id="B947EA9FFF9CFFCED1CFFCB8CE376E2D" box="[684,862,869,894]" italics="true" pageId="14" pageNumber="42">A. lepidogaster</emphasis>
|
||
</taxonomicName>
|
||
by having more subdigital lamellae on the fourth toe (26–28 vs. 22–24 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED2DEFC51CF486EF6" box="[445,545,908,933]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="capra">
|
||
<emphasis id="B947EA9FFF9CFFCED2DEFC51CF486EF6" box="[445,545,908,933]" italics="true" pageId="14" pageNumber="42">A. capra</emphasis>
|
||
</taxonomicName>
|
||
, 15–22 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED1CAFC51CE5B6EF6" box="[681,818,908,933]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="coronata">
|
||
<emphasis id="B947EA9FFF9CFFCED1CAFC51CE5B6EF6" box="[681,818,908,933]" italics="true" pageId="14" pageNumber="42">A. coronata</emphasis>
|
||
</taxonomicName>
|
||
, 19–24 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED0DAFC51C9256EF6" box="[953,1100,908,933]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="crucigera">
|
||
<emphasis id="B947EA9FFF9CFFCED0DAFC51C9256EF6" box="[953,1100,908,933]" italics="true" pageId="14" pageNumber="42">A. crucigera</emphasis>
|
||
</taxonomicName>
|
||
, and 23 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED7BBFC51C8E36EF6" box="[1240,1418,908,933]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="lepidogaster">
|
||
<emphasis id="B947EA9FFF9CFFCED7BBFC51C8E36EF6" box="[1240,1418,908,933]" italics="true" pageId="14" pageNumber="42">A. lepidogaster</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C334D0EFF9CFFCED3F4FC6FCC846E98" box="[151,493,946,971]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="bintangensis">
|
||
<emphasis id="B947EA9FFF9CFFCED3F4FC6FCC846E98" box="[151,493,946,971]" italics="true" pageId="14" pageNumber="42">Acanthosaura bintangensis</emphasis>
|
||
</taxonomicName>
|
||
can be differentiated from all other species of
|
||
<taxonomicName id="4C334D0EFF9CFFCED721FC6FC9986E98" box="[1090,1265,946,971]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="B947EA9FFF9CFFCED721FC6FC9986E98" box="[1090,1265,946,971]" italics="true" pageId="14" pageNumber="42">Acanthosaura</emphasis>
|
||
</taxonomicName>
|
||
except for
|
||
<taxonomicName id="4C334D0EFF9CFFCED6E1FC6FCC256EA1" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="titiwangsaensis" status="sp. nov.">
|
||
<emphasis id="B947EA9FFF9CFFCED6E1FC6FCC256EA1" italics="true" pageId="14" pageNumber="42">A. titiwangsaensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B947EA9FFF9CFFCED230FC05CCD96EA1" bold="true" box="[339,432,984,1010]" pageId="14" pageNumber="42">
|
||
<taxonomicNameLabel id="A27457E4FF9CFFCED230FC05CCD96EA1" box="[339,432,984,1010]" pageId="14" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by having more canthus rostralis-supracilliary scales (14–15 vs. 11–13 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED643FC04C8FD6EA1" box="[1312,1428,985,1010]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="armata">
|
||
<emphasis id="B947EA9FFF9CFFCED643FC04C8FD6EA1" box="[1312,1428,985,1010]" italics="true" pageId="14" pageNumber="42">A. armata</emphasis>
|
||
</taxonomicName>
|
||
, 13 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED385FBDDCC19694A" box="[230,368,1024,1049]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="coronata">
|
||
<emphasis id="B947EA9FFF9CFFCED385FBDDCC19694A" box="[230,368,1024,1049]" italics="true" pageId="14" pageNumber="42">A. coronata</emphasis>
|
||
</taxonomicName>
|
||
, 12–13 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED298FBDDCFE7694A" box="[507,654,1024,1049]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="crucigera">
|
||
<emphasis id="B947EA9FFF9CFFCED298FBDDCFE7694A" box="[507,654,1024,1049]" italics="true" pageId="14" pageNumber="42">A. crucigera</emphasis>
|
||
</taxonomicName>
|
||
, 13 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED18FFBDDCE07694A" box="[748,878,1024,1049]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="nataliae">
|
||
<emphasis id="B947EA9FFF9CFFCED18FFBDDCE07694A" box="[748,878,1024,1049]" italics="true" pageId="14" pageNumber="42">A. nataliae</emphasis>
|
||
</taxonomicName>
|
||
, and 10–12 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED748FBDDC9B7694A" box="[1067,1246,1024,1049]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="lepidogaster">
|
||
<emphasis id="B947EA9FFF9CFFCED748FBDDC9B7694A" box="[1067,1246,1024,1049]" italics="true" pageId="14" pageNumber="42">A. lepidogaster</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C334D0EFF9CFFCED794FBDDCC426913" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="bintangensis">
|
||
<emphasis id="B947EA9FFF9CFFCED794FBDDCC426913" italics="true" pageId="14" pageNumber="42">Acanthosaura bintangensis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C334D0EFF9CFFCED2A7FBFACF516913" box="[452,568,1063,1088]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="armata">
|
||
<emphasis id="B947EA9FFF9CFFCED2A7FBFACF516913" box="[452,568,1063,1088]" italics="true" pageId="14" pageNumber="42">A. armata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C334D0EFF9CFFCED125FBFACFB96913" box="[582,720,1063,1088]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="coronata">
|
||
<emphasis id="B947EA9FFF9CFFCED125FBFACFB96913" box="[582,720,1063,1088]" italics="true" pageId="14" pageNumber="42">A. coronata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C334D0EFF9CFFCED1BEFBFACE066913" box="[733,879,1063,1088]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="crucigera">
|
||
<emphasis id="B947EA9FFF9CFFCED1BEFBFACE066913" box="[733,879,1063,1088]" italics="true" pageId="14" pageNumber="42">A. crucigera</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C334D0EFF9CFFCED0CDFBFAC9096913" box="[942,1120,1063,1088]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="lepidogaster">
|
||
<emphasis id="B947EA9FFF9CFFCED0CDFBFAC9096913" box="[942,1120,1063,1088]" italics="true" pageId="14" pageNumber="42">A. lepidogaster</emphasis>
|
||
</taxonomicName>
|
||
by lacking a light patch on the knee. It can be differentiated from
|
||
<taxonomicName id="4C334D0EFF9CFFCED1EDFB90CE4F6935" box="[654,806,1101,1126]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="coronata">
|
||
<emphasis id="B947EA9FFF9CFFCED1EDFB90CE4F6935" box="[654,806,1101,1126]" italics="true" pageId="14" pageNumber="42">A. coronata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C334D0EFF9CFFCED005FB90CE9C6935" box="[870,1013,1101,1126]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="nataliae">
|
||
<emphasis id="B947EA9FFF9CFFCED005FB90CE9C6935" box="[870,1013,1101,1126]" italics="true" pageId="14" pageNumber="42">A. nataliae</emphasis>
|
||
</taxonomicName>
|
||
by having a black nuchal collar.
|
||
<taxonomicName id="4C334D0EFF9CFFCED3F4FBA9CC8D69DE" box="[151,484,1140,1165]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="bintangensis">
|
||
<emphasis id="B947EA9FFF9CFFCED3F4FBA9CC8D69DE" box="[151,484,1140,1165]" italics="true" pageId="14" pageNumber="42">Acanthosaura bintangensis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C334D0EFF9CFFCED1EBFBA9CE6B69DE" box="[648,770,1140,1165]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="armata">
|
||
<emphasis id="B947EA9FFF9CFFCED1EBFBA9CE6B69DE" box="[648,770,1140,1165]" italics="true" pageId="14" pageNumber="42">A. armata</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C334D0EFF9CFFCED024FBA9CEA769DE" box="[839,974,1140,1165]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="nataliae">
|
||
<emphasis id="B947EA9FFF9CFFCED024FBA9CEA769DE" box="[839,974,1140,1165]" italics="true" pageId="14" pageNumber="42">A. nataliae</emphasis>
|
||
</taxonomicName>
|
||
, by having a higher number of scales between the diastema (11–15 vs. 1–8 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED114FB46CF8569E7" box="[631,748,1179,1204]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="armata">
|
||
<emphasis id="B947EA9FFF9CFFCED114FB46CF8569E7" box="[631,748,1179,1204]" italics="true" pageId="14" pageNumber="42">A. armata</emphasis>
|
||
</taxonomicName>
|
||
and 10 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED011FB46CE9B69E7" box="[882,1010,1179,1204]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="nataliae">
|
||
<emphasis id="B947EA9FFF9CFFCED011FB46CE9B69E7" box="[882,1010,1179,1204]" italics="true" pageId="14" pageNumber="42">A. nataliae</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C334D0EFF9CFFCED769FB46C82069E7" box="[1034,1353,1179,1204]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="bintangensis">
|
||
<emphasis id="B947EA9FFF9CFFCED769FB46C82069E7" box="[1034,1353,1179,1204]" italics="true" pageId="14" pageNumber="42">Acanthosaura bintangensis</emphasis>
|
||
</taxonomicName>
|
||
can be separated from
|
||
<taxonomicName id="4C334D0EFF9CFFCED22FFB1CCCA86989" box="[332,449,1217,1242]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="armata">
|
||
<emphasis id="B947EA9FFF9CFFCED22FFB1CCCA86989" box="[332,449,1217,1242]" italics="true" pageId="14" pageNumber="42">A. armata</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C334D0EFF9CFFCED162FB1CCFE86989" box="[513,641,1217,1242]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="nataliae">
|
||
<emphasis id="B947EA9FFF9CFFCED162FB1CCFE86989" box="[513,641,1217,1242]" italics="true" pageId="14" pageNumber="42">A. nataliae</emphasis>
|
||
</taxonomicName>
|
||
by having fewer ventral scales (47–57 vs. 59–68 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED78DFB1CC80A6989" box="[1262,1379,1217,1242]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="armata">
|
||
<emphasis id="B947EA9FFF9CFFCED78DFB1CC80A6989" box="[1262,1379,1217,1242]" italics="true" pageId="14" pageNumber="42">A. armata</emphasis>
|
||
</taxonomicName>
|
||
, and 64–71 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED274FB35CCF76852" box="[279,414,1256,1281]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="nataliae">
|
||
<emphasis id="B947EA9FFF9CFFCED274FB35CCF76852" box="[279,414,1256,1281]" italics="true" pageId="14" pageNumber="42">A. nataliae</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C334D0EFF9CFFCED2DBFB35CE6C6852" box="[440,773,1256,1281]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="bintangensis">
|
||
<emphasis id="B947EA9FFF9CFFCED2DBFB35CE6C6852" box="[440,773,1256,1281]" italics="true" pageId="14" pageNumber="42">Acanthosaura bintangensis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C334D0EFF9CFFCED0CAFB35C9596852" box="[937,1072,1256,1281]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="nataliae">
|
||
<emphasis id="B947EA9FFF9CFFCED0CAFB35C9596852" box="[937,1072,1256,1281]" italics="true" pageId="14" pageNumber="42">A. nataliae</emphasis>
|
||
</taxonomicName>
|
||
by having a lower number of scales between the seventh supralabial and the sixth canthal (9–14 vs. 16 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED733FAD2C9BE687B" box="[1104,1239,1295,1320]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="nataliae">
|
||
<emphasis id="B947EA9FFF9CFFCED733FAD2C9BE687B" box="[1104,1239,1295,1320]" italics="true" pageId="14" pageNumber="42">A. nataliae</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C334D0EFF9CFFCED791FAD2CC47681D" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="bintangensis">
|
||
<emphasis id="B947EA9FFF9CFFCED791FAD2CC47681D" italics="true" pageId="14" pageNumber="42">Acanthosaura bintangensis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C334D0EFF9CFFCED2AEFAE8CFED681D" box="[461,644,1333,1358]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="lepidogaster">
|
||
<emphasis id="B947EA9FFF9CFFCED2AEFAE8CFED681D" box="[461,644,1333,1358]" italics="true" pageId="14" pageNumber="42">A. lepidogaster</emphasis>
|
||
</taxonomicName>
|
||
by having a smaller diastema length divided by snout vent length (4.3–6.7% vs. 7.8–8.2% for
|
||
<taxonomicName id="4C334D0EFF9CFFCED281FA81CFFC6826" box="[482,661,1372,1397]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="lepidogaster">
|
||
<emphasis id="B947EA9FFF9CFFCED281FA81CFFC6826" box="[482,661,1372,1397]" italics="true" pageId="14" pageNumber="42">A. lepidogaster</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<tableCitation id="C6B10336FF9CFFCED1C7FA81CF936825" box="[676,762,1372,1398]" captionStart="TABLE 1" captionStartId="3.[151,239,847,871]" captionText="TABLE 1. All measurements (in mm) for all currently recognized species of Acanthosaura. PS – postorbital spine length, measured from the base to the tip of the spine; NS – the maximum length of the largest spine measured from the base to the tip; NSL – longest nuchal scale, measured at the base; DS – maximum length of longest spine on the body measured from the base to the tip; DSL – longest dorsal scale, measured at the base below the dorsal crest; WNC – maximum width of the spines in the nuchal crest, measured at the base; WDS – maximum width of the largest dorsal scale below the dorsal crest, measured at the base; DIAS – length of the diastema, measured from the posterior end of the nuchal crest to the anterior end of the nuchal crest; FOREL – forelimb length, measured from axilla to the proximal edge of the palmar region; HINDL – hindlimb length, measured from groin to the proximal edge plantar region; RS – number of scales bordering the rostral scale; NCS – number of scales between the fifth canthals; NSCSL – number of scales from the fifth canthal to the fifth supralabial; NR – number of scales between the nasals and the rostrals; NSSLC – number of scales between the seventh supralabial and the sixth canthal; LKP – presence or absence of light knee patch; GP – size of gular pouch scored as absent, small, medium or large; OF – presence or absence of oblique fold anterior to the fore limb insertions. For additional character abbreviations see materials and methods. / Indicates missing data. All data were taken from specimens or from the literature." pageId="14" pageNumber="42">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8C368DFF9CFFCFD3A6FA5ECF1C6CC9" blockId="14.[151,1437,675,2016]" lastBlockId="15.[151,1437,152,719]" lastPageId="15" lastPageNumber="43" pageId="14" pageNumber="42">
|
||
<taxonomicName id="4C334D0EFF9CFFCED3A6FA5ECF4168CF" box="[197,552,1411,1436]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="titiwangsaensis">
|
||
<emphasis id="B947EA9FFF9CFFCED3A6FA5ECF4168CF" box="[197,552,1411,1436]" italics="true" pageId="14" pageNumber="42">Acanthosaura titiwangsaensis</emphasis>
|
||
</taxonomicName>
|
||
can be differentiated from all other species of
|
||
<taxonomicName id="4C334D0EFF9CFFCED736FA5EC99368CF" box="[1109,1274,1411,1436]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="genus">
|
||
<emphasis id="B947EA9FFF9CFFCED736FA5EC99368CF" box="[1109,1274,1411,1436]" italics="true" pageId="14" pageNumber="42">Acanthosaura</emphasis>
|
||
</taxonomicName>
|
||
except for
|
||
<taxonomicName id="4C334D0EFF9CFFCED6E0FA5ECC5B6891" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="bintangensis">
|
||
<emphasis id="B947EA9FFF9CFFCED6E0FA5ECC5B6891" italics="true" pageId="14" pageNumber="42">A. bintangensis</emphasis>
|
||
</taxonomicName>
|
||
by having a higher number of canthus rostralis-supracilliary scales (14–15 vs. 11–13 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED6E1FA74CD9A68BA" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="armata">
|
||
<emphasis id="B947EA9FFF9CFFCED6E1FA74CD9A68BA" italics="true" pageId="14" pageNumber="42">A. armata</emphasis>
|
||
</taxonomicName>
|
||
, 13 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED23FFA0DCC9B68BA" box="[348,498,1488,1513]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="coronata">
|
||
<emphasis id="B947EA9FFF9CFFCED23FFA0DCC9B68BA" box="[348,498,1488,1513]" italics="true" pageId="14" pageNumber="42">A. coronata</emphasis>
|
||
</taxonomicName>
|
||
, 12–13 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED1E8FA0DCE4568BA" box="[651,812,1488,1513]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="crucigera">
|
||
<emphasis id="B947EA9FFF9CFFCED1E8FA0DCE4568BA" box="[651,812,1488,1513]" italics="true" pageId="14" pageNumber="42">A. crucigera</emphasis>
|
||
</taxonomicName>
|
||
, 13
|
||
<taxonomicName id="4C334D0EFF9CFFCED004FA0DCE9C68BA" box="[871,1013,1488,1513]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="nataliae">
|
||
<emphasis id="B947EA9FFF9CFFCED004FA0DCE9C68BA" box="[871,1013,1488,1513]" italics="true" pageId="14" pageNumber="42">A. nataliae</emphasis>
|
||
</taxonomicName>
|
||
, and 10–12 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED7A7FA0DC8E068BA" box="[1220,1417,1488,1513]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="lepidogaster">
|
||
<emphasis id="B947EA9FFF9CFFCED7A7FA0DC8E068BA" box="[1220,1417,1488,1513]" italics="true" pageId="14" pageNumber="42">A. lepidogaster</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C334D0EFF9CFFCED3F4FA2ACF696B43" box="[151,512,1527,1552]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="titiwangsaensis">
|
||
<emphasis id="B947EA9FFF9CFFCED3F4FA2ACF696B43" box="[151,512,1527,1552]" italics="true" pageId="14" pageNumber="42">Acanthosaura titiwangsaensis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C334D0EFF9CFFCED1C3FA2ACE4C6B43" box="[672,805,1527,1552]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="nataliae">
|
||
<emphasis id="B947EA9FFF9CFFCED1C3FA2ACE4C6B43" box="[672,805,1527,1552]" italics="true" pageId="14" pageNumber="42">A. nataliae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C334D0EFF9CFFCED003FA2ACEA16B43" box="[864,968,1527,1552]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="capra">
|
||
<emphasis id="B947EA9FFF9CFFCED003FA2ACEA16B43" box="[864,968,1527,1552]" italics="true" pageId="14" pageNumber="42">A. capra</emphasis>
|
||
</taxonomicName>
|
||
by having an occipital spine. It can be differentiated from
|
||
<taxonomicName id="4C334D0EFF9CFFCED21CF9C0CC9C6B65" box="[383,501,1565,1590]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="armata">
|
||
<emphasis id="B947EA9FFF9CFFCED21CF9C0CC9C6B65" box="[383,501,1565,1590]" italics="true" pageId="14" pageNumber="42">A. armata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C334D0EFF9CFFCED166F9C0CFF96B65" box="[517,656,1565,1590]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="coronata">
|
||
<emphasis id="B947EA9FFF9CFFCED166F9C0CFF96B65" box="[517,656,1565,1590]" italics="true" pageId="14" pageNumber="42">A. coronata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C334D0EFF9CFFCED1FCF9C0CE5A6B65" box="[671,819,1565,1590]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="crucigera">
|
||
<emphasis id="B947EA9FFF9CFFCED1FCF9C0CE5A6B65" box="[671,819,1565,1590]" italics="true" pageId="14" pageNumber="42">A. crucigera</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C334D0EFF9CFFCED016F9C0C9436B65" box="[885,1066,1565,1590]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="lepidogaster">
|
||
<emphasis id="B947EA9FFF9CFFCED016F9C0C9436B65" box="[885,1066,1565,1590]" italics="true" pageId="14" pageNumber="42">A. lepidogaster</emphasis>
|
||
</taxonomicName>
|
||
by lacking a light patch on the knee.
|
||
<taxonomicName id="4C334D0EFF9CFFCED3BCF999CF206B0E" box="[223,585,1604,1629]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="titiwangsaensis">
|
||
<emphasis id="B947EA9FFF9CFFCED3BCF999CF206B0E" box="[223,585,1604,1629]" italics="true" pageId="14" pageNumber="42">Acanthosaura titiwangsaensis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C334D0EFF9CFFCED189F999CE0A6B0E" box="[746,867,1604,1629]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="armata">
|
||
<emphasis id="B947EA9FFF9CFFCED189F999CE0A6B0E" box="[746,867,1604,1629]" italics="true" pageId="14" pageNumber="42">A. armata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C334D0EFF9CFFCED010F999CEB26B0E" box="[883,987,1604,1629]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="capra">
|
||
<emphasis id="B947EA9FFF9CFFCED010F999CEB26B0E" box="[883,987,1604,1629]" italics="true" pageId="14" pageNumber="42">A. capra</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C334D0EFF9CFFCED088F999C9EB6B0E" box="[1003,1154,1604,1629]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="crucigera">
|
||
<emphasis id="B947EA9FFF9CFFCED088F999C9EB6B0E" box="[1003,1154,1604,1629]" italics="true" pageId="14" pageNumber="42">A. crucigera</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C334D0EFF9CFFCED7DEF999C81C6B0E" box="[1213,1397,1604,1629]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="lepidogaster">
|
||
<emphasis id="B947EA9FFF9CFFCED7DEF999C81C6B0E" box="[1213,1397,1604,1629]" italics="true" pageId="14" pageNumber="42">A. lepidogaster</emphasis>
|
||
</taxonomicName>
|
||
by having a higher number of subdigital lamellae on the fourth finger (20–21 vs. 16–18 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED7C6F9B6C8706BD7" box="[1189,1305,1643,1668]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="armata">
|
||
<emphasis id="B947EA9FFF9CFFCED7C6F9B6C8706BD7" box="[1189,1305,1643,1668]" italics="true" pageId="14" pageNumber="42">A. armata</emphasis>
|
||
</taxonomicName>
|
||
, 16–17 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED3F4F94CCD946BF9" box="[151,253,1681,1706]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="capra">
|
||
<emphasis id="B947EA9FFF9CFFCED3F4F94CCD946BF9" box="[151,253,1681,1706]" italics="true" pageId="14" pageNumber="42">A. capra</emphasis>
|
||
</taxonomicName>
|
||
, 14–18
|
||
<taxonomicName id="4C334D0EFF9CFFCED23CF94CCC9B6BF9" box="[351,498,1681,1706]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="crucigera">
|
||
<emphasis id="B947EA9FFF9CFFCED23CF94CCC9B6BF9" box="[351,498,1681,1706]" italics="true" pageId="14" pageNumber="42">A. crucigera</emphasis>
|
||
</taxonomicName>
|
||
, and 17–19 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED1D1F94CCE0E6BF9" box="[690,871,1681,1706]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="lepidogaster">
|
||
<emphasis id="B947EA9FFF9CFFCED1D1F94CCE0E6BF9" box="[690,871,1681,1706]" italics="true" pageId="14" pageNumber="42">A. lepidogaster</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C334D0EFF9CFFCED0E3F94CC98D6BF9" box="[896,1252,1681,1706]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="titiwangsaensis">
|
||
<emphasis id="B947EA9FFF9CFFCED0E3F94CC98D6BF9" box="[896,1252,1681,1706]" italics="true" pageId="14" pageNumber="42">Acanthosaura titiwangsaensis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C334D0EFF9CFFCED6E0F94CCD866B82" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="armata">
|
||
<emphasis id="B947EA9FFF9CFFCED6E0F94CCD866B82" italics="true" pageId="14" pageNumber="42">A. armata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C334D0EFF9CFFCED263F965CCF86B82" box="[256,401,1720,1745]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="coronata">
|
||
<emphasis id="B947EA9FFF9CFFCED263F965CCF86B82" box="[256,401,1720,1745]" italics="true" pageId="14" pageNumber="42">A. coronata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C334D0EFF9CFFCED2C2F965CF556B82" box="[417,572,1720,1745]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="crucigera">
|
||
<emphasis id="B947EA9FFF9CFFCED2C2F965CF556B82" box="[417,572,1720,1745]" italics="true" pageId="14" pageNumber="42">A. crucigera</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C334D0EFF9CFFCED12FF965CFBC6B82" box="[588,725,1720,1745]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="nataliae">
|
||
<emphasis id="B947EA9FFF9CFFCED12FF965CFBC6B82" box="[588,725,1720,1745]" italics="true" pageId="14" pageNumber="42">A. nataliae</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C334D0EFF9CFFCED079F965CEB16B82" box="[794,984,1720,1745]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="lepidogaster">
|
||
<emphasis id="B947EA9FFF9CFFCED079F965CEB16B82" box="[794,984,1720,1745]" italics="true" pageId="14" pageNumber="42">A. lepidogaster</emphasis>
|
||
</taxonomicName>
|
||
in having a higher number of scales bordering the rostral (9 vs. 5–7 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED155F902CFC26BAB" box="[566,683,1759,1784]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="armata">
|
||
<emphasis id="B947EA9FFF9CFFCED155F902CFC26BAB" box="[566,683,1759,1784]" italics="true" pageId="14" pageNumber="42">A. armata</emphasis>
|
||
</taxonomicName>
|
||
, 8 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED198F902CEEC6BAB" box="[763,901,1759,1784]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="coronata">
|
||
<emphasis id="B947EA9FFF9CFFCED198F902CEEC6BAB" box="[763,901,1759,1784]" italics="true" pageId="14" pageNumber="42">A. coronata</emphasis>
|
||
</taxonomicName>
|
||
, 6–7 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED092F902C9ED6BAB" box="[1009,1156,1759,1784]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="crucigera">
|
||
<emphasis id="B947EA9FFF9CFFCED092F902C9ED6BAB" box="[1009,1156,1759,1784]" italics="true" pageId="14" pageNumber="42">A. crucigera</emphasis>
|
||
</taxonomicName>
|
||
) and by having a lower number of scales bordering the nostrils, (9 vs. 13 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED078F8D8CEF66A4D" box="[795,927,1797,1822]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="nataliae">
|
||
<emphasis id="B947EA9FFF9CFFCED078F8D8CEF66A4D" box="[795,927,1797,1822]" italics="true" pageId="14" pageNumber="42">A. nataliae</emphasis>
|
||
</taxonomicName>
|
||
, and 10–12 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED702F8D8C8736A4D" box="[1121,1306,1797,1822]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="lepidogaster">
|
||
<emphasis id="B947EA9FFF9CFFCED702F8D8C8736A4D" box="[1121,1306,1797,1822]" italics="true" pageId="14" pageNumber="42">A. lepidogaster</emphasis>
|
||
</taxonomicName>
|
||
). It can be differentiated from
|
||
<taxonomicName id="4C334D0EFF9CFFCED219F8F1CC866A16" box="[378,495,1836,1861]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="armata">
|
||
<emphasis id="B947EA9FFF9CFFCED219F8F1CC866A16" box="[378,495,1836,1861]" italics="true" pageId="14" pageNumber="42">A. armata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C334D0EFF9CFFCED145F8F1CFCF6A16" box="[550,678,1836,1861]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="nataliae">
|
||
<emphasis id="B947EA9FFF9CFFCED145F8F1CFCF6A16" box="[550,678,1836,1861]" italics="true" pageId="14" pageNumber="42">A. nataliae</emphasis>
|
||
</taxonomicName>
|
||
on having a lower number of ventral scales (47–57 vs. 59–68 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED3F4F88ECC626A3F" box="[151,267,1875,1900]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="armata">
|
||
<emphasis id="B947EA9FFF9CFFCED3F4F88ECC626A3F" box="[151,267,1875,1900]" italics="true" pageId="14" pageNumber="42">A. armata</emphasis>
|
||
</taxonomicName>
|
||
, and 64–71 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED2A5F88ECF2E6A3F" box="[454,583,1875,1900]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="nataliae">
|
||
<emphasis id="B947EA9FFF9CFFCED2A5F88ECF2E6A3F" box="[454,583,1875,1900]" italics="true" pageId="14" pageNumber="42">A. nataliae</emphasis>
|
||
</taxonomicName>
|
||
). It can be differentiated from
|
||
<taxonomicName id="4C334D0EFF9CFFCED0D3F88EC9536A3F" box="[944,1082,1875,1900]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="coronata">
|
||
<emphasis id="B947EA9FFF9CFFCED0D3F88EC9536A3F" box="[944,1082,1875,1900]" italics="true" pageId="14" pageNumber="42">A. coronata</emphasis>
|
||
</taxonomicName>
|
||
by having a higher number of subdigital lamellae on the fourth toe (23–27 vs. 15–22 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED019F8A4C9416AC0" authority="It" authorityName="It" box="[890,1064,1913,1939]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="coronata">
|
||
<emphasis id="B947EA9FFF9CFFCED019F8A4C9656AC1" box="[890,1036,1913,1938]" italics="true" pageId="14" pageNumber="42">A. coronata</emphasis>
|
||
It
|
||
</taxonomicName>
|
||
can be differentiated from
|
||
<taxonomicName id="4C334D0EFF9CFFCED6E1F8A4CC6B6AEA" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="coronata">
|
||
<emphasis id="B947EA9FFF9CFFCED6E1F8A4CC6B6AEA" italics="true" pageId="14" pageNumber="42">A. coronata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B947EA9FFF9CFFCED272F87DCCFD6AEA" box="[273,404,1952,1977]" italics="true" pageId="14" pageNumber="42">
|
||
A,
|
||
<taxonomicName id="4C334D0EFF9CFFCED250F87DCCFD6AEA" box="[307,404,1952,1977]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="nataliae">nataliae</taxonomicName>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="4C334D0EFF9CFFCED2BBF87DCFE66AEA" box="[472,655,1952,1977]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="lepidogaster">
|
||
<emphasis id="B947EA9FFF9CFFCED2BBF87DCFE66AEA" box="[472,655,1952,1977]" italics="true" pageId="14" pageNumber="42">A. lepidogaster</emphasis>
|
||
</taxonomicName>
|
||
in having a higher number of scales from the fifth canthal to the fifth supra labial (9–11 vs. 6–7 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED15EF81ACFA06AB3" box="[573,713,1991,2016]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="coronata">
|
||
<emphasis id="B947EA9FFF9CFFCED15EF81ACFA06AB3" box="[573,713,1991,2016]" italics="true" pageId="14" pageNumber="42">A. coronata</emphasis>
|
||
</taxonomicName>
|
||
, 8 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED078F81ACEF66AB3" box="[795,927,1991,2016]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="nataliae">
|
||
<emphasis id="B947EA9FFF9CFFCED078F81ACEF66AB3" box="[795,927,1991,2016]" italics="true" pageId="14" pageNumber="42">A. nataliae</emphasis>
|
||
</taxonomicName>
|
||
, and 8 for
|
||
<taxonomicName id="4C334D0EFF9CFFCED747F81AC9B26AB3" box="[1060,1243,1991,2016]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="lepidogaster">
|
||
<emphasis id="B947EA9FFF9CFFCED747F81AC9B26AB3" box="[1060,1243,1991,2016]" italics="true" pageId="14" pageNumber="42">A. lepidogaster</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C334D0EFF9CFFCFD796F81ACC276DE2" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" lastPageId="15" lastPageNumber="43" order="Squamata" pageId="14" pageNumber="42" phylum="Chordata" rank="species" species="titiwangsaensis">
|
||
<emphasis id="B947EA9FFF9CFFCFD796F81ACC276DE2" italics="true" lastPageId="15" lastPageNumber="43" pageId="14" pageNumber="42">Acanthosaura titiwangsaensis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C334D0EFF9DFFCFD28FFF45CF116DE2" box="[492,632,152,177]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="43" phylum="Chordata" rank="species" species="coronata">
|
||
<emphasis id="B947EA9FFF9DFFCFD28FFF45CF116DE2" box="[492,632,152,177]" italics="true" pageId="15" pageNumber="43">A. coronata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C334D0EFF9DFFCFD1EBFF45CE756DE2" box="[648,796,152,177]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="43" phylum="Chordata" rank="species" species="crucigera">
|
||
<emphasis id="B947EA9FFF9DFFCFD1EBFF45CE756DE2" box="[648,796,152,177]" italics="true" pageId="15" pageNumber="43">A. crucigera</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C334D0EFF9DFFCFD03CFF45CE8B6DE2" box="[863,994,152,177]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="43" phylum="Chordata" rank="species" species="nataliae">
|
||
<emphasis id="B947EA9FFF9DFFCFD03CFF45CE8B6DE2" box="[863,994,152,177]" italics="true" pageId="15" pageNumber="43">A. nataliae</emphasis>
|
||
</taxonomicName>
|
||
by having a higher number of scales bordering the mental (5 vs. 4 for
|
||
<taxonomicName id="4C334D0EFF9DFFCFD15AFF62CFA56D8B" box="[569,716,191,216]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="43" phylum="Chordata" rank="species" species="coronata">
|
||
<emphasis id="B947EA9FFF9DFFCFD15AFF62CFA56D8B" box="[569,716,191,216]" italics="true" pageId="15" pageNumber="43">A. coronata</emphasis>
|
||
</taxonomicName>
|
||
, 4 for
|
||
<taxonomicName id="4C334D0EFF9DFFCFD041FF62CED66D8B" box="[802,959,191,216]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="43" phylum="Chordata" rank="species" species="crucigera">
|
||
<emphasis id="B947EA9FFF9DFFCFD041FF62CED66D8B" box="[802,959,191,216]" italics="true" pageId="15" pageNumber="43">A. crucigera</emphasis>
|
||
</taxonomicName>
|
||
, and 4 for
|
||
<taxonomicName id="4C334D0EFF9DFFCFD729FF62C9BA6D8B" box="[1098,1235,191,216]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="43" phylum="Chordata" rank="species" species="nataliae">
|
||
<emphasis id="B947EA9FFF9DFFCFD729FF62C9BA6D8B" box="[1098,1235,191,216]" italics="true" pageId="15" pageNumber="43">A. nataliae</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C334D0EFF9DFFCFD78CFF62CC256DAD" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="43" phylum="Chordata" rank="species" species="titiwangsaensis">
|
||
<emphasis id="B947EA9FFF9DFFCFD78CFF62CC256DAD" italics="true" pageId="15" pageNumber="43">Acanthosaura titiwangsaensis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C334D0EFF9DFFCFD285FF38CFF36DAD" box="[486,666,229,254]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="43" phylum="Chordata" rank="species" species="lepidogaster">
|
||
<emphasis id="B947EA9FFF9DFFCFD285FF38CFF36DAD" box="[486,666,229,254]" italics="true" pageId="15" pageNumber="43">A. lepidogaster</emphasis>
|
||
</taxonomicName>
|
||
by having a smaller diastema length divided by snout vent length (4.9–6.7% vs. 7.8–8.2% for
|
||
<taxonomicName id="4C334D0EFF9DFFCFD280FED1CFFF6C76" box="[483,662,268,293]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="43" phylum="Chordata" rank="species" species="lepidogaster">
|
||
<emphasis id="B947EA9FFF9DFFCFD280FED1CFFF6C76" box="[483,662,268,293]" italics="true" pageId="15" pageNumber="43">A. lepidogaster</emphasis>
|
||
</taxonomicName>
|
||
). This species differs from
|
||
<taxonomicName id="4C334D0EFF9DFFCFD0BFFED1C90F6C76" box="[988,1126,268,293]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="43" phylum="Chordata" rank="species" species="coronata">
|
||
<emphasis id="B947EA9FFF9DFFCFD0BFFED1C90F6C76" box="[988,1126,268,293]" italics="true" pageId="15" pageNumber="43">A. coronata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C334D0EFF9DFFCFD7FCFED1C8766C76" box="[1183,1311,268,293]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="43" phylum="Chordata" rank="species" species="nataliae">
|
||
<emphasis id="B947EA9FFF9DFFCFD7FCFED1C8766C76" box="[1183,1311,268,293]" italics="true" pageId="15" pageNumber="43">A. nataliae</emphasis>
|
||
</taxonomicName>
|
||
by having a nuchal diamond present on the nape.
|
||
<taxonomicName id="4C334D0EFF9DFFCFD114FEEFCE8F6C18" box="[631,998,306,331]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="43" phylum="Chordata" rank="species" species="titiwangsaensis">
|
||
<emphasis id="B947EA9FFF9DFFCFD114FEEFCE8F6C18" box="[631,998,306,331]" italics="true" pageId="15" pageNumber="43">Acanthosaura titiwangsaensis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C334D0EFF9DFFCFD7E9FEEFC86C6C18" box="[1162,1285,306,331]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="43" phylum="Chordata" rank="species" species="armata">
|
||
<emphasis id="B947EA9FFF9DFFCFD7E9FEEFC86C6C18" box="[1162,1285,306,331]" italics="true" pageId="15" pageNumber="43">A. armata</emphasis>
|
||
</taxonomicName>
|
||
by having a higher number of scales between the diastema (10–13 vs. 1–8 for
|
||
<taxonomicName id="4C334D0EFF9DFFCFD0C4FE84C9746C21" box="[935,1053,345,370]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="43" phylum="Chordata" rank="species" species="armata">
|
||
<emphasis id="B947EA9FFF9DFFCFD0C4FE84C9746C21" box="[935,1053,345,370]" italics="true" pageId="15" pageNumber="43">A. armata</emphasis>
|
||
</taxonomicName>
|
||
). It differs from
|
||
<taxonomicName id="4C334D0EFF9DFFCFD784FE84C8F56C21" box="[1255,1436,345,370]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="43" phylum="Chordata" rank="species" species="lepidogaster">
|
||
<emphasis id="B947EA9FFF9DFFCFD784FE84C8F56C21" box="[1255,1436,345,370]" italics="true" pageId="15" pageNumber="43">A. lepidogaster</emphasis>
|
||
</taxonomicName>
|
||
by having a larger gular pouch (
|
||
<tableCitation id="C6B10336FF9DFFCFD16FFE5DCF0D6CC9" box="[524,612,384,410]" captionStart="TABLE 1" captionStartId="3.[151,239,847,871]" captionText="TABLE 1. All measurements (in mm) for all currently recognized species of Acanthosaura. PS – postorbital spine length, measured from the base to the tip of the spine; NS – the maximum length of the largest spine measured from the base to the tip; NSL – longest nuchal scale, measured at the base; DS – maximum length of longest spine on the body measured from the base to the tip; DSL – longest dorsal scale, measured at the base below the dorsal crest; WNC – maximum width of the spines in the nuchal crest, measured at the base; WDS – maximum width of the largest dorsal scale below the dorsal crest, measured at the base; DIAS – length of the diastema, measured from the posterior end of the nuchal crest to the anterior end of the nuchal crest; FOREL – forelimb length, measured from axilla to the proximal edge of the palmar region; HINDL – hindlimb length, measured from groin to the proximal edge plantar region; RS – number of scales bordering the rostral scale; NCS – number of scales between the fifth canthals; NSCSL – number of scales from the fifth canthal to the fifth supralabial; NR – number of scales between the nasals and the rostrals; NSSLC – number of scales between the seventh supralabial and the sixth canthal; LKP – presence or absence of light knee patch; GP – size of gular pouch scored as absent, small, medium or large; OF – presence or absence of oblique fold anterior to the fore limb insertions. For additional character abbreviations see materials and methods. / Indicates missing data. All data were taken from specimens or from the literature." pageId="15" pageNumber="43">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8C368DFF9DFFCFD3A6FE7ACC636F9C" blockId="15.[151,1437,152,719]" pageId="15" pageNumber="43">
|
||
<taxonomicName id="4C334D0EFF9DFFCFD3A6FE7ACF4C6C93" box="[197,549,423,448]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="43" phylum="Chordata" rank="species" species="bintangensis">
|
||
<emphasis id="B947EA9FFF9DFFCFD3A6FE7ACF4C6C93" box="[197,549,423,448]" italics="true" pageId="15" pageNumber="43">Acanthosaura bintangensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C334D0EFF9DFFCFD105FE7ACE3A6C93" box="[614,851,423,448]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="43" phylum="Chordata" rank="species" species="titiwangsaensis">
|
||
<emphasis id="B947EA9FFF9DFFCFD105FE7ACE3A6C93" box="[614,851,423,448]" italics="true" pageId="15" pageNumber="43">A. titiwangsaensis</emphasis>
|
||
</taxonomicName>
|
||
most closely resemble each other but can be differentiated by their number of subdigital lamellae under the fourth finger (
|
||
<quantity id="4CCB9B68FF9DFFCFD745FE10C90D6CB4" box="[1062,1124,461,487]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" pageId="15" pageNumber="43" unit="in" value="23.0">23 in</quantity>
|
||
<taxonomicName id="4C334D0EFF9DFFCFD70FFE10C8486CB5" box="[1132,1313,461,486]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="43" phylum="Chordata" rank="species" species="bintangensis">
|
||
<emphasis id="B947EA9FFF9DFFCFD70FFE10C8486CB5" box="[1132,1313,461,486]" italics="true" pageId="15" pageNumber="43">A. bintangensis</emphasis>
|
||
</taxonomicName>
|
||
vs. 20–21 for
|
||
<taxonomicName id="4C334D0EFF9DFFCFD3A2FE29CCF06F5E" box="[193,409,500,525]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="43" phylum="Chordata" rank="species" species="titiwangsaensis">
|
||
<emphasis id="B947EA9FFF9DFFCFD3A2FE29CCF06F5E" box="[193,409,500,525]" italics="true" pageId="15" pageNumber="43">A. titiwangsaensis</emphasis>
|
||
</taxonomicName>
|
||
); number of scales surrounding the occipital spine (
|
||
<taxonomicName id="4C334D0EFF9DFFCFD765FE29C9D46F5E" box="[1030,1213,500,525]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="43" phylum="Chordata" rank="species" species="bintangensis">
|
||
<emphasis id="B947EA9FFF9DFFCFD765FE29C9D46F5E" box="[1030,1213,500,525]" italics="true" pageId="15" pageNumber="43">A. bintangensis</emphasis>
|
||
</taxonomicName>
|
||
6–7 vs. 4–5 for
|
||
<taxonomicName id="4C334D0EFF9DFFCFD6E0FE29CC276F60" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="43" phylum="Chordata" rank="species" species="titiwangsaensis">
|
||
<emphasis id="B947EA9FFF9DFFCFD6E0FE29CC276F60" italics="true" pageId="15" pageNumber="43">A. titiwangsaensis</emphasis>
|
||
</taxonomicName>
|
||
); number of scales between the fifth canthal to the fifth supralabial (
|
||
<taxonomicName id="4C334D0EFF9DFFCFD7E5FDC7C8576F60" box="[1158,1342,538,563]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="43" phylum="Chordata" rank="species" species="bintangensis">
|
||
<emphasis id="B947EA9FFF9DFFCFD7E5FDC7C8576F60" box="[1158,1342,538,563]" italics="true" pageId="15" pageNumber="43">A. bintangensis</emphasis>
|
||
</taxonomicName>
|
||
7–8 vs. 9–11 for
|
||
<taxonomicName id="4C334D0EFF9DFFCFD262FD9CCCBF6F09" box="[257,470,577,602]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="43" phylum="Chordata" rank="species" species="titiwangsaensis">
|
||
<emphasis id="B947EA9FFF9DFFCFD262FD9CCCBF6F09" box="[257,470,577,602]" italics="true" pageId="15" pageNumber="43">A. titiwangsaensis</emphasis>
|
||
</taxonomicName>
|
||
); by the presence of two elliptical keeled scales below the orbit in
|
||
<taxonomicName id="4C334D0EFF9DFFCFD784FD9CC8F56F09" box="[1255,1436,577,602]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="43" phylum="Chordata" rank="species" species="bintangensis">
|
||
<emphasis id="B947EA9FFF9DFFCFD784FD9CC8F56F09" box="[1255,1436,577,602]" italics="true" pageId="15" pageNumber="43">A. bintangensis</emphasis>
|
||
</taxonomicName>
|
||
and their absence in
|
||
<taxonomicName id="4C334D0EFF9DFFCFD2EDFDB5CF0E6FD2" box="[398,615,616,641]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="43" phylum="Chordata" rank="species" species="titiwangsaensis">
|
||
<emphasis id="B947EA9FFF9DFFCFD2EDFDB5CF0E6FD2" box="[398,615,616,641]" italics="true" pageId="15" pageNumber="43">A. titiwangsaensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13082A08FF9DFFCFD11BFDB5CFA86FD1" box="[632,705,616,642]" captionStart="FIGURE 7" captionStartId="12.[151,255,1879,1903]" captionTargetBox="[151,1434,1371,1854]" captionTargetId="figure@12.[151,1436,1371,1855]" captionTargetPageId="12" captionText="FIGURE 7. Illustration of the two enlarged, keeled, elliptical suborbital scales in Acanthosaura bintangensis (right) and their absence in A. titiwangsaensis (left)." httpUri="https://zenodo.org/record/185829/files/figure.png" pageId="15" pageNumber="43">Fig. 7</figureCitation>
|
||
); by the yellow medial gular stripe in
|
||
<taxonomicName id="4C334D0EFF9DFFCFD7EEFDB5C82D6FD2" box="[1165,1348,616,641]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="43" phylum="Chordata" rank="species" species="bintangensis">
|
||
<emphasis id="B947EA9FFF9DFFCFD7EEFDB5C82D6FD2" box="[1165,1348,616,641]" italics="true" pageId="15" pageNumber="43">A. bintangensis</emphasis>
|
||
</taxonomicName>
|
||
and its absence in
|
||
<taxonomicName id="4C334D0EFF9DFFCFD278FD52CC986FFB" box="[283,497,655,680]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="43" phylum="Chordata" rank="species" species="titiwangsaensis">
|
||
<emphasis id="B947EA9FFF9DFFCFD278FD52CC986FFB" box="[283,497,655,680]" italics="true" pageId="15" pageNumber="43">A. titiwangsaensis</emphasis>
|
||
</taxonomicName>
|
||
; and
|
||
<taxonomicName id="4C334D0EFF9DFFCFD156FD52CF826FFB" box="[565,747,655,680]" class="Reptilia" family="Agamidae" genus="Acanthosaura" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="43" phylum="Chordata" rank="species" species="bintangensis">
|
||
<emphasis id="B947EA9FFF9DFFCFD156FD52CF826FFB" box="[565,747,655,680]" italics="true" pageId="15" pageNumber="43">A. bintangensis</emphasis>
|
||
</taxonomicName>
|
||
having more spots on the dorsum than the latter (
|
||
<tableCitation id="C6B10336FF9DFFCFD659FD53C8FD6FFB" box="[1338,1428,654,680]" captionStart="TABLE 1" captionStartId="3.[151,239,847,871]" captionText="TABLE 1. All measurements (in mm) for all currently recognized species of Acanthosaura. PS – postorbital spine length, measured from the base to the tip of the spine; NS – the maximum length of the largest spine measured from the base to the tip; NSL – longest nuchal scale, measured at the base; DS – maximum length of longest spine on the body measured from the base to the tip; DSL – longest dorsal scale, measured at the base below the dorsal crest; WNC – maximum width of the spines in the nuchal crest, measured at the base; WDS – maximum width of the largest dorsal scale below the dorsal crest, measured at the base; DIAS – length of the diastema, measured from the posterior end of the nuchal crest to the anterior end of the nuchal crest; FOREL – forelimb length, measured from axilla to the proximal edge of the palmar region; HINDL – hindlimb length, measured from groin to the proximal edge plantar region; RS – number of scales bordering the rostral scale; NCS – number of scales between the fifth canthals; NSCSL – number of scales from the fifth canthal to the fifth supralabial; NR – number of scales between the nasals and the rostrals; NSSLC – number of scales between the seventh supralabial and the sixth canthal; LKP – presence or absence of light knee patch; GP – size of gular pouch scored as absent, small, medium or large; OF – presence or absence of oblique fold anterior to the fore limb insertions. For additional character abbreviations see materials and methods. / Indicates missing data. All data were taken from specimens or from the literature." pageId="15" pageNumber="43">Table 1</tableCitation>
|
||
;
|
||
<figureCitation id="13082A08FF9DFFCFD3F4FD68CDB46F9C" box="[151,221,693,719]" captionStart="FIGURE 2" captionStartId="5.[151,258,1083,1107]" captionTargetBox="[151,1434,197,1057]" captionTargetId="figure@5.[151,1436,197,1058]" captionTargetPageId="5" captionText="FIGURE 2. Adult male holotype (ZRC 2.6804) of Acanthosaura bintangensis from Bukit Larut, Perak, Peninsular Malaysia. Upper left: in life. Lower left: lateral view of the scalation on the left side of the body. Upper right: lateral view of the head. Lower right: dorsal view." httpUri="https://zenodo.org/record/185823/files/figure.png" pageId="15" pageNumber="43">Fig. 2</figureCitation>
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,
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<figureCitation id="13082A08FF9DFFCFD389FD68CD926F9C" box="[234,251,693,719]" captionStart="FIGURE 5" captionStartId="11.[151,258,1152,1176]" captionTargetBox="[151,1434,194,1128]" captionTargetId="figure@11.[151,1436,194,1128]" captionTargetPageId="11" captionText="FIGURE 5. Adult male holotype (ZRC 2.6805) of Acanthosaura titiwangsaensis from Bishop’s trail Fraser’s Hill, Pahang, Peninsular Malaysia. Upper left: in life. Lower left: lateral view of the scalation on the left side of the body. Upper right: lateral view of the head. Lower right: dorsal view." httpUri="https://zenodo.org/record/185827/files/figure.png" pageId="15" pageNumber="43">5</figureCitation>
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).
|
||
</paragraph>
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</subSubSection>
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</treatment>
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</document> |